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<document id="1219449F9804F3B5D77A4363D392CBDC" ID-DOI="10.11646/zootaxa.4991.3.8" ID-ISSN="1175-5326" ID-Zenodo-Dep="5042656" ID-ZooBank="F0FAE07E-4A90-4602-BCC5-6D80E158BB6C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624989159860" checkinUser="plazi" docAuthor="Suksai, Benjamart, Gustafson, Grey T., Sites, Robert W. &amp; Sangpradub, Narumon" docDate="2021" docId="03BC87C1FFFF225C0CC07E38FE79501B" docLanguage="en" docName="zootaxa.4991.3.8.pdf" docOrigin="Zootaxa 4991 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Patrus salvazai" docType="treatment" docVersion="3" lastPageNumber="584" masterDocId="FF85FFB9FFEA224B0C577A23FFDF537D" masterDocTitle="Review of the Patrus landaisi species group from Thailand with description of three new species (Coleoptera, Gyrinidae)" masterLastPageNumber="591" masterPageNumber="561" pageNumber="582" updateTime="1698978743576" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="539BD2FEA1A5DE1663A381C58BE55576">Review of the Patrus landaisi species group from Thailand with description of three new species (Coleoptera, Gyrinidae)</mods:title>
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<mods:namePart id="9AFFB592D0F1FDC48F84222DFCBDF5FB">Suksai, Benjamart</mods:namePart>
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<mods:namePart id="91D81D82C7B079AA84FCAA36AF2200F4">Gustafson, Grey T.</mods:namePart>
<mods:affiliation id="A3523D9703B8EDE6C678242185CE795E">Department of Ecology and Evolutionary Biology, University of Kansas, Lawrence, Kansas 66045, USA. &amp; Department of Biological Sciences, Northern Arizona University, Flagstaff, Arizona, 86011, USA.</mods:affiliation>
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<mods:namePart id="B009157B05BC786217B114D693C2A3A6">Sites, Robert W.</mods:namePart>
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<mods:affiliation id="380C97388AADB7AB583AB8604838D08B">Enns Entomology Museum, Division of Plant Science, University of Missouri-Columbia, Missouri 65211, USA. SitesR @ missouri. edu; https: // orcid. org / 0000 - 0002 - 3895 - 813 X</mods:affiliation>
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<mods:namePart id="79D17A6FCEBEEA6DD42D496263C08E7C">Sangpradub, Narumon</mods:namePart>
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<mods:affiliation id="4AA37929705F3F96FA26B4E532EEA004">Applied Taxonomic Research Center, Faculty of Science, Khon Kaen University, Khon Kaen 40002, Thailand &amp; Corresponding Author: narumon @ kku. ac. th; https: // orcid. org / 0000 - 0003 - 0228 - 8696</mods:affiliation>
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<mods:title id="CA8084EC97C1740366308D6CAFE683FB">Zootaxa</mods:title>
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<mods:date id="9D277F4325929FCEBC82D7096071CDFC">2021</mods:date>
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<heading id="D0E281BBFFFF225E0CC07E38FE2D5749" bold="true" box="[151,498,1050,1077]" fontSize="11" level="1" pageId="21" pageNumber="582" reason="1">
<taxonomicName id="4C154D54FFFF225E0CC07E38FE2D5749" authority="(Ochs, 1940)" baseAuthorityName="Ochs" baseAuthorityYear="1940" box="[151,498,1050,1077]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="582" phylum="Arthropoda" rank="species" species="salvazai">
<emphasis id="B961EAC5FFFF225E0CC07E38FE2D5749" bold="true" box="[151,498,1050,1077]" pageId="21" pageNumber="582">
<emphasis id="B961EAC5FFFF225E0CC07E38FE915748" bold="true" box="[151,334,1050,1077]" italics="true" pageId="21" pageNumber="582">Patrus salvazai</emphasis>
(
<bibRefCitation id="EF844B26FFFF225E0D097E39FE355749" author="Ochs" box="[350,490,1050,1076]" pageId="21" pageNumber="582" pagination="1 - 22" refId="ref22563" refString="Ochs, G. (1940) Entomological results from the Swedish Expedition 1934 to Burma and British India. Coleoptera: Gyrinidae, gesammelt von Rene Malaise. Arkiv for Zoologi, 32 A, 1 - 22." type="journal article" year="1940">Ochs, 1940</bibRefCitation>
)
</emphasis>
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<paragraph id="8BAA36D7FFFF225E0CC07E1CFD635724" blockId="21.[151,700,1050,1113]" box="[151,700,1087,1113]" pageId="21" pageNumber="582">
(
<figureCitation id="132E2A52FFFF225E0CC87E1CFEDC5724" box="[159,259,1087,1113]" captionStart="FIGURE 2" captionStartId="6.[151,260,1861,1887]" captionTargetBox="[151,1434,179,1836]" captionTargetId="figure-25@6.[151,1435,179,1836]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042660" httpUri="https://zenodo.org/record/5042660/files/figure.png" pageId="21" pageNumber="582">Figs. 2G</figureCitation>
,
<figureCitation id="132E2A52FFFF225E0D587E1CFEEC5724" box="[271,307,1087,1113]" captionStart="FIGURE 3" captionStartId="7.[151,260,1865,1891]" captionTargetBox="[151,1434,182,1841]" captionTargetId="figure-18@7.[151,1435,179,1841]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal habitus of females of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042662" httpUri="https://zenodo.org/record/5042662/files/figure.png" pageId="21" pageNumber="582">3G</figureCitation>
,
<figureCitation id="132E2A52FFFF225E0D687E1CFEBB5724" box="[319,356,1087,1113]" captionStart="FIGURE 4" captionStartId="8.[151,260,707,733]" captionTargetBox="[158,1432,185,677]" captionTargetId="figure-25@8.[152,1436,179,683]" captionTargetPageId="8" captionText="FIGURE 4. Left protibia and protarsus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5042666" httpUri="https://zenodo.org/record/5042666/files/figure.png" pageId="21" pageNumber="582">4G</figureCitation>
,
<figureCitation id="132E2A52FFFF225E0D387E1CFE535724" box="[367,396,1087,1113]" captionStart="FIGURE 11" captionStartId="22.[151,260,1770,1796]" captionTargetBox="[159,1430,816,1737]" captionTargetId="figure-281@22.[151,1436,810,1745]" captionTargetPageId="22" captionText="FIGURE 11. Patrus salvazai: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042686" httpUri="https://zenodo.org/record/5042686/files/figure.png" pageId="21" pageNumber="582">11</figureCitation>
,
<figureCitation id="132E2A52FFFF225E0DCF7E1CFDA75724" box="[408,632,1087,1113]" captionStart="FIGURE 13" captionStartId="25.[151,260,1683,1709]" captionTargetBox="[169,1418,181,1656]" captionTargetId="figure-18@25.[169,1418,179,1659]" captionTargetPageId="25" captionText="FIGURE 13. Habitats of P. landaisi species group: (A) stream at Siriphum Waterfall, Doi Inthanon National Park, Chiang Mai Province, habitat of P. garuda sp. nov. and P. subapicalis; (B) Benjamart Suksai in habitat of P. apicalis and P. salvazai at Nam Kuen stream, Chiang Rai Province; (C) shallow stream close to Ban Thai Samakkhi, Chiang Rai Province, habitat of P. landaisi and P. salvazai; (D) Tham Yai Waterfall Phu Kradueng National Park, Loei Province, habitat of P. phetchabunensis sp. nov.; (E) puddle in dry stream bed close to Ban Phithak Thai, Chiang Rai Province, habitat of P. landaisi and P. salvazai; (F) P. landaisi and P. salvazai from puddle in dry stream habitat close to Ban Thai Samakkhi, Chiang Rai Province." figureDoi="http://doi.org/10.5281/zenodo.5042690" httpUri="https://zenodo.org/record/5042690/files/figure.png" pageId="21" pageNumber="582">13B, 13C, 13E, 13F</figureCitation>
,
<figureCitation id="132E2A52FFFF225E0ED37E1CFD6B5724" box="[644,692,1087,1113]" captionStart="FIGURE 14" captionStartId="26.[151,260,743,769]" captionTargetBox="[152,1436,179,719]" captionTargetId="figure-25@26.[151,1436,179,719]" captionTargetPageId="26" captionText="FIGURE 14. (AB) Distribution records of the P. landaisi species group in Thailand." figureDoi="http://doi.org/10.5281/zenodo.5042692" httpUri="https://zenodo.org/record/5042692/files/figure.png" pageId="21" pageNumber="582">14B</figureCitation>
)
</paragraph>
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<paragraph id="8BAA36D7FFFF225E0CC07EA0FC4357E6" blockId="21.[151,924,1155,1241]" box="[151,924,1155,1179]" pageId="21" pageNumber="582">
<treatmentCitationGroup id="AB0511F9FFFF225E0CC07EA0FC4357E6" box="[151,924,1155,1179]" pageId="21" pageNumber="582">
<taxonomicName id="4C154D54FFFF225E0CC07EA0FDC357E6" authority="Regimbart" authorityName="Regimbart" authorityYear="1888" box="[151,540,1155,1179]" class="Insecta" family="Gyrinidae" genus="Orectochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="582" phylum="Arthropoda" rank="species" species="cardiophorus">
<emphasis id="B961EAC5FFFF225E0CC07EA0FE7857E6" box="[151,423,1155,1179]" italics="true" pageId="21" pageNumber="582">Orectochilus cardiophorus</emphasis>
Régimbart
</taxonomicName>
:
<treatmentCitation id="0AB410C6FFFF225E0E717EA0FD0657E6" author="Peschet, R." box="[550,729,1155,1179]" page="125" pageId="21" pageNumber="582" year="1923">
<bibRefCitation id="EF844B26FFFF225E0E717EA0FD0657E6" author="Peschet, R." box="[550,729,1155,1179]" pageId="21" pageNumber="582" pagination="121 - 136" refId="ref22628" refString="Peschet, R. (1923) Fauna entomologique de l'Indochine Gyrinidae. Opuscules de l'Institut Scientifique de l'Indochine Saigon, 6, 121 - 136." type="journal article" year="1923">Peschet 1923:125</bibRefCitation>
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(misidentification)
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</paragraph>
<paragraph id="8BAA36D7FFFF225E0CC07E81FCD257C7" blockId="21.[151,924,1155,1241]" box="[151,781,1186,1210]" pageId="21" pageNumber="582">
<treatmentCitationGroup id="AB0511F9FFFF225E0CC07E81FCD257C7" box="[151,781,1186,1210]" pageId="21" pageNumber="582">
<taxonomicName id="4C154D54FFFF225E0CC07E81FE1557C7" authority="Peschet" authorityName="Peschet" box="[151,458,1186,1210]" class="Insecta" family="Gyrinidae" genus="Orectochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="582" phylum="Arthropoda" rank="species" species="salvazai">
<emphasis id="B961EAC5FFFF225E0CC07E81FEAD57C7" box="[151,370,1186,1210]" italics="true" pageId="21" pageNumber="582">Orectochilus salvazai</emphasis>
Peschet
</taxonomicName>
:
<treatmentCitation id="0AB410C6FFFF225E0D837E81FDB857C7" author="Ochs" box="[468,615,1186,1210]" page="33" pageId="21" pageNumber="582" year="1930">
<bibRefCitation id="EF844B26FFFF225E0D837E81FDB857C7" author="Ochs" box="[468,615,1186,1210]" pageId="21" pageNumber="582" pagination="1 - 39" refId="ref22521" refString="Ochs, G. (1930) Gyrinoidea. Catalogue of Indian Insects, 19, 1 - 39." type="journal article" year="1930">Ochs 1930: 33</bibRefCitation>
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(
<emphasis id="B961EAC5FFFF225E0E217E80FCDA57C7" box="[630,773,1186,1210]" italics="true" pageId="21" pageNumber="582">nomen nudum</emphasis>
)
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<paragraph id="8BAA36D7FFFF225E0CC07EE3FCF657A5" blockId="21.[151,924,1155,1241]" box="[151,809,1216,1241]" pageId="21" pageNumber="582">
<treatmentCitationGroup id="AB0511F9FFFF225E0CC07EE3FCF657A5" box="[151,809,1216,1241]" pageId="21" pageNumber="582">
<treatmentCitation id="0AB410C6FFFF225E0CC07EE3FDCC57A4" author="Ochs" box="[151,531,1216,1241]" page="19" pageId="21" pageNumber="582" year="1940">
<taxonomicName id="4C154D54FFFF225E0CC07EE3FDCC57A4" authority="Ochs, 1940: 19" authorityName="Ochs" authorityPageNumber="19" authorityYear="1940" box="[151,531,1216,1241]" class="Insecta" family="Gyrinidae" genus="Orectochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="582" phylum="Arthropoda" rank="species" species="salvazai">
<emphasis id="B961EAC5FFFF225E0CC07EE3FEAD57A5" box="[151,370,1216,1240]" italics="true" pageId="21" pageNumber="582">Orectochilus salvazai</emphasis>
<bibRefCitation id="EF844B26FFFF225E0D2E7EE3FDCC57A4" author="Ochs" box="[377,531,1216,1241]" pageId="21" pageNumber="582" pagination="1 - 22" refId="ref22563" refString="Ochs, G. (1940) Entomological results from the Swedish Expedition 1934 to Burma and British India. Coleoptera: Gyrinidae, gesammelt von Rene Malaise. Arkiv for Zoologi, 32 A, 1 - 22." type="journal article" year="1940">Ochs, 1940: 19</bibRefCitation>
</taxonomicName>
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(valid original description)
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</paragraph>
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<materialsCitation id="3B7D3C8AFFFF225E0CC07F22FD315661" box="[151,750,1281,1308]" country="Laos" location="Luang Prabang" pageId="21" pageNumber="582" specimenCount="1" stateProvince="Louangphabang" typeStatus="holotype">
<emphasis id="B961EAC5FFFF225E0CC07F22FEE95666" bold="true" box="[151,310,1281,1307]" pageId="21" pageNumber="582">
<typeStatus id="54AE8875FFFF225E0CC07F22FF0D5666" box="[151,210,1281,1307]" pageId="21" pageNumber="582">Type</typeStatus>
locality.
</emphasis>
Sala-Nam-Pik,
<collectingRegion id="49D1F835FFFF225E0DAB7F21FD765666" box="[508,681,1281,1308]" country="Laos" name="Louangphabang" pageId="21" pageNumber="582">Luang Prabang</collectingRegion>
,
<collectingCountry id="F3027647FFFF225E0EE37F21FD315661" box="[692,750,1282,1308]" name="Laos" pageId="21" pageNumber="582">Laos</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8BAA36D7FFFF225D0C907F06FAB55384" blockId="21.[151,1437,1281,2028]" lastBlockId="22.[151,1437,150,753]" lastPageId="22" lastPageNumber="583" pageId="21" pageNumber="582">
<emphasis id="B961EAC5FFFF225E0C907F06FE725642" bold="true" box="[199,429,1317,1343]" pageId="21" pageNumber="582">Material examined.</emphasis>
<materialsCitation id="3B7D3C8AFFFF225E0DE77F06FAF7561E" collectingDate="2002-05-08" collectionCode="THNHM" country="Thailand" elevation="1060" latitude="18.516666" location="Huai Sai Luang Waterfall" longLatPrecision="1275" longitude="98.45" municipality="Doi Inthanon Natl. Park." pageId="21" pageNumber="582" specimenCount="1" specimenCount-male="1" stateProvince="Chiang Mai">
<emphasis id="B961EAC5FFFF225E0DE77F06FC8A5642" bold="true" box="[432,853,1317,1343]" pageId="21" pageNumber="582">
<collectingCountry id="F3027647FFFF225E0DE77F06FD945642" box="[432,587,1317,1343]" name="Thailand" pageId="21" pageNumber="582">THAILAND</collectingCountry>
:
<collectingRegion id="49D1F835FFFF225E0E037F06FC8F5642" box="[596,848,1317,1343]" country="Thailand" name="Chiang Mai" pageId="21" pageNumber="582">Chiang Mai Province</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6BCEACADFFFF225E0F0F7F06FBBA5642" box="[856,1125,1317,1343]" pageId="21" pageNumber="582">Doi Inthanon Natl.Park.</collectingMunicipality>
, stream from
<location id="8ECA600CFFFF225E08A37F06FEDD561E" LSID="urn:lsid:plazi:treatment:03BC87C1FFFF225C0CC07E38FE79501B:8ECA600CFFFF225E08A37F06FEDD561E" country="Thailand" latitude="18.516666" longLatPrecision="1275" longitude="98.45" municipality="Doi Inthanon Natl. Park." name="Huai Sai Luang Waterfall" pageId="21" pageNumber="582" stateProvince="Chiang Mai">Huai Sai Luang Waterfall</location>
,
<geoCoordinate id="EE215010FFFF225E0D5C7F6AFEB3561E" box="[267,364,1353,1379]" degrees="18" direction="north" minutes="31" orientation="latitude" pageId="21" pageNumber="582" precision="925" value="18.516666">18°31N</geoCoordinate>
<geoCoordinate id="EE215010FFFF225E0D257F6AFE0D561E" box="[370,466,1353,1379]" degrees="98" direction="east" minutes="27" orientation="longitude" pageId="21" pageNumber="582" precision="925" value="98.45">98°27E</geoCoordinate>
,
<elevation id="0038D1E4FFFF225E0D8B7F6AFDB4561E" box="[476,619,1353,1380]" metricMagnitude="3" metricUnit="m" metricValue="1.06" pageId="21" pageNumber="582" unit="m" value="1060.0">
<quantity id="4CED9B32FFFF225E0D8B7F6AFDEF5619" box="[476,560,1353,1380]" metricMagnitude="3" metricUnit="m" metricValue="1.06" pageId="21" pageNumber="582" unit="m" value="1060.0">1060 m</quantity>
a.s.l.
</elevation>
,
<date id="FFAB1017FFFF225E0E217F6AFD02561E" box="[630,733,1353,1379]" pageId="21" pageNumber="582" value="2002-05-08">
<collectingDate id="EFEFE9FFFFFF225E0E217F6AFD02561E" box="[630,733,1353,1379]" pageId="21" pageNumber="582" value="2002-05-08">8.V.2002</collectingDate>
</date>
, leg. UMC &amp; CMU teams, L-396,
<specimenCount id="9D13FD5EFFFF225E080B7F6AFB7A561E" box="[1116,1189,1353,1379]" pageId="21" pageNumber="582" type="male">1 male</specimenCount>
(
<collectionCode id="ED04AE12FFFF225E08E47F6AFAFF561E" box="[1203,1312,1353,1379]" pageId="21" pageNumber="582">THNHM</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B7D3C8AFFFF225E09667F6AFEBC56FA" collectingDate="2002-05-08" collectionCode="THNHM, UMC" country="Thailand" elevation="1060" isTailEnumeration="true" latitude="18.516666" location="Huai Sai Luang Waterfall" longLatPrecision="1275" longitude="98.45" municipality="Doi Inthanon Natl. Park." pageId="21" pageNumber="582" specimenCount="2" specimenCount-female="2" stateProvince="Chiang Mai">
<specimenCount id="9D13FD5EFFFF225E09667F6AFA43561E" box="[1329,1436,1353,1379]" pageId="21" pageNumber="582" type="female">2 females</specimenCount>
(
<collectionCode id="ED04AE12FFFF225E0CC87F4EFED556FA" box="[159,266,1389,1415]" pageId="21" pageNumber="582">THNHM</collectionCode>
,
<collectionCode id="ED04AE12FFFF225E0D417F4EFE8456FA" box="[278,347,1389,1415]" pageId="21" pageNumber="582">UMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D3C8AFFFF225E0D387F4EFE0556D6" collectingDate="2003-02-14" collectionCode="UMC" country="Thailand" elevation="700" latitude="18.816668" location="Monthathan Waterfall" longLatPrecision="1274" longitude="98.916664" municipality="Doi Suthep Natl. Park." pageId="21" pageNumber="582" specimenCount="1" specimenCount-female="1" stateProvince="Chiang Mai">
<collectingMunicipality id="6BCEACADFFFF225E0D387F4EFDAB56FA" box="[367,628,1389,1415]" pageId="21" pageNumber="582">Doi Suthep Natl. Park.</collectingMunicipality>
,
<location id="8ECA600CFFFF225E0ED77F4EFCA556FA" LSID="urn:lsid:plazi:treatment:03BC87C1FFFF225C0CC07E38FE79501B:8ECA600CFFFF225E0ED77F4EFCA556FA" box="[640,890,1389,1415]" country="Thailand" latitude="18.816668" longLatPrecision="1274" longitude="98.916664" municipality="Doi Suthep Natl. Park." name="Monthathan Waterfall" pageId="21" pageNumber="582" stateProvince="Chiang Mai">Monthathan Waterfall</location>
,
<geoCoordinate id="EE215010FFFF225E0FD17F4EFC3856FA" box="[902,999,1389,1415]" degrees="18" direction="north" minutes="49" orientation="latitude" pageId="21" pageNumber="582" precision="925" value="18.816668">18°49N</geoCoordinate>
<geoCoordinate id="EE215010FFFF225E0FB87F4EFB9056F5" box="[1007,1103,1389,1416]" degrees="98" direction="east" minutes="55" orientation="longitude" pageId="21" pageNumber="582" precision="925" value="98.916664">98°55E</geoCoordinate>
,
<elevation id="0038D1E4FFFF225E080B7F4EFB3D56FA" box="[1116,1250,1389,1416]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="21" pageNumber="582" unit="m" value="700.0">
<quantity id="4CED9B32FFFF225E080B7F4EFB7B56F5" box="[1116,1188,1389,1416]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="21" pageNumber="582" unit="m" value="700.0">700 m</quantity>
a.s.l.
</elevation>
,
<date id="FFAB1017FFFF225E08B87F4EFAB856FA" box="[1263,1383,1389,1415]" pageId="21" pageNumber="582" value="2003-02-14">
<collectingDate id="EFEFE9FFFFFF225E08B87F4EFAB856FA" box="[1263,1383,1389,1415]" pageId="21" pageNumber="582" value="2003-02-14">14.II.2003</collectingDate>
</date>
, leg. CMU team,
<specimenCount id="9D13FD5EFFFF225E0D487FB2FEA156D6" box="[287,382,1425,1451]" pageId="21" pageNumber="582" type="female">1 female</specimenCount>
(
<collectionCode id="ED04AE12FFFF225E0DDB7FB2FE0D56D6" box="[396,466,1425,1451]" pageId="21" pageNumber="582">UMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D3C8AFFFF225E0DB47FB1FBD256B2" collectingDate="2018-01-08" collectionCode="KKU" collectorName="B. Suksai &amp; C. Yatfung &amp; S. Yoolong" country="Thailand" county="Haow River" elevation="505" latitude="19.122223" location="Mae Taeng District" longLatPrecision="1" longitude="98.77908" municipality="Sop Poeng Subdistrict" pageId="21" pageNumber="582" specimenCount="1" specimenCount-male="1" stateProvince="Chiang Mai">
<collectingCounty id="62CB4E5BFFFF225E0DB47FB1FDB456D6" box="[483,619,1425,1452]" pageId="21" pageNumber="582">Haow River</collectingCounty>
,
<collectingMunicipality id="6BCEACADFFFF225E0E227FB2FCB256D6" box="[629,877,1425,1452]" pageId="21" pageNumber="582">Sop Poeng Subdistrict</collectingMunicipality>
,
<location id="8ECA600CFFFF225E0F207FB1FB9256D6" LSID="urn:lsid:plazi:treatment:03BC87C1FFFF225C0CC07E38FE79501B:8ECA600CFFFF225E0F207FB1FB9256D6" box="[887,1101,1425,1452]" country="Thailand" county="Haow River" latitude="19.122223" longLatPrecision="1" longitude="98.77908" municipality="Sop Poeng Subdistrict" name="Mae Taeng District" pageId="21" pageNumber="582" stateProvince="Chiang Mai">Mae Taeng District</location>
,
<geoCoordinate id="EE215010FFFF225E08007FB2FB2B56D6" box="[1111,1268,1425,1451]" degrees="19" direction="north" minutes="07" orientation="latitude" pageId="21" pageNumber="582" precision="1" seconds="20.0" value="19.122223">19°0720.0”N</geoCoordinate>
<geoCoordinate id="EE215010FFFF225E08AD7FB2FA4756D6" box="[1274,1432,1425,1451]" degrees="98" direction="east" minutes="46" orientation="longitude" pageId="21" pageNumber="582" precision="1" seconds="44.7" value="98.77908">98°4644.7”E</geoCoordinate>
,
<elevation id="0038D1E4FFFF225E0CC07F96FEC456B2" box="[151,283,1461,1488]" metricMagnitude="2" metricUnit="m" metricValue="5.05" pageId="21" pageNumber="582" unit="m" value="505.0">
<quantity id="4CED9B32FFFF225E0CC07F96FF0056AD" box="[151,223,1461,1488]" metricMagnitude="2" metricUnit="m" metricValue="5.05" pageId="21" pageNumber="582" unit="m" value="505.0">505 m</quantity>
a.s.l.
</elevation>
,
<date id="FFAB1017FFFF225E0D7F7F96FE5756B2" box="[296,392,1461,1487]" pageId="21" pageNumber="582" value="2018-01-08">
<collectingDate id="EFEFE9FFFFFF225E0D7F7F96FE5756B2" box="[296,392,1461,1487]" pageId="21" pageNumber="582" value="2018-01-08">8.I.2018</collectingDate>
</date>
, leg.
<collectorName id="26E05301FFFF225E0D927F95FDEC56B2" box="[453,563,1461,1488]" pageId="21" pageNumber="582">B. Suksai</collectorName>
,
<collectorName id="26E05301FFFF225E0E687F96FD6756B2" box="[575,696,1461,1487]" pageId="21" pageNumber="582">C. Yatfung</collectorName>
&amp;
<collectorName id="26E05301FFFF225E0E8B7F96FC8456B2" box="[732,859,1461,1487]" pageId="21" pageNumber="582">S. Yoolong</collectorName>
,
<specimenCount id="9D13FD5EFFFF225E0F307F96FC6D56B2" box="[871,946,1461,1487]" pageId="21" pageNumber="582" type="male">1 male</specimenCount>
(
<collectionCode id="ED04AE12FFFF225E0F957F96FBDA56B2" box="[962,1029,1461,1487]" country="Thailand" httpUri="http://grbio.org/cool/pwjq-e3rs" name="Herbarium, Department of Biology, Khon Kaen University" pageId="21" pageNumber="582">KKU</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7D3C8AFFFF225E084F7F96FB02556A" collectingDate="2018-11-11" collectionCode="THNHM" collectorName="B. Suksai &amp; S. Kullasoot &amp; P. Sapewisut" country="Thailand" county="Nam Kuen Stream" elevation="1327" latitude="19.122862" location="Mae Chedi Subdistrict" longLatPrecision="1" longitude="99.3287" municipality="Ban Huai Nam Kuen" pageId="21" pageNumber="582" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Chiang Rai">
<emphasis id="B961EAC5FFFF225E084F7F96FAC556B2" bold="true" box="[1048,1306,1461,1487]" pageId="21" pageNumber="582">
<collectingRegion id="49D1F835FFFF225E084F7F96FAC956B2" box="[1048,1302,1461,1487]" country="Thailand" name="Chiang Rai" pageId="21" pageNumber="582">Chiang Rai Province</collectingRegion>
:
</emphasis>
<collectingCounty id="62CB4E5BFFFF225E09757F95FF36568E" pageId="21" pageNumber="582">Nam Kuen Stream</collectingCounty>
,
<collectingMunicipality id="6BCEACADFFFF225E0CA17FF9FE365689" box="[246,489,1497,1524]" pageId="21" pageNumber="582">Ban Huai Nam Kuen</collectingMunicipality>
,
<location id="8ECA600CFFFF225E0DA07FF9FD26568E" LSID="urn:lsid:plazi:treatment:03BC87C1FFFF225C0CC07E38FE79501B:8ECA600CFFFF225E0DA07FF9FD26568E" box="[503,761,1497,1524]" country="Thailand" county="Nam Kuen Stream" latitude="19.122862" longLatPrecision="1" longitude="99.3287" municipality="Ban Huai Nam Kuen" name="Mae Chedi Subdistrict" pageId="21" pageNumber="582" stateProvince="Chiang Rai">Mae Chedi Subdistrict</location>
,
<location id="8ECA600CFFFF225E0F527FFAFBD9568E" LSID="urn:lsid:plazi:treatment:03BC87C1FFFF225C0CC07E38FE79501B:8ECA600CFFFF225E0F527FFAFBD9568E" box="[773,1030,1497,1524]" country="Thailand" county="Nam Kuen Stream" latitude="19.122862" longLatPrecision="1" longitude="99.3287" municipality="Ban Huai Nam Kuen" name="Wiang Pa Pao District" pageId="21" pageNumber="582" stateProvince="Chiang Rai">Wiang Pa Pao District</location>
,
<geoCoordinate id="EE215010FFFF225E08447FFAFB6E568E" box="[1043,1201,1497,1523]" degrees="19" direction="north" minutes="07" orientation="latitude" pageId="21" pageNumber="582" precision="1" seconds="22.3" value="19.122862">19°0722.3”N</geoCoordinate>
<geoCoordinate id="EE215010FFFF225E08EE7FFAFA88568E" box="[1209,1367,1497,1523]" degrees="99" direction="east" minutes="19" orientation="longitude" pageId="21" pageNumber="582" precision="1" seconds="43.3" value="99.3287">99°1943.3”E</geoCoordinate>
,
<elevation id="0038D1E4FFFF225E09337FFAFF33556A" metricMagnitude="3" metricUnit="m" metricValue="1.327" pageId="21" pageNumber="582" unit="m" value="1327.0">
<quantity id="4CED9B32FFFF225E09337FFAFF725565" metricMagnitude="3" metricUnit="m" metricValue="1.327" pageId="21" pageNumber="582" unit="m" value="1327.0">1327 m</quantity>
a.s.l.
</elevation>
,
<date id="FFAB1017FFFF225E0CAC7FDEFEA2556A" box="[251,381,1533,1559]" pageId="21" pageNumber="582" value="2018-11-11">
<collectingDate id="EFEFE9FFFFFF225E0CAC7FDEFEA2556A" box="[251,381,1533,1559]" pageId="21" pageNumber="582" value="2018-11-11">11.XI.2018</collectingDate>
</date>
, leg.
<collectorName id="26E05301FFFF225E0DEA7FDDFDF1556A" box="[445,558,1533,1560]" pageId="21" pageNumber="582">B. Suksai</collectorName>
,
<collectorName id="26E05301FFFF225E0E6B7FDEFD16556A" box="[572,713,1533,1559]" pageId="21" pageNumber="582">S. Kullasoot</collectorName>
&amp;
<collectorName id="26E05301FFFF225E0EA57FDDFC5A556A" box="[754,901,1533,1560]" pageId="21" pageNumber="582">P. Sapewisut</collectorName>
,
<specimenCount id="9D13FD5EFFFF225E0FC47FDEFC3B556A" box="[915,996,1533,1559]" pageId="21" pageNumber="582" type="male">1 male</specimenCount>
,
<specimenCount id="9D13FD5EFFFF225E0FA57FDEFB89556A" box="[1010,1110,1533,1559]" pageId="21" pageNumber="582" type="female">1 female</specimenCount>
(
<collectionCode id="ED04AE12FFFF225E08307FDEFB0B556A" box="[1127,1236,1533,1559]" pageId="21" pageNumber="582">THNHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D3C8AFFFF225E08BD7FDEFC725522" collectingDate="2019-05-02" collectionCode="THNHM" collectorName="B. Suksai &amp; P. Sapewisut &amp; N. Plangklang" country="Thailand" county="Stream" elevation="1089" latitude="19.750153" location="Tap Tao Subdistrict" longLatPrecision="1" longitude="100.399864" municipality="Ban Phithak Thai" pageId="21" pageNumber="582" specimenCount="1" specimenCount-female="1" stateProvince="Chiang Rai">
<collectingCounty id="62CB4E5BFFFF225E08BD7FDEFAE6556A" box="[1258,1337,1533,1559]" pageId="21" pageNumber="582">Stream</collectingCounty>
close to
<collectingMunicipality id="6BCEACADFFFF225E0CC07C01FE805546" box="[151,351,1569,1596]" pageId="21" pageNumber="582">Ban Phithak Thai</collectingMunicipality>
,
<location id="8ECA600CFFFF225E0D3C7C01FD955546" LSID="urn:lsid:plazi:treatment:03BC87C1FFFF225C0CC07E38FE79501B:8ECA600CFFFF225E0D3C7C01FD955546" box="[363,586,1569,1596]" country="Thailand" county="Stream" latitude="19.750153" longLatPrecision="1" longitude="100.399864" municipality="Ban Phithak Thai" name="Tap Tao Subdistrict" pageId="21" pageNumber="582" stateProvince="Chiang Rai">Tap Tao Subdistrict</location>
,
<location id="8ECA600CFFFF225E0E027C02FCD55546" LSID="urn:lsid:plazi:treatment:03BC87C1FFFF225C0CC07E38FE79501B:8ECA600CFFFF225E0E027C02FCD55546" box="[597,778,1569,1595]" country="Thailand" county="Stream" latitude="19.750153" longLatPrecision="1" longitude="100.399864" municipality="Ban Phithak Thai" name="Thoeng District" pageId="21" pageNumber="582" stateProvince="Chiang Rai">Thoeng District</location>
,
<geoCoordinate id="EE215010FFFF225E0F417C02FC1D5541" box="[790,962,1569,1596]" degrees="19" direction="north" minutes="45" orientation="latitude" pageId="21" pageNumber="582" precision="1" seconds="00.55" value="19.750153">19°4500.55”N</geoCoordinate>
<geoCoordinate id="EE215010FFFF225E0F9D7C02FB5B5541" box="[970,1156,1569,1596]" degrees="100" direction="east" minutes="23" orientation="longitude" pageId="21" pageNumber="582" precision="1" seconds="59.53" value="100.399864">100°2359.53”E</geoCoordinate>
,
<elevation id="0038D1E4FFFF225E08C77C02FAFC5546" box="[1168,1315,1569,1596]" metricMagnitude="3" metricUnit="m" metricValue="1.089" pageId="21" pageNumber="582" unit="m" value="1089.0">
<quantity id="4CED9B32FFFF225E08C77C02FB395541" box="[1168,1254,1569,1596]" metricMagnitude="3" metricUnit="m" metricValue="1.089" pageId="21" pageNumber="582" unit="m" value="1089.0">1089 m</quantity>
a.s.l.
</elevation>
,
<date id="FFAB1017FFFF225E09667C02FA475546" box="[1329,1432,1569,1595]" pageId="21" pageNumber="582" value="2019-05-02">
<collectingDate id="EFEFE9FFFFFF225E09667C02FA475546" box="[1329,1432,1569,1595]" pageId="21" pageNumber="582" value="2019-05-02">2.V.2019</collectingDate>
</date>
, leg.
<collectorName id="26E05301FFFF225E0C9C7C65FEE25522" box="[203,317,1605,1632]" pageId="21" pageNumber="582">B. Suksai</collectorName>
,
<collectorName id="26E05301FFFF225E0D1A7C65FE015522" box="[333,478,1605,1632]" pageId="21" pageNumber="582">P. Sapewisut</collectorName>
&amp;
<collectorName id="26E05301FFFF225E0E5D7C65FD6E5522" box="[522,689,1605,1632]" pageId="21" pageNumber="582">N. Plangklang</collectorName>
,
<specimenCount id="9D13FD5EFFFF225E0E977C66FCFA5522" box="[704,805,1605,1631]" pageId="21" pageNumber="582" type="female">1 female</specimenCount>
(
<collectionCode id="ED04AE12FFFF225E0F6F7C66FC7A5522" box="[824,933,1605,1631]" pageId="21" pageNumber="582">THNHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D3C8AFFFF225E0FEB7C66FEF655DA" collectingDate="2017-05-28" collectionCode="KKU" collectorName="P. Sapewisut" country="Thailand" county="Stream" elevation="1299" latitude="19.737028" location="Tap Tao Subdistrict" longLatPrecision="1" longitude="100.40061" municipality="Ban Thai Samakkhi" pageId="21" pageNumber="582" specimenCount="1" specimenCount-male="1" stateProvince="Chiang Rai">
<collectingCounty id="62CB4E5BFFFF225E0FEB7C66FBD45522" box="[956,1035,1605,1631]" pageId="21" pageNumber="582">Stream</collectingCounty>
close to
<collectingMunicipality id="6BCEACADFFFF225E082C7C65FABB5522" box="[1147,1380,1605,1632]" pageId="21" pageNumber="582">Ban Thai Samakkhi</collectingMunicipality>
,
<location id="8ECA600CFFFF225E09247C65FE9755FE" LSID="urn:lsid:plazi:treatment:03BC87C1FFFF225C0CC07E38FE79501B:8ECA600CFFFF225E09247C65FE9755FE" country="Thailand" county="Stream" latitude="19.737028" longLatPrecision="1" longitude="100.40061" municipality="Ban Thai Samakkhi" name="Tap Tao Subdistrict" pageId="21" pageNumber="582" stateProvince="Chiang Rai">Tap Tao Subdistrict</location>
,
<location id="8ECA600CFFFF225E0D0F7C4AFDCF55FE" LSID="urn:lsid:plazi:treatment:03BC87C1FFFF225C0CC07E38FE79501B:8ECA600CFFFF225E0D0F7C4AFDCF55FE" box="[344,528,1641,1667]" country="Thailand" county="Stream" latitude="19.737028" longLatPrecision="1" longitude="100.40061" municipality="Ban Thai Samakkhi" name="Thoeng District" pageId="21" pageNumber="582" stateProvince="Chiang Rai">Thoeng District</location>
,
<geoCoordinate id="EE215010FFFF225E0E487C4AFD6255FE" box="[543,701,1641,1667]" degrees="19" direction="north" minutes="44" orientation="latitude" pageId="21" pageNumber="582" precision="1" seconds="13.3" value="19.737028">19°4413.3”N</geoCoordinate>
<geoCoordinate id="EE215010FFFF225E0E9E7C4AFCAA55FE" box="[713,885,1641,1667]" degrees="100" direction="east" minutes="24" orientation="longitude" pageId="21" pageNumber="582" precision="1" seconds="02.2" value="100.40061">100°2402.2”E</geoCoordinate>
,
<elevation id="0038D1E4FFFF225E0FD37C4AFBC055FE" box="[900,1055,1641,1668]" metricMagnitude="3" metricUnit="m" metricValue="1.299" pageId="21" pageNumber="582" unit="m" value="1299.0">
<quantity id="4CED9B32FFFF225E0FD37C4AFC0155F9" box="[900,990,1641,1668]" metricMagnitude="3" metricUnit="m" metricValue="1.299" pageId="21" pageNumber="582" unit="m" value="1299.0">1299 m</quantity>
a.s.l.
</elevation>
,
<date id="FFAB1017FFFF225E08677C4AFB7A55FE" box="[1072,1189,1641,1667]" pageId="21" pageNumber="582" value="2017-05-28">
<collectingDate id="EFEFE9FFFFFF225E08677C4AFB7A55FE" box="[1072,1189,1641,1667]" pageId="21" pageNumber="582" value="2017-05-28">28.V.2017</collectingDate>
</date>
, leg.
<collectorName id="26E05301FFFF225E08BD7C49FAA055FE" box="[1258,1407,1641,1668]" pageId="21" pageNumber="582">P. Sapewisut</collectorName>
,
<specimenCount id="9D13FD5EFFFF225E09D97C4AFF1255DA" pageId="21" pageNumber="582" type="male">1 male</specimenCount>
(
<collectionCode id="ED04AE12FFFF225E0C887CAEFEFE55DA" box="[223,289,1677,1703]" country="Thailand" httpUri="http://grbio.org/cool/pwjq-e3rs" name="Herbarium, Department of Biology, Khon Kaen University" pageId="21" pageNumber="582">KKU</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B7D3C8AFFFF225E0D6F7CAEFDA455D5" box="[312,635,1677,1704]" collectingDate="2017-05-28" collectionCode="KKU, THNHM" collectorName="P. Sapewisut" country="Thailand" county="Stream" elevation="1299" isTailEnumeration="true" latitude="19.737028" location="Tap Tao Subdistrict" longLatPrecision="1" longitude="100.40061" municipality="Ban Thai Samakkhi" pageId="21" pageNumber="582" specimenCount="2" specimenCount-female="2" stateProvince="Chiang Rai">
<specimenCount id="9D13FD5EFFFF225E0D6F7CAEFE7955DA" box="[312,422,1677,1703]" pageId="21" pageNumber="582" type="female">2 females</specimenCount>
(
<collectionCode id="ED04AE12FFFF225E0DEF7CAEFE2655DA" box="[440,505,1677,1703]" country="Thailand" httpUri="http://grbio.org/cool/pwjq-e3rs" name="Herbarium, Department of Biology, Khon Kaen University" pageId="21" pageNumber="582">KKU</collectionCode>
,
<collectionCode id="ED04AE12FFFF225E0E507CADFDAC55D5" box="[519,627,1678,1704]" pageId="21" pageNumber="582">THNHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D3C8AFFFF225E0EDF7CADFD5C55B6" collectingDate="2018-11-10" collectionCode="THNHM" collectorName="B. Suksai &amp; S. Kullasoot &amp; P. Sapewisut" country="Thailand" county="Stream" elevation="1299" isTailEnumeration="true" latitude="19.737028" location="Tap Tao Subdistrict" longLatPrecision="1" longitude="100.40061" municipality="Ban Thai Samakkhi" pageId="21" pageNumber="582" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Chiang Rai">
same locality except for,
<date id="FFAB1017FFFF225E0FFA7CAEFBF055DA" box="[941,1071,1677,1703]" pageId="21" pageNumber="582" value="2018-11-10">
<collectingDate id="EFEFE9FFFFFF225E0FFA7CAEFBF055DA" box="[941,1071,1677,1703]" pageId="21" pageNumber="582" value="2018-11-10">10.XI.2018</collectingDate>
</date>
, leg.
<collectorName id="26E05301FFFF225E08277CADFB3E55DA" box="[1136,1249,1677,1704]" pageId="21" pageNumber="582">B. Suksai</collectorName>
,
<collectorName id="26E05301FFFF225E08B87CAEFAA255DA" box="[1263,1405,1677,1703]" pageId="21" pageNumber="582">S. Kullasoot</collectorName>
&amp;
<collectorName id="26E05301FFFF225E0CC07C91FEF655B6" box="[151,297,1713,1740]" pageId="21" pageNumber="582">P. Sapewisut</collectorName>
,
<specimenCount id="9D13FD5EFFFF225E0D627C92FE5B55B6" box="[309,388,1713,1739]" pageId="21" pageNumber="582" type="male">1 male</specimenCount>
,
<specimenCount id="9D13FD5EFFFF225E0DC77C92FE2255B6" box="[400,509,1713,1739]" pageId="21" pageNumber="582" type="female">2 females</specimenCount>
(
<collectionCode id="ED04AE12FFFF225E0E5A7C92FDA555B6" box="[525,634,1713,1739]" pageId="21" pageNumber="582">THNHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D3C8AFFFF225E0ED87C91FD99558D" collectingDate="2019-05-02" collectionCode="KKU, THNHM" collectorName="B. Suksai &amp; P. Sapewisut &amp; N. Plangklang" country="Thailand" county="Stream" elevation="1299" isTailEnumeration="true" latitude="19.737028" location="Tap Tao Subdistrict" longLatPrecision="1" longitude="100.40061" municipality="Ban Thai Samakkhi" pageId="21" pageNumber="582" specimenCount="8" specimenCount-male="8" stateProvince="Chiang Rai">
same locality except for,
<date id="FFAB1017FFFF225E0FFB7C92FBCB55B6" box="[940,1044,1713,1739]" pageId="21" pageNumber="582" value="2019-05-02">
<collectingDate id="EFEFE9FFFFFF225E0FFB7C92FBCB55B6" box="[940,1044,1713,1739]" pageId="21" pageNumber="582" value="2019-05-02">2.V.2019</collectingDate>
</date>
, leg.
<collectorName id="26E05301FFFF225E08067C91FB1F55B6" box="[1105,1216,1713,1740]" pageId="21" pageNumber="582">B. Suksai</collectorName>
,
<collectorName id="26E05301FFFF225E089B7C91FA8455B6" box="[1228,1371,1713,1740]" pageId="21" pageNumber="582">P. Sapewisut</collectorName>
&amp;
<collectorName id="26E05301FFFF225E09D67C91FEC75592" pageId="21" pageNumber="582">N. Plangklang</collectorName>
,
<specimenCount id="9D13FD5EFFFF225E0D747CF6FEA75592" box="[291,376,1749,1775]" pageId="21" pageNumber="582" type="male">8 males</specimenCount>
(
<collectionCode id="ED04AE12FFFF225E0DD07CF6FE175592" box="[391,456,1749,1775]" country="Thailand" httpUri="http://grbio.org/cool/pwjq-e3rs" name="Herbarium, Department of Biology, Khon Kaen University" pageId="21" pageNumber="582">KKU</collectionCode>
,
<collectionCode id="ED04AE12FFFF225E0D847CF5FDE1558D" box="[467,574,1750,1776]" pageId="21" pageNumber="582">THNHM</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B7D3C8AFFFF225E0E077CF6FCFA5592" box="[592,805,1749,1775]" collectingDate="2019-05-02" collectionCode="GTGC" collectorName="B. Suksai &amp; P. Sapewisut &amp; N. Plangklang" country="Thailand" county="Stream" elevation="1299" isTailEnumeration="true" latitude="19.737028" location="Tap Tao Subdistrict" longLatPrecision="1" longitude="100.40061" municipality="Ban Thai Samakkhi" pageId="21" pageNumber="582" specimenCount="2" specimenCount-female="2" stateProvince="Chiang Rai">
<specimenCount id="9D13FD5EFFFF225E0E077CF6FD635592" box="[592,700,1749,1775]" pageId="21" pageNumber="582" type="female">2 females</specimenCount>
(
<collectionCode id="ED04AE12FFFF225E0E9C7CF6FCC15592" box="[715,798,1749,1775]" pageId="21" pageNumber="582">GTGC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7D3C8AFFFF225E0F787CF6FB22546E" collectingDate="2002-03-13" collectionCode="UMC" country="Thailand" elevation="900" latitude="19.2" location="Namtok Ton Tong" longLatPrecision="1273" longitude="101.066666" municipality="Doi Phuka Natl. Park." pageId="21" pageNumber="582" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Nan">
<emphasis id="B961EAC5FFFF225E0F787CF6FC055592" bold="true" box="[815,986,1749,1775]" pageId="21" pageNumber="582">
<collectingRegion id="49D1F835FFFF225E0F787CF6FC095592" box="[815,982,1749,1775]" country="Thailand" name="Nan" pageId="21" pageNumber="582">Nan Province</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6BCEACADFFFF225E0FB67CF6FB065592" box="[993,1241,1749,1775]" pageId="21" pageNumber="582">Doi Phuka Natl. Park.</collectingMunicipality>
, creek at
<location id="8ECA600CFFFF225E09157CF6FED95469" LSID="urn:lsid:plazi:treatment:03BC87C1FFFF225C0CC07E38FE79501B:8ECA600CFFFF225E09157CF6FED95469" country="Thailand" latitude="19.2" longLatPrecision="1273" longitude="101.066666" municipality="Doi Phuka Natl. Park." name="Namtok Ton Tong" pageId="21" pageNumber="582" stateProvince="Nan">Namtok Ton Tong</location>
,
<geoCoordinate id="EE215010FFFF225E0D457CDAFEAC546E" box="[274,371,1785,1811]" degrees="19" direction="north" minutes="12" orientation="latitude" pageId="21" pageNumber="582" precision="925" value="19.2">19°12N</geoCoordinate>
<geoCoordinate id="EE215010FFFF225E0D2C7CDAFE36546E" box="[379,489,1785,1811]" degrees="101" direction="east" minutes="04" orientation="longitude" pageId="21" pageNumber="582" precision="925" value="101.066666">101°04E</geoCoordinate>
,
<elevation id="0038D1E4FFFF225E0DA17CDAFDA3546E" box="[502,636,1785,1812]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="21" pageNumber="582" unit="m" value="900.0">
<quantity id="4CED9B32FFFF225E0DA17CDAFDE15469" box="[502,574,1785,1812]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="21" pageNumber="582" unit="m" value="900.0">900 m</quantity>
a.s.l.
</elevation>
,
<date id="FFAB1017FFFF225E0EDD7CDAFCD5546E" box="[650,778,1785,1811]" pageId="21" pageNumber="582" value="2002-03-13">
<collectingDate id="EFEFE9FFFFFF225E0EDD7CDAFCD5546E" box="[650,778,1785,1811]" pageId="21" pageNumber="582" value="2002-03-13">13.III.2002</collectingDate>
</date>
, leg. CMU teams,
<specimenCount id="9D13FD5EFFFF225E0FB67CDAFBEF546E" box="[993,1072,1785,1811]" pageId="21" pageNumber="582" type="male">1 male</specimenCount>
,
<specimenCount id="9D13FD5EFFFF225E086A7CDAFB40546E" box="[1085,1183,1785,1811]" pageId="21" pageNumber="582" type="female">1 female</specimenCount>
(
<collectionCode id="ED04AE12FFFF225E08F87CDAFB2A546E" box="[1199,1269,1785,1811]" pageId="21" pageNumber="582">UMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D3C8AFFFF225E095D7CDAFCE85421" collectingDate="2017-05-30" collectionCode="KKU, THNHM" collectorName="P. Sapewisut" country="Thailand" county="Khun Sathan Natl. Park." elevation="1474" latitude="18.273367" location="San Ta Subdistrict" longLatPrecision="1" longitude="100.48633" municipality="Ban Khun Sathan" pageId="21" pageNumber="582" specimenCount="3" specimenCount-female="3" stateProvince="Nan">
<collectingCounty id="62CB4E5BFFFF225E095D7CDAFECC544A" pageId="21" pageNumber="582">Khun Sathan Natl. Park.</collectingCounty>
,
<collectingMunicipality id="6BCEACADFFFF225E0D487D3DFE37544A" box="[287,488,1821,1848]" pageId="21" pageNumber="582">Ban Khun Sathan</collectingMunicipality>
,
<location id="8ECA600CFFFF225E0DA37D3EFD1C544A" LSID="urn:lsid:plazi:treatment:03BC87C1FFFF225C0CC07E38FE79501B:8ECA600CFFFF225E0DA37D3EFD1C544A" box="[500,707,1821,1848]" country="Thailand" county="Khun Sathan Natl. Park." latitude="18.273367" longLatPrecision="1" longitude="100.48633" municipality="Ban Khun Sathan" name="San Ta Subdistrict" pageId="21" pageNumber="582" stateProvince="Nan">San Ta Subdistrict</location>
,
<location id="8ECA600CFFFF225E0E987D3DFC5F544A" LSID="urn:lsid:plazi:treatment:03BC87C1FFFF225C0CC07E38FE79501B:8ECA600CFFFF225E0E987D3DFC5F544A" box="[719,896,1821,1848]" country="Thailand" county="Khun Sathan Natl. Park." latitude="18.273367" longLatPrecision="1" longitude="100.48633" municipality="Ban Khun Sathan" name="Na Noi District" pageId="21" pageNumber="582" stateProvince="Nan">Na Noi District</location>
,
<geoCoordinate id="EE215010FFFF225E0FDC7D3EFBE8544A" box="[907,1079,1821,1847]" degrees="18" direction="north" minutes="16" orientation="latitude" pageId="21" pageNumber="582" precision="1" seconds="24.12" value="18.273367">18°1624.12”N</geoCoordinate>
<geoCoordinate id="EE215010FFFF225E08687D3EFB26544A" box="[1087,1273,1821,1847]" degrees="100" direction="east" minutes="29" orientation="longitude" pageId="21" pageNumber="582" precision="1" seconds="10.80" value="100.48633">100°2910.80”E</geoCoordinate>
,
<elevation id="0038D1E4FFFF225E09537D3EFA48544A" box="[1284,1431,1821,1848]" metricMagnitude="3" metricUnit="m" metricValue="1.474" pageId="21" pageNumber="582" unit="m" value="1474.0">
<quantity id="4CED9B32FFFF225E09537D3EFA855445" box="[1284,1370,1821,1848]" metricMagnitude="3" metricUnit="m" metricValue="1.474" pageId="21" pageNumber="582" unit="m" value="1474.0">1474 m</quantity>
a.s.l.
</elevation>
,
<date id="FFAB1017FFFF225E0CC07D62FED25426" box="[151,269,1857,1883]" pageId="21" pageNumber="582" value="2017-05-30">
<collectingDate id="EFEFE9FFFFFF225E0CC07D62FED25426" box="[151,269,1857,1883]" pageId="21" pageNumber="582" value="2017-05-30">30.V.2017</collectingDate>
</date>
, leg.
<collectorName id="26E05301FFFF225E0D187D61FE3C5426" box="[335,483,1857,1884]" pageId="21" pageNumber="582">P. Sapewisut</collectorName>
,
<specimenCount id="9D13FD5EFFFF225E0DA57D62FDBD5426" box="[498,610,1857,1883]" pageId="21" pageNumber="582" type="female">3 females</specimenCount>
(
<collectionCode id="ED04AE12FFFF225E0E237D62FD6A5426" box="[628,693,1857,1883]" country="Thailand" httpUri="http://grbio.org/cool/pwjq-e3rs" name="Herbarium, Department of Biology, Khon Kaen University" pageId="21" pageNumber="582">KKU</collectionCode>
,
<collectionCode id="ED04AE12FFFF225E0E947D61FCF05421" box="[707,815,1858,1884]" pageId="21" pageNumber="582">THNHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D3C8AFFFF225E0F117D61FF3654DE" collectingDate="2017-05-29" collectionCode="KKU" collectorName="P. Sapewisut" country="Thailand" county="Ban Mueng Li" elevation="786" latitude="18.252344" location="Na Muen District" longLatPrecision="1" longitude="100.49326" municipality="Mueng Li Subdistrict" pageId="21" pageNumber="582" specimenCount="1" specimenCount-male="1" stateProvince="Nan">
same locality except for,
<collectingCounty id="62CB4E5BFFFF225E083A7D61FAC55426" box="[1133,1306,1857,1884]" pageId="21" pageNumber="582">Ban Mueng Li</collectingCounty>
,
<collectingMunicipality id="6BCEACADFFFF225E097E7D61FECC5402" pageId="21" pageNumber="582">Mueng Li Subdistrict</collectingMunicipality>
,
<location id="8ECA600CFFFF225E0D487D45FE375402" LSID="urn:lsid:plazi:treatment:03BC87C1FFFF225C0CC07E38FE79501B:8ECA600CFFFF225E0D487D45FE375402" box="[287,488,1893,1920]" country="Thailand" county="Ban Mueng Li" latitude="18.252344" longLatPrecision="1" longitude="100.49326" municipality="Mueng Li Subdistrict" name="Na Muen District" pageId="21" pageNumber="582" stateProvince="Nan">Na Muen District</location>
,
<geoCoordinate id="EE215010FFFF225E0DA37D46FD7F54FD" box="[500,672,1893,1920]" degrees="18" direction="north" minutes="15" orientation="latitude" pageId="21" pageNumber="582" precision="1" seconds="08.44" value="18.252344">18°1508.44”N</geoCoordinate>
<geoCoordinate id="EE215010FFFF225E0EFF7D46FCBD54FD" box="[680,866,1893,1920]" degrees="100" direction="east" minutes="29" orientation="longitude" pageId="21" pageNumber="582" precision="1" seconds="35.77" value="100.49326">100°2935.77”E</geoCoordinate>
,
<elevation id="0038D1E4FFFF225E0F397D46FC2C5402" box="[878,1011,1893,1920]" metricMagnitude="2" metricUnit="m" metricValue="7.859999999999999" pageId="21" pageNumber="582" unit="m" value="786.0">
<quantity id="4CED9B32FFFF225E0F397D46FC6954FD" box="[878,950,1893,1920]" metricMagnitude="2" metricUnit="m" metricValue="7.859999999999999" pageId="21" pageNumber="582" unit="m" value="786.0">786 m</quantity>
a.s.l.
</elevation>
,
<date id="FFAB1017FFFF225E08577D46FBAA5402" box="[1024,1141,1893,1919]" pageId="21" pageNumber="582" value="2017-05-29">
<collectingDate id="EFEFE9FFFFFF225E08577D46FBAA5402" box="[1024,1141,1893,1919]" pageId="21" pageNumber="582" value="2017-05-29">29.V.2017</collectingDate>
</date>
, leg.
<collectorName id="26E05301FFFF225E08E47D45FA9B5402" box="[1203,1348,1893,1920]" pageId="21" pageNumber="582">P. Sapewisut</collectorName>
,
<specimenCount id="9D13FD5EFFFF225E09077D46FA435402" box="[1360,1436,1893,1919]" pageId="21" pageNumber="582" type="male">1 male</specimenCount>
(
<collectionCode id="ED04AE12FFFF225E0CC87DAAFF3E54DE" box="[159,225,1929,1955]" country="Thailand" httpUri="http://grbio.org/cool/pwjq-e3rs" name="Herbarium, Department of Biology, Khon Kaen University" pageId="21" pageNumber="582">KKU</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B7D3C8AFFFF225E0CA17DAAFDEA54D9" box="[246,565,1929,1956]" collectingDate="2017-05-29" collectionCode="KKU, THNHM" collectorName="P. Sapewisut" country="Thailand" county="Ban Mueng Li" elevation="786" isTailEnumeration="true" latitude="18.252344" location="Na Muen District" longLatPrecision="1" longitude="100.49326" municipality="Mueng Li Subdistrict" pageId="21" pageNumber="582" specimenCount="3" specimenCount-female="3" stateProvince="Nan">
<specimenCount id="9D13FD5EFFFF225E0CA17DAAFEBC54DE" box="[246,355,1929,1955]" pageId="21" pageNumber="582" type="female">3 females</specimenCount>
(
<collectionCode id="ED04AE12FFFF225E0D237DAAFE6A54DE" box="[372,437,1929,1955]" country="Thailand" httpUri="http://grbio.org/cool/pwjq-e3rs" name="Herbarium, Department of Biology, Khon Kaen University" pageId="21" pageNumber="582">KKU</collectionCode>
,
<collectionCode id="ED04AE12FFFF225E0D967DA9FDF254D9" box="[449,557,1930,1956]" pageId="21" pageNumber="582">THNHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D3C8AFFFF225E0E167DA9FECB5496" collectingDate="2017-05-30" collectionCode="THNHM" collectorName="P. Sapewisut" country="Thailand" county="Khun Sathan Watershed Management Station" elevation="1334" isTailEnumeration="true" latitude="18.278797" location="Na Noi District" longLatPrecision="1" longitude="100.50398" municipality="San Ta Subdistrict" pageId="21" pageNumber="582" specimenCount="1" specimenCount-male="1" stateProvince="Nan">
same locality except for,
<collectingCounty id="62CB4E5BFFFF225E0F377DAAFABA54DE" box="[864,1381,1929,1956]" pageId="21" pageNumber="582">Khun Sathan Watershed Management Station</collectingCounty>
,
<collectingMunicipality id="6BCEACADFFFF225E09257DAAFEEA54BA" pageId="21" pageNumber="582">San Ta Subdistrict</collectingMunicipality>
,
<location id="8ECA600CFFFF225E0D687D8DFE3254BA" LSID="urn:lsid:plazi:treatment:03BC87C1FFFF225C0CC07E38FE79501B:8ECA600CFFFF225E0D687D8DFE3254BA" box="[319,493,1965,1992]" country="Thailand" county="Khun Sathan Watershed Management Station" latitude="18.278797" longLatPrecision="1" longitude="100.50398" municipality="San Ta Subdistrict" name="Na Noi District" pageId="21" pageNumber="582" stateProvince="Nan">Na Noi District</location>
,
<geoCoordinate id="EE215010FFFF225E0DA07D8EFD7B54BA" box="[503,676,1965,1991]" degrees="18" direction="north" minutes="16" orientation="latitude" pageId="21" pageNumber="582" precision="1" seconds="43.67" value="18.278797">18°1643.67”N</geoCoordinate>
<geoCoordinate id="EE215010FFFF225E0EFD7D8EFCBC54BA" box="[682,867,1965,1991]" degrees="100" direction="east" minutes="30" orientation="longitude" pageId="21" pageNumber="582" precision="1" seconds="14.34" value="100.50398">100°3014.34”E</geoCoordinate>
,
<elevation id="0038D1E4FFFF225E0F397D8EFC2254BA" box="[878,1021,1965,1992]" metricMagnitude="3" metricUnit="m" metricValue="1.334" pageId="21" pageNumber="582" unit="m" value="1334.0">
<quantity id="4CED9B32FFFF225E0F397D8EFC1D54B5" box="[878,962,1965,1992]" metricMagnitude="3" metricUnit="m" metricValue="1.334" pageId="21" pageNumber="582" unit="m" value="1334.0">1334 m</quantity>
a.s.l.
</elevation>
,
<date id="FFAB1017FFFF225E085E7D8EFBA154BA" box="[1033,1150,1965,1991]" pageId="21" pageNumber="582" value="2017-05-30">
<collectingDate id="EFEFE9FFFFFF225E085E7D8EFBA154BA" box="[1033,1150,1965,1991]" pageId="21" pageNumber="582" value="2017-05-30">30.V.2017</collectingDate>
</date>
, leg.
<collectorName id="26E05301FFFF225E08EF7D8DFA9854BA" box="[1208,1351,1965,1992]" pageId="21" pageNumber="582">P. Sapewisut</collectorName>
,
<specimenCount id="9D13FD5EFFFF225E09057D8EFA4354BA" box="[1362,1436,1965,1991]" pageId="21" pageNumber="582" type="male">1 male</specimenCount>
(
<collectionCode id="ED04AE12FFFF225E0CC87DF2FED35496" box="[159,268,2001,2027]" pageId="21" pageNumber="582">THNHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D3C8AFFFF225D0D767DF1FED453A8" collectingDate="2017-05-30" collectionCode="KKU, THNHM" collectorName="P. Sapewisut" country="Thailand" county="Water" elevation="1356" isTailEnumeration="true" lastPageId="22" lastPageNumber="583" latitude="18.278349" location="Khun Sathan National Park" longLatPrecision="1" longitude="100.507484" municipality="Weir" pageId="21" pageNumber="582" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Nan">
same locality except for,
<collectingCounty id="62CB4E5BFFFF225E0E177DF1FD5D5491" box="[576,642,2002,2028]" pageId="21" pageNumber="582">Water</collectingCounty>
supply
<collectingMunicipality id="6BCEACADFFFF225E0E897DF2FCC95496" box="[734,790,2001,2027]" pageId="21" pageNumber="582">Weir</collectingMunicipality>
,
<location id="8ECA600CFFFF225E0F757DF2FB825496" LSID="urn:lsid:plazi:treatment:03BC87C1FFFF225C0CC07E38FE79501B:8ECA600CFFFF225E0F757DF2FB825496" box="[802,1117,2001,2027]" country="Thailand" county="Water" latitude="18.278349" longLatPrecision="1" longitude="100.507484" municipality="Weir" name="Khun Sathan National Park" pageId="21" pageNumber="582" stateProvince="Nan">Khun Sathan National Park</location>
,
<location id="8ECA600CFFFF225E083D7DF2FAE35496" LSID="urn:lsid:plazi:treatment:03BC87C1FFFF225C0CC07E38FE79501B:8ECA600CFFFF225E083D7DF2FAE35496" box="[1130,1340,2001,2028]" country="Thailand" county="Water" latitude="18.278349" longLatPrecision="1" longitude="100.507484" municipality="Weir" name="San Ta Subdistrict" pageId="21" pageNumber="582" stateProvince="Nan">San Ta Subdistrict</location>
,
<location id="8ECA600CFFFF225D091E7DF1FF3153CC" LSID="urn:lsid:plazi:treatment:03BC87C1FFFF225C0CC07E38FE79501B:8ECA600CFFFF225D091E7DF1FF3153CC" country="Thailand" county="Water" lastPageId="22" lastPageNumber="583" latitude="18.278349" longLatPrecision="1" longitude="100.507484" municipality="Weir" name="Na Noi District" pageId="21" pageNumber="582" stateProvince="Nan">Na Noi District</location>
,
<geoCoordinate id="EE215010FFFC225D0CAC7AB4FE7853CC" box="[251,423,151,177]" degrees="18" direction="north" minutes="16" orientation="latitude" pageId="22" pageNumber="583" precision="1" seconds="42.06" value="18.278349">18°1642.06”N</geoCoordinate>
<geoCoordinate id="EE215010FFFC225D0DE77AB5FDB553CC" box="[432,618,150,177]" degrees="100" direction="east" minutes="30" orientation="longitude" pageId="22" pageNumber="583" precision="1" seconds="26.95" value="100.507484">100°3026.95”E</geoCoordinate>
,
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<quantity id="4CED9B32FFFC225D0E207AB5FD1153CC" box="[631,718,150,177]" metricMagnitude="3" metricUnit="m" metricValue="1.3559999999999999" pageId="22" pageNumber="583" unit="m" value="1356.0">1356 m</quantity>
a.s.l.
</elevation>
,
<date id="FFAB1017FFFC225D0F4C7AB4FC4E53CC" box="[795,913,151,177]" pageId="22" pageNumber="583" value="2017-05-30">
<collectingDate id="EFEFE9FFFFFC225D0F4C7AB4FC4E53CC" box="[795,913,151,177]" pageId="22" pageNumber="583" value="2017-05-30">30.V.2017</collectingDate>
</date>
, leg.
<collectorName id="26E05301FFFC225D0F877AB4FBBC53CC" box="[976,1123,151,177]" pageId="22" pageNumber="583">P. Sapewisut</collectorName>
,
<specimenCount id="9D13FD5EFFFC225D08277AB4FB1453CC" box="[1136,1227,151,177]" pageId="22" pageNumber="583" type="male">2 males</specimenCount>
,
<specimenCount id="9D13FD5EFFFC225D088F7AB4FA9953CC" box="[1240,1350,151,177]" pageId="22" pageNumber="583" type="female">2 females</specimenCount>
(
<collectionCode id="ED04AE12FFFC225D09007AB4FA4753CC" box="[1367,1432,151,177]" country="Thailand" httpUri="http://grbio.org/cool/pwjq-e3rs" name="Herbarium, Department of Biology, Khon Kaen University" pageId="22" pageNumber="583">KKU</collectionCode>
,
<collectionCode id="ED04AE12FFFC225D0CC07A98FEDC53A8" box="[151,259,187,213]" pageId="22" pageNumber="583">THNHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D3C8AFFFC225D0D417A98FAB95384" collectingDate="2013-01-02" collectionCode="KKU" collectorName="K. Rattanachan" country="Thailand" county="Mae Charim Natl. Park." elevation="898" latitude="18.583704" location="Beside of Ban Nampang-Romklao Road" longLatPrecision="1" longitude="101.06459" municipality="Huay Tao" pageId="22" pageNumber="583" specimenCount="1" specimenCount-female="1" stateProvince="Nan">
<collectingCounty id="62CB4E5BFFFC225D0D417A98FDF953A8" box="[278,550,187,213]" pageId="22" pageNumber="583">Mae Charim Natl. Park.</collectingCounty>
,
<collectingMunicipality id="6BCEACADFFFC225D0E667A98FD4153A8" box="[561,670,187,213]" pageId="22" pageNumber="583">Huay Tao</collectingMunicipality>
stream,
<location id="8ECA600CFFFC225D0EA97A98FB1953A8" LSID="urn:lsid:plazi:treatment:03BC87C1FFFF225C0CC07E38FE79501B:8ECA600CFFFC225D0EA97A98FB1953A8" box="[766,1222,187,213]" country="Thailand" county="Mae Charim Natl. Park." latitude="18.583704" longLatPrecision="1" longitude="101.06459" municipality="Huay Tao" name="Beside of Ban Nampang-Romklao Road" pageId="22" pageNumber="583" stateProvince="Nan">Beside of Ban Nampang-Romklao Road</location>
, close to
<location id="8ECA600CFFFC225D09787A98FE9E5384" LSID="urn:lsid:plazi:treatment:03BC87C1FFFF225C0CC07E38FE79501B:8ECA600CFFFC225D09787A98FE9E5384" country="Thailand" county="Mae Charim Natl. Park." latitude="18.583704" longLatPrecision="1" longitude="101.06459" municipality="Huay Tao" name="Huay Tao Ranger Station" pageId="22" pageNumber="583" stateProvince="Nan">Huay Tao Ranger Station</location>
,
<geoCoordinate id="EE215010FFFC225D0D1B7AFDFE275384" box="[332,504,222,249]" degrees="18" direction="north" minutes="35" orientation="latitude" pageId="22" pageNumber="583" precision="1" seconds="01.33" value="18.583704">18°3501.33”N</geoCoordinate>
<geoCoordinate id="EE215010FFFC225D0E577AFDFD665384" box="[512,697,222,249]" degrees="101" direction="east" minutes="03" orientation="longitude" pageId="22" pageNumber="583" precision="1" seconds="52.52" value="101.06459">101°0352.52”E</geoCoordinate>
,
<elevation id="0038D1E4FFFC225D0E927AFCFC975384" box="[709,840,223,249]" metricMagnitude="2" metricUnit="m" metricValue="8.98" pageId="22" pageNumber="583" unit="m" value="898.0">
<quantity id="4CED9B32FFFC225D0E927AFCFCD35384" box="[709,780,223,249]" metricMagnitude="2" metricUnit="m" metricValue="8.98" pageId="22" pageNumber="583" unit="m" value="898.0">898 m</quantity>
a.s.l.
</elevation>
,
<date id="FFAB1017FFFC225D0F027AFCFC6B5384" box="[853,948,223,249]" pageId="22" pageNumber="583" value="2013-01-02">
<collectingDate id="EFEFE9FFFFFC225D0F027AFCFC6B5384" box="[853,948,223,249]" pageId="22" pageNumber="583" value="2013-01-02">2.I.2013</collectingDate>
</date>
, leg.
<collectorName id="26E05301FFFC225D0FA77AFCFB405384" box="[1008,1183,223,249]" pageId="22" pageNumber="583">K. Rattanachan</collectorName>
,
<specimenCount id="9D13FD5EFFFC225D08FC7AFCFAD35384" box="[1195,1292,223,249]" pageId="22" pageNumber="583" type="female">1 female</specimenCount>
(
<collectionCode id="ED04AE12FFFC225D094C7AFCFA815384" box="[1307,1374,223,249]" country="Thailand" httpUri="http://grbio.org/cool/pwjq-e3rs" name="Herbarium, Department of Biology, Khon Kaen University" pageId="22" pageNumber="583">KKU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30F655CFFFC225D0C907B21FC69518C" pageId="22" pageNumber="583" type="diagnosis">
<paragraph id="8BAA36D7FFFC225D0C907B21FD2351D4" blockId="22.[151,1437,150,753]" pageId="22" pageNumber="583">
<emphasis id="B961EAC5FFFC225D0C907B21FE9D5261" bold="true" box="[199,322,258,284]" pageId="22" pageNumber="583">Diagnosis.</emphasis>
Size: Male: 12.0
<quantity id="4CED9B32FFFC225D0E5A7B20FDAE5260" box="[525,625,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="1.3699999999999999" pageId="22" pageNumber="583" unit="mm" value="13.7">13.7 mm</quantity>
, female:
<quantity id="4CED9B32FFFC225D0E827B20FCA55260" box="[725,890,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="1.175" metricValueMax="1.29" metricValueMin="1.06" pageId="22" pageNumber="583" unit="mm" value="11.75" valueMax="12.9" valueMin="10.6">10.612.9 mm</quantity>
; body form in lateral view weakly dorsoventrally convex, more depressed anteriorly than posteriorly; elytral glabrous region of male (
<figureCitation id="132E2A52FFFC225D083B7B04FB0C523C" box="[1132,1235,295,321]" captionStart="FIGURE 2" captionStartId="6.[151,260,1861,1887]" captionTargetBox="[151,1434,179,1836]" captionTargetId="figure-25@6.[151,1435,179,1836]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042660" httpUri="https://zenodo.org/record/5042660/files/figure.png" pageId="22" pageNumber="583">Figs. 2G</figureCitation>
,
<figureCitation id="132E2A52FFFC225D08B57B04FACC523C" box="[1250,1299,295,321]" captionStart="FIGURE 11" captionStartId="22.[151,260,1770,1796]" captionTargetBox="[159,1430,816,1737]" captionTargetId="figure-281@22.[151,1436,810,1745]" captionTargetPageId="22" captionText="FIGURE 11. Patrus salvazai: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042686" httpUri="https://zenodo.org/record/5042686/files/figure.png" pageId="22" pageNumber="583">11A</figureCitation>
) cordiform, along suture extending ca. 7/9 length of elytra, laterally expanded in anterior 2/3 to similar width as pronotal glabrous region, posterior 1/3 acuminate with weakly rounded margins; female elytral glabrous region (
<figureCitation id="132E2A52FFFC225D09657B4CFA4852F4" box="[1330,1431,367,393]" captionStart="FIGURE 3" captionStartId="7.[151,260,1865,1891]" captionTargetBox="[151,1434,182,1841]" captionTargetId="figure-18@7.[151,1435,179,1841]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal habitus of females of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042662" httpUri="https://zenodo.org/record/5042662/files/figure.png" pageId="22" pageNumber="583">Figs. 3G</figureCitation>
,
<figureCitation id="132E2A52FFFC225D0CC07BB0FF1B52D0" box="[151,196,403,429]" captionStart="FIGURE 11" captionStartId="22.[151,260,1770,1796]" captionTargetBox="[159,1430,816,1737]" captionTargetId="figure-281@22.[151,1436,810,1745]" captionTargetPageId="22" captionText="FIGURE 11. Patrus salvazai: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042686" httpUri="https://zenodo.org/record/5042686/files/figure.png" pageId="22" pageNumber="583">11E</figureCitation>
) similar to male, except lateral expansion of glabrous region is stronger, surpassing lateral margin of pronotal glabrous region, apical acumination weaker; elytral apex of both sexes similar, weakly rounded in appearance, without apicolateral sinuation, epipleural angle indistinct; male protibia (
<figureCitation id="132E2A52FFFC225D0F8A7BF8FBE85288" box="[989,1079,475,501]" captionStart="FIGURE 4" captionStartId="8.[151,260,707,733]" captionTargetBox="[158,1432,185,677]" captionTargetId="figure-25@8.[152,1436,179,683]" captionTargetPageId="8" captionText="FIGURE 4. Left protibia and protarsus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5042666" httpUri="https://zenodo.org/record/5042666/files/figure.png" pageId="22" pageNumber="583">Fig. 4G</figureCitation>
) cleaver shaped, constricted in basal 1/2, protibial apex truncate with prominent distolateral angle; male protarsus (
<figureCitation id="132E2A52FFFC225D083B7BDCFB175164" box="[1132,1224,511,537]" captionStart="FIGURE 4" captionStartId="8.[151,260,707,733]" captionTargetBox="[158,1432,185,677]" captionTargetId="figure-25@8.[152,1436,179,683]" captionTargetPageId="8" captionText="FIGURE 4. Left protibia and protarsus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5042666" httpUri="https://zenodo.org/record/5042666/files/figure.png" pageId="22" pageNumber="583">Fig. 4G</figureCitation>
) ca. 1/2 length of protibia, protarsomeres width weakly decreasing apically; mesoventrite without medial pit; aedeagus (
<figureCitation id="132E2A52FFFC225D097D7800FF73511C" captionStart="FIGURE 11" captionStartId="22.[151,260,1770,1796]" captionTargetBox="[159,1430,816,1737]" captionTargetId="figure-281@22.[151,1436,810,1745]" captionTargetPageId="22" captionText="FIGURE 11. Patrus salvazai: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042686" httpUri="https://zenodo.org/record/5042686/files/figure.png" pageId="22" pageNumber="583">Fig. 11B D</figureCitation>
) with median lobe constricted medially, apex with rounded lateral margins, tip papilliform, in lateral view tip dorsally up-turned (
<figureCitation id="132E2A52FFFC225D0D247848FE0851F8" box="[371,471,619,645]" captionStart="FIGURE 11" captionStartId="22.[151,260,1770,1796]" captionTargetBox="[159,1430,816,1737]" captionTargetId="figure-281@22.[151,1436,810,1745]" captionTargetPageId="22" captionText="FIGURE 11. Patrus salvazai: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042686" httpUri="https://zenodo.org/record/5042686/files/figure.png" pageId="22" pageNumber="583">Fig. 11C</figureCitation>
); female reproductive tract (
<figureCitation id="132E2A52FFFC225D0F457848FCA951F8" box="[786,886,619,645]" captionStart="FIGURE 11" captionStartId="22.[151,260,1770,1796]" captionTargetBox="[159,1430,816,1737]" captionTargetId="figure-281@22.[151,1436,810,1745]" captionTargetPageId="22" captionText="FIGURE 11. Patrus salvazai: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042686" httpUri="https://zenodo.org/record/5042686/files/figure.png" pageId="22" pageNumber="583">Fig. 11G</figureCitation>
) without additional sclerotized structure between gonocoxae (
<figureCitation id="132E2A52FFFC225D0D7778ACFE5E51D4" box="[288,385,655,681]" captionStart="FIGURE 11" captionStartId="22.[151,260,1770,1796]" captionTargetBox="[159,1430,816,1737]" captionTargetId="figure-281@22.[151,1436,810,1745]" captionTargetPageId="22" captionText="FIGURE 11. Patrus salvazai: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042686" httpUri="https://zenodo.org/record/5042686/files/figure.png" pageId="22" pageNumber="583">Fig. 11F</figureCitation>
), fertilization duct as in
<figureCitation id="132E2A52FFFC225D0EC578ACFD2851D4" box="[658,759,655,681]" captionStart="FIGURE 11" captionStartId="22.[151,260,1770,1796]" captionTargetBox="[159,1430,816,1737]" captionTargetId="figure-281@22.[151,1436,810,1745]" captionTargetPageId="22" captionText="FIGURE 11. Patrus salvazai: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042686" httpUri="https://zenodo.org/record/5042686/files/figure.png" pageId="22" pageNumber="583">Fig. 11H</figureCitation>
.
</paragraph>
<paragraph id="8BAA36D7FFFC225D0C907891FC69518C" blockId="22.[151,1437,150,753]" pageId="22" pageNumber="583">
<emphasis id="B961EAC5FFFC225D0C907891FE1A51B1" bold="true" box="[199,453,690,716]" pageId="22" pageNumber="583">Differential diagnosis</emphasis>
.
<taxonomicName id="4C154D54FFFC225D0D847890FD5F51B0" baseAuthorityName="Ochs" baseAuthorityYear="1940" box="[467,640,691,717]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="583" phylum="Arthropoda" rank="species" species="salvazai">
<emphasis id="B961EAC5FFFC225D0D847890FD5F51B0" box="[467,640,691,717]" italics="true" pageId="22" pageNumber="583">Patrus salvazai</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C154D54FFFC225D0F1A7890FBFD51B0" authority="Suksai &amp; Gustafson &amp; Sites &amp; Sangpradub, 2021" authorityName="Suksai &amp; Gustafson &amp; Sites &amp; Sangpradub" authorityYear="2021" box="[845,1058,691,717]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="583" phylum="Arthropoda" rank="species" species="phetchabunensis" status="sp. nov.">
<emphasis id="B961EAC5FFFC225D0F1A7890FBFD51B0" box="[845,1058,691,717]" italics="true" pageId="22" pageNumber="583">P. phetchabunensis</emphasis>
</taxonomicName>
<emphasis id="B961EAC5FFFC225D087F7890FB5E51B0" bold="true" box="[1064,1153,691,717]" pageId="22" pageNumber="583">
<taxonomicNameLabel id="A25257BEFFFC225D087F7890FB5E51B0" box="[1064,1153,691,717]" pageId="22" pageNumber="583" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but can be distinguished from it by the characters given in the differential diagnosis of the latter.
</paragraph>
</subSubSection>
<caption id="DF6A665FFFFC225D0CC07CC9FDC154E9" ID-DOI="http://doi.org/10.5281/zenodo.5042686" ID-Zenodo-Dep="5042686" httpUri="https://zenodo.org/record/5042686/files/figure.png" pageId="22" pageNumber="583" startId="22.[151,260,1770,1796]" targetBox="[159,1430,816,1737]" targetPageId="22">
<paragraph id="8BAA36D7FFFC225D0CC07CC9FDC154E9" blockId="22.[151,1436,1770,1940]" pageId="22" pageNumber="583">
<emphasis id="B961EAC5FFFC225D0CC07CC9FEF25479" bold="true" box="[151,301,1770,1796]" pageId="22" pageNumber="583">FIGURE 11.</emphasis>
<taxonomicName id="4C154D54FFFC225D0D627CC8FE3C5479" baseAuthorityName="Ochs" baseAuthorityYear="1940" box="[309,483,1770,1796]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="583" phylum="Arthropoda" rank="species" species="salvazai">
<emphasis id="B961EAC5FFFC225D0D627CC8FE3C5479" box="[309,483,1770,1796]" italics="true" pageId="22" pageNumber="583">Patrus salvazai</emphasis>
</taxonomicName>
: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view.
</paragraph>
</caption>
<subSubSection id="C30F655CFFFD225C0C907AB5FE6F52FC" pageId="23" pageNumber="584" type="distribution">
<paragraph id="8BAA36D7FFFD225C0C907AB5FE6F52FC" blockId="23.[151,1437,150,871]" pageId="23" pageNumber="584">
<emphasis id="B961EAC5FFFD225C0C907AB5FEBE53CD" bold="true" box="[199,353,150,176]" pageId="23" pageNumber="584">Distribution.</emphasis>
This species was originally described from
<collectingRegion id="49D1F835FFFD225C0F3B7AB4FB5553CC" box="[876,1162,151,177]" country="Laos" name="Louangphabang" pageId="23" pageNumber="584">Luang Prabang Province</collectingRegion>
with several additional records from
<collectingCountry id="F3027647FFFD225C0D787A99FEBA53A9" box="[303,357,186,212]" name="Laos" pageId="23" pageNumber="584">Laos</collectingCountry>
(
<bibRefCitation id="EF844B26FFFD225C0D227A9AFDD053AE" author="Peschet, R." box="[373,527,185,211]" pageId="23" pageNumber="584" pagination="121 - 136" refId="ref22628" refString="Peschet, R. (1923) Fauna entomologique de l'Indochine Gyrinidae. Opuscules de l'Institut Scientifique de l'Indochine Saigon, 6, 121 - 136." type="journal article" year="1923">Peschet 1923</bibRefCitation>
;
<bibRefCitation id="EF844B26FFFD225C0E4C7A9AFD4753AE" author="Ochs" box="[539,664,185,211]" pageId="23" pageNumber="584" pagination="1 - 39" refId="ref22521" refString="Ochs, G. (1930) Gyrinoidea. Catalogue of Indian Insects, 19, 1 - 39." type="journal article" year="1930">Ochs 1930</bibRefCitation>
).
<bibRefCitation id="EF844B26FFFD225C0EFB7A9AFCE753AE" author="Ochs" box="[684,824,185,211]" pageId="23" pageNumber="584" pagination="1 - 39" refId="ref22521" refString="Ochs, G. (1930) Gyrinoidea. Catalogue of Indian Insects, 19, 1 - 39." type="journal article" year="1930">Ochs (1930)</bibRefCitation>
reported the first records of
<taxonomicName id="4C154D54FFFD225C082C7A99FB2B53AE" baseAuthorityName="Ochs" baseAuthorityYear="1940" box="[1147,1268,185,211]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="584" phylum="Arthropoda" rank="species" species="salvazai">
<emphasis id="B961EAC5FFFD225C082C7A99FB2B53AE" box="[1147,1268,185,211]" italics="true" pageId="23" pageNumber="584">P. salvazai</emphasis>
</taxonomicName>
from Doi Prae Muang mountain in
<collectingRegion id="49D1F835FFFD225C0DD57AFFFDA6538B" box="[386,633,220,246]" country="Thailand" name="Chiang Mai" pageId="23" pageNumber="584">Chiang Mai Province</collectingRegion>
,
<collectingCountry id="F3027647FFFD225C0ED17AFFFD32538B" box="[646,749,220,246]" name="Thailand" pageId="23" pageNumber="584">Thailand</collectingCountry>
. Here we add additional localities for
<taxonomicName id="4C154D54FFFD225C08FF7AFEFAFD538B" baseAuthorityName="Ochs" baseAuthorityYear="1940" box="[1192,1314,220,246]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="584" phylum="Arthropoda" rank="species" species="salvazai">
<emphasis id="B961EAC5FFFD225C08FF7AFEFAFD538B" box="[1192,1314,220,246]" italics="true" pageId="23" pageNumber="584">P. salvazai</emphasis>
</taxonomicName>
across the mountains of northern
<collectingCountry id="F3027647FFFD225C0DC97ADCFDDD5264" box="[414,514,255,281]" name="Thailand" pageId="23" pageNumber="584">Thailand</collectingCountry>
in
<collectingRegion id="49D1F835FFFD225C0E7C7ADCFD735264" box="[555,684,255,281]" country="Thailand" name="Chiang Rai" pageId="23" pageNumber="584">Chiang Rai</collectingRegion>
and
<collectingRegion id="49D1F835FFFD225C0EBF7ADCFCC85264" box="[744,791,255,281]" country="Thailand" name="Nan" pageId="23" pageNumber="584">Nan</collectingRegion>
Provinces (
<figureCitation id="132E2A52FFFD225C0FF67ADCFBD55264" box="[929,1034,255,281]" captionStart="FIGURE 14" captionStartId="26.[151,260,743,769]" captionTargetBox="[152,1436,179,719]" captionTargetId="figure-25@26.[151,1436,179,719]" captionTargetPageId="26" captionText="FIGURE 14. (AB) Distribution records of the P. landaisi species group in Thailand." figureDoi="http://doi.org/10.5281/zenodo.5042692" httpUri="https://zenodo.org/record/5042692/files/figure.png" pageId="23" pageNumber="584">Fig. 14B</figureCitation>
). In
<collectingCountry id="F3027647FFFD225C08167ADCFB785264" box="[1089,1191,255,281]" name="Thailand" pageId="23" pageNumber="584">Thailand</collectingCountry>
,
<taxonomicName id="4C154D54FFFD225C08E27ADCFAF05264" baseAuthorityName="Ochs" baseAuthorityYear="1940" box="[1205,1327,255,281]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="584" phylum="Arthropoda" rank="species" species="salvazai">
<emphasis id="B961EAC5FFFD225C08E27ADCFAF05264" box="[1205,1327,255,281]" italics="true" pageId="23" pageNumber="584">P. salvazai</emphasis>
</taxonomicName>
has been found coexisting with
<taxonomicName id="4C154D54FFFD225C0DC57B01FDD95246" baseAuthorityName="Regimbart" baseAuthorityYear="1891" box="[402,518,289,315]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="584" phylum="Arthropoda" rank="species" species="apicalis">
<emphasis id="B961EAC5FFFD225C0DC57B01FDD95246" box="[402,518,289,315]" italics="true" pageId="23" pageNumber="584">P. apicalis</emphasis>
</taxonomicName>
in streams at high elevations (
<figureCitation id="132E2A52FFFD225C0F0D7B02FC615246" box="[858,958,289,315]" captionStart="FIGURE 13" captionStartId="25.[151,260,1683,1709]" captionTargetBox="[169,1418,181,1656]" captionTargetId="figure-18@25.[169,1418,179,1659]" captionTargetPageId="25" captionText="FIGURE 13. Habitats of P. landaisi species group: (A) stream at Siriphum Waterfall, Doi Inthanon National Park, Chiang Mai Province, habitat of P. garuda sp. nov. and P. subapicalis; (B) Benjamart Suksai in habitat of P. apicalis and P. salvazai at Nam Kuen stream, Chiang Rai Province; (C) shallow stream close to Ban Thai Samakkhi, Chiang Rai Province, habitat of P. landaisi and P. salvazai; (D) Tham Yai Waterfall Phu Kradueng National Park, Loei Province, habitat of P. phetchabunensis sp. nov.; (E) puddle in dry stream bed close to Ban Phithak Thai, Chiang Rai Province, habitat of P. landaisi and P. salvazai; (F) P. landaisi and P. salvazai from puddle in dry stream habitat close to Ban Thai Samakkhi, Chiang Rai Province." figureDoi="http://doi.org/10.5281/zenodo.5042690" httpUri="https://zenodo.org/record/5042690/files/figure.png" pageId="23" pageNumber="584">Fig. 13B</figureCitation>
) as well as with
<taxonomicName id="4C154D54FFFD225C082F7B01FB315246" authorityName="Regimbart" authorityYear="1892" box="[1144,1262,289,315]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="584" phylum="Arthropoda" rank="species" species="landaisi">
<emphasis id="B961EAC5FFFD225C082F7B01FB315246" box="[1144,1262,289,315]" italics="true" pageId="23" pageNumber="584">P. landaisi</emphasis>
</taxonomicName>
from a shallow stream with stagnant water or with slow current (
<figureCitation id="132E2A52FFFD225C0EE57B67FCC85223" box="[690,791,324,350]" captionStart="FIGURE 13" captionStartId="25.[151,260,1683,1709]" captionTargetBox="[169,1418,181,1656]" captionTargetId="figure-18@25.[169,1418,179,1659]" captionTargetPageId="25" captionText="FIGURE 13. Habitats of P. landaisi species group: (A) stream at Siriphum Waterfall, Doi Inthanon National Park, Chiang Mai Province, habitat of P. garuda sp. nov. and P. subapicalis; (B) Benjamart Suksai in habitat of P. apicalis and P. salvazai at Nam Kuen stream, Chiang Rai Province; (C) shallow stream close to Ban Thai Samakkhi, Chiang Rai Province, habitat of P. landaisi and P. salvazai; (D) Tham Yai Waterfall Phu Kradueng National Park, Loei Province, habitat of P. phetchabunensis sp. nov.; (E) puddle in dry stream bed close to Ban Phithak Thai, Chiang Rai Province, habitat of P. landaisi and P. salvazai; (F) P. landaisi and P. salvazai from puddle in dry stream habitat close to Ban Thai Samakkhi, Chiang Rai Province." figureDoi="http://doi.org/10.5281/zenodo.5042690" httpUri="https://zenodo.org/record/5042690/files/figure.png" pageId="23" pageNumber="584">Fig. 13C</figureCitation>
). These two species were also found in puddles in a drying stream bed (
<figureCitation id="132E2A52FFFD225C0D767B44FE7B52FC" box="[289,420,359,385]" captionStart="FIGURE 13" captionStartId="25.[151,260,1683,1709]" captionTargetBox="[169,1418,181,1656]" captionTargetId="figure-18@25.[169,1418,179,1659]" captionTargetPageId="25" captionText="FIGURE 13. Habitats of P. landaisi species group: (A) stream at Siriphum Waterfall, Doi Inthanon National Park, Chiang Mai Province, habitat of P. garuda sp. nov. and P. subapicalis; (B) Benjamart Suksai in habitat of P. apicalis and P. salvazai at Nam Kuen stream, Chiang Rai Province; (C) shallow stream close to Ban Thai Samakkhi, Chiang Rai Province, habitat of P. landaisi and P. salvazai; (D) Tham Yai Waterfall Phu Kradueng National Park, Loei Province, habitat of P. phetchabunensis sp. nov.; (E) puddle in dry stream bed close to Ban Phithak Thai, Chiang Rai Province, habitat of P. landaisi and P. salvazai; (F) P. landaisi and P. salvazai from puddle in dry stream habitat close to Ban Thai Samakkhi, Chiang Rai Province." figureDoi="http://doi.org/10.5281/zenodo.5042690" httpUri="https://zenodo.org/record/5042690/files/figure.png" pageId="23" pageNumber="584">Fig. 13EF</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30F655CFFFD225C0C907BAAFE79501B" pageId="23" pageNumber="584" type="discussion">
<paragraph id="8BAA36D7FFFD225C0C907BAAFE79501B" blockId="23.[151,1437,150,871]" pageId="23" pageNumber="584">
<emphasis id="B961EAC5FFFD225C0C907BAAFE9352DE" bold="true" box="[199,332,393,419]" pageId="23" pageNumber="584">Discussion.</emphasis>
The name
<taxonomicName id="4C154D54FFFD225C0D927BAAFD6852DE" authorityName="Ochs" authorityYear="1940" box="[453,695,393,419]" class="Insecta" family="Gyrinidae" genus="Orectochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="584" phylum="Arthropoda" rank="species" species="salvazai">
<emphasis id="B961EAC5FFFD225C0D927BAAFD6852DE" box="[453,695,393,419]" italics="true" pageId="23" pageNumber="584">Orectochilus salvazai</emphasis>
</taxonomicName>
was first published by
<bibRefCitation id="EF844B26FFFD225C0FE17BAAFB9F52DE" author="Ochs" box="[950,1088,393,419]" pageId="23" pageNumber="584" pagination="1 - 39" refId="ref22521" refString="Ochs, G. (1930) Gyrinoidea. Catalogue of Indian Insects, 19, 1 - 39." type="journal article" year="1930">Ochs (1930)</bibRefCitation>
in his catalog of Indian insects, for specimens that
<bibRefCitation id="EF844B26FFFD225C0D207B8FFDFD52BB" author="Peschet, R." box="[375,546,428,454]" pageId="23" pageNumber="584" pagination="121 - 136" refId="ref22628" refString="Peschet, R. (1923) Fauna entomologique de l'Indochine Gyrinidae. Opuscules de l'Institut Scientifique de l'Indochine Saigon, 6, 121 - 136." type="journal article" year="1923">Peschet (1923)</bibRefCitation>
misidentified as
<taxonomicName id="4C154D54FFFD225C0EB97B8FFB2052BB" authority="(Regimbart, 1888)" baseAuthorityName="Regimbart" baseAuthorityYear="1888" box="[750,1279,428,454]" class="Insecta" family="Gyrinidae" genus="Orectochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="584" phylum="Arthropoda" rank="species" species="cardiophorus">
<emphasis id="B961EAC5FFFD225C0EB97B8FFBC152BB" box="[750,1054,428,454]" italics="true" pageId="23" pageNumber="584">Orectochilus cardiophorus</emphasis>
(
<bibRefCitation id="EF844B26FFFD225C08667B8FFB2952BB" author="Regimbart, M." box="[1073,1270,428,454]" pageId="23" pageNumber="584" pagination="609 - 623" refId="ref22960" refString="Regimbart, M. (1888) Viaggio di Leonardo Fea in Birmania e regioni vicine. X. Dytiscidae et Gyrinidae. Annali del Museo Civico di Storia Naturale di Genova, Serie 2 a, 6, 609 - 623." type="journal article" year="1888">Régimbart, 1888</bibRefCitation>
)
</taxonomicName>
. This renders
<taxonomicName id="4C154D54FFFD225C0CC07BECFDEE5294" authority="Ochs, 1930 a" authorityName="Ochs" authorityYear="1930" box="[151,561,463,489]" class="Insecta" family="Gyrinidae" genus="Orectochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="584" phylum="Arthropoda" rank="species" species="salvazai">
<emphasis id="B961EAC5FFFD225C0CC07BECFE525294" box="[151,397,463,489]" italics="true" pageId="23" pageNumber="584">Orectochilus salvazai</emphasis>
<bibRefCitation id="EF844B26FFFD225C0DC07BECFDC55294" author="Ochs" box="[407,538,463,489]" pageId="23" pageNumber="584" pagination="1 - 39" refId="ref22521" refString="Ochs, G. (1930) Gyrinoidea. Catalogue of Indian Insects, 19, 1 - 39." type="journal article" year="1930">Ochs, 1930</bibRefCitation>
a
</taxonomicName>
<emphasis id="B961EAC5FFFD225C0E6B7BECFD025294" box="[572,733,463,489]" italics="true" pageId="23" pageNumber="584">nomen nudum</emphasis>
as it was not accompanied by a description.
<bibRefCitation id="EF844B26FFFD225C08A77BECFAA65294" author="Ochs" box="[1264,1401,463,489]" pageId="23" pageNumber="584" pagination="1 - 22" refId="ref22563" refString="Ochs, G. (1940) Entomological results from the Swedish Expedition 1934 to Burma and British India. Coleoptera: Gyrinidae, gesammelt von Rene Malaise. Arkiv for Zoologi, 32 A, 1 - 22." type="journal article" year="1940">Ochs (1940</bibRefCitation>
, p. 19), explains later that he attributed the name
<taxonomicName id="4C154D54FFFD225C0EC87BD1FCC15176" authorityName="Ochs" authorityYear="1930" box="[671,798,497,524]" class="Insecta" family="Gyrinidae" genus="Orectochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="584" phylum="Arthropoda" rank="species" species="salvazai">
<emphasis id="B961EAC5FFFD225C0EC87BD1FCC15176" box="[671,798,497,524]" italics="true" pageId="23" pageNumber="584">O. salvazai</emphasis>
</taxonomicName>
to Peschet in both his prior checklist (1930) and present work (1940) because, subsequent to the aforementioned misidentification (
<bibRefCitation id="EF844B26FFFD225C0F887837FB5F5153" author="Peschet, R." box="[991,1152,532,558]" pageId="23" pageNumber="584" pagination="121 - 136" refId="ref22628" refString="Peschet, R. (1923) Fauna entomologique de l'Indochine Gyrinidae. Opuscules de l'Institut Scientifique de l'Indochine Saigon, 6, 121 - 136." type="journal article" year="1923">Peschet, 1923</bibRefCitation>
), Peschet had recognized that these specimens actually belonged to a new species he dubbed “
<taxonomicName id="4C154D54FFFD225C0FF07814FBDC512C" authorityName="Ochs" authorityYear="1930" box="[935,1027,567,593]" class="Insecta" family="Gyrinidae" genus="Orectochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="584" phylum="Arthropoda" rank="species" species="salvazai">
<emphasis id="B961EAC5FFFD225C0FF07814FBDC512C" box="[935,1027,567,593]" italics="true" pageId="23" pageNumber="584">salvazai</emphasis>
</taxonomicName>
” and had provided Ochs specimens identified as such. Because Peschet had not previously published a description of
<emphasis id="B961EAC5FFFD225C08167879FB1E510E" box="[1089,1217,601,627]" italics="true" pageId="23" pageNumber="584">
<taxonomicName id="4C154D54FFFD225C08167879FB62510E" authorityName="Ochs" authorityYear="1930" box="[1089,1213,601,627]" class="Insecta" family="Gyrinidae" genus="Orectochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="584" phylum="Arthropoda" rank="species" species="salvazai">P. salvazai</taxonomicName>
,
</emphasis>
when
<bibRefCitation id="EF844B26FFFD225C0958787AFA43510E" author="Ochs" box="[1295,1436,601,627]" pageId="23" pageNumber="584" pagination="1 - 22" refId="ref22563" refString="Ochs, G. (1940) Entomological results from the Swedish Expedition 1934 to Burma and British India. Coleoptera: Gyrinidae, gesammelt von Rene Malaise. Arkiv for Zoologi, 32 A, 1 - 22." type="journal article" year="1940">Ochs (1940)</bibRefCitation>
used the name
<taxonomicName id="4C154D54FFFD225C0D12785FFDE651EB" authorityName="Ochs" authorityYear="1930" box="[325,569,636,662]" class="Insecta" family="Gyrinidae" genus="Orectochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="584" phylum="Arthropoda" rank="species" species="salvazai">
<emphasis id="B961EAC5FFFD225C0D12785FFDE651EB" box="[325,569,636,662]" italics="true" pageId="23" pageNumber="584">Orectochilus salvazai</emphasis>
</taxonomicName>
and included both a description of the species, a differential diagnosis, and a mention of examined material, the name was made available and is attributable to him following Article 13.1 of
<emphasis id="B961EAC5FFFD225C0CC078E2FD3551A6" box="[151,746,705,731]" italics="true" pageId="23" pageNumber="584">The International Code of Zoological Nomenclature</emphasis>
(
<bibRefCitation id="EF844B26FFFD225C0EAE78E2FC5D51A6" author="ICZN" box="[761,898,705,731]" pageId="23" pageNumber="584" refId="ref21889" refString="ICZN (1999) International Code of Zoological Nomenclature, Fourth Edition, adopted by the International Union of Biological Sciences. International Trust for Zoological Nomenclature, London, 106 pp." type="book" year="1999">ICZN 1999</bibRefCitation>
), not Peschet. A similar situation surrounds the name
<taxonomicName id="4C154D54FFFD225C0C8C78C7FDA35183" authority="(Ochs, 1940)" baseAuthorityName="Ochs" baseAuthorityYear="1940" box="[219,636,740,766]" class="Insecta" family="Gyrinidae" genus="Orectochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="584" phylum="Arthropoda" rank="species" species="peguensis">
<emphasis id="B961EAC5FFFD225C0C8C78C7FE3F5183" box="[219,480,740,766]" italics="true" pageId="23" pageNumber="584">Orectochilus peguensis</emphasis>
(
<bibRefCitation id="EF844B26FFFD225C0DA778C7FDAA5183" author="Ochs" box="[496,629,740,766]" pageId="23" pageNumber="584" pagination="1 - 22" refId="ref22563" refString="Ochs, G. (1940) Entomological results from the Swedish Expedition 1934 to Burma and British India. Coleoptera: Gyrinidae, gesammelt von Rene Malaise. Arkiv for Zoologi, 32 A, 1 - 22." type="journal article" year="1940">Ochs, 1940</bibRefCitation>
)
</taxonomicName>
, another member of the
<taxonomicName id="4C154D54FFFD225C0FD978C6FBDB5183" authorityName="Regimbart" authorityYear="1892" box="[910,1028,740,766]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="584" phylum="Arthropoda" rank="species" species="landaisi">
<emphasis id="B961EAC5FFFD225C0FD978C6FBDB5183" box="[910,1028,740,766]" italics="true" pageId="23" pageNumber="584">P. landaisi</emphasis>
</taxonomicName>
group (see checklist here below), in that its first use is in Ochs 1930 catalog as
<taxonomicName id="4C154D54FFFD225C0E297924FC73505D" box="[638,940,775,801]" class="Insecta" family="Gyrinidae" genus="Orectochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="584" phylum="Arthropoda" rank="subSpecies" species="cardiophorus" subSpecies="peguensis">
<emphasis id="B961EAC5FFFD225C0E297924FC73505D" box="[638,940,775,801]" italics="true" pageId="23" pageNumber="584">O. cardiophorus peguensis</emphasis>
</taxonomicName>
(a
<emphasis id="B961EAC5FFFD225C0F877924FBB1505C" box="[976,1134,775,801]" italics="true" pageId="23" pageNumber="584">nomen nudum</emphasis>
) attributed to Peschet with the name being made available subsequently by the description given in
<bibRefCitation id="EF844B26FFFD225C0F97790AFB9B503E" author="Ochs" box="[960,1092,809,835]" pageId="23" pageNumber="584" pagination="1 - 22" refId="ref22563" refString="Ochs, G. (1940) Entomological results from the Swedish Expedition 1934 to Burma and British India. Coleoptera: Gyrinidae, gesammelt von Rene Malaise. Arkiv for Zoologi, 32 A, 1 - 22." type="journal article" year="1940">Ochs (1940</bibRefCitation>
, p. 17) and is thus attributed to
<bibRefCitation id="EF844B26FFFD225C0CC0796FFECC501B" author="Ochs" box="[151,275,844,870]" pageId="23" pageNumber="584" pagination="1 - 22" refId="ref22563" refString="Ochs, G. (1940) Entomological results from the Swedish Expedition 1934 to Burma and British India. Coleoptera: Gyrinidae, gesammelt von Rene Malaise. Arkiv for Zoologi, 32 A, 1 - 22." type="journal article" year="1940">Ochs 1940</bibRefCitation>
, not Peschet.
</paragraph>
</subSubSection>
</treatment>
</document>