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<document id="B33D0F9AFD368B296628599E27D6EB33" ID-CLB-Dataset="31855" ID-DOI="10.11646/zootaxa.4331.1.1" ID-GBIF-Dataset="14c29731-224f-43a4-ae52-f76823177ce5" ID-ISSN="1175-5326" ID-Zenodo-Dep="1009725" ID-ZooBank="FB50725E-4D03-4507-9CAF-EB939C7F548F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1507876567978" checkinUser="plazi" docAuthor="Asami, Takahiro" docDate="2017" docId="03CA87D8FFAAFF8CFF0BF961FAEDF850" docLanguage="en" docName="zootaxa.4331.1.1.pdf" docOrigin="Zootaxa 4331 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Metalycaeus muciferus" docType="treatment" docVersion="10" lastPageNumber="91" masterDocId="FFF3FFA0FFFDFFD6FF9CFFC0FF80FFFF" masterDocTitle="Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus" masterLastPageNumber="124" masterPageNumber="1" pageNumber="88" updateTime="1698469440756" updateUser="ExternalLinkService">
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<mods:title id="3E027ABFB2562A2CC62BBC249EFC12D8">Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus</mods:title>
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<treatment id="03CA87D8FFAAFF8CFF0BF961FAEDF850" ID-DOI="http://doi.org/10.5281/zenodo.6004488" ID-GBIF-Taxon="135435123" ID-Zenodo-Dep="6004488" LSID="urn:lsid:plazi:treatment:03CA87D8FFAAFF8CFF0BF961FAEDF850" httpUri="http://treatment.plazi.org/id/03CA87D8FFAAFF8CFF0BF961FAEDF850" lastPageId="90" lastPageNumber="91" pageId="87" pageNumber="88">
<subSubSection id="C3796545FFAAFF81FF0BF961FE13F922" pageId="87" pageNumber="88" type="nomenclature">
<paragraph id="8BDC36CEFFAAFF81FF0BF961FDE4F944" blockId="87.[151,612,1697,1757]" box="[151,612,1697,1723]" pageId="87" pageNumber="88">
<heading id="D09481A2FFAAFF81FF0BF961FDE4F944" bold="true" box="[151,612,1697,1723]" fontSize="11" level="1" pageId="87" pageNumber="88" reason="1">
<taxonomicName id="4C634D4DFFAAFF81FF0BF961FDE4F944" ID-CoL="7S579" authority="(Heude, 1885)" baseAuthorityName="Heude" baseAuthorityYear="1885" box="[151,612,1697,1723]" class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="87" pageNumber="88" phylum="Mollusca" rank="species" species="muciferus">
<emphasis id="B917EADCFFAAFF81FF0BF961FDE4F944" bold="true" box="[151,612,1697,1723]" pageId="87" pageNumber="88">
<emphasis id="B917EADCFFAAFF81FF0BF961FE2FF944" bold="true" box="[151,431,1697,1723]" italics="true" pageId="87" pageNumber="88">Metalycaeus muciferus</emphasis>
(
<bibRefCitation id="EFF24B3FFFAAFF81FE22F961FDDCF944" author="Heude" box="[446,604,1697,1723]" pageId="87" pageNumber="88" refString="Heude, R. P. M. (1885) Memoires concernant l'histoire naturelle de l'empire chinois par des peres de la Compagnie de Jesus. Notes sur les Mollusques terrestres de la vallee du Fleuve Bleu. Vol. III. Mission Catholique, Chang-Hai, 44 pp. [pp. 89 - 132]" type="book chapter" year="1885">Heude, 1885</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BDC36CEFFAAFF81FF0BF904FE13F922" blockId="87.[151,612,1697,1757]" box="[151,403,1732,1757]" pageId="87" pageNumber="88">
<figureCitation id="13582A4BFFAAFF81FF0BF904FE8EF922" box="[151,270,1732,1757]" captionStart="FIGURE 58" captionStartId="89.[151,250,1695,1717]" captionTargetBox="[151,1436,195,1671]" captionTargetId="figure@89.[151,1436,195,1671]" captionTargetPageId="89" captionText="FIGURE 58. Shells of Metalycaeus muciferus (Heude, 1885). A. syntype. B. 20070714 B, coll. PGB. C. 2011 / 15, coll. HA. D. 2015 / 48, coll. HA. Note that the black &quot; operculum &quot; of Fig. D is the posterior end of a predator fly larva. Scale represents 5 mm. All photos: B. Páll-Gergely." httpUri="https://zenodo.org/record/1009841/files/figure.png" pageId="87" pageNumber="88">FIGUrEs 58</figureCitation>
–59, 60, 61.
</paragraph>
</subSubSection>
<subSubSection id="C3796545FFAAFF8EFF0BF8CCFE82FE62" lastPageId="88" lastPageNumber="89" pageId="87" pageNumber="88" type="reference_group">
<paragraph id="8BDC36CEFFAAFF81FF0BF8CCFD41F8DD" blockId="87.[151,891,1804,2010]" box="[151,705,1804,1826]" pageId="87" pageNumber="88">
<treatmentCitationGroup id="AB7311E0FFAAFF81FF0BF8CCFD41F8DD" box="[151,705,1804,1826]" pageId="87" pageNumber="88">
<taxonomicName id="4C634D4DFFAAFF81FF0BF8CCFD8DF8DD" authority="Heude, 1885: 96" authorityName="Heude" authorityPageNumber="96" authorityYear="1885" box="[151,525,1804,1826]" class="Gastropoda" family="Cyclophoridae" genus="Alycoeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="87" pageNumber="88" phylum="Mollusca" rank="species" species="muciferus">
<emphasis id="B917EADCFFAAFF81FF0BF8CCFEDDF8DD" box="[151,349,1804,1826]" italics="true" pageId="87" pageNumber="88">Alycoeus muciferus</emphasis>
<treatmentCitation id="0AC210DFFFAAFF81FEFFF8CCFD8DF8DD" author="Heude" box="[355,525,1804,1826]" page="96" pageId="87" pageNumber="88" year="1885">
<bibRefCitation id="EFF24B3FFFAAFF81FEFFF8CCFE67F8DD" author="Heude" box="[355,487,1804,1826]" pageId="87" pageNumber="88" refString="Heude, R. P. M. (1885) Memoires concernant l'histoire naturelle de l'empire chinois par des peres de la Compagnie de Jesus. Notes sur les Mollusques terrestres de la vallee du Fleuve Bleu. Vol. III. Mission Catholique, Chang-Hai, 44 pp. [pp. 89 - 132]" type="book chapter" year="1885">Heude, 1885</bibRefCitation>
: 96
</treatmentCitation>
</taxonomicName>
, pl. 24, figs 1, 1a.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDC36CEFFAAFF81FF0BF8EAFD43F8A0" blockId="87.[151,891,1804,2010]" pageId="87" pageNumber="88">
<treatmentCitationGroup id="AB7311E0FFAAFF81FF0BF8EAFD43F8A0" pageId="87" pageNumber="88">
<taxonomicName id="4C634D4DFFAAFF81FF0BF8EAFE7CF8BF" authorityName="Heude" authorityYear="1885" box="[151,508,1834,1856]" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Architaenioglossa" pageId="87" pageNumber="88" phylum="Mollusca" rank="species" species="muciferus" subGenus="Orthalycaeus">
<emphasis id="B917EADCFFAAFF81FF0BF8EAFF73F8BF" box="[151,243,1834,1856]" italics="true" pageId="87" pageNumber="88">Alycaeus</emphasis>
(
<emphasis id="B917EADCFFAAFF81FE9DF8EAFE09F8BF" box="[257,393,1834,1856]" italics="true" pageId="87" pageNumber="88">Orthalycaeus</emphasis>
)
<emphasis id="B917EADCFFAAFF81FE04F8EAFE7CF8BF" box="[408,508,1834,1856]" italics="true" pageId="87" pageNumber="88">muciferus</emphasis>
</taxonomicName>
—
<treatmentCitation id="0AC210DFFFAAFF81FD8AF8EAFC86F8BF" author="Mollendorff" box="[534,774,1834,1856]" page="171" pageId="87" pageNumber="88" year="1886">
<bibRefCitation id="EFF24B3FFFAAFF81FD8AF8EAFD53F8BF" author="Mollendorff" box="[534,723,1834,1856]" pageId="87" pageNumber="88" refString="Mollendorff, O. F. von (1886) Materialen zur Fauna for China. Jahrbucher der Deutschen Malakozoologischen Gesellschaft, 13, 156 - 210." type="journal article" year="1886">Möllendorff, 1886</bibRefCitation>
: 171
</treatmentCitation>
.
<taxonomicName id="4C634D4DFFAAFF81FF0BF889FD9FF8A0" authority="Gredler, 1887: 365" authorityName="Gredler" authorityPageNumber="365" authorityYear="1887" box="[151,543,1865,1887]" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Architaenioglossa" pageId="87" pageNumber="88" phylum="Mollusca" rank="species" species="helicodes">
<emphasis id="B917EADCFFAAFF81FF0BF889FED8F8A0" box="[151,344,1865,1887]" italics="true" pageId="87" pageNumber="88">Alycaeus helicodes</emphasis>
<treatmentCitation id="0AC210DFFFAAFF81FEC2F889FD9FF8A0" author="Gredler" box="[350,543,1865,1887]" page="365" pageId="87" pageNumber="88" year="1887">
<bibRefCitation id="EFF24B3FFFAAFF81FEC2F889FE6DF8A0" author="Gredler" box="[350,493,1865,1887]" pageId="87" pageNumber="88" refString="Gredler, V. (1887) Zur Conchylien-Fauna von China. XIII. Stuck. Jahrbucher der Deutschen Malakozoologischen Gesellschaft, 14, 343 - 373." type="journal article" year="1887">Gredler, 1887</bibRefCitation>
: 365
</treatmentCitation>
</taxonomicName>
,
<taxonomicNameLabel id="A22457A7FFAAFF81FDB6F889FD3DF8A0" box="[554,701,1865,1887]" pageId="87" pageNumber="88">
<emphasis id="B917EADCFFAAFF81FDB6F889FD3DF8A0" bold="true" box="[554,701,1865,1887]" pageId="87" pageNumber="88">new synonym</emphasis>
</taxonomicNameLabel>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDC36CEFFAAFF81FF0BF8A8FD5CF844" blockId="87.[151,891,1804,2010]" pageId="87" pageNumber="88">
<treatmentCitationGroup id="AB7311E0FFAAFF81FF0BF8A8FD5CF844" pageId="87" pageNumber="88">
<taxonomicName id="4C634D4DFFAAFF81FF0BF8A8FD84F881" authorityName="Gredler" authorityYear="1887" box="[151,516,1896,1918]" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Architaenioglossa" pageId="87" pageNumber="88" phylum="Mollusca" rank="species" species="helicodes" subGenus="Chamalycaeus">
<emphasis id="B917EADCFFAAFF81FF0BF8A8FF73F881" box="[151,243,1896,1918]" italics="true" pageId="87" pageNumber="88">Alycaeus</emphasis>
(
<emphasis id="B917EADCFFAAFF81FE9DF8A8FE16F881" box="[257,406,1896,1918]" italics="true" pageId="87" pageNumber="88">Chamalycaeus</emphasis>
)
<emphasis id="B917EADCFFAAFF81FE39F8A8FD84F881" box="[421,516,1896,1918]" italics="true" pageId="87" pageNumber="88">helicodes</emphasis>
</taxonomicName>
—
<treatmentCitation id="0AC210DFFFAAFF81FD82F8A8FCF4F881" author="Kobelt" box="[542,884,1896,1918]" page="148" pageId="87" pageNumber="88" year="1897">
<bibRefCitation id="EFF24B3FFFAAFF81FD82F8A8FCC1F881" author="Kobelt" box="[542,833,1896,1918]" pageId="87" pageNumber="88" refString="Kobelt, W. &amp; Mollendorff, O. von (1897) Catalog der gegenvartig lebend bekannten Pneumonopomen. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft, 29, 73 - 88, 105 - 120, 137 - 152." type="journal article" year="1897">Kobelt &amp; Möllendorff, 1897</bibRefCitation>
: 148
</treatmentCitation>
.
<taxonomicName id="4C634D4DFFAAFF81FF0BF846FD89F863" authorityName="Heude" authorityYear="1885" box="[151,521,1926,1948]" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Architaenioglossa" pageId="87" pageNumber="88" phylum="Mollusca" rank="species" species="muciferus" subGenus="Chamalycaeus">
<emphasis id="B917EADCFFAAFF81FF0BF846FF73F863" box="[151,243,1926,1948]" italics="true" pageId="87" pageNumber="88">Alycaeus</emphasis>
(
<emphasis id="B917EADCFFAAFF81FE9DF846FE16F863" box="[257,406,1926,1948]" italics="true" pageId="87" pageNumber="88">Chamalycaeus</emphasis>
)
<emphasis id="B917EADCFFAAFF81FE39F846FD89F863" box="[421,521,1926,1948]" italics="true" pageId="87" pageNumber="88">muciferus</emphasis>
</taxonomicName>
—
<treatmentCitation id="0AC210DFFFAAFF81FDBEF846FCF8F863" author="Kobelt" box="[546,888,1926,1948]" page="149" pageId="87" pageNumber="88" year="1897">
<bibRefCitation id="EFF24B3FFFAAFF81FDBEF846FCC5F863" author="Kobelt" box="[546,837,1926,1948]" pageId="87" pageNumber="88" refString="Kobelt, W. &amp; Mollendorff, O. von (1897) Catalog der gegenvartig lebend bekannten Pneumonopomen. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft, 29, 73 - 88, 105 - 120, 137 - 152." type="journal article" year="1897">Kobelt &amp; Möllendorff, 1897</bibRefCitation>
: 149
</treatmentCitation>
.
<taxonomicName id="4C634D4DFFAAFF81FF0BF865FD84F844" authorityName="Gredler" authorityYear="1887" box="[151,516,1957,1979]" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Architaenioglossa" pageId="87" pageNumber="88" phylum="Mollusca" rank="species" species="helicodes" subGenus="Chamalycaeus">
<emphasis id="B917EADCFFAAFF81FF0BF865FF73F844" box="[151,243,1957,1979]" italics="true" pageId="87" pageNumber="88">Alycaeus</emphasis>
(
<emphasis id="B917EADCFFAAFF81FE9DF865FE16F844" box="[257,406,1957,1979]" italics="true" pageId="87" pageNumber="88">Chamalycaeus</emphasis>
)
<emphasis id="B917EADCFFAAFF81FE39F865FD84F844" box="[421,516,1957,1979]" italics="true" pageId="87" pageNumber="88">helicodes</emphasis>
</taxonomicName>
—
<treatmentCitation id="0AC210DFFFAAFF81FD83F865FD58F844" author="Kobelt" box="[543,728,1957,1979]" page="356" pageId="87" pageNumber="88" year="1902">
<bibRefCitation id="EFF24B3FFFAAFF81FD83F865FD26F844" author="Kobelt" box="[543,678,1957,1979]" pageId="87" pageNumber="88" refString="Kobelt, W. (1902) Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen. In Verbindung mit der Deutschen Zoologischen Gesellschaft herausgegeben von der Koniglich Preussischen Akademie der Wissenschaften zu Berlin. Mollusca: Cyclophoridae. R. Friedlander und Sohn, Berlin, 662 pp." type="book" year="1902">Kobelt, 1902</bibRefCitation>
: 356
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDC36CEFFAAFF81FF0BF804FD60F825" blockId="87.[151,891,1804,2010]" box="[151,736,1988,2010]" pageId="87" pageNumber="88">
<treatmentCitationGroup id="AB7311E0FFAAFF81FF0BF804FD60F825" box="[151,736,1988,2010]" pageId="87" pageNumber="88">
<taxonomicName id="4C634D4DFFAAFF81FF0BF804FD89F825" authorityName="Heude" authorityYear="1885" box="[151,521,1988,2010]" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Architaenioglossa" pageId="87" pageNumber="88" phylum="Mollusca" rank="species" species="muciferus" subGenus="Chamalycaeus">
<emphasis id="B917EADCFFAAFF81FF0BF804FF73F825" box="[151,243,1988,2010]" italics="true" pageId="87" pageNumber="88">Alycaeus</emphasis>
(
<emphasis id="B917EADCFFAAFF81FE9DF804FE16F825" box="[257,406,1988,2010]" italics="true" pageId="87" pageNumber="88">Chamalycaeus</emphasis>
)
<emphasis id="B917EADCFFAAFF81FE39F804FD89F825" box="[421,521,1988,2010]" italics="true" pageId="87" pageNumber="88">muciferus</emphasis>
</taxonomicName>
—
<treatmentCitation id="0AC210DFFFAAFF81FDBFF804FD5CF825" author="Kobelt" box="[547,732,1988,2010]" page="359" pageId="87" pageNumber="88" year="1902">
<bibRefCitation id="EFF24B3FFFAAFF81FDBFF804FD29F825" author="Kobelt" box="[547,681,1988,2010]" pageId="87" pageNumber="88" refString="Kobelt, W. (1902) Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen. In Verbindung mit der Deutschen Zoologischen Gesellschaft herausgegeben von der Koniglich Preussischen Akademie der Wissenschaften zu Berlin. Mollusca: Cyclophoridae. R. Friedlander und Sohn, Berlin, 662 pp." type="book" year="1902">Kobelt, 1902</bibRefCitation>
: 359
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDC36CEFFA5FF8EFF0BFF58FD55FF14" blockId="88.[151,867,152,235]" pageId="88" pageNumber="89">
<treatmentCitationGroup id="AB7311E0FFA5FF8EFF0BFF58FD55FF14" pageId="88" pageNumber="89">
<taxonomicName id="4C634D4DFFA5FF8EFF0BFF58FE32FF51" box="[151,434,152,174]" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="88" pageNumber="89" phylum="Mollusca" rank="species" species="dolichodirus">
<emphasis id="B917EADCFFA5FF8EFF0BFF58FE32FF51" box="[151,434,152,174]" italics="true" pageId="88" pageNumber="89">Chamalycaeus dolichodirus</emphasis>
</taxonomicName>
(sic!)—
<treatmentCitation id="0AC210DFFFA5FF8EFD9FFF58FD17FF51" author="Yen" box="[515,663,152,174]" page="30" pageId="88" pageNumber="89" year="1939">
<bibRefCitation id="EFF24B3FFFA5FF8EFD9FFF58FDF1FF51" author="Yen" box="[515,625,152,174]" pageId="88" pageNumber="89" refString="Yen, T. - C. (1939) Die Chinesischen Land-und Susswasser-Gastropoden des Natur-Museums Senckenberg. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, Frankfurt am Main, 234 pp." type="book" year="1939">Yen, 1939</bibRefCitation>
: 30
</treatmentCitation>
, pl. 2, fig. 37.
<taxonomicName id="4C634D4DFFA5FF8EFF0BFF76FDDEFF33" box="[151,606,182,204]" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="88" pageNumber="89" phylum="Mollusca" rank="species" species="dolichodirus" subGenus="Chamalycaeus">
<emphasis id="B917EADCFFA5FF8EFF0BFF76FEACFF33" box="[151,300,182,204]" italics="true" pageId="88" pageNumber="89">Chamalycaeus</emphasis>
(
<emphasis id="B917EADCFFA5FF8EFEA7FF76FE50FF33" box="[315,464,182,204]" italics="true" pageId="88" pageNumber="89">Chamalycaeus</emphasis>
)
<emphasis id="B917EADCFFA5FF8EFE42FF76FDDEFF33" box="[478,606,182,204]" italics="true" pageId="88" pageNumber="89">dolichodirus</emphasis>
</taxonomicName>
(sic!)—
<treatmentCitation id="0AC210DFFFA5FF8EFD2CFF76FCDEFF33" author="Zilch" box="[688,862,182,204]" page="142" pageId="88" pageNumber="89" year="1957">
<bibRefCitation id="EFF24B3FFFA5FF8EFD2CFF76FCABFF33" author="Zilch" box="[688,811,182,204]" pageId="88" pageNumber="89" refString="Zilch, A. (1957) Die Typen und Typoide der Natur-Museums Senckenberg, 20: Mollusca, Cyclophoridae, Alycaeinae. Archiv fur Molluskenkunde, 86 (4 / 6), 141 - 150." type="journal article" year="1957">Zilch, 1957</bibRefCitation>
: 142
</treatmentCitation>
.
<taxonomicName id="4C634D4DFFA5FF8EFF0BFF15FE11FF14" authorityName="Gredler" authorityYear="1887" box="[151,401,213,235]" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="88" pageNumber="89" phylum="Mollusca" rank="species" species="helicodes">
<emphasis id="B917EADCFFA5FF8EFF0BFF15FE11FF14" box="[151,401,213,235]" italics="true" pageId="88" pageNumber="89">Chamalycaeus helicodes</emphasis>
</taxonomicName>
—
<treatmentCitation id="0AC210DFFFA5FF8EFE30FF15FDD5FF14" author="Zilch" box="[428,597,213,235]" page="193" pageId="88" pageNumber="89" year="1974">
<bibRefCitation id="EFF24B3FFFA5FF8EFE30FF15FDA3FF14" author="Zilch" box="[428,547,213,235]" pageId="88" pageNumber="89" refString="Zilch, A. (1974) Vinzenz Gredler und die Erforschung der Weichtiere Chinas durch Franziskaner aus Tirol. Archiv fur Molluskenkunde, 104, 171 - 228." type="journal article" year="1974">Zilch, 1974</bibRefCitation>
: 193
</treatmentCitation>
, pl. 7, fig. 4.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDC36CEFFA5FF8EFF0BFED7FE82FE62" blockId="88.[151,1437,279,1997]" pageId="88" pageNumber="89">
<emphasis id="B917EADCFFA5FF8EFF0BFED7FEE4FECF" bold="true" box="[151,356,279,304]" pageId="88" pageNumber="89">Types examined.</emphasis>
7 syNTypEs (MCZ 167230) Of
<taxonomicName id="4C634D4DFFA5FF8EFD7CFED8FCF4FECF" authorityName="Heude" authorityYear="1885" box="[736,884,280,304]" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Architaenioglossa" pageId="88" pageNumber="89" phylum="Mollusca" rank="species" species="muciferus">
<emphasis id="B917EADCFFA5FF8EFD7CFED8FCF4FECF" box="[736,884,280,304]" italics="true" pageId="88" pageNumber="89">A. muciferus</emphasis>
</taxonomicName>
CHINa, 2 paraLECTOTypEs (SMF 192225) Of
<taxonomicName id="4C634D4DFFA5FF8EFA18FED8FE81FEAB" authorityName="Gredler" authorityYear="1887" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="88" pageNumber="89" phylum="Mollusca" rank="species" species="helicodes">
<emphasis id="B917EADCFFA5FF8EFA18FED8FE81FEAB" italics="true" pageId="88" pageNumber="89">A. helicodes</emphasis>
</taxonomicName>
, TCHEN-k'EOU, CHINa, PE-sHaNG, HUNaN, LEG. FUCHs, COLL. GrEdLEr. ACCOrdING TO
<bibRefCitation id="EFF24B3FFFA5FF8EFBE7FEFCFAA9FEAB" author="JOHNsON" box="[1147,1321,316,340]" pageId="88" pageNumber="89" refString="Johnson, R. J. (1973) Heude's Molluscan Types: or, Asian land and fresh water mollusks, mostly from the People's Republic of China, described by P. M. Heude. Special Occasional Publication, Department of Mollusks, Museum of Comparative Zoology, Harvard University, Cambridge, Massachusetts, 111 pp." type="book" year="1973">JOHNsON (1973)</bibRefCitation>
paraTypEs Of
<taxonomicName id="4C634D4DFFA5FF8EFF25FEA0FEA7FE87" authorityName="Heude" authorityYear="1885" box="[185,295,352,376]" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Architaenioglossa" pageId="88" pageNumber="89" phylum="Mollusca" rank="species" species="muciferus">
<emphasis id="B917EADCFFA5FF8EFF25FEA0FEA7FE87" box="[185,295,352,376]" italics="true" pageId="88" pageNumber="89">muciferus</emphasis>
</taxonomicName>
arE aLsO prEsENT IN THE NMNH (INv. NUmbEr: 472340). THEsE wErE NOT ExamINEd by Us. SEE aLsO REmarks.
</paragraph>
</subSubSection>
<subSubSection id="C3796545FFA5FF8EFF5BFE67FACCFADF" pageId="88" pageNumber="89" type="materials_examined">
<paragraph id="8BDC36CEFFA5FF8EFF5BFE67FD7DFDD3" blockId="88.[151,1437,279,1997]" pageId="88" pageNumber="89">
<emphasis id="B917EADCFFA5FF8EFF5BFE67FDA9FE3F" bold="true" box="[199,553,423,448]" pageId="88" pageNumber="89">Museum material examined.</emphasis>
C SMF 39254/1 (fIGUrEd IN
<bibRefCitation id="EFF24B3FFFA5FF8EFC0CFE68FB85FE3F" author="YEN" box="[912,1029,424,448]" pageId="88" pageNumber="89" refString="Yen, T. - C. (1939) Die Chinesischen Land-und Susswasser-Gastropoden des Natur-Museums Senckenberg. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, Frankfurt am Main, 234 pp." type="book" year="1939">YEN 1939</bibRefCitation>
, pL. 2, fIG. 37, as &quot;
<taxonomicName id="4C634D4DFFA5FF8EFB6BFE68FEA4FE1B" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="88" pageNumber="89" phylum="Mollusca" rank="species" species="dolichodirus">
<emphasis id="B917EADCFFA5FF8EFB6BFE68FEA4FE1B" italics="true" pageId="88" pageNumber="89">Chamalycaeus dolichodirus</emphasis>
</taxonomicName>
&quot;), HaNG-yaNG, HUpEI, CHINa, LEG. FULTON, 1894, COLL. O. BOETTGEr; SMF 346399/1 (mIxEd sampLE wITH
<taxonomicName id="4C634D4DFFA5FF8EFF0BFE31FE9FFDF8" baseAuthorityName="Mollendorff" baseAuthorityYear="1886" box="[151,287,496,520]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="88" pageNumber="89" phylum="Mollusca" rank="species" species="cristatus">
<emphasis id="B917EADCFFA5FF8EFF0BFE31FE9FFDF8" box="[151,287,496,520]" italics="true" pageId="88" pageNumber="89">D. cristatus</emphasis>
</taxonomicName>
: SMF 39250), PaTUNG, HUpEI, CHINa, COLL. MöLLENdOrff; NHMUK 20150487/2 (TypICaL
<taxonomicName id="4C634D4DFFA5FF8EFAB7FE30FA15FDF7" authorityName="Gredler" authorityYear="1887" box="[1323,1429,496,520]" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="88" pageNumber="89" phylum="Mollusca" rank="species" species="helicodes">
<emphasis id="B917EADCFFA5FF8EFAB7FE30FA15FDF7" box="[1323,1429,496,520]" italics="true" pageId="88" pageNumber="89">helicodes</emphasis>
</taxonomicName>
, LabELLEd as
<taxonomicName id="4C634D4DFFA5FF8EFE89FDD4FE22FDD3" box="[277,418,532,556]" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="88" pageNumber="89" phylum="Mollusca" rank="species" species="dolichodirus">
<emphasis id="B917EADCFFA5FF8EFE89FDD4FE22FDD3" box="[277,418,532,556]" italics="true" pageId="88" pageNumber="89">dolichodirus</emphasis>
</taxonomicName>
), CHaNGyaNG, E.R. SykEs COLL.
</paragraph>
<paragraph id="8BDC36CEFFA5FF8EFF5BFDF7FACCFADF" blockId="88.[151,1437,279,1997]" pageId="88" pageNumber="89">
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material examined.
</emphasis>
HA/2,
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,
<collectingRegion id="49A7F82CFFA5FF8EFC99FDF8FC05FDAF" box="[773,901,568,592]" country="China" name="Chongqing" pageId="88" pageNumber="89">CHONGqING</collectingRegion>
, CHENGkOU XIaN, DONGaN XIaNG, apprOx.
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DONGaN, arOUNd GUaNyINyaN,
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,
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, LEG. A. HUNyadI, 0
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HA/1,
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,
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, CHENGkOU XIaN, HEyU XIaNG,
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HEyU CUN, vICINITy
<location id="8EBC6015FFA5FF8EFB31FDBFFB44FD67" LSID="urn:lsid:plazi:treatment:03CA87D8FFAAFF8CFF0BF961FAEDF850:8EBC6015FFA5FF8EFB31FDBFFB44FD67" box="[1197,1220,639,664]" country="China" latitude="31.904766" longLatPrecision="1" longitude="108.94012" municipality="NOrTH Of" name="Of" pageId="88" pageNumber="89" stateProvince="CHONGqING">Of</location>
CUOjIkOU,
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,
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, LEG. A. HUNyadI,
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12
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;
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HA/2,
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,
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, CHENGkOU XIaN, LONGTIaN XIaNG, NOrTHErN EdGE
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YOUfaNG,
<geoCoordinate id="EE575009FFA5FF8EFD0AFD07FCA6FD1F" box="[662,806,711,736]" direction="north" orientation="latitude" pageId="88" pageNumber="89" precision="1" value="32.020084">32°01.205'N</geoCoordinate>
,
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, LEG. A. HUNyadI,
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HA/4,
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<collectingCountry id="F374765EFFA5FF8EFF62FD2CFEC4FCFB" box="[254,324,748,772]" name="China" pageId="88" pageNumber="89">CHINa</collectingCountry>
,
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, CHENGdU
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, DUjIaNGyaN
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, QINGCHENGHOUsHaN, UNdEr BaIzHaNG BrIdGE,
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,
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,
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, LEG. A. HUNyadI &amp; M. SzEkErEs, 0
<collectingDate id="EF99E9E6FFA5FF8EFCC5FCCFFC64FCD7" box="[857,996,783,808]" pageId="88" pageNumber="89" value="2015-06-05">
<date id="FFDD100EFFA5FF8EFCC5FCCFFC64FCD7" box="[857,996,783,808]" pageId="88" pageNumber="89" value="2015-06-05">
5
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</date>
</collectingDate>
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,
<materialsCitation id="3B0B3C93FFA5FF8EFC73FCD0FB75FC8F" ID-GBIF-Occurrence="1671715071" collectingDate="2007-07-14" collectionCode="COLL, SHI, ZHEN" collectorName="Y. NakaHara &amp; JUTTING" country="China" elevation="821" latitude="29.578857" location="Ex" longLatPrecision="1" longitude="103.38874" pageId="88" pageNumber="89" specimenCode="HNHM 99729" specimenCount="1" stateProvince="SICHUaN">
<specimenCode id="DBC59EB5FFA5FF8EFC73FCD0FB16FCD7" box="[1007,1174,784,808]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="88" pageNumber="89">HNHM 99729</specimenCode>
/1 (fIGUrEd sHELL); PGB/ 1 (
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. K. OHara),
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, EmEIsHaN
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, EsHaN
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, EmEIsHaN, BaILONGdONG,
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, LEG.
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, K. OHara, K. OkUbO &amp;
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OTaNI,
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;
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JUO/1, OK/1,
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, HUaNGpING XIaN, FEIyUNdaxIaGU,
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, LEG. K. OHara, K. OkUbO &amp;
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OTaNI, 0
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;
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HA/2,
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<collectingCountry id="F374765EFFA5FF8EFC80FC60FCE2FC47" box="[796,866,928,952]" name="China" pageId="88" pageNumber="89">CHINa</collectingCountry>
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, WENsHaN ZHUaNGzU MIaOzU ZIzHIzHOU, XICHOU XIaN, XINGjIE
<collectionCode id="ED72AE0BFFA5FF8EFE0FFC05FE50FC23" box="[403,464,965,988]" pageId="88" pageNumber="89">ZHEN</collectionCode>
, DadONG
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TOwards BaTaNG, LaOLONG,
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, LEG. A. HUNyadI,
<collectingDate id="EF99E9E6FFA5FF8EFE8DFC28FE3CFBFF" box="[273,444,1000,1024]" pageId="88" pageNumber="89" value="2011-03-22">
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;
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OK/2,
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<collectingCountry id="F374765EFFA5FF8EFD58FC28FC8AFBFF" box="[708,778,1000,1024]" name="China" pageId="88" pageNumber="89">CHINa</collectingCountry>
,
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, NapO XIaN,
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XIaNG, NaCHI,
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, LEG. K. OHara, K. OkUbO &amp;
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OTaNI,
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</collectingDate>
; HA/8, 3 jUvENILE/brOkEN sHELLs
</materialsCitation>
,
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/1,
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<collectingCountry id="F374765EFFA5FF8EFDBDFBF0FDE7FBB7" box="[545,615,1072,1096]" name="China" pageId="88" pageNumber="89">CHINa</collectingCountry>
,
<collectingRegion id="49A7F82CFFA5FF8EFDE6FBF0FD5EFBB7" box="[634,734,1072,1096]" country="China" name="Guangxi" pageId="88" pageNumber="89">GUaNGxI</collectingRegion>
</materialsCitation>
,
<materialsCitation id="3B0B3C93FFA5FF8EFD6DFBF0FC55FB93" ID-GBIF-Occurrence="1671715077" collectingDate="2013-09-11" collectionCode="HECHI, SHI" country="China" elevation="630" latitude="25.069384" location="Of" longLatPrecision="1" longitude="107.602936" pageId="88" pageNumber="89" specimenCount="1" stateProvince="GUaNGxI">
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<collectionCode id="ED72AE0BFFA5FF8EFCDFFBF0FCEBFBB7" box="[835,875,1072,1096]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14032" name="Shihezi Agricultural College" pageId="88" pageNumber="89">SHI</collectionCode>
, NaNdaN XIaN, CLIffs SE
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EN CUN,
<elevation id="004ED1FDFFA5FF8EFADAFBEFFA17FBB8" box="[1350,1431,1071,1096]" metricMagnitude="2" metricUnit="m" metricValue="6.3" pageId="88" pageNumber="89" unit="m" value="630.0">
<quantity id="4C9B9B2BFFA5FF8EFADAFBEFFA17FBB8" box="[1350,1431,1071,1096]" metricMagnitude="2" metricUnit="m" metricValue="6.3" pageId="88" pageNumber="89" unit="m" value="630.0">630 m</quantity>
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,
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,
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, LEG. A. HUNyadI &amp; M. SzEkErEs,
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<date id="FFDD100EFFA5FF8EFCDBFB94FC55FB93" box="[839,981,1108,1132]" pageId="88" pageNumber="89" value="2013-09-11">11 SEp 2013</date>
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;
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PGB/1 (
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. K. OkUbO),
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<collectingCountry id="F374765EFFA5FF8EFF0BFBB8FF5DFB6F" box="[151,221,1144,1168]" name="China" pageId="88" pageNumber="89">CHINa</collectingCountry>
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, NapO XIaN, LONGHE XIaNG, INNEr LONGHE,
<elevation id="004ED1FDFFA5FF8EFCACFBB8FCF9FB70" box="[816,889,1144,1168]" metricMagnitude="2" metricUnit="m" metricValue="9.9" pageId="88" pageNumber="89" unit="m" value="990.0">
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,
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,
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, LEG. K. OHara, K. OkUbO &amp;
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OTaNI,
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</collectingDate>
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;
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HE/1,
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11-141
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, km 105.5 ON rOad 4C, bETwEEN YêN
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aNd ĐồNG VăN (NE Of
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TOwN),
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, LEG
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<collectingCountry id="F374765EFFA5FF8EFBC6FB00FBFFFB27" box="[1114,1151,1216,1240]" name="Switzerland" pageId="88" pageNumber="89">CH</collectingCountry>
. HEmmEN,
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<date id="FFDD100EFFA5FF8EFA9DFB00FA17FB27" box="[1281,1431,1216,1240]" pageId="88" pageNumber="89" value="2011-03-21">21 Mar 2011</date>
</collectingDate>
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;
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JUO/4, PGB/1 (
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<collectionCode id="ED72AE0BFFA5FF8EFEE2FB24FE2DFB03" box="[382,429,1252,1276]" pageId="88" pageNumber="89">COLL</collectionCode>
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<collectorName id="26965318FFA5FF8EFE21FB25FE6AFB03" box="[445,490,1253,1276]" name="JUTTING" pageId="88" pageNumber="89">J.U.</collectorName>
OTaNI),
<emphasis id="B917EADCFFA5FF8EFDCFFB24FD52FB02" bold="true" box="[595,722,1252,1277]" pageId="88" pageNumber="89">
<date id="FFDD100EFFA5FF8EFDCFFB24FD42FB02" box="[595,706,1252,1277]" pageId="88" pageNumber="89" value="1113-08-20">20081113</date>
B
</emphasis>
<collectingCountry id="F374765EFFA5FF8EFD7CFB25FCC3FB03" box="[736,835,1253,1276]" name="Vietnam" pageId="88" pageNumber="89">VIETNam</collectingCountry>
,
<collectingRegion id="49A7F82CFFA5FF8EFCCFFB25FBB9FB03" box="[851,1081,1252,1276]" country="Vietnam" name="Ha Giang" pageId="88" pageNumber="89">Ha GIaNG PrOvINCE</collectingRegion>
, VI
<collectionCode id="ED72AE0BFFA5FF8EFBEDFB24FB3CFB03" box="[1137,1212,1252,1276]" pageId="88" pageNumber="89">XUGEN</collectionCode>
DIsTrICT
</materialsCitation>
,
<materialsCitation id="3B0B3C93FFA5FF8EFAADFB24FAC9FADF" ID-GBIF-Occurrence="1671715112" collectingDate="2008-11-13" collectionCode="HUNG" collectorName="JUTTING" country="Vietnam" elevation="216" latitude="22.867733" location="Nam" longLatPrecision="1" longitude="105.06973" municipality="Ba" pageId="88" pageNumber="89" specimenCount="1" stateProvince="Ha GIaNG PrOvINCE">
<collectionCode id="ED72AE0BFFA5FF8EFAADFB24FAEFFB03" box="[1329,1391,1252,1276]" pageId="88" pageNumber="89">HUNG</collectionCode>
<collectingMunicipality id="6BB8ACB4FFA5FF8EFAE1FB25FA1CFB03" box="[1405,1436,1253,1276]" pageId="88" pageNumber="89">Ba</collectingMunicipality>
COmmUNE,
<location id="8EBC6015FFA5FF8EFE86FAC8FED0FAE0" LSID="urn:lsid:plazi:treatment:03CA87D8FFAAFF8CFF0BF961FAEDF850:8EBC6015FFA5FF8EFE86FAC8FED0FAE0" box="[282,336,1288,1311]" country="Vietnam" latitude="22.867733" longLatPrecision="1" longitude="105.06973" municipality="Ba" name="Nam" pageId="88" pageNumber="89" stateProvince="Ha GIaNG PrOvINCE">Nam</location>
PIa VILLaGE,
<elevation id="004ED1FDFFA5FF8EFE7CFAC7FDA9FAE0" box="[480,553,1287,1312]" metricMagnitude="2" metricUnit="m" metricValue="2.16" pageId="88" pageNumber="89" unit="m" value="216.0">
<quantity id="4C9B9B2BFFA5FF8EFE7CFAC7FDA9FAE0" box="[480,553,1287,1312]" metricMagnitude="2" metricUnit="m" metricValue="2.16" pageId="88" pageNumber="89" unit="m" value="216.0">216 m</quantity>
</elevation>
,
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,
<geoCoordinate id="EE575009FFA5FF8EFD4DFAC7FCECFADF" box="[721,876,1287,1312]" direction="east" orientation="longitude" pageId="88" pageNumber="89" precision="1" value="105.06973">105°04.184'E</geoCoordinate>
, LEG. K. OHara &amp;
<collectorName id="26965318FFA5FF8EFBAEFAC8FBDFFAE0" box="[1074,1119,1288,1311]" name="JUTTING" pageId="88" pageNumber="89">J.U.</collectorName>
OTaNI,
<collectingDate id="EF99E9E6FFA5FF8EFB2EFAC8FAC9FADF" box="[1202,1353,1288,1312]" pageId="88" pageNumber="89" value="2008-11-13">
<date id="FFDD100EFFA5FF8EFB2EFAC8FAC9FADF" box="[1202,1353,1288,1312]" pageId="88" pageNumber="89" value="2008-11-13">13 NOv 2008</date>
</collectingDate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3796545FFA5FF8EFF5BFAECFE60FA97" pageId="88" pageNumber="89" type="diagnosis">
<paragraph id="8BDC36CEFFA5FF8EFF5BFAECFE60FA97" blockId="88.[151,1437,279,1997]" pageId="88" pageNumber="89">
<emphasis id="B917EADCFFA5FF8EFF5BFAECFEC1FABA" bold="true" box="[199,321,1324,1349]" pageId="88" pageNumber="89">Diagnosis.</emphasis>
A smaLL TO mEdIUm sIzEd spECIEs wITH vEry sHOrT R2 aNd R3, R2 HavING TypICaL
<taxonomicName id="4C634D4DFFA5FF8EFB49FAECFAE2FABB" authorityName="Pilsbry" authorityYear="1900" box="[1237,1378,1324,1348]" class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="88" pageNumber="89" phylum="Mollusca" rank="genus">
<emphasis id="B917EADCFFA5FF8EFB49FAECFAE2FABB" box="[1237,1378,1324,1348]" italics="true" pageId="88" pageNumber="89">Metalycaeus</emphasis>
</taxonomicName>
-
<typeStatus id="54D8886CFFA5FF8EFAF4FAEDFA1CFABA" box="[1384,1436,1325,1349]" pageId="88" pageNumber="89">TypE</typeStatus>
rIbs aNd R3 LaCkING swELLING.
</paragraph>
</subSubSection>
<subSubSection id="C3796545FFA5FF8EFF5BFAB4FB33F87B" pageId="88" pageNumber="89" type="description">
<paragraph id="8BDC36CEFFA5FF8EFF5BFAB4FB88F92F" blockId="88.[151,1437,279,1997]" pageId="88" pageNumber="89">
<emphasis id="B917EADCFFA5FF8EFF5BFAB4FDCAFA72" bold="true" box="[199,586,1395,1421]" pageId="88" pageNumber="89">
Description of typical
<taxonomicName id="4C634D4DFFA5FF8EFE52FAB3FDC2FA73" authorityName="Heude" authorityYear="1885" box="[462,578,1395,1420]" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Architaenioglossa" pageId="88" pageNumber="89" phylum="Mollusca" rank="species" species="muciferus">
<emphasis id="B917EADCFFA5FF8EFE52FAB3FDC2FA73" bold="true" box="[462,578,1395,1420]" italics="true" pageId="88" pageNumber="89">muciferus</emphasis>
</taxonomicName>
.
</emphasis>
SHELL (
<figureCitation id="13582A4BFFA5FF8EFD0BFAB4FC9BFA72" box="[663,795,1396,1421]" captionStart="FIGURE 58" captionStartId="89.[151,250,1695,1717]" captionTargetBox="[151,1436,195,1671]" captionTargetId="figure@89.[151,1436,195,1671]" captionTargetPageId="89" captionText="FIGURE 58. Shells of Metalycaeus muciferus (Heude, 1885). A. syntype. B. 20070714 B, coll. PGB. C. 2011 / 15, coll. HA. D. 2015 / 48, coll. HA. Note that the black &quot; operculum &quot; of Fig. D is the posterior end of a predator fly larva. Scale represents 5 mm. All photos: B. Páll-Gergely." httpUri="https://zenodo.org/record/1009841/files/figure.png" pageId="88" pageNumber="89">FIGUrE 58A</figureCitation>
) wHITE; sHELL OUTLINE sLIGHTLy OvaL frOm dOrsaL vIEw, spIrE sLIGHTLy ELEvaTEd, EdGE Of bOdy wHOrL rOUNdEd; prOTOCONCH (
<figureCitation id="13582A4BFFA5FF8EFCD9FA58FC78FA4F" box="[837,1016,1431,1456]" captionStart="FIGURE 61" captionStartId="92.[151,250,1270,1292]" captionTargetBox="[178,1422,193,1266]" captionTargetId="figure@92.[178,1422,193,1266]" captionTargetPageId="92" captionText="FIGURE 61. Protoconchs of Metalycaeus muciferus (Heude, 1885). A – B. 2013 / 7. C – D. syntype (MCZ 167230)." httpUri="https://zenodo.org/record/1009845/files/figure.png" pageId="88" pageNumber="89">FIGUrEs 61C–D</figureCitation>
) CONspICUOUsLy ELEvaTEd frOm dOrsaL sUrfaCE, CONsIsTING Of 1.75 wHOrLs (N = 2), raTHEr maTTE, ITs ENd wITH INCONspICUOUs spIraL sTrIaTIONs; R1 CONsIsTs Of 1.5–1.75 wHOrLs, wITH IrrEGULar, wIdELy-spaCEd rIbs aNd fINE spIraL sCULpTUrE; rIbs sLIGHTLy bECOmING mOrE wIdELyspaCEd TOwards ENd Of R1; TraNsITION bETwEEN R1 aNd R2 NOT CONspICUOUs; R2 wITH dENsE, sHarp, ELEvaTEd rIbs (
<figureCitation id="13582A4BFFA5FF8EFF01F9E8FECCF9BF" box="[157,332,1575,1600]" captionStart="FIGURE 60" captionStartId="91.[151,250,1942,1964]" captionTargetBox="[151,1436,215,1888]" captionTargetId="figure@91.[151,1436,215,1888]" captionTargetPageId="91" captionText="FIGURE 60. R 2 ribs of Metalycaeus muciferus (Heude, 1885). A – D. 2015 / 48. E – F. MCZ 167230. Arrows show the edge of the overfolding anterior crust. All images: B. Páll-Gergely." httpUri="https://zenodo.org/record/1009843/files/figure.png" pageId="88" pageNumber="89">FIGUrEs 60E–F</figureCitation>
); CrOss sECTION Of rIbs Of TypICaL
<taxonomicName id="4C634D4DFFA5FF8EFD20F9E8FCAAF9BF" authorityName="Heude" authorityYear="1885" box="[700,810,1576,1600]" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Architaenioglossa" pageId="88" pageNumber="89" phylum="Mollusca" rank="species" species="muciferus">
<emphasis id="B917EADCFFA5FF8EFD20F9E8FCAAF9BF" box="[700,810,1576,1600]" italics="true" pageId="88" pageNumber="89">muciferus</emphasis>
</taxonomicName>
COULd NOT bE ExamINEd (bUT sEE rEmarks); LENGTH Of R2 aNd R3 vEry sLIGHTLy mOrE THaN a qUarTEr wHOrL COmbINEd (apprOxImaTELy 100 dEGrEEs); R2 LONGEr THaN R3; TraNsITION bETwEEN R2 aNd R3 raTHEr smOOTH, NO CONsTrICTION vIsIbLE; R3 wITH fEw wIdELy-spaCEd wrINkLEs, apErTUrE aLmOsT ExaCTLy rOUNdEd; OUTEr LIp vEry HIGH aNd sHarp, sIGNIfICaNTLy ExpaNdEd aNd sLIGHTLy rEfLECTEd; INNEr LIp sLIGHTLy THICkENEd, NOT CONspICUOUs; bOUNdary bETwEEN TwO LIps HardLy vIsIbLE.
</paragraph>
<paragraph id="8BDC36CEFFA5FF8EFF5BF91CFA1CF90A" blockId="88.[151,1437,279,1997]" box="[199,1436,1756,1781]" pageId="88" pageNumber="89">
<emphasis id="B917EADCFFA5FF8EFF5BF91CFE5DF90A" bold="true" box="[199,477,1756,1781]" pageId="88" pageNumber="89">Measurements (in mm).</emphasis>
D = 3.3–5.3, H = 2.1–3.4 (N = 18, spECImENs frOm dIffErENT LOCaLITIEs). SEE aLsO
<tableCitation id="C6E10375FFA5FF8EFADAF91CFA18F90A" box="[1350,1432,1756,1781]" captionStart="TABLE 5" captionStartId="92.[151,239,1332,1355]" captionText="TABLE 5. Variability of Metalycaeus muciferus populations." pageId="88" pageNumber="89">TabLE 5</tableCitation>
.
</paragraph>
<paragraph id="8BDC36CEFFA5FF8EFF5BF93FFB33F87B" blockId="88.[151,1437,279,1997]" pageId="88" pageNumber="89">
<emphasis id="B917EADCFFA5FF8EFF5BF93FFED6F8E7" bold="true" box="[199,342,1791,1816]" pageId="88" pageNumber="89">Operculum.</emphasis>
THE OUTEr sUrfaCE Of THE OpErCULUm Of THE
<typeStatus id="54D8886CFFA5FF8EFCAFF8C0FCE3F8E7" box="[819,867,1792,1816]" pageId="88" pageNumber="89">TypE</typeStatus>
sErIEs Of
<taxonomicName id="4C634D4DFFA5FF8EFC53F8C0FBBDF8E7" authorityName="Heude" authorityYear="1885" box="[975,1085,1792,1816]" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Architaenioglossa" pageId="88" pageNumber="89" phylum="Mollusca" rank="species" species="muciferus">
<emphasis id="B917EADCFFA5FF8EFC53F8C0FBBDF8E7" box="[975,1085,1792,1816]" italics="true" pageId="88" pageNumber="89">muciferus</emphasis>
</taxonomicName>
aNd
<taxonomicName id="4C634D4DFFA5FF8EFBEFF8C0FB5DF8E7" authorityName="Gredler" authorityYear="1887" box="[1139,1245,1792,1816]" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="88" pageNumber="89" phylum="Mollusca" rank="species" species="helicodes">
<emphasis id="B917EADCFFA5FF8EFBEFF8C0FB5DF8E7" box="[1139,1245,1792,1816]" italics="true" pageId="88" pageNumber="89">helicodes</emphasis>
</taxonomicName>
wErE COvErEd by mUd. UNdEr THE mUd, HOwEvEr, aN ELEvaTEd, mULTIspIraL, wrINkLEd LamELLa was vIsIbLE. BOTH sIdEs Of THE OpErCULUm Of sampLE
<date id="FFDD100EFFA5FF8EFE96F888FEFAF89F" box="[266,378,1864,1888]" pageId="88" pageNumber="89" value="1113-08-20">20081113</date>
B wErE ExamINEd. THE OUTEr sUrfaCE sHOwEd mULTIspIraL CHaraCTEr wITHOUT ELEvaTEd LamELLa; THE INNEr sIdE Had a vEry smaLL, LOw
<collectingRegion id="49A7F82CFFA5FF8EFD9FF8ACFDCFF87B" box="[515,591,1900,1924]" country="Sri Lanka" name="Central" pageId="88" pageNumber="89">CENTraL</collectingRegion>
NIppLE sITTING ON THE TOp Of a GradUaLLy ELEvaTEd &quot;HILL&quot;.
</paragraph>
</subSubSection>
<subSubSection id="C3796545FFA5FF8EFF5BF84FFB33F833" pageId="88" pageNumber="89" type="materials_examined">
<paragraph id="8BDC36CEFFA5FF8EFF5BF84FFB33F833" blockId="88.[151,1437,279,1997]" pageId="88" pageNumber="89">
<emphasis id="B917EADCFFA5FF8EFF5BF84FFE63F857" bold="true" box="[199,483,1935,1960]" pageId="88" pageNumber="89">Variation in specimens.</emphasis>
<taxonomicName id="4C634D4DFFA5FF8EFE70F850FC8FF858" authority="Is" authorityName="Is" box="[492,783,1936,1960]" class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="88" pageNumber="89" phylum="Mollusca" rank="species" species="muciferus">
<emphasis id="B917EADCFFA5FF8EFE70F850FD72F857" box="[492,754,1936,1960]" italics="true" pageId="88" pageNumber="89">Metalycaeus muciferus</emphasis>
Is
</taxonomicName>
a vEry varIabLE spECIEs IN sHELL sIzE, sHELL sCULpTUrE aNd COLOUr.
<tableCitation id="C6E10375FFA5FF8EFF77F874FEBEF832" box="[235,318,1972,1997]" captionStart="TABLE 5" captionStartId="92.[151,239,1332,1355]" captionText="TABLE 5. Variability of Metalycaeus muciferus populations." pageId="88" pageNumber="89">TabLE 5</tableCitation>
COmpILEs
<collectionCode id="ED72AE0BFFA5FF8EFE2DF875FE53F833" box="[433,467,1973,1996]" pageId="88" pageNumber="89">THE</collectionCode>
varIabLE TraITs aNd THEIr CHaraCTEr sTaTEs IN ExamINEd pOpULaTIONs.
</paragraph>
</subSubSection>
<caption id="DF1C6646FFA4FF8FFF0BF95FFE33F90D" httpUri="https://zenodo.org/record/1009841/files/figure.png" pageId="89" pageNumber="90" targetBox="[151,1436,195,1671]" targetPageId="89">
<paragraph id="8BDC36CEFFA4FF8FFF0BF95FFE33F90D" blockId="89.[151,1437,1695,1778]" pageId="89" pageNumber="90">
<emphasis id="B917EADCFFA4FF8FFF0BF95FFEA0F94A" bold="true" box="[151,288,1695,1717]" pageId="89" pageNumber="90">FIGURE 58.</emphasis>
Shells of
<taxonomicName id="4C634D4DFFA4FF8FFE19F95FFC8CF94A" authority="(Heude, 1885)" authorityName="Heude" authorityYear="1885" baseAuthorityName="Heude" baseAuthorityYear="1885" box="[389,780,1695,1717]" class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="89" pageNumber="90" phylum="Mollusca" rank="species" species="muciferus">
<emphasis id="B917EADCFFA4FF8FFE19F95FFDF0F94A" box="[389,624,1695,1717]" italics="true" pageId="89" pageNumber="90">Metalycaeus muciferus</emphasis>
(Heude, 1885)
</taxonomicName>
.
<emphasis id="B917EADCFFA4FF8FFC89F95FFCAEF94A" bold="true" box="[789,814,1695,1717]" pageId="89" pageNumber="90">A.</emphasis>
syntype.
<emphasis id="B917EADCFFA4FF8FFC0CF95FFC27F94A" bold="true" box="[912,935,1695,1717]" pageId="89" pageNumber="90">B.</emphasis>
20070714B, coll. PGB.
<emphasis id="B917EADCFFA4FF8FFB3DF95FFB3AF94A" bold="true" box="[1185,1210,1695,1717]" pageId="89" pageNumber="90">C.</emphasis>
2011/15, coll. HA.
<emphasis id="B917EADCFFA4FF8FFA1FF95FFA1CF94A" bold="true" box="[1411,1436,1695,1717]" pageId="89" pageNumber="90">D.</emphasis>
2015/48, coll. HA. Note that the black &quot;operculum&quot; of Fig. D is the posterior end of a predator fly larva. Scale represents 5 mm. All photos: B. Páll-Gergely.
</paragraph>
</caption>
<subSubSection id="C3796545FFA4FF8FFF5BF8D9FB83F886" pageId="89" pageNumber="90" type="diagnosis">
<paragraph id="8BDC36CEFFA4FF8FFF5BF8D9FB83F886" blockId="89.[151,1437,1817,1986]" pageId="89" pageNumber="90">
<emphasis id="B917EADCFFA4FF8FFF5BF8D9FE51F8CD" bold="true" box="[199,465,1817,1842]" pageId="89" pageNumber="90">Differential diagnosis.</emphasis>
<taxonomicName id="4C634D4DFFA4FF8FFE46F8DAFD5FF8CD" baseAuthorityName="Heude" baseAuthorityYear="1885" box="[474,735,1818,1842]" class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="89" pageNumber="90" phylum="Mollusca" rank="species" species="muciferus">
<emphasis id="B917EADCFFA4FF8FFE46F8DAFD5FF8CD" box="[474,735,1818,1842]" italics="true" pageId="89" pageNumber="90">Metalycaeus muciferus</emphasis>
</taxonomicName>
CaN bE dIsTINGUIsHEd frOm OTHEr
<taxonomicName id="4C634D4DFFA4FF8FFBFEF8DAFB6FF8CD" authorityName="Pilsbry" authorityYear="1900" box="[1122,1263,1818,1842]" class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="89" pageNumber="90" phylum="Mollusca" rank="genus">
<emphasis id="B917EADCFFA4FF8FFBFEF8DAFB6FF8CD" box="[1122,1263,1818,1842]" italics="true" pageId="89" pageNumber="90">Metalycaeus</emphasis>
</taxonomicName>
spECIEs ON THE basIs Of THE sHOrT R2, aNd THE aLsO sHOrT R3, wHICH LaCks a swELLING IN ITs mIddLE. SEE UNdEr
<taxonomicName id="4C634D4DFFA4FF8FFB3DF8FEFACFF8A9" authority="Asami, 2017" authorityName="Asami" authorityYear="2017" box="[1185,1359,1854,1878]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="89" pageNumber="90" phylum="Mollusca" rank="species" species="bison" status="sp. nov.">
<emphasis id="B917EADCFFA4FF8FFB3DF8FEFACFF8A9" box="[1185,1359,1854,1878]" italics="true" pageId="89" pageNumber="90">Dicharax bison</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22457A7FFA4FF8FFAC9F8FDFA15F8A9" box="[1365,1429,1853,1878]" pageId="89" pageNumber="90" rank="species">
<emphasis id="B917EADCFFA4FF8FFAC9F8FDFA15F8A9" bold="true" box="[1365,1429,1853,1878]" pageId="89" pageNumber="90">n. sp.</emphasis>
</taxonomicNameLabel>
,
<taxonomicName id="4C634D4DFFA4FF8FFF0BF8A2FEF6F885" box="[151,374,1890,1914]" class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="89" pageNumber="90" phylum="Mollusca" rank="species" species="heudei">
<emphasis id="B917EADCFFA4FF8FFF0BF8A2FEF6F885" box="[151,374,1890,1914]" italics="true" pageId="89" pageNumber="90">Metalycaeus heudei</emphasis>
</taxonomicName>
,
<taxonomicName id="4C634D4DFFA4FF8FFE18F8A2FD90F885" authority="Asami, 2017" authorityName="Asami" authorityYear="2017" box="[388,528,1890,1914]" class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="89" pageNumber="90" phylum="Mollusca" rank="species" species="laosensis" status="sp. nov.">
<emphasis id="B917EADCFFA4FF8FFE18F8A2FD90F885" box="[388,528,1890,1914]" italics="true" pageId="89" pageNumber="90">M. laosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22457A7FFA4FF8FFD8BF8A1FDD5F885" box="[535,597,1889,1914]" pageId="89" pageNumber="90" rank="species">
<emphasis id="B917EADCFFA4FF8FFD8BF8A1FDD5F885" bold="true" box="[535,597,1889,1914]" pageId="89" pageNumber="90">n. sp.</emphasis>
</taxonomicNameLabel>
,
<taxonomicName id="4C634D4DFFA4FF8FFDF8F8A2FD54F885" authority="Asami, 2017" authorityName="Asami" authorityYear="2017" box="[612,724,1890,1914]" class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="89" pageNumber="90" phylum="Mollusca" rank="species" species="oharai" status="sp. nov.">
<emphasis id="B917EADCFFA4FF8FFDF8F8A2FD54F885" box="[612,724,1890,1914]" italics="true" pageId="89" pageNumber="90">M. oharai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22457A7FFA4FF8FFD47F8A1FC9AF885" box="[731,794,1889,1914]" pageId="89" pageNumber="90" rank="species">
<emphasis id="B917EADCFFA4FF8FFD47F8A1FC9AF885" bold="true" box="[731,794,1889,1914]" pageId="89" pageNumber="90">n. sp.</emphasis>
</taxonomicNameLabel>
aNd
<taxonomicName id="4C634D4DFFA4FF8FFCCCF8A2FC7CF885" baseAuthorityName="Moellendorff" baseAuthorityYear="1882" box="[848,1020,1890,1914]" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="89" pageNumber="90" phylum="Mollusca" rank="species" species="latecostatus">
<emphasis id="B917EADCFFA4FF8FFCCCF8A2FC7CF885" box="[848,1020,1890,1914]" italics="true" pageId="89" pageNumber="90">M. latecostatus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3796545FFA4FF8FFF5BF845FB1BF83D" pageId="89" pageNumber="90" type="distribution">
<paragraph id="8BDC36CEFFA4FF8FFF5BF845FB1BF83D" blockId="89.[151,1437,1817,1986]" pageId="89" pageNumber="90">
<emphasis id="B917EADCFFA4FF8FFF5BF845FEE0F861" bold="true" box="[199,352,1925,1950]" pageId="89" pageNumber="90">Distribution.</emphasis>
SHELLs assIGNEd TO THIs spECIEs wErE COLLECTEd frOm mULTIpLE CHINEsE prOvINCEs (
<collectingRegion id="49A7F82CFFA4FF8FFAA0F846FA19F861" box="[1340,1433,1926,1950]" country="China" name="Yunnan" pageId="89" pageNumber="90">YUNNaN</collectingRegion>
,
<collectingRegion id="49A7F82CFFA4FF8FFF0BF86AFF7BF83D" box="[151,251,1962,1986]" country="China" name="Guangxi" pageId="89" pageNumber="90">GUaNGxI</collectingRegion>
,
<collectingRegion id="49A7F82CFFA4FF8FFE99F86AFED3F83E" box="[261,339,1962,1985]" country="China" name="Hunan" pageId="89" pageNumber="90">HUNaN</collectingRegion>
,
<collectingRegion id="49A7F82CFFA4FF8FFEC2F869FE26F83D" box="[350,422,1961,1986]" country="China" name="Hubei" pageId="89" pageNumber="90">HUbEI</collectingRegion>
,
<collectingRegion id="49A7F82CFFA4FF8FFE2CF86AFD8EF83D" box="[432,526,1962,1986]" country="China" name="Sichuan" pageId="89" pageNumber="90">SICHUaN</collectingRegion>
,
<collectingRegion id="49A7F82CFFA4FF8FFD84F86AFD18F83D" box="[536,664,1962,1986]" country="China" name="Chongqing" pageId="89" pageNumber="90">CHONGqING</collectingRegion>
,
<collectingRegion id="49A7F82CFFA4FF8FFD3FF86AFC87F83D" box="[675,775,1962,1986]" country="China" name="Guizhou" pageId="89" pageNumber="90">GUIzHOU</collectingRegion>
) aNd
<collectingRegion id="49A7F82CFFA4FF8FFCDFF86AFC28F83D" box="[835,936,1962,1986]" country="Sri Lanka" name="Northern" pageId="89" pageNumber="90">NOrTHErN</collectingRegion>
<collectingCountry id="F374765EFFA4FF8FFC33F86AFB8EF83E" box="[943,1038,1962,1985]" name="Vietnam" pageId="89" pageNumber="90">VIETNam</collectingCountry>
(
<figureCitation id="13582A4BFFA4FF8FFB87F86AFB0FF83D" box="[1051,1167,1961,1986]" captionStart="FIGURE 57" captionStartId="86.[151,250,1803,1825]" captionTargetBox="[151,1436,844,1782]" captionTargetId="figure@86.[151,1436,844,1782]" captionTargetPageId="86" captionText="FIGURE 57. Distribution of Metalycaeus species. Empty diamond. Metalycaeus zayuensis (Zhang, Chen &amp; Zhou, 2008). Filled triangle. Metalycaeus oharai Páll-Gergely &amp; Hunyadi n. sp. Filled square. Metalycaeus (?) awalycaeoides Páll- Gergely &amp; Hunyadi, n. sp. Circles. Metalycaeus muciferus (Heude, 1885). Empty triangle, top down. Metalycaeus laosensis n. sp. Empty circles indicate figured populations of Metalycaeus muciferus. 1. Tchen K’eou (Chengkou), the type locality of Alycaeus muciferus Heude, 1885. 2. Changyang (SMF 39254). 3. 2015 / 48. 4. 20070714 B. 5. Pe-shang (type locality of Alycaeus helicodes Gredler, 1887). 6. 2013 / 7. 7. 2011 / 15." httpUri="https://zenodo.org/record/1009839/files/figure.png" pageId="89" pageNumber="90">FIGUrE 57</figureCitation>
).
</paragraph>
</subSubSection>
<paragraph id="8BDC36CEFFA7FF8CFF0BFB1CFE0CFAD0" pageId="90" pageNumber="91">
<table id="F963C46EFFA70029FF0BFB1CFA1CFAD0" box="[151,1436,1244,1327]" gridcols="4" gridrows="3" pageId="90" pageNumber="91">
<tr id="3553348CFFA70029FF0BFB1CFA1CFB0D" box="[151,1436,1244,1266]" gridrow="0" pageId="90" pageNumber="91">
<th id="76825DF0FFA70029FF0BFB1CFE12FB0D" box="[151,402,1244,1266]" gridcol="0" gridrow="0" pageId="90" pageNumber="91">
<emphasis id="B917EADCFFA7FF8CFF0BFB1CFEA5FB0D" bold="true" box="[151,293,1244,1266]" pageId="90" pageNumber="91">FIGURE 59.</emphasis>
Shells of
</th>
<taxonomicName id="4C634D4DFFA7FF8CFE00FB1CFCB3FB0D" authority="(Heude, 1885)" baseAuthorityName="Heude" baseAuthorityYear="1885" box="[412,819,1244,1266]" class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="90" pageNumber="91" phylum="Mollusca" rank="species" species="muciferus">
<th id="76825DF0FFA70029FE05FB1CFD0DFB0D" box="[409,653,1244,1266]" gridcol="1" gridrow="0" pageId="90" pageNumber="91">
<emphasis id="B917EADCFFA7FF8CFE00FB1CFD0CFB0D" box="[412,652,1244,1266]" italics="true" pageId="90" pageNumber="91">Metalycaeus muciferus</emphasis>
</th>
<th id="76825DF0FFA70029FD08FB1CFC5CFB0D" box="[660,988,1244,1266]" gridcol="2" gridrow="0" pageId="90" pageNumber="91">
(Heude, 1885).
<emphasis id="B917EADCFFA7FF8CFCDFFB1CFCDBFB0D" bold="true" box="[835,859,1244,1266]" pageId="90" pageNumber="91">A.</emphasis>
2013/7.
<emphasis id="B917EADCFFA7FF8CFC5CFB1CFC57FB0D" bold="true" box="[960,983,1244,1266]" pageId="90" pageNumber="91">B.</emphasis>
</th>
</taxonomicName>
<th id="76825DF0FFA70029FC7FFB1CFA1CFB0D" box="[995,1436,1244,1266]" gridcol="3" gridrow="0" pageId="90" pageNumber="91">
paralectotype (SMF 192225) of
<taxonomicName id="4C634D4DFFA7FF8CFADCFB1CFA1CFB0D" box="[1344,1436,1244,1266]" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Architaenioglossa" pageId="90" pageNumber="91" phylum="Mollusca" rank="genus">
<emphasis id="B917EADCFFA7FF8CFADCFB1CFA1CFB0D" box="[1344,1436,1244,1266]" italics="true" pageId="90" pageNumber="91">Alycaeus</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="3553348CFFA70029FF0BFB3BFA1CFAEE" box="[151,1436,1275,1297]" gridrow="1" pageId="90" pageNumber="91">
<th id="76825DF0FFA70029FF0BFB3BFE12FAEE" box="[151,402,1275,1297]" gridcol="0" gridrow="1" pageId="90" pageNumber="91">
<taxonomicName id="4C634D4DFFA7FF8CFF0BFB3BFE0FFAEE" authority="Gredler, 1887" authorityName="Gredler" authorityYear="1887" box="[151,399,1275,1297]" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="90" pageNumber="91" phylum="Mollusca" rank="species" species="helicodes">
<emphasis id="B917EADCFFA7FF8CFF0BFB3BFF77FAEE" box="[151,247,1275,1297]" italics="true" pageId="90" pageNumber="91">helicodes</emphasis>
Gredler, 1887
</taxonomicName>
.
</th>
<td id="76825DF0FFA70029FE05FB3BFD0DFAEE" box="[409,653,1275,1297]" gridcol="1" gridrow="1" pageId="90" pageNumber="91">
<emphasis id="B917EADCFFA7FF8CFE05FB3BFE32FAEE" bold="true" box="[409,434,1275,1297]" pageId="90" pageNumber="91">C.</emphasis>
SMF 39254 (figured
</td>
<td id="76825DF0FFA70029FD08FB3BFC5CFAEE" box="[660,988,1275,1297]" gridcol="2" gridrow="1" pageId="90" pageNumber="91">
in Yen 1939, as &quot;
<taxonomicName id="4C634D4DFFA7FF8CFCDBFB3BFC5CFAEE" authorityName="Mollendorff" authorityYear="1897" box="[839,988,1275,1297]" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="90" pageNumber="91" phylum="Mollusca" rank="genus">
<emphasis id="B917EADCFFA7FF8CFCDBFB3BFC5CFAEE" box="[839,988,1275,1297]" italics="true" pageId="90" pageNumber="91">Chamalycaeus</emphasis>
</taxonomicName>
</td>
<td id="76825DF0FFA70029FC7FFB3BFA1CFAEE" box="[995,1436,1275,1297]" gridcol="3" gridrow="1" pageId="90" pageNumber="91">
<taxonomicName id="4C634D4DFFA7FF8CFC78FB3BFBE4FAEE" box="[996,1124,1275,1297]" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="90" pageNumber="91" phylum="Mollusca" rank="species" species="dolichodirus">
<emphasis id="B917EADCFFA7FF8CFC78FB3BFBE4FAEE" box="[996,1124,1275,1297]" italics="true" pageId="90" pageNumber="91">dolichodirus</emphasis>
</taxonomicName>
&quot;). Scale represents 5 mm. All
</td>
</tr>
<tr id="3553348CFFA70029FF0BFAD9FA1CFAD0" box="[151,1436,1305,1327]" gridrow="2" pageId="90" pageNumber="91" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="76825DF0FFA70029FF0BFAD9FE12FAD0" box="[151,402,1305,1327]" gridcol="0" gridrow="2" pageId="90" pageNumber="91">photos: B. Páll-Gergely.</th>
</tr>
</table>
</paragraph>
<subSubSection id="C3796545FFA7FF8CFF5BFA96FAEDF850" pageId="90" pageNumber="91" type="discussion">
<paragraph id="8BDC36CEFFA7FF8CFF5BFA96FE34F94C" blockId="90.[151,1438,1366,1967]" pageId="90" pageNumber="91">
<emphasis id="B917EADCFFA7FF8CFF5BFA96FEBBFA90" bold="true" box="[199,315,1366,1391]" pageId="90" pageNumber="91">Remarks.</emphasis>
TypICaL
<taxonomicName id="4C634D4DFFA7FF8CFE00FA97FD1FFA90" baseAuthorityName="Heude" baseAuthorityYear="1885" box="[412,671,1367,1391]" class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="90" pageNumber="91" phylum="Mollusca" rank="species" species="muciferus">
<emphasis id="B917EADCFFA7FF8CFE00FA97FD1FFA90" box="[412,671,1367,1391]" italics="true" pageId="90" pageNumber="91">Metalycaeus muciferus</emphasis>
</taxonomicName>
(
<figureCitation id="13582A4BFFA7FF8CFD30FA97FCB1FA90" box="[684,817,1366,1391]" captionStart="FIGURE 58" captionStartId="89.[151,250,1695,1717]" captionTargetBox="[151,1436,195,1671]" captionTargetId="figure@89.[151,1436,195,1671]" captionTargetPageId="89" captionText="FIGURE 58. Shells of Metalycaeus muciferus (Heude, 1885). A. syntype. B. 20070714 B, coll. PGB. C. 2011 / 15, coll. HA. D. 2015 / 48, coll. HA. Note that the black &quot; operculum &quot; of Fig. D is the posterior end of a predator fly larva. Scale represents 5 mm. All photos: B. Páll-Gergely." httpUri="https://zenodo.org/record/1009841/files/figure.png" pageId="90" pageNumber="91">FIGUrE 58A</figureCitation>
) aNd
<taxonomicName id="4C634D4DFFA7FF8CFCF3FA98FC7EFA90" authorityName="Gredler" authorityYear="1887" box="[879,1022,1367,1391]" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="90" pageNumber="91" phylum="Mollusca" rank="species" species="helicodes">
<emphasis id="B917EADCFFA7FF8CFCF3FA98FC7EFA90" box="[879,1022,1367,1391]" italics="true" pageId="90" pageNumber="91">M. helicodes</emphasis>
</taxonomicName>
(FIGUrE 59B) spECImENs aGrEE IN THE fOLLOwING TraITs: bOTH R2 aNd R3 arE vEry sHOrT; THErE Is NO rECOGNIsabLE CONsTrICTION aT THE bOrdEr Of THE TwO rEGIONs; THE rIbs ON R2 arE wIdELy-spaCEd aNd sHarp; pErIsTOmE rELaTIvELy THIN (aLTHOUGH sOmETImEs ExpaNdEd). WE ExamINEd a NUmbEr Of NEwLy COLLECTEd CHINEsE sampLEs, wHICH aGrEE wITH
<taxonomicName id="4C634D4DFFA7FF8CFC6FFA04FB09FA24" baseAuthorityName="Heude" baseAuthorityYear="1885" box="[1011,1161,1475,1499]" class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="90" pageNumber="91" phylum="Mollusca" rank="species" species="muciferus">
<emphasis id="B917EADCFFA7FF8CFC6FFA04FB09FA24" box="[1011,1161,1475,1499]" italics="true" pageId="90" pageNumber="91">M. muciferus</emphasis>
</taxonomicName>
aNd
<taxonomicName id="4C634D4DFFA7FF8CFB5EFA04FAD3FA24" authorityName="Gredler" authorityYear="1887" box="[1218,1363,1475,1499]" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="90" pageNumber="91" phylum="Mollusca" rank="species" species="helicodes">
<emphasis id="B917EADCFFA7FF8CFB5EFA04FAD3FA24" box="[1218,1363,1475,1499]" italics="true" pageId="90" pageNumber="91">M. helicodes</emphasis>
</taxonomicName>
IN THE abOvE mENTIONEd TraITs, aLTHOUGH sHOw dIffErENCEs IN TErms Of spIrE HEIGHT, sCULpTUrE Of R1 aNd sIzE (sEE
<tableCitation id="C6E10375FFA7FF8CFAD4FA26FA1CFA00" box="[1352,1436,1510,1535]" captionStart="TABLE 5" captionStartId="92.[151,239,1332,1355]" captionText="TABLE 5. Variability of Metalycaeus muciferus populations." pageId="90" pageNumber="91">TabLE 5</tableCitation>
aNd
<figureCitation id="13582A4BFFA7FF8CFF51F9CBFECDF9DB" box="[205,333,1547,1572]" captionStart="FIGURE 58" captionStartId="89.[151,250,1695,1717]" captionTargetBox="[151,1436,195,1671]" captionTargetId="figure@89.[151,1436,195,1671]" captionTargetPageId="89" captionText="FIGURE 58. Shells of Metalycaeus muciferus (Heude, 1885). A. syntype. B. 20070714 B, coll. PGB. C. 2011 / 15, coll. HA. D. 2015 / 48, coll. HA. Note that the black &quot; operculum &quot; of Fig. D is the posterior end of a predator fly larva. Scale represents 5 mm. All photos: B. Páll-Gergely." httpUri="https://zenodo.org/record/1009841/files/figure.png" pageId="90" pageNumber="91">FIGUrEs 58</figureCitation>
–59). NEarLy EvEry sINGLE pOpULaTION wE ExamINEd Has CHaraCTErIsTIC fEaTUrEs, wHICH aLLOws dIsTINGUIsHING IT frOm OTHErs. THEsE dIffErENCEs arE raTHEr mINOr. MOrEOvEr, THEsE sampLEs arE smaLL aNd maINLy CONTaIN wEaTHErEd sHELLs wITHOUT OpErCULa. THErEfOrE wE dO NOT fINd IT rEasONabLE TO dEsCrIbE THEm fOrmaLLy. INsTEad, UNTIL mOrE maTErIaL bECOmEs avaILabLE, wE LIsT aLL THEsE sampLEs aNd HaNdLE THEm UNdEr a sINGLE NamE (
<taxonomicName id="4C634D4DFFA7FF8CFF3CF95BFE22F94C" baseAuthorityName="Heude" baseAuthorityYear="1885" box="[160,418,1691,1715]" class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="90" pageNumber="91" phylum="Mollusca" rank="species" species="muciferus">
<emphasis id="B917EADCFFA7FF8CFF3CF95BFE22F94C" box="[160,418,1691,1715]" italics="true" pageId="90" pageNumber="91">Metalycaeus muciferus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BDC36CEFFA7FF8CFF5BF97EFDEBF8E0" blockId="90.[151,1438,1366,1967]" pageId="90" pageNumber="91">
AdOLf ZILCH brOUGHT TwO spECImENs frOm GrEdLEr's COLLECTION (BOzEN = BOLzaNO) TO THE SENCkENbErG MUsEUm. HE mENTIONEd THaT THE
<typeStatus id="54D8886CFFA7FF8CFE3CF923FD87F904" box="[416,519,1763,1787]" pageId="90" pageNumber="91" type="lectotype">LECTOTypE</typeStatus>
rEmaINEd IN BOLzaNO (
<bibRefCitation id="EFF24B3FFFA7FF8CFC92F923FC12F904" author="ZILCH" box="[782,914,1763,1787]" pageId="90" pageNumber="91" refString="Zilch, A. (1974) Vinzenz Gredler und die Erforschung der Weichtiere Chinas durch Franziskaner aus Tirol. Archiv fur Molluskenkunde, 104, 171 - 228." type="journal article" year="1974">ZILCH 1974</bibRefCitation>
), THErEfOrE THE spECImENs IN THE SENCkENbErG MUsEUm arE
<typeStatus id="54D8886CFFA7FF8CFEB5F8C7FE4BF8E0" box="[297,459,1799,1823]" pageId="90" pageNumber="91" type="paralectotype">paraLECTOTypEs</typeStatus>
(FIGUrE 59B).
</paragraph>
<paragraph id="8BDC36CEFFA7FF8CFF5BF8ECFAEDF850" blockId="90.[151,1438,1366,1967]" pageId="90" pageNumber="91">
THE CrOss sECTION Of R2 was ExamINEd IN ONE spECImEN Of THE pOpULaTION 2015/48 (
<figureCitation id="13582A4BFFA7FF8CFBFAF8EBFB6DF8BB" box="[1126,1261,1835,1860]" captionStart="FIGURE 61" captionStartId="92.[151,250,1270,1292]" captionTargetBox="[178,1422,193,1266]" captionTargetId="figure@92.[178,1422,193,1266]" captionTargetPageId="92" captionText="FIGURE 61. Protoconchs of Metalycaeus muciferus (Heude, 1885). A – B. 2013 / 7. C – D. syntype (MCZ 167230)." httpUri="https://zenodo.org/record/1009845/files/figure.png" pageId="90" pageNumber="91">FIGUrE 61D</figureCitation>
). ANTErIOr pLaTE raTHEr sTraIGHT, HOLLOw, IT Has a LOw, bUT sHarp rIb; pOsTErIOr CrUsT sTrONG, ONLy sLIGHTLy LOwEr THaN THE aNTErIOr ONE; THE mICrOTUNNEL Has a TEardrOp-sHapEd CrOss sECTION, ITs dIamETEr Is CONspICUOUsLy LarGE IN rELaTION TO THE sIzE Of THE CrUsTs. THE aNTErIOr CrUsT Of TypICaL
<taxonomicName id="4C634D4DFFA7FF8CFDB9F857FD13F850" baseAuthorityName="Heude" baseAuthorityYear="1885" box="[549,659,1943,1967]" class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="90" pageNumber="91" phylum="Mollusca" rank="species" species="muciferus">
<emphasis id="B917EADCFFA7FF8CFDB9F857FD13F850" box="[549,659,1943,1967]" italics="true" pageId="90" pageNumber="91">muciferus</emphasis>
</taxonomicName>
sHELLs Is HOLLOw. SEE aLsO rEmarks UNdEr
<taxonomicName id="4C634D4DFFA7FF8CFBFDF857FAE6F850" authorityName="Heude" authorityYear="1890" box="[1121,1382,1943,1967]" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Architaenioglossa" pageId="90" pageNumber="91" phylum="Mollusca" rank="species" species="dolichodeiros">
<emphasis id="B917EADCFFA7FF8CFBFDF857FAE6F850" box="[1121,1382,1943,1967]" italics="true" pageId="90" pageNumber="91">Alycaeus dolichodeiros</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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