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<document id="353EC7EA4B568E41B3A8B476124A232D" ID-DOI="10.5281/zenodo.204914" ID-GBIF-Dataset="035006f1-9010-4f0b-a5fb-83fed142d245" ID-ISSN="1175-5326" ID-Zenodo-Dep="204914" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460221844229" checkinUser="plazi" docAuthor="Ciccotto, Patrick J., Konings, Adrianus &amp; Stauffer, Jay R." docDate="2011" docId="03BB942A0D7C836C9DEFFC73FD6EFA2A" docLanguage="en" docName="zt02738p025.pdf" docOrigin="Zootaxa 2738" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Metriaclima glaucos Ciccotto, Konings &amp; Stauffer, 2011, new species" docType="treatment" docVersion="10" lastPageNumber="11" masterDocId="FF82EC520D7483669D78FF83FFCBFFFE" masterDocTitle="Descriptions of five new species in the genus Metriaclima (Teleostei: Cichlidae) from Lake Malaŵi, Africa" masterLastPageNumber="25" masterPageNumber="1" pageNumber="9" updateTime="1698237790047" updateUser="plazi">
<mods:mods id="180CF9312EAB160AF135CDE3D521DE5A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B2B09EC6C845C9CB0E6932CE4F175755">
<mods:title id="C2377252655080C751B8311AECE0C7A4">Descriptions of five new species in the genus Metriaclima (Teleostei: Cichlidae) from Lake Malaŵi, Africa</mods:title>
</mods:titleInfo>
<mods:name id="26CABE988CE64929422EA3A72F376892" type="personal">
<mods:role id="03A1E6CAB4BFA8B2728EBAB67A6FDCC0">
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<mods:namePart id="0439A056E4404ABCE3FD65D1D1E4604B">Ciccotto, Patrick J.</mods:namePart>
</mods:name>
<mods:name id="E5AE77C9B07F4355D0AA527C966BE1C8" type="personal">
<mods:role id="18490E6F0C2B0F3D6F7EC9F9BB98F428">
<mods:roleTerm id="126DBCCA0AA84FAD3104268112B38063">Author</mods:roleTerm>
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<mods:namePart id="83CA4A94DA35A161CD4C91737574AA03">Konings, Adrianus</mods:namePart>
</mods:name>
<mods:name id="37A130D4D4B281F4CC02AFDD1C89DE9E" type="personal">
<mods:role id="F3877FD4B1A2A56619064303DB6D7CF9">
<mods:roleTerm id="4DEE4A2E75585A7ABC7521497215BAB1">Author</mods:roleTerm>
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<mods:namePart id="C91A8E9F8C750ECBF2624833EC36C563">Stauffer, Jay R.</mods:namePart>
</mods:name>
<mods:typeOfResource id="B85E7A557E1B7DB06BB96C6E753AFA91">text</mods:typeOfResource>
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<mods:title id="67BACA481BD6583E822FB6CB40563FF5">Zootaxa</mods:title>
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<mods:part id="9CBDA641A10EFC144CACF7C2ED80EA48">
<mods:date id="B34A04FED217C58BE009DCF7C7EAC929">2011</mods:date>
<mods:detail id="DDBBDD9F501CBC88E28617610AF60AF8" type="volume">
<mods:number id="1572326D682C8E4D60519475E197C47D">2738</mods:number>
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<mods:classification id="361F3ACFF0B2D5FF548EF5D5E2591434">journal article</mods:classification>
<mods:identifier id="86EFDF61D85F1EA4A3E5E9CCC8284598" type="DOI">10.5281/zenodo.204914</mods:identifier>
<mods:identifier id="4CBEACB62DFC0129AAF793236D139B28" type="GBIF-Dataset">035006f1-9010-4f0b-a5fb-83fed142d245</mods:identifier>
<mods:identifier id="9E1584FF77885CE213070D818F0C7B37" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="5C884EE031736AEF7F4D80EE81A2C910" type="Zenodo-Dep">204914</mods:identifier>
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<treatment id="03BB942A0D7C836C9DEFFC73FD6EFA2A" ID-DOI="http://doi.org/10.5281/zenodo.6192252" ID-GBIF-Taxon="119385414" ID-Zenodo-Dep="6192252" LSID="urn:lsid:plazi:treatment:03BB942A0D7C836C9DEFFC73FD6EFA2A" httpUri="http://treatment.plazi.org/id/03BB942A0D7C836C9DEFFC73FD6EFA2A" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9">
<subSubSection id="C30876B70D7C836E9DEFFC73FEA4FBD4" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="8BAD253C0D7C836E9DEFFC73FDE6FBF4" blockId="8.[151,557,1008,1066]" box="[151,557,1008,1034]" pageId="8" pageNumber="9">
<heading id="D0E592500D7C836E9DEFFC73FDE6FBF4" bold="true" box="[151,557,1008,1034]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
<emphasis id="B966F92E0D7C836E9DEFFC73FDE6FBF4" bold="true" box="[151,557,1008,1034]" pageId="8" pageNumber="9">
<emphasis id="B966F92E0D7C836E9DEFFC73FE5DFBF4" bold="true" box="[151,406,1008,1034]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="4C125EBF0D7C836E9DEFFC73FE5BFBF4" ID-CoL="42HW5" box="[151,400,1008,1034]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="glaucos" status="sp. nov.">Metriaclima glaucos</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A25544550D7C836E9CE6FC73FDE6FBF4" box="[414,557,1008,1034]" pageId="8" pageNumber="9" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAD253C0D7C836E9DEFFB91FEA4FBD4" blockId="8.[151,557,1008,1066]" box="[151,367,1042,1066]" pageId="8" pageNumber="9">
<figureCitation id="132939B90D7C836E9DEFFB91FF25FBD4" box="[151,238,1042,1066]" captionStart="FIGURE 4. A" captionStartId="10.[151,250,1052,1075]" captionTargetBox="[276,1311,195,1030]" captionTargetId="figure@10.[276,1312,195,1031]" captionTargetPageId="10" captionText="FIGURE 4. A. Metriaclima glaucos, holotype, PSU 4487, 69.2 mm SL; Mozambique: Lake MalaŴi: Cobwé. B. M. glaucos, male in habitat at Cobwé, Mozambique; approx. 65 mm SL; not preserved. C. M. glaucos, female in habitat at Cobwé, Mozambique; approx. 55 mm SL; not preserved." httpUri="https://zenodo.org/record/204918/files/figure.png" pageId="8" pageNumber="9">Fig. 4A</figureCitation>
–C;
<tableCitation id="C69010870D7C836E9C64FB91FEA4FBD4" box="[284,367,1042,1066]" captionStart="TABLE 3" captionStartId="9.[151,235,154,175]" captionTargetBox="[151,1350,286,1962]" captionTargetPageId="9" captionText="TABLE 3. Morphometric and meristic values of Metriaclima glaucos from Cobwé. Mean and range include holotype." httpUri="http://table.plazi.org/id/DF6D75B40D7D836F9DEFFF19FA8DFF51" pageId="8" pageNumber="9" tableUuid="DF6D75B40D7D836F9DEFFF19FA8DFF51">Table 3</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30876B70D7C836E9DEFFBD8FD50FB8E" box="[151,667,1114,1137]" pageId="8" pageNumber="9" type="description">
<paragraph id="8BAD253C0D7C836E9DEFFBD8FD50FB8E" blockId="8.[151,667,1114,1137]" box="[151,667,1114,1137]" pageId="8" pageNumber="9">
<treatmentCitationGroup id="AB0202120D7C836E9DEFFBD8FD50FB8E" box="[151,667,1114,1137]" pageId="8" pageNumber="9">
<taxonomicName id="4C125EBF0D7C836E9DEFFBD8FEDFFB8F" box="[151,276,1115,1137]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">
<emphasis id="B966F92E0D7C836E9DEFFBD8FEDFFB8F" box="[151,276,1115,1137]" italics="true" pageId="8" pageNumber="9">Metriaclima</emphasis>
</taxonomicName>
sp. &quot;
<taxonomicName id="4C125EBF0D7C836E9C3EFBD8FE42FB8E" box="[326,393,1115,1136]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="aurora">aurora</taxonomicName>
blue&quot;:
<treatmentCitation id="0AB3032D0D7C836E9CAAFBD9FD5CFB8E" author="Konings" box="[466,663,1114,1136]" page="155" pageId="8" pageNumber="9" year="2001">
<bibRefCitation id="EF8358CD0D7C836E9CAAFBD9FDAFFB8E" author="Konings" box="[466,612,1114,1136]" pageId="8" pageNumber="25" refString="Konings, A. (2001) Mala w i cichlids in their natural habitat, Third edition. Cichlid Press, El Paso, Texas, 352 pp." type="book" year="2001">Konings 2001</bibRefCitation>
: 155
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30876B70D7C836E9DEFFB1EFAFAFB03" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="8BAD253C0D7C836E9DEFFB1EFE2BFB27" blockId="8.[151,1437,1181,2034]" pageId="8" pageNumber="9">
<emphasis id="B966F92E0D7C836E9DEFFB1EFEC0FB4B" bold="true" box="[151,267,1181,1205]" pageId="8" pageNumber="9">
<typeStatus id="54A99B9E0D7C836E9DEFFB1EFECDFB4B" box="[151,262,1181,1205]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED03BDF90D7C836E9C69FB1EFE8FFB4B" box="[273,324,1181,1205]" httpUri="http://grbio.org/cool/5mg9-yyc1" name="Portland State University, Vertebrate Biology Museum" pageId="8" pageNumber="9">PSU</collectionCode>
4487,
<quantity id="4CEA88D90D7C836E9CE9FB1EFE3FFB4B" box="[401,500,1181,1205]" metricMagnitude="-2" metricUnit="m" metricValue="6.92" pageId="8" pageNumber="9" unit="mm" value="69.2">69.2 mm</quantity>
SL;
<collectingCountry id="F30565AC0D7C836E9F52FB1EFD09FB4B" box="[554,706,1181,1205]" name="Mozambique" pageId="8" pageNumber="9">Mozambique</collectingCountry>
: Lake
<collectingCountry id="F30565AC0D7C836E9E73FB1EFCAEFB48" box="[779,869,1181,1206]" name="Malawi" pageId="8" pageNumber="9">Malaŵi</collectingCountry>
: Cobwé,
<geoCoordinate id="EE2643FB0D7C836E9EB3FB1EFB9EFB4B" box="[971,1109,1181,1205]" direction="south" orientation="latitude" pageId="8" pageNumber="9" precision="1" value="-12.137383">12° 8.243' S</geoCoordinate>
,
<geoCoordinate id="EE2643FB0D7C836E9919FB1EFB30FB4B" box="[1121,1275,1181,1205]" direction="east" orientation="longitude" pageId="8" pageNumber="9" precision="1" value="34.756516">34° 45.391' E</geoCoordinate>
; A.F. Konings &amp; J.R. Stauffer,
<date id="FFAC03FC0D7C836E9C34FB42FE10FB27" box="[332,475,1217,1241]" pageId="8" pageNumber="9" value="2002-02-18">18 Feb 2002</date>
.
</paragraph>
<paragraph id="8BAD253C0D7C836E9DBFFB66FAFAFB03" blockId="8.[151,1437,1181,2034]" box="[199,1329,1253,1278]" pageId="8" pageNumber="9">
<emphasis id="B966F92E0D7C836E9DBFFB66FE8DFB03" bold="true" box="[199,326,1253,1277]" pageId="8" pageNumber="9">
<typeStatus id="54A99B9E0D7C836E9DBFFB66FE8AFB03" box="[199,321,1253,1277]" pageId="8" pageNumber="9" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="ED03BDF90D7C836E9C34FB66FEB4FB03" box="[332,383,1253,1277]" httpUri="http://grbio.org/cool/5mg9-yyc1" name="Portland State University, Vertebrate Biology Museum" pageId="8" pageNumber="9">PSU</collectionCode>
4488, 16;
<collectionCode id="ED03BDF90D7C836E9C80FB66FD80FB03" box="[504,587,1253,1277]" httpUri="http://grbio.org/cool/ymef-zyvh" name="Univeristy of Malawi" pageId="8" pageNumber="9">UMBC</collectionCode>
15, 1;
<collectionCode id="ED03BDF90D7C836E9FE0FB65FD25FB00" LSID="urn:lsid:biocol.org:col:34925" box="[664,750,1254,1278]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="8" pageNumber="9">AMNH</collectionCode>
246003, 2; 53.0–
<quantity id="4CEA88D90D7C836E9ECAFB66FBDDFB03" box="[946,1046,1253,1277]" metricMagnitude="-2" metricUnit="m" metricValue="6.74" pageId="8" pageNumber="9" unit="mm" value="67.4">67.4 mm</quantity>
SL; data as for
<typeStatus id="54A99B9E0D7C836E99BFFB66FAE6FB03" box="[1223,1325,1253,1277]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C30876B70D7C836E9DBFFA8AFEDBF99B" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="8BAD253C0D7C836E9DBFFA8AFEDBF99B" blockId="8.[151,1437,1181,2034]" pageId="8" pageNumber="9">
<emphasis id="B966F92E0D7C836E9DBFFA8AFE8AFADC" bold="true" box="[199,321,1289,1314]" pageId="8" pageNumber="9">Diagnosis.</emphasis>
A moderately-sloped head and bicuspid teeth in the outer row of the jaws place this species in
<taxonomicName id="4C125EBF0D7C836E9DEFFAADFEEAFAB8" box="[151,289,1326,1350]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">
<emphasis id="B966F92E0D7C836E9DEFFAADFEEAFAB8" box="[151,289,1326,1350]" italics="true" pageId="8" pageNumber="9">Metriaclima</emphasis>
</taxonomicName>
. The absence of a black submarginal band in the dorsal fin in conjunction with a yellow gular region and branchiostegal membranes, and the light-brown colored females that have yellow fin margins distinguish
<taxonomicName id="4C125EBF0D7C836E9805FAD1FF25FA70" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="glaucos">
<emphasis id="B966F92E0D7C836E9805FAD1FF25FA70" italics="true" pageId="8" pageNumber="9">M. glaucos</emphasis>
</taxonomicName>
from other members of
<taxonomicName id="4C125EBF0D7C836E9F70FAF5FD59FA70" box="[520,658,1398,1422]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">
<emphasis id="B966F92E0D7C836E9F70FAF5FD59FA70" box="[520,658,1398,1422]" italics="true" pageId="8" pageNumber="9">Metriaclima</emphasis>
</taxonomicName>
, except
<taxonomicName id="4C125EBF0D7C836E9F8CFAF4FC7AFA70" box="[756,945,1398,1422]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="chrysomallos">
<emphasis id="B966F92E0D7C836E9F8CFAF4FC7AFA70" box="[756,945,1398,1422]" italics="true" pageId="8" pageNumber="9">M. chrysomallos</emphasis>
</taxonomicName>
and
<taxonomicName id="4C125EBF0D7C836E9E92FAF4FBADFA70" box="[1002,1126,1398,1422]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="benetos">
<emphasis id="B966F92E0D7C836E9E92FAF4FBADFA70" box="[1002,1126,1398,1422]" italics="true" pageId="8" pageNumber="9">M. benetos</emphasis>
</taxonomicName>
. There are generally fewer teeth in the outer row of the lower left jaw (mode 16, range 14–19) of
<taxonomicName id="4C125EBF0D7C836E9EBBFA19FB8AFA4F" box="[963,1089,1433,1457]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="glaucos">
<emphasis id="B966F92E0D7C836E9EBBFA19FB8AFA4F" box="[963,1089,1433,1457]" italics="true" pageId="8" pageNumber="9">M. glaucos</emphasis>
</taxonomicName>
compared to
<taxonomicName id="4C125EBF0D7C836E99A7FA19FA50FA4F" box="[1247,1435,1433,1457]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="chrysomallos">
<emphasis id="B966F92E0D7C836E99A7FA19FA50FA4F" box="[1247,1435,1433,1457]" italics="true" pageId="8" pageNumber="9">M. chrysomallos</emphasis>
</taxonomicName>
(mode 22, range 15–25) and more compared to
<taxonomicName id="4C125EBF0D7C836E9FC0FA3CFCFEFA28" box="[696,821,1470,1494]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="benetos">
<emphasis id="B966F92E0D7C836E9FC0FA3CFCFEFA28" box="[696,821,1470,1494]" italics="true" pageId="8" pageNumber="9">M. benetos</emphasis>
</taxonomicName>
(mode 12, range 9–13). On the basis of color pattern, females of
<taxonomicName id="4C125EBF0D7C836E9C6BFA61FE44FA07" box="[275,399,1505,1529]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="glaucos">
<emphasis id="B966F92E0D7C836E9C6BFA61FE44FA07" box="[275,399,1505,1529]" italics="true" pageId="8" pageNumber="9">M. glaucos</emphasis>
</taxonomicName>
cannot reliably be distinguished from those of most other members of the
<taxonomicName id="4C125EBF0D7C836E99AAFA61FAEAFA04" box="[1234,1313,1506,1530]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="aurora">Aurora</taxonomicName>
group, but they differ from those of
<taxonomicName id="4C125EBF0D7C836E9CC2F984FDF3F9E0" box="[442,568,1542,1566]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="xanthos" status="sp. nov.">
<emphasis id="B966F92E0D7C836E9CC2F984FDF3F9E0" box="[442,568,1542,1566]" italics="true" pageId="8" pageNumber="9">M. xanthos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25544550D7C836E9F39F985FD01F9E3" box="[577,714,1541,1565]" pageId="8" pageNumber="9" rank="species">new species</taxonomicNameLabel>
by a shorter lower jaw (mean 30.8, range 28.8–32.8 % HL vs. mean 35.4, range 34.6–36.7 % HL) and from those of
<taxonomicName id="4C125EBF0D7C836E9F80F9A9FCA1F9BF" box="[760,874,1578,1601]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="aurora">
<emphasis id="B966F92E0D7C836E9F80F9A9FCA1F9BF" box="[760,874,1578,1601]" italics="true" pageId="8" pageNumber="9">M. aurora</emphasis>
</taxonomicName>
by 8–11 vertical bars below the dorsal fin vs.
<quantity id="4CEA88D90D7C836E9809F9AAFA57F9BF" box="[1393,1436,1577,1601]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="8" pageNumber="9" unit="in" value="6.0">6 in</quantity>
<taxonomicName id="4C125EBF0D7C836E9DEFF9CCFEC2F998" box="[151,265,1615,1638]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="aurora">
<emphasis id="B966F92E0D7C836E9DEFF9CCFEC2F998" box="[151,265,1615,1638]" italics="true" pageId="8" pageNumber="9">M. aurora</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C30876B70D7C836C9DBFF9F2FE0CFABA" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9" type="description">
<paragraph id="8BAD253C0D7C836E9DBFF9F2FEF0F89F" blockId="8.[151,1437,1181,2034]" pageId="8" pageNumber="9">
<emphasis id="B966F92E0D7C836E9DBFF9F2FE92F974" bold="true" box="[199,345,1649,1674]" pageId="8" pageNumber="9">Description.</emphasis>
Morphometric and meristic data in
<tableCitation id="C69010870D7C836E9E78F9F2FC92F977" box="[768,857,1649,1673]" captionStart="TABLE 3" captionStartId="9.[151,235,154,175]" captionTargetBox="[151,1350,286,1962]" captionTargetPageId="9" captionText="TABLE 3. Morphometric and meristic values of Metriaclima glaucos from Cobwé. Mean and range include holotype." httpUri="http://table.plazi.org/id/DF6D75B40D7D836F9DEFFF19FA8DFF51" pageId="8" pageNumber="9" tableUuid="DF6D75B40D7D836F9DEFFF19FA8DFF51">Table 3</tableCitation>
. Dorsal snout profile slightly concave to straight; mouth cleft slightly downward to horizontal; jaws isognathous. Teeth on dentary in 2-3 rows, on premaxilla in 2 or 3 rows; outer row teeth typically bicuspid anteriorly and unicuspid posteriorly, middle rows when present tricuspid, innermost row unicuspid; lower pharyngeal jaw with numerous slender teeth with teeth in posterior row slightly larger. Portion of upper dental arcade normally exposed when mouth closed. Tips of teeth in premaxilla and dentary in V-shaped line with anteriormost in upper and lower jaw furthest apart and not touching in closed mouth. Lateral scales ctenoid.
</paragraph>
<paragraph id="8BAD253C0D7C836E9DBFF8EEFABEF80F" blockId="8.[151,1437,1181,2034]" pageId="8" pageNumber="9">Breeding males light blue/gray laterally with 7 dark gray bars below dorsal fin and white belly. Head light blue/gray with preorbital, cheek, and operculum light blue; darker blue opercular spot; yellow gular region and brachiostegals. Dorsal fin blue with white lappets. Caudal-fin rays gray with white/blue membranes. Anal fin gray with 1–4 yellow ocelli. Pelvic fin with white leading edge, first ray gray and remainder clear. Pectoral fin clear.</paragraph>
<caption id="DF6D75B40D7D836F9DEFFF19FA8DFF51" ID-Table-UUID="DF6D75B40D7D836F9DEFFF19FA8DFF51" box="[151,1350,153,175]" httpUri="http://table.plazi.org/id/DF6D75B40D7D836F9DEFFF19FA8DFF51" pageId="9" pageNumber="10" targetBox="[151,1350,286,1962]" targetIsTable="true" targetPageId="9">
<paragraph id="8BAD253C0D7D836F9DEFFF19FA8DFF51" blockId="9.[151,1350,153,266]" box="[151,1350,153,175]" pageId="9" pageNumber="10">
<emphasis id="B966F92E0D7D836F9DEFFF19FECFFF51" bold="true" box="[151,260,154,175]" pageId="9" pageNumber="10">TABLE 3.</emphasis>
Morphometric and meristic values of
<taxonomicName id="4C125EBF0D7D836F9FF0FF1AFC90FF51" box="[648,859,153,175]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="glaucos">
<emphasis id="B966F92E0D7D836F9FF0FF1AFC90FF51" box="[648,859,153,175]" italics="true" pageId="9" pageNumber="10">Metriaclima glaucos</emphasis>
</taxonomicName>
from Cobwé. Mean and range include holotype.
</paragraph>
</caption>
<paragraph id="8BAD253C0D7D836F9F84FF4FFC04FF1C" blockId="9.[151,1350,153,266]" box="[764,975,204,226]" pageId="9" pageNumber="10">
<taxonomicName id="4C125EBF0D7D836F9F84FF4FFC04FF1C" box="[764,975,204,226]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="glaucos">
<emphasis id="B966F92E0D7D836F9F84FF4FFC04FF1C" box="[764,975,204,226]" italics="true" pageId="9" pageNumber="10">Metriaclima glaucos</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BAD253C0D7D836F9F84FF76FCFAFEF4" blockId="9.[151,1350,153,266]" box="[764,817,245,266]" pageId="9" pageNumber="10">n=20</paragraph>
<paragraph id="8BAD253C0D7D836F9F84FE9DFB75F857" pageId="9" pageNumber="10">
<table id="F912D79C0D7D7C999DEFFE9DFA8DF854" box="[151,1350,286,1962]" gridcols="4" gridrows="11" pageId="9" pageNumber="10">
<tr id="3522277E0D7D7C999DEFFE9DFA8DFECA" box="[151,1350,286,308]" gridrow="0" pageId="9" pageNumber="10" rowspan-0="1">
<th id="76F34E020D7D7C999F84FE9DFC9EFECA" box="[764,853,286,308]" gridcol="1" gridrow="0" pageId="9" pageNumber="10">holotype</th>
<th id="76F34E020D7D7C999E95FE9DFBEEFECA" box="[1005,1061,286,308]" gridcol="2" gridrow="0" pageId="9" pageNumber="10">mean</th>
<th id="76F34E020D7D7C9999E0FE9DFA8DFECA" box="[1176,1350,286,308]" gridcol="3" gridrow="0" pageId="9" pageNumber="10">range</th>
</tr>
<tr id="3522277E0D7D7C999DEFFECBFA8DFE27" box="[151,1350,328,473]" gridrow="1" pageId="9" pageNumber="10">
<th id="76F34E020D7D7C999DEFFECBFDC2FE27" box="[151,521,328,473]" gridcol="0" gridrow="1" pageId="9" pageNumber="10">Standard length, mm Head length, mm Percentage of standard length Head length</th>
<td id="76F34E020D7D7C999F84FECBFC9EFE27" box="[764,853,328,473]" gridcol="1" gridrow="1" pageId="9" pageNumber="10">69.2 21.0 30.3</td>
<td id="76F34E020D7D7C999E95FECBFBEEFE27" box="[1005,1061,328,473]" gridcol="2" gridrow="1" pageId="9" pageNumber="10">61.4 18.7 30.5</td>
<td id="76F34E020D7D7C9999E0FECBFA8DFE27" box="[1176,1350,328,473]" gridcol="3" gridrow="1" pageId="9" pageNumber="10">53.0–69.2 16.1–21.0 29.4–31.9</td>
</tr>
<tr id="3522277E0D7D7C999DEFFE6EFA8DFDD2" box="[151,1350,493,556]" gridrow="2" pageId="9" pageNumber="10">
<th id="76F34E020D7D7C999DEFFE6EFDC2FDD2" box="[151,521,493,556]" gridcol="0" gridrow="2" pageId="9" pageNumber="10">Snout to dorsal Snout to pelvic</th>
<td id="76F34E020D7D7C999F84FE6EFC9EFDD2" box="[764,853,493,556]" gridcol="1" gridrow="2" pageId="9" pageNumber="10">32.7 37.4</td>
<td id="76F34E020D7D7C999E95FE6EFBEEFDD2" box="[1005,1061,493,556]" gridcol="2" gridrow="2" pageId="9" pageNumber="10">33.4 38.2</td>
<td id="76F34E020D7D7C9999E0FE6EFA8DFDD2" box="[1176,1350,493,556]" gridcol="3" gridrow="2" pageId="9" pageNumber="10">31.7–34.6 35.9–40.4</td>
</tr>
<tr id="3522277E0D7D7C999DEFFDC3FA8DFD2F" box="[151,1350,576,721]" gridrow="3" pageId="9" pageNumber="10">
<th id="76F34E020D7D7C999DEFFDC3FDC2FD2F" box="[151,521,576,721]" gridcol="0" gridrow="3" pageId="9" pageNumber="10">Greatest body depth Caudal peduncle length Least caudal peduncle depth Dorsal-fin base length</th>
<td id="76F34E020D7D7C999F84FDC3FC9EFD2F" box="[764,853,576,721]" gridcol="1" gridrow="3" pageId="9" pageNumber="10">31.9 14.6 10.9 64.4</td>
<td id="76F34E020D7D7C999E95FDC3FBEEFD2F" box="[1005,1061,576,721]" gridcol="2" gridrow="3" pageId="9" pageNumber="10">32.8 14.2 11.0 61.9</td>
<td id="76F34E020D7D7C9999E0FDC3FA8DFD2F" box="[1176,1350,576,721]" gridcol="3" gridrow="3" pageId="9" pageNumber="10">31.8–34.0 12.5–15.5 9.9–12.3 59.8–64.4</td>
</tr>
<tr id="3522277E0D7D7C999DEFFD66FA8DFCDA" box="[151,1350,741,804]" gridrow="4" pageId="9" pageNumber="10">
<th id="76F34E020D7D7C999DEFFD66FDC2FCDA" box="[151,521,741,804]" gridcol="0" gridrow="4" pageId="9" pageNumber="10">Anterior dorsal to anterior anal Anterior dorsal to posterior anal</th>
<td id="76F34E020D7D7C999F84FD66FC9EFCDA" box="[764,853,741,804]" gridcol="1" gridrow="4" pageId="9" pageNumber="10">52.5 66.3</td>
<td id="76F34E020D7D7C999E95FD66FBEEFCDA" box="[1005,1061,741,804]" gridcol="2" gridrow="4" pageId="9" pageNumber="10">53.6 64.4</td>
<td id="76F34E020D7D7C9999E0FD66FA8DFCDA" box="[1176,1350,741,804]" gridcol="3" gridrow="4" pageId="9" pageNumber="10">50.8–55.8 62.9–66.5</td>
</tr>
<tr id="3522277E0D7D7C999DEFFCBBFA8DFBE2" box="[151,1350,824,1052]" gridrow="5" pageId="9" pageNumber="10">
<th id="76F34E020D7D7C999DEFFCBBFDC2FBE2" box="[151,521,824,1052]" gridcol="0" gridrow="5" pageId="9" pageNumber="10">Posterior dorsal to anterior anal Posterior dorsal to posterior anal Anterior dorsal to pelvic-fin origin Posterior dorsal to pelvic-fin origin Percentage of head length Horizontal eye diameter</th>
<td id="76F34E020D7D7C999F84FCBBFC9EFBE2" box="[764,853,824,1052]" gridcol="1" gridrow="5" pageId="9" pageNumber="10">29.6 14.9 36.6 59.1 30.8</td>
<td id="76F34E020D7D7C999E95FCBBFBEEFBE2" box="[1005,1061,824,1052]" gridcol="2" gridrow="5" pageId="9" pageNumber="10">29.3 15.1 37.2 58.7 33.7</td>
<td id="76F34E020D7D7C9999E0FCBBFA8DFBE2" box="[1176,1350,824,1052]" gridcol="3" gridrow="5" pageId="9" pageNumber="10">27.1–30.9 14.4–16.5 35.2–39.5 55.6–60.7 30.7–36.6</td>
</tr>
<tr id="3522277E0D7D7C999DEFFBB3FA8DFB3F" box="[151,1350,1072,1217]" gridrow="6" pageId="9" pageNumber="10">
<th id="76F34E020D7D7C999DEFFBB3FDC2FB3F" box="[151,521,1072,1217]" gridcol="0" gridrow="6" pageId="9" pageNumber="10">Vertical eye diameter Snout length Postorbital head length Preorbital depth</th>
<td id="76F34E020D7D7C999F84FBB3FC9EFB3F" box="[764,853,1072,1217]" gridcol="1" gridrow="6" pageId="9" pageNumber="10">32.0 28.1 42.0 21.5</td>
<td id="76F34E020D7D7C999E95FBB3FBEEFB3F" box="[1005,1061,1072,1217]" gridcol="2" gridrow="6" pageId="9" pageNumber="10">33.7 28.9 41.8 21.9</td>
<td id="76F34E020D7D7C9999E0FBB3FA8DFB3F" box="[1176,1350,1072,1217]" gridcol="3" gridrow="6" pageId="9" pageNumber="10">31.0–36.9 25.4–30.8 38.5–43.3 19.2–23.6</td>
</tr>
<tr id="3522277E0D7D7C999DEFFB56FA8DFAEA" box="[151,1350,1237,1300]" gridrow="7" pageId="9" pageNumber="10">
<th id="76F34E020D7D7C999DEFFB56FDC2FAEA" box="[151,521,1237,1300]" gridcol="0" gridrow="7" pageId="9" pageNumber="10">Lower-jaw length Cheek depth</th>
<td id="76F34E020D7D7C999F84FB56FC9EFAEA" box="[764,853,1237,1300]" gridcol="1" gridrow="7" pageId="9" pageNumber="10">29.3 25.0</td>
<td id="76F34E020D7D7C999E95FB56FBEEFAEA" box="[1005,1061,1237,1300]" gridcol="2" gridrow="7" pageId="9" pageNumber="10">30.8 23.8</td>
<td id="76F34E020D7D7C9999E0FB56FA8DFAEA" box="[1176,1350,1237,1300]" gridcol="3" gridrow="7" pageId="9" pageNumber="10">28.8–32.8 21.4–26.7</td>
</tr>
<tr id="3522277E0D7D7C999DEFFAABFA8DF9F2" box="[151,1350,1320,1548]" gridrow="8" pageId="9" pageNumber="10">
<th id="76F34E020D7D7C999DEFFAABFDC2F9F2" box="[151,521,1320,1548]" gridcol="0" gridrow="8" pageId="9" pageNumber="10">Head depth Meristics Dorsal-fin spines Dorsal-fin rays Anal-fin spines Anal-fin rays</th>
<td id="76F34E020D7D7C999F84FAABFC9EF9F2" box="[764,853,1320,1548]" gridcol="1" gridrow="8" pageId="9" pageNumber="10">92.5 18 8 3 8</td>
<td id="76F34E020D7D7C999E95FAABFBEEF9F2" box="[1005,1061,1320,1548]" gridcol="2" gridrow="8" pageId="9" pageNumber="10">91.0 mode 18 9 3 8</td>
<td id="76F34E020D7D7C9999E0FAABFA8DF9F2" box="[1176,1350,1320,1548]" gridcol="3" gridrow="8" pageId="9" pageNumber="10">86.5–95.9 range 17–19 8–10 3–3 7–8</td>
</tr>
<tr id="3522277E0D7D7C999DEFF9A3FA8DF8FA" box="[151,1350,1568,1796]" gridrow="9" pageId="9" pageNumber="10">
<th id="76F34E020D7D7C999DEFF9A3FDC2F8FA" box="[151,521,1568,1796]" gridcol="0" gridrow="9" pageId="9" pageNumber="10">Pelvic-fin rays Pectoral-fin rays Lateral line scales Pored scales posterior to lateral line Cheek scales Gill rakers on first ceratobranchial</th>
<td id="76F34E020D7D7C999F84F9A3FC9EF8FA" box="[764,853,1568,1796]" gridcol="1" gridrow="9" pageId="9" pageNumber="10">5 14 30 1 6 11</td>
<td id="76F34E020D7D7C999E95F9A3FBEEF8FA" box="[1005,1061,1568,1796]" gridcol="2" gridrow="9" pageId="9" pageNumber="10">5 14 30 2 6 11</td>
<td id="76F34E020D7D7C9999E0F9A3FA8DF8FA" box="[1176,1350,1568,1796]" gridcol="3" gridrow="9" pageId="9" pageNumber="10">4–5 14–15 29–32 0–2 4–7 10–11</td>
</tr>
<tr id="3522277E0D7D7C999DEFF89BFA8DF854" box="[151,1350,1816,1962]" gridrow="10" pageId="9" pageNumber="10">
<th id="76F34E020D7D7C999DEFF89BFDC2F854" box="[151,521,1816,1962]" gridcol="0" gridrow="10" pageId="9" pageNumber="10">Gill rakers on first epibranchial Teeth in outer row of left lower jaw Teeth rows on upper jaw Teeth rows on lower jaw</th>
<td id="76F34E020D7D7C999F84F89BFC9EF854" box="[764,853,1816,1962]" gridcol="1" gridrow="10" pageId="9" pageNumber="10">3 16 3 3</td>
<td id="76F34E020D7D7C999E95F89BFBEEF854" box="[1005,1061,1816,1962]" gridcol="2" gridrow="10" pageId="9" pageNumber="10">3 16 3 3</td>
<td id="76F34E020D7D7C9999E0F89BFA8DF854" box="[1176,1350,1816,1962]" gridcol="3" gridrow="10" pageId="9" pageNumber="10">2–4 14–19 2–3 2–3</td>
</tr>
</table>
</paragraph>
<caption id="DF6D75B40D7E836C9DEFFB9FFDF3FB8E" httpUri="https://zenodo.org/record/204918/files/figure.png" pageId="10" pageNumber="11" targetBox="[276,1311,195,1030]" targetPageId="10">
<paragraph id="8BAD253C0D7E836C9DEFFB9FFDF3FB8E" blockId="10.[151,1436,1051,1136]" pageId="10" pageNumber="11">
<emphasis id="B966F92E0D7E836C9DEFFB9FFEFFFBCF" bold="true" box="[151,308,1052,1075]" pageId="10" pageNumber="11">FIGURE 4. A.</emphasis>
<taxonomicName id="4C125EBF0D7E836C9C43FB9FFDC5FBCC" box="[315,526,1052,1074]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="glaucos">
<emphasis id="B966F92E0D7E836C9C43FB9FFDC5FBCC" box="[315,526,1052,1074]" italics="true" pageId="10" pageNumber="11">Metriaclima glaucos</emphasis>
</taxonomicName>
, holotype, PSU 4487, 69.2 mm SL; Mozambique: Lake MalaŴi: Cobwé.
<emphasis id="B966F92E0D7E836C987DFB9EFAD6FBCC" bold="true" box="[1285,1309,1053,1074]" pageId="10" pageNumber="11">B.</emphasis>
<taxonomicName id="4C125EBF0D7E836C985CFB9EFA5EFBCC" box="[1316,1429,1052,1074]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="glaucos">
<emphasis id="B966F92E0D7E836C985CFB9EFA5EFBCC" box="[1316,1429,1052,1074]" italics="true" pageId="10" pageNumber="11">M. glaucos</emphasis>
</taxonomicName>
, male in habitat at Cobwé, Mozambique; approx. 65 mm SL; not preserved.
<emphasis id="B966F92E0D7E836C9EE8FBB9FC63FBAF" bold="true" box="[912,936,1082,1105]" pageId="10" pageNumber="11">C.</emphasis>
<taxonomicName id="4C125EBF0D7E836C9ED6FBBFFBD5FBAF" box="[942,1054,1083,1105]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="glaucos">
<emphasis id="B966F92E0D7E836C9ED6FBBFFBD5FBAF" box="[942,1054,1083,1105]" italics="true" pageId="10" pageNumber="11">M. glaucos</emphasis>
</taxonomicName>
, female in habitat at Cobwé, Mozambique; approx. 55 mm SL; not preserved.
</paragraph>
</caption>
<paragraph id="8BAD253C0D7E836C9DBFFB1FFE0CFABA" blockId="10.[151,1436,1180,1492]" pageId="10" pageNumber="11">Lateral coloration of females gray dorsally and white ventrally. Head gray with gray cheeks, blue/gray operculum with green highlights, gray/black opercular spot, and white gular region. Dorsal fin gray with yellow/orange lappets. Caudal-fin rays gray with clear membranes and faint yellow/orange spots. Anal fin gray proximally and yellow distally without ocelli. Pelvic fin with white leading edge and black submarginal band, membranes yellow to clear. Pectoral fins clear.</paragraph>
</subSubSection>
<subSubSection id="C30876B70D7E836C9DBFFAD3FE86FA72" pageId="10" pageNumber="11" type="distribution">
<paragraph id="8BAD253C0D7E836C9DBFFAD3FE86FA72" blockId="10.[151,1436,1180,1492]" pageId="10" pageNumber="11">
<emphasis id="B966F92E0D7E836C9DBFFAD3FEABFA97" bold="true" box="[199,352,1360,1385]" pageId="10" pageNumber="11">Distribution.</emphasis>
<taxonomicName id="4C125EBF0D7E836C9C11FAD3FD9FFA96" box="[361,596,1360,1384]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="glaucos">
<emphasis id="B966F92E0D7E836C9C11FAD3FD9FFA96" box="[361,596,1360,1384]" italics="true" pageId="10" pageNumber="11">Metriaclima glaucos</emphasis>
</taxonomicName>
was collected at Cobwé,
<collectingCountry id="F30565AC0D7E836C9E06FAD3FBDEFA96" box="[894,1045,1360,1384]" name="Mozambique" pageId="10" pageNumber="11">Mozambique</collectingCountry>
, and is only known from the
<typeStatus id="54A99B9E0D7E836C9814FAD2FA57FA97" box="[1388,1436,1361,1385]" pageId="10" pageNumber="11">type</typeStatus>
locality (
<figureCitation id="132939B90D7E836C9D82FAF7FEF0FA72" box="[250,315,1396,1420]" captionStart="FIGURE 1" captionStartId="1.[151,250,1266,1289]" captionTargetBox="[154,1435,196,1245]" captionTargetId="figure@1.[151,1436,193,1245]" captionTargetPageId="1" captionText="FIGURE 1. Map of Lake MalaŴi showing the locations of species discussed throughout the text." httpUri="https://zenodo.org/record/204915/files/figure.png" pageId="10" pageNumber="11">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30876B70D7E836C9DBFFA1AFD6EFA2A" pageId="10" pageNumber="11" type="etymology">
<paragraph id="8BAD253C0D7E836C9DBFFA1AFD6EFA2A" blockId="10.[151,1436,1180,1492]" pageId="10" pageNumber="11">
<emphasis id="B966F92E0D7E836C9DBFFA1AFE85FA4F" bold="true" box="[199,334,1433,1457]" pageId="10" pageNumber="11">Etymology.</emphasis>
The specific epithet,
<emphasis id="B966F92E0D7E836C9F3BFA1BFD6AFA4E" box="[579,673,1432,1456]" italics="true" pageId="10" pageNumber="11">
<taxonomicName id="4C125EBF0D7E836C9F3BFA1BFD56FA4E" box="[579,669,1432,1456]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="glaucos">glaucos</taxonomicName>
,
</emphasis>
is Greek for bluish gray, in reference to the blue-gray body and fin coloration. It is treated as a noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>
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