247 lines
31 KiB
XML
247 lines
31 KiB
XML
<document id="77F2E8A44CD8AF75A63FB79A55F75EBD" ID-DOI="10.11646/zootaxa.3620.1.2" ID-GBIF-Dataset="60ae7d68-88b7-4da4-acae-6e4cda073ea0" ID-ISSN="1175-5326" ID-Zenodo-Dep="283400" ID-ZooBank="3870AC9F-A4EE-4E36-86C7-E1D6967A2A2E" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460578993957" checkinUser="plazi" docAuthor="Wu, Min & Xu, Qin" docDate="2013" docId="03B8B70EFF89FFCFFF05FB72FBEBB5E8" docLanguage="en" docName="zt03620p066.pdf" docOrigin="Zootaxa 3620 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Serina denticulata Wu & Xu, 2013, new species" docType="treatment" docVersion="5" lastPageNumber="63" masterDocId="FF81CF76FF98FFDBFF92FFF2FFB8B51F" masterDocTitle="Serina Gredler (Gastropoda, Stylommatophora: Enidae), the continuous-peristomed mountain snails endemic to the eastern slope of the Qinghai-Xizang Plateau" masterLastPageNumber="66" masterPageNumber="43" pageNumber="60" updateTime="1698323809859" updateUser="plazi">
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<mods:title id="EC43FCFDD9EAF347B7C3E2C8BC27EF26">Serina Gredler (Gastropoda, Stylommatophora: Enidae), the continuous-peristomed mountain snails endemic to the eastern slope of the Qinghai-Xizang Plateau</mods:title>
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<mods:namePart id="4E43FBCB836429D08382D081C3C61339">Wu, Min</mods:namePart>
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<mods:roleTerm id="38A692F3A316C8EFF601220829D71825">Author</mods:roleTerm>
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<mods:namePart id="B03C0DA4CB57EAFE9C87CFAA25F9E65A">Xu, Qin</mods:namePart>
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<mods:typeOfResource id="86F25E15BDE3383A13BD70AB9720CC16">text</mods:typeOfResource>
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<mods:title id="8E4843F3E07A8A20F5569B1C4B5D3FE7">Zootaxa</mods:title>
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<mods:part id="DB96A848736F97A2B4864FD97FEE9297">
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<mods:date id="E5CE314067A4CA5967657E94D9A4B7E9">2013</mods:date>
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<mods:detail id="06E5DFE8BE218A03EF459F284D472F42" type="volume">
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<mods:number id="E3F394F24B4FE39CD9A58F96A97FD823">3620</mods:number>
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<mods:classification id="9895022E28FBDB2CE6C5821BF70392E5">journal article</mods:classification>
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<mods:identifier id="B98DD5878814A7FCB06A1CB525084D7A" type="DOI">10.11646/zootaxa.3620.1.2</mods:identifier>
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<mods:identifier id="91589A74AA9F1679C34C8C7091F17A0B" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="5DC582939E69903F56884ED22C0F2839" type="Zenodo-Dep">283400</mods:identifier>
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<treatment id="03B8B70EFF89FFCFFF05FB72FBEBB5E8" ID-DOI="http://doi.org/10.5281/zenodo.6162575" ID-GBIF-Taxon="127663380" ID-Zenodo-Dep="6162575" LSID="urn:lsid:plazi:treatment:03B8B70EFF89FFCFFF05FB72FBEBB5E8" httpUri="http://treatment.plazi.org/id/03B8B70EFF89FFCFFF05FB72FBEBB5E8" lastPageId="20" lastPageNumber="63" pageId="17" pageNumber="60">
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<subSubSection id="C30B5593FF89FFCAFF05FB72FB4DB057" pageId="17" pageNumber="60" type="nomenclature">
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<paragraph id="8BAE0618FF89FFCAFF05FB72FD85B185" blockId="17.[151,573,1152,1211]" box="[151,573,1152,1178]" pageId="17" pageNumber="60">
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<heading id="D0E6B174FF89FFCAFF05FB72FD85B185" bold="true" box="[151,573,1152,1178]" fontSize="11" level="1" pageId="17" pageNumber="60" reason="1">
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<emphasis id="B965DA0AFF89FFCAFF05FB72FD85B185" bold="true" box="[151,573,1152,1178]" pageId="17" pageNumber="60">
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11.
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<taxonomicName id="4C117D9BFF89FFCAFF53FB72FE27B185" ID-CoL="7T4SH" box="[193,415,1152,1178]" class="Gastropoda" family="Enidae" genus="Serina" kingdom="Animalia" order="Stylommatophora" pageId="17" pageNumber="60" phylum="Mollusca" rank="species" species="denticulata" status="sp. nov.">
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<emphasis id="B965DA0AFF89FFCAFF53FB72FE27B185" bold="true" box="[193,415,1152,1178]" italics="true" pageId="17" pageNumber="60">Serina denticulata</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A2566771FF89FFCAFE3FFB72FD85B185" box="[429,573,1152,1178]" pageId="17" pageNumber="60" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BAE0618FF89FFCAFF05FB51FDAFB1A4" blockId="17.[151,573,1152,1211]" box="[151,535,1187,1211]" pageId="17" pageNumber="60">
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<figureCitation id="132A1A9DFF89FFCAFF05FB51FF5BB1A4" box="[151,227,1187,1211]" captionStart="FIGURE 1" captionStartId="2.[151,250,1303,1326]" captionTargetBox="[153,1432,201,1255]" captionTargetId="figure@2.[151,1436,171,1279]" captionTargetPageId="2" captionText="FIGURE 1. Distribution of Serina species. S. soluta inflata Yen, 1939 is also indicated although synonymized in this paper. S. soluta stenochila (Möllendorff, 1901) is not mapped due to imprecise locality information." httpUri="https://zenodo.org/record/283401/files/figure.png" pageId="17" pageNumber="60">Figs. 1</figureCitation>
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,
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<figureCitation id="132A1A9DFF89FFCAFF63FB51FF46B1A4" box="[241,254,1187,1211]" captionStart="FIGURE 5. A, B" captionStartId="11.[151,250,1764,1787]" captionTargetBox="[151,1435,195,1740]" captionTargetId="figure@11.[151,1436,195,1740]" captionTargetPageId="11" captionText="FIGURE 5. A, B. Serina ser Gredler, 1898. A. SMF- 42036, Wangjiaba. B. ZI RAS N 76. C-E. Serina subser Gredler, 1898. C. NHM- 1902.5.13.23 – 24, Gansu. D. SMF- 42045, Wudu. E. HBUMM- 05392 – specimen 22, Lixian. F-K. Serina soluta (Möllendorff, 1901). F. NHM-Trechmann Acc. No. 2176, Dangchang. G. SMF- 42047, Wasigou. H, I. Serina soluta inflata Yen, 1939. H. HBUMM- 04318 – specimen 2, Luding. I. SMF- 42095, holotype, Kangding. J, K. Buliminus soluta stenochila Möllendorff, 1901. J. SMF- 42048, lectotype, valley of Daduhe River. K. SMF- 42049, paralectotype. L. Serina vincentii (Gredler, 1898), SMF- 42032, between Tshun-dshia-wan and Wangjiaba. M, N. Serina denticulata n. sp. M. HBUMM- 00281 – specimen 5, holotype, Daigusi. N. HBUMM- 00281 – specimen 6, paratype, Daigusi." httpUri="https://zenodo.org/record/283405/files/figure.png" pageId="17" pageNumber="60">5</figureCitation>
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M–N, 8, 9;
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<tableCitation id="C69333A3FF89FFCAFE13FB51FE44B1A4" box="[385,508,1187,1211]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStart-2="TABLE 3" captionStart-3="TABLE 4" captionStartId-0="5.[151,235,152,173]" captionStartId-1="5.[151,235,1295,1316]" captionStartId-2="6.[151,235,298,319]" captionStartId-3="20.[151,235,296,317]" captionTargetBox-0="[151,1436,323,1223]" captionTargetBox-1="[151,1435,1374,1654]" captionTargetBox-2="[159,1399,347,882]" captionTargetBox-3="[159,1228,345,703]" captionTargetPageId-0="5" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="20" captionText-0="TABLE 1. Shell measurements of Serina species (specimen lots: Serina belae, SMF- 42069; S. cathaica, SMF- 42046; S. egressa, HBUMM- 05426 + 05464 + 05538 + 05563 + 05648 + 0 5673, SMF- 42038; S. prostoma, NHM- 1912.6.27.69 - 71 + 1923.5.24.17 - 36, SMF- 42050, HBUMM- 04287 - + 04296; S. ser, SMF- 42036; S. subser, HBUMM- 00287 + 05796 + 05709 + 0 5392, SMF- 42041; S. soluta, SMF- 42095 + 42096 (S. soluta “ inflata ”), SMF- 42048 + 42049 (S. soluta “ stenochila ”), SMF 42047, HBUMM- 04318; S. vincentii, SMF 42032; S. denticulata n. sp., HBUMM- 00281)." captionText-1="TABLE 2. Measurements of genitalia of some Serina species (unit: mm; measured specimens: S. egressa, 2; S. prostoma, 2; S. subser, 2; S. denticulata n. sp., 1)." captionText-2="TABLE 3. Diagnostics of Serina species. Anatomy data from this study and Schileyko, 1998 (Serina ser Gredler, 1898)." captionText-3="TABLE 4. Descriptive statistics of Serina denticulata n. sp." httpUri-0="http://table.plazi.org/id/DF6E5690FF9DFFDEFF05FF6AFBD5B436" httpUri-1="http://table.plazi.org/id/DF6E5690FF9DFFDEFF05FAFDFE4AB05B" httpUri-2="http://table.plazi.org/id/DF6E5690FF9EFFDDFF05FED8FAE5B45F" httpUri-3="http://table.plazi.org/id/DF6E5690FF8CFFCFFF05FEDAFD40B422" pageId="17" pageNumber="60" tableUuid-0="DF6E5690FF9DFFDEFF05FF6AFBD5B436" tableUuid-1="DF6E5690FF9DFFDEFF05FAFDFE4AB05B" tableUuid-2="DF6E5690FF9EFFDDFF05FED8FAE5B45F" tableUuid-3="DF6E5690FF8CFFCFFF05FEDAFD40B422">Tables 1–4</tableCitation>
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,
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<tableCitation id="C69333A3FF89FFCAFD9BFB51FDAFB1A4" box="[521,535,1187,1211]" captionStart="TABLE 6" captionStartId="22.[151,235,293,314]" captionTargetBox="[156,1416,936,1454]" captionTargetPageId="22" captionText="TABLE 6. Data set containing 18 characters (0 - 17) and 11 taxa used in the phylogenetic analyses. Information of the data contributors – S. ser (Gredler, 1898): shell characters from SMF- 42036, genitalian characters after Schileyko (1998, Fig. 235 B); S. egressa Sturany, 1900: HBUMM- 05426, 0 5464, 0 5538, 0 5563, 0 5648, 0 5673, SMF- 42038; S. subser Gredler, 1898: HBUMM- 00287, 0 5796, 0 5709, 05392; S. prostoma (Ancey, 1884): HBUMM- 04287, 0 4296, SMF- 042050; S. denticulata n. sp.: HBUMM- 0 0 281, types; Pupinidius latilabrum (Annandale, 1924): HBUMM- 04328; Pupinidius melinostoma (Möllendorff, 1901): HBUMM- 05480; Pupinidius nanpingensis (Möllendorff, 1901): HBUMM- 05497; Pupinidius wenxian Wu & Zheng, 2009: HBUMM- 05432; Pupopsis pupopsis (Gredler, 1898): HBUMM- 04298; Clausiliopsis szechenyi (Böttger, 1883): HBUMM- 05478; Holcauchen sulcata (Möllendorff, 1901): HBUMM- 05706. Measurements are from 334 shells of above species. Description of character and character states: Character 0, whorl number of embryonic shell; Character 1, whorl number; Character 2, shell height; Character 3, shell breadth; Character 4, aperture height; Character 5, aperture width; Character 6, ratio of shell height to shell breadth; Character 7, ratio of shell height to aperture height. For Characters 0 ~ 7, grouped by cluster center 1 – 1, grouped by cluster center 2 – 2, grouped by cluster center 1 or 2 – 3 (the clustering centers for character states 1 and 2 of Characters 0 - 7 see in Table 5). Character 8, peristome continuous with a channel – 1, aperture not continuous – 0; Character 9, palatal tooth present – 1, palatal tooth absent – 0; Character 10, parietal tooth present – 1, parietal tooth absent – 0; Character 11, with 2 columellar teeth – 2, with 1 columellar tooth – 1, columellar tooth absent – 0; Character 12, epiphallus twisted – 1, epiphallus straight – 0; Character 13, epiphallic caecum present – 1, epiphallic caecum absent – 0; Character 14, penial caecum present – 1, penical caecum absent – 0; Character 15, bursa copulatrix duct twisted – 1, bursa copulatrix duct straight – 0; Character 16, diverticle absent – 0, diverticle not terminally expanded – 1, diverticle terminally expanded – 2; Character 17, free oviduct longer than vagina – 1, free oviduct subequal to or shorter than vagina – 0." httpUri="http://table.plazi.org/id/DF6E5690FF8EFFCDFF05FED7FB38B67C" pageId="17" pageNumber="60" tableUuid="DF6E5690FF8EFFCDFF05FED7FB38B67C">6</tableCitation>
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</paragraph>
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<paragraph id="8BAE0618FF89FFCAFF05FB14FB4DB057" blockId="17.[151,1437,1253,1751]" pageId="17" pageNumber="60">
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<emphasis id="B965DA0AFF89FFCAFF05FB14FF59B1E1" bold="true" box="[151,225,1254,1278]" pageId="17" pageNumber="60">
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<typeStatus id="54AAB8BAFF89FFCAFF05FB14FF62B1E1" box="[151,218,1254,1278]" pageId="17" pageNumber="60">Types</typeStatus>
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.
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</emphasis>
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<typeStatus id="54AAB8BAFF89FFCAFF63FB17FEE6B1E2" box="[241,350,1253,1277]" pageId="17" pageNumber="60" type="holotype">Holotype</typeStatus>
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: HBUMM-00281–specimen 5; Daigusi Village, Luoda (33°54ʹN, 103°54ʹE), Diebu County, Gansu,
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<date id="FFAF20D8FF89FFCAFF7EFAFEFEC5B03B" box="[236,381,1292,1316]" pageId="17" pageNumber="60" value="2004-04-29">2004–IV–29</date>
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, colln. MW; measurements: ewh 1 3/4, wh 8 1/2, h
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<quantity id="4CE9ABFDFF89FFCAFC1EFAFEFC49B03B" box="[908,1009,1292,1316]" metricMagnitude="-2" metricUnit="m" metricValue="1.09" pageId="17" pageNumber="60" unit="mm" value="10.9">10.9 mm</quantity>
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, w
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<quantity id="4CE9ABFDFF89FFCAFB85FAFEFBD6B03B" box="[1047,1134,1292,1316]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="17" pageNumber="60" unit="mm" value="3.5">3.5 mm</quantity>
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, aph
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<quantity id="4CE9ABFDFF89FFCAFB3AFAFEFAB8B03B" box="[1192,1280,1292,1316]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="17" pageNumber="60" unit="mm" value="3.3">3.3 mm</quantity>
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, apw
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<quantity id="4CE9ABFDFF89FFCAFAD2FAFEFA20B03B" box="[1344,1432,1292,1316]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="17" pageNumber="60" unit="mm" value="2.5">2.5 mm</quantity>
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.
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<typeStatus id="54AAB8BAFF89FFCAFF05FAC3FEBFB056" box="[151,263,1329,1353]" pageId="17" pageNumber="60" type="paratype">Paratypes</typeStatus>
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, 25 fma, HBUMM- 0 0 281 –specimens 1~4 & 6~26; the same collection data as
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<typeStatus id="54AAB8BAFF89FFCAFB1EFAC2FB4AB057" box="[1164,1266,1328,1352]" pageId="17" pageNumber="60" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30B5593FF89FFCAFF5BFAA7FD95B073" box="[201,557,1364,1389]" pageId="17" pageNumber="60" type="materials_examined">
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<paragraph id="8BAE0618FF89FFCAFF5BFAA7FD95B073" blockId="17.[151,1437,1253,1751]" box="[201,557,1364,1389]" pageId="17" pageNumber="60">
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<emphasis id="B965DA0AFF89FFCAFF5BFAA7FEDDB072" bold="true" box="[201,357,1364,1389]" pageId="17" pageNumber="60">
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<typeStatus id="54AAB8BAFF89FFCAFF5BFAA7FEBAB073" box="[201,258,1365,1388]" pageId="17" pageNumber="60">Type</typeStatus>
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locality.
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</emphasis>
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Daigusi (Gansu).
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</paragraph>
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</subSubSection>
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<subSubSection id="C30B5593FF89FFCAFF5BFA8BFAF2B08F" box="[201,1354,1400,1425]" pageId="17" pageNumber="60" type="etymology">
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<paragraph id="8BAE0618FF89FFCAFF5BFA8BFAF2B08F" blockId="17.[151,1437,1253,1751]" box="[201,1354,1400,1425]" pageId="17" pageNumber="60">
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<emphasis id="B965DA0AFF89FFCAFF5BFA8BFEF7B08E" bold="true" box="[201,335,1401,1425]" pageId="17" pageNumber="60">Etymology.</emphasis>
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The species name derives from the columellar tooth which can be observed from aperture.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30B5593FF89FFCAFF5BFA6EFC0BB0AB" box="[201,947,1436,1461]" pageId="17" pageNumber="60" type="distribution">
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<paragraph id="8BAE0618FF89FFCAFF5BFA6EFC0BB0AB" blockId="17.[151,1437,1253,1751]" box="[201,947,1436,1461]" pageId="17" pageNumber="60">
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<emphasis id="B965DA0AFF89FFCAFF5BFA6EFEDAB0AA" bold="true" box="[201,354,1436,1461]" pageId="17" pageNumber="60">Distribution.</emphasis>
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Gansu (only known from the
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<typeStatus id="54AAB8BAFF89FFCAFD26FA6FFD5CB0AA" box="[692,740,1437,1461]" pageId="17" pageNumber="60">type</typeStatus>
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locality, Daigusi).
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</paragraph>
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</subSubSection>
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<subSubSection id="C30B5593FF89FFC8FF5BFA32FAA1B5CA" lastPageId="19" lastPageNumber="62" pageId="17" pageNumber="60" type="diagnosis">
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<paragraph id="8BAE0618FF89FFCAFF5BFA32FE0AB35B" blockId="17.[151,1437,1253,1751]" pageId="17" pageNumber="60">
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<emphasis id="B965DA0AFF89FFCAFF5BFA32FEFBB0C6" bold="true" box="[201,323,1472,1497]" pageId="17" pageNumber="60">Diagnosis.</emphasis>
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Shell with the most swollen part occurred at penultimate whorl. Suture simple. Last whorl straight in front or ascending immediately near aperture; rounded at periphery. Aperture completely attached to body whorl. Aperture not duplicate. Columellar margin with a weak oblique spiral baso-columellar tooth and a more developed tooth above it.
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</paragraph>
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<paragraph id="8BAE0618FF89FFC8FF5BF9A2FAA1B5CA" blockId="17.[151,1437,1253,1751]" lastBlockId="19.[151,1437,151,752]" lastPageId="19" lastPageNumber="62" pageId="17" pageNumber="60">
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<emphasis id="B965DA0AFF89FFCAFF5BF9A2FEB3B376" bold="true" box="[201,267,1616,1641]" pageId="17" pageNumber="60">Shell.</emphasis>
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Fusiform; with apex slightly abruptly pointed; dextral; with the most swollen part occurred at penultimate whorl; height 10.2–11.5–13.9, diameter major 3.5–3.9–4.5, ratio of height /diameter major 2.66–2.94–3.35. Growth lines usually not very clear. Whorls 8 1/8–8 3/4-–9 1/4, convex. Embryonic shell 1 5/8–1 3/4+–2 whorls. Postnuclear whorls smooth. Suture simple. Last whorl straight in front, or ascending immediately near aperture; rounded at periphery; abaperturally with a rugate region with crowded and/or thickened growthline-like folds. Aperture in a plane and straight; ovate, or subcircular; slightly oblique; completely attached to body whorl; armed; height 3.0–3.5–3.8, width 2.5–2.8–3.0, ratio of shell height /apertural height 2.96-3.30-3.82. Aperture not duplicate. Palatal tooth absent. Palatal margin rounded; toothless. Peristome and parietal callus almost completely fusing except for a channel at upper insertion of peristome. Reflexed part of peristome straight and not curved backward. Parietal callus distinct. Parietal tooth absent. Palatal wall without deep depression or tooth of irregular shape. Columellar margin reflexed; with a weak oblique baso-columellar spiral tooth and a more developed tooth above it; not sinuous. Columella arched. Umbilicus widely open. Shell light brown with a broad dirty white spiral band nearly occupying whole whorls; aperture white. Measured specimens:
|
||
<typeStatus id="54AAB8BAFF8BFFC8FC79FF4EFBF5B5CB" box="[1003,1101,188,212]" pageId="19" pageNumber="62" type="holotype">holotype</typeStatus>
|
||
and 19
|
||
<typeStatus id="54AAB8BAFF8BFFC8FB35FF4FFAAEB5CA" box="[1191,1302,189,213]" pageId="19" pageNumber="62" type="paratype">paratypes</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF6E5690FF8AFFC9FF05F94CFAA4B3CB" box="[151,1308,1726,1749]" httpUri="https://zenodo.org/record/283410/files/figure.png" pageId="18" pageNumber="61" targetBox="[212,1365,218,1700]" targetPageId="18">
|
||
<paragraph id="8BAE0618FF8AFFC9FF05F94CFAA4B3CB" blockId="18.[151,1308,1726,1749]" box="[151,1308,1726,1749]" pageId="18" pageNumber="61">
|
||
<emphasis id="B965DA0AFF8AFFC9FF05F94CFEAAB3CC" bold="true" box="[151,274,1726,1749]" pageId="18" pageNumber="61">FIGURE 8.</emphasis>
|
||
<taxonomicName id="4C117D9BFF8AFFC9FE8BF94DFE6BB3CB" box="[281,467,1726,1748]" class="Gastropoda" family="Enidae" genus="Serina" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="61" phylum="Mollusca" rank="species" species="denticulata" status="sp. nov.">
|
||
<emphasis id="B965DA0AFF8AFFC9FE8BF94DFE6BB3CB" box="[281,467,1726,1748]" italics="true" pageId="18" pageNumber="61">Serina denticulata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B965DA0AFF8AFFC9FE4BF94DFDAAB3CB" bold="true" box="[473,530,1727,1748]" pageId="18" pageNumber="61">
|
||
<taxonomicNameLabel id="A2566771FF8AFFC9FE4BF94DFDAAB3CB" box="[473,530,1727,1748]" pageId="18" pageNumber="61" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, HBUMM-00281–specimen 1, paratype, Daigusi. General view of genitalia.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C30B5593FF8BFFCFFF5BFF2DFBEBB5E8" lastPageId="20" lastPageNumber="63" pageId="19" pageNumber="62" type="description">
|
||
<paragraph id="8BAE0618FF8BFFC8FF5BFF2DFDC5B7F0" blockId="19.[151,1437,151,752]" pageId="19" pageNumber="62">
|
||
<emphasis id="B965DA0AFF8BFFC8FF5BFF2DFE87B5E7" bold="true" box="[201,319,223,248]" pageId="19" pageNumber="62">Genitalia.</emphasis>
|
||
Vas deferens short; evenly thick; entering epiphallus laterally with distinct demarcation. Epiphallus long; narrowed towards distal end; forming a few loops. Epiphallic caecum absent. Flagellum short; conic; proximally normal; with tip blunt. Penis clavate and distally enlarged; terminally entering epiphallus; thin-walled. Longitudinal pilasters more than two; fused at epiphallic pore forming velum; forming 1 V-shaped structure. Vshaped pilaster with proximal free end approaching at penial retractor insertion; without papilla formed by fused distal pilasters. Penial process absent. Penial appendix moderately long; branched off from proximal portion of penis; with A-1 fused with A-2, and with distinct A-3, A-4 and A-5. A-1 very short; without verge toward penial chamber; with longitudinal pilasters. A-2 internally with longitudinal pilasters; near A-3 internally with a ring of papilla. A-3 opening into A-2 by a long needle-like papilla. A-5 short; convoluted. Penial retractor biramous; with arms arising from diaphragm separately; with penial branch attaching to medial penis; with appendical branch attaching to A-1+A-2. Atrium short; with weak atrial retractor. Free oviduct moderately long; longer than vagina. Vagina very short; not swollen; straight. Bursa copulatrix duct very long; proximally twisted. Bursa copulatrix small; well defined; shortly with stalk; without apical ligament. Diverticle longer than reservoir; expanded. Bursa copulatrix and diverticle distinguishable; forked more distally from their base. Measurement of genitalia: See
|
||
<tableCitation id="C69333A3FF8BFFC8FACCFD46FF1DB7F0" captionStart="TABLE 2" captionStartId="5.[151,235,1295,1316]" captionTargetBox="[151,1435,1374,1654]" captionTargetPageId="5" captionText="TABLE 2. Measurements of genitalia of some Serina species (unit: mm; measured specimens: S. egressa, 2; S. prostoma, 2; S. subser, 2; S. denticulata n. sp., 1)." httpUri="http://table.plazi.org/id/DF6E5690FF9DFFDEFF05FAFDFE4AB05B" pageId="19" pageNumber="62" tableUuid="DF6E5690FF9DFFDEFF05FAFDFE4AB05B">Table 2</tableCitation>
|
||
(
|
||
<typeStatus id="54AAB8BAFF8BFFC8FF21FD25FEADB7F0" box="[179,277,727,751]" pageId="19" pageNumber="62" type="paratype">paratype</typeStatus>
|
||
HBUMM- 00281–specimen 1).
|
||
</paragraph>
|
||
<caption id="DF6E5690FF8BFFC8FF05F8FAFF5EB286" httpUri="https://zenodo.org/record/283411/files/figure.png" pageId="19" pageNumber="62" targetBox="[170,1414,859,1761]" targetPageId="19">
|
||
<paragraph id="8BAE0618FF8BFFC8FF05F8FAFF5EB286" blockId="19.[151,1437,1800,1945]" pageId="19" pageNumber="62">
|
||
<emphasis id="B965DA0AFF8BFFC8FF05F8FAFEAEB201" bold="true" box="[151,278,1800,1823]" pageId="19" pageNumber="62">FIGURE 9.</emphasis>
|
||
Rooted cladogram showing phylogenetic relationships among four genera, i.e.
|
||
<emphasis id="B965DA0AFF8BFFC8FBCCF8F8FB18B200" box="[1118,1184,1802,1823]" italics="true" pageId="19" pageNumber="62">Serina</emphasis>
|
||
(
|
||
<emphasis id="B965DA0AFF8BFFC8FB20F8F8FB07B200" box="[1202,1215,1802,1823]" italics="true" pageId="19" pageNumber="62">S</emphasis>
|
||
.
|
||
<emphasis id="B965DA0AFF8BFFC8FB5DF8F8FB55B200" box="[1231,1261,1802,1823]" italics="true" pageId="19" pageNumber="62">ser</emphasis>
|
||
,
|
||
<taxonomicName id="4C117D9BFF8BFFC8FB6FF8F8FAC1B200" box="[1277,1401,1802,1823]" class="Gastropoda" family="Enidae" genus="Serina" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="62" phylum="Mollusca" rank="species" species="prostoma">
|
||
<emphasis id="B965DA0AFF8BFFC8FB6FF8F8FAB2B200" box="[1277,1290,1802,1823]" italics="true" pageId="19" pageNumber="62">S</emphasis>
|
||
.
|
||
<emphasis id="B965DA0AFF8BFFC8FA88F8F8FAC1B200" box="[1306,1401,1802,1823]" italics="true" pageId="19" pageNumber="62">prostoma</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C117D9BFF8BFFC8FA1BF8F8FF60B222" class="Gastropoda" family="Enidae" genus="Serina" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="62" phylum="Mollusca" rank="species" species="subser">
|
||
<emphasis id="B965DA0AFF8BFFC8FA1BF8F8FA2DB200" box="[1417,1429,1802,1823]" italics="true" pageId="19" pageNumber="62">S</emphasis>
|
||
.
|
||
<emphasis id="B965DA0AFF8BFFC8FF05F8D5FF60B222" box="[151,216,1831,1853]" italics="true" pageId="19" pageNumber="62">subser</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C117D9BFF8BFFC8FF74F8DAFEF6B222" box="[230,334,1832,1853]" class="Gastropoda" family="Enidae" genus="Serina" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="62" phylum="Mollusca" rank="species" species="egressa">
|
||
<emphasis id="B965DA0AFF8BFFC8FF74F8DAFF4BB222" box="[230,243,1832,1853]" italics="true" pageId="19" pageNumber="62">S</emphasis>
|
||
.
|
||
<emphasis id="B965DA0AFF8BFFC8FE90F8DAFEF6B222" box="[258,334,1832,1853]" italics="true" pageId="19" pageNumber="62">egressa</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C117D9BFF8BFFC8FECFF8DAFE51B222" box="[349,489,1831,1853]" class="Gastropoda" family="Enidae" genus="Serina" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="62" phylum="Mollusca" rank="species" species="denticulate" status="sp. nov.">
|
||
<emphasis id="B965DA0AFF8BFFC8FECFF8DAFED1B222" box="[349,361,1832,1853]" italics="true" pageId="19" pageNumber="62">S</emphasis>
|
||
.
|
||
<emphasis id="B965DA0AFF8BFFC8FEEBF8D5FE51B222" box="[377,489,1831,1853]" italics="true" pageId="19" pageNumber="62">denticulate</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B965DA0AFF8BFFC8FE63F8DAFD94B222" bold="true" box="[497,556,1832,1853]" pageId="19" pageNumber="62">
|
||
<taxonomicNameLabel id="A2566771FF8BFFC8FE63F8DAFD94B222" box="[497,556,1832,1853]" pageId="19" pageNumber="62" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
),
|
||
<emphasis id="B965DA0AFF8BFFC8FDD6F8D5FD70B222" box="[580,712,1831,1853]" italics="true" pageId="19" pageNumber="62">Clausiliopsis</emphasis>
|
||
(
|
||
<emphasis id="B965DA0AFF8BFFC8FD4BF8DAFD52B222" box="[729,746,1832,1853]" italics="true" pageId="19" pageNumber="62">C</emphasis>
|
||
.
|
||
<taxonomicName id="4C117D9BFF8BFFC8FD6AF8D5FCE1B222" box="[760,857,1831,1853]" class="Gastropoda" family="Enidae" genus="Buliminus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="62" phylum="Mollusca" rank="species" species="szechenyi">
|
||
<emphasis id="B965DA0AFF8BFFC8FD6AF8D5FCE1B222" box="[760,857,1831,1853]" italics="true" pageId="19" pageNumber="62">szechenyi</emphasis>
|
||
</taxonomicName>
|
||
),
|
||
<taxonomicName id="4C117D9BFF8BFFC8FCE3F8DAFC76B222" box="[881,974,1832,1853]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="19" pageNumber="65" phylum="Tracheophyta" rank="species" species="pupopsis">
|
||
<emphasis id="B965DA0AFF8BFFC8FCE3F8DAFC76B222" box="[881,974,1832,1853]" italics="true" pageId="19" pageNumber="62">Pupopsis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C117D9BFF8BFFC8FC4CF8DAFBECB222" box="[990,1108,1832,1853]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="19" pageNumber="62" phylum="Tracheophyta" rank="species" species="pupopsis">
|
||
<emphasis id="B965DA0AFF8BFFC8FC4CF8DAFC53B222" box="[990,1003,1832,1853]" italics="true" pageId="19" pageNumber="62">P</emphasis>
|
||
.
|
||
<emphasis id="B965DA0AFF8BFFC8FC68F8DAFBECB222" box="[1018,1108,1832,1853]" italics="true" pageId="19" pageNumber="62">pupopsis</emphasis>
|
||
</taxonomicName>
|
||
), and
|
||
<emphasis id="B965DA0AFF8BFFC8FB0AF8D5FAAAB222" box="[1176,1298,1831,1853]" italics="true" pageId="19" pageNumber="62">Holcauchen</emphasis>
|
||
(
|
||
<emphasis id="B965DA0AFF8BFFC8FAB1F8DAFA8DB222" box="[1315,1333,1832,1853]" italics="true" pageId="19" pageNumber="62">H</emphasis>
|
||
.
|
||
<emphasis id="B965DA0AFF8BFFC8FAD6F8D5FA35B222" box="[1348,1421,1831,1853]" italics="true" pageId="19" pageNumber="62">sulcata</emphasis>
|
||
). Outgroup
|
||
<taxonomicName id="4C117D9BFF8BFFC8FE91F8B4FEC8B243" box="[259,368,1862,1884]" class="Gastropoda" family="Enidae" genus="Pupinidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="62" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B965DA0AFF8BFFC8FE91F8B4FEC8B243" box="[259,368,1862,1884]" italics="true" pageId="19" pageNumber="62">Pupinidius</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C117D9BFF8BFFC8FE16F8B5FE4FB243" box="[388,503,1863,1884]" class="Gastropoda" family="Enidae" genus="Pupinidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="62" phylum="Mollusca" rank="species" species="wenxian">
|
||
<emphasis id="B965DA0AFF8BFFC8FE16F8B5FE28B243" box="[388,400,1863,1884]" italics="true" pageId="19" pageNumber="62">P</emphasis>
|
||
.
|
||
<emphasis id="B965DA0AFF8BFFC8FE31F8B5FE4FB243" box="[419,503,1863,1884]" italics="true" pageId="19" pageNumber="62">wenxian</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C117D9BFF8BFFC8FD9BF8B5FD2AB243" box="[521,658,1862,1884]" class="Gastropoda" family="Enidae" genus="Pupinidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="62" phylum="Mollusca" rank="species" species="latilabrum">
|
||
<emphasis id="B965DA0AFF8BFFC8FD9BF8B5FDADB243" box="[521,533,1863,1884]" italics="true" pageId="19" pageNumber="62">P</emphasis>
|
||
.
|
||
<emphasis id="B965DA0AFF8BFFC8FDB5F8B4FD2AB243" box="[551,658,1862,1884]" italics="true" pageId="19" pageNumber="62">latilabrum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C117D9BFF8BFFC8FD31F8B5FCFBB243" box="[675,835,1862,1884]" class="Gastropoda" family="Enidae" genus="Pupinidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="62" phylum="Mollusca" rank="species" species="melinostoma">
|
||
<emphasis id="B965DA0AFF8BFFC8FD31F8B5FD08B243" box="[675,688,1863,1884]" italics="true" pageId="19" pageNumber="62">P</emphasis>
|
||
.
|
||
<emphasis id="B965DA0AFF8BFFC8FD50F8B4FCFBB243" box="[706,835,1862,1884]" italics="true" pageId="19" pageNumber="62">melinostoma</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C117D9BFF8BFFC8FCC6F8B5FC40B243" box="[852,1016,1863,1884]" class="Gastropoda" family="Enidae" genus="Pupinidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="62" phylum="Mollusca" rank="species" species="nanpingensis">
|
||
<emphasis id="B965DA0AFF8BFFC8FCC6F8B5FCD9B243" box="[852,865,1863,1884]" italics="true" pageId="19" pageNumber="62">P</emphasis>
|
||
.
|
||
<emphasis id="B965DA0AFF8BFFC8FCE1F8B5FC40B243" box="[883,1016,1863,1884]" italics="true" pageId="19" pageNumber="62">nanpingensis</emphasis>
|
||
</taxonomicName>
|
||
). Branch length 45, CI=0.60, RI=0.70. Explanation for characters and character states see in Table 6. Solid circle: nonhomoplasious change; open circle: homoplasious change.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAE0618FF8CFFCFFF5BFF65FBEBB5E8" blockId="20.[151,1436,151,247]" pageId="20" pageNumber="63">
|
||
<emphasis id="B965DA0AFF8CFFCFFF5BFF65FE7DB5AF" bold="true" box="[201,453,151,176]" pageId="20" pageNumber="63">Taxonomic remarks.</emphasis>
|
||
This species can be easily distinguished from all the other
|
||
<emphasis id="B965DA0AFF8CFFCFFB0EFF6AFB5DB5B0" box="[1180,1253,152,175]" italics="true" pageId="20" pageNumber="63">Serina</emphasis>
|
||
species by the presence of two columellar teeth, one larger, occupying almost 1/2 whorl and can usually be partially observed through aperture, another one much smaller but distinct, just beneath the larger tooth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |