361 lines
50 KiB
XML
361 lines
50 KiB
XML
<document id="24F0B5D714B0F6F76D1D89A0FA173623" ID-DOI="10.11646/zootaxa.4991.3.8" ID-ISSN="1175-5326" ID-Zenodo-Dep="5042656" ID-ZooBank="F0FAE07E-4A90-4602-BCC5-6D80E158BB6C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624989159860" checkinUser="plazi" docAuthor="Suksai, Benjamart, Gustafson, Grey T., Sites, Robert W. & Sangpradub, Narumon" docDate="2021" docId="03BC87C1FFE322420CC07AB4FB2A54D4" docLanguage="en" docName="zootaxa.4991.3.8.pdf" docOrigin="Zootaxa 4991 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Patrus corpulentus Borneo" docType="treatment" docVersion="3" lastPageNumber="570" masterDocId="FF85FFB9FFEA224B0C577A23FFDF537D" masterDocTitle="Review of the Patrus landaisi species group from Thailand with description of three new species (Coleoptera, Gyrinidae)" masterLastPageNumber="591" masterPageNumber="561" pageNumber="570" updateTime="1698978743576" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="8845B01C9A81F8C9E8A9A474A607914C">Review of the Patrus landaisi species group from Thailand with description of three new species (Coleoptera, Gyrinidae)</mods:title>
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<mods:namePart id="729BEF008D669A9747232FD390AF2529">Suksai, Benjamart</mods:namePart>
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<mods:nameIdentifier id="32B57F15398D49BFF8B6BD85874C388B" type="email">benjamartbenjamart@gmail.com</mods:nameIdentifier>
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<mods:namePart id="9984954E30575455B871B7393BA0CE7F">Gustafson, Grey T.</mods:namePart>
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<mods:affiliation id="2B59DDCAD6BFB769A9DBE2808895E150">Department of Ecology and Evolutionary Biology, University of Kansas, Lawrence, Kansas 66045, USA. & Department of Biological Sciences, Northern Arizona University, Flagstaff, Arizona, 86011, USA.</mods:affiliation>
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<mods:namePart id="CD7F40D271B5B79CC18798F7E8735947">Sites, Robert W.</mods:namePart>
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<mods:affiliation id="38EFCA41A5B067E3B715596190B2E7F6">Enns Entomology Museum, Division of Plant Science, University of Missouri-Columbia, Missouri 65211, USA. SitesR @ missouri. edu; https: // orcid. org / 0000 - 0002 - 3895 - 813 X</mods:affiliation>
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<mods:namePart id="5A4DD8AB3958CE9B43F9F535AD63E09F">Sangpradub, Narumon</mods:namePart>
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<mods:affiliation id="F0B51A9F34D90F72FA8D18C1F4929F93">Applied Taxonomic Research Center, Faculty of Science, Khon Kaen University, Khon Kaen 40002, Thailand & Corresponding Author: narumon @ kku. ac. th; https: // orcid. org / 0000 - 0003 - 0228 - 8696</mods:affiliation>
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<mods:nameIdentifier id="65E4AB41767756A20827A4F5347E3332" type="email">benjamartbenjamart@gmail.com</mods:nameIdentifier>
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<mods:title id="73857C3D8C3D95713331ECC48A054707">Zootaxa</mods:title>
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<mods:part id="F2F8F57A6127E548FAA4C6514AFECD95">
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<mods:date id="3E9508EC906729445949DE7D686DE43E">2021</mods:date>
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<mods:number id="3C0E284226BC4EDE2E373C6D9119B0B1">2021-06-25</mods:number>
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<mods:number id="1FC49A29C6F0DF015AA79C2346C813A7">4991</mods:number>
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<taxonomicName id="4C154D54FFE322420CC07AB4FDB653CC" authority="(Regimbart, 1884)" authorityName="Borneo" baseAuthorityName="Regimbart" baseAuthorityYear="1884" box="[151,617,150,178]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="570" phylum="Arthropoda" rank="species" species="corpulentus">
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<emphasis id="B961EAC5FFE322420CC07AB4FEA253CF" bold="true" box="[151,381,151,178]" italics="true" pageId="9" pageNumber="570">Patrus corpulentus</emphasis>
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(
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<bibRefCitation id="EF844B26FFE322420DDA7AB5FDBE53CC" author="Regimbart, M." box="[397,609,150,177]" pageId="9" pageNumber="570" pagination="381 - 482" refId="ref22884" refString="Regimbart, M. (1884) Essai monographique de la famille des Gyrinidae. 3 e partie. Annales de la Societe entomologique de France, 6, 381 - 482, pls. 311 - 314." type="journal article" year="1884">Régimbart, 1884</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</paragraph>
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<subSubSection id="C30F655CFFE322420CC07A98FE1F53A8" box="[151,448,187,213]" pageId="9" pageNumber="570" type="description">
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<paragraph id="8BAA36D7FFE322420CC07A98FE1F53A8" blockId="9.[151,617,150,213]" box="[151,448,187,213]" pageId="9" pageNumber="570">
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(
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<figureCitation id="132E2A52FFE322420CC87A98FEDD53A8" box="[159,258,187,213]" captionStart="FIGURE 2" captionStartId="6.[151,260,1861,1887]" captionTargetBox="[151,1434,179,1836]" captionTargetId="figure-25@6.[151,1435,179,1836]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042660" httpUri="https://zenodo.org/record/5042660/files/figure.png" pageId="9" pageNumber="570">Figs. 2B</figureCitation>
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,
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<figureCitation id="132E2A52FFE322420D5A7A98FEEF53A8" box="[269,304,187,213]" captionStart="FIGURE 3" captionStartId="7.[151,260,1865,1891]" captionTargetBox="[151,1434,182,1841]" captionTargetId="figure-18@7.[151,1435,179,1841]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal habitus of females of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042662" httpUri="https://zenodo.org/record/5042662/files/figure.png" pageId="9" pageNumber="570">3B</figureCitation>
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,
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<figureCitation id="132E2A52FFE322420D6B7A98FE8053A8" box="[316,351,187,213]" captionStart="FIGURE 4" captionStartId="8.[151,260,707,733]" captionTargetBox="[158,1432,185,677]" captionTargetId="figure-25@8.[152,1436,179,683]" captionTargetPageId="8" captionText="FIGURE 4. Left protibia and protarsus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5042666" httpUri="https://zenodo.org/record/5042666/files/figure.png" pageId="9" pageNumber="570">4B</figureCitation>
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,
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<figureCitation id="132E2A52FFE322420D3C7A98FEA453A8" box="[363,379,187,213]" captionStart="FIGURE 6" captionStartId="10.[151,260,1117,1143]" captionTargetBox="[180,1401,191,1085]" captionTargetId="figure-25@10.[176,1410,179,1093]" captionTargetPageId="10" captionText="FIGURE 6. Patrus corpulentus: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) female pronotum and elytra, dorsal view, scale bar = 4 mm; (D) gonocoxae, ventral view; (E) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (F) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042674" httpUri="https://zenodo.org/record/5042674/files/figure.png" pageId="9" pageNumber="570">6</figureCitation>
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,
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<figureCitation id="132E2A52FFE322420DD07A98FE6753A8" box="[391,440,187,213]" captionStart="FIGURE 14" captionStartId="26.[151,260,743,769]" captionTargetBox="[152,1436,179,719]" captionTargetId="figure-25@26.[151,1436,179,719]" captionTargetPageId="26" captionText="FIGURE 14. (A–B) Distribution records of the P. landaisi species group in Thailand." figureDoi="http://doi.org/10.5281/zenodo.5042692" httpUri="https://zenodo.org/record/5042692/files/figure.png" pageId="9" pageNumber="570">14A</figureCitation>
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)
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</paragraph>
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<subSubSection id="C30F655CFFE322420CC07B23FC865228" pageId="9" pageNumber="570" type="reference_group">
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<paragraph id="8BAA36D7FFE322420CC07B23FC885265" blockId="9.[151,1031,256,341]" box="[151,855,256,280]" pageId="9" pageNumber="570">
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<treatmentCitationGroup id="AB0511F9FFE322420CC07B23FC885265" box="[151,855,256,280]" pageId="9" pageNumber="570">
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<treatmentCitation id="0AB410C6FFE322420CC07B23FDA55265" author="Regimbart, M." box="[151,634,256,280]" page="411" pageId="9" pageNumber="570" year="1884">
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<taxonomicName id="4C154D54FFE322420CC07B23FDA55265" authority="Regimbart, 1884: 411" authorityName="Regimbart" authorityPageNumber="411" authorityYear="1884" box="[151,634,256,280]" class="Insecta" family="Gyrinidae" genus="Orectochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="570" phylum="Arthropoda" rank="species" species="corpulentus">
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<emphasis id="B961EAC5FFE322420CC07B23FE485265" box="[151,407,256,280]" italics="true" pageId="9" pageNumber="570">Orectochilus corpulentus</emphasis>
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<bibRefCitation id="EF844B26FFE322420DCA7B23FDA55265" author="Regimbart, M." box="[413,634,256,280]" pageId="9" pageNumber="570" pagination="381 - 482" refId="ref22884" refString="Regimbart, M. (1884) Essai monographique de la famille des Gyrinidae. 3 e partie. Annales de la Societe entomologique de France, 6, 381 - 482, pls. 311 - 314." type="journal article" year="1884">Régimbart, 1884: 411</bibRefCitation>
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(original description)
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<paragraph id="8BAA36D7FFE322420CC07B3DFBD8524B" blockId="9.[151,1031,256,341]" box="[151,1031,286,310]" pageId="9" pageNumber="570">
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<taxonomicName id="4C154D54FFE322420CC07B3DFD94524B" authority="Regimbart, 1884" authorityName="Regimbart" authorityYear="1884" box="[151,587,286,310]" class="Insecta" family="Gyrinidae" genus="Orectochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="570" phylum="Arthropoda" rank="species" species="corpulentus">
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<emphasis id="B961EAC5FFE322420CC07B3DFE48524B" box="[151,407,286,310]" italics="true" pageId="9" pageNumber="570">Orectochilus corpulentus</emphasis>
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<bibRefCitation id="EF844B26FFE322420DCA7B3DFD94524B" author="Regimbart, M." box="[413,587,286,310]" pageId="9" pageNumber="570" pagination="381 - 482" refId="ref22884" refString="Regimbart, M. (1884) Essai monographique de la famille des Gyrinidae. 3 e partie. Annales de la Societe entomologique de France, 6, 381 - 482, pls. 311 - 314." type="journal article" year="1884">Régimbart, 1884</bibRefCitation>
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</taxonomicName>
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:
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<treatmentCitation id="0AB410C6FFE322420E027B3DFCF3524B" author="Regimbart, M." box="[597,812,286,310]" page="619" pageId="9" pageNumber="570" year="1888">
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<bibRefCitation id="EF844B26FFE322420E027B3DFCF3524B" author="Regimbart, M." box="[597,812,286,310]" pageId="9" pageNumber="570" pagination="609 - 623" refId="ref22960" refString="Regimbart, M. (1888) Viaggio di Leonardo Fea in Birmania e regioni vicine. X. Dytiscidae et Gyrinidae. Annali del Museo Civico di Storia Naturale di Genova, Serie 2 a, 6, 609 - 623." type="journal article" year="1888">Régimbart 1888: 619</bibRefCitation>
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(description of male)
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<paragraph id="8BAA36D7FFE322420CC07B1EFC865228" blockId="9.[151,1031,256,341]" box="[151,857,317,341]" pageId="9" pageNumber="570">
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<treatmentCitationGroup id="AB0511F9FFE322420CC07B1EFC865228" box="[151,857,317,341]" pageId="9" pageNumber="570">
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<taxonomicName id="4C154D54FFE322420CC07B1EFD795228" authority="Regimbart, 1884" authorityName="Regimbart" authorityYear="1884" box="[151,678,317,341]" class="Insecta" family="Gyrinidae" genus="Orectochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="570" phylum="Arthropoda" rank="species" species="corpulentus" subGenus="Patrus">
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<emphasis id="B961EAC5FFE322420CC07B1EFEC65228" box="[151,281,317,341]" italics="true" pageId="9" pageNumber="570">Orectochilus</emphasis>
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(
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<emphasis id="B961EAC5FFE322420D7F7B1DFEB35228" box="[296,364,318,341]" italics="true" pageId="9" pageNumber="570">Patrus</emphasis>
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)
|
||
<emphasis id="B961EAC5FFE322420D2D7B1EFE2D5228" box="[378,498,317,341]" italics="true" pageId="9" pageNumber="570">corpulentus</emphasis>
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<bibRefCitation id="EF844B26FFE322420DAF7B1EFD795228" author="Regimbart, M." box="[504,678,317,341]" pageId="9" pageNumber="570" pagination="381 - 482" refId="ref22884" refString="Regimbart, M. (1884) Essai monographique de la famille des Gyrinidae. 3 e partie. Annales de la Societe entomologique de France, 6, 381 - 482, pls. 311 - 314." type="journal article" year="1884">Régimbart, 1884</bibRefCitation>
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</taxonomicName>
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:
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<treatmentCitation id="0AB410C6FFE322420EE77B1EFC865228" author="Hatch, M. H." box="[688,857,317,341]" page="453" pageId="9" pageNumber="570" year="1926">
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<bibRefCitation id="EF844B26FFE322420EE77B1EFC865228" author="Hatch, M. H." box="[688,857,317,341]" pageId="9" pageNumber="570" pagination="429 - 467" refId="ref21854" refString="Hatch, M. H. (1926) The phylogeny and phylogenetic tendencies of Gyrinidae. Papers of the Michigan Academy of Science, Arts and Letters, 5, 429 - 467." type="journal article" year="1926">Hatch 1926: 453</bibRefCitation>
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</treatmentCitation>
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</treatmentCitationGroup>
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</paragraph>
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<typeStatus id="54AE8875FFE322420CC07BA0FF0D52E0" box="[151,210,387,413]" pageId="9" pageNumber="570">Type</typeStatus>
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localities.
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</emphasis>
|
||
<collectingRegion id="49D1F835FFE322420D187BA7FE5E52E3" box="[335,385,388,414]" country="Indonesia" name="Jawa Timur" pageId="9" pageNumber="570">Java</collectingRegion>
|
||
and
|
||
<location id="8ECA600CFFE322420DE07BA7FDD652E3" LSID="urn:lsid:plazi:treatment:03BC87C1FFE322420CC07AB4FB2A54D4:8ECA600CFFE322420DE07BA7FDD652E3" box="[439,521,388,414]" name="Borneo" pageId="9" pageNumber="570" stateProvince="Jawa Timur">Borneo</location>
|
||
[both likely in error]
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BAA36D7FFE322420C907B84FDBE5108" blockId="9.[151,1437,386,1962]" pageId="9" pageNumber="570">
|
||
<emphasis id="B961EAC5FFE322420C907B84FE3152BC" bold="true" box="[199,494,423,449]" pageId="9" pageNumber="570">Type material examined.</emphasis>
|
||
<typeStatus id="54AE8875FFE322420DA37B84FDAD52BC" box="[500,626,423,449]" pageId="9" pageNumber="570" type="syntype">SYNTYPE</typeStatus>
|
||
female ‘
|
||
<collectingRegion id="49D1F835FFE322420E877B84FCD852BC" box="[720,775,423,449]" country="Indonesia" name="Jawa Timur" pageId="9" pageNumber="570">Java</collectingRegion>
|
||
[yellow label, black border, handwritten black ink]’ ‘Badiu [white label, black border, handwritten black ink]’ ‘Corpulentus / Regb. Type [white label, handwritten black ink]’ ‘
|
||
<superScript id="7C609B9FFFE322420CCB7BCCFF655280" attach="left" box="[156,186,495,521]" fontSize="6" pageId="9" pageNumber="570">Dr</superScript>
|
||
Régimb. / vidit 1883. [white label, typed black ink]’ ‘Ex COLL. / E. WEHNCKE [white label, typed black ink]’ ‘
|
||
<taxonomicName id="4C154D54FFE322420CCC7830FEA95150" authority="Regbt." authorityName="Regimbart" authorityYear="1884" box="[155,374,531,557]" class="Insecta" family="Gyrinidae" genus="Orectochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="570" phylum="Arthropoda" rank="species" species="corpulentus">corpulentus Régbt.</taxonomicName>
|
||
/ java type! [being label, yellow border, handwritten black ink]’ ‘
|
||
<typeStatus id="54AE8875FFE32242080F7830FB045150" box="[1112,1243,531,557]" pageId="9" pageNumber="570" type="syntype">SYNTYPE</typeStatus>
|
||
[red label, typed black ink]’ ‘
|
||
<typeStatus id="54AE8875FFE322420D487814FE7D512C" box="[287,418,567,593]" pageId="9" pageNumber="570" type="syntype">SYNTYPE</typeStatus>
|
||
/
|
||
<emphasis id="B961EAC5FFE322420DEB7814FD83512F" box="[444,604,567,594]" italics="true" pageId="9" pageNumber="570">
|
||
<taxonomicName id="4C154D54FFE322420DEB7814FD93512C" authorityName="Dejean" authorityYear="1833" box="[444,588,567,593]" class="Insecta" family="Gyrinidae" genus="Orectochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="570" phylum="Arthropoda" rank="genus">Orectochilus</taxonomicName>
|
||
/
|
||
</emphasis>
|
||
<taxonomicName id="4C154D54FFE322420E327814FCB3512C" authority="Regimbart" authorityName="Regimbart" authorityYear="1884" box="[613,876,567,593]" class="Insecta" family="Gyrinidae" genus="Orectochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="570" phylum="Arthropoda" rank="species" species="corpulentus">
|
||
<emphasis id="B961EAC5FFE322420E327814FD36512C" box="[613,745,567,593]" italics="true" pageId="9" pageNumber="570">corpulentus</emphasis>
|
||
Régimbart
|
||
</taxonomicName>
|
||
, / 1884 [white label, typed black ink]’ ‘MNHN / EC10141 [white label, typed black ink]’.
|
||
</paragraph>
|
||
<paragraph id="8BAA36D7FFE322420C90785CFB6C519F" blockId="9.[151,1437,386,1962]" pageId="9" pageNumber="570">
|
||
<emphasis id="B961EAC5FFE322420C90785CFDDB51E4" bold="true" box="[199,516,639,665]" pageId="9" pageNumber="570">Other material examined.</emphasis>
|
||
<materialsCitation id="3B7D3C8AFFE322420E59785CFB3151E4" box="[526,1262,639,665]" collectionCode="NMPC" country="Myanmar" location="Tenasserim region" pageId="9" pageNumber="570" specimenCount="1" specimenCount-male="1">
|
||
<emphasis id="B961EAC5FFE322420E59785CFD7051E4" bold="true" box="[526,687,639,665]" pageId="9" pageNumber="570">
|
||
<collectingCountry id="F3027647FFE322420E59785CFD7551E4" box="[526,682,639,665]" name="Myanmar" pageId="9" pageNumber="570">MYANMAR</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<location id="8ECA600CFFE322420EED785CFC5151E4" LSID="urn:lsid:plazi:treatment:03BC87C1FFE322420CC07AB4FB2A54D4:8ECA600CFFE322420EED785CFC5151E4" box="[698,910,639,665]" country="Myanmar" name="Tenasserim region" pageId="9" pageNumber="570">Tenasserim region</location>
|
||
, 1838–1839,
|
||
<specimenCount id="9D13FD5EFFE322420879785CFBA251E4" box="[1070,1149,639,665]" pageId="9" pageNumber="570" type="male">1 male</specimenCount>
|
||
(
|
||
<collectionCode id="ED04AE12FFE3224208C7785CFB3951E4" box="[1168,1254,639,665]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="9" pageNumber="570">NMPC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B7D3C8AFFE3224208AB785CFB70519F" collectingDate="2005-05-25" collectionCode="UMC, THNHM" collectorName="A. Vitheepradit & Vitheepradit, R. W & T. Prommi" country="Thailand" elevation="287" latitude="8.883223" location="Mae Yai Waterfall" longLatPrecision="1" longitude="98.49955" pageId="9" pageNumber="570" specimenCount="2" specimenCount-female="2" stateProvince="Surat Thani">
|
||
<emphasis id="B961EAC5FFE3224208AB785CFE7151C0" bold="true" pageId="9" pageNumber="570">
|
||
<collectingCountry id="F3027647FFE3224208AB785CFA4851E4" box="[1276,1431,639,665]" name="Thailand" pageId="9" pageNumber="570">THAILAND</collectingCountry>
|
||
:
|
||
<collectingRegion id="49D1F835FFE322420CC07880FE7651C0" box="[151,425,675,701]" country="Thailand" name="Surat Thani" pageId="9" pageNumber="570">Surat Thani Province</collectingRegion>
|
||
:
|
||
</emphasis>
|
||
Khao Sok Natl. Park.,
|
||
<location id="8ECA600CFFE322420E877887FC7451C0" LSID="urn:lsid:plazi:treatment:03BC87C1FFE322420CC07AB4FB2A54D4:8ECA600CFFE322420E877887FC7451C0" box="[720,939,675,702]" country="Thailand" latitude="8.883223" longLatPrecision="1" longitude="98.49955" name="Mae Yai Waterfall" pageId="9" pageNumber="570" stateProvince="Surat Thani">Mae Yai Waterfall</location>
|
||
,
|
||
<geoCoordinate id="EE215010FFE322420FEC7880FB9451C3" box="[955,1099,675,702]" degrees="8" direction="north" minutes="52" orientation="latitude" pageId="9" pageNumber="570" precision="1" seconds="59.6" value="8.883223">8°52’59.6”N</geoCoordinate>
|
||
<geoCoordinate id="EE215010FFE32242080F7880FB2951C3" box="[1112,1270,675,702]" degrees="98" direction="east" minutes="29" orientation="longitude" pageId="9" pageNumber="570" precision="1" seconds="58.4" value="98.49955">98°29’58.4”E</geoCoordinate>
|
||
,
|
||
<elevation id="0038D1E4FFE3224209507880FA4851C0" box="[1287,1431,675,702]" metricMagnitude="2" metricUnit="m" metricValue="2.87" pageId="9" pageNumber="570" unit="m" value="287.0">
|
||
<quantity id="4CED9B32FFE3224209507880FA8B51C3" box="[1287,1364,675,702]" metricMagnitude="2" metricUnit="m" metricValue="2.87" pageId="9" pageNumber="570" unit="m" value="287.0">287 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<date id="FFAB1017FFE322420CC078E4FED3519F" box="[151,268,711,738]" pageId="9" pageNumber="570" value="2005-05-25">
|
||
<collectingDate id="EFEFE9FFFFE322420CC078E4FED3519F" box="[151,268,711,738]" pageId="9" pageNumber="570" value="2005-05-25">25.V.2005</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26E05301FFE322420D1178E4FE2A519C" box="[326,501,711,737]" pageId="9" pageNumber="570">A. Vitheepradit</collectorName>
|
||
, R.W. Sites &
|
||
<collectorName id="26E05301FFE322420ECD78EBFCCB519C" box="[666,788,711,738]" pageId="9" pageNumber="570">T. Prommi</collectorName>
|
||
, L-797,
|
||
<specimenCount id="9D13FD5EFFE322420F2678E4FC02519C" box="[881,989,711,737]" pageId="9" pageNumber="570" type="female">2 females</specimenCount>
|
||
(
|
||
<collectionCode id="ED04AE12FFE322420FBB78E4FBEF519C" box="[1004,1072,711,737]" pageId="9" pageNumber="570">UMC</collectionCode>
|
||
,
|
||
<collectionCode id="ED04AE12FFE32242086C78EBFB78519F" box="[1083,1191,712,738]" pageId="9" pageNumber="570">THNHM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30F655CFFE322420C9078C8FEEB5614" pageId="9" pageNumber="570" type="diagnosis">
|
||
<paragraph id="8BAA36D7FFE322420C9078C8FEF75710" blockId="9.[151,1437,386,1962]" pageId="9" pageNumber="570">
|
||
<emphasis id="B961EAC5FFE322420C9078C8FE9D5078" bold="true" box="[199,322,747,773]" pageId="9" pageNumber="570">Diagnosis.</emphasis>
|
||
Size: Male: 12.0 mm; females: 9.0–
|
||
<quantity id="4CED9B32FFE322420E9D78C8FCC0507B" box="[714,799,747,774]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="9" pageNumber="570" unit="mm" value="9.5">9.5 mm</quantity>
|
||
; body form in lateral view strongly dorsoventrally convex, evenly weakly depressed anteriorly and posteriorly; elytral glabrous region of male not cordiform (
|
||
<figureCitation id="132E2A52FFE3224208AD792CFA825054" box="[1274,1373,783,809]" captionStart="FIGURE 2" captionStartId="6.[151,260,1861,1887]" captionTargetBox="[151,1434,179,1836]" captionTargetId="figure-25@6.[151,1435,179,1836]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042660" httpUri="https://zenodo.org/record/5042660/files/figure.png" pageId="9" pageNumber="570">Figs. 2B</figureCitation>
|
||
,
|
||
<figureCitation id="132E2A52FFE32242093D792CFA505057" box="[1386,1423,783,810]" captionStart="FIGURE 6" captionStartId="10.[151,260,1117,1143]" captionTargetBox="[180,1401,191,1085]" captionTargetId="figure-25@10.[176,1410,179,1093]" captionTargetPageId="10" captionText="FIGURE 6. Patrus corpulentus: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) female pronotum and elytra, dorsal view, scale bar = 4 mm; (D) gonocoxae, ventral view; (E) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (F) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042674" httpUri="https://zenodo.org/record/5042674/files/figure.png" pageId="9" pageNumber="570">6A</figureCitation>
|
||
), along suture extending ca. 6/7 length of elytra, lateral margins rounded, being expanded after humeral region to just wider than pronotal glabrous region then attenuated apically; female elytral glabrous region (
|
||
<figureCitation id="132E2A52FFE3224208CF7974FB26500C" box="[1176,1273,855,881]" captionStart="FIGURE 3" captionStartId="7.[151,260,1865,1891]" captionTargetBox="[151,1434,182,1841]" captionTargetId="figure-18@7.[151,1435,179,1841]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal habitus of females of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042662" httpUri="https://zenodo.org/record/5042662/files/figure.png" pageId="9" pageNumber="570">Figs. 3B</figureCitation>
|
||
,
|
||
<figureCitation id="132E2A52FFE3224209547974FAFA500F" box="[1283,1317,855,882]" captionStart="FIGURE 6" captionStartId="10.[151,260,1117,1143]" captionTargetBox="[180,1401,191,1085]" captionTargetId="figure-25@10.[176,1410,179,1093]" captionTargetPageId="10" captionText="FIGURE 6. Patrus corpulentus: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) female pronotum and elytra, dorsal view, scale bar = 4 mm; (D) gonocoxae, ventral view; (E) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (F) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042674" httpUri="https://zenodo.org/record/5042674/files/figure.png" pageId="9" pageNumber="570">6C</figureCitation>
|
||
) similar to male; elytral apex weakly rounded (female) or narrowly rounded (male) with apicolateral sinuation, epipleural angle distinct and rounded; male protibia club-shaped (
|
||
<figureCitation id="132E2A52FFE322420E9E79BCFCFD50C4" box="[713,802,927,953]" captionStart="FIGURE 4" captionStartId="8.[151,260,707,733]" captionTargetBox="[158,1432,185,677]" captionTargetId="figure-25@8.[152,1436,179,683]" captionTargetPageId="8" captionText="FIGURE 4. Left protibia and protarsus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5042666" httpUri="https://zenodo.org/record/5042666/files/figure.png" pageId="9" pageNumber="570">Fig. 4B</figureCitation>
|
||
) strongly curved basally, without constriction, protibial apex truncate with prominent distolateral angle; male protarsus ca. 1/2 of protibia (
|
||
<figureCitation id="132E2A52FFE32242081079E0FB7F50A0" box="[1095,1184,963,989]" captionStart="FIGURE 4" captionStartId="8.[151,260,707,733]" captionTargetBox="[158,1432,185,677]" captionTargetId="figure-25@8.[152,1436,179,683]" captionTargetPageId="8" captionText="FIGURE 4. Left protibia and protarsus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5042666" httpUri="https://zenodo.org/record/5042666/files/figure.png" pageId="9" pageNumber="570">Fig. 4B</figureCitation>
|
||
), protarsomeres width gradually decreasing apically, first protarsomere 2× length of second; mesoventrite of female without medial pit (not examined in male); aedeagus (
|
||
<figureCitation id="132E2A52FFE322420E637E28FD51575B" box="[564,654,1035,1062]" captionStart="FIGURE 6" captionStartId="10.[151,260,1117,1143]" captionTargetBox="[180,1401,191,1085]" captionTargetId="figure-25@10.[176,1410,179,1093]" captionTargetPageId="10" captionText="FIGURE 6. Patrus corpulentus: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) female pronotum and elytra, dorsal view, scale bar = 4 mm; (D) gonocoxae, ventral view; (E) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (F) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042674" httpUri="https://zenodo.org/record/5042674/files/figure.png" pageId="9" pageNumber="570">Fig. 6B</figureCitation>
|
||
) with median lobe evenly tapered, apex narrowly rounded; female reproductive tract (
|
||
<figureCitation id="132E2A52FFE322420D3B7E0CFE1E5737" box="[364,449,1071,1098]" captionStart="FIGURE 6" captionStartId="10.[151,260,1117,1143]" captionTargetBox="[180,1401,191,1085]" captionTargetId="figure-25@10.[176,1410,179,1093]" captionTargetPageId="10" captionText="FIGURE 6. Patrus corpulentus: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) female pronotum and elytra, dorsal view, scale bar = 4 mm; (D) gonocoxae, ventral view; (E) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (F) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042674" httpUri="https://zenodo.org/record/5042674/files/figure.png" pageId="9" pageNumber="570">Fig. 6E</figureCitation>
|
||
) without additional sclerotized structure between gonocoxae (
|
||
<figureCitation id="132E2A52FFE3224208217E0CFB105737" box="[1142,1231,1071,1098]" captionStart="FIGURE 6" captionStartId="10.[151,260,1117,1143]" captionTargetBox="[180,1401,191,1085]" captionTargetId="figure-25@10.[176,1410,179,1093]" captionTargetPageId="10" captionText="FIGURE 6. Patrus corpulentus: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) female pronotum and elytra, dorsal view, scale bar = 4 mm; (D) gonocoxae, ventral view; (E) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (F) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042674" httpUri="https://zenodo.org/record/5042674/files/figure.png" pageId="9" pageNumber="570">Fig. 6D</figureCitation>
|
||
), fertilization duct as in
|
||
<figureCitation id="132E2A52FFE322420C857E70FEFB5710" box="[210,292,1107,1133]" captionStart="FIGURE 6" captionStartId="10.[151,260,1117,1143]" captionTargetBox="[180,1401,191,1085]" captionTargetId="figure-25@10.[176,1410,179,1093]" captionTargetPageId="10" captionText="FIGURE 6. Patrus corpulentus: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) female pronotum and elytra, dorsal view, scale bar = 4 mm; (D) gonocoxae, ventral view; (E) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (F) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042674" httpUri="https://zenodo.org/record/5042674/files/figure.png" pageId="9" pageNumber="570">
|
||
Fig.
|
||
<specimenCount id="9D13FD5EFFE322420D517E70FEFB5710" box="[262,292,1107,1133]" pageId="9" pageNumber="570" type="generic">6F</specimenCount>
|
||
</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BAA36D7FFE322420C907E54FEEB5614" blockId="9.[151,1437,386,1962]" pageId="9" pageNumber="570">
|
||
<emphasis id="B961EAC5FFE322420C907E54FE0E57EC" bold="true" box="[199,465,1143,1169]" pageId="9" pageNumber="570">Differential diagnosis.</emphasis>
|
||
This species can be distinguished from all other known members of the
|
||
<taxonomicName id="4C154D54FFE3224209737E5BFA4357EC" baseAuthorityName="Regimbart" baseAuthorityYear="1892" box="[1316,1436,1143,1169]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="570" phylum="Arthropoda" rank="species" species="landaisi">
|
||
<emphasis id="B961EAC5FFE3224209737E5BFA4357EC" box="[1316,1436,1143,1169]" italics="true" pageId="9" pageNumber="570">P. landaisi</emphasis>
|
||
</taxonomicName>
|
||
species group in
|
||
<collectingCountry id="F3027647FFE322420D047EB8FE6857C8" box="[339,439,1179,1205]" name="Thailand" pageId="9" pageNumber="570">Thailand</collectingCountry>
|
||
by the shape of the glabrous region of the elytra (
|
||
<figureCitation id="132E2A52FFE322420FB57EB8FB9B57C8" box="[994,1092,1179,1205]" captionStart="FIGURE 2" captionStartId="6.[151,260,1861,1887]" captionTargetBox="[151,1434,179,1836]" captionTargetId="figure-25@6.[151,1435,179,1836]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042660" httpUri="https://zenodo.org/record/5042660/files/figure.png" pageId="9" pageNumber="570">Figs. 2B</figureCitation>
|
||
,
|
||
<figureCitation id="132E2A52FFE3224208187EB8FBAD57C8" box="[1103,1138,1179,1205]" captionStart="FIGURE 3" captionStartId="7.[151,260,1865,1891]" captionTargetBox="[151,1434,182,1841]" captionTargetId="figure-18@7.[151,1435,179,1841]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal habitus of females of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042662" httpUri="https://zenodo.org/record/5042662/files/figure.png" pageId="9" pageNumber="570">3B</figureCitation>
|
||
), and by having the elytral apex narrowly or weakly rounded with visible apicolateral sinuation (
|
||
<figureCitation id="132E2A52FFE322420FF07E9CFBEE57A7" box="[935,1073,1215,1242]" captionStart="FIGURE 6" captionStartId="10.[151,260,1117,1143]" captionTargetBox="[180,1401,191,1085]" captionTargetId="figure-25@10.[176,1410,179,1093]" captionTargetPageId="10" captionText="FIGURE 6. Patrus corpulentus: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) female pronotum and elytra, dorsal view, scale bar = 4 mm; (D) gonocoxae, ventral view; (E) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (F) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042674" httpUri="https://zenodo.org/record/5042674/files/figure.png" pageId="9" pageNumber="570">Fig. 6A, 6C</figureCitation>
|
||
). Both males and females of the other species lack apicolateral sinuation to the elytral apex and they have their glabrous region extending a relatively shorter distance along the elytra suture, up to 7/9 elytral length. These features, combined with
|
||
<taxonomicName id="4C154D54FFE3224208AE7F2BFA43565C" authorityName="Regimbart" authorityYear="1884" box="[1273,1436,1287,1313]" class="Insecta" family="Gyrinidae" genus="Orectochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="570" phylum="Arthropoda" rank="species" species="corpulentus">
|
||
<emphasis id="B961EAC5FFE3224208AE7F2BFA43565C" box="[1273,1436,1287,1313]" italics="true" pageId="9" pageNumber="570">P. corpulentus</emphasis>
|
||
</taxonomicName>
|
||
being the only
|
||
<taxonomicName id="4C154D54FFE322420D1A7F0FFE185638" baseAuthorityName="Regimbart" baseAuthorityYear="1892" box="[333,455,1323,1349]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="570" phylum="Arthropoda" rank="species" species="landaisi">
|
||
<emphasis id="B961EAC5FFE322420D1A7F0FFE185638" box="[333,455,1323,1349]" italics="true" pageId="9" pageNumber="570">P. landaisi</emphasis>
|
||
</taxonomicName>
|
||
group species found at lower elevation in
|
||
<collectingCountry id="F3027647FFE322420F9F7F08FBF15638" box="[968,1070,1323,1349]" name="Thailand" pageId="9" pageNumber="570">Thailand</collectingCountry>
|
||
, should allow its unambiguous identification.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30F655CFFE322420C907F50FD75551B" pageId="9" pageNumber="570" type="distribution">
|
||
<paragraph id="8BAA36D7FFE322420C907F50FD75551B" blockId="9.[151,1437,386,1962]" pageId="9" pageNumber="570">
|
||
<emphasis id="B961EAC5FFE322420C907F50FEBE56F0" bold="true" box="[199,353,1395,1421]" pageId="9" pageNumber="570">Distribution.</emphasis>
|
||
<taxonomicName id="4C154D54FFE322420D3D7F57FD9C56F0" authorityName="Borneo" baseAuthorityName="Regimbart" baseAuthorityYear="1884" box="[362,579,1395,1421]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="570" phylum="Arthropoda" rank="species" species="corpulentus">
|
||
<emphasis id="B961EAC5FFE322420D3D7F57FD9C56F0" box="[362,579,1395,1421]" italics="true" pageId="9" pageNumber="570">Patrus corpulentus</emphasis>
|
||
</taxonomicName>
|
||
was originally described from Java, potentially in error (
|
||
<bibRefCitation id="EF844B26FFE32242088D7F50FA4856F0" author="Regimbart, M." box="[1242,1431,1395,1421]" pageId="9" pageNumber="570" pagination="381 - 482" refId="ref22884" refString="Regimbart, M. (1884) Essai monographique de la famille des Gyrinidae. 3 e partie. Annales de la Societe entomologique de France, 6, 381 - 482, pls. 311 - 314." type="journal article" year="1884">Régimbart 1884</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF844B26FFE322420CC07FB4FEC856CC" author="Ochs" box="[151,279,1431,1457]" pageId="9" pageNumber="570" pagination="1 - 39" refId="ref22521" refString="Ochs, G. (1930) Gyrinoidea. Catalogue of Indian Insects, 19, 1 - 39." type="journal article" year="1930">Ochs 1930</bibRefCitation>
|
||
), and Borneo, potentially in reference to a separate species (see below). Subsequent records for this species are from Tenasserim and the Karen hills regions of
|
||
<collectingCountry id="F3027647FFE322420F7A7F9FFC4556AB" box="[813,922,1468,1494]" name="Myanmar" pageId="9" pageNumber="570">Myanmar</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF844B26FFE322420FFF7F98FBBD56A8" author="Regimbart, M." box="[936,1122,1467,1493]" pageId="9" pageNumber="570" pagination="537 - 554" refId="ref23003" refString="Regimbart, M. (1891) Viaggio di Leonardo Fea in Birmania e regioni vicine. XXXIII. Enumeration des Dytiscidae et Gyrinidae recueillis par M. Leonardo Fea, dans ses voyages en Birmanie et regions voisines. Annali del Museo Civico di Storia Naturale Giacomo Doria Genova, 10 (2), 537 - 554." type="journal article" year="1891">Régimbart 1891</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF844B26FFE32242083A7F98FA9656A8" author="Zimmermann, A." box="[1133,1353,1467,1493]" pageId="9" pageNumber="570" pagination="135 - 170" refId="ref23783" refString="Zimmermann, A. (1917) Der derzeitige Bestand der Gyrinidensammlung des Deutschen Entomologischen Museums in Berlin- Dahlem und die wissenschaftlichen Ergebnisse ihrer Durcharbeitung (Col.). Entomologische Mitteilungen aus dem Zoologischen Staatsinstitut und Zoologischen Museum Hamburg, 6, 135 - 170. https: // doi. org / 10.5962 / bhl. part. 10104" type="journal article" year="1917">Zimmermann 1917</bibRefCitation>
|
||
). It had also been recorded in
|
||
<collectingCountry id="F3027647FFE322420DD87FFCFE2C5684" box="[399,499,1503,1529]" name="Thailand" pageId="9" pageNumber="570">Thailand</collectingCountry>
|
||
from
|
||
<collectingRegion id="49D1F835FFE322420E6F7FC3FD255684" box="[568,762,1503,1530]" country="Thailand" name="Ranong" pageId="9" pageNumber="570">Ranong Province</collectingRegion>
|
||
by
|
||
<bibRefCitation id="EF844B26FFE322420F727FFCFC2F5684" author="Regimbart, M." box="[805,1008,1503,1529]" pageId="9" pageNumber="570" pagination="663 - 752" refId="ref23063" refString="Regimbart, M. (1892) Essai monographique de la famille des Gyrinidae. 2 e supplement. Annales de la Societe entomologique de France, 60 (1891), 663 - 752, pls. 618 - 619." type="journal article" year="1892">Régimbart (1892)</bibRefCitation>
|
||
, and subsequently in
|
||
<collectingRegion id="49D1F835FFE3224208B47FFCFA425684" box="[1251,1437,1503,1529]" country="Thailand" name="Phuket" pageId="9" pageNumber="570">Phuket Province</collectingRegion>
|
||
by
|
||
<bibRefCitation id="EF844B26FFE322420CE97C20FE905560" author="Ochs" box="[190,335,1539,1565]" pageId="9" pageNumber="570" pagination="242 - 245" refId="ref22453" refString="Ochs, G. (1927) XII. Papers on Malayan aquatic biology. iv. On the Gyrinidae of the Malay Peninsula. Journal of the Federated Malay States Museums, 13, 242 - 245." type="journal article" year="1927">Ochs (1927)</bibRefCitation>
|
||
. Recently
|
||
<specimenCount id="9D13FD5EFFE322420D9B7C27FD925560" box="[460,589,1539,1566]" pageId="9" pageNumber="570" type="female">two female</specimenCount>
|
||
specimens were found together within
|
||
<collectingRegion id="49D1F835FFE32242084B7C20FACE5560" box="[1052,1297,1539,1565]" country="Thailand" name="Surat Thani" pageId="9" pageNumber="570">Surat Thani Province</collectingRegion>
|
||
(
|
||
<figureCitation id="132E2A52FFE3224209737C20FA4F5560" box="[1316,1424,1539,1565]" captionStart="FIGURE 14" captionStartId="26.[151,260,743,769]" captionTargetBox="[152,1436,179,719]" captionTargetId="figure-25@26.[151,1436,179,719]" captionTargetPageId="26" captionText="FIGURE 14. (A–B) Distribution records of the P. landaisi species group in Thailand." figureDoi="http://doi.org/10.5281/zenodo.5042692" httpUri="https://zenodo.org/record/5042692/files/figure.png" pageId="9" pageNumber="570">Fig. 14A</figureCitation>
|
||
). Unlike the other
|
||
<taxonomicName id="4C154D54FFE322420D017C0BFE12553C" baseAuthorityName="Regimbart" baseAuthorityYear="1892" box="[342,461,1575,1601]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="570" phylum="Arthropoda" rank="species" species="landaisi">
|
||
<emphasis id="B961EAC5FFE322420D017C0BFE12553C" box="[342,461,1575,1601]" italics="true" pageId="9" pageNumber="570">P. landaisi</emphasis>
|
||
</taxonomicName>
|
||
species group members in
|
||
<collectingCountry id="F3027647FFE322420F557C04FCB7553C" box="[770,872,1575,1601]" name="Thailand" pageId="9" pageNumber="570">Thailand</collectingCountry>
|
||
,
|
||
<taxonomicName id="4C154D54FFE322420F237C0BFBCB553C" authorityName="Borneo" baseAuthorityName="Regimbart" baseAuthorityYear="1884" box="[884,1044,1575,1601]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="570" phylum="Arthropoda" rank="species" species="corpulentus">
|
||
<emphasis id="B961EAC5FFE322420F237C0BFBCB553C" box="[884,1044,1575,1601]" italics="true" pageId="9" pageNumber="570">P. corpulentus</emphasis>
|
||
</taxonomicName>
|
||
appears to occur at low elevations rather than in high-elevation mountain streams.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30F655CFFE322420C907C4CFB2A54D4" pageId="9" pageNumber="570" type="discussion">
|
||
<paragraph id="8BAA36D7FFE322420C907C4CFB2A54D4" blockId="9.[151,1437,386,1962]" pageId="9" pageNumber="570">
|
||
<emphasis id="B961EAC5FFE322420C907C4CFE9355F4" bold="true" box="[199,332,1647,1673]" pageId="9" pageNumber="570">Discussion.</emphasis>
|
||
The original description of
|
||
<taxonomicName id="4C154D54FFE322420ED17C53FC2955F4" authority="(Regimbart 1884)" authorityName="Borneo" baseAuthorityName="Regimbart" baseAuthorityYear="1884" box="[646,1014,1647,1673]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="570" phylum="Arthropoda" rank="species" species="corpulentus">
|
||
<emphasis id="B961EAC5FFE322420ED17C53FCF955F4" box="[646,806,1647,1673]" italics="true" pageId="9" pageNumber="570">P. corpulentus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF844B26FFE322420F627C4CFC3255F4" author="Regimbart, M." box="[821,1005,1647,1673]" pageId="9" pageNumber="570" pagination="381 - 482" refId="ref22884" refString="Regimbart, M. (1884) Essai monographique de la famille des Gyrinidae. 3 e partie. Annales de la Societe entomologique de France, 6, 381 - 482, pls. 311 - 314." type="journal article" year="1884">Régimbart 1884</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
was based on
|
||
<specimenCount id="9D13FD5EFFE3224208CD7C4CFAC755F4" box="[1178,1304,1647,1673]" pageId="9" pageNumber="570" type="female">five female</specimenCount>
|
||
specimens: one from Java, one from Borneo, and three without locality indications. Régimbart later (1888, 1892) described specimens of what he considered to be males of
|
||
<taxonomicName id="4C154D54FFE322420E847C9BFCA955AC" authorityName="Borneo" baseAuthorityName="Regimbart" baseAuthorityYear="1884" box="[723,886,1719,1745]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="570" phylum="Arthropoda" rank="species" species="corpulentus">
|
||
<emphasis id="B961EAC5FFE322420E847C9BFCA955AC" box="[723,886,1719,1745]" italics="true" pageId="9" pageNumber="570">P. corpulentus</emphasis>
|
||
</taxonomicName>
|
||
from Tenasserim and the Karen hills region of
|
||
<collectingCountry id="F3027647FFE322420CC07CFFFED9558B" box="[151,262,1756,1782]" name="Myanmar" pageId="9" pageNumber="570">Myanmar</collectingCountry>
|
||
. In the same publications he clearly distinguishes these last specimens, the female from Java and the unlabeled females, from the single female from Borneo as belonging to different “varieties”. Therefore, it seems likely that the
|
||
<typeStatus id="54AE8875FFE322420D6F7D07FE505443" box="[312,399,1828,1854]" pageId="9" pageNumber="570" type="syntype">syntype</typeStatus>
|
||
series may consist of two different species, one from the Malay peninsula (accepting “Java” as an erroneous locality), and another from Borneo (P. Mazzoldi pers. comm.). We examined the
|
||
<typeStatus id="54AE8875FFE3224208A77D6BFA98541F" box="[1264,1351,1864,1890]" pageId="9" pageNumber="570" type="syntype">syntype</typeStatus>
|
||
female labeled as being from Java and it is indeed conspecific to the specimens we collected from Surat Thani Province, thus we consider them as belonging to
|
||
<taxonomicName id="4C154D54FFE322420E1C7DB3FC2A54D4" authority="sensu Regimbart (1884" authorityName="sensu Regimbart" authorityYear="1884" box="[587,1013,1935,1962]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="570" phylum="Arthropoda" rank="species" species="corpulentus">
|
||
<emphasis id="B961EAC5FFE322420E1C7DB3FD3454D4" box="[587,747,1935,1961]" italics="true" pageId="9" pageNumber="570">P. corpulentus</emphasis>
|
||
sensu
|
||
<bibRefCitation id="EF844B26FFE322420F607DACFC2A54D4" author="Regimbart, M." box="[823,1013,1935,1961]" pageId="9" pageNumber="570" pagination="381 - 482" refId="ref22884" refString="Regimbart, M. (1884) Essai monographique de la famille des Gyrinidae. 3 e partie. Annales de la Societe entomologique de France, 6, 381 - 482, pls. 311 - 314." type="journal article" year="1884">Régimbart (1884</bibRefCitation>
|
||
</taxonomicName>
|
||
[in part], 1888, 1892).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |