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<mods:title id="880D5D3C6CCCF0A2652E49A75A9D550A">New synonymies of Coprophilus (Zonyptilus) pentatoma Fauvel, 1897 (Coleoptera: Staphylinidae: Oxytelinae)</mods:title>
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<treatment id="03BC87D2CA1FBA6F1BECCBED0958FEB5" ID-DOI="http://doi.org/10.5281/zenodo.6000216" ID-GBIF-Taxon="132650202" ID-Zenodo-Dep="6000216" LSID="urn:lsid:plazi:treatment:03BC87D2CA1FBA6F1BECCBED0958FEB5" httpUri="http://treatment.plazi.org/id/03BC87D2CA1FBA6F1BECCBED0958FEB5" lastPageId="6" lastPageNumber="135" pageId="4" pageNumber="134">
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<paragraph id="8BAA36C4CA1FBA6D1BECCBED0B3FFD49" blockId="4.[151,883,680,741]" box="[151,883,680,707]" pageId="4" pageNumber="134">
<heading id="D0E281A8CA1FBA6D1BECCBED0B3FFD49" bold="true" box="[151,883,680,707]" fontSize="11" level="1" pageId="4" pageNumber="134" reason="1">
<emphasis id="B961EAD6CA1FBA6D1BECCBED0B3FFD49" bold="true" box="[151,883,680,707]" pageId="4" pageNumber="134">
<taxonomicName id="4C154D47CA1FBA6D1BECCBED0B59FD49" authority="Kashcheev, 1999" authorityName="Kashcheev" authorityYear="1999" box="[151,789,680,707]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="134" phylum="Arthropoda" rank="species" species="rufitarsis" subGenus="Zonyptilus">
<emphasis id="B961EAD6CA1FBA6D1BECCBED0A71FD48" bold="true" box="[151,573,680,706]" italics="true" pageId="4" pageNumber="134">Coprophilus (Zonyptilus) rufitarsis</emphasis>
<bibRefCitation id="EF844B35CA1FBA6D193ECBEC0B59FD49" author="Kashcheev" box="[581,789,681,707]" pageId="4" pageNumber="134" refString="Kashcheev, V. A. (1999) Novye Oxytelinae (Coleoptera, Staphylinidae) Srednei Azii i Kazakhstana. Tethys Entomological Research, 1, 141 - 156. [in Russian]" type="journal article" year="1999">Kashcheev, 1999</bibRefCitation>
</taxonomicName>
,
<taxonomicNameLabel id="A25257ADCA1FBA6D1859CBEC0B3FFD49" box="[802,883,681,707]" pageId="4" pageNumber="134">syn. n.</taxonomicNameLabel>
</emphasis>
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<paragraph id="8BAA36C4CA1FBA6D1BECCB89096CFD6E" blockId="4.[151,883,680,741]" box="[151,288,716,741]" pageId="4" pageNumber="134">
(
<figureCitation id="132E2A41CA1FBA6D1BE4CB8908A2FD6F" box="[159,238,716,741]" captionStart="FIGURES 6 7" captionStartId="4.[151,264,1982,2005]" captionTargetBox="[164,1422,949,1948]" captionTargetId="figure@4.[151,1437,934,1961]" captionTargetPageId="4" captionText="FIGURES 6 7. Coprophilus spp.: 6 — C. (Z.) rufitarsis Kashcheev, 1999 (holotype?, female), body, dorsal view; 7 — C. (Z.) pentatoma Fauvel, 1897 (“ Kebin ”, female), body, dorsal view. Scale bar: figs. 6, 7 — 1 mm." httpUri="https://zenodo.org/record/843169/files/figure.png" pageId="4" pageNumber="134">Figs. 6</figureCitation>
, 13)
</paragraph>
</subSubSection>
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<paragraph id="8BAA36C4CA1FBA6D1BECCA510A19FCC2" blockId="4.[151,851,788,902]" pageId="4" pageNumber="134">
<treatmentCitationGroup id="AB0511EACA1FBA6D1BECCA510A19FCC2" pageId="4" pageNumber="134">
<taxonomicName id="4C154D47CA1FBA6D1BECCA510AADFCA0" authority="Kashcheev, 1999: 149" authorityName="Kashcheev" authorityPageNumber="149" authorityYear="1999" box="[151,737,788,810]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="134" phylum="Arthropoda" rank="species" species="rufitarsis" subGenus="Zonoptilus">
<emphasis id="B961EAD6CA1FBA6D1BECCA5109B7FCA0" box="[151,507,788,810]" italics="true" pageId="4" pageNumber="134">Coprophilus (Zonoptilus) rufitarsis</emphasis>
<treatmentCitation id="0AB410D5CA1FBA6D197ACA510AADFCA0" author="Kashcheev" box="[513,737,788,810]" page="149" pageId="4" pageNumber="134" year="1999">
<bibRefCitation id="EF844B35CA1FBA6D197ACA510AFDFCA0" author="Kashcheev" box="[513,689,788,810]" pageId="4" pageNumber="134" refString="Kashcheev, V. A. (1999) Novye Oxytelinae (Coleoptera, Staphylinidae) Srednei Azii i Kazakhstana. Tethys Entomological Research, 1, 141 - 156. [in Russian]" type="journal article" year="1999">Kashcheev, 1999</bibRefCitation>
: 149
</treatmentCitation>
</taxonomicName>
<taxonomicName id="4C154D47CA1FBA6D1BECCA770A19FCC2" authority=": Herman, 2001: 1316" authorityName="Herman" authorityPageNumber="1316" authorityYear="2001" box="[151,597,818,840]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="134" phylum="Arthropoda" rank="species" species="rufitarsis">
<emphasis id="B961EAD6CA1FBA6D1BECCA770934FCC2" box="[151,376,818,840]" italics="true" pageId="4" pageNumber="134">Coprophilus rufitarsis</emphasis>
:
<treatmentCitation id="0AB410D5CA1FBA6D1AFECA760A19FCC2" author="Herman" box="[389,597,818,840]" page="1316" pageId="4" pageNumber="134" year="2001">
<bibRefCitation id="EF844B35CA1FBA6D1AFECA760A55FCC2" author="Herman" box="[389,537,818,840]" pageId="4" pageNumber="134" refString="Herman, L. H. (2001) Catalog of the Staphylinidae (Insecta: Coleoptera). 1785 to the end of the second millennium. Bulletin of the American Museum of Natural History, 265, 1309 - 1319." type="journal article" year="2001">Herman, 2001</bibRefCitation>
: 1316
</treatmentCitation>
</taxonomicName>
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</paragraph>
<paragraph id="8BAA36C4CA1FBA6C1BECCA140A85F9EA" blockId="4.[151,851,788,902]" lastBlockId="5.[151,1437,1355,2029]" lastPageId="5" lastPageNumber="135" pageId="4" pageNumber="134">
<treatmentCitationGroup id="AB0511EACA1FBA6C1BECCA140A85F9EA" infered="true" lastPageId="5" lastPageNumber="135" pageId="4" pageNumber="134">
<taxonomicName id="4C154D47CA1FBA6D1BECCA140A9CFCED" authority=": Smetana, 2004: 511" authorityName="Smetana" authorityPageNumber="511" authorityYear="2004" box="[151,720,849,871]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="134" phylum="Arthropoda" rank="species" species="rufitarsis" subGenus="Zonyptilus">
<emphasis id="B961EAD6CA1FBA6D1BECCA1409B5FCED" box="[151,505,849,871]" italics="true" pageId="4" pageNumber="134">Coprophilus (Zonyptilus) rufitarsis</emphasis>
:
<treatmentCitation id="0AB410D5CA1FBA6D197CCA140A9CFCED" author="Smetana" box="[519,720,849,871]" page="511" pageId="4" pageNumber="134" year="2004">
<bibRefCitation id="EF844B35CA1FBA6D197CCA140AEEFCED" author="Smetana" box="[519,674,849,871]" pageId="4" pageNumber="134" refString="Smetana, A. (2004) Oxytelinae. In Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera, Hydrophiloidea- Histeroidea-Staphylinoidea, 5 (2), 511 - 535. [Apollo Books, Stenstrup]" type="journal article" year="2004">Smetana, 2004</bibRefCitation>
: 511
</treatmentCitation>
</taxonomicName>
<taxonomicName id="4C154D47CA1FBA6D1BECCA350B1FFC0C" authority=": Schulke and Smetana, 2015: 767" authorityName="Schulke and Smetana" authorityPageNumber="767" authorityYear="2015" box="[151,851,880,902]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="134" phylum="Arthropoda" rank="species" species="rufitarsis" subGenus="Zonyptilus">
<emphasis id="B961EAD6CA1FBA6D1BECCA3509B5FC0C" box="[151,505,880,902]" italics="true" pageId="4" pageNumber="134">Coprophilus (Zonyptilus) rufitarsis</emphasis>
:
<treatmentCitation id="0AB410D5CA1FBA6D197CCA350B1FFC0C" author="Schulke" box="[519,851,880,902]" page="767" pageId="4" pageNumber="134" year="2015">
<bibRefCitation id="EF844B35CA1FBA6D197CCA350B68FC0C" author="Schulke" box="[519,804,880,902]" pageId="4" pageNumber="134" refString="Schulke, M. &amp; Smetana, A. (2015) Oxytelinae. In Lobl, I. &amp; Lobl, D. (Ed.), Catalogue of Palaearctic Coleoptera, Hydrophiloidea-Staphylinoidea, 5 (1), pp. 757 - 795. [Brill, Leiden &amp; Boston]" type="journal article" year="2015">Schülke and Smetana, 2015</bibRefCitation>
: 767
</treatmentCitation>
</taxonomicName>
<emphasis id="B961EAD6CA1EBA6C1BECCC0E098FFAEE" bold="true" box="[151,451,1355,1380]" pageId="5" pageNumber="135">Type material examined.</emphasis>
Paratypes: 1 ♀, Kazakhstan “[in Russian] Aksu Dzhabagly
<date id="FFAB1004CA1EBA6C1F01CC0E0CA6FAEE" box="[1146,1258,1355,1380]" pageId="5" pageNumber="135" value="1985-06-22">22.6.1985</date>
V. Kashcheev” “Holotypus
<taxonomicName id="4C154D47CA1EBA6C1A5DCC2A0AE0FA02" authority="Kastcheev" authorityName="Kastcheev" box="[294,684,1391,1416]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="135" phylum="Arthropoda" rank="species" species="rufitarsis">
<emphasis id="B961EAD6CA1EBA6C1A5DCC2A0A6BFA02" box="[294,551,1391,1416]" italics="true" pageId="5" pageNumber="135">Coprophilus rufitarsis</emphasis>
Kastcheev
</taxonomicName>
” “
<taxonomicName id="4C154D47CA1EBA6C19A9CC2A0D6DFA02" ID-CoL="5ZYJ6" authority="Fauvel, 1897" authorityName="Fauvel" authorityYear="1897" box="[722,1313,1391,1416]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="135" phylum="Arthropoda" rank="species" species="pentatoma" subGenus="Zonyptilus">
<emphasis id="B961EAD6CA1EBA6C19A9CC2A0C35FA02" box="[722,1145,1391,1416]" italics="true" pageId="5" pageNumber="135">Coprophilus (Zonyptilus) pentatoma</emphasis>
<bibRefCitation id="EF844B35CA1EBA6C1FFCCC2A0D6DFA02" author="Fauvel" box="[1159,1313,1391,1416]" pageId="5" pageNumber="135" refString="Fauvel, A. (1897) Tableau des especes palearctiques du genre Coprophilus Latr. (Zonoptilus Mots. - Homalotrichus Sol.). Revue d'Entomologie, 16, 226 - 229." type="journal article" year="1897">Fauvel, 1897</bibRefCitation>
</taxonomicName>
| det. M. Gildenkov, 2014” “ZOOLOGICAL INSTITUTE RAS ST. PETERSBURG” (ZMAS); 1 ♀ “[in Russian] Aksu Dzhabagly
<date id="FFAB1004CA1EBA6C1A66CCF209C0FA5A" box="[285,396,1463,1488]" pageId="5" pageNumber="135" value="1985-06-22">22.6.1985</date>
V. Kashcheev” “Paratypus
<taxonomicName id="4C154D47CA1EBA6C19ACCCF20C15FA5A" authority="Kastcheev" authorityName="Kastcheev" box="[727,1113,1463,1488]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="135" phylum="Arthropoda" rank="species" species="rufitarsis">
<emphasis id="B961EAD6CA1EBA6C19ACCCF20B9AFA5A" box="[727,982,1463,1488]" italics="true" pageId="5" pageNumber="135">Coprophilus rufitarsis</emphasis>
Kastcheev
</taxonomicName>
” “
<taxonomicName id="4C154D47CA1EBA6C1F07CCF209E3FA7E" ID-CoL="5ZYJ6" authority="Fauvel, 1897" authorityName="Fauvel" authorityYear="1897" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="135" phylum="Arthropoda" rank="species" species="pentatoma" subGenus="Zonyptilus">
<emphasis id="B961EAD6CA1EBA6C1F07CCF20941FA7E" italics="true" pageId="5" pageNumber="135">Coprophilus (Zonyptilus) pentatoma</emphasis>
<bibRefCitation id="EF844B35CA1EBA6C1A63CC9E09E3FA7E" author="Fauvel" box="[280,431,1499,1524]" pageId="5" pageNumber="135" refString="Fauvel, A. (1897) Tableau des especes palearctiques du genre Coprophilus Latr. (Zonoptilus Mots. - Homalotrichus Sol.). Revue d'Entomologie, 16, 226 - 229." type="journal article" year="1897">Fauvel, 1897</bibRefCitation>
</taxonomicName>
| det. M. Gildenkov, 2014” “ZOOLOGICAL INSTITUTE RAS ST. PETERSBURG” (ZMAS); 5 ♀♀ “[in Russian] Aksu Dzhabagly
<date id="FFAB1004CA1EBA6C19A2CCBA0B04F992" box="[729,840,1535,1560]" pageId="5" pageNumber="135" value="1985-06-22">22.6.1985</date>
V. Kashcheev” “Paratypus
<taxonomicName id="4C154D47CA1EBA6C1FECCCBA0942F9B6" authority="Kastcheev" authorityName="Kastcheev" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="135" phylum="Arthropoda" rank="species" species="rufotarsis">
<emphasis id="B961EAD6CA1EBA6C1FECCCBA0DD0F992" box="[1175,1436,1535,1560]" italics="true" pageId="5" pageNumber="135">Coprophilus rufotarsis</emphasis>
Kastcheev
</taxonomicName>
” “
<taxonomicName id="4C154D47CA1EBA6C1A49CF660B31F9B6" ID-CoL="5ZYJ6" authority="Fauvel, 1897" authorityName="Fauvel" authorityYear="1897" box="[306,893,1571,1596]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="135" phylum="Arthropoda" rank="species" species="pentatoma" subGenus="Zonyptilus">
<emphasis id="B961EAD6CA1EBA6C1A49CF660A9AF9B6" box="[306,726,1571,1596]" italics="true" pageId="5" pageNumber="135">Coprophilus (Zonyptilus) pentatoma</emphasis>
<bibRefCitation id="EF844B35CA1EBA6C199FCF660B31F9B6" author="Fauvel" box="[740,893,1571,1596]" pageId="5" pageNumber="135" refString="Fauvel, A. (1897) Tableau des especes palearctiques du genre Coprophilus Latr. (Zonoptilus Mots. - Homalotrichus Sol.). Revue d'Entomologie, 16, 226 - 229." type="journal article" year="1897">Fauvel, 1897</bibRefCitation>
</taxonomicName>
| det. M. Gildenkov, 2014” “ZOOLOGICAL INSTITUTE RAS ST. PETERSBURG” (ZMAS).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="DF6A664CCA1FBA6D1BECCEFB0C7EF879" httpUri="https://zenodo.org/record/843169/files/figure.png" pageId="4" pageNumber="134" targetBox="[164,1422,949,1948]" targetPageId="4">
<paragraph id="8BAA36C4CA1FBA6D1BECCEFB0C7EF879" blockId="4.[151,1436,1982,2036]" pageId="4" pageNumber="134">
<emphasis id="B961EAD6CA1FBA6D1BECCEFB0970F85E" bold="true" box="[151,316,1982,2005]" pageId="4" pageNumber="134">FIGURES 67.</emphasis>
<taxonomicName id="4C154D47CA1FBA6D1A38CEFA098CF85F" authorityName="Latreille" authorityYear="1829" box="[323,448,1983,2005]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAD6CA1FBA6D1A38CEFA098CF85F" box="[323,448,1983,2005]" italics="true" pageId="4" pageNumber="134">Coprophilus</emphasis>
</taxonomicName>
spp.: 6—
<taxonomicName id="4C154D47CA1FBA6D195CCE850BCBF85F" authority="Kashcheev, 1999" authorityName="Kashcheev" authorityYear="1999" box="[551,903,1983,2005]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="134" phylum="Arthropoda" rank="species" species="rufitarsis" subGenus="Zonyptilus">
<emphasis id="B961EAD6CA1FBA6D195CCE850A74F85F" box="[551,568,1984,2005]" italics="true" pageId="4" pageNumber="134">C</emphasis>
. (
<emphasis id="B961EAD6CA1FBA6D1934CE850A11F85F" box="[591,605,1984,2005]" italics="true" pageId="4" pageNumber="134">Z</emphasis>
.)
<emphasis id="B961EAD6CA1FBA6D1908CEFA0A9CF85F" box="[627,720,1983,2005]" italics="true" pageId="4" pageNumber="134">rufitarsis</emphasis>
Kashcheev, 1999
</taxonomicName>
(holotype?, female), body, dorsal view; 7—
<taxonomicName id="4C154D47CA1FBA6D1E2CCE8509C1F879" authority="Fauvel, 1897" authorityName="Fauvel" authorityYear="1897" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="134" phylum="Arthropoda" rank="species" species="pentatoma" subGenus="Zonyptilus">
<emphasis id="B961EAD6CA1FBA6D1E2CCE850D24F85F" box="[1367,1384,1984,2005]" italics="true" pageId="4" pageNumber="134">C</emphasis>
. (
<emphasis id="B961EAD6CA1FBA6D1E04CE850DC1F85F" box="[1407,1421,1984,2005]" italics="true" pageId="4" pageNumber="134">Z</emphasis>
.)
<emphasis id="B961EAD6CA1FBA6D1BECCE9A094EF87E" box="[151,258,2015,2036]" italics="true" pageId="4" pageNumber="134">pentatoma</emphasis>
Fauvel, 1897
</taxonomicName>
(“Kebin”, female), body, dorsal view. Scale bar: figs. 6, 7—1 mm.
</paragraph>
</caption>
<caption id="DF6A664CCA1EBA6C1BECCD8E0DDFFA95" httpUri="https://zenodo.org/record/843171/files/figure.png" pageId="5" pageNumber="135" targetBox="[170,1410,206,1188]" targetPageId="5">
<paragraph id="8BAA36C4CA1EBA6C1BECCD8E0DDFFA95" blockId="5.[151,1427,1227,1311]" pageId="5" pageNumber="135">
<emphasis id="B961EAD6CA1EBA6C1BECCD8E0976FB6B" bold="true" box="[151,314,1227,1250]" pageId="5" pageNumber="135">FIGURES 89.</emphasis>
<taxonomicName id="4C154D47CA1EBA6C1A3BCD8909F1FB68" authorityName="Latreille" authorityYear="1829" box="[320,445,1228,1250]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAD6CA1EBA6C1A3BCD8909F1FB68" box="[320,445,1228,1250]" italics="true" pageId="5" pageNumber="135">Coprophilus</emphasis>
</taxonomicName>
spp., the labels: 8—
<taxonomicName id="4C154D47CA1EBA6C19F5CD890B5AFB68" authorityName="Fauvel" authorityYear="1897" box="[654,790,1228,1250]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="135" phylum="Arthropoda" rank="species" species="pentatoma">
<emphasis id="B961EAD6CA1EBA6C19F5CD890B5AFB68" box="[654,790,1228,1250]" italics="true" pageId="5" pageNumber="135">C. pentatoma</emphasis>
</taxonomicName>
(lectotype, male); 9—
<taxonomicName id="4C154D47CA1EBA6C1887CD890CC8FB68" authorityName="Fauvel" authorityYear="1897" box="[1020,1156,1228,1250]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="135" phylum="Arthropoda" rank="species" species="pentatoma">
<emphasis id="B961EAD6CA1EBA6C1887CD890CC8FB68" box="[1020,1156,1228,1250]" italics="true" pageId="5" pageNumber="135">C. pentatoma</emphasis>
</taxonomicName>
(paralectotype, female).
<emphasis id="B961EAD6CA1EBA6C1BECCDAF091EFB75" bold="true" box="[151,338,1258,1281]" pageId="5" pageNumber="135">FIGURES 1011.</emphasis>
<taxonomicName id="4C154D47CA1EBA6C1A23CDAF099AFA8A" authorityName="Latreille" authorityYear="1829" box="[344,470,1258,1280]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAD6CA1EBA6C1A23CDAF099AFA8A" box="[344,470,1258,1280]" italics="true" pageId="5" pageNumber="135">Coprophilus</emphasis>
</taxonomicName>
spp., the labels: 10—
<taxonomicName id="4C154D47CA1EBA6C19C8CDAE0B77FA8A" authorityName="Fauvel" authorityYear="1897" box="[691,827,1259,1280]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="135" phylum="Arthropoda" rank="species" species="pentatoma">
<emphasis id="B961EAD6CA1EBA6C19C8CDAE0B77FA8A" box="[691,827,1259,1280]" italics="true" pageId="5" pageNumber="135">C. pentatoma</emphasis>
</taxonomicName>
(“Kebin”, female); 11—
<taxonomicName id="4C154D47CA1EBA6C1F4CCDAE0C84FA8A" authorityName="Bernhauer" authorityYear="1903" box="[1079,1224,1258,1280]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="135" phylum="Arthropoda" rank="species" species="longicornis">
<emphasis id="B961EAD6CA1EBA6C1F4CCDAE0C84FA8A" box="[1079,1224,1258,1280]" italics="true" pageId="5" pageNumber="135">C. longicornis</emphasis>
</taxonomicName>
(holotype, male).
<emphasis id="B961EAD6CA1EBA6C1BECCC4D091FFA94" bold="true" box="[151,339,1288,1311]" pageId="5" pageNumber="135">FIGURES 1213.</emphasis>
<taxonomicName id="4C154D47CA1EBA6C1A21CC4C099BFA95" authorityName="Latreille" authorityYear="1829" box="[346,471,1289,1311]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAD6CA1EBA6C1A21CC4C099BFA95" box="[346,471,1289,1311]" italics="true" pageId="5" pageNumber="135">Coprophilus</emphasis>
</taxonomicName>
spp., the labels: 12—
<taxonomicName id="4C154D47CA1EBA6C19CFCC4F0B02FA95" authorityName="Luze" authorityYear="1904" box="[692,846,1289,1311]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="135" phylum="Arthropoda" rank="species" species="bimaculatus">
<emphasis id="B961EAD6CA1EBA6C19CFCC4F0B02FA95" box="[692,846,1289,1311]" italics="true" pageId="5" pageNumber="135">C. bimaculatus</emphasis>
</taxonomicName>
(lectotype, male); 13—
<taxonomicName id="4C154D47CA1EBA6C1F44CC4F0CF5FA95" authorityName="Kashcheev" authorityYear="1999" box="[1087,1209,1289,1311]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="135" phylum="Arthropoda" rank="species" species="rufitarsis">
<emphasis id="B961EAD6CA1EBA6C1F44CC4F0CF5FA95" box="[1087,1209,1289,1311]" italics="true" pageId="5" pageNumber="135">C. rufitarsis</emphasis>
</taxonomicName>
(holotype?, female).
</paragraph>
</caption>
<subSubSection id="C30F654FCA1EBA6F1BBCCF2E0958FEB5" lastPageId="6" lastPageNumber="136" pageId="5" pageNumber="135" type="discussion">
<paragraph id="8BAA36C4CA1EBA6C1BBCCF2E0BD1F946" blockId="5.[151,1437,1355,2029]" pageId="5" pageNumber="135">
<emphasis id="B961EAD6CA1EBA6C1BBCCF2E0909F90E" bold="true" box="[199,325,1643,1668]" pageId="5" pageNumber="135">Discussion</emphasis>
. The body structure and coloration of the
<typeStatus id="54AE8866CA1EBA6C1865CF290B02F90E" box="[798,846,1644,1668]" pageId="5" pageNumber="135">type</typeStatus>
specimens of
<taxonomicName id="4C154D47CA1EBA6C1894CF290C3BF90E" authorityName="Kashcheev" authorityYear="1999" box="[1007,1143,1644,1668]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="135" phylum="Arthropoda" rank="species" species="rufitarsis">
<emphasis id="B961EAD6CA1EBA6C1894CF290C3BF90E" box="[1007,1143,1644,1668]" italics="true" pageId="5" pageNumber="135">C. rufitarsis</emphasis>
</taxonomicName>
are similar to those of the
<typeStatus id="54AE8866CA1EBA6C1BECCFCA08B2F922" box="[151,254,1679,1704]" pageId="5" pageNumber="135" type="lectotype">lectotype</typeStatus>
and
<typeStatus id="54AE8866CA1EBA6C1A3ACFCA0994F922" box="[321,472,1679,1704]" pageId="5" pageNumber="135" type="paralectotype">paralectotype</typeStatus>
(female) of
<taxonomicName id="4C154D47CA1EBA6C190CCFD50B58F922" authorityName="Fauvel" authorityYear="1897" box="[631,788,1680,1704]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="135" phylum="Arthropoda" rank="species" species="pentatoma">
<emphasis id="B961EAD6CA1EBA6C190CCFD50B58F922" box="[631,788,1680,1704]" italics="true" pageId="5" pageNumber="135">C. pentatoma</emphasis>
</taxonomicName>
. Thus, and the new synonymy is established:
<taxonomicName id="4C154D47CA1EBA6C1E30CFD509EAF946" authority="Fauvel, 1897" authorityName="Fauvel" authorityYear="1897" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="135" phylum="Arthropoda" rank="species" species="pentatoma" subGenus="Zonyptilus">
<emphasis id="B961EAD6CA1EBA6C1E30CFD50D12F92D" box="[1355,1374,1680,1703]" italics="true" pageId="5" pageNumber="135">C</emphasis>
. (
<emphasis id="B961EAD6CA1EBA6C1E07CFD40DC7F922" box="[1404,1419,1681,1704]" italics="true" pageId="5" pageNumber="135">Z</emphasis>
.)
<emphasis id="B961EAD6CA1EBA6C1BECCFF00941F946" box="[151,269,1717,1740]" italics="true" pageId="5" pageNumber="135">pentatoma</emphasis>
<bibRefCitation id="EF844B35CA1EBA6C1A6FCFF609EAF946" author="Fauvel" box="[276,422,1715,1740]" pageId="5" pageNumber="135" refString="Fauvel, A. (1897) Tableau des especes palearctiques du genre Coprophilus Latr. (Zonoptilus Mots. - Homalotrichus Sol.). Revue d'Entomologie, 16, 226 - 229." type="journal article" year="1897">Fauvel, 1897</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="4C154D47CA1EBA6C1ABFCFF10B09F946" authority="Kashcheev, 1999" authorityName="Kashcheev" authorityYear="1999" box="[452,837,1715,1740]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="135" phylum="Arthropoda" rank="species" species="rufitarsis" subGenus="Zonyptilus">
<emphasis id="B961EAD6CA1EBA6C1ABFCFF1099BF941" box="[452,471,1716,1739]" italics="true" pageId="5" pageNumber="135">C</emphasis>
. (
<emphasis id="B961EAD6CA1EBA6C1A95CFF009B1F946" box="[494,509,1717,1740]" italics="true" pageId="5" pageNumber="135">Z</emphasis>
.)
<emphasis id="B961EAD6CA1EBA6C196ECFF10A30F946" box="[533,636,1716,1740]" italics="true" pageId="5" pageNumber="135">rufitarsis</emphasis>
<bibRefCitation id="EF844B35CA1EBA6C19F8CFF60B09F946" author="Kashcheev" box="[643,837,1715,1740]" pageId="5" pageNumber="135" refString="Kashcheev, V. A. (1999) Novye Oxytelinae (Coleoptera, Staphylinidae) Srednei Azii i Kazakhstana. Tethys Entomological Research, 1, 141 - 156. [in Russian]" type="journal article" year="1999">Kashcheev, 1999</bibRefCitation>
</taxonomicName>
,
<taxonomicNameLabel id="A25257ADCA1EBA6C1834CFF60BD1F946" box="[847,925,1715,1740]" pageId="5" pageNumber="135">
<emphasis id="B961EAD6CA1EBA6C1834CFF60BD1F946" bold="true" box="[847,925,1715,1740]" pageId="5" pageNumber="135">syn. n.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="8BAA36C4CA1EBA6F1BBCCF920958FEB5" blockId="5.[151,1437,1355,2029]" lastBlockId="6.[151,1436,151,320]" lastPageId="6" lastPageNumber="136" pageId="5" pageNumber="135">
<emphasis id="B961EAD6CA1EBA6C1BBCCF920978F97A" bold="true" box="[199,308,1751,1776]" pageId="5" pageNumber="135">Remarks</emphasis>
. In the original description (
<bibRefCitation id="EF844B35CA1EBA6C190CCF920B7BF97A" author="Kashcheev" box="[631,823,1751,1776]" pageId="5" pageNumber="135" refString="Kashcheev, V. A. (1999) Novye Oxytelinae (Coleoptera, Staphylinidae) Srednei Azii i Kazakhstana. Tethys Entomological Research, 1, 141 - 156. [in Russian]" type="journal article" year="1999">Kashcheev 1999</bibRefCitation>
), the holotype was stated to be a male (10 males were also designated as paratypes). The examination of the material from ZMAS revealed seven type specimens of
<taxonomicName id="4C154D47CA1EBA6C1EF9CFB908B2F8B2" authorityName="Kashcheev" authorityYear="1999" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="135" phylum="Arthropoda" rank="species" species="rufitarsis">
<emphasis id="B961EAD6CA1EBA6C1EF9CFB908B2F8B2" italics="true" pageId="5" pageNumber="135">C. rufitarsis</emphasis>
</taxonomicName>
, including one specimen labeled as “Holotypus
<taxonomicName id="4C154D47CA1EBA6C185CCE5A0CE9F8B2" authority="Kastcheev" authorityName="Kastcheev" box="[807,1189,1823,1848]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="135" phylum="Arthropoda" rank="species" species="rufitarsis">
<emphasis id="B961EAD6CA1EBA6C185CCE5A0C6EF8B2" box="[807,1058,1823,1848]" italics="true" pageId="5" pageNumber="135">Coprophilus rufitarsis</emphasis>
Kastcheev
</taxonomicName>
”. All type specimens, however, were found to be females. The location of males indicated in the original description (
<bibRefCitation id="EF844B35CA1EBA6C1FC5CE060D37F8D6" author="Kashcheev" box="[1214,1403,1859,1884]" pageId="5" pageNumber="135" refString="Kashcheev, V. A. (1999) Novye Oxytelinae (Coleoptera, Staphylinidae) Srednei Azii i Kazakhstana. Tethys Entomological Research, 1, 141 - 156. [in Russian]" type="journal article" year="1999">Kashcheev 1999</bibRefCitation>
) is unknown. The labels of the type specimens studied were written by hand of the author and their content completely matches with the data from the original description (
<bibRefCitation id="EF844B35CA1EBA6C1990CEC90BE0F82E" author="Kashcheev" box="[747,940,1932,1957]" pageId="5" pageNumber="135" refString="Kashcheev, V. A. (1999) Novye Oxytelinae (Coleoptera, Staphylinidae) Srednei Azii i Kazakhstana. Tethys Entomological Research, 1, 141 - 156. [in Russian]" type="journal article" year="1999">Kashcheev 1999</bibRefCitation>
). The morphological structure of these type specimens leaves no doubt of their synonymy. This conclusion is also indirectly supported by my study of numerous specimens of
<taxonomicName id="4C154D47CA1EBA6C1AC8CE900A03F866" authorityName="Fauvel" authorityYear="1897" box="[435,591,2005,2028]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="135" phylum="Arthropoda" rank="species" species="pentatoma">
<emphasis id="B961EAD6CA1EBA6C1AC8CE900A03F866" box="[435,591,2005,2028]" italics="true" pageId="5" pageNumber="135">C. pentatoma</emphasis>
</taxonomicName>
from Kazakhstan, from the collection of Vitaliy Kashcheev, which was recently donated to ZMAS. The figures of the aedeagus of
<taxonomicName id="4C154D47CA1DBA6F184BC9DD0D69FF3A" authority="in Kashcheev (1999, fig. 1: 12)" authorityName="in Kashcheev" authorityYear="1999" box="[816,1317,151,176]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="136" phylum="Arthropoda" rank="species" species="pentatoma">
<emphasis id="B961EAD6CA1DBA6F184BC9DD0B8AFF3A" box="[816,966,152,176]" italics="true" pageId="6" pageNumber="136">C. pentatoma</emphasis>
in
<bibRefCitation id="EF844B35CA1DBA6F1890C9D20D69FF3A" author="Kashcheev" box="[1003,1317,151,176]" pageId="6" pageNumber="136" refString="Kashcheev, V. A. (1999) Novye Oxytelinae (Coleoptera, Staphylinidae) Srednei Azii i Kazakhstana. Tethys Entomological Research, 1, 141 - 156. [in Russian]" type="journal article" year="1999">Kashcheev (1999, fig.1: 12)</bibRefCitation>
</taxonomicName>
contradict the structure of the aedeagus of the lectotype of
<taxonomicName id="4C154D47CA1DBA6F19C3C9F80B1CFF5E" authorityName="in Kashcheev" authorityYear="1999" box="[696,848,189,212]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="136" phylum="Arthropoda" rank="species" species="pentatoma">
<emphasis id="B961EAD6CA1DBA6F19C3C9F80B1CFF5E" box="[696,848,189,212]" italics="true" pageId="6" pageNumber="136">C. pentatoma</emphasis>
</taxonomicName>
(figs. 23). I believe that the figure of the aedeagus of
<taxonomicName id="4C154D47CA1DBA6F1BC2C9A50908FF72" box="[185,324,224,248]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="136" phylum="Arthropoda" rank="species" species="rufitarsis">
<emphasis id="B961EAD6CA1DBA6F1BC2C9A50908FF72" box="[185,324,224,248]" italics="true" pageId="6" pageNumber="136">C. rufitarsis</emphasis>
</taxonomicName>
that differs (
<bibRefCitation id="EF844B35CA1DBA6F1AA6C99A0B5DFF72" author="Kashcheev" box="[477,785,223,248]" pageId="6" pageNumber="136" refString="Kashcheev, V. A. (1999) Novye Oxytelinae (Coleoptera, Staphylinidae) Srednei Azii i Kazakhstana. Tethys Entomological Research, 1, 141 - 156. [in Russian]" type="journal article" year="1999">Kashcheev 1999, fig.1: 13</bibRefCitation>
) from the illustrations of the aedeagus of
<taxonomicName id="4C154D47CA1DBA6F1E79C9A50DD0FF72" authorityName="in Kashcheev" authorityYear="1999" box="[1282,1436,224,248]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="136" phylum="Arthropoda" rank="species" species="pentatoma">
<emphasis id="B961EAD6CA1DBA6F1E79C9A50DD0FF72" box="[1282,1436,224,248]" italics="true" pageId="6" pageNumber="136">C. pentatoma</emphasis>
</taxonomicName>
should unfortunately be regarded as erroneous. It is clear that Kashcheev had an incorrect understanding of
<taxonomicName id="4C154D47CA1DBA6F1EF9C8400941FECA" authorityName="in Kashcheev" authorityYear="1999" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="136" phylum="Arthropoda" rank="species" species="pentatoma">
<emphasis id="B961EAD6CA1DBA6F1EF9C8400941FECA" italics="true" pageId="6" pageNumber="136">C. pentatoma</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>