243 lines
29 KiB
XML
243 lines
29 KiB
XML
<document id="4A6F867F1305BCAEB4761250D1BD1A4E" ID-CLB-Dataset="8579" ID-DOI="10.11646/zootaxa.4850.1.1" ID-GBIF-Dataset="686337a2-1528-43d0-ae32-68ca023d8df3" ID-ISSN="1175-5326" ID-Zenodo-Dep="4407072" ID-ZooBank="18AA0411-0E54-4922-84C7-608EAC68D281" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1609428297648" checkinUser="plazi" docAuthor="Guéorguiev, Borislav & Liang, Hongbin" docDate="2020" docId="03BC5E5B2958FFA5FF4BF899EE08FBC8" docLanguage="en" docName="zootaxa.4850.1.1.pdf" docOrigin="Zootaxa 4850 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Holosoma rambouseki" docType="treatment" docVersion="8" lastPageNumber="27" masterDocId="FF8526232940FFBFFFDCFFB4EC2EFFFD" masterDocTitle="Revision of the Palaearctic and Oriental representatives of Lachnocrepis LeConte and Oodes Bonelli (Coleoptera: Carabidae), with special account on Chinese species" masterLastPageNumber="89" masterPageNumber="1" pageNumber="25" updateTime="1698852686469" updateUser="ExternalLinkService">
|
||
<mods:mods id="FD3B9502D9309EB892C575F48D79BDE8" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="4722639DAE211F52ABD03F7E96DC537D">
|
||
<mods:title id="9EC00A40B545B85FF360ACFAB64B4040">Revision of the Palaearctic and Oriental representatives of Lachnocrepis LeConte and Oodes Bonelli (Coleoptera: Carabidae), with special account on Chinese species</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="887F743BF0C2343B23302DC98704DF5F" type="personal">
|
||
<mods:role id="E5DE414F692283783D351878E925FFD7">
|
||
<mods:roleTerm id="21356AB5A4532619C36CD663A4BC14AB">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="5AB7D23366D2D31E0C81C92FCC6F5D6C">Guéorguiev, Borislav</mods:namePart>
|
||
<mods:nameIdentifier id="CAD3DF7E7B1BD175E00CD087036D5944" type="ORCID">0000-0002-8532-0657</mods:nameIdentifier>
|
||
<mods:nameIdentifier id="4526D658E4B7059E704000BDC7A12AF7" type="email">gueorguiev@nmnhs.com</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="26939E4D75880F95DA41CD3446683090" type="personal">
|
||
<mods:role id="004B4AEF3520F4DB9182A3AA0F38F5A2">
|
||
<mods:roleTerm id="00D0D1EC6B4DBAA07DDBB7D12D00D35A">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="73380271682A04741F3C88D4CC3505CB">Liang, Hongbin</mods:namePart>
|
||
<mods:nameIdentifier id="AC3DFFA655514404BD2D7CA798BBBA2F" type="ORCID">0000-0002-9668-1167</mods:nameIdentifier>
|
||
<mods:nameIdentifier id="63E3363D750446B28382C7486C3B9E05" type="email">lianghb@ioz.ac.cn</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:typeOfResource id="C9C3035E3574F62870E8B651BA8CC347">text</mods:typeOfResource>
|
||
<mods:relatedItem id="993D09686773F12F273E637D67344221" type="host">
|
||
<mods:titleInfo id="121F7D4FB92B3A5700E3C3035432DCEA">
|
||
<mods:title id="018B8DA14EBF9A504956FCEC07970F3A">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="F62A6260F3E87BF81AE402B93A9C5D65">
|
||
<mods:date id="51E91B47EAE3F8792E455D1302386DC3">2020</mods:date>
|
||
<mods:detail id="2ADDA02ECE5C8205DC4F3F4A86697174" type="pubDate">
|
||
<mods:number id="D4040D9A55191C04AD2F08DDEAB0A69C">2020-09-08</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="760105B0079157C713D17F601F774A3C" type="volume">
|
||
<mods:number id="570A341620BC5D075F5F3567D1592429">4850</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="509DE180E4C91DD98000C553908DF3C4" type="issue">
|
||
<mods:number id="78C989EAB17775CFF745C12E97E1686F">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="340B06C8ED465C6B2FEA7AE75013FCEF" unit="page">
|
||
<mods:start id="BC0B049EC3CE832C83D912A3C96A012E">1</mods:start>
|
||
<mods:end id="A6CF29D84ED2AFCB6A988AF2EADE5A05">89</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="211B8AD164F7E378A582FD1C0341030E">journal article</mods:classification>
|
||
<mods:identifier id="5E19C558C828E82FC28C69DDBE162AAF" type="CLB-Dataset">8579</mods:identifier>
|
||
<mods:identifier id="35866427D70C2D68F9640D303B2ED4BE" type="DOI">10.11646/zootaxa.4850.1.1</mods:identifier>
|
||
<mods:identifier id="08FBF924362BF5CB3A2C8E162D264C8A" type="GBIF-Dataset">686337a2-1528-43d0-ae32-68ca023d8df3</mods:identifier>
|
||
<mods:identifier id="D00CB3521E8BB79E402BDD7170525324" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="9A5813CC20F53A1F9E24195B80CCE32D" type="Zenodo-Dep">4407072</mods:identifier>
|
||
<mods:identifier id="EDC7CB41362D69FD05435492939A978B" type="ZooBank">18AA0411-0E54-4922-84C7-608EAC68D281</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03BC5E5B2958FFA5FF4BF899EE08FBC8" ID-DOI="http://doi.org/10.5281/zenodo.4479999" ID-GBIF-Taxon="177139741" ID-Zenodo-Dep="4479999" LSID="urn:lsid:plazi:treatment:03BC5E5B2958FFA5FF4BF899EE08FBC8" httpUri="http://treatment.plazi.org/id/03BC5E5B2958FFA5FF4BF899EE08FBC8" lastPageId="26" lastPageNumber="27" pageId="24" pageNumber="25">
|
||
<subSubSection id="C30FBCC62958FFA7FF4BF899EDDFF8BA" box="[151,497,1837,1864]" pageId="24" pageNumber="25" type="nomenclature">
|
||
<paragraph id="8BAAEF4D2958FFA7FF4BF899EDDFF8BA" blockId="24.[151,497,1837,1864]" box="[151,497,1837,1864]" pageId="24" pageNumber="25">
|
||
<emphasis id="B961335F2958FFA7FF4BF899EDDFF8BA" bold="true" box="[151,497,1837,1864]" pageId="24" pageNumber="25">
|
||
“
|
||
<taxonomicName id="4C1594CE2958FFA7FF7AF899ED1AF8B5" ID-CoL="8LGGX" baseAuthorityName="Jedlicka" baseAuthorityYear="1931" box="[166,308,1837,1864]" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="rambouseki">
|
||
<emphasis id="B961335F2958FFA7FF7AF899ED1AF8B5" bold="true" box="[166,308,1837,1864]" italics="true" pageId="24" pageNumber="25">rambouseki</emphasis>
|
||
</taxonomicName>
|
||
” species group
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30FBCC62958FFA5FF4BF8C4E85FFEBC" lastPageId="26" lastPageNumber="27" pageId="24" pageNumber="25" type="diagnosis">
|
||
<paragraph id="8BAAEF4D2958FFA7FF4BF8C4ED89F80B" blockId="24.[151,1437,1904,2038]" pageId="24" pageNumber="25">
|
||
<emphasis id="B961335F2958FFA7FF4BF8C4ED24F877" bold="true" box="[151,266,1904,1930]" pageId="24" pageNumber="25">Diagnosis</emphasis>
|
||
. The species group includes five wingless species that possess the following synapomorphies: (1) metepisternum one-and-a-half times or more wider than long; (2) extraordinarily elongate spermatheca; and (3) apical gonocoxite short, subtriangular, with dorsomedial ensiform seta situated near the middle or equidistant from the middle and the base.
|
||
</paragraph>
|
||
<caption id="DF6ABFC52959FFA6FF4BF8D4EF84F818" ID-DOI="http://doi.org/10.5281/zenodo.4407088" ID-Zenodo-Dep="4407088" httpUri="https://zenodo.org/record/4407088/files/figure.png" pageId="25" pageNumber="26" startId="25.[151,250,1888,1912]" targetBox="[283,1289,208,1837]" targetPageId="25">
|
||
<paragraph id="8BAAEF4D2959FFA6FF4BF8D4EF84F818" blockId="25.[151,1436,1888,2021]" pageId="25" pageNumber="26">
|
||
<emphasis id="B961335F2959FFA6FF4BF8D4ED39F885" bold="true" box="[151,279,1888,1912]" pageId="25" pageNumber="26">FIGURE 8.</emphasis>
|
||
<taxonomicName id="4C1594CE2959FFA6FEFDF8D4EE04F885" authority="Guéorguiev & Liang, 2020" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[289,554,1888,1912]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="ampliusculus" status="sp. nov.">
|
||
<emphasis id="B961335F2959FFA6FEFDF8D4EE04F885" box="[289,554,1888,1912]" italics="true" pageId="25" pageNumber="26">Pseudoodes ampliusculus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B961335F2959FFA6FDE8F8D5EE5EF884" bold="true" box="[564,624,1889,1913]" pageId="25" pageNumber="26">
|
||
<taxonomicNameLabel id="A2528E242959FFA6FDE8F8D5EE5EF884" box="[564,624,1889,1913]" pageId="25" pageNumber="26" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype. A: habitus and labels; B: mentum (white spots show locations of submental setiferous punctures); C: prosternal process; D: right metepisternum and metacoxa (arrow shows lateral end of metacoxal basal sulcus); E: left gonocoxite, ventral view; F: spermathecal complex and gonocoxites, ventral view. Scale lines = 2 mm (Fig. A); = 0.5 mm (Fig. B, F); = 1 mm (Figs C–D); = 0.2 mm (Fig. E).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAAEF4D295AFFA5FF1BFF23E85FFEBC" blockId="26.[151,1437,151,1077]" pageId="26" pageNumber="27">
|
||
In addition, the species from the “
|
||
<taxonomicName id="4C1594CE295AFFA5FD8AFF23EEF7FF4C" baseAuthorityName="Jedlicka" baseAuthorityYear="1931" box="[598,729,151,177]" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="rambouseki">
|
||
<emphasis id="B961335F295AFFA5FD8AFF23EEF7FF4C" box="[598,729,151,177]" italics="true" pageId="26" pageNumber="27">rambouseki</emphasis>
|
||
</taxonomicName>
|
||
” group have the anterior angles of pronotum more projected and pointed, as well as the parascutellar stria more obliterated than in other representatives of the genus (except
|
||
<taxonomicName id="4C1594CE295AFFA5FF4BFF6BED15FF04" authorityName="Chaudoir" authorityYear="1882" box="[151,315,223,249]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="coelestinus">
|
||
<emphasis id="B961335F295AFFA5FF4BFF6BEC86FF05" box="[151,168,223,248]" italics="true" pageId="26" pageNumber="27">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F295AFFA5FF63FF6BED15FF04" box="[191,315,223,249]" italics="true" pageId="26" pageNumber="27">coelestinus</emphasis>
|
||
</taxonomicName>
|
||
which also has an obliterated parascutellar stria). Further, three species (
|
||
<taxonomicName id="4C1594CE295AFFA5FB14FF6BE940FF04" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[1224,1390,223,249]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="hunanensis">
|
||
<emphasis id="B961335F295AFFA5FB14FF6BE8F7FF05" box="[1224,1241,223,248]" italics="true" pageId="26" pageNumber="27">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F295AFFA5FB2CFF6BE940FF04" box="[1264,1390,223,249]" italics="true" pageId="26" pageNumber="27">hunanensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1594CE295AFFA5FA58FF6BED63FEE0" authorityName="Guéorguiev & Liang" authorityYear="2020" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="leigongshanicus">
|
||
<emphasis id="B961335F295AFFA5FA58FF6BE9BBFF05" box="[1412,1429,223,248]" italics="true" pageId="26" pageNumber="27">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F295AFFA5FF4BFEB7ED63FEE0" box="[151,333,259,285]" italics="true" pageId="26" pageNumber="27">leigongshanicus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C1594CE295AFFA5FE56FEB7EE02FEE0" baseAuthorityName="Jedlicka" baseAuthorityYear="1931" box="[394,556,259,285]" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="rambouseki">
|
||
<emphasis id="B961335F295AFFA5FE56FEB7EDB5FEE1" box="[394,411,259,284]" italics="true" pageId="26" pageNumber="27">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F295AFFA5FE75FEB7EE02FEE0" box="[425,556,259,285]" italics="true" pageId="26" pageNumber="27">rambouseki</emphasis>
|
||
</taxonomicName>
|
||
) have a metatrochanter more or less pointed at the apex (
|
||
<figureCitation id="132EF3C8295AFFA5FB77FEB7E93DFEE0" box="[1195,1299,259,285]" captionStart="FIGURE 10" captionStartId="29.[151,250,1887,1911]" captionTargetBox="[238,1325,200,1849]" captionTargetId="figure-17@29.[222,1365,181,1863]" captionTargetPageId="29" captionText="FIGURE 10. Pseudoodes hunanensis sp. n. (A, C–G: holotype; B, H–I: female paratype). A: habitus and labels; B: habitus; C: mentum; D: prosternal process; E: right metepisternum, metacoxa and metatrochanter; F: median lobe of aedeagus, left lateral view; G: same, dorsal view; H: left gonocoxite, ventral view; I: spermathecal complex and gonocoxites, ventral view. Scale lines = 2 mm (Figs A–B); = 0.5 mm (Fig. C–D, F–G, I); = 1 mm (Fig. E); = 0.2 mm (Fig. H)." figureDoi="http://doi.org/10.5281/zenodo.4407092" httpUri="https://zenodo.org/record/4407092/files/figure.png" pageId="26" pageNumber="27">Figs 10E</figureCitation>
|
||
,
|
||
<figureCitation id="132EF3C8295AFFA5FAC3FEB7E97EFEE0" box="[1311,1360,259,285]" captionStart="FIGURE 11" captionStartId="31.[151,250,1886,1910]" captionTargetBox="[241,1335,196,1850]" captionTargetId="figure-17@31.[222,1365,181,1862]" captionTargetPageId="31" captionText="FIGURE 11. Pseudoodes leigongshanicus sp. n. (A–G:holotype; H–J:paratype).A:habitus andlabels; B: mentum; C: prosternal process; D: right metepisternum and metacoxa; E: median lobe of aedeagus, left lateral view; F: same, anterior dorsal view; G: same, posterior dorsal view; H: habitus of paratype; I: left gonocoxite, ventral view; J: spermathecal complex and gonocoxites, ventral view. Scale lines = 2 mm (Figs A, H); = 0.5 mm (Figs B–C, E–G, J); = 1 mm (Figs D); = 0.2 mm (Fig. I)." figureDoi="http://doi.org/10.5281/zenodo.4407094" httpUri="https://zenodo.org/record/4407094/files/figure.png" pageId="26" pageNumber="27">11D</figureCitation>
|
||
,
|
||
<figureCitation id="132EF3C8295AFFA5FA80FEB7E9BEFEE0" box="[1372,1424,259,285]" captionStart="FIGURE 12" captionStartId="33.[151,250,1820,1844]" captionTargetBox="[261,1305,201,1783]" captionTargetId="figure-17@33.[244,1342,181,1796]" captionTargetPageId="33" captionText="FIGURE 12. Pseudoodes rambouseki (Jedlička, 1931) (A: lectotype of Holosoma rambouseki and its labels; B–D:topotype male specimen, China, Jinfo Shan, 29°01N 107°14E; E–H: topotype female specimen, China Jinfo Shan, N29.04043 E107.18410; I–J: topotype female specimen, SE Sichuan Jinfo Shan, 29°01N 107°14E). A–B, Е: habitus; C: median lobe of aedeagus, left lateral view; D: same, dorsal view; F: mentum; G: prosternal process; H: left metepisternum and metacoxa; I: left gonocoxite, ventral view; J: spermathecal complex and gonocoxites, ventral view. Scale lines = 2 mm (Figs A–B, E); = 1 mm (Fig. H); = 0.5 mm (Figs C–D, F–G, J); = 0.2 mm (Fig. I)." figureDoi="http://doi.org/10.5281/zenodo.4407096" httpUri="https://zenodo.org/record/4407096/files/figure.png" pageId="26" pageNumber="27">12H</figureCitation>
|
||
). All five representatives are wingless, with fused elytra and reduced metathoracic wings.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30FBCC6295AFFA5FF1BFEFFEE08FBC8" pageId="26" pageNumber="27" type="discussion">
|
||
<paragraph id="8BAAEF4D295AFFA5FF1BFEFFE902FD54" blockId="26.[151,1437,151,1077]" pageId="26" pageNumber="27">
|
||
<emphasis id="B961335F295AFFA5FF1BFEFFED24FE98" bold="true" box="[199,266,331,357]" pageId="26" pageNumber="27">Notes</emphasis>
|
||
.
|
||
<taxonomicName id="4C1594CE295AFFA5FECAFEFFEE30FE98" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[278,542,331,357]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="hunanensis">
|
||
<emphasis id="B961335F295AFFA5FECAFEFFEE30FE98" box="[278,542,331,357]" italics="true" pageId="26" pageNumber="27">Pseudoodes hunanensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1594CE295AFFA5FD8CFEFFEF3BFE98" box="[592,789,331,357]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="leigonshanicus">
|
||
<emphasis id="B961335F295AFFA5FD8CFEFFEE4FFE99" box="[592,609,331,356]" italics="true" pageId="26" pageNumber="27">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F295AFFA5FDB1FEFFEF3BFE98" box="[621,789,331,357]" italics="true" pageId="26" pageNumber="27">leigonshanicus</emphasis>
|
||
</taxonomicName>
|
||
(supposedly also
|
||
<taxonomicName id="4C1594CE295AFFA5FC0BFEFFE8A5FE98" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[983,1163,331,357]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="emeishanicus">
|
||
<emphasis id="B961335F295AFFA5FC0BFEFFEFC6FE99" box="[983,1000,331,356]" italics="true" pageId="26" pageNumber="27">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F295AFFA5FC28FEFFE8A5FE98" box="[1012,1163,331,357]" italics="true" pageId="26" pageNumber="27">emeishanicus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1594CE295AFFA5FB61FEFFE97EFE98" box="[1213,1360,331,357]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="tianlinenis">
|
||
<emphasis id="B961335F295AFFA5FB61FEFFE8E0FE99" box="[1213,1230,331,356]" italics="true" pageId="26" pageNumber="27">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F295AFFA5FB06FEFFE97EFE98" box="[1242,1360,331,357]" italics="true" pageId="26" pageNumber="27">tianlinenis</emphasis>
|
||
</taxonomicName>
|
||
whose females are still unknown) have very long, undifferentiated spermatheca. On the other hand,
|
||
<taxonomicName id="4C1594CE295AFFA5FB6BFEDBE974FE74" baseAuthorityName="Jedlicka" baseAuthorityYear="1931" box="[1207,1370,367,393]" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="rambouseki">
|
||
<emphasis id="B961335F295AFFA5FB6BFEDBE8E6FE75" box="[1207,1224,367,392]" italics="true" pageId="26" pageNumber="27">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F295AFFA5FB0BFEDBE974FE74" box="[1239,1370,367,393]" italics="true" pageId="26" pageNumber="27">rambouseki</emphasis>
|
||
</taxonomicName>
|
||
has a spermatheca also quite long but slightly differentiated (like
|
||
<taxonomicName id="4C1594CE295AFFA5FCFFFE27EFFAFE50" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[803,980,403,429]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="ampliusculus">
|
||
<emphasis id="B961335F295AFFA5FCFFFE27EF1AFE51" box="[803,820,403,428]" italics="true" pageId="26" pageNumber="27">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F295AFFA5FC9CFE27EFFAFE50" box="[832,980,403,429]" italics="true" pageId="26" pageNumber="27">ampliusculus</emphasis>
|
||
</taxonomicName>
|
||
) into a seminal canal and a receptaculum. Tendencies to slightly differentiated and undifferentiated spermatheca are aberrant conditions within
|
||
<taxonomicName id="4C1594CE295AFFA5FAD3FE03E9BBFE2C" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[1295,1429,439,465]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B961335F295AFFA5FAD3FE03E9BBFE2C" box="[1295,1429,439,465]" italics="true" pageId="26" pageNumber="27">Pseudoodes</emphasis>
|
||
</taxonomicName>
|
||
. They occur in species from the “
|
||
<taxonomicName id="4C1594CE295AFFA5FDCFFE6FEE89FE08" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[531,679,475,501]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="ampliusculus">
|
||
<emphasis id="B961335F295AFFA5FDCFFE6FEE89FE08" box="[531,679,475,501]" italics="true" pageId="26" pageNumber="27">ampliusculus</emphasis>
|
||
</taxonomicName>
|
||
” and “
|
||
<taxonomicName id="4C1594CE295AFFA5FD26FE6FEF53FE08" baseAuthorityName="Jedlicka" baseAuthorityYear="1931" box="[762,893,475,501]" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="rambouseki">
|
||
<emphasis id="B961335F295AFFA5FD26FE6FEF53FE08" box="[762,893,475,501]" italics="true" pageId="26" pageNumber="27">rambouseki</emphasis>
|
||
</taxonomicName>
|
||
” groups, and are considered a reversal (loss of derived trait present in the common ancestor) due to considerable lenghtening of the apical part of spermatheca (i.e., receptaculum). Similarly, there are two kinds of attachment of the spermathecal gland to spermatheca among species of the “
|
||
<taxonomicName id="4C1594CE295AFFA5FE95FDF3EDE2FD9C" baseAuthorityName="Jedlicka" baseAuthorityYear="1931" box="[329,460,583,609]" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="rambouseki">
|
||
<emphasis id="B961335F295AFFA5FE95FDF3EDE2FD9C" box="[329,460,583,609]" italics="true" pageId="26" pageNumber="27">rambouseki</emphasis>
|
||
</taxonomicName>
|
||
” group, each linked, respectively, to the above mentioned two kinds of spermatheca.
|
||
<taxonomicName id="4C1594CE295AFFA5FF4BFDDFED86FD78" authorityName="Guéorguiev & Liang" authorityYear="2020" baseAuthorityName="Jedlicka" baseAuthorityYear="1931" box="[151,424,619,645]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="rambouseki">
|
||
<emphasis id="B961335F295AFFA5FF4BFDDFED86FD78" box="[151,424,619,645]" italics="true" pageId="26" pageNumber="27">Pseudoodes rambouseki</emphasis>
|
||
</taxonomicName>
|
||
has a spermathecal gland attached at the apical third of spermatheca while
|
||
<taxonomicName id="4C1594CE295AFFA5FB22FDDFE9B2FD78" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[1278,1436,619,645]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="hunanensis">
|
||
<emphasis id="B961335F295AFFA5FB22FDDFE921FD79" box="[1278,1295,619,644]" italics="true" pageId="26" pageNumber="27">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F295AFFA5FAC2FDDFE9B2FD78" box="[1310,1436,619,645]" italics="true" pageId="26" pageNumber="27">hunanensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1594CE295AFFA5FF1BFD3BEDB2FD54" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[199,412,655,681]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="leigongshanicus">
|
||
<emphasis id="B961335F295AFFA5FF1BFD3BECF6FD55" box="[199,216,655,680]" italics="true" pageId="26" pageNumber="27">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F295AFFA5FF3AFD3BEDB2FD54" box="[230,412,655,681]" italics="true" pageId="26" pageNumber="27">leigongshanicus</emphasis>
|
||
</taxonomicName>
|
||
have the spermathecal gland connected at the proximal half of the seminal canal.
|
||
</paragraph>
|
||
<paragraph id="8BAAEF4D295AFFA5FF1BFD07EE08FBC8" blockId="26.[151,1437,151,1077]" pageId="26" pageNumber="27">
|
||
Likely the species of this group derived from a common ancestor (supposed form close to
|
||
<taxonomicName id="4C1594CE295AFFA5FB7BFD07E961FD30" authorityName="Bates" authorityYear="1892" box="[1191,1359,691,717]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="cribristernis">
|
||
<emphasis id="B961335F295AFFA5FB7BFD07E896FD31" box="[1191,1208,691,716]" italics="true" pageId="26" pageNumber="27">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F295AFFA5FB19FD07E961FD30" box="[1221,1359,691,717]" italics="true" pageId="26" pageNumber="27">cribristernis</emphasis>
|
||
</taxonomicName>
|
||
and/or
|
||
<taxonomicName id="4C1594CE295AFFA5FF4BFD63ED67FD0C" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[151,329,727,753]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="ampliusculus">
|
||
<emphasis id="B961335F295AFFA5FF4BFD63EC86FD0D" box="[151,168,727,752]" italics="true" pageId="26" pageNumber="27">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F295AFFA5FF69FD63ED67FD0C" box="[181,329,727,753]" italics="true" pageId="26" pageNumber="27">ampliusculus</emphasis>
|
||
</taxonomicName>
|
||
) via isolation in the high mountainous areas of south and southwest
|
||
<collectingCountry id="F302AFDD295AFFA5FB9CFD63E8A9FD0C" box="[1088,1159,727,753]" name="China" pageId="26" pageNumber="27">China</collectingCountry>
|
||
. Two groups are distinct: (1)
|
||
<taxonomicName id="4C1594CE295AFFA5FF63FD4FED4CFCE8" authorityName="Guéorguiev & Liang" authorityYear="2020" baseAuthorityName="Jedlicka" baseAuthorityYear="1931" box="[191,354,763,789]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="rambouseki">
|
||
<emphasis id="B961335F295AFFA5FF63FD4FECFEFCE9" box="[191,208,763,788]" italics="true" pageId="26" pageNumber="27">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F295AFFA5FF03FD4FED4CFCE8" box="[223,354,763,789]" italics="true" pageId="26" pageNumber="27">rambouseki</emphasis>
|
||
</taxonomicName>
|
||
, having bursa copulatrix with a small dorsal sclerite, spermathecal gland attached at apical third of seminal canal, apical lamella of median lobe narrow, bent to left and rounded at tip, male mesotibia considerably curved and dilated distally, male protarsomere 2 very broad, and MA/MM relatively low; and (2)
|
||
<taxonomicName id="4C1594CE295AFFA5FB21FCF7E9B2FCA0" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[1277,1436,835,861]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="hunanensis">
|
||
<emphasis id="B961335F295AFFA5FB21FCF7E920FCA1" box="[1277,1294,835,860]" italics="true" pageId="26" pageNumber="27">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F295AFFA5FAC2FCF7E9B2FCA0" box="[1310,1436,835,861]" italics="true" pageId="26" pageNumber="27">hunanensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1594CE295AFFA5FF1AFCD3EDB5FC7C" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[198,411,871,897]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="leigongshanicus">
|
||
<emphasis id="B961335F295AFFA5FF1AFCD3ECF9FC7D" box="[198,215,871,896]" italics="true" pageId="26" pageNumber="27">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F295AFFA5FF39FCD3EDB5FC7C" box="[229,411,871,897]" italics="true" pageId="26" pageNumber="27">leigongshanicus</emphasis>
|
||
</taxonomicName>
|
||
, having bursa copulatrix without sclerite, spermathecal gland attached at middle of seminal canal, apical lamella of median lobe wide, straight and more or less pointed at tip, male mesotibia almost straight, slightly dilated distally, male protarsomere 2 moderately broadened, and MA/MM higher. Though females of
|
||
<taxonomicName id="4C1594CE295AFFA5FA58FC1BED00FC10" authorityName="Guéorguiev & Liang" authorityYear="2020" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="emeishanicus">
|
||
<emphasis id="B961335F295AFFA5FA58FC1BE9BBFC35" box="[1412,1429,943,968]" italics="true" pageId="26" pageNumber="27">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F295AFFA5FF4BFC67ED00FC10" box="[151,302,979,1005]" italics="true" pageId="26" pageNumber="27">emeishanicus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1594CE295AFFA5FEBFFC67EE2CFC10" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[355,514,979,1005]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="tianlinensis">
|
||
<emphasis id="B961335F295AFFA5FEBFFC67ED5AFC11" box="[355,372,979,1004]" italics="true" pageId="26" pageNumber="27">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F295AFFA5FE5DFC67EE2CFC10" box="[385,514,979,1005]" italics="true" pageId="26" pageNumber="27">tianlinensis</emphasis>
|
||
</taxonomicName>
|
||
are unknown, judging from the structure of the median lobe (apical lamella similar to that of the two species from
|
||
<collectingRegion id="49D121AF295AFFA5FDD9FC43EE7EFBEC" box="[517,592,1015,1041]" country="China" name="Hunan" pageId="26" pageNumber="27">Hunan</collectingRegion>
|
||
and
|
||
<collectingRegion id="49D121AF295AFFA5FD57FC43EEC3FBEF" box="[651,749,1015,1042]" country="China" name="Guizhou" pageId="26" pageNumber="27">Guizhou</collectingRegion>
|
||
) and less broadened male protarsomere 2, we believe these species belong to the second group.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |