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<document id="739E25C4708A2FC13F9ACACCC0716F76" ID-DOI="10.25674/so95iss1id311" ID-ISSN="2509-9523" ID-Zenodo-Dep="10724236" checkinTime="1709147239332" checkinUser="felipe" docAuthor="Frisch, Johannes &amp; Narakusumo, Raden Pramesa" docDate="2022" docId="03B7C77AFFD9FFE8BF0AFB700E550699" docLanguage="en" docName="SoilOrg.95.1.23-73.pdf" docOrigin="Soil Organisms 95 (1)" docSource="http://dx.doi.org/10.25674/so95iss1id311" docStyle="DocumentStyle:EA2ECAC87EC21C07AB805D5B9B4601F2:SoilOrg.2018-.journal_article" docStyleId="EA2ECAC87EC21C07AB805D5B9B4601F2" docStyleName="SoilOrg.2018-.journal_article" docTitle="Scopaeus elegantulus Cameron 1930" docType="treatment" docVersion="2" lastPageNumber="51" masterDocId="FF8EBF02FFC0FFF4BC5BFFF80D1D040D" masterDocTitle="Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species" masterLastPageNumber="73" masterPageNumber="23" pageNumber="48" updateTime="1709147743548" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="D6BF7B78DBBB3F8600C5EBB163BC6466">Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species</mods:title>
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<mods:namePart id="9E59CD8F586015D15BA05DFE2851B292">Frisch, Johannes</mods:namePart>
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<mods:namePart id="842CCD5BD9FE5098AD9265095DD3E65E">Narakusumo, Raden Pramesa</mods:namePart>
<mods:affiliation id="AEA0C071BA440D6D27E3B2812F4EBEBA">MUseUm ZoologicUm Bogoriense, Research Center for Biosystematics and EvolUtion, National Research and Innovation Agency (BRIN), Jl. Raya Jakarta-Bogor KM 46, Cibinong, West Java, 16911, Indonesia</mods:affiliation>
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<mods:date id="55C6DC4351CED34DFB09B4ED1C9A4F00">2022</mods:date>
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<treatment id="03B7C77AFFD9FFE8BF0AFB700E550699" LSID="urn:lsid:plazi:treatment:03B7C77AFFD9FFE8BF0AFB700E550699" httpUri="http://treatment.plazi.org/id/03B7C77AFFD9FFE8BF0AFB700E550699" lastPageId="28" lastPageNumber="51" pageId="25" pageNumber="48">
<subSubSection id="C30425E7FFD9FFEDBF0AFB70083E00AF" box="[849,1315,1160,1187]" pageId="25" pageNumber="48" type="nomenclature">
<paragraph id="8BA1766CFFD9FFEDBF0AFB70083E00AF" blockId="25.[849,1315,1160,1220]" box="[849,1315,1160,1187]" pageId="25" pageNumber="48">
<heading id="D0E9C100FFD9FFEDBF0AFB70083E00AF" box="[849,1315,1160,1187]" fontSize="11" level="2" pageId="25" pageNumber="48" reason="5">
<taxonomicName id="4C1E0DEFFFD9FFEDBF0AFB70083E00AF" ID-CoL="6XXCD" authority="Cameron, 1930" authorityName="Cameron" authorityYear="1930" box="[849,1315,1160,1187]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="48" phylum="Arthropoda" rank="species" species="elegantulus">
<emphasis id="B96AAA7EFFD9FFEDBF0AFB70083E00AF" bold="true" box="[849,1315,1160,1187]" pageId="25" pageNumber="48">
<emphasis id="B96AAA7EFFD9FFEDBF0AFB70094600AE" bold="true" box="[849,1115,1160,1187]" italics="true" pageId="25" pageNumber="48">Scopaeus elegantulus</emphasis>
<bibRefCitation id="EF8F0B9DFFD9FFEDB838FB70083E00AF" author="Cameron, M." box="[1123,1315,1160,1186]" pageId="25" pageNumber="48" pagination="325 - 348" refId="ref31003" refString="Cameron, M. (1930): Fauna Sumatrensis. Bijdrage No. 67, Staphylinidae (Col.). - Tijdschrift voor Entomologie 73: 325 - 348." type="journal article" year="1930">Cameron, 1930</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFD9FFEDBF0AFB5109F400CE" box="[849,1257,1193,1220]" pageId="25" pageNumber="48" type="description">
<paragraph id="8BA1766CFFD9FFEDBF0AFB5109F400CE" blockId="25.[849,1315,1160,1220]" box="[849,1257,1193,1220]" pageId="25" pageNumber="48">
(
<figureCitation id="13256AE9FFD9FFEDBF02FB510EAF00CE" box="[857,946,1193,1219]" captionStart="Figures 17–22" captionStartId="11.[189,262,1908,1930]" captionTargetBox="[279,1382,235,1877]" captionTargetId="figure-90@11.[206,1416,206,1889]" captionTargetPageId="11" captionText="Figures 17–22. Habitus of Scopaeusvelifer, paratype,Banten, Lebak:Citorek Kidul (17); S. elegantulus, Jawa Barat, Sukabumi:Djampang Kulon (18); S. uncinatus, paratype, Jawa Barat, Sukabumi: Cidahu (19); S. bipenicillatus, paratype, Sulawesi Tengah, Sigi: Pagana (20); S. bipectenatus, paratype, Sulawesi Tengah: Lake Poso (21); S. crassipunctatus, paratype, Jawa Barat, Sukabumi: Sirnarasa (22)." figureDoi="http://doi.org/10.5281/zenodo.10724250" httpUri="https://zenodo.org/record/10724250/files/figure.png" pageId="25" pageNumber="48">Figs 18</figureCitation>
,
<figureCitation id="13256AE9FFD9FFEDBFE4FB510EC000CE" box="[959,989,1193,1219]" captionStart="Figures 42–49" captionStartId="17.[190,263,1886,1908]" captionTargetBox="[263,1341,889,1832]" captionTargetId="figure-379@17.[189,1436,851,1867]" captionTargetPageId="17" captionText="Figures 42–49. Aedeagus in lateral view of Scopaeus velifer, holotype, Banten, Lebak: Citorek Kidul (42); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (43); S. uncinatus, holotype,Banten, Lebak:Citorek Kidul (44); S. bipenicillatus, holotype,Sulawesi Tengah, Sigi: Pagana (45); S. bipectenatus, holotype, Sulawesi Tengah, Poso: Lake Poso (46); S. cuspidatus, holotype, Bali, Jembrana: Cekik (47); S. crassipunctatus, holotype, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (48); S. sulawesianus, holotype, Sulawesi Selatan: Engrekang (49)." figureDoi="http://doi.org/10.5281/zenodo.10724262" httpUri="https://zenodo.org/record/10724262/files/figure.png" pageId="25" pageNumber="48">43</figureCitation>
,
<figureCitation id="13256AE9FFD9FFEDBFB2FB51092600CE" box="[1001,1083,1193,1220]" captionStart="Figures 92–97" captionStartId="28.[151,224,1886,1908]" captionTargetBox="[223,1329,734,1834]" captionTargetId="figure-235@28.[151,1398,685,1867]" captionTargetPageId="28" captionText="Figures 92–97. Apex of aedeagus in lateral (92, 95), ventral (93, 96), dorsal view (94, 97) of Scopaeus cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (92–94); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa: Cisarua (95– 97). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: ventral tooth of dextral apical lobe (I), lateral ridges of median foramen (II, III)." figureDoi="http://doi.org/10.5281/zenodo.10724282" httpUri="https://zenodo.org/record/10724282/files/figure.png" pageId="25" pageNumber="48">95 – 97</figureCitation>
, 136, 170, 171)
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFD9FFEDBF0AFB1508460108" box="[849,1371,1261,1285]" pageId="25" pageNumber="48" type="reference_group">
<paragraph id="8BA1766CFFD9FFEDBF0AFB1508460108" blockId="25.[849,1371,1261,1285]" box="[849,1371,1261,1285]" pageId="25" pageNumber="48">
<treatmentCitationGroup id="AB0E5142FFD9FFEDBF0AFB1508460108" box="[849,1371,1261,1285]" pageId="25" pageNumber="48">
<taxonomicName id="4C1E0DEFFFD9FFEDBF0AFB15092F0108" authorityName="Cameron" authorityYear="1930" box="[849,1074,1261,1285]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="48" phylum="Arthropoda" rank="species" species="elegantulu">
<emphasis id="B96AAA7EFFD9FFEDBF0AFB15092F0108" box="[849,1074,1261,1285]" italics="true" pageId="25" pageNumber="48">Scopaeus elegantulu</emphasis>
</taxonomicName>
s
<treatmentCitation id="0ABF507DFFD9FFEDB81FFB15083E0108" author="Cameron, M." box="[1092,1315,1261,1285]" page="346" pageId="25" pageNumber="48" year="1930">
<bibRefCitation id="EF8F0B9DFFD9FFEDB81FFB15083E0108" author="Cameron, M." box="[1092,1315,1261,1285]" pageId="25" pageNumber="48" pagination="325 - 348" refId="ref31003" refString="Cameron, M. (1930): Fauna Sumatrensis. Bijdrage No. 67, Staphylinidae (Col.). - Tijdschrift voor Entomologie 73: 325 - 348." type="journal article" year="1930">Cameron, 1930: 346</bibRefCitation>
</treatmentCitation>
, 347.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFD9FFEEBF0AFAD70E6306FA" lastPageId="26" lastPageNumber="49" pageId="25" pageNumber="48" type="materials_examined">
<paragraph id="8BA1766CFFD9FFEDBF0AFAD7091C02DC" blockId="25.[826,1437,1326,1976]" pageId="25" pageNumber="48">
<emphasis id="B96AAA7EFFD9FFEDBF0AFAD7098D014B" bold="true" box="[849,1168,1326,1351]" pageId="25" pageNumber="48">Type specimens examined:</emphasis>
<typeStatus id="54A5C8CEFFD9FFEDB8C7FAD6081C014B" box="[1180,1281,1326,1350]" pageId="25" pageNumber="48" type="holotype">Holotype</typeStatus>
<specimenCount id="9D18BDE5FFD9FFEDB956FAD7083C014B" box="[1293,1313,1327,1350]" count="1" pageId="25" pageNumber="48" type="male">♂</specimenCount>
,
<collectingCountry id="F30936FCFFD9FFEDB969FAD60885014B" box="[1330,1432,1326,1350]" name="Malaysia" pageId="25" pageNumber="48">Malaysia</collectingCountry>
,
<collectingRegion id="49DAB88EFFD9FFEDBF61FAB70E93016A" box="[826,910,1359,1383]" country="Malaysia" name="Pahang" pageId="25" pageNumber="48">Pahang</collectingRegion>
: Raub; labeled ‘Type’ (red edged, round, printed label), ‘Raub /
<collectingRegion id="49DAB88EFFD9FFEDB819FA88098F0185" box="[1090,1170,1392,1416]" country="Malaysia" name="Pahang" pageId="25" pageNumber="48">Pahang</collectingRegion>
F.M.S. / Dr. Cameron’ (printed), ‘S. /
<taxonomicName id="4C1E0DEFFFD9FFEDBF80FA69098401A4" authority="Cam." authorityName="Cameron" authorityYear="1930" box="[987,1177,1425,1449]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="48" phylum="Arthropoda" rank="species" species="elegantulus">elegantulus Cam.</taxonomicName>
/ TYPE’ (handwritten), ‘M. Cameron / Bequest / B.M.1955-147.’ (printed), ‘NHMUK014663052’ (printed with QR-code), ‘
<typeStatus id="54A5C8CEFFD9FFEDB968FA2B088101E6" box="[1331,1436,1491,1515]" pageId="25" pageNumber="48" type="holotype">Holotype</typeStatus>
/
<taxonomicName id="4C1E0DEFFFD9FFEDBF15FA0C095D0201" authorityName="Cameron" authorityYear="1930" box="[846,1088,1524,1548]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="48" phylum="Arthropoda" rank="species" species="elegantulus">
<emphasis id="B96AAA7EFFD9FFEDBF15FA0C095D0201" box="[846,1088,1524,1548]" italics="true" pageId="25" pageNumber="48">Scopaeus elegantulus</emphasis>
</taxonomicName>
/
<bibRefCitation id="EF8F0B9DFFD9FFEDB83BFA0C08190201" author="Cameron, M." box="[1120,1284,1524,1548]" pageId="25" pageNumber="48" pagination="325 - 348" refId="ref31003" refString="Cameron, M. (1930): Fauna Sumatrensis. Bijdrage No. 67, Staphylinidae (Col.). - Tijdschrift voor Entomologie 73: 325 - 348." type="journal article" year="1930">Cameron 1930</bibRefCitation>
/ label by J.
<bibRefCitation id="EF8F0B9DFFD9FFEDBF61F9ED0ED70220" author="Frisch, J. &amp; T. Mainda" box="[826,970,1557,1581]" pageId="25" pageNumber="48" pagination="139 - 147" refId="ref31869" refString="Frisch, J. &amp; T. Mainda (2022): The Scopaeus kokodanus species group (Coleoptera: Staphylinidae: Paederinae) from New Guinea and the Solomon Islands, with description of three new species. - Soil Organisms 94 (3): 139 - 147." type="journal article" year="2022">Frisch, 2022</bibRefCitation>
’ (red, printed) (
<collectionCode id="ED0FEEA9FFD9FFEDB8DBF9ED09CB0220" box="[1152,1238,1557,1581]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="25" pageNumber="48">NHML</collectionCode>
).
<typeStatus id="54A5C8CEFFD9FFEDB8B6F9ED08410220" box="[1261,1372,1557,1581]" pageId="25" pageNumber="48" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D18BDE5FFD9FFEDB930F9ED088A0221" box="[1387,1431,1557,1581]" count="4" pageId="25" pageNumber="48" type="male">4 ♂</specimenCount>
, labeled ‘Raub /
<collectingRegion id="49DAB88EFFD9FFEDBFBEF9CE09280243" box="[997,1077,1590,1614]" country="Malaysia" name="Pahang" pageId="25" pageNumber="48">Pahang</collectingRegion>
F.M.S. / Dr. Cameron’ (printed), ‘M. Cameron / Bequest / B.M.1955-147.’ (printed), ‘
<typeStatus id="54A5C8CEFFD9FFEDBF65F98F0EBE0282" box="[830,931,1655,1679]" pageId="25" pageNumber="48" type="paratype">Paratype</typeStatus>
/
<taxonomicName id="4C1E0DEFFFD9FFEDBF93F98009A10282" authorityName="Cameron" authorityYear="1930" box="[968,1212,1655,1679]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="48" phylum="Arthropoda" rank="species" species="elegantulus">
<emphasis id="B96AAA7EFFD9FFEDBF93F98009A10282" box="[968,1212,1655,1679]" italics="true" pageId="25" pageNumber="48">Scopaeus elegantulus</emphasis>
</taxonomicName>
/
<bibRefCitation id="EF8F0B9DFFD9FFEDB8BBF98F089B0282" author="Cameron, M." box="[1248,1414,1655,1679]" pageId="25" pageNumber="48" pagination="325 - 348" refId="ref31003" refString="Cameron, M. (1930): Fauna Sumatrensis. Bijdrage No. 67, Staphylinidae (Col.). - Tijdschrift voor Entomologie 73: 325 - 348." type="journal article" year="1930">Cameron 1930</bibRefCitation>
/ label by J.
<bibRefCitation id="EF8F0B9DFFD9FFEDBFF6F960092502BD" author="Frisch, J. &amp; T. Mainda" box="[941,1080,1688,1712]" pageId="25" pageNumber="48" pagination="139 - 147" refId="ref31869" refString="Frisch, J. &amp; T. Mainda (2022): The Scopaeus kokodanus species group (Coleoptera: Staphylinidae: Paederinae) from New Guinea and the Solomon Islands, with description of three new species. - Soil Organisms 94 (3): 139 - 147." type="journal article" year="2022">Frisch, 2022</bibRefCitation>
’ (subsequent
<typeStatus id="54A5C8CEFFD9FFEDB891F961083702BC" box="[1226,1322,1689,1713]" pageId="25" pageNumber="48" type="paratype">paratype</typeStatus>
label, red, printed) (
<collectionCode id="ED0FEEA9FFD9FFEDBFFBF9410EEB02DC" box="[928,1014,1721,1745]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="25" pageNumber="48">NHML</collectionCode>
).
</paragraph>
<paragraph id="8BA1766CFFD9FFEEBF0AF9220CFA05A0" blockId="25.[826,1437,1326,1976]" lastBlockId="26.[151,762,241,759]" lastPageId="26" lastPageNumber="49" pageId="25" pageNumber="48">
Comment: In the original description of
<taxonomicName id="4C1E0DEFFFD9FFEDB96FF9220EA5031E" authorityName="Cameron" authorityYear="1930" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="48" phylum="Arthropoda" rank="species" species="elegantulus">
<emphasis id="B96AAA7EFFD9FFEDB96FF9220EA5031E" italics="true" pageId="25" pageNumber="48">Scopaeus elegantulus</emphasis>
</taxonomicName>
, the number of underlying specimens is missing, but
<bibRefCitation id="EF8F0B9DFFD9FFEDBFA9F8E409F90339" author="Cameron, M." box="[1010,1252,1820,1844]" pageId="25" pageNumber="48" pagination="325 - 348" refId="ref31003" refString="Cameron, M. (1930): Fauna Sumatrensis. Bijdrage No. 67, Staphylinidae (Col.). - Tijdschrift voor Entomologie 73: 325 - 348." type="journal article" year="1930">Cameron (1930: 347)</bibRefCitation>
wrote the type was from Raub and labeled it as the ‘type’. Thus, the specimen is a
<typeStatus id="54A5C8CEFFD9FFEDBF85F8A509200378" box="[990,1085,1885,1909]" pageId="25" pageNumber="48" type="holotype">holotype</typeStatus>
by original designation (
<collectionCode id="ED0FEEA9FFD9FFEDB900F8A60881037B" box="[1371,1436,1886,1910]" pageId="25" pageNumber="48">ICZN</collectionCode>
1999: Article 73.1.1.). In the same paper,
<bibRefCitation id="EF8F0B9DFFD9FFEDB962F8860E6003BA" author="Cameron, M." pageId="25" pageNumber="48" pagination="325 - 348" refId="ref31003" refString="Cameron, M. (1930): Fauna Sumatrensis. Bijdrage No. 67, Staphylinidae (Col.). - Tijdschrift voor Entomologie 73: 325 - 348." type="journal article" year="1930">Cameron (1930)</bibRefCitation>
several times termed
<typeStatus id="54A5C8CEFFD9FFEDB8D7F86709EB03BA" box="[1164,1270,1951,1975]" pageId="25" pageNumber="48" type="holotype">holotypes</typeStatus>
by monotypy (
<collectionCode id="ED0FEEA9FFDAFFEEBCC5FF0A0DC20507" box="[158,223,242,266]" pageId="26" pageNumber="49">ICZN</collectionCode>
1999: Article 73.1.2.) as ‘unique’, but not for
<taxonomicName id="4C1E0DEFFFDAFFEEBEBEFF0A0C080527" authorityName="Cameron" authorityYear="1930" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="49" phylum="Arthropoda" rank="species" species="elegantulus">
<emphasis id="B96AAA7EFFDAFFEEBEBEFF0A0C080527" italics="true" pageId="26" pageNumber="49">S. elegantulus</emphasis>
</taxonomicName>
, which is why I conclude that the species was described after more than
<specimenCount id="9D18BDE5FFDAFFEEBDA7FECC0F840546" box="[508,665,307,331]" count="1" pageId="26" pageNumber="49" type="generic">one specimen</specimenCount>
. Thus, I consider the four remaining specimens as
<typeStatus id="54A5C8CEFFDAFFEEBED3FEAD0FE80560" box="[648,757,341,365]" pageId="26" pageNumber="49" type="paratype">paratypes</typeStatus>
, because they originate from the type locality and the Cameron collection at NHML.
</paragraph>
<paragraph id="8BA1766CFFDAFFEEBCF5FE4F0E6306FA" blockId="26.[151,762,241,759]" lastBlockId="26.[788,1399,241,759]" pageId="26" pageNumber="49">
<emphasis id="B96AAA7EFFDAFFEEBCF5FE4F0CF605C2" bold="true" box="[174,491,439,463]" pageId="26" pageNumber="49">New Indonesian records:</emphasis>
<materialsCitation id="3B767C31FFDAFFEEBDA4FE4F0FA405FD" collectionCode="PWCM" collectorName="Wunderle" elevation="1994300" location="Cekik" municipality="Jembrana" pageId="26" pageNumber="49" specimenCount="1" stateProvince="Bali">
<collectingRegion id="49DAB88EFFDAFFEEBDA4FE4F0F2C05C2" box="[511,561,439,463]" country="Indonesia" name="Bali" pageId="26" pageNumber="49">Bali</collectingRegion>
:
<collectingMunicipality id="6BC5EC16FFDAFFEEBE13FE4F0FAB05C2" box="[584,694,439,463]" pageId="26" pageNumber="49">Jembrana</collectingMunicipality>
: env.
<location id="8EC120B7FFDAFFEEBCCCFE200DC005FD" LSID="urn:lsid:plazi:treatment:03B7C77AFFD9FFE8BF0AFB700E550699:8EC120B7FFDAFFEEBCCCFE200DC005FD" box="[151,221,472,496]" municipality="Jembrana" name="Cekik" pageId="26" pageNumber="49" stateProvince="Bali">Cekik</location>
,
<date id="FFA050ACFFDAFFEEBCB3FE200C4905FD" box="[232,340,472,496]" pageId="26" pageNumber="49" value="1994-06-20">
20.6.
<quantity id="4CE6DB89FFDAFFEEBD46FE200CBA05E2" box="[285,423,472,496]" metricMagnitude="6" metricUnit="m" metricValue="1.9943" pageId="26" pageNumber="49" unit="m" value="1994300.0">
<elevation id="0033915FFFDAFFEEBD46FE200CBA05E2" box="[285,423,472,496]" metricMagnitude="6" metricUnit="m" metricValue="1.9943" pageId="26" pageNumber="49" unit="m" value="1994300.0">1994, 300 m</elevation>
</quantity>
</date>
, leg.
<collectorName id="26EB13BAFFDAFFEEBDBBFE200F5605FD" box="[480,587,472,496]" pageId="26" pageNumber="49">Wunderle</collectorName>
(
<collectionCode id="ED0FEEA9FFDAFFEEBE01FE200FAF05FD" box="[602,690,472,496]" pageId="26" pageNumber="49">PWCM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B767C31FFDAFFEEBE9FFE200C54065F" collectingDate="2015-09-20" collectionCode="MFNB, MZB" collectorName="Frisch" county="Sukabumi" elevation="680" latitude="-6.860833" location="Cisarua" longLatPrecision="21" longitude="106.513336" municipality="Sirnarasa" pageId="26" pageNumber="49" specimenCount="1" stateProvince="Jawa Barat">
<collectingRegion id="49DAB88EFFDAFFEEBE9FFE200DCA061D" country="Indonesia" name="Jawa Barat" pageId="26" pageNumber="49">Jawa Barat</collectingRegion>
:
<collectingCounty id="62C00EE0FFDAFFEEBCBDFE000C44061D" box="[230,345,504,528]" pageId="26" pageNumber="49">Sukabumi</collectingCounty>
,
<collectingMunicipality id="6BC5EC16FFDAFFEEBD32FE000CC8061D" box="[361,469,504,528]" pageId="26" pageNumber="49">Sirnarasa</collectingMunicipality>
,
<location id="8EC120B7FFDAFFEEBDBFFE000F23061D" LSID="urn:lsid:plazi:treatment:03B7C77AFFD9FFE8BF0AFB700E550699:8EC120B7FFDAFFEEBDBFFE000F23061D" box="[484,574,504,528]" county="Sukabumi" latitude="-6.860833" longLatPrecision="21" longitude="106.513336" municipality="Sirnarasa" name="Cisarua" pageId="26" pageNumber="49" stateProvince="Jawa Barat">Cisarua</location>
:
<location id="8EC120B7FFDAFFEEBE16FE000FE4061D" LSID="urn:lsid:plazi:treatment:03B7C77AFFD9FFE8BF0AFB700E550699:8EC120B7FFDAFFEEBE16FE000FE4061D" box="[589,761,504,528]" county="Sukabumi" latitude="-6.860833" longLatPrecision="21" longitude="106.513336" municipality="Sirnarasa" name="Ciawitali River" pageId="26" pageNumber="49" stateProvince="Jawa Barat">Ciawitali River</location>
(
<geoCoordinate id="EE2A10ABFFDAFFEEBCFBFDE10C3C063C" box="[160,289,537,561]" degrees="06" direction="south" minutes="51" orientation="latitude" pageId="26" pageNumber="49" precision="15" seconds="39" value="-6.860833">06°51’39’’S</geoCoordinate>
,
<geoCoordinate id="EE2A10ABFFDAFFEEBD72FDE10CA6063C" box="[297,443,537,561]" degrees="106" direction="east" minutes="30" orientation="longitude" pageId="26" pageNumber="49" precision="15" seconds="48" value="106.513336">106°30’48’’E</geoCoordinate>
),
<quantity id="4CE6DB89FFDAFFEEBD90FDE10F12063C" box="[459,527,537,561]" metricMagnitude="2" metricUnit="m" metricValue="6.8" pageId="26" pageNumber="49" unit="m" value="680.0">
<elevation id="0033915FFFDAFFEEBD90FDE10F12063C" box="[459,527,537,561]" metricMagnitude="2" metricUnit="m" metricValue="6.8" pageId="26" pageNumber="49" unit="m" value="680.0">680 m</elevation>
</quantity>
,
<date id="FFA050ACFFDAFFEEBE4CFDE10F9F063C" box="[535,642,537,561]" pageId="26" pageNumber="49" value="2015-09-20">
<collectingDate id="EFE4A944FFDAFFEEBE4CFDE10F9F063C" box="[535,642,537,561]" pageId="26" pageNumber="49" value="2015-09-20">20.9.2015</collectingDate>
</date>
, leg.
<collectorName id="26EB13BAFFDAFFEEBEEEFDE10FE4063C" box="[693,761,537,561]" pageId="26" pageNumber="49">Frisch</collectorName>
(
<collectionCode id="ED0FEEA9FFDAFFEEBCC4FDC20DEF065F" box="[159,242,570,594]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="26" pageNumber="49">MFNB</collectionCode>
,
<collectionCode id="ED0FEEA9FFDAFFEEBD59FDC20C5C065F" box="[258,321,570,594]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="26" pageNumber="49" type="Museum">MZB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B767C31FFDAFFEEBD02FDC20C680699" collectingDate="1999-04-26" collectionCode="MZB" collectorName="Halimun-Salak NP" county="Sukabumi" elevation="950" location="Cikaniki River" municipality="Cikaniki" pageId="26" pageNumber="49" specimenCount="1" stateProvince="Jawa Barat">
<collectingCounty id="62C00EE0FFDAFFEEBD02FDC20CD1065F" box="[345,460,570,594]" pageId="26" pageNumber="49">Sukabumi</collectingCounty>
,
<collectingMunicipality id="6BC5EC16FFDAFFEEBD87FDC20F5C065F" box="[476,577,570,594]" pageId="26" pageNumber="49">Cikaniki</collectingMunicipality>
:
<location id="8EC120B7FFDAFFEEBE0BFDC20FE4065F" LSID="urn:lsid:plazi:treatment:03B7C77AFFD9FFE8BF0AFB700E550699:8EC120B7FFDAFFEEBE0BFDC20FE4065F" box="[592,761,570,594]" county="Sukabumi" municipality="Cikaniki" name="Cikaniki River" pageId="26" pageNumber="49" stateProvince="Jawa Barat">Cikaniki River</location>
(
<location id="8EC120B7FFDAFFEEBCC4FDA30C32067E" LSID="urn:lsid:plazi:treatment:03B7C77AFFD9FFE8BF0AFB700E550699:8EC120B7FFDAFFEEBCC4FDA30C32067E" box="[159,303,603,627]" county="Sukabumi" municipality="Cikaniki" name="Mt Halimun" pageId="26" pageNumber="49" stateProvince="Jawa Barat">Mt Halimun</location>
,
<collectorName id="26EB13BAFFDAFFEEBD64FDA30F05067E" box="[319,536,603,627]" pageId="26" pageNumber="49">Halimun-Salak NP</collectorName>
),
<quantity id="4CE6DB89FFDAFFEEBE74FDA30F66067E" box="[559,635,603,627]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="26" pageNumber="49" unit="m" value="950.0">
<elevation id="0033915FFFDAFFEEBE74FDA30F66067E" box="[559,635,603,627]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="26" pageNumber="49" unit="m" value="950.0">950 m</elevation>
</quantity>
,
<date id="FFA050ACFFDAFFEEBED1FDA30FE8067E" box="[650,757,603,627]" pageId="26" pageNumber="49" value="1999-04-26">
<collectingDate id="EFE4A944FFDAFFEEBED1FDA30FE8067E" box="[650,757,603,627]" pageId="26" pageNumber="49" value="1999-04-26">26.4.1999</collectingDate>
</date>
, leg. MZB (
<collectionCode id="ED0FEEA9FFDAFFEEBD76FD840C700699" box="[301,365,636,660]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="26" pageNumber="49" type="Museum">MZB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B767C31FFDAFFEEBDD5FD840FE806DB" collectingDate="2015-09-17" collectionCode="MZB" collectorName="Frisch" county="Sukabumi" elevation="1020" latitude="-6.7461114" location="Cikaniki River" longLatPrecision="21" longitude="106.540276" municipality="Cikaniki" pageId="26" pageNumber="49" specimenCount="1" stateProvince="Jawa Barat">
<collectingCounty id="62C00EE0FFDAFFEEBDD5FD840F1C0699" box="[398,513,636,660]" pageId="26" pageNumber="49">Sukabumi</collectingCounty>
,
<collectingMunicipality id="6BC5EC16FFDAFFEEBE40FD840F9D0699" box="[539,640,636,660]" pageId="26" pageNumber="49">Cikaniki</collectingMunicipality>
:
<location id="8EC120B7FFDAFFEEBEC2FD840DC906B8" LSID="urn:lsid:plazi:treatment:03B7C77AFFD9FFE8BF0AFB700E550699:8EC120B7FFDAFFEEBEC2FD840DC906B8" county="Sukabumi" latitude="-6.7461114" longLatPrecision="21" longitude="106.540276" municipality="Cikaniki" name="Cikaniki River" pageId="26" pageNumber="49" stateProvince="Jawa Barat">Cikaniki River</location>
(
<location id="8EC120B7FFDAFFEEBCBFFD650C6F06B8" LSID="urn:lsid:plazi:treatment:03B7C77AFFD9FFE8BF0AFB700E550699:8EC120B7FFDAFFEEBCBFFD650C6F06B8" box="[228,370,669,693]" county="Sukabumi" latitude="-6.7461114" longLatPrecision="21" longitude="106.540276" municipality="Cikaniki" name="Mt Halimun" pageId="26" pageNumber="49" stateProvince="Jawa Barat">Mt Halimun</location>
,
<location id="8EC120B7FFDAFFEEBD24FD650F3A06B8" LSID="urn:lsid:plazi:treatment:03B7C77AFFD9FFE8BF0AFB700E550699:8EC120B7FFDAFFEEBD24FD650F3A06B8" box="[383,551,669,693]" county="Sukabumi" latitude="-6.7461114" longLatPrecision="21" longitude="106.540276" municipality="Cikaniki" name="Halimun-Salak" pageId="26" pageNumber="49" stateProvince="Jawa Barat">Halimun-Salak</location>
NP), (
<geoCoordinate id="EE2A10ABFFDAFFEEBE28FD650FE806B8" box="[627,757,669,693]" degrees="06" direction="south" minutes="44" orientation="latitude" pageId="26" pageNumber="49" precision="15" seconds="46" value="-6.7461114">06°44’46’’S</geoCoordinate>
,
<geoCoordinate id="EE2A10ABFFDAFFEEBCCCFD460C3406DB" box="[151,297,702,726]" degrees="106" direction="east" minutes="32" orientation="longitude" pageId="26" pageNumber="49" precision="15" seconds="25" value="106.540276">106°32’25’’E</geoCoordinate>
),
<quantity id="4CE6DB89FFDAFFEEBD19FD460C8706D8" box="[322,410,702,726]" metricMagnitude="3" metricUnit="m" metricValue="1.02" pageId="26" pageNumber="49" unit="m" value="1020.0">
<elevation id="0033915FFFDAFFEEBD19FD460C8706D8" box="[322,410,702,726]" metricMagnitude="3" metricUnit="m" metricValue="1.02" pageId="26" pageNumber="49" unit="m" value="1020.0">1020 m</elevation>
</quantity>
,
<date id="FFA050ACFFDAFFEEBDF0FD460F0C06DB" box="[427,529,702,726]" pageId="26" pageNumber="49" value="2015-09-17">
<collectingDate id="EFE4A944FFDAFFEEBDF0FD460F0C06DB" box="[427,529,702,726]" pageId="26" pageNumber="49" value="2015-09-17">17.9.2015</collectingDate>
</date>
, leg.
<collectorName id="26EB13BAFFDAFFEEBE0EFD460F8406DB" box="[597,665,702,726]" pageId="26" pageNumber="49">Frisch</collectorName>
(
<collectionCode id="ED0FEEA9FFDAFFEEBEF6FD460FF006DB" box="[685,749,702,726]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="26" pageNumber="49" type="Museum">MZB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B767C31FFDAFFEEBCCCFD2709420527" collectingDate="2015-09-21" collectionCode="MFNB, MZB" collectorName="Frisch" county="Sukabumi" elevation="670" latitude="-6.8588886" location="Cimaja River" longLatPrecision="21" longitude="106.51833" municipality="Sirnarasa" pageId="26" pageNumber="49" specimenCount="1" stateProvince="Jawa Barat">
<collectingCounty id="62C00EE0FFDAFFEEBCCCFD270C1706FA" box="[151,266,735,759]" pageId="26" pageNumber="49">Sukabumi</collectingCounty>
,
<collectingMunicipality id="6BC5EC16FFDAFFEEBD7DFD270C8E06FA" box="[294,403,735,759]" pageId="26" pageNumber="49">Sirnarasa</collectingMunicipality>
:
<location id="8EC120B7FFDAFFEEBDF5FD270F4C06FA" LSID="urn:lsid:plazi:treatment:03B7C77AFFD9FFE8BF0AFB700E550699:8EC120B7FFDAFFEEBDF5FD270F4C06FA" box="[430,593,735,759]" county="Sukabumi" latitude="-6.8588886" longLatPrecision="21" longitude="106.51833" municipality="Sirnarasa" name="Cimaja River" pageId="26" pageNumber="49" stateProvince="Jawa Barat">Cimaja River</location>
(S-slope
<location id="8EC120B7FFDAFFEEBE80FD270E650504" LSID="urn:lsid:plazi:treatment:03B7C77AFFD9FFE8BF0AFB700E550699:8EC120B7FFDAFFEEBE80FD270E650504" box="[731,888,241,758]" county="Sukabumi" latitude="-6.8588886" longLatPrecision="21" longitude="106.51833" municipality="Sirnarasa" name="Mt Halimun" pageId="26" pageNumber="49" stateProvince="Jawa Barat">Mt Halimun</location>
), (
<geoCoordinate id="EE2A10ABFFDAFFEEBFCAFF09090F0504" box="[913,1042,241,265]" degrees="06" direction="south" minutes="51" orientation="latitude" pageId="26" pageNumber="49" precision="15" seconds="32" value="-6.8588886">06°51’32’’S</geoCoordinate>
,
<geoCoordinate id="EE2A10ABFFDAFFEEB840FF0909B60504" box="[1051,1195,241,265]" degrees="106" direction="east" minutes="31" orientation="longitude" pageId="26" pageNumber="49" precision="15" seconds="06" value="106.51833">106°31’06’’E</geoCoordinate>
),
<quantity id="4CE6DB89FFDAFFEEB8E6FF09081C0504" box="[1213,1281,241,265]" metricMagnitude="2" metricUnit="m" metricValue="6.7" pageId="26" pageNumber="49" unit="m" value="670.0">
<elevation id="0033915FFFDAFFEEB8E6FF09081C0504" box="[1213,1281,241,265]" metricMagnitude="2" metricUnit="m" metricValue="6.7" pageId="26" pageNumber="49" unit="m" value="670.0">670 m</elevation>
</quantity>
,
<date id="FFA050ACFFDAFFEEB950FF09086F0504" box="[1291,1394,241,265]" pageId="26" pageNumber="49" value="2015-09-21">
<collectingDate id="EFE4A944FFDAFFEEB950FF09086F0504" box="[1291,1394,241,265]" pageId="26" pageNumber="49" value="2015-09-21">21.9.2015</collectingDate>
</date>
, leg.
<collectorName id="26EB13BAFFDAFFEEBF15FEEA0E8F0527" box="[846,914,274,298]" pageId="26" pageNumber="49">Frisch</collectorName>
(
<collectionCode id="ED0FEEA9FFDAFFEEBFF5FEEA091C0527" box="[942,1025,274,298]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="26" pageNumber="49">MFNB</collectionCode>
,
<collectionCode id="ED0FEEA9FFDAFFEEB843FEEA094A0527" box="[1048,1111,274,298]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="26" pageNumber="49" type="Museum">MZB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B767C31FFDAFFEEB82EFEEA09F20561" collectingDate="2015-09-22" collectionCode="MFNB, MZB" collectorName="Frisch" county="Sukabumi" elevation="1190" latitude="-6.828889" location="Halimun-Salak" longLatPrecision="21" longitude="106.50056" municipality="Simaresmi" pageId="26" pageNumber="49" specimenCount="1" stateProvince="Jawa Barat">
<collectingCounty id="62C00EE0FFDAFFEEB82EFEEA09F40527" box="[1141,1257,274,298]" pageId="26" pageNumber="49">Sukabumi</collectingCounty>
:
<collectingMunicipality id="6BC5EC16FFDAFFEEB95BFEEA086B0527" box="[1280,1398,274,298]" pageId="26" pageNumber="49">Simaresmi</collectingMunicipality>
(
<location id="8EC120B7FFDAFFEEBF40FECB0ED80546" LSID="urn:lsid:plazi:treatment:03B7C77AFFD9FFE8BF0AFB700E550699:8EC120B7FFDAFFEEBF40FECB0ED80546" box="[795,965,307,331]" county="Sukabumi" latitude="-6.828889" longLatPrecision="21" longitude="106.50056" municipality="Simaresmi" name="Halimun-Salak" pageId="26" pageNumber="49" stateProvince="Jawa Barat">Halimun-Salak</location>
NP), (
<geoCoordinate id="EE2A10ABFFDAFFEEB855FECB098D0546" box="[1038,1168,307,331]" degrees="06" direction="south" minutes="49" orientation="latitude" pageId="26" pageNumber="49" precision="15" seconds="44" value="-6.828889">06°49’44’’S</geoCoordinate>
,
<geoCoordinate id="EE2A10ABFFDAFFEEB8C7FECB08320546" box="[1180,1327,307,331]" degrees="106" direction="east" minutes="30" orientation="longitude" pageId="26" pageNumber="49" precision="15" seconds="02" value="106.50056">106°30’02’’E</geoCoordinate>
),
<quantity id="4CE6DB89FFDAFFEEB918FECB0E310561" metricMagnitude="3" metricUnit="m" metricValue="1.19" pageId="26" pageNumber="49" unit="m" value="1190.0">
<elevation id="0033915FFFDAFFEEB918FECB0E310561" metricMagnitude="3" metricUnit="m" metricValue="1.19" pageId="26" pageNumber="49" unit="m" value="1190.0">1190 m</elevation>
</quantity>
,
<date id="FFA050ACFFDAFFEEBF66FEAC0EBA0561" box="[829,935,340,364]" pageId="26" pageNumber="49" value="2015-09-22">
<collectingDate id="EFE4A944FFDAFFEEBF66FEAC0EBA0561" box="[829,935,340,364]" pageId="26" pageNumber="49" value="2015-09-22">22.9.2015</collectingDate>
</date>
, leg.
<collectorName id="26EB13BAFFDAFFEEBFB7FEAC092D0561" box="[1004,1072,340,364]" pageId="26" pageNumber="49">Frisch</collectorName>
(
<collectionCode id="ED0FEEA9FFDAFFEEB81FFEAC098A0561" box="[1092,1175,340,364]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="26" pageNumber="49">MFNB</collectionCode>
,
<collectionCode id="ED0FEEA9FFDAFFEEB8F3FEAC09FA0561" box="[1192,1255,340,364]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="26" pageNumber="49" type="Museum">MZB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B767C31FFDAFFEEB8A4FEAC09F705A3" collectingDate="2015-09-24" collectionCode="MFNB" collectorName="Frisch" county="Sukabumi" elevation="710" latitude="-7.1408334" location="Ciletuh River" longLatPrecision="21" longitude="106.62945" municipality="Kiara Dua" pageId="26" pageNumber="49" specimenCount="1" stateProvince="Jawa Barat">
<collectingCounty id="62C00EE0FFDAFFEEB8A4FEAC086F0561" box="[1279,1394,340,364]" pageId="26" pageNumber="49">Sukabumi</collectingCounty>
,
<collectingMunicipality id="6BC5EC16FFDAFFEEBF4FFE8D0E930580" box="[788,910,373,397]" pageId="26" pageNumber="49">Kiara Dua</collectingMunicipality>
:
<location id="8EC120B7FFDAFFEEBFC0FE8D092E0580" LSID="urn:lsid:plazi:treatment:03B7C77AFFD9FFE8BF0AFB700E550699:8EC120B7FFDAFFEEBFC0FE8D092E0580" box="[923,1075,373,397]" county="Sukabumi" latitude="-7.1408334" longLatPrecision="21" longitude="106.62945" municipality="Kiara Dua" name="Ciletuh River" pageId="26" pageNumber="49" stateProvince="Jawa Barat">Ciletuh River</location>
(
<geoCoordinate id="EE2A10ABFFDAFFEEB81EFE8D09D60580" box="[1093,1227,373,397]" degrees="07" direction="south" minutes="08" orientation="latitude" pageId="26" pageNumber="49" precision="15" seconds="27" value="-7.1408334">07°08’27’’S</geoCoordinate>
,
<geoCoordinate id="EE2A10ABFFDAFFEEB882FE8D08770580" box="[1241,1386,373,397]" degrees="106" direction="east" minutes="37" orientation="longitude" pageId="26" pageNumber="49" precision="15" seconds="46" value="106.62945">106°37’46’’E</geoCoordinate>
),
<quantity id="4CE6DB89FFDAFFEEBF4FFE6E0E7E05A0" box="[788,867,406,430]" metricMagnitude="2" metricUnit="m" metricValue="7.1" pageId="26" pageNumber="49" unit="m" value="710.0">
<elevation id="0033915FFFDAFFEEBF4FFE6E0E7E05A0" box="[788,867,406,430]" metricMagnitude="2" metricUnit="m" metricValue="7.1" pageId="26" pageNumber="49" unit="m" value="710.0">710 m</elevation>
</quantity>
,
<date id="FFA050ACFFDAFFEEBF22FE6E0EFE05A3" box="[889,995,406,430]" pageId="26" pageNumber="49" value="2015-09-24">
<collectingDate id="EFE4A944FFDAFFEEBF22FE6E0EFE05A3" box="[889,995,406,430]" pageId="26" pageNumber="49" value="2015-09-24">24.9.2015</collectingDate>
</date>
, leg.
<collectorName id="26EB13BAFFDAFFEEB86AFE6E096805A3" box="[1073,1141,406,430]" pageId="26" pageNumber="49">Frisch</collectorName>
(
<collectionCode id="ED0FEEA9FFDAFFEEB8D5FE6E09FC05A3" box="[1166,1249,406,430]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="26" pageNumber="49">MFNB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B767C31FFDAFFEEB8A4FE6E0EB005C2" collectorName="N Djampang" location="Sukabumi" pageId="26" pageNumber="49" specimenCount="1" stateProvince="Jawa Barat">
<location id="8EC120B7FFDAFFEEB8A4FE6E086F05A3" LSID="urn:lsid:plazi:treatment:03B7C77AFFD9FFE8BF0AFB700E550699:8EC120B7FFDAFFEEB8A4FE6E086F05A3" box="[1279,1394,406,430]" name="Sukabumi" pageId="26" pageNumber="49" stateProvince="Jawa Barat">Sukabumi</location>
,
<collectorName id="26EB13BAFFDAFFEEBF4FFE4F0EB005C2" box="[788,941,439,463]" pageId="26" pageNumber="49">N Djampang</collectorName>
</materialsCitation>
<materialsCitation id="3B767C31FFDAFFEEBF9BFE4F0E46061C" collectingDate="2015-09-25" collectionCode="MFNB, MZB" collectorName="Frisch" elevation="250" latitude="-7.2341666" location="Cikarang River" longLatPrecision="21" longitude="106.61361" pageId="26" pageNumber="49" specimenCount="1" stateProvince="Mjini Magharibi">
<collectingRegion id="49DAB88EFFDAFFEEBF9BFE4F091505C2" box="[960,1032,439,463]" country="Tanzania" name="Mjini Magharibi" pageId="26" pageNumber="49">Kulon</collectingRegion>
:
<location id="8EC120B7FFDAFFEEB844FE4F09C805C2" LSID="urn:lsid:plazi:treatment:03B7C77AFFD9FFE8BF0AFB700E550699:8EC120B7FFDAFFEEB844FE4F09C805C2" box="[1055,1237,439,463]" latitude="-7.2341666" longLatPrecision="21" longitude="106.61361" name="Cikarang River" pageId="26" pageNumber="49" stateProvince="Mjini Magharibi">Cikarang River</location>
(
<geoCoordinate id="EE2A10ABFFDAFFEEB8AAFE4F086F05C2" box="[1265,1394,439,463]" degrees="07" direction="south" minutes="14" orientation="latitude" pageId="26" pageNumber="49" precision="15" seconds="03" value="-7.2341666">07°14’03’’S</geoCoordinate>
,
<geoCoordinate id="EE2A10ABFFDAFFEEBF4FFE200EB905FD" box="[788,932,472,496]" degrees="106" direction="east" minutes="36" orientation="longitude" pageId="26" pageNumber="49" precision="15" seconds="49" value="106.61361">106°36’49’’E</geoCoordinate>
),
<quantity id="4CE6DB89FFDAFFEEBFE6FE2F091405E2" box="[957,1033,471,495]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="26" pageNumber="49" unit="m" value="250.0">
<elevation id="0033915FFFDAFFEEBFE6FE2F091405E2" box="[957,1033,471,495]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="26" pageNumber="49" unit="m" value="250.0">250 m</elevation>
</quantity>
,
<date id="FFA050ACFFDAFFEEB842FE2F099E05E2" box="[1049,1155,471,495]" pageId="26" pageNumber="49" value="2015-09-25">
<collectingDate id="EFE4A944FFDAFFEEB842FE2F099E05E2" box="[1049,1155,471,495]" pageId="26" pageNumber="49" value="2015-09-25">25.9.2015</collectingDate>
</date>
, leg.
<collectorName id="26EB13BAFFDAFFEEB89CFE20081605FD" box="[1223,1291,472,496]" pageId="26" pageNumber="49">Frisch</collectorName>
(
<collectionCode id="ED0FEEA9FFDAFFEEB944FE20086F05FD" box="[1311,1394,472,496]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="26" pageNumber="49">MFNB</collectionCode>
,
<collectionCode id="ED0FEEA9FFDAFFEEBF4FFE010E4E061C" box="[788,851,505,529]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="26" pageNumber="49" type="Museum">MZB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B767C31FFDAFFEEBF33FE01086E061C" box="[872,1395,504,529]" collectionCode="SAMA" collectorName="Lea" location="Bogor" pageId="26" pageNumber="49" specimenCount="1" stateProvince="Kulon">
<location id="8EC120B7FFDAFFEEBF33FE010EB6061C" LSID="urn:lsid:plazi:treatment:03B7C77AFFD9FFE8BF0AFB700E550699:8EC120B7FFDAFFEEBF33FE010EB6061C" box="[872,939,505,529]" name="Bogor" pageId="26" pageNumber="49" stateProvince="Kulon">Bogor</location>
[Buitenzorg], leg.
<collectorName id="26EB13BAFFDAFFEEB8DEFE0109B3061D" box="[1157,1198,505,528]" pageId="26" pageNumber="49">Lea</collectorName>
&amp; wife (
<collectionCode id="ED0FEEA9FFDAFFEEB943FE010876061C" box="[1304,1387,505,529]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="26" pageNumber="49" type="Museum">SAMA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B767C31FFDAFFEEBF4FFDE2086F065F" collectingDate="2001-06-28" collectionCode="SMNS" collectorName="Baluran NP &amp; Bolm" elevation="50" location="Situbondo" pageId="26" pageNumber="49" specimenCount="1" stateProvince="Jawa Timur">
<collectingRegion id="49DAB88EFFDAFFEEBF4FFDE20E83063C" box="[788,926,537,561]" country="Indonesia" name="Jawa Timur" pageId="26" pageNumber="49">Jawa Timur</collectingRegion>
:
<location id="8EC120B7FFDAFFEEBFF7FDE1093D063C" LSID="urn:lsid:plazi:treatment:03B7C77AFFD9FFE8BF0AFB700E550699:8EC120B7FFDAFFEEBFF7FDE1093D063C" box="[940,1056,537,561]" name="Situbondo" pageId="26" pageNumber="49" stateProvince="Jawa Timur">Situbondo</location>
:
<locationDeviation id="9A136F4EFFDAFFEEB875FDE10817063C" box="[1070,1290,537,561]" location="Wonorejo" pageId="26" pageNumber="49">
<quantity id="4CE6DB89FFDAFFEEB875FDE1096B063C" box="[1070,1142,537,561]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="26" pageNumber="49" unit="km" value="15.0">15 km</quantity>
N Wonorejo
</locationDeviation>
(
<collectorName id="26EB13BAFFDAFFEEB946FDE10E27065F" pageId="26" pageNumber="49">Baluran NP</collectorName>
),
<quantity id="4CE6DB89FFDAFFEEBF06FDC20EBA065F" box="[861,935,570,594]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="26" pageNumber="49" unit="m" value="50.0">
<elevation id="0033915FFFDAFFEEBF06FDC20EBA065F" box="[861,935,570,594]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="26" pageNumber="49" unit="m" value="50.0">50 m</elevation>
</quantity>
, 24. –
<date id="FFA050ACFFDAFFEEBFA2FDC2097B065F" box="[1017,1126,570,594]" pageId="26" pageNumber="49" value="2001-06-28">
<collectingDate id="EFE4A944FFDAFFEEBFA2FDC2097B065F" box="[1017,1126,570,594]" pageId="26" pageNumber="49" value="2001-06-28">28.6.2001</collectingDate>
</date>
, leg.
<collectorName id="26EB13BAFFDAFFEEB89AFDC209E0065F" box="[1217,1277,570,594]" pageId="26" pageNumber="49">Bolm</collectorName>
(
<collectionCode id="ED0FEEA9FFDAFFEEB947FDC20877065F" box="[1308,1386,570,594]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="26" pageNumber="49">SMNS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B767C31FFDAFFEEBF4FFDA30E670699" collectionCode="NHML" collectorName="Louwerens" elevation="84" location="Tulungagung" pageId="26" pageNumber="49" specimenCount="1" stateProvince="Jawa Timur">
<location id="8EC120B7FFDAFFEEBF4FFDA30EB8067E" LSID="urn:lsid:plazi:treatment:03B7C77AFFD9FFE8BF0AFB700E550699:8EC120B7FFDAFFEEBF4FFDA30EB8067E" box="[788,933,603,627]" name="Tulungagung" pageId="26" pageNumber="49" stateProvince="Jawa Timur">Tulungagung</location>
[Toeloengagoeng],
<quantity id="4CE6DB89FFDAFFEEB8D8FDA309A2067E" box="[1155,1215,603,627]" metricMagnitude="1" metricUnit="m" metricValue="8.4" pageId="26" pageNumber="49" unit="m" value="84.0">
<elevation id="0033915FFFDAFFEEB8D8FDA309A2067E" box="[1155,1215,603,627]" metricMagnitude="1" metricUnit="m" metricValue="8.4" pageId="26" pageNumber="49" unit="m" value="84.0">84 m</elevation>
</quantity>
, leg.
<collectorName id="26EB13BAFFDAFFEEB8A6FDA4086B067E" box="[1277,1398,604,627]" pageId="26" pageNumber="49">Louwerens</collectorName>
(
<collectionCode id="ED0FEEA9FFDAFFEEBF47FD840E6E0699" box="[796,883,636,660]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="26" pageNumber="49">NHML</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B767C31FFDAFFEEBFD9FD8409AC06B8" collectingDate="2000-01-25" collectionCode="PKCC" collectorName="Bezdek" latitude="-0.8333333" location="Bukit Tigapuluh" longLatPrecision="1308" longitude="102.433334" pageId="26" pageNumber="49" specimenCount="1" stateProvince="Riau">
<collectingRegion id="49DAB88EFFDAFFEEBFD9FD840EA70699" box="[898,954,636,660]" country="Indonesia" name="Riau" pageId="26" pageNumber="49">Riau</collectingRegion>
:
<location id="8EC120B7FFDAFFEEBF98FD84096B0699" LSID="urn:lsid:plazi:treatment:03B7C77AFFD9FFE8BF0AFB700E550699:8EC120B7FFDAFFEEBF98FD84096B0699" box="[963,1142,636,660]" latitude="-0.8333333" longLatPrecision="1308" longitude="102.433334" name="Bukit Tigapuluh" pageId="26" pageNumber="49" stateProvince="Riau">Bukit Tigapuluh</location>
NP (
<geoCoordinate id="EE2A10ABFFDAFFEEB8F0FD8409E50699" box="[1195,1272,636,660]" degrees="0" direction="south" minutes="50" orientation="latitude" pageId="26" pageNumber="49" precision="925" value="-0.8333333">0°50’S</geoCoordinate>
,
<geoCoordinate id="EE2A10ABFFDAFFEEB95AFD8408740699" box="[1281,1385,636,660]" degrees="102" direction="east" minutes="26" orientation="longitude" pageId="26" pageNumber="49" precision="925" value="102.433334">102°26’E</geoCoordinate>
), 18. –
<date id="FFA050ACFFDAFFEEBF12FD650EAB06B8" box="[841,950,669,693]" pageId="26" pageNumber="49" value="2000-01-25">
<collectingDate id="EFE4A944FFDAFFEEBF12FD650EAB06B8" box="[841,950,669,693]" pageId="26" pageNumber="49" value="2000-01-25">25.1.2000</collectingDate>
</date>
, leg.
<collectorName id="26EB13BAFFDAFFEEBFA3FD65095706B8" box="[1016,1098,669,693]" pageId="26" pageNumber="49">Bezdĕk</collectorName>
(
<collectionCode id="ED0FEEA9FFDAFFEEB806FD6509B406B8" box="[1117,1193,669,693]" pageId="26" pageNumber="49">PKCC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B767C31FFDAFFEEB8E4FD650E6706FA" collectionCode="NHML" collectorName="Jacobson" elevation="920" location="Fort de Kock" municipality="Bukittingii" pageId="26" pageNumber="49" specimenCount="1" stateProvince="Riau">
<taxonomicName id="4C1E0DEFFFDAFFEEB8E4FD65086E06B9" authority="Barat" authorityName="Barat" box="[1215,1395,669,693]" class="Insecta" family="Staphylinidae" genus="Sumatera" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="49" phylum="Arthropoda" rank="genus">Sumatera Barat</taxonomicName>
:
<collectingMunicipality id="6BC5EC16FFDAFFEEBF4FFD460E8F06DB" box="[788,914,702,726]" pageId="26" pageNumber="49">Bukittingii</collectingMunicipality>
:
<location id="8EC120B7FFDAFFEEBFC4FD46092A06DB" LSID="urn:lsid:plazi:treatment:03B7C77AFFD9FFE8BF0AFB700E550699:8EC120B7FFDAFFEEBFC4FD46092A06DB" box="[927,1079,702,726]" municipality="Bukittingii" name="Fort de Kock" pageId="26" pageNumber="49" stateProvince="Riau">Fort de Kock</location>
,
<quantity id="4CE6DB89FFDAFFEEB81FFD46099306D8" box="[1092,1166,702,726]" metricMagnitude="2" metricUnit="m" metricValue="9.2" pageId="26" pageNumber="49" unit="m" value="920.0">
<elevation id="0033915FFFDAFFEEB81FFD46099306D8" box="[1092,1166,702,726]" metricMagnitude="2" metricUnit="m" metricValue="9.2" pageId="26" pageNumber="49" unit="m" value="920.0">920 m</elevation>
</quantity>
, 1926, leg.
<collectorName id="26EB13BAFFDAFFEEB949FD46086B06DB" box="[1298,1398,702,726]" pageId="26" pageNumber="49">Jacobson</collectorName>
(
<collectionCode id="ED0FEEA9FFDAFFEEBF47FD270E6E06FA" box="[796,883,735,759]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="26" pageNumber="49">NHML</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF6126E4FFDAFFEEBCCCF8A40C2F03BA" ID-DOI="http://doi.org/10.5281/zenodo.10724276" ID-Zenodo-Dep="10724276" httpUri="https://zenodo.org/record/10724276/files/figure.png" pageId="26" pageNumber="49" startId="26.[151,224,1884,1906]" targetBox="[223,1322,801,1824]" targetPageId="26" targetType="figure">
<paragraph id="8BA1766CFFDAFFEEBCCCF8A40C2F03BA" blockId="26.[151,1399,1884,1975]" pageId="26" pageNumber="49">
<emphasis id="B96AAA7EFFDAFFEEBCCCF8A40C3B037F" bold="true" box="[151,294,1884,1906]" pageId="26" pageNumber="49">Figures 80–85.</emphasis>
Apex of aedeagus in lateral (
<emphasis id="B96AAA7EFFDAFFEEBE60F8A50F4F037F" bold="true" box="[571,594,1885,1906]" pageId="26" pageNumber="49">80</emphasis>
,
<emphasis id="B96AAA7EFFDAFFEEBE04F8A50F6B037F" bold="true" box="[607,630,1885,1906]" pageId="26" pageNumber="49">83</emphasis>
), ventral (
<emphasis id="B96AAA7EFFDAFFEEBE83F8A50FF2037F" bold="true" box="[728,751,1885,1906]" pageId="26" pageNumber="49">81</emphasis>
,
<emphasis id="B96AAA7EFFDAFFEEBEA7F8A50E0E037F" bold="true" box="[764,787,1885,1906]" pageId="26" pageNumber="49">84</emphasis>
), dorsal view (
<emphasis id="B96AAA7EFFDAFFEEBFFAF8A50EA5037F" bold="true" box="[929,952,1885,1906]" pageId="26" pageNumber="49">82</emphasis>
,
<emphasis id="B96AAA7EFFDAFFEEBF9FF8A50EC6037F" bold="true" box="[964,987,1885,1906]" pageId="26" pageNumber="49">85</emphasis>
) of
<taxonomicName id="4C1E0DEFFFDAFFEEB85FF8A509BB037F" authorityName="Frisch" authorityYear="2022" box="[1028,1190,1885,1906]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="49" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="B96AAA7EFFDAFFEEB85FF8A509BB037F" box="[1028,1190,1885,1906]" italics="true" pageId="26" pageNumber="49">Scopaeus grandis</emphasis>
</taxonomicName>
, holotype, Java Barat, Sukabumi: Cidahu (
<emphasis id="B96AAA7EFFDAFFEEBD14F88B0C930385" bold="true" box="[335,398,1907,1929]" pageId="26" pageNumber="49">80 – 82</emphasis>
);
<taxonomicName id="4C1E0DEFFFDAFFEEBDF9F88C0CE30385" authorityName="Frisch" authorityYear="2022" box="[418,510,1907,1929]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="49" phylum="Arthropoda" rank="species" species="pulcher">
<emphasis id="B96AAA7EFFDAFFEEBDF9F88C0CE30385" box="[418,510,1907,1929]" italics="true" pageId="26" pageNumber="49">S. pulcher</emphasis>
</taxonomicName>
, holotype, Jawa Barat, Sukabumi: Cikaniki (
<emphasis id="B96AAA7EFFDAFFEEBFC0F88B0EC70385" bold="true" box="[923,986,1907,1929]" pageId="26" pageNumber="49">83 – 85</emphasis>
).
<emphasis id="B96AAA7EFFDAFFEEBFB6F88B096B0384" bold="true" box="[1005,1142,1907,1929]" pageId="26" pageNumber="49">Abbreviations</emphasis>
:
<emphasis id="B96AAA7EFFDAFFEEB8D8F88B09AD0384" bold="true" box="[1155,1200,1907,1929]" pageId="26" pageNumber="49">al(d)</emphasis>
– dextral apical lobe,
<emphasis id="B96AAA7EFFDAFFEEBCCCF8720DDC03AD" bold="true" box="[151,193,1930,1952]" pageId="26" pageNumber="49">al(s)</emphasis>
– sinistral apical lobe,
<emphasis id="B96AAA7EFFDAFFEEBDCCF8720CB70392" bold="true" box="[407,426,1930,1951]" pageId="26" pageNumber="49">dl</emphasis>
– dorsal lobe,
<emphasis id="B96AAA7EFFDAFFEEBE69F8720F2703AD" bold="true" box="[562,570,1930,1952]" pageId="26" pageNumber="49">f</emphasis>
– flagellum,
<emphasis id="B96AAA7EFFDAFFEEBEEFF8720FDC0392" bold="true" box="[692,705,1930,1951]" pageId="26" pageNumber="49">ll</emphasis>
– lateral lobes,
<emphasis id="B96AAA7EFFDAFFEEBF08F8720E7003AD" bold="true" box="[851,877,1930,1952]" pageId="26" pageNumber="49">mf</emphasis>
– median foramen,
<emphasis id="B96AAA7EFFDAFFEEB87CF87209240392" bold="true" box="[1063,1081,1930,1951]" pageId="26" pageNumber="49">tr</emphasis>
– transverse ridge of median lobe,
<emphasis id="B96AAA7EFFDAFFEEBCCCF8590DB403BB" bold="true" box="[151,169,1953,1974]" pageId="26" pageNumber="49">vl</emphasis>
– ventral lobe.
</paragraph>
</caption>
<subSubSection id="C30425E7FFDBFFE8BC8FFF090839061C" lastPageId="28" lastPageNumber="51" pageId="27" pageNumber="50" type="description">
<paragraph id="8BA1766CFFDBFFEFBC8FFF09088D0580" blockId="27.[189,800,241,528]" lastBlockId="27.[826,1436,241,529]" pageId="27" pageNumber="50">
<emphasis id="B96AAA7EFFDBFFEFBC8FFF090C600504" bold="true" box="[212,381,241,265]" pageId="27" pageNumber="50">Redescription:</emphasis>
Habitus and coloring as in
<figureCitation id="13256AE9FFDBFFEFBEADFF090DC40527" captionStart="Figures 17–22" captionStartId="11.[189,262,1908,1930]" captionTargetBox="[279,1382,235,1877]" captionTargetId="figure-90@11.[206,1416,206,1889]" captionTargetPageId="11" captionText="Figures 17–22. Habitus of Scopaeusvelifer, paratype,Banten, Lebak:Citorek Kidul (17); S. elegantulus, Jawa Barat, Sukabumi:Djampang Kulon (18); S. uncinatus, paratype, Jawa Barat, Sukabumi: Cidahu (19); S. bipenicillatus, paratype, Sulawesi Tengah, Sigi: Pagana (20); S. bipectenatus, paratype, Sulawesi Tengah: Lake Poso (21); S. crassipunctatus, paratype, Jawa Barat, Sukabumi: Sirnarasa (22)." figureDoi="http://doi.org/10.5281/zenodo.10724250" httpUri="https://zenodo.org/record/10724250/files/figure.png" pageId="27" pageNumber="50">Fig. 18</figureCitation>
. Head subcircular. Penultimate antennal segment quadrate or slightly elongate. Mesotibia strongly thickened. Body surface shiny with fine, spacious, setose punctation; microreticulation absent. Pubescence of body surface short, erect. Body orange-brown to black-brown with reddish tinge, usually reddish dark brown with pronotum and abdomen somewhat lighter than head and elytra; tip of abdomen and posterior sixth of elytra yellow-brown; maxillary palpi light medium brown; antennae with scapus medium brown and pedicellus and subsequent segments more or less blackened, gradually lighter towards light brown terminal segments. Total body length
<quantity id="4CE6DB89FFDBFFEFBF9AFE8D09550580" box="[961,1096,373,397]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" metricValueMax="3.7" metricValueMin="3.1" pageId="27" pageNumber="50" unit="mm" value="3.4" valueMax="3.7" valueMin="3.1">3.1 – 3.7 mm</quantity>
; forebody length
<quantity id="4CE6DB89FFDBFFEFB95FFE8D08910580" box="[1284,1420,373,397]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="27" pageNumber="50" unit="mm" value="2.0" valueMax="2.1" valueMin="1.9">1.9 – 2.1 mm</quantity>
.
</paragraph>
<paragraph id="8BA1766CFFDBFFE8BF0AFE6E09870546" blockId="27.[826,1436,241,529]" lastBlockId="28.[786,1399,241,660]" lastPageId="28" lastPageNumber="51" pageId="27" pageNumber="50">
Male: Abdominal sternite VII with unmodified posterior margin. Abdominal sternite VIII in about posterior fifth with wide, triangular emargination straight in median fourth (Fig. 136). Aedeagus (
<figureCitation id="13256AE9FFDBFFEFB918FE00088A061C" box="[1347,1431,504,529]" captionStart="Figures 42–49" captionStartId="17.[190,263,1886,1908]" captionTargetBox="[263,1341,889,1832]" captionTargetId="figure-379@17.[189,1436,851,1867]" captionTargetPageId="17" captionText="Figures 42–49. Aedeagus in lateral view of Scopaeus velifer, holotype, Banten, Lebak: Citorek Kidul (42); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (43); S. uncinatus, holotype,Banten, Lebak:Citorek Kidul (44); S. bipenicillatus, holotype,Sulawesi Tengah, Sigi: Pagana (45); S. bipectenatus, holotype, Sulawesi Tengah, Poso: Lake Poso (46); S. cuspidatus, holotype, Bali, Jembrana: Cekik (47); S. crassipunctatus, holotype, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (48); S. sulawesianus, holotype, Sulawesi Selatan: Engrekang (49)." figureDoi="http://doi.org/10.5281/zenodo.10724262" httpUri="https://zenodo.org/record/10724262/files/figure.png" pageId="27" pageNumber="50">Figs 43</figureCitation>
,
<figureCitation id="13256AE9FFDCFFE8BCCCFF090DFE0507" box="[151,227,241,266]" captionStart="Figures 92–97" captionStartId="28.[151,224,1886,1908]" captionTargetBox="[223,1329,734,1834]" captionTargetId="figure-235@28.[151,1398,685,1867]" captionTargetPageId="28" captionText="Figures 92–97. Apex of aedeagus in lateral (92, 95), ventral (93, 96), dorsal view (94, 97) of Scopaeus cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (92–94); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa: Cisarua (95– 97). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: ventral tooth of dextral apical lobe (I), lateral ridges of median foramen (II, III)." figureDoi="http://doi.org/10.5281/zenodo.10724282" httpUri="https://zenodo.org/record/10724282/files/figure.png" pageId="28" pageNumber="51">95 – 97</figureCitation>
) about
<quantity id="4CE6DB89FFDCFFE8BD1BFF0A0C8E0504" box="[320,403,242,266]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="28" pageNumber="51" unit="mm" value="0.7">0.7 mm</quantity>
long, with short, asymmetrical distal lobes; apical lobes rectangular with wide, truncate apex and small, acute, apicoventral tooth (
<figureCitation id="13256AE9FFDCFFE8BE37FECB0FDC0546" box="[620,705,307,331]" captionStart="Figures 42–49" captionStartId="17.[190,263,1886,1908]" captionTargetBox="[263,1341,889,1832]" captionTargetId="figure-379@17.[189,1436,851,1867]" captionTargetPageId="17" captionText="Figures 42–49. Aedeagus in lateral view of Scopaeus velifer, holotype, Banten, Lebak: Citorek Kidul (42); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (43); S. uncinatus, holotype,Banten, Lebak:Citorek Kidul (44); S. bipenicillatus, holotype,Sulawesi Tengah, Sigi: Pagana (45); S. bipectenatus, holotype, Sulawesi Tengah, Poso: Lake Poso (46); S. cuspidatus, holotype, Bali, Jembrana: Cekik (47); S. crassipunctatus, holotype, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (48); S. sulawesianus, holotype, Sulawesi Selatan: Engrekang (49)." figureDoi="http://doi.org/10.5281/zenodo.10724262" httpUri="https://zenodo.org/record/10724262/files/figure.png" pageId="28" pageNumber="51">Figs 43</figureCitation>
,
<figureCitation id="13256AE9FFDCFFE8BE94FECB0FF00546" box="[719,749,307,331]" captionStart="Figures 92–97" captionStartId="28.[151,224,1886,1908]" captionTargetBox="[223,1329,734,1834]" captionTargetId="figure-235@28.[151,1398,685,1867]" captionTargetPageId="28" captionText="Figures 92–97. Apex of aedeagus in lateral (92, 95), ventral (93, 96), dorsal view (94, 97) of Scopaeus cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (92–94); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa: Cisarua (95– 97). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: ventral tooth of dextral apical lobe (I), lateral ridges of median foramen (II, III)." figureDoi="http://doi.org/10.5281/zenodo.10724282" httpUri="https://zenodo.org/record/10724282/files/figure.png" pageId="28" pageNumber="51">95</figureCitation>
), in ventral strongly curved distomediad and slightly overlapping distomedially (
<figureCitation id="13256AE9FFDCFFE8BD92FE8D0F3F0580" box="[457,546,373,397]" captionStart="Figures 92–97" captionStartId="28.[151,224,1886,1908]" captionTargetBox="[223,1329,734,1834]" captionTargetId="figure-235@28.[151,1398,685,1867]" captionTargetPageId="28" captionText="Figures 92–97. Apex of aedeagus in lateral (92, 95), ventral (93, 96), dorsal view (94, 97) of Scopaeus cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (92–94); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa: Cisarua (95– 97). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: ventral tooth of dextral apical lobe (I), lateral ridges of median foramen (II, III)." figureDoi="http://doi.org/10.5281/zenodo.10724282" httpUri="https://zenodo.org/record/10724282/files/figure.png" pageId="28" pageNumber="51">Figs 96</figureCitation>
); dorsal lobe stout, transverse, with apex emarginate (
<figureCitation id="13256AE9FFDCFFE8BE5EFE6E0F9C05A3" box="[517,641,406,430]" captionStart="Figures 92–97" captionStartId="28.[151,224,1886,1908]" captionTargetBox="[223,1329,734,1834]" captionTargetId="figure-235@28.[151,1398,685,1867]" captionTargetPageId="28" captionText="Figures 92–97. Apex of aedeagus in lateral (92, 95), ventral (93, 96), dorsal view (94, 97) of Scopaeus cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (92–94); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa: Cisarua (95– 97). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: ventral tooth of dextral apical lobe (I), lateral ridges of median foramen (II, III)." figureDoi="http://doi.org/10.5281/zenodo.10724282" httpUri="https://zenodo.org/record/10724282/files/figure.png" pageId="28" pageNumber="51">Figs 96, 97</figureCitation>
); flagellum short, thin, projecting from apical lobes ventrally as well as small, proximad curved tooth proximal of flagellum (
<figureCitation id="13256AE9FFDCFFE8BCC4FE000DE8061C" box="[159,245,504,529]" captionStart="Figures 42–49" captionStartId="17.[190,263,1886,1908]" captionTargetBox="[263,1341,889,1832]" captionTargetId="figure-379@17.[189,1436,851,1867]" captionTargetPageId="17" captionText="Figures 42–49. Aedeagus in lateral view of Scopaeus velifer, holotype, Banten, Lebak: Citorek Kidul (42); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (43); S. uncinatus, holotype,Banten, Lebak:Citorek Kidul (44); S. bipenicillatus, holotype,Sulawesi Tengah, Sigi: Pagana (45); S. bipectenatus, holotype, Sulawesi Tengah, Poso: Lake Poso (46); S. cuspidatus, holotype, Bali, Jembrana: Cekik (47); S. crassipunctatus, holotype, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (48); S. sulawesianus, holotype, Sulawesi Selatan: Engrekang (49)." figureDoi="http://doi.org/10.5281/zenodo.10724262" httpUri="https://zenodo.org/record/10724262/files/figure.png" pageId="28" pageNumber="51">Figs 43</figureCitation>
,
<figureCitation id="13256AE9FFDCFFE8BD58FE000C3C061D" box="[259,289,504,528]" captionStart="Figures 92–97" captionStartId="28.[151,224,1886,1908]" captionTargetBox="[223,1329,734,1834]" captionTargetId="figure-235@28.[151,1398,685,1867]" captionTargetPageId="28" captionText="Figures 92–97. Apex of aedeagus in lateral (92, 95), ventral (93, 96), dorsal view (94, 97) of Scopaeus cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (92–94); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa: Cisarua (95– 97). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: ventral tooth of dextral apical lobe (I), lateral ridges of median foramen (II, III)." figureDoi="http://doi.org/10.5281/zenodo.10724282" httpUri="https://zenodo.org/record/10724282/files/figure.png" pageId="28" pageNumber="51">95</figureCitation>
); setose lateral lobes absent (
<figureCitation id="13256AE9FFDCFFE8BE31FE000FF1061C" box="[618,748,504,529]" captionStart="Figures 92–97" captionStartId="28.[151,224,1886,1908]" captionTargetBox="[223,1329,734,1834]" captionTargetId="figure-235@28.[151,1398,685,1867]" captionTargetPageId="28" captionText="Figures 92–97. Apex of aedeagus in lateral (92, 95), ventral (93, 96), dorsal view (94, 97) of Scopaeus cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (92–94); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa: Cisarua (95– 97). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: ventral tooth of dextral apical lobe (I), lateral ridges of median foramen (II, III)." figureDoi="http://doi.org/10.5281/zenodo.10724282" httpUri="https://zenodo.org/record/10724282/files/figure.png" pageId="28" pageNumber="51">Figs 96, 97</figureCitation>
); ventral lobe about twice as long as wide with apex acute in lateral view (
<figureCitation id="13256AE9FFDCFFE8BD0EFDC20CAD065F" box="[341,432,570,594]" captionStart="Figures 42–49" captionStartId="17.[190,263,1886,1908]" captionTargetBox="[263,1341,889,1832]" captionTargetId="figure-379@17.[189,1436,851,1867]" captionTargetPageId="17" captionText="Figures 42–49. Aedeagus in lateral view of Scopaeus velifer, holotype, Banten, Lebak: Citorek Kidul (42); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (43); S. uncinatus, holotype,Banten, Lebak:Citorek Kidul (44); S. bipenicillatus, holotype,Sulawesi Tengah, Sigi: Pagana (45); S. bipectenatus, holotype, Sulawesi Tengah, Poso: Lake Poso (46); S. cuspidatus, holotype, Bali, Jembrana: Cekik (47); S. crassipunctatus, holotype, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (48); S. sulawesianus, holotype, Sulawesi Selatan: Engrekang (49)." figureDoi="http://doi.org/10.5281/zenodo.10724262" httpUri="https://zenodo.org/record/10724262/files/figure.png" pageId="28" pageNumber="51">Figs 43</figureCitation>
,
<figureCitation id="13256AE9FFDCFFE8BD98FDC20CFC065F" box="[451,481,570,594]" captionStart="Figures 92–97" captionStartId="28.[151,224,1886,1908]" captionTargetBox="[223,1329,734,1834]" captionTargetId="figure-235@28.[151,1398,685,1867]" captionTargetPageId="28" captionText="Figures 92–97. Apex of aedeagus in lateral (92, 95), ventral (93, 96), dorsal view (94, 97) of Scopaeus cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (92–94); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa: Cisarua (95– 97). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: ventral tooth of dextral apical lobe (I), lateral ridges of median foramen (II, III)." figureDoi="http://doi.org/10.5281/zenodo.10724282" httpUri="https://zenodo.org/record/10724282/files/figure.png" pageId="28" pageNumber="51">95</figureCitation>
), but deeply emarginate in ventral view (
<figureCitation id="13256AE9FFDCFFE8BD33FDA30CD9067E" box="[360,452,603,627]" captionStart="Figures 92–97" captionStartId="28.[151,224,1886,1908]" captionTargetBox="[223,1329,734,1834]" captionTargetId="figure-235@28.[151,1398,685,1867]" captionTargetPageId="28" captionText="Figures 92–97. Apex of aedeagus in lateral (92, 95), ventral (93, 96), dorsal view (94, 97) of Scopaeus cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (92–94); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa: Cisarua (95– 97). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: ventral tooth of dextral apical lobe (I), lateral ridges of median foramen (II, III)." figureDoi="http://doi.org/10.5281/zenodo.10724282" httpUri="https://zenodo.org/record/10724282/files/figure.png" pageId="28" pageNumber="51">Fig. 96</figureCitation>
); median foramen oblong with acute proximal end, occupying more than half of phallobase length (
<figureCitation id="13256AE9FFDCFFE8BF85FF0909330504" box="[990,1070,241,265]" captionStart="Figures 92–97" captionStartId="28.[151,224,1886,1908]" captionTargetBox="[223,1329,734,1834]" captionTargetId="figure-235@28.[151,1398,685,1867]" captionTargetPageId="28" captionText="Figures 92–97. Apex of aedeagus in lateral (92, 95), ventral (93, 96), dorsal view (94, 97) of Scopaeus cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (92–94); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa: Cisarua (95– 97). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: ventral tooth of dextral apical lobe (I), lateral ridges of median foramen (II, III)." figureDoi="http://doi.org/10.5281/zenodo.10724282" httpUri="https://zenodo.org/record/10724282/files/figure.png" pageId="28" pageNumber="51">Fig. 96</figureCitation>
), with transverse ridge weakly sclerotized and lateral ridges of median foramen bent up ventrad (
<figureCitation id="13256AE9FFDCFFE8BF28FECB0EEF0546" box="[883,1010,307,331]" captionStart="Figures 92–97" captionStartId="28.[151,224,1886,1908]" captionTargetBox="[223,1329,734,1834]" captionTargetId="figure-235@28.[151,1398,685,1867]" captionTargetPageId="28" captionText="Figures 92–97. Apex of aedeagus in lateral (92, 95), ventral (93, 96), dorsal view (94, 97) of Scopaeus cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (92–94); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa: Cisarua (95– 97). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: ventral tooth of dextral apical lobe (I), lateral ridges of median foramen (II, III)." figureDoi="http://doi.org/10.5281/zenodo.10724282" httpUri="https://zenodo.org/record/10724282/files/figure.png" pageId="28" pageNumber="51">Figs 95, 96</figureCitation>
: arrows
<collectionCode id="ED0FEEA9FFDCFFE8B815FECC097F0546" box="[1102,1122,308,331]" pageId="28" pageNumber="51">II</collectionCode>
,
<collectionCode id="ED0FEEA9FFDCFFE8B835FECB09900546" box="[1134,1165,307,331]" pageId="28" pageNumber="51">III</collectionCode>
).
</paragraph>
<caption id="DF6126E4FFDBFFEFBCE6F8BF0CF603B5" ID-DOI="http://doi.org/10.5281/zenodo.10724278" ID-Zenodo-Dep="10724278" httpUri="https://zenodo.org/record/10724278/files/figure.png" pageId="27" pageNumber="50" startId="27.[189,262,1863,1885]" targetBox="[261,1365,582,1806]" targetPageId="27" targetType="figure">
<paragraph id="8BA1766CFFDBFFEFBCE6F8BF0CF603B5" blockId="27.[189,1437,1863,1976]" pageId="27" pageNumber="50">
<emphasis id="B96AAA7EFFDBFFEFBCE6F8BF0C510351" bold="true" box="[189,332,1863,1885]" pageId="27" pageNumber="50">Figures 86–91.</emphasis>
Apex of aedeagus in lateral (
<emphasis id="B96AAA7EFFDBFFEFBE3CF8BF0F630351" bold="true" box="[615,638,1863,1884]" pageId="27" pageNumber="50">86</emphasis>
,
<emphasis id="B96AAA7EFFDBFFEFBED7F8BF0FBE0351" bold="true" box="[652,675,1863,1884]" pageId="27" pageNumber="50">89</emphasis>
), ventral (
<emphasis id="B96AAA7EFFDBFFEFBF5CF8BF0E030351" bold="true" box="[775,798,1863,1884]" pageId="27" pageNumber="50">87</emphasis>
,
<emphasis id="B96AAA7EFFDBFFEFBF70F8BF0E5F0351" bold="true" box="[811,834,1863,1884]" pageId="27" pageNumber="50">90</emphasis>
), dorsal view (
<emphasis id="B96AAA7EFFDBFFEFBF88F8BF0EF70351" bold="true" box="[979,1002,1863,1884]" pageId="27" pageNumber="50">88</emphasis>
,
<emphasis id="B96AAA7EFFDBFFEFBFA3F8BF09120351" bold="true" box="[1016,1039,1863,1884]" pageId="27" pageNumber="50">91</emphasis>
) of
<taxonomicName id="4C1E0DEFFFDBFFEFB861F8B009F10351" authorityName="Frisch" authorityYear="2022" box="[1082,1260,1863,1885]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="50" phylum="Arthropoda" rank="species" species="heronifer">
<emphasis id="B96AAA7EFFDBFFEFB861F8B009F10351" box="[1082,1260,1863,1885]" italics="true" pageId="27" pageNumber="50">Scopaeus heronifer</emphasis>
</taxonomicName>
, holotype, Banten, Lebak: Majasari (
<emphasis id="B96AAA7EFFDBFFEFBD04F8A60C83037E" bold="true" box="[351,414,1886,1908]" pageId="27" pageNumber="50">86– 88</emphasis>
);
<taxonomicName id="4C1E0DEFFFDBFFEFBDEAF8A60F55037E" authorityName="Frisch" authorityYear="2022" box="[433,584,1886,1907]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="50" phylum="Arthropoda" rank="species" species="diversilobatus">
<emphasis id="B96AAA7EFFDBFFEFBDEAF8A60F55037E" box="[433,584,1886,1907]" italics="true" pageId="27" pageNumber="50">S. diversilobatus</emphasis>
</taxonomicName>
, holotype, Jawa Barat, Sukabumi: Djampang Kulon (
<emphasis id="B96AAA7EFFDBFFEFB869F8A6096C037E" bold="true" box="[1074,1137,1886,1908]" pageId="27" pageNumber="50">89 – 91</emphasis>
).
<emphasis id="B96AAA7EFFDBFFEFB8DFF8A608100379" bold="true" box="[1156,1293,1886,1908]" pageId="27" pageNumber="50">Abbreviations</emphasis>
:
<emphasis id="B96AAA7EFFDBFFEFB942F8A6085B0379" bold="true" box="[1305,1350,1886,1908]" pageId="27" pageNumber="50">al(d)</emphasis>
– dextral apical lobe,
<emphasis id="B96AAA7EFFDBFFEFBD75F88C0C4A0387" bold="true" box="[302,343,1908,1930]" pageId="27" pageNumber="50">al(s)</emphasis>
– sinistral apical lobe,
<emphasis id="B96AAA7EFFDBFFEFBE74F88D0F5F0387" bold="true" box="[559,578,1909,1930]" pageId="27" pageNumber="50">dl</emphasis>
– dorsal lobe,
<emphasis id="B96AAA7EFFDBFFEFBE96F88C0FC80387" bold="true" box="[717,725,1908,1930]" pageId="27" pageNumber="50">f</emphasis>
– flagellum,
<emphasis id="B96AAA7EFFDBFFEFBF0BF88D0E400387" bold="true" box="[848,861,1909,1930]" pageId="27" pageNumber="50">ll</emphasis>
– lateral lobes,
<emphasis id="B96AAA7EFFDBFFEFBFAAF88C09160387" bold="true" box="[1009,1035,1908,1930]" pageId="27" pageNumber="50">mf</emphasis>
– median foramen,
<emphasis id="B96AAA7EFFDBFFEFB89CF88D09C40387" bold="true" box="[1223,1241,1909,1930]" pageId="27" pageNumber="50">tr</emphasis>
– transverse ridge of median foramen,
<emphasis id="B96AAA7EFFDBFFEFBD05F8740C6D03AC" bold="true" box="[350,368,1932,1953]" pageId="27" pageNumber="50">vl</emphasis>
– ventral lobe.
<emphasis id="B96AAA7EFFDBFFEFBE5BF8730F5403AD" bold="true" box="[512,585,1931,1952]" pageId="27" pageNumber="50">Arrows</emphasis>
: distoventral incision of dextral apical lobe (
<emphasis id="B96AAA7EFFDBFFEFBFBDF8740EF203AC" bold="true" box="[998,1007,1932,1953]" pageId="27" pageNumber="50">I</emphasis>
); sinistral, apical tooth of flagellum (
<emphasis id="B96AAA7EFFDBFFEFB91EF874084A03AC" bold="true" box="[1349,1367,1932,1953]" pageId="27" pageNumber="50">II</emphasis>
); distal emargination of dorsal lobe (
<emphasis id="B96AAA7EFFDBFFEFBD9FF85A0CC303BA" bold="true" box="[452,478,1954,1975]" pageId="27" pageNumber="50">III</emphasis>
).
</paragraph>
</caption>
<paragraph id="8BA1766CFFDCFFE8BF72FEAC0E9805C2" blockId="28.[786,1399,241,660]" pageId="28" pageNumber="51">
Female: Chamber of sperm pump elongate, with apophysis adjacent to bursal duct (
<figureCitation id="13256AE9FFDCFFE8B8D4FE8D09F50580" box="[1167,1256,373,397]" captionStart="Figures 163–172" captionStartId="42.[151,224,1839,1861]" captionTargetBox="[225,1322,230,1817]" captionTargetId="figure-168@42.[157,1398,210,1820]" captionTargetPageId="42" captionText="Figures 163–172. Sperm pump (163–172) and bursa (165, 167, 170, 172) of ♀ of Scopaeus grandis, paratype, Jawa Barat, Sukabumi: Cidahu (163); S.heronifer, paratype, Jawa Barat, Sukabumi:Sirnarasa (164); S. diversilobatus, paratype, Jawa Barat, Sukabumi, Djampang Kulon (165, with lateral gonocoxal plates IX folded laterally; 166); S. cuspilobatus, paratype, Jawa Barat, Sukabumi: Sirnarasa (167, with lateral gonocoxal plates IX, 168); S. velifer, paratype, Banten, Lebak: Citorek Kidul (169); S. elegantulus, Jawa Barat, Sukabumi: Djampang Kulon (170, with lateral gonocoxal plates IX, 171), S. uncinatus, paratype, Banten, Lebak: Citorek Kidul (172). Long scale bar: 163, 164, 166, 168, 169, 171, 172; short scale bar: 165, 167, 170." figureDoi="http://doi.org/10.5281/zenodo.10724315" httpUri="https://zenodo.org/record/10724315/files/figure.png" pageId="28" pageNumber="51">Fig. 171</figureCitation>
); bursal duct about twice as long as sperm pump; bursa membranous (
<figureCitation id="13256AE9FFDCFFE8BF41FE4F0E6405C2" box="[794,889,439,463]" captionStart="Figures 163–172" captionStartId="42.[151,224,1839,1861]" captionTargetBox="[225,1322,230,1817]" captionTargetId="figure-168@42.[157,1398,210,1820]" captionTargetPageId="42" captionText="Figures 163–172. Sperm pump (163–172) and bursa (165, 167, 170, 172) of ♀ of Scopaeus grandis, paratype, Jawa Barat, Sukabumi: Cidahu (163); S.heronifer, paratype, Jawa Barat, Sukabumi:Sirnarasa (164); S. diversilobatus, paratype, Jawa Barat, Sukabumi, Djampang Kulon (165, with lateral gonocoxal plates IX folded laterally; 166); S. cuspilobatus, paratype, Jawa Barat, Sukabumi: Sirnarasa (167, with lateral gonocoxal plates IX, 168); S. velifer, paratype, Banten, Lebak: Citorek Kidul (169); S. elegantulus, Jawa Barat, Sukabumi: Djampang Kulon (170, with lateral gonocoxal plates IX, 171), S. uncinatus, paratype, Banten, Lebak: Citorek Kidul (172). Long scale bar: 163, 164, 166, 168, 169, 171, 172; short scale bar: 165, 167, 170." figureDoi="http://doi.org/10.5281/zenodo.10724315" httpUri="https://zenodo.org/record/10724315/files/figure.png" pageId="28" pageNumber="51">Figs 170</figureCitation>
).
</paragraph>
<paragraph id="8BA1766CFFDCFFE8BF72FE200839061C" blockId="28.[786,1399,241,660]" pageId="28" pageNumber="51">
<emphasis id="B96AAA7EFFDCFFE8BF72FE200EB405FD" bold="true" box="[809,937,472,496]" pageId="28" pageNumber="51">Phylogeny:</emphasis>
<taxonomicName id="4C1E0DEFFFDCFFE8BFEAFE20098205FD" authorityName="Cameron" authorityYear="1930" box="[945,1183,472,496]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="51" phylum="Arthropoda" rank="species" species="elegantulus">
<emphasis id="B96AAA7EFFDCFFE8BFEAFE20098205FD" box="[945,1183,472,496]" italics="true" pageId="28" pageNumber="51">Scopaeus elegantulus</emphasis>
</taxonomicName>
is a member of the
<taxonomicName id="4C1E0DEFFFDCFFE8BF49FE010E9F061D" authorityName="Oke" authorityYear="1933" box="[786,898,504,528]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="51" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B96AAA7EFFDCFFE8BF49FE010E9F061D" box="[786,898,504,528]" italics="true" pageId="28" pageNumber="51">S. gracilis</emphasis>
</taxonomicName>
species group (
<bibRefCitation id="EF8F0B9DFFDCFFE8B876FE000805061C" author="Frisch, J. &amp; D. Burckhardt &amp; V. Wolters" box="[1069,1304,504,529]" pageId="28" pageNumber="51" pagination="27 - 53" refId="ref31920" refString="Frisch, J., D. Burckhardt &amp; V. Wolters (2002): Rove beetles of the subtribe Scopaeina Mulsant &amp; Rey (Coleoptera: Staphylinidae) in the West Palaearctic: Phylogeny, biogeography and species catalogue. - Organisms, Diversity &amp; Evolution 2: 27 - 53." type="journal article" year="2002">Frisch et al. 2002: 39</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFDCFFE8BF72FDE10E550699" pageId="28" pageNumber="51" type="distribution">
<paragraph id="8BA1766CFFDCFFE8BF72FDE10E550699" blockId="28.[786,1399,241,660]" pageId="28" pageNumber="51">
<emphasis id="B96AAA7EFFDCFFE8BF72FDE10ED9063C" bold="true" box="[809,964,537,561]" pageId="28" pageNumber="51">Distribution:</emphasis>
Described from Pahang in the Malay Peninsula,
<taxonomicName id="4C1E0DEFFFDCFFE8BFCBFDC30999065F" authorityName="Cameron" authorityYear="1930" box="[912,1156,570,594]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="51" phylum="Arthropoda" rank="species" species="elegantulus">
<emphasis id="B96AAA7EFFDCFFE8BFCBFDC30999065F" box="[912,1156,570,594]" italics="true" pageId="28" pageNumber="51">Scopaeus elegantulus</emphasis>
</taxonomicName>
is widely distributed across
<collectingCountry id="F30936FCFFDCFFE8BF38FDA30ED2067E" box="[867,975,603,627]" name="Indonesia" pageId="28" pageNumber="51">Indonesia</collectingCountry>
and recorded for
<collectingRegion id="49DAB88EFFDCFFE8B8FFFDA409CA067E" box="[1188,1239,604,627]" country="Indonesia" name="Jawa Timur" pageId="28" pageNumber="51">Java</collectingRegion>
,
<collectingRegion id="49DAB88EFFDCFFE8B8BDFDA3085E067E" box="[1254,1347,603,627]" country="Indonesia" name="Sumatera Utara" pageId="28" pageNumber="51">Sumatra</collectingRegion>
and
<collectingRegion id="49DAB88EFFDCFFE8BF49FD840E590699" box="[786,836,636,660]" country="Indonesia" name="Bali" pageId="28" pageNumber="51">Bali</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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