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<document id="C36419CB498F069730117451160878C7" ID-DOI="10.5281/zenodo.5331014" ID-ISSN="0374-1036" ID-Zenodo-Dep="5331014" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630168815873" checkinUser="diego" docAuthor="Megna, Yoandri S. &amp; Epler, John H." docDate="2012" docId="03CA7F7DFFEC4C59FE4099C84BBFF985" docLanguage="en" docName="ActaEntMusNatPra.2012.52.2.383-410.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 52 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Copelatus : GUEORGUIEV 1968" docType="key" docVersion="4" lastPageNumber="387" masterDocId="FFF30705FFE84C5DFFAE99134914FFB2" masterDocTitle="A review of Copelatus from Cuba, with the description of two new species (Coleoptera: Dytiscidae: Copelatinae)" masterLastPageNumber="410" masterPageNumber="383" pageNumber="387" updateTime="1699171915230" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="7F6D15AF96E9334D0FD844D7C09088C1">A review of Copelatus from Cuba, with the description of two new species (Coleoptera: Dytiscidae: Copelatinae)</mods:title>
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<mods:namePart id="AFF21EF8A704C67FBCCB6F8E19C14325">Megna, Yoandri S.</mods:namePart>
<mods:affiliation id="C71EED88E028D8522185216D1D3602AA">Departamento de Biología, Universidad de Oriente. Patricio Lumumba s / n, Santiago, Santiago de Cuba, Cuba; e-mail: ysuarez 1976 @ yahoo. es</mods:affiliation>
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<mods:namePart id="21F47E26B91A7C53033565678A2672ED">Epler, John H.</mods:namePart>
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Key to 
<taxonomicName id="4C63B5E8FFEC4C59FDEE99C94BBAFF41" ID-CoL="3TN2" authorityName=": GUEORGUIEV" authorityYear="1968" box="[576,686,218,243]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="387" phylum="Arthropoda" rank="genus">
<emphasis id="B9171279FFEC4C59FDEE99C94BBAFF41" bold="true" box="[576,686,218,243]" italics="true" pageId="4" pageNumber="387">Copelatus</emphasis>
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of 
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1 Elytra with well-defined striae, either linear or undulating/anastomosing (
<figureCitation id="1358D2EEFFEC4C59FC5698194811FEF0" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="6.[151,189,1533,1552]" captionStartId-1="11.[113,152,1502,1521]" captionTargetBox-0="[166,1114,432,1505]" captionTargetBox-1="[151,1057,222,1450]" captionTargetId-0="figure-11@6.[151,1126,417,1515]" captionTargetId-1="figure-13@11.[113,1088,216,1465]" captionTargetPageId-0="6" captionTargetPageId-1="11" captionText-0="Figs.1–8. Habitus and color patterns of Copelatus species from Cuba.1 – C.barbouri Young, 1942; 2 – C. caelatipennis angustatus Chevrolat, 1863; 3 – C. cordovai sp. nov.; 4 – C. cubaensis Schaeffer, 1908; 5 – Copelatus darlingtoni Young, 1942; 6 – C. insolitus Chevrolat, 1863; 7 – C. montivagus Young, 1942; 8 – C. posticatus (Fabricius, 1801)." captionText-1="Figs. 9–11. Copelatus danyi sp. nov. 9 – female habitus; 10 – male habitus; 11 – male genitalia (a – median lobe in right lateral view; b – same in left lateral view; c – left paramere in lateral view)." figureDoi-0="http://doi.org/10.5281/zenodo.5331018" figureDoi-1="http://doi.org/10.5281/zenodo.5331020" httpUri-0="https://zenodo.org/record/5331018/files/figure.png" httpUri-1="https://zenodo.org/record/5331020/files/figure.png" pageId="4" pageNumber="387">Figs. 2–4, 8–10</figureCitation>
). ........................................................................................................................... 2
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<paragraph id="8BDCCE6BFFEC4C59FF3998584D72FE30" blockId="4.[151,1127,266,1533]" pageId="4" pageNumber="387">
<keyLead id="30926D5EFFEC4C59FF3998584D72FE30" pageId="4" pageNumber="387">
- Elytra mostly smooth, at most with longitudinal series of weak punctures (
<figureCitation id="1358D2EEFFEC4C59FC4C985949C1FE30" captionStart="Figs" captionStartId="6.[151,189,1533,1552]" captionTargetBox="[166,1114,432,1505]" captionTargetId="figure-11@6.[151,1126,417,1515]" captionTargetPageId="6" captionText="Figs.1–8. Habitus and color patterns of Copelatus species from Cuba.1 – C.barbouri Young, 1942; 2 – C. caelatipennis angustatus Chevrolat, 1863; 3 – C. cordovai sp. nov.; 4 – C. cubaensis Schaeffer, 1908; 5 – Copelatus darlingtoni Young, 1942; 6 – C. insolitus Chevrolat, 1863; 7 – C. montivagus Young, 1942; 8 – C. posticatus (Fabricius, 1801)." figureDoi="http://doi.org/10.5281/zenodo.5331018" httpUri="https://zenodo.org/record/5331018/files/figure.png" pageId="4" pageNumber="387">Figs. 1, 5, 6, 7</figureCitation>
). .................................................................................................................................. 9
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2 Elytra with 5–10 well defined linear striae; males and females (
<figureCitation id="1358D2EEFFEC4C59FCC998994D13FE10" box="[871,1031,394,418]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="6.[151,189,1533,1552]" captionStartId-1="11.[113,152,1502,1521]" captionTargetBox-0="[166,1114,432,1505]" captionTargetBox-1="[151,1057,222,1450]" captionTargetId-0="figure-11@6.[151,1126,417,1515]" captionTargetId-1="figure-13@11.[113,1088,216,1465]" captionTargetPageId-0="6" captionTargetPageId-1="11" captionText-0="Figs.1–8. Habitus and color patterns of Copelatus species from Cuba.1 – C.barbouri Young, 1942; 2 – C. caelatipennis angustatus Chevrolat, 1863; 3 – C. cordovai sp. nov.; 4 – C. cubaensis Schaeffer, 1908; 5 – Copelatus darlingtoni Young, 1942; 6 – C. insolitus Chevrolat, 1863; 7 – C. montivagus Young, 1942; 8 – C. posticatus (Fabricius, 1801)." captionText-1="Figs. 9–11. Copelatus danyi sp. nov. 9 – female habitus; 10 – male habitus; 11 – male genitalia (a – median lobe in right lateral view; b – same in left lateral view; c – left paramere in lateral view)." figureDoi-0="http://doi.org/10.5281/zenodo.5331018" figureDoi-1="http://doi.org/10.5281/zenodo.5331020" httpUri-0="https://zenodo.org/record/5331018/files/figure.png" httpUri-1="https://zenodo.org/record/5331020/files/figure.png" pageId="4" pageNumber="387">Fig. 2–4, 8–10</figureCitation>
). ........ 3
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<keyLead id="30926D5EFFEC4C59FF3998B84D73FE70" box="[151,1127,424,450]" pageId="4" pageNumber="387">
- Elytra striae fine, undulating/anastomosing; females 
<superScript id="7C166323FFEC4C59FD4198BB4BE3FE05" attach="left" box="[751,759,424,439]" fontSize="6" pageId="4" pageNumber="387">1</superScript>
(
<figureCitation id="1358D2EEFFEC4C59FCA898B94A53FE70" box="[774,839,426,450]" captionStart="Figs" captionStartId="11.[113,152,1502,1521]" captionTargetBox="[151,1057,222,1450]" captionTargetId="figure-13@11.[113,1088,216,1465]" captionTargetPageId="11" captionText="Figs. 9–11. Copelatus danyi sp. nov. 9 – female habitus; 10 – male habitus; 11 – male genitalia (a – median lobe in right lateral view; b – same in left lateral view; c – left paramere in lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5331020" httpUri="https://zenodo.org/record/5331020/files/figure.png" pageId="4" pageNumber="387">Fig. 9</figureCitation>
). .................................. 12
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3 Each elytron with 5 or 6 discal striae (
<figureCitation id="1358D2EEFFEC4C59FDF598D94B89FE50" box="[603,669,458,482]" captionStart="Figs" captionStartId="6.[151,189,1533,1552]" captionTargetBox="[166,1114,432,1505]" captionTargetId="figure-11@6.[151,1126,417,1515]" captionTargetPageId="6" captionText="Figs.1–8. Habitus and color patterns of Copelatus species from Cuba.1 – C.barbouri Young, 1942; 2 – C. caelatipennis angustatus Chevrolat, 1863; 3 – C. cordovai sp. nov.; 4 – C. cubaensis Schaeffer, 1908; 5 – Copelatus darlingtoni Young, 1942; 6 – C. insolitus Chevrolat, 1863; 7 – C. montivagus Young, 1942; 8 – C. posticatus (Fabricius, 1801)." figureDoi="http://doi.org/10.5281/zenodo.5331018" httpUri="https://zenodo.org/record/5331018/files/figure.png" pageId="4" pageNumber="387">Fig. 4</figureCitation>
). .............................................................. 4
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<paragraph id="8BDCCE6BFFEC4C59FF3998F84D72FDB0" blockId="4.[151,1127,266,1533]" box="[151,1126,490,514]" pageId="4" pageNumber="387">
<keyLead id="30926D5EFFEC4C59FF3998F84D72FDB0" box="[151,1126,490,514]" pageId="4" pageNumber="387">
- Each elytron with 10 discal striae (
<figureCitation id="1358D2EEFFEC4C59FD9998F94B6CFDB0" box="[567,632,490,514]" captionStart="Figs" captionStartId="6.[151,189,1533,1552]" captionTargetBox="[166,1114,432,1505]" captionTargetId="figure-11@6.[151,1126,417,1515]" captionTargetPageId="6" captionText="Figs.1–8. Habitus and color patterns of Copelatus species from Cuba.1 – C.barbouri Young, 1942; 2 – C. caelatipennis angustatus Chevrolat, 1863; 3 – C. cordovai sp. nov.; 4 – C. cubaensis Schaeffer, 1908; 5 – Copelatus darlingtoni Young, 1942; 6 – C. insolitus Chevrolat, 1863; 7 – C. montivagus Young, 1942; 8 – C. posticatus (Fabricius, 1801)." figureDoi="http://doi.org/10.5281/zenodo.5331018" httpUri="https://zenodo.org/record/5331018/files/figure.png" pageId="4" pageNumber="387">Fig. 8</figureCitation>
). .................................................................... 5
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<keyStep id="3097D6CEFFEC4C59FF399B194D73FD29" pageId="4" pageNumber="387">
<paragraph id="8BDCCE6BFFEC4C59FF399B194D72FDF0" blockId="4.[151,1127,266,1533]" pageId="4" pageNumber="387">
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4 Each elytron with a posterolateral submarginal stria and 6 discal striae (
<figureCitation id="1358D2EEFFEC4C59FC6E9B194D16FD90" box="[960,1026,522,546]" captionStart="Figs" captionStartId="6.[151,189,1533,1552]" captionTargetBox="[166,1114,432,1505]" captionTargetId="figure-11@6.[151,1126,417,1515]" captionTargetPageId="6" captionText="Figs.1–8. Habitus and color patterns of Copelatus species from Cuba.1 – C.barbouri Young, 1942; 2 – C. caelatipennis angustatus Chevrolat, 1863; 3 – C. cordovai sp. nov.; 4 – C. cubaensis Schaeffer, 1908; 5 – Copelatus darlingtoni Young, 1942; 6 – C. insolitus Chevrolat, 1863; 7 – C. montivagus Young, 1942; 8 – C. posticatus (Fabricius, 1801)." figureDoi="http://doi.org/10.5281/zenodo.5331018" httpUri="https://zenodo.org/record/5331018/files/figure.png" pageId="4" pageNumber="387">Fig. 4</figureCitation>
). ........... ........................................................................................... 
<taxonomicName id="4C63B5E8FFEC4C59FC839B394D72FDF0" ID-CoL="Y36G" authority="Schaeffer, 1908" authorityName="Schaeffer" authorityYear="1908" box="[813,1126,553,578]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="387" phylum="Arthropoda" rank="species" species="cubaensis">
<emphasis id="B9171279FFEC4C59FC839B394AACFDF0" bold="true" box="[813,952,553,578]" italics="true" pageId="4" pageNumber="387">C. cubaensis</emphasis>
<bibRefCitation id="EFF2B39AFFEC4C59FC119B394D72FDF0" author="SCHAEFFER C." box="[959,1126,554,578]" pageId="4" pageNumber="387" pagination="16 - 18" refId="ref13877" refString="SCHAEFFER C. 1908: On North American and some Cuban Copelatus. Journal of the New York Entomological Society 16: 16 - 18." type="journal article" year="1908">Schaeffer, 1908</bibRefCitation>
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- Each elytron with a posterolateral submarginal stria and 5 discal striae. ........................ ................................................................................................ 
<taxonomicName id="4C63B5E8FFEC4C59FCE39B794D72FD30" ID-CoL="5ZX7D" authority="Young, 1953" authorityName="Young" authorityYear="1953" box="[845,1126,617,643]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="387" phylum="Arthropoda" rank="species" species="blatchleyi">
<emphasis id="B9171279FFEC4C59FCE39B794AC3FD30" bold="true" box="[845,983,617,642]" italics="true" pageId="4" pageNumber="387">C. blatchleyi</emphasis>
Young, 1953
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(not recorded from 
<collectingCountry id="F3748EFBFFEC4C59FE769B9B4B13FD29" box="[472,519,648,667]" name="Cuba" pageId="4" pageNumber="387">Cuba</collectingCountry>
, but may occur there due to its presence in Florida Keys and 
<collectingCountry id="F3748EFBFFEC4C59FBBF9B9B4D75FD29" box="[1041,1121,648,667]" name="Bahamas" pageId="4" pageNumber="387">Bahamas</collectingCountry>
)
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<paragraph id="8BDCCE6BFFEC4C59FF399BB64D72FD6F" blockId="4.[151,1127,266,1533]" pageId="4" pageNumber="387">
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5 Each elytron with striae 1, 3, 5, and 9 extending to apical third (
<figureCitation id="1358D2EEFFEC4C59FCCF9BB64AB5FD0F" box="[865,929,677,701]" captionStart="Figs" captionStartId="6.[151,189,1533,1552]" captionTargetBox="[166,1114,432,1505]" captionTargetId="figure-11@6.[151,1126,417,1515]" captionTargetPageId="6" captionText="Figs.1–8. Habitus and color patterns of Copelatus species from Cuba.1 – C.barbouri Young, 1942; 2 – C. caelatipennis angustatus Chevrolat, 1863; 3 – C. cordovai sp. nov.; 4 – C. cubaensis Schaeffer, 1908; 5 – Copelatus darlingtoni Young, 1942; 6 – C. insolitus Chevrolat, 1863; 7 – C. montivagus Young, 1942; 8 – C. posticatus (Fabricius, 1801)." figureDoi="http://doi.org/10.5281/zenodo.5331018" httpUri="https://zenodo.org/record/5331018/files/figure.png" pageId="4" pageNumber="387">Fig. 8</figureCitation>
). Body size larger, TL&gt; 6.0 mm; EW&gt; 3.0 mm. ........................................ 
<taxonomicName id="4C63B5E8FFEC4C59FCB59BD64D72FD6F" ID-CoL="5ZX8Q" authority="(Fabricius, 1801)" baseAuthorityName="Fabricius" baseAuthorityYear="1801" box="[795,1126,709,733]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="387" phylum="Arthropoda" rank="species" species="posticatus">
<emphasis id="B9171279FFEC4C59FCB59BD64ABCFD6F" bold="true" box="[795,936,709,733]" italics="true" pageId="4" pageNumber="387">C. posticatus</emphasis>
(
<bibRefCitation id="EFF2B39AFFEC4C59FC189BD64D4AFD6F" author="FABRICIUS J. C." box="[950,1118,709,733]" pageId="4" pageNumber="387" refId="ref13078" refString="FABRICIUS J. C. 1801: Systema Eleutheratorum secundum ordines, genera, species; adiectis synonymis, locis, observationibus, descriptionibus. Tomus I. Bibliopolii Academici Novi, Kiliae, xxiv + 506 pp." type="book" year="1801">Fabricius, 1801</bibRefCitation>
)
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<keyLead id="30926D5EFFEC4C59FF399BF64D72FD4F" box="[151,1126,741,765]" pageId="4" pageNumber="387">
- Each elytron with all striae almost complete (
<figureCitation id="1358D2EEFFEC4C59FD079BF64BFFFD4F" box="[681,747,741,765]" captionStart="Figs" captionStartId="6.[151,189,1533,1552]" captionTargetBox="[166,1114,432,1505]" captionTargetId="figure-11@6.[151,1126,417,1515]" captionTargetPageId="6" captionText="Figs.1–8. Habitus and color patterns of Copelatus species from Cuba.1 – C.barbouri Young, 1942; 2 – C. caelatipennis angustatus Chevrolat, 1863; 3 – C. cordovai sp. nov.; 4 – C. cubaensis Schaeffer, 1908; 5 – Copelatus darlingtoni Young, 1942; 6 – C. insolitus Chevrolat, 1863; 7 – C. montivagus Young, 1942; 8 – C. posticatus (Fabricius, 1801)." figureDoi="http://doi.org/10.5281/zenodo.5331018" httpUri="https://zenodo.org/record/5331018/files/figure.png" pageId="4" pageNumber="387">Fig. 2</figureCitation>
); TL &lt;
<quantity id="4C9B638EFFEC4C59FC949BF64A9BFD4E" box="[826,911,741,765]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="4" pageNumber="387" unit="mm" value="6.5">6.5 mm</quantity>
. ........................... 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3097D6CEFFEC4C59FF399A164D72FC8F" pageId="4" pageNumber="387">
<paragraph id="8BDCCE6BFFEC4C59FF399A164D72FCAE" blockId="4.[151,1127,266,1533]" box="[151,1126,773,797]" pageId="4" pageNumber="387">
<keyLead id="30926D5EFFEC4C59FF399A164D72FCAE" box="[151,1126,773,797]" pageId="4" pageNumber="387">
6 Median lobe with apex similar to a bird head (
<figureCitation id="1358D2EEFFEC4C59FD1C9A164A1FFCAF" box="[690,779,773,797]" captionStart="Figs" captionStartId="14.[151,190,1544,1563]" captionTargetBox="[209,1062,729,1508]" captionTargetId="figure-11@14.[179,1099,723,1525]" captionTargetPageId="14" captionText="Figs. 20–23. Male genitalia of Cuban species of Copelatus. 20 – C. caelatipennis angustatus author, year; 21 – C. cordovai sp. nov.; 22 – C. cubaensis Schaeffer, 1908; 23 – C. posticatus (Fabricius, 1801). Parts of male genitalia: a – median lobe of aedeagus in left lateral view, b – left paramere in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5331026" httpUri="https://zenodo.org/record/5331026/files/figure.png" pageId="4" pageNumber="387">Fig. 20a</figureCitation>
) (
<taxonomicName id="4C63B5E8FFEC4C59FC8C9A154ADBFCAF" ID-CoL="Y35F" authorityName=": SCHAEFFER" authorityYear="1908" box="[802,975,774,797]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="387" phylum="Arthropoda" rank="species" species="caelatipennis">
<emphasis id="B9171279FFEC4C59FC8C9A154ADBFCAF" box="[802,975,774,797]" italics="true" pageId="4" pageNumber="387">C. caelatipennis</emphasis>
</taxonomicName>
). ................ 7
</keyLead>
</paragraph>
<paragraph id="8BDCCE6BFFEC4C59FF399A364D72FC8F" blockId="4.[151,1127,266,1533]" box="[151,1126,805,829]" pageId="4" pageNumber="387">
<keyLead id="30926D5EFFEC4C59FF399A364D72FC8F" box="[151,1126,805,829]" pageId="4" pageNumber="387">- Median lobe of different shape, not similar to a bird head. .......................................... 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="3097D6CEFFEC4C59FF399A564D72FC0F" pageId="4" pageNumber="387">
<paragraph id="8BDCCE6BFFEC4C59FF399A564D72FCCF" blockId="4.[151,1127,266,1533]" pageId="4" pageNumber="387">
<keyLead id="30926D5EFFEC4C59FF399A564D72FCCF" pageId="4" pageNumber="387">
7 Pronotum in both sexes usually without strioles on disc, if strioles are present, they are restricted to pronotal sides. ..................... 
<taxonomicName id="4C63B5E8FFEC4C59FD2B9A764D72FCCF" ID-CoL="5GLFM" authority="Chevrolat, 1863" authorityName="Chevrolat" authorityYear="1863" box="[645,1126,868,893]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="387" phylum="Arthropoda" rank="subSpecies" species="caelatipennis" subSpecies="angustatus">
<emphasis id="B9171279FFEC4C59FD2B9A764AA0FCCE" bold="true" box="[645,948,868,893]" italics="true" pageId="4" pageNumber="387">C. caelatipennis angustatus</emphasis>
<bibRefCitation id="EFF2B39AFFEC4C59FC149A764D72FCCF" author="CHEVROLAT L. A. A." box="[954,1126,869,893]" pageId="4" pageNumber="387" pagination="183 - 210" refId="ref12897" refString="CHEVROLAT L. A. A. 1863: Coleopteres de l'IIe de Cuba. Notes, synonymies et descriptions d'especes nouvelles. Familles des cicindeletes, carabiques, dytiscides, gyrinides et palpicornes. Annales de la Societe Entomologique de France 3: 183 - 210." type="journal article" year="1863">Chevrolat, 1863</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDCCE6BFFEC4C59FF399A964D72FC0F" blockId="4.[151,1127,266,1533]" pageId="4" pageNumber="387">
<keyLead id="30926D5EFFEC4C59FF399A964D72FC0F" pageId="4" pageNumber="387">
- Pronotum in both sexes usually with short strioles on disc. ............................................ ........................................................................... 
<taxonomicName id="4C63B5E8FFEC4C59FD6D9AB64D72FC0F" ID-CoL="5GLFQ" authority="Young, 1963" authorityName="Young" authorityYear="1963" box="[707,1126,932,957]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="387" phylum="Arthropoda" rank="subSpecies" species="caelatipennis" subSpecies="princeps">
<emphasis id="B9171279FFEC4C59FD6D9AB64AC3FC0F" bold="true" box="[707,983,932,957]" italics="true" pageId="4" pageNumber="387">C. caelatipennis princeps</emphasis>
<bibRefCitation id="EFF2B39AFFEC4C59FC709AB64D72FC0F" author="YOUNG F. N." box="[990,1126,933,957]" pageId="4" pageNumber="387" pagination="56 - 77" refId="ref14144" refString="YOUNG F. N. 1963: The Nearctic species of Copelatus Erichson (Coleoptera: Dytiscidae). Quarterly Journal of the Florida Academy of Sciences 26: 56 - 77." type="journal article" year="1963">Young, 1963</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3097D6CEFFEC4C59FF399AD64D73FB2F" pageId="4" pageNumber="387">
<paragraph id="8BDCCE6BFFEC4C59FF399AD64D73FBAF" blockId="4.[151,1127,266,1533]" pageId="4" pageNumber="387">
<keyLead id="30926D5EFFEC4C59FF399AD64D73FBAF" pageId="4" pageNumber="387">
<specimenCount id="9D6505E2FFEC4C59FF399AD649E8FC6F" box="[151,252,965,989]" pageId="4" pageNumber="387" type="male">8 Male</specimenCount>
protibia with two ventrobasal emarginations; median lobe with outside margin abruptly expanded at midlength (
<figureCitation id="1358D2EEFFEC4C59FDB09AF64B8FFC4F" box="[542,667,997,1021]" captionStart="Figs" captionStartId="14.[151,190,1544,1563]" captionTargetBox="[209,1062,729,1508]" captionTargetId="figure-11@14.[179,1099,723,1525]" captionTargetPageId="14" captionText="Figs. 20–23. Male genitalia of Cuban species of Copelatus. 20 – C. caelatipennis angustatus author, year; 21 – C. cordovai sp. nov.; 22 – C. cubaensis Schaeffer, 1908; 23 – C. posticatus (Fabricius, 1801). Parts of male genitalia: a – median lobe of aedeagus in left lateral view, b – left paramere in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5331026" httpUri="https://zenodo.org/record/5331026/files/figure.png" pageId="4" pageNumber="387">Figs. 21a, b</figureCitation>
); parameres broad, with short setae on inner margin (
<figureCitation id="1358D2EEFFEC4C59FEB09D16486EFBAF" box="[286,378,1029,1053]" captionStart="Figs" captionStartId="14.[151,190,1544,1563]" captionTargetBox="[209,1062,729,1508]" captionTargetId="figure-11@14.[179,1099,723,1525]" captionTargetPageId="14" captionText="Figs. 20–23. Male genitalia of Cuban species of Copelatus. 20 – C. caelatipennis angustatus author, year; 21 – C. cordovai sp. nov.; 22 – C. cubaensis Schaeffer, 1908; 23 – C. posticatus (Fabricius, 1801). Parts of male genitalia: a – median lobe of aedeagus in left lateral view, b – left paramere in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5331026" httpUri="https://zenodo.org/record/5331026/files/figure.png" pageId="4" pageNumber="387">Fig. 21c</figureCitation>
); female with interstrial space mostly smooth. .... 
<emphasis id="B9171279FFEC4C59FC229D164D73FBAF" bold="true" box="[908,1127,1028,1053]" pageId="4" pageNumber="387">
<taxonomicName id="4C63B5E8FFEC4C59FC229D164D1FFBAF" ID-CoL="8LZPN" authority="Megna &amp; Epler, 2012" authorityName="Megna &amp; Epler" authorityYear="2012" box="[908,1035,1028,1053]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="387" phylum="Arthropoda" rank="species" species="cordovai" status="sp. nov.">
<emphasis id="B9171279FFEC4C59FC229D164D1FFBAF" bold="true" box="[908,1035,1028,1053]" italics="true" pageId="4" pageNumber="387">C. cordovai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A224AF02FFEC4C59FBBF9D164D73FBAF" box="[1041,1127,1029,1053]" pageId="4" pageNumber="387" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BDCCE6BFFEC4C59FF399D364D73FB2F" blockId="4.[151,1127,266,1533]" pageId="4" pageNumber="387">
<keyLead id="30926D5EFFEC4C59FF399D364D73FB2F" pageId="4" pageNumber="387">
- Male protibia without emarginations ventrobasally; median lobe with inner margin strongly concave at midlength (
<figureCitation id="1358D2EEFFEC4C59FDB39D564B6DFBEF" box="[541,633,1093,1117]" captionStart="Figs" captionStartId="11.[113,152,1502,1521]" captionTargetBox="[151,1057,222,1450]" captionTargetId="figure-13@11.[113,1088,216,1465]" captionTargetPageId="11" captionText="Figs. 9–11. Copelatus danyi sp. nov. 9 – female habitus; 10 – male habitus; 11 – male genitalia (a – median lobe in right lateral view; b – same in left lateral view; c – left paramere in lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5331020" httpUri="https://zenodo.org/record/5331020/files/figure.png" pageId="4" pageNumber="387">Fig. 11a</figureCitation>
); parameres narrow, with long setae on inner margin (
<figureCitation id="1358D2EEFFEC4C59FEB09D76486CFBCF" box="[286,376,1125,1149]" captionStart="Figs" captionStartId="11.[113,152,1502,1521]" captionTargetBox="[151,1057,222,1450]" captionTargetId="figure-13@11.[113,1088,216,1465]" captionTargetPageId="11" captionText="Figs. 9–11. Copelatus danyi sp. nov. 9 – female habitus; 10 – male habitus; 11 – male genitalia (a – median lobe in right lateral view; b – same in left lateral view; c – left paramere in lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5331020" httpUri="https://zenodo.org/record/5331020/files/figure.png" pageId="4" pageNumber="387">Fig. 11c</figureCitation>
); female with interstrial space with minute, complex striation (
<figureCitation id="1358D2EEFFEC4C59FBB69D764D4EFBCF" box="[1048,1114,1125,1149]" captionStart="Figs" captionStartId="11.[113,152,1502,1521]" captionTargetBox="[151,1057,222,1450]" captionTargetId="figure-13@11.[113,1088,216,1465]" captionTargetPageId="11" captionText="Figs. 9–11. Copelatus danyi sp. nov. 9 – female habitus; 10 – male habitus; 11 – male genitalia (a – median lobe in right lateral view; b – same in left lateral view; c – left paramere in lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5331020" httpUri="https://zenodo.org/record/5331020/files/figure.png" pageId="4" pageNumber="387">Fig. 9</figureCitation>
). .............................................................................................................. 
<emphasis id="B9171279FFEC4C59FC019D964D73FB2F" bold="true" box="[943,1127,1156,1181]" pageId="4" pageNumber="387">
<taxonomicName id="4C63B5E8FFEC4C59FC019D964D1FFB2F" ID-CoL="8LZ8F" authority="Megna &amp; Epler, 2012" authorityName="Megna &amp; Epler" authorityYear="2012" box="[943,1035,1156,1181]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="387" phylum="Arthropoda" rank="species" species="danyi" status="sp. nov.">
<emphasis id="B9171279FFEC4C59FC019D964D1FFB2F" bold="true" box="[943,1035,1156,1181]" italics="true" pageId="4" pageNumber="387">C. danyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A224AF02FFEC4C59FBBF9D964D73FB2F" box="[1041,1127,1157,1181]" pageId="4" pageNumber="387" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3097D6CEFFEC4C59FF399DB64D73FB4F" pageId="4" pageNumber="387">
<paragraph id="8BDCCE6BFFEC4C59FF399DB64D72FB6F" blockId="4.[151,1127,266,1533]" pageId="4" pageNumber="387">
<keyLead id="30926D5EFFEC4C59FF399DB64D72FB6F" pageId="4" pageNumber="387">
9 Larger, TL = 
<quantity id="4C9B638EFFEC4C59FEFD9DB648C9FB0E" box="[339,477,1189,1213]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" metricValueMax="6.3" metricValueMin="5.9" pageId="4" pageNumber="387" unit="mm" value="6.1" valueMax="6.3" valueMin="5.9">5.9– 6.3 mm</quantity>
; elytra with apical fasciae reduced (
<figureCitation id="1358D2EEFFEC4C59FCF59DB64A8EFB0F" box="[859,922,1189,1213]" captionStart="Figs" captionStartId="6.[151,189,1533,1552]" captionTargetBox="[166,1114,432,1505]" captionTargetId="figure-11@6.[151,1126,417,1515]" captionTargetPageId="6" captionText="Figs.1–8. Habitus and color patterns of Copelatus species from Cuba.1 – C.barbouri Young, 1942; 2 – C. caelatipennis angustatus Chevrolat, 1863; 3 – C. cordovai sp. nov.; 4 – C. cubaensis Schaeffer, 1908; 5 – Copelatus darlingtoni Young, 1942; 6 – C. insolitus Chevrolat, 1863; 7 – C. montivagus Young, 1942; 8 – C. posticatus (Fabricius, 1801)." figureDoi="http://doi.org/10.5281/zenodo.5331018" httpUri="https://zenodo.org/record/5331018/files/figure.png" pageId="4" pageNumber="387">Fig. 1</figureCitation>
). ........................... .............................................................................................. ... 
<taxonomicName id="4C63B5E8FFEC4C59FCF69DD64D72FB6F" ID-CoL="Y349" authority="Young, 1942" authorityName="Young" authorityYear="1942" box="[856,1126,1220,1245]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="387" phylum="Arthropoda" rank="species" species="barbouri">
<emphasis id="B9171279FFEC4C59FCF69DD64AC3FB6F" bold="true" box="[856,983,1220,1245]" italics="true" pageId="4" pageNumber="387">C. barbouri</emphasis>
<bibRefCitation id="EFF2B39AFFEC4C59FC709DD64D72FB6F" author="YOUNG F. N." box="[990,1126,1221,1245]" pageId="4" pageNumber="387" pagination="79 - 94" refId="ref14016" refString="YOUNG F. N. 1942: New species of Copelatus from the West Indies (Coleoptera: Dytiscidae). Proceedings of the New England Zoological Club 20: 79 - 94." type="journal article" year="1942">Young, 1942</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDCCE6BFFEC4C59FF399DF64D73FB4F" blockId="4.[151,1127,266,1533]" box="[151,1127,1253,1277]" pageId="4" pageNumber="387">
<keyLead id="30926D5EFFEC4C59FF399DF64D73FB4F" box="[151,1127,1253,1277]" pageId="4" pageNumber="387">
- Smaller, TL = 
<quantity id="4C9B638EFFEC4C59FEF19DF648F7FB4E" box="[351,483,1253,1277]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" metricValueMax="5.9" metricValueMin="5.3" pageId="4" pageNumber="387" unit="mm" value="5.6" valueMax="5.9" valueMin="5.3">5.3–5.9 mm</quantity>
; elytra with apical fasciae large (
<figureCitation id="1358D2EEFFEC4C59FCEF9DF64ABDFB4F" box="[833,937,1253,1277]" captionStart="Figs" captionStartId="6.[151,189,1533,1552]" captionTargetBox="[166,1114,432,1505]" captionTargetId="figure-11@6.[151,1126,417,1515]" captionTargetPageId="6" captionText="Figs.1–8. Habitus and color patterns of Copelatus species from Cuba.1 – C.barbouri Young, 1942; 2 – C. caelatipennis angustatus Chevrolat, 1863; 3 – C. cordovai sp. nov.; 4 – C. cubaensis Schaeffer, 1908; 5 – Copelatus darlingtoni Young, 1942; 6 – C. insolitus Chevrolat, 1863; 7 – C. montivagus Young, 1942; 8 – C. posticatus (Fabricius, 1801)." figureDoi="http://doi.org/10.5281/zenodo.5331018" httpUri="https://zenodo.org/record/5331018/files/figure.png" pageId="4" pageNumber="387">Figs. 5–7</figureCitation>
). .................... 10
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3097D6CEFFEC4C59FF399C164D73FACF" pageId="4" pageNumber="387">
<paragraph id="8BDCCE6BFFEC4C59FF399C164D72FA8F" blockId="4.[151,1127,266,1533]" pageId="4" pageNumber="387">
<keyLead id="30926D5EFFEC4C59FF399C164D72FA8F" pageId="4" pageNumber="387">
<specimenCount id="9D6505E2FFEC4C59FF399C1649E8FAAF" box="[151,252,1285,1309]" pageId="4" pageNumber="387" type="male">10 Male</specimenCount>
protibiae not modified, their posterior margin straight (
<figureCitation id="1358D2EEFFEC4C59FCE49C164A8DFAAF" box="[842,921,1285,1309]" captionStart="Figs" captionStartId="13.[113,152,1517,1536]" captionTargetBox="[145,1055,494,1485]" captionTargetId="figure-13@13.[133,1069,485,1499]" captionTargetPageId="13" captionText="Figs. 12–19. Male genitalia (12–15) and left protibia (16–19) of Cuban species of Copelatus. 12, 16 – C. barbouri Young, 1942; 13, 17 – C. darlingtoni Young, 1942; 14, 18 – C. insolitus Chevrolat, 1863; 15, 19 – C. montivagus Young, 1942. Parts of male genitalia: a – median lobe of the aedeagus in left lateral view, b – same in ventral lateral view, c – left paramere in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5331022" httpUri="https://zenodo.org/record/5331022/files/figure.png" pageId="4" pageNumber="387">Fig. 17</figureCitation>
). ........................... .............................................................................................. 
<taxonomicName id="4C63B5E8FFEC4C59FC939C364D72FA8F" ID-CoL="Y36P" authority="Young, 1942" authorityName="Young" authorityYear="1942" box="[829,1126,1316,1341]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="387" phylum="Arthropoda" rank="species" species="darlingtoni">
<emphasis id="B9171279FFEC4C59FC939C364AC3FA8F" bold="true" box="[829,983,1316,1341]" italics="true" pageId="4" pageNumber="387">C. darlingtoni</emphasis>
<bibRefCitation id="EFF2B39AFFEC4C59FC709C364D72FA8F" author="YOUNG F. N." box="[990,1126,1317,1341]" pageId="4" pageNumber="387" pagination="79 - 94" refId="ref14016" refString="YOUNG F. N. 1942: New species of Copelatus from the West Indies (Coleoptera: Dytiscidae). Proceedings of the New England Zoological Club 20: 79 - 94." type="journal article" year="1942">Young, 1942</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDCCE6BFFEC4C59FF399C564D73FACF" blockId="4.[151,1127,266,1533]" pageId="4" pageNumber="387">
<keyLead id="30926D5EFFEC4C59FF399C564D73FACF" pageId="4" pageNumber="387">
- Male protibiae modified, their anterior margin weakly to strongly bowed, posterior margin with ventrobasal emargination (
<figureCitation id="1358D2EEFFEC4C59FDB99C764B72FACF" box="[535,614,1381,1405]" captionStart="Figs" captionStartId="13.[113,152,1517,1536]" captionTargetBox="[145,1055,494,1485]" captionTargetId="figure-13@13.[133,1069,485,1499]" captionTargetPageId="13" captionText="Figs. 12–19. Male genitalia (12–15) and left protibia (16–19) of Cuban species of Copelatus. 12, 16 – C. barbouri Young, 1942; 13, 17 – C. darlingtoni Young, 1942; 14, 18 – C. insolitus Chevrolat, 1863; 15, 19 – C. montivagus Young, 1942. Parts of male genitalia: a – median lobe of the aedeagus in left lateral view, b – same in ventral lateral view, c – left paramere in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5331022" httpUri="https://zenodo.org/record/5331022/files/figure.png" pageId="4" pageNumber="387">Fig. 18</figureCitation>
). .................................................................... 11
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3097D6CEFFEC4C59FF399C964D72FA4F" pageId="4" pageNumber="387">
<paragraph id="8BDCCE6BFFEC4C59FF399C964D72FA0F" blockId="4.[151,1127,266,1533]" pageId="4" pageNumber="387">
<keyLead id="30926D5EFFEC4C59FF399C964D72FA0F" pageId="4" pageNumber="387">
11 Elytra fusco-testaceous with base and apex lighter (
<figureCitation id="1358D2EEFFEC4C59FD7D9C964A05FA2F" box="[723,785,1413,1437]" captionStart="Figs" captionStartId="6.[151,189,1533,1552]" captionTargetBox="[166,1114,432,1505]" captionTargetId="figure-11@6.[151,1126,417,1515]" captionTargetPageId="6" captionText="Figs.1–8. Habitus and color patterns of Copelatus species from Cuba.1 – C.barbouri Young, 1942; 2 – C. caelatipennis angustatus Chevrolat, 1863; 3 – C. cordovai sp. nov.; 4 – C. cubaensis Schaeffer, 1908; 5 – Copelatus darlingtoni Young, 1942; 6 – C. insolitus Chevrolat, 1863; 7 – C. montivagus Young, 1942; 8 – C. posticatus (Fabricius, 1801)." figureDoi="http://doi.org/10.5281/zenodo.5331018" httpUri="https://zenodo.org/record/5331018/files/figure.png" pageId="4" pageNumber="387">Fig. 6</figureCitation>
); median lobe with apex strongly curved in lateral aspect (
<figureCitation id="1358D2EEFFEC4C59FE649CB64B31FA0F" box="[458,549,1445,1469]" captionStart="Figs" captionStartId="13.[113,152,1517,1536]" captionTargetBox="[145,1055,494,1485]" captionTargetId="figure-13@13.[133,1069,485,1499]" captionTargetPageId="13" captionText="Figs. 12–19. Male genitalia (12–15) and left protibia (16–19) of Cuban species of Copelatus. 12, 16 – C. barbouri Young, 1942; 13, 17 – C. darlingtoni Young, 1942; 14, 18 – C. insolitus Chevrolat, 1863; 15, 19 – C. montivagus Young, 1942. Parts of male genitalia: a – median lobe of the aedeagus in left lateral view, b – same in ventral lateral view, c – left paramere in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5331022" httpUri="https://zenodo.org/record/5331022/files/figure.png" pageId="4" pageNumber="387">Fig. 14a</figureCitation>
). ..................................... 
<taxonomicName id="4C63B5E8FFEC4C59FC999CB64D72FA0F" ID-CoL="Y39Q" authority="Chevrolat, 1863" authorityName="Chevrolat" authorityYear="1863" box="[823,1126,1444,1469]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="387" phylum="Arthropoda" rank="species" species="insolitus">
<emphasis id="B9171279FFEC4C59FC999CB64AA7FA0F" bold="true" box="[823,947,1444,1469]" italics="true" pageId="4" pageNumber="387">C. insolitus</emphasis>
<bibRefCitation id="EFF2B39AFFEC4C59FC149CB64D72FA0F" author="CHEVROLAT L. A. A." box="[954,1126,1445,1469]" pageId="4" pageNumber="387" pagination="183 - 210" refId="ref12897" refString="CHEVROLAT L. A. A. 1863: Coleopteres de l'IIe de Cuba. Notes, synonymies et descriptions d'especes nouvelles. Familles des cicindeletes, carabiques, dytiscides, gyrinides et palpicornes. Annales de la Societe Entomologique de France 3: 183 - 210." type="journal article" year="1863">Chevrolat, 1863</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDCCE6BFFEC4C59FF399CD64D72FA4F" blockId="4.[151,1127,266,1533]" pageId="4" pageNumber="387">
<keyLead id="30926D5EFFEC4C59FF399CD64D72FA4F" pageId="4" pageNumber="387">
- Elytra fuscous with well-defined basal markings and testaceous apex (
<figureCitation id="1358D2EEFFEC4C59FC3B9CD64AC0FA6F" box="[917,980,1477,1501]" captionStart="Figs" captionStartId="6.[151,189,1533,1552]" captionTargetBox="[166,1114,432,1505]" captionTargetId="figure-11@6.[151,1126,417,1515]" captionTargetPageId="6" captionText="Figs.1–8. Habitus and color patterns of Copelatus species from Cuba.1 – C.barbouri Young, 1942; 2 – C. caelatipennis angustatus Chevrolat, 1863; 3 – C. cordovai sp. nov.; 4 – C. cubaensis Schaeffer, 1908; 5 – Copelatus darlingtoni Young, 1942; 6 – C. insolitus Chevrolat, 1863; 7 – C. montivagus Young, 1942; 8 – C. posticatus (Fabricius, 1801)." figureDoi="http://doi.org/10.5281/zenodo.5331018" httpUri="https://zenodo.org/record/5331018/files/figure.png" pageId="4" pageNumber="387">Fig. 7</figureCitation>
); median lobe with apex slightly curved in lateral aspect (
<figureCitation id="1358D2EEFFEC4C59FD2A9CF64BC8FA4F" box="[644,732,1509,1533]" captionStart="Figs" captionStartId="13.[113,152,1517,1536]" captionTargetBox="[145,1055,494,1485]" captionTargetId="figure-13@13.[133,1069,485,1499]" captionTargetPageId="13" captionText="Figs. 12–19. Male genitalia (12–15) and left protibia (16–19) of Cuban species of Copelatus. 12, 16 – C. barbouri Young, 1942; 13, 17 – C. darlingtoni Young, 1942; 14, 18 – C. insolitus Chevrolat, 1863; 15, 19 – C. montivagus Young, 1942. Parts of male genitalia: a – median lobe of the aedeagus in left lateral view, b – same in ventral lateral view, c – left paramere in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5331022" httpUri="https://zenodo.org/record/5331022/files/figure.png" pageId="4" pageNumber="387">Fig. 15a</figureCitation>
). .......... 
<taxonomicName id="4C63B5E8FFEC4C59FC969CF64D72FA4F" ID-CoL="Y3CG" authority="Young, 1942" authorityName="Young" authorityYear="1942" box="[824,1126,1509,1533]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="387" phylum="Arthropoda" rank="species" species="montivagus">
<emphasis id="B9171279FFEC4C59FC969CF64AC3FA4F" bold="true" box="[824,983,1509,1533]" italics="true" pageId="4" pageNumber="387">C. montivagus</emphasis>
<bibRefCitation id="EFF2B39AFFEC4C59FC709CF64D72FA4F" author="YOUNG F. N." box="[990,1126,1509,1533]" pageId="4" pageNumber="387" pagination="79 - 94" refId="ref14016" refString="YOUNG F. N. 1942: New species of Copelatus from the West Indies (Coleoptera: Dytiscidae). Proceedings of the New England Zoological Club 20: 79 - 94." type="journal article" year="1942">Young, 1942</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<paragraph id="8BDCCE6BFFEC4C59FF319F304BBFF985" blockId="4.[159,683,1571,1591]" box="[159,683,1571,1591]" pageId="4" pageNumber="387">
<keyLead id="30926D5EFFEC4C59FF319F304BBFF985" box="[159,683,1571,1591]" pageId="4" pageNumber="387">
<keyStep id="3097D6CEFFEC4C59FF319F304BBFF985" box="[159,683,1571,1591]" pageId="4" pageNumber="387">
<superScript id="7C166323FFEC4C59FF319F3049B1F99D" attach="left" box="[159,165,1571,1583]" fontSize="5" pageId="4" pageNumber="387">1</superScript>
Identification of isolated females is problematic/impossible.
</keyStep>
</keyLead>
</paragraph>
</subSubSection>
</treatment>
</document>