treatments-xml/data/03/C9/6F/03C96F50FFA6FFCD75DFE74AFDB2C4BB.xml
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<mods:title id="C742C9AE01EDAFC3DC414444BD4C93B3">Late Neogene Lophophaenidae (Nassellaria, Radiolaria) from the eastern equatorial Pacific</mods:title>
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<subSubSection id="C37A8DCDFFA6FFCD75DFE74AFE27C02B" box="[151,480,829,855]" pageId="53" pageNumber="54" type="nomenclature">
<paragraph id="8BDFDE46FFA6FFCD75DFE74AFE27C02B" blockId="53.[151,480,829,891]" box="[151,480,829,855]" pageId="53" pageNumber="54">
<taxonomicName id="4C60A5C5FFA6FFCD75DFE74AFE51C02B" authority="Trubovitz, Renaudie, Lazarus &amp; Noble, 2022" box="[151,406,829,855]" class="Polycystina" family="Plagiacanthidae" genus="Lophophaena" kingdom="Chromista" order="Nassellaria" pageId="53" pageNumber="54" phylum="Radiozoa" rank="species" species="casperi" status="sp. nov.">
<emphasis id="B9140254FFA6FFCD75DFE74AFE51C02B" box="[151,406,829,855]" italics="true" pageId="53" pageNumber="54">Lophophaena casperi</emphasis>
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<emphasis id="B9140254FFA6FFCD74D6E74AFE27C02B" bold="true" box="[414,480,829,855]" pageId="53" pageNumber="54">
<taxonomicNameLabel id="A227BF2FFFA6FFCD74D6E74AFE27C02B" box="[414,480,829,855]" pageId="53" pageNumber="54" rank="species">n. sp.</taxonomicNameLabel>
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<subSubSection id="C37A8DCDFFA6FFCD75DFE716FCB6C0C1" pageId="53" pageNumber="54" type="description">
<paragraph id="8BDFDE46FFA6FFCD75DFE716FE50C007" blockId="53.[151,480,829,891]" box="[151,407,865,891]" pageId="53" pageNumber="54">
Plate 28,
<figureCitation id="135BC2C3FFA6FFCD7448E716FE98C007" box="[256,351,865,891]" captionStart="FIGURE 1" captionStartId="6.[151,250,1340,1364]" captionTargetBox="[151,1430,743,1316]" captionTargetId="figure-284@6.[151,1436,741,1317]" captionTargetPageId="6" captionText="FIGURE 1. Diagram illustrating the general skeletal morphology of Lophophaenidae, and terminology used in this manuscript. Dashed lines indicate spines that are not visible from the angle shown, as they extend into the skeleton. An example specimen of Lophophaena casperi n. sp. is included for comparison to the diagrams." figureDoi="http://doi.org/10.5281/zenodo.10544060" httpUri="https://zenodo.org/record/10544060/files/figure.png" pageId="53" pageNumber="54">Figs. 1A</figureCitation>
<figureCitation id="135BC2C3FFA6FFCD7438E716FE54C007" box="[368,403,865,891]" captionStart="FIGURE 9" captionStartId="30.[151,250,1556,1580]" captionTargetBox="[152,1436,742,1532]" captionTargetId="figure-252@30.[151,1436,741,1532]" captionTargetPageId="30" captionText="FIGURE 9. Internal skeletal structure of Ceratocyrtis sp. C." figureDoi="http://doi.org/10.5281/zenodo.10544076" httpUri="https://zenodo.org/record/10544076/files/figure.png" pageId="53" pageNumber="54">9B</figureCitation>
.
</paragraph>
<paragraph id="8BDFDE46FFA6FFCD75DFE7D2FCB6C0C1" blockId="53.[151,957,933,1019]" box="[151,881,933,957]" pageId="53" pageNumber="54">
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urn:lsid:zoobank.org:act:
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<subSubSection id="C37A8DCDFFA6FFCD75DFE7B3FC7AC086" pageId="53" pageNumber="54" type="reference_group">
<paragraph id="8BDFDE46FFA6FFCD75DFE7B3FD14C0A0" blockId="53.[151,957,933,1019]" box="[151,723,964,988]" pageId="53" pageNumber="54">
?
<taxonomicName id="4C60A5C5FFA6FFCD75E1E7B3FEA8C0A0" box="[169,367,964,988]" genus="Tripodocorys" kingdom="Chromista" pageId="53" pageNumber="54" phylum="Radiozoa" rank="species" species="undefined-a">
<emphasis id="B9140254FFA6FFCD75E1E7B3FEF7C0A0" box="[169,304,964,988]" italics="true" pageId="53" pageNumber="54">Tripodocorys</emphasis>
sp. A
</taxonomicName>
.,
<bibRefCitation id="EFF1A3B7FFA6FFCD7435E7B3FDE3C0A0" author="Sugiyama, K." box="[381,548,964,988]" pageId="53" pageNumber="54" pagination="1 - 11" refId="ref95347" refString="Sugiyama, K. (1994) Lower Miocene New Nassellarians (Radiolaria) from the Toyohama Formation, Morozaki Group, Central Japan. Bulletin of the Mizunami Fossil Museum, 21, 1 - 11." type="journal article" year="1994">Sugiyama, 1994</bibRefCitation>
, pl. 5, figs. 3ad.
</paragraph>
<paragraph id="8BDFDE46FFA6FFCD75DFE794FC7AC086" blockId="53.[151,957,933,1019]" box="[151,957,995,1019]" pageId="53" pageNumber="54">
unknown plagonid group C sp 28,
<bibRefCitation id="EFF1A3B7FFA6FFCD74BEE794FD13C087" author="Trubovitz, S. &amp; Lazarus, D. &amp; Renaudie, J. &amp; Noble, P. J." box="[502,724,995,1019]" pageId="53" pageNumber="54" pagination="1 - 10" refId="ref96304" refString="Trubovitz, S., Lazarus, D., Renaudie, J. &amp; Noble, P. J. (2020) Marine plankton show threshold extinction response to Neogene climate change. Nature Communications, 11 (5069), 1 - 10. https: // doi. org / 10.1038 / s 41467 - 020 - 18879 - 7" type="journal article" year="2020">
Trubovitz
<emphasis id="B9140254FFA6FFCD7717E794FD5FC087" box="[607,664,995,1019]" italics="true" pageId="53" pageNumber="54">et al.,</emphasis>
2020
</bibRefCitation>
, supplementary data 7.
</paragraph>
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<subSubSection id="C37A8DCDFFA6FFCD75DFE05EFE0EC71B" pageId="53" pageNumber="54" type="diagnosis">
<paragraph id="8BDFDE46FFA6FFCD75DFE05EFE0EC71B" blockId="53.[151,1437,1065,1991]" pageId="53" pageNumber="54">
<emphasis id="B9140254FFA6FFCD75DFE05EFED5C73F" bold="true" box="[151,274,1065,1091]" pageId="53" pageNumber="54">Diagnosis.</emphasis>
<taxonomicName id="4C60A5C5FFA6FFCD7451E05EFE74C73F" authorityName="Ehrenberg, 1847 emend. Petrushevskaya" authorityYear="1971" box="[281,435,1065,1091]" class="Polycystina" family="Plagiacanthidae" genus="Lophophaena" kingdom="Chromista" order="Nassellaria" pageId="53" pageNumber="54" phylum="Radiozoa" rank="genus">
<emphasis id="B9140254FFA6FFCD7451E05EFE74C73F" box="[281,435,1065,1091]" italics="true" pageId="53" pageNumber="54">Lophophaena</emphasis>
</taxonomicName>
with an hourglass shape in outline, exhibiting many thin spines around the cephalis and the shoulders of the thorax.
</paragraph>
</subSubSection>
<subSubSection id="C37A8DCDFFA6FFCD7586E006FBEFC61F" pageId="53" pageNumber="54" type="description">
<paragraph id="8BDFDE46FFA6FFCD7586E006FBEFC61F" blockId="53.[151,1437,1065,1991]" pageId="53" pageNumber="54">
<emphasis id="B9140254FFA6FFCD7586E006FEA7C7F7" bold="true" box="[206,352,1137,1163]" pageId="53" pageNumber="54">Description.</emphasis>
This species has a high, balloon- shaped cephalis that reaches its widest point about ⅔ of the way up. From the circumference of this wide point, many thin spines extend upward from the cephalis. Pores on the cephalis are small and round. The neck area is proportionally narrow, and includes several short, thin spines in addition to the horizontally-orientated ventral spine. The thorax flares widely at the shoulders, contributing to the skeletons overall hourglass shape. When well-preserved, many thin spines extend down and outward from the widest point of the shoulders. Below the shoulders, the thorax wall tends to become thinner and have smaller, more irregular pores. The base of the thorax is smooth, lacking teeth at the termination.
</paragraph>
</subSubSection>
<subSubSection id="C37A8DCDFFA6FFCD7586E11AFAC2C68F" pageId="53" pageNumber="54" type="discussion">
<paragraph id="8BDFDE46FFA6FFCD7586E11AFAC2C68F" blockId="53.[151,1437,1065,1991]" pageId="53" pageNumber="54">
<emphasis id="B9140254FFA6FFCD7586E11AFE85C6FB" bold="true" box="[206,322,1389,1415]" pageId="53" pageNumber="54">Remarks.</emphasis>
This species shows a relatively high amount of size variation, but the shell shape and proportions are consistent. It differs from
<taxonomicName id="4C60A5C5FFA6FFCD74AEE1E6FD2FC6D7" box="[486,744,1425,1451]" class="Polycystina" family="Plagiacanthidae" genus="Lophophaena" kingdom="Chromista" order="Nassellaria" pageId="53" pageNumber="54" phylum="Radiozoa" rank="species" species="rusalkae" status="sp. nov.">
<emphasis id="B9140254FFA6FFCD74AEE1E6FD2FC6D7" box="[486,744,1425,1451]" italics="true" pageId="53" pageNumber="54">Lophophaena rusalkae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A227BF2FFFA6FFCD77B8E1E6FCEBC6D7" box="[752,812,1425,1451]" pageId="53" pageNumber="54" rank="species">n. sp.</taxonomicNameLabel>
(Pl. 29,
<figureCitation id="135BC2C3FFA6FFCD76C6E1E6FC2AC6D7" box="[910,1005,1425,1451]" captionStart="FIGURE 1" captionStartId="6.[151,250,1340,1364]" captionTargetBox="[151,1430,743,1316]" captionTargetId="figure-284@6.[151,1436,741,1317]" captionTargetPageId="6" captionText="FIGURE 1. Diagram illustrating the general skeletal morphology of Lophophaenidae, and terminology used in this manuscript. Dashed lines indicate spines that are not visible from the angle shown, as they extend into the skeleton. An example specimen of Lophophaena casperi n. sp. is included for comparison to the diagrams." figureDoi="http://doi.org/10.5281/zenodo.10544060" httpUri="https://zenodo.org/record/10544060/files/figure.png" pageId="53" pageNumber="54">Figs. 1A</figureCitation>
<figureCitation id="135BC2C3FFA6FFCD76B5E1E6FBCCC6D7" box="[1021,1035,1425,1451]" captionStart="FIGURE 6" captionStartId="15.[151,250,1463,1487]" captionTargetBox="[153,1436,737,1439]" captionTargetId="figure-139@15.[151,1436,736,1440]" captionTargetPageId="15" captionText="FIGURE 6. Internal skeletal structure of Arachnocorallium calvata (Haeckel, 1887) Petrushevskaya, 1971." figureDoi="http://doi.org/10.5281/zenodo.10544070" httpUri="https://zenodo.org/record/10544070/files/figure.png" pageId="53" pageNumber="54">6</figureCitation>
) in its larger overall size, and more pronounced hourglass shape, with a long and narrow neck, and high, inflated cephalis.
<figureCitation id="135BC2C3FFA6FFCD7116E1C2FB0DC6B3" box="[1118,1226,1461,1487]" captionStart="FIGURE 12" captionStartId="54.[151,250,1031,1055]" captionTargetBox="[162,1427,188,1002]" captionTargetId="figure-22@54.[155,1432,181,1007]" captionTargetPageId="54" captionText="FIGURE 12. Chart illustrating the different cephalis dimensions of Lophophaena casperi n. sp. (blue squares) and L. rusalkae n. sp. (red triangles). L. rusalkae has a cephalis height ranging from 34-40 microns, and an average cephalis width:neck ratio of 1. By contrast, L. casperi has a higher cephalis, ranging from 44- 58 microns, and a cephalis width:neck ratio averaging 1.3. The specimen with a cephalis width:neck ratio of 1.7, which is visually the outlier on this chart, has a particularly narrow neck (23 microns), but a cephalis of typical width for this species (40 microns)." figureDoi="http://doi.org/10.5281/zenodo.10544080" httpUri="https://zenodo.org/record/10544080/files/figure.png" pageId="53" pageNumber="54">Figure 12</figureCitation>
illustrates how the cephalis dimensions are different between these two species, which can appear similar at first glance.
</paragraph>
</subSubSection>
<subSubSection id="C37A8DCDFFA6FFCD758FE18AFBECC44B" pageId="53" pageNumber="54" type="materials_examined">
<paragraph id="8BDFDE46FFA6FFCD758FE18AFB60C547" blockId="53.[151,1437,1065,1991]" pageId="53" pageNumber="54">
<emphasis id="B9140254FFA6FFCD758FE18AFE75C56B" bold="true" box="[199,434,1533,1559]" pageId="53" pageNumber="54">Material examined.</emphasis>
<specimenCount id="9D6615CFFFA6FFCD74F3E18AFD93C56B" box="[443,596,1533,1560]" pageId="53" pageNumber="54" type="generic">85 specimens</specimenCount>
observed from Late Miocene samples 321-1337A-21H-1,
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, 321- 1337A-18H-6,
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, 321-1337A-16H-
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, and 321-1337A-14H-7,
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.
</paragraph>
<paragraph id="8BDFDE46FFA6FFCD758FE232FB45C523" blockId="53.[151,1437,1065,1991]" box="[199,1154,1605,1632]" pageId="53" pageNumber="54">
<materialsCitation id="3B08D41BFFA6FFCD758FE232FBBAC523" box="[199,1149,1605,1632]" pageId="53" pageNumber="54" specimenCode="321-1337A-21H-1" specimenCount="1" typeStatus="holotype">
<emphasis id="B9140254FFA6FFCD758FE232FEFCC523" bold="true" box="[199,315,1605,1631]" pageId="53" pageNumber="54">
<typeStatus id="54DB60E4FFA6FFCD758FE232FEF1C523" box="[199,310,1605,1631]" pageId="53" pageNumber="54" type="holotype">Holotype</typeStatus>
.
</emphasis>
Pl. 28, figs. 1AB; sample
<specimenCode id="DBC6763DFFA6FFCD7739E232FC83C523" box="[625,836,1605,1631]" pageId="53" pageNumber="54">321-1337A-21H-1</specimenCode>
,
<quantity id="4C9873A3FFA6FFCD7618E232FC7BC51C" box="[848,956,1605,1632]" metricMagnitude="-1" metricUnit="m" metricValue="3.4" metricValueMax="3.5" metricValueMin="3.3" pageId="53" pageNumber="54" unit="cm" value="34.0" valueMax="35.0" valueMin="33.0">3335cm</quantity>
; ECO-144; G3-1
</materialsCitation>
.
</paragraph>
<paragraph id="8BDFDE46FFA6FFCD758FE21EFBECC44B" blockId="53.[151,1437,1065,1991]" pageId="53" pageNumber="54">
<emphasis id="B9140254FFA6FFCD758FE21EFE81C5FF" bold="true" box="[199,326,1641,1667]" pageId="53" pageNumber="54">
<typeStatus id="54DB60E4FFA6FFCD758FE21EFE86C5FF" box="[199,321,1641,1667]" pageId="53" pageNumber="54" type="paratype">Paratypes</typeStatus>
.
</emphasis>
(1) Pl. 28, figs. 4AB; sample 321-1337A-21H-1,
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; ECO-144; S16-2. (2) Pl. 28, fig. 2; sample 321-1337A-14H-7,
<quantity id="4C9873A3FFA6FFCD749AE2FAFDF9C5DB" box="[466,574,1677,1703]" metricMagnitude="-1" metricUnit="m" metricValue="4.05" metricValueMax="4.2" metricValueMin="3.9" pageId="53" pageNumber="54" unit="cm" value="40.5" valueMax="42.0" valueMin="39.0">3942cm</quantity>
; ECO-139; W39-4. (3) Pl. 28, figs. 8AB; sample 321-1337A-14H-7,
<quantity id="4C9873A3FFA6FFCD703AE2FAFF1EC5B7" metricMagnitude="-1" metricUnit="m" metricValue="4.05" metricValueMax="4.2" metricValueMin="3.9" pageId="53" pageNumber="54" unit="cm" value="40.5" valueMax="42.0" valueMin="39.0">39 42cm</quantity>
; ECO-138; Q30-1. (4) Pl. 28, figs. 7AB; sample 321-1337A-16H-
<quantity id="4C9873A3FFA6FFCD769BE2C6FBB3C5B7" box="[979,1140,1713,1739]" metricMagnitude="-1" metricUnit="m" metricValue="6.50605" metricValueMax="12.4" metricValueMin="0.6121" pageId="53" pageNumber="54" unit="cm" value="65.0605" valueMax="124.0" valueMin="6.121">6, 121124cm</quantity>
; ECO-141; W22-1. (5) Pl. 28, figs. 5AB; sample 321-1337A-16H-
<quantity id="4C9873A3FFA6FFCD7725E2A2FCC8C593" box="[621,783,1749,1775]" metricMagnitude="-1" metricUnit="m" metricValue="6.50605" metricValueMax="12.4" metricValueMin="0.6121" pageId="53" pageNumber="54" unit="cm" value="65.0605" valueMax="124.0" valueMin="6.121">6, 121124cm</quantity>
; ECO-142; M17-1. (6) Pl. 28, fig. 3; sample 321-1337A-16H-
<quantity id="4C9873A3FFA6FFCD759BE28EFEB1C46F" box="[211,374,1785,1811]" metricMagnitude="-1" metricUnit="m" metricValue="6.50605" metricValueMax="12.4" metricValueMin="0.6121" pageId="53" pageNumber="54" unit="cm" value="65.0605" valueMax="124.0" valueMin="6.121">6, 121124cm</quantity>
; ECO-141; J15-4. (7) Pl. 28, figs. 6AB; sample 321-1337A-16H-
<quantity id="4C9873A3FFA6FFCD7131E28EFADCC46F" box="[1145,1307,1785,1811]" metricMagnitude="-1" metricUnit="m" metricValue="6.50605" metricValueMax="12.4" metricValueMin="0.6121" pageId="53" pageNumber="54" unit="cm" value="65.0605" valueMax="124.0" valueMin="6.121">6, 121124cm</quantity>
; ECO-141; H39-4. (8) [not figured] sample 321-1337A-16H-
<quantity id="4C9873A3FFA6FFCD778DE36AFCA1C44B" box="[709,870,1821,1847]" metricMagnitude="-1" metricUnit="m" metricValue="6.50605" metricValueMax="12.4" metricValueMin="0.6121" pageId="53" pageNumber="54" unit="cm" value="65.0605" valueMax="124.0" valueMin="6.121">6, 121124cm</quantity>
; ECO-141; C2-2.
</paragraph>
</subSubSection>
<subSubSection id="C37A8DCDFFA6FFCD7586E336FC4DC403" pageId="53" pageNumber="54" type="description">
<paragraph id="8BDFDE46FFA6FFCD7586E336FC4DC403" blockId="53.[151,1437,1065,1991]" pageId="53" pageNumber="54">
<emphasis id="B9140254FFA6FFCD7586E336FE42C427" bold="true" box="[206,389,1857,1883]" pageId="53" pageNumber="54">Measurements.</emphasis>
Height of cephalis 4458 (52)μm; width of cephalis 3041 (38)μm; width at neck 2333 (29)μm; width at shoulders 4252 (48)μm. Based on
<specimenCount id="9D6615CFFFA6FFCD77A3E312FC41C403" box="[747,902,1893,1919]" pageId="53" pageNumber="54" type="generic">16 specimens</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection id="C37A8DCDFFA6FFCD7586E3FEFB25C4DF" box="[206,1250,1929,1955]" pageId="53" pageNumber="54" type="etymology">
<paragraph id="8BDFDE46FFA6FFCD7586E3FEFB25C4DF" blockId="53.[151,1437,1065,1991]" box="[206,1250,1929,1955]" pageId="53" pageNumber="54">
<emphasis id="B9140254FFA6FFCD7586E3FEFE92C4DF" bold="true" box="[206,341,1929,1955]" pageId="53" pageNumber="54">Etymology.</emphasis>
Named for Casper the friendly ghost, a mid-century American cartoon character.
</paragraph>
</subSubSection>
<subSubSection id="C37A8DCDFFA6FFCD7586E3DAFDB2C4BB" box="[206,629,1965,1991]" pageId="53" pageNumber="54" type="biology_ecology">
<paragraph id="8BDFDE46FFA6FFCD7586E3DAFDB2C4BB" blockId="53.[151,1437,1065,1991]" box="[206,629,1965,1991]" pageId="53" pageNumber="54">
<emphasis id="B9140254FFA6FFCD7586E3DAFEE6C4BB" bold="true" box="[206,289,1965,1991]" pageId="53" pageNumber="54">Range.</emphasis>
Late Miocene, EEP (Table 1).
</paragraph>
</subSubSection>
</treatment>
</document>