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<document id="A4DA28977397B96FDA44FA00FC282C5F" ID-DOI="10.5281/zenodo.213710" ID-GBIF-Dataset="7d3de354-12a7-488e-9cc0-df0b7ebb60ae" ID-ISSN="1175-5326" ID-Zenodo-Dep="213710" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460520904993" checkinUser="plazi" docAuthor="Lin, Yucheng, Li, Shuqiang &amp; Jäger, Peter" docDate="2012" docId="03BB644A0342866BFF393DCE35C14737" docLanguage="en" docName="zt03475p064.pdf" docOrigin="Zootaxa 3475" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tetrablemma namkhan Lin, Li &amp; Jäger, 2012, sp. nov." docType="treatment" docVersion="9" lastPageNumber="60" masterDocId="FF821C320347866EFFAE3D59372E411D" masterDocTitle="Two new species of the family Tetrablemmidae (Araneae) from Laos and Malaysia" masterLastPageNumber="64" masterPageNumber="55" pageNumber="60" updateTime="1698314626194" updateUser="plazi">
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<mods:title id="59BF0B1EF48FCCCCEB4740EC2895D245">Two new species of the family Tetrablemmidae (Araneae) from Laos and Malaysia</mods:title>
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<mods:namePart id="F4609DBDECE0F12CEE032BCA343CCC83">Lin, Yucheng</mods:namePart>
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<mods:namePart id="C9364941BA544EDDED6A23E790250FD2">Li, Shuqiang</mods:namePart>
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<mods:namePart id="33F3DC54DB5769934FC9F1D0A6194251">Jäger, Peter</mods:namePart>
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<mods:typeOfResource id="872586B769B468447BED7C0F6D1BCD90">text</mods:typeOfResource>
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<mods:title id="E327A435123A171FC344809630DBD29E">Zootaxa</mods:title>
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<mods:date id="AEE614C45C8F06D8A49D660A58B9751A">2012</mods:date>
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<treatment id="03BB644A0342866BFF393DCE35C14737" ID-DOI="http://doi.org/10.5281/zenodo.6180991" ID-GBIF-Taxon="119525733" ID-Zenodo-Dep="6180991" LSID="urn:lsid:plazi:treatment:03BB644A0342866BFF393DCE35C14737" httpUri="http://treatment.plazi.org/id/03BB644A0342866BFF393DCE35C14737" lastPageNumber="60" pageId="5" pageNumber="60">
<subSubSection id="C30886D70342866BFF393DCE37D641CF" pageId="5" pageNumber="60" type="nomenclature">
<paragraph id="8BADD55C0342866BFF393DCE353B41AC" blockId="5.[151,533,151,210]" box="[151,533,151,177]" pageId="5" pageNumber="60">
<heading id="D0E562300342866BFF393DCE353B41AC" bold="true" box="[151,533,151,177]" fontSize="11" level="1" pageId="5" pageNumber="60" reason="1">
<emphasis id="B966094E0342866BFF393DCE353B41AC" bold="true" box="[151,533,151,177]" pageId="5" pageNumber="60">
<taxonomicName id="4C12AEDF0342866BFF393DCE369E41AC" ID-CoL="55KWS" box="[151,432,151,177]" class="Arachnida" family="Tetrablemmidae" genus="Tetrablemma" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="60" phylum="Arthropoda" rank="species" species="namkhan" status="sp. nov.">
<emphasis id="B966094E0342866BFF393DCE369E41AC" bold="true" box="[151,432,151,177]" italics="true" pageId="5" pageNumber="60">Tetrablemma namkhan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A255B4350342866BFE193DCE353B41AC" box="[439,533,151,177]" pageId="5" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BADD55C0342866BFF393DE337D641CF" blockId="5.[151,533,151,210]" box="[151,248,186,210]" pageId="5" pageNumber="60">
<figureCitation id="1329C9D90342866BFF393DE337D641CF" box="[151,248,186,210]" captionStart-0="FIGURES 4 A G" captionStart-1="FIGURES 5 A C" captionStart-2="FIGURES 6 A C" captionStartId-0="6.[151,264,1937,1960]" captionStartId-1="7.[151,264,1940,1963]" captionStartId-2="8.[151,264,1958,1981]" captionTargetBox-0="[213,1376,195,1913]" captionTargetBox-1="[226,1375,195,1911]" captionTargetBox-2="[209,1390,196,1936]" captionTargetId-0="figure@6.[209,1378,195,1913]" captionTargetId-1="figure@7.[209,1378,195,1913]" captionTargetId-2="figure@8.[204,1390,195,1940]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="FIGURES 4 A G. Tetrablemma namkhan sp. nov., male (holotype). A D Habitus; E F Left chelicera; G Left leg I; H Metatarsus I (A dorsal, B ventral, C, E ectal, D frontal, H dorsal, G retrolateral). CA = cheliceral apophysis, CT = cheliceral tubercle, OT = ocular tubercle." captionText-1="FIGURES 5 A C. Tetrablemma namkhan sp. nov., male (holotype). A B Left pedipalp; C Distal end of bulb (A prolateral, B retrolateral, C dorsal). Em = embolus, Fe = femur, Pa = patella, Pb = bulb, Sd = sperm duct, Ti = tibia." captionText-2="FIGURES 6 A C. Tetrablemma namkhan sp. nov., male (holotype). A B Left pedipalp; C Distal end of bulb (A prolateral, B retrolateral, C dorsal)." httpUri-0="https://zenodo.org/record/213714/files/figure.png" httpUri-1="https://zenodo.org/record/213715/files/figure.png" httpUri-2="https://zenodo.org/record/213716/files/figure.png" pageId="5" pageNumber="60">Figs 46</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30886D70342866BFF393C5A34AA407F" pageId="5" pageNumber="60" type="materials_examined">
<paragraph id="8BADD55C0342866BFF393C5A34AA407F" blockId="5.[151,1437,258,1579]" pageId="5" pageNumber="60">
<emphasis id="B966094E0342866BFF393C5A352A4006" bold="true" box="[151,516,258,283]" pageId="5" pageNumber="60">
<typeStatus id="54A96BFE0342866BFF393C5A37FC4006" box="[151,210,259,283]" pageId="5" pageNumber="60">Type</typeStatus>
material:
<typeStatus id="54A96BFE0342866BFEE33C5A36904006" box="[333,446,259,283]" pageId="5" pageNumber="60" type="holotype">Holotype</typeStatus>
: male
</emphasis>
(
<collectionCode id="ED034D990342866BFDBC3C5B35614007" LSID="urn:lsid:biocol.org:col:34838" box="[530,591,258,282]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="60">SMF</collectionCode>
),
<collectingCountry id="F30595CC0342866BFDCD3C5A35B24007" box="[611,668,259,282]" name="Laos" pageId="5" pageNumber="60">Laos</collectingCountry>
, Luang Prabang Prov., SE Luang Prabang, Xiang Ngeun Distr., Nam Khan, Ban Keng Koung, 19°40ʹ963ʺN, 102°18ʹ442ʺE, altitude
<quantity id="4CEA78B90342866BFC2C3C7F34FE4023" box="[898,976,294,318]" metricMagnitude="2" metricUnit="m" metricValue="3.72" pageId="5" pageNumber="60" unit="m" value="372.0">372 m</quantity>
, disturbed forest, valley, waterfall, dry stream bed, P. Jäger leg.
<date id="FFACF39C0342866BFE023C13358E407F" box="[428,672,330,354]" pageId="5" pageNumber="60" value="2008-02-21" valueMax="2008-02-23" valueMin="2008-02-21">2123 February 2008</date>
, Winkler extraction.
</paragraph>
</subSubSection>
<subSubSection id="C30886D70342866BFF6B3C3632B9409B" box="[197,1431,366,391]" pageId="5" pageNumber="60" type="etymology">
<paragraph id="8BADD55C0342866BFF6B3C3632B9409B" blockId="5.[151,1437,258,1579]" box="[197,1431,366,391]" pageId="5" pageNumber="60">
<emphasis id="B966094E0342866BFF6B3C363665409A" bold="true" box="[197,331,367,391]" pageId="5" pageNumber="60">Etymology.</emphasis>
The specific epithet refers to the
<typeStatus id="54A96BFE0342866BFD6D3C3635DD409A" box="[707,755,367,391]" pageId="5" pageNumber="60">type</typeStatus>
locality, the Nam Khan (= River Khan); noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C30886D70342866BFF6B3CCB32AF4343" pageId="5" pageNumber="60" type="diagnosis">
<paragraph id="8BADD55C0342866BFF6B3CCB32AF4343" blockId="5.[151,1437,258,1579]" pageId="5" pageNumber="60">
<emphasis id="B966094E0342866BFF6B3CCB361140B6" bold="true" box="[197,319,402,427]" pageId="5" pageNumber="60">Diagnosis.</emphasis>
This new species is similar to
<taxonomicName id="4C12AEDF0342866BFD193CCD341640B6" box="[695,824,404,427]" class="Arachnida" family="Tetrablemmidae" genus="Tetrablemma" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="60" phylum="Arthropoda" rank="species" species="magister">
<emphasis id="B966094E0342866BFD193CCD341640B6" box="[695,824,404,427]" italics="true" pageId="5" pageNumber="60">T. magister</emphasis>
</taxonomicName>
from Queenland (see Burger 2008: figs. 1520),
<taxonomicName id="4C12AEDF0342866BFA293CCD362B40D3" class="Arachnida" family="Tetrablemmidae" genus="Tetrablemma" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="60" phylum="Arthropoda" rank="species" species="brevidens">
<emphasis id="B966094E0342866BFA293CCD362B40D3" italics="true" pageId="5" pageNumber="60">T. brevidens</emphasis>
</taxonomicName>
from
<collectingCountry id="F30595CC0342866BFEFE3CEF36BD40D3" box="[336,403,438,462]" name="China" pageId="5" pageNumber="60">China</collectingCountry>
(see
<bibRefCitation id="EF83A8AD0342866BFE7A3CEE35B940D3" author="Tong" box="[468,663,438,463]" pageId="5" pageNumber="60" refString="Tong, Y. F. &amp; Li, S. (2008) Tetrablemmidae (Arachnida, Araneae), a spider family newly recorded from China. Organisms, Diversity &amp; Evolution, 8, 84 - 98." type="journal article" year="2008">Tong &amp; Li 2008</bibRefCitation>
: figs. 5AC, FH) and
<taxonomicName id="4C12AEDF0342866BFC023CE1333740D3" box="[940,1049,438,463]" class="Arachnida" family="Tetrablemmidae" genus="Tetrablemma" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="60" phylum="Arthropoda" rank="species" species="thamin">
<emphasis id="B966094E0342866BFC023CE1333740D3" box="[940,1049,438,463]" italics="true" pageId="5" pageNumber="60">T. thamin</emphasis>
</taxonomicName>
from
<collectingCountry id="F30595CC0342866BFBCA3CEE33FF40D2" box="[1124,1233,439,463]" name="Myanmar" pageId="5" pageNumber="60">Myanmar</collectingCountry>
(see
<bibRefCitation id="EF83A8AD0342866BFABC3CEF366540EF" author="Labarque" pageId="5" pageNumber="60" refString="Labarque, F. M. &amp; Grismado, C. J. (2009) Description of a new species of armored spider from Myanmar (Araneae: Tetrablemmidae). Zootaxa, 2143, 55 - 58." type="journal article" year="2009">Labarque &amp; Grismado 2009</bibRefCitation>
: figs. 1, 3, 69) in the shape of the prosoma in lateral view, but males can be distinguished by the presence of an anterior dorsal lump on the male prosoma (
<figureCitation id="1329C9D90342866BFC863CA734AE430B" box="[808,896,510,534]" captionStart="FIGURES 4 A G" captionStartId="6.[151,264,1937,1960]" captionTargetBox="[213,1376,195,1913]" captionTargetId="figure@6.[209,1378,195,1913]" captionTargetPageId="6" captionText="FIGURES 4 A G. Tetrablemma namkhan sp. nov., male (holotype). A D Habitus; E F Left chelicera; G Left leg I; H Metatarsus I (A dorsal, B ventral, C, E ectal, D frontal, H dorsal, G retrolateral). CA = cheliceral apophysis, CT = cheliceral tubercle, OT = ocular tubercle." httpUri="https://zenodo.org/record/213714/files/figure.png" pageId="5" pageNumber="60">Fig. 4C</figureCitation>
), the short and S-shaped cheliceral apophysis in dorsal view (
<figureCitation id="1329C9D90342866BFE873F7B368D4327" box="[297,419,546,570]" captionStart="FIGURES 4 A G" captionStartId="6.[151,264,1937,1960]" captionTargetBox="[213,1376,195,1913]" captionTargetId="figure@6.[209,1378,195,1913]" captionTargetPageId="6" captionText="FIGURES 4 A G. Tetrablemma namkhan sp. nov., male (holotype). A D Habitus; E F Left chelicera; G Left leg I; H Metatarsus I (A dorsal, B ventral, C, E ectal, D frontal, H dorsal, G retrolateral). CA = cheliceral apophysis, CT = cheliceral tubercle, OT = ocular tubercle." httpUri="https://zenodo.org/record/213714/files/figure.png" pageId="5" pageNumber="60">Figs 4EF</figureCitation>
), the tortuous and six curves sperm duct (
<figureCitation id="1329C9D90342866BFCDD3F7B34DF4327" box="[883,1009,546,570]" captionStart="FIGURES 5 A C" captionStartId="7.[151,264,1940,1963]" captionTargetBox="[226,1375,195,1911]" captionTargetId="figure@7.[209,1378,195,1913]" captionTargetPageId="7" captionText="FIGURES 5 A C. Tetrablemma namkhan sp. nov., male (holotype). A B Left pedipalp; C Distal end of bulb (A prolateral, B retrolateral, C dorsal). Em = embolus, Fe = femur, Pa = patella, Pb = bulb, Sd = sperm duct, Ti = tibia." httpUri="https://zenodo.org/record/213715/files/figure.png" pageId="5" pageNumber="60">Figs 5A, B</figureCitation>
; 6A, B), and the slightly curved, wide embolus (
<figureCitation id="1329C9D90342866BFEA93F1F364D4343" box="[263,355,582,606]" captionStart="FIGURES 5 A C" captionStartId="7.[151,264,1940,1963]" captionTargetBox="[226,1375,195,1911]" captionTargetId="figure@7.[209,1378,195,1913]" captionTargetPageId="7" captionText="FIGURES 5 A C. Tetrablemma namkhan sp. nov., male (holotype). A B Left pedipalp; C Distal end of bulb (A prolateral, B retrolateral, C dorsal). Em = embolus, Fe = femur, Pa = patella, Pb = bulb, Sd = sperm duct, Ti = tibia." httpUri="https://zenodo.org/record/213715/files/figure.png" pageId="5" pageNumber="60">Figs 5C</figureCitation>
; 6C vs Burger 2008: fig. 19,
<bibRefCitation id="EF83A8AD0342866BFD063F1E344F4343" author="Tong" box="[680,865,582,607]" pageId="5" pageNumber="60" refString="Tong, Y. F. &amp; Li, S. (2008) Tetrablemmidae (Arachnida, Araneae), a spider family newly recorded from China. Organisms, Diversity &amp; Evolution, 8, 84 - 98." type="journal article" year="2008">Tong &amp; Li 2008</bibRefCitation>
: fig. 5F and
<bibRefCitation id="EF83A8AD0342866BFC433F1F32024343" author="Labarque" box="[1005,1324,582,606]" pageId="5" pageNumber="60" refString="Labarque, F. M. &amp; Grismado, C. J. (2009) Description of a new species of armored spider from Myanmar (Araneae: Tetrablemmidae). Zootaxa, 2143, 55 - 58." type="journal article" year="2009">Labarque &amp; Grismado 2009</bibRefCitation>
: fig. 3).
</paragraph>
</subSubSection>
<subSubSection id="C30886D70342866BFF6B3F33368C471B" pageId="5" pageNumber="60" type="description">
<paragraph id="8BADD55C0342866BFF6B3F33347843BB" blockId="5.[151,1437,258,1579]" pageId="5" pageNumber="60">
<emphasis id="B966094E0342866BFF6B3F33368F439E" bold="true" box="[197,417,618,643]" pageId="5" pageNumber="60">Description. Male</emphasis>
(
<typeStatus id="54A96BFE0342866BFE183F333530439F" box="[438,542,618,642]" pageId="5" pageNumber="60" type="holotype">holotype</typeStatus>
). Colouration: Prosoma reddish-brown, opisthosoma and legs pale reddishbrown to yellowish-brown. Somatic characters see
<figureCitation id="1329C9D90342866BFD783FD7347C43BB" box="[726,850,654,678]" captionStart="FIGURES 4 A G" captionStartId="6.[151,264,1937,1960]" captionTargetBox="[213,1376,195,1913]" captionTargetId="figure@6.[209,1378,195,1913]" captionTargetPageId="6" captionText="FIGURES 4 A G. Tetrablemma namkhan sp. nov., male (holotype). A D Habitus; E F Left chelicera; G Left leg I; H Metatarsus I (A dorsal, B ventral, C, E ectal, D frontal, H dorsal, G retrolateral). CA = cheliceral apophysis, CT = cheliceral tubercle, OT = ocular tubercle." httpUri="https://zenodo.org/record/213714/files/figure.png" pageId="5" pageNumber="60">Figs 4AG</figureCitation>
.
</paragraph>
<paragraph id="8BADD55C0342866BFF6B3FEA36CE422B" blockId="5.[151,1437,258,1579]" pageId="5" pageNumber="60">Measurements: Total length 1.16. Prosoma length 0.56, width 0.47, height 0.53. Opisthosoma length 0.77, width 0.61, height 0.46. Clypeus height 0.36. ALE 0.071, PLE 0.054. Sternum length 0.32, width 0.36. Legs: I 1.28 (0.43, 0.13, 0.30, 0.21, 0.21); II 1.19 (0.38, 0.13, 0.27, 0.20, 0.21); III 1.07 (0.32, 0.13, 0.23, 0.20, 0.20); IV 1.38 (0.43, 0.13, 0.36, 0.25, 0.21).</paragraph>
<paragraph id="8BADD55C0342866BFF6B3E1A37C5452E" blockId="5.[151,1437,258,1579]" pageId="5" pageNumber="60">Prosoma: Dorsal shield sclerotised, finely reticulate, except smooth radial region. Cephalic part elevated, surface smooth, planar, anteriorly sloping forward. Thoracic part marginally rugose with smooth striae. Ocular area situated centrally on prosoma, with four eyes in a tetrad, eye base black, AME and PME absent. ALE&gt;PLE. Clypeus very wide, completely reticulate, margin rounded. Chelicerae surface smooth, cheliceral horn robust, Sshaped in dorsal view, a small tubercle at cheliceral horn basally. Cheliceral laminae developed. Gnathocoxae basally wide, distally narrow. Labium subtriangular. Sternum wider than long, with fine reticulation, marginally rugose.</paragraph>
<paragraph id="8BADD55C0342866BFF6B396634AC4583" blockId="5.[151,1437,258,1579]" pageId="5" pageNumber="60">Legs: All femora with granules and annular veins. Other segments sculptured. Femur I slightly swollen. Metatarsus I proximo-mesially contracted, with two small tubercles. Tibiae IIV with three trichobothria, metatarsus IIV with one trichobothrium. Leg formula: IV-I-II-III.</paragraph>
<paragraph id="8BADD55C0342866BFF6B39F3338A4433" blockId="5.[151,1437,258,1579]" pageId="5" pageNumber="60">Opisthosoma: Dorsal scutum ovoid, finely reticulate, covered with long setae. Ventral scutum reticulate, posterior region slightly crinkled, booklung cover smooth. Lateral scutum I long, exceeding posterior margin of pre-anal plate. Postgenital plate present, as wide as pre-anal plate. Pre-anal plate rectangular, wider than long, covered sparsely with plumose setae, overlapping with posterior margin of postgenital plate.</paragraph>
<paragraph id="8BADD55C0342866BFF6B386337DF44FF" blockId="5.[151,1437,258,1579]" pageId="5" pageNumber="60">
Pedipalp (
<figureCitation id="1329C9D90342866BFE943863368B444F" box="[314,421,1338,1362]" captionStart-0="FIGURES 5 A C" captionStart-1="FIGURES 6 A C" captionStartId-0="7.[151,264,1940,1963]" captionStartId-1="8.[151,264,1958,1981]" captionTargetBox-0="[226,1375,195,1911]" captionTargetBox-1="[209,1390,196,1936]" captionTargetId-0="figure@7.[209,1378,195,1913]" captionTargetId-1="figure@8.[204,1390,195,1940]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURES 5 A C. Tetrablemma namkhan sp. nov., male (holotype). A B Left pedipalp; C Distal end of bulb (A prolateral, B retrolateral, C dorsal). Em = embolus, Fe = femur, Pa = patella, Pb = bulb, Sd = sperm duct, Ti = tibia." captionText-1="FIGURES 6 A C. Tetrablemma namkhan sp. nov., male (holotype). A B Left pedipalp; C Distal end of bulb (A prolateral, B retrolateral, C dorsal)." httpUri-0="https://zenodo.org/record/213715/files/figure.png" httpUri-1="https://zenodo.org/record/213716/files/figure.png" pageId="5" pageNumber="60">Figs 56</figureCitation>
): Femur finely granulated, slightly swollen, subequal to tibia in length, with two long ventral setae proximally. Patella short, subequal to 1/2 width of tibia, without modification. Tibia strongly swollen, with distal trichobothrium dorsally. Cymbium shorter than patella, with few long setae dorsally. Bulb pear-shaped. Spermophor course with several bends. Embolus short, strongly sclerotised, curving mesially and weakly furcate distally.
</paragraph>
<paragraph id="8BADD55C0342866BFF6B38B6368C471B" blockId="5.[151,1437,258,1579]" box="[197,418,1518,1543]" pageId="5" pageNumber="60">
<emphasis id="B966094E0342866BFF6B38B6360B471A" bold="true" box="[197,293,1519,1543]" pageId="5" pageNumber="60">Female:</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30886D70342866BFF6B3B4B35C14737" box="[197,751,1554,1579]" pageId="5" pageNumber="60" type="distribution">
<paragraph id="8BADD55C0342866BFF6B3B4B35C14737" blockId="5.[151,1437,258,1579]" box="[197,751,1554,1579]" pageId="5" pageNumber="60">
<emphasis id="B966094E0342866BFF6B3B4B36704736" bold="true" box="[197,350,1554,1579]" pageId="5" pageNumber="60">Distribution.</emphasis>
Known only from the
<typeStatus id="54A96BFE0342866BFDF33B4A35A34736" box="[605,653,1555,1579]" pageId="5" pageNumber="60">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>