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<document id="A65E32784C425534D5565A18C2B5C6BD" ID-DOI="10.11646/zootaxa.5374.4.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10158722" ID-ZooBank="EEFF13BD-664A-4FD8-9FED-4E4B21A475E4" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700469135046" checkinUser="plazi" docAuthor="Ahrens, Dirk, Lukic, Daniel, Pham, Phu, Li, Wei &amp; Liu, Wangang" docDate="2023" docId="03B73169BE792741D9E40BC2FA82FEB4" docLanguage="en" docName="zootaxa.5374.4.1.pdf" docOrigin="Zootaxa 5374 (4)" docSource="https://mapress.com/zt/article/download/zootaxa.5374.4.1/52301" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tetraserica yucheni Liu, Li &amp; Ahrens 2023, new species" docType="treatment" docVersion="2" lastPageNumber="463" masterDocId="FF8E4911BE73274DD9730A19FFFAFFD1" masterDocTitle="Tetraserica Ahrens, 2004 of continental Southeast Asia: new records, new species and an updated key to species (Coleoptera: Scarabaeidae: Sericinae: Sericini)" masterLastPageNumber="486" masterPageNumber="451" pageNumber="461" updateTime="1700469620343" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="522483DA8DE6828D246B267B1518BFDF">Tetraserica Ahrens, 2004 of continental Southeast Asia: new records, new species and an updated key to species (Coleoptera: Scarabaeidae: Sericinae: Sericini)</mods:title>
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<mods:namePart id="23D8255597E889CAE86A4F0E2C5DD76C">Ahrens, Dirk</mods:namePart>
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<mods:affiliation id="48C0191596C286793577C47FA7CE1162">Museum A. Koenig, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Adenauerallee 127, 53113 Bonn, Germany &amp; ahrens. dirk _ col @ gmx. de; https: // orcid. org / 0000 - 0003 - 3524 - 7153</mods:affiliation>
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<mods:namePart id="18B2AF6509D414D750DEC6D5F4C8144E">Lukic, Daniel</mods:namePart>
<mods:affiliation id="4D66417014036480A6766A665459DA47">Museum A. Koenig, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Adenauerallee 127, 53113 Bonn, Germany &amp; University of Salzburg, Salzburg, Austria</mods:affiliation>
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<mods:affiliation id="D92F962D5695DD2802D6512EAD2D935D">Museum A. Koenig, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Adenauerallee 127, 53113 Bonn, Germany &amp; Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam</mods:affiliation>
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<mods:nameIdentifier id="07A1AB0A005B021A1FA9AE95609B25CA" type="ORCID">0009-0001-0195-7889</mods:nameIdentifier>
<mods:affiliation id="9B483E9419D1D94786EE77EA05692D29">Institute of Earth and Environment, Chinese Academy of Sciences, Yanxiang Road 97 #, Yanta District, Xi’an 710061, P. R. China. &amp; liweiv @ stumail. nwu. edu. cn; https: // orcid. org / 0009 - 0001 - 0195 - 7889</mods:affiliation>
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<mods:affiliation id="5D059DC8FC27153EB929FC62C6483E06">Museum A. Koenig, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Adenauerallee 127, 53113 Bonn, Germany &amp; Institute of Earth and Environment, Chinese Academy of Sciences, Yanxiang Road 97 #, Yanta District, Xi’an 710061, P. R. China. &amp; liuwangang @ ieecas. cn; https: // orcid. org / 0000 - 0003 - 4788 - 7967</mods:affiliation>
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<mods:date id="ECEE8E1B2117BB340D8D7E1813D3284A">2023</mods:date>
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<taxonomicName id="4C1EFBFCBE792747D9E40BC2FD9EFE25" authority="Liu, Li &amp; Ahrens" authorityName="Liu, Li &amp; Ahrens" authorityYear="2023" box="[151,612,474,501]" class="Insecta" family="Scarabaeidae" genus="Tetraserica" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="yucheni" status="sp. nov.">
<emphasis id="B96A5C6DBE792747D9E40BC2FE7FFE24" bold="true" box="[151,389,474,501]" italics="true" pageId="10" pageNumber="461">Tetraserica yucheni</emphasis>
Liu, Li
<emphasis id="B96A5C6DBE792747D8990BC2FDFBFE24" bold="true" box="[490,513,475,501]" italics="true" pageId="10" pageNumber="461">&amp;</emphasis>
Ahrens
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,
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<figureCitation id="13259CFABE792747D9E40BE6FEF1FDC8" box="[151,267,511,537]" captionStart="FIGURE 4" captionStartId="11.[152,255,1781,1805]" captionTargetBox="[162,1430,283,1748]" captionTargetId="figure-54@11.[151,1436,273,1758]" captionTargetPageId="11" captionText="FIGURE 4. A–F: Tetraserica yucheni Liu, Li &amp; Ahrens, new species (holotype); G–L: T. fabriziae Liu, Li &amp; Ahrens, new species (holotype). A, G: aedeagus, left side lateral view; B, C, H, I: parameres, dorsal view; D, J: aedeagus, right side lateral view; E, K: habitus; F, L: hind leg. Scale: 0.5 mm. Legs and habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.10158730" httpUri="https://zenodo.org/record/10158730/files/figure.png" pageId="10" pageNumber="461">Fig. 4A–F</figureCitation>
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<emphasis id="B96A5C6DBE792747D9E4085EFE30FDB1" bold="true" box="[151,458,582,609]" pageId="10" pageNumber="461">Type material examined.</emphasis>
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<typeStatus id="54A53EDDBE792747D8A4085EFDBEFDB0" box="[471,580,583,609]" pageId="10" pageNumber="461" type="holotype">Holotype</typeStatus>
: ♁ “
<collectingMunicipality id="6BC51A05BE792747DB0C085EFCCFFDB0" box="[639,821,583,609]" pageId="10" pageNumber="461">Mengdui Town</collectingMunicipality>
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<collectingRegion id="49DA4E9DBE792747DDA3085EFA62FDB0" box="[1232,1432,583,609]" country="China" name="Yunnan" pageId="10" pageNumber="461">Yunnan Province</collectingRegion>
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<collectingDate id="EFE45F57BE792747D9E40872FEF7FD54" box="[151,269,619,645]" pageId="10" pageNumber="461" value="2022-05-27">27.V.2022</collectingDate>
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<location id="8EC1D6A4BE792747D86E0872FE43FD54" LSID="urn:lsid:plazi:treatment:03B73169BE792741D9E40BC2FA82FEB4:8EC1D6A4BE792747D86E0872FE43FD54" box="[285,441,619,645]" county="Zhenkang County" municipality="Mengdui Town" name="Zhao Yuchen" pageId="10" pageNumber="461" stateProvince="Yunnan">Zhao Yuchen</location>
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<location id="8EC1D6A4BE792747D8BA0872FDC9FD54" LSID="urn:lsid:plazi:treatment:03B73169BE792741D9E40BC2FA82FEB4:8EC1D6A4BE792747D8BA0872FDC9FD54" box="[457,563,619,645]" county="Zhenkang County" municipality="Mengdui Town" name="Lu Jinbo" pageId="10" pageNumber="461" stateProvince="Yunnan">Lu Jinbo</location>
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<collectingDate id="EFE45F57BE792747DB740896FD87FD78" box="[519,637,655,681]" pageId="10" pageNumber="461" value="2022-05-27">27.V.2022</collectingDate>
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<location id="8EC1D6A4BE792747DBFA0896FCDAFD78" LSID="urn:lsid:plazi:treatment:03B73169BE792741D9E40BC2FA82FEB4:8EC1D6A4BE792747DBFA0896FCDAFD78" box="[649,800,655,681]" country="China" county="Zhenkang County" municipality="Mengdui Town" name="Zhao Yuchen" pageId="10" pageNumber="461" stateProvince="Yunnan">Zhao Yuchen</location>
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<location id="8EC1D6A4BE792747DA5F0896FC68FD78" LSID="urn:lsid:plazi:treatment:03B73169BE792741D9E40BC2FA82FEB4:8EC1D6A4BE792747DA5F0896FC68FD78" box="[812,914,655,681]" country="China" county="Zhenkang County" municipality="Mengdui Town" name="Lu Jinbo" pageId="10" pageNumber="461" stateProvince="Yunnan">Lu Jinbo</location>
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<collectingMethod id="525FF868BE792747DAED0896FBE3FD78" box="[926,1049,655,681]" pageId="10" pageNumber="461">Light Trap</collectingMethod>
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, 3 ♁♁,
<materialsCitation id="3B768A22BE792747DDAA0896FAE1FD1C" collectingDate="2022-05-24" collectionCode="IZAS" country="China" county="Yao'an County" location="Chuxiong" municipality="Gao Shan Trough Village" pageId="10" pageNumber="461" specimenCount="2" specimenCount-female="2" stateProvince="Yunnan" typeStatus="paratype">
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<collectingMunicipality id="6BC51A05BE792747DC5F0896FEB9FD1C" pageId="10" pageNumber="461">Gao Shan Trough Village</collectingMunicipality>
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<collectingDate id="EFE45F57BE792747DA3A08AAFC44FD1C" box="[841,958,691,717]" pageId="10" pageNumber="461" value="2022-05-24">24.V.2022</collectingDate>
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<location id="8EC1D6A4BE792747DABA08AAFBBEFD1C" LSID="urn:lsid:plazi:treatment:03B73169BE792741D9E40BC2FA82FEB4:8EC1D6A4BE792747DABA08AAFBBEFD1C" box="[969,1092,691,717]" country="China" county="Yao'an County" municipality="Gao Shan Trough Village" name="Xu Rentao" pageId="10" pageNumber="461" stateProvince="Yunnan">Xu Rentao</location>
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<collectingMethod id="525FF868BE792747DD3C08AAFB4DFD1C" box="[1103,1207,691,717]" pageId="10" pageNumber="461">light trap</collectingMethod>
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, 1 ♁,
<materialsCitation id="3B768A22BE792747DC2E08AAFAA8FD20" collectingDate="2022-05-26" collectionCode="IZAS" country="China" county="Zhenkang County" location="Lincang City" municipality="Fengwei Town" pageId="10" pageNumber="461" specimenCount="4" specimenCount-female="4" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D184BF6BE792747DC2E08AAFA66FD1C" box="[1373,1436,691,717]" count="4" pageId="10" pageNumber="461" type="female">4 ♀♀</specimenCount>
“
<collectingMunicipality id="6BC51A05BE792747D9D208CEFEB6FD20" box="[161,332,727,753]" pageId="10" pageNumber="461">Fengwei Town</collectingMunicipality>
,
<collectingCounty id="62C0F8F3BE792747D82408CEFDD8FD20" box="[343,546,727,753]" pageId="10" pageNumber="461">Zhenkang County</collectingCounty>
,
<location id="8EC1D6A4BE792747DB5F08CEFD45FD20" LSID="urn:lsid:plazi:treatment:03B73169BE792741D9E40BC2FA82FEB4:8EC1D6A4BE792747DB5F08CEFD45FD20" box="[556,703,727,753]" country="China" county="Zhenkang County" municipality="Fengwei Town" name="Lincang City" pageId="10" pageNumber="461" stateProvince="Yunnan">Lincang City</location>
,
<collectingRegion id="49DA4E9DBE792747DBBA08CEFC71FD20" box="[713,907,727,753]" country="China" name="Yunnan" pageId="10" pageNumber="461">Yunnan Province</collectingRegion>
,
<date id="FFA0A6BFBE792747DAE508CEFBF1FD20" box="[918,1035,727,753]" pageId="10" pageNumber="461" value="2022-05-26">
<collectingDate id="EFE45F57BE792747DAE508CEFBF1FD20" box="[918,1035,727,753]" pageId="10" pageNumber="461" value="2022-05-26">26.V.2022</collectingDate>
</date>
,
<location id="8EC1D6A4BE792747DD6408CEFB81FD20" LSID="urn:lsid:plazi:treatment:03B73169BE792741D9E40BC2FA82FEB4:8EC1D6A4BE792747DD6408CEFB81FD20" box="[1047,1147,727,753]" country="China" county="Zhenkang County" municipality="Fengwei Town" name="Lu Jinbo" pageId="10" pageNumber="461" stateProvince="Yunnan">Lu Jinbo</location>
,
<collectingMethod id="525FF868BE792747DDF408CEFB15FD20" box="[1159,1263,727,753]" pageId="10" pageNumber="461">light trap</collectingMethod>
” (
<collectionCode id="ED0F18BABE792747DC7408CEFAB0FD20" box="[1287,1354,727,753]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="10" pageNumber="461">IZAS</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B768A22BE792747DC2D08CEFBDFFCE8" collectingDate="2019-06-28" collectingDateMax="2019-06-30" collectingDateMin="2019-06-28" collectionCode="NMPC, ZFMK" country="China" county="Kral" location="Sommer" longitude="98.85167" municipality="Ruzicka" pageId="10" pageNumber="461" specimenCount="1" stateProvince="Yunnan" typeStatus="paratype">
2 ♁♁ “
<collectingCountry id="F309C0EFBE792747D9D208E2FEFAFCC4" box="[161,256,763,789]" name="China" pageId="10" pageNumber="461">CHINA</collectingCountry>
:
<collectingRegion id="49DA4E9DBE792747D87908E2FE5BFCC4" box="[266,417,763,789]" country="China" name="Yunnan" pageId="10" pageNumber="461">Yunnan Prov.</collectingRegion>
Lushui Co., Liuku city (park) Secondary broadleaved forest 25°51’.1’N,
<geoCoordinate id="EE2AE6B8BE792747DDA608E3FAB0FCC4" box="[1237,1354,762,789]" degrees="98" direction="east" minutes="51.1" orientation="longitude" pageId="10" pageNumber="461" precision="92" value="98.85167">98°51.1’E</geoCoordinate>
,
<locationDeviation id="9A13995DBE792747DC2508E3FF27FCE8" location="Hajek" pageId="10" pageNumber="461">
<quantity id="4CE62D9ABE792747DC2508E3FA66FCC4" box="[1366,1436,762,789]" metricMagnitude="2" metricUnit="m" metricValue="8.85" pageId="10" pageNumber="461" unit="m" value="885.0">
<locationDeviation id="9A13995DBE792747DC2508E3FA66FCC4" box="[1366,1436,762,789]" metricMagnitude="2" metricUnit="m" metricValue="8.85" pageId="10" pageNumber="461" unit="m" value="885.0">885 m</locationDeviation>
</quantity>
<locationDeviation id="9A13995DBE792747D9E40906FF27FCE8" box="[151,221,799,825]" pageId="10" pageNumber="461">Hájek</locationDeviation>
</locationDeviation>
,
<locationDeviation id="9A13995DBE792747D99B0907FEB1FCE8" box="[232,331,798,825]" pageId="10" pageNumber="461">Hrůzová</locationDeviation>
,
<collectingCounty id="62C0F8F3BE792747D8250906FE71FCE8" box="[342,395,799,825]" pageId="10" pageNumber="461">Král</collectingCounty>
,
<collectingMunicipality id="6BC51A05BE792747D8E50907FE08FCE8" box="[406,498,798,825]" pageId="10" pageNumber="461">Růzička</collectingMunicipality>
&amp;
<location id="8EC1D6A4BE792747DB650906FD8FFCE8" LSID="urn:lsid:plazi:treatment:03B73169BE792741D9E40BC2FA82FEB4:8EC1D6A4BE792747DB650906FD8FFCE8" box="[534,629,799,825]" country="China" county="Kral" longitude="98.85167" municipality="Ruzicka" name="Sommer" pageId="10" pageNumber="461" stateProvince="Yunnan">Sommer</location>
lgt.
<date id="FFA0A6BFBE792747DBD40906FCA8FCE8" box="[679,850,799,825]" pageId="10" pageNumber="461" value="2019-06-28" valueMax="2019-06-30" valueMin="2019-06-28">
<collectingDate id="EFE45F57BE792747DBD40906FCA8FCE8" box="[679,850,799,825]" pageId="10" pageNumber="461" value="2019-06-28" valueMax="2019-06-30" valueMin="2019-06-28">28.–30.vi.2019</collectingDate>
</date>
” (
<collectionCode id="ED0F18BABE792747DA180906FC45FCE8" box="[875,959,799,825]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="10" pageNumber="461">NMPC</collectionCode>
,
<collectionCode id="ED0F18BABE792747DAB90906FBE7FCE8" box="[970,1053,799,825]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="10" pageNumber="461" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C304D3F4BE792746D9B4095BFD73FF28" lastPageId="11" lastPageNumber="462" pageId="10" pageNumber="461" type="description">
<paragraph id="8BA1807FBE792747D9B4095BFC9AFC50" blockId="10.[151,1437,582,2013]" pageId="10" pageNumber="461">
<emphasis id="B96A5C6DBE792747D9B4095BFDE6FC8C" bold="true" box="[199,540,834,861]" pageId="10" pageNumber="461">
Description of the
<typeStatus id="54A53EDDBE792747D8DD095AFDEDFC8C" box="[430,535,835,861]" pageId="10" pageNumber="461" type="holotype">holotype</typeStatus>
.
</emphasis>
Length:
<quantity id="4CE62D9ABE792747DBFA095AFD1EFC8C" box="[649,740,835,861]" metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="10" pageNumber="461" unit="mm" value="9.9">9.9 mm</quantity>
, length of elytra: 7.0 mm, width:
<quantity id="4CE62D9ABE792747DD05095BFB29FC8C" box="[1142,1235,834,861]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="10" pageNumber="461" unit="mm" value="5.4">5.4 mm</quantity>
. Body oval, dark reddish brown, labroclypeus and dorsal surface shiny, glabrous.
</paragraph>
<paragraph id="8BA1807FBE792747D9B40992FC47FB14" blockId="10.[151,1437,582,2013]" pageId="10" pageNumber="461">
Labroclypeus subtrapezoidal, wider than long, widest at base, lateral margins weakly convex, anterior angles strongly convex, anterior margin shallowly emarginate medially, lateral margin and ocular canthus producing a blunt angle, margins moderately reflexed; surface weakly convex medially, shiny, moderately and densely punctate, without setae; frontoclypeal suture feebly impressed and moderately angled medially; smooth area in front of eye approximately four times as wide as long; ocular canthus moderate and triangular(1/3 of ocular diameter), weakly punctate, without setae. Frons weakly shiny and smooth, bearing only a few erect setae beside eyes. Eyes moderately small, ratio of diameter/ interocular width: 0.57. Antenna yellowish brown, with ten antennomeres, club in male with four antennomeres, slightly longer than remaining antennomeres combined, first joint of club about as long as club.
<collectingRegion id="49DA4E9DBE792747D9A60EB2FECEFB14" box="[213,308,1195,1221]" country="Myanmar" name="Chin" pageId="10" pageNumber="461">Mentum</collectingRegion>
anteriorly weakly elevated and weakly convex in middle.
</paragraph>
<paragraph id="8BA1807FBE792747D9B40ED6FC8BFAA8" blockId="10.[151,1437,582,2013]" pageId="10" pageNumber="461">Pronotum strongly transverse, widest just before base, lateral margins in basal quarter strongly convex, in anterior portion moderately convex and evenly convergent anteriorly, anterior angles distinctly produced, posterior angles convex. Anterior margin medially almost straight, with a fine, complete marginal line. Surface densely and finely punctate, without minute setae in punctures; lateral and anterior lateral margins sparsely setose. Hypomeron not carinate. Scutellum triangular, with fine and dense punctures.</paragraph>
<paragraph id="8BA1807FBE792747D9B40F9AFE17F9D8" blockId="10.[151,1437,582,2013]" pageId="10" pageNumber="461">Elytra oblong, widest at middle, striae moderately impressed, finely and densely punctate, intervals moderately convex, with fine, dense punctures, without erect, short setae, punctures without minute setae; epipleural edge robust, ending at convex external apical angle of elytra, epipleura sparsely setose, apical border membraneous, with a rim of short microtrichomes.</paragraph>
<paragraph id="8BA1807FBE792747D9B40C0AFAE6F96C" blockId="10.[151,1437,582,2013]" pageId="10" pageNumber="461">Ventral surface dull, yellowish brown, with large, dense punctures, sparsely setose, metasternum sparsely covered with fine, short, or minute setae; metacoxa glabrous, with a few single setae laterally; abdominal sternites finely and densely punctate, with a transverse row of coarse punctures each bearing a robust seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/ metacoxa: 1/1.8. Pygidium weakly convex and dull, coarsely densely punctate, without smooth midline, punctures with fine setae along apical margin.</paragraph>
<paragraph id="8BA1807FBE792746D9B40CDEFB82FF04" blockId="10.[151,1437,582,2013]" lastBlockId="11.[151,1436,151,249]" lastPageId="11" lastPageNumber="462" pageId="10" pageNumber="461">
Legs wide; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur wide, moderately shiny, its anterior margin acute, lacking an adjacent serrated line; ventral posterior margin straight, smooth and moderately widened in apical half (
<figureCitation id="13259CFABE792747DBD90D16FCFAF8F8" box="[682,768,1807,1833]" captionStart="FIGURE 4" captionStartId="11.[152,255,1781,1805]" captionTargetBox="[162,1430,283,1748]" captionTargetId="figure-54@11.[151,1436,273,1758]" captionTargetPageId="11" captionText="FIGURE 4. A–F: Tetraserica yucheni Liu, Li &amp; Ahrens, new species (holotype); G–L: T. fabriziae Liu, Li &amp; Ahrens, new species (holotype). A, G: aedeagus, left side lateral view; B, C, H, I: parameres, dorsal view; D, J: aedeagus, right side lateral view; E, K: habitus; F, L: hind leg. Scale: 0.5 mm. Legs and habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.10158730" httpUri="https://zenodo.org/record/10158730/files/figure.png" pageId="10" pageNumber="461">Fig. 4F</figureCitation>
); dorsal posterior margin smooth, with moderate short setae basally. Metatibia short, widest at middle; ratio width/ length: 1/ 2.9; dorsal margin weakly longitudinally convex, with two groups of spines, basal one at 1/3 first third, apical one at two thirds of metatibial length, basally with a few short single setae; lateral face weakly finely punctate; ventral margin finely serrate, with four robust equidistant setae; medial face smooth; apex interiorly near tarsal articulation moderately concave. Tarsomeres in distal half with fine, very dense setae ventrally, dorsally smooth; metatarsomeres with a strong serrated ridge, glabrous; first metatarsomere as long as following two tarsomeres combined, slightly longer than dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex.
</paragraph>
<paragraph id="8BA1807FBE782746D9B40AC6FD73FF28" blockId="11.[151,1436,151,249]" box="[199,649,223,249]" pageId="11" pageNumber="462">
Aedeagus.
<figureCitation id="13259CFABE782746D8370AC6FE45FF28" box="[324,447,223,249]" captionStart="FIGURE 4" captionStartId="11.[152,255,1781,1805]" captionTargetBox="[162,1430,283,1748]" captionTargetId="figure-54@11.[151,1436,273,1758]" captionTargetPageId="11" captionText="FIGURE 4. A–F: Tetraserica yucheni Liu, Li &amp; Ahrens, new species (holotype); G–L: T. fabriziae Liu, Li &amp; Ahrens, new species (holotype). A, G: aedeagus, left side lateral view; B, C, H, I: parameres, dorsal view; D, J: aedeagus, right side lateral view; E, K: habitus; F, L: hind leg. Scale: 0.5 mm. Legs and habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.10158730" httpUri="https://zenodo.org/record/10158730/files/figure.png" pageId="11" pageNumber="462">Fig. 4A–D</figureCitation>
. Habitus:
<figureCitation id="13259CFABE782746DB5C0AC6FD7FFF28" box="[559,645,223,249]" captionStart="FIGURE 4" captionStartId="11.[152,255,1781,1805]" captionTargetBox="[162,1430,283,1748]" captionTargetId="figure-54@11.[151,1436,273,1758]" captionTargetPageId="11" captionText="FIGURE 4. A–F: Tetraserica yucheni Liu, Li &amp; Ahrens, new species (holotype); G–L: T. fabriziae Liu, Li &amp; Ahrens, new species (holotype). A, G: aedeagus, left side lateral view; B, C, H, I: parameres, dorsal view; D, J: aedeagus, right side lateral view; E, K: habitus; F, L: hind leg. Scale: 0.5 mm. Legs and habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.10158730" httpUri="https://zenodo.org/record/10158730/files/figure.png" pageId="11" pageNumber="462">Fig. 4E</figureCitation>
.
</paragraph>
</subSubSection>
<caption id="DF61D0F7BE782746D9EB0CECFC2AF884" ID-DOI="http://doi.org/10.5281/zenodo.10158730" ID-Zenodo-Dep="10158730" httpUri="https://zenodo.org/record/10158730/files/figure.png" pageId="11" pageNumber="462" startId="11.[152,255,1781,1805]" targetBox="[162,1430,283,1748]" targetPageId="11" targetType="figure">
<paragraph id="8BA1807FBE782746D9EB0CECFC2AF884" blockId="11.[151,1436,1781,1878]" pageId="11" pageNumber="462">
<emphasis id="B96A5C6DBE782746D9EB0CECFEA2F8DC" bold="true" box="[152,344,1781,1805]" pageId="11" pageNumber="462">FIGURE 4. A–F:</emphasis>
<taxonomicName id="4C1EFBFCBE782746D8130CEFFD13F8DC" authority="Liu, Li &amp; Ahrens" authorityName="Liu, Li &amp; Ahrens" authorityYear="2023" box="[352,745,1781,1805]" class="Insecta" family="Scarabaeidae" genus="Tetraserica" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="yucheni" status="sp. nov.">
<emphasis id="B96A5C6DBE782746D8130CEFFDD3F8DC" box="[352,553,1781,1805]" italics="true" pageId="11" pageNumber="462">Tetraserica yucheni</emphasis>
Liu, Li
<emphasis id="B96A5C6DBE782746DBF20CEFFD6FF8DC" box="[641,661,1782,1805]" italics="true" pageId="11" pageNumber="462">&amp;</emphasis>
Ahrens
</taxonomicName>
,
<taxonomicNameLabel id="A259E116BE782746DB870CEFFC95F8DC" box="[756,879,1781,1805]" pageId="11" pageNumber="462" rank="species">new species</taxonomicNameLabel>
(holotype);
<emphasis id="B96A5C6DBE782746DA830CECFBD0F8DC" bold="true" box="[1008,1066,1781,1805]" pageId="11" pageNumber="462">G–L:</emphasis>
<taxonomicName id="4C1EFBFCBE782746DD410CEFFA9CF8DC" authority="Liu, Li &amp; Ahrens" authorityName="Liu, Li &amp; Ahrens" authorityYear="2023" box="[1074,1382,1781,1805]" class="Insecta" family="Scarabaeidae" genus="Tetraserica" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="fabriziae" status="sp. nov.">
<emphasis id="B96A5C6DBE782746DD410CEFFB5DF8DC" box="[1074,1191,1781,1805]" italics="true" pageId="11" pageNumber="462">T. fabriziae</emphasis>
Liu, Li
<emphasis id="B96A5C6DBE782746DD8C0CEFFAE9F8DC" box="[1279,1299,1782,1805]" italics="true" pageId="11" pageNumber="462">&amp;</emphasis>
Ahrens
</taxonomicName>
,
<taxonomicNameLabel id="A259E116BE782746DC010CEFFF1AF8E0" pageId="11" pageNumber="462" rank="species">new species</taxonomicNameLabel>
(holotype).
<emphasis id="B96A5C6DBE782746D82F0D00FE6DF8E0" bold="true" box="[348,407,1817,1841]" pageId="11" pageNumber="462">A, G:</emphasis>
aedeagus, left side lateral view;
<emphasis id="B96A5C6DBE782746DB970D00FCAEF8E0" bold="true" box="[740,852,1817,1841]" pageId="11" pageNumber="462">B, C, H, I:</emphasis>
parameres, dorsal view;
<emphasis id="B96A5C6DBE782746DD200D00FB7DF8E0" bold="true" box="[1107,1159,1817,1841]" pageId="11" pageNumber="462">D, J:</emphasis>
aedeagus, right side lateral view;
<emphasis id="B96A5C6DBE782746D9A50D24FEF5F884" bold="true" box="[214,271,1853,1877]" pageId="11" pageNumber="462">E, K:</emphasis>
habitus;
<emphasis id="B96A5C6DBE782746D81F0D24FE65F884" bold="true" box="[364,415,1853,1877]" pageId="11" pageNumber="462">F, L:</emphasis>
hind leg. Scale: 0.5 mm. Legs and habitus not to scale.
</paragraph>
</caption>
<subSubSection id="C304D3F4BE782741D9B40DBEFE9CFF04" lastPageId="12" lastPageNumber="463" pageId="11" pageNumber="462" type="diagnosis">
<paragraph id="8BA1807FBE782746D9B40DBEFA66F810" blockId="11.[151,1436,1959,2022]" box="[199,1436,1959,1986]" pageId="11" pageNumber="462">
<emphasis id="B96A5C6DBE782746D9B40DBEFEB8F810" bold="true" box="[199,322,1959,1985]" pageId="11" pageNumber="462">Diagnosis.</emphasis>
<taxonomicName id="4C1EFBFCBE782746D83A0DB1FD0DF810" authority="Liu, Li &amp; Ahrens" authorityName="Liu, Li &amp; Ahrens" authorityYear="2023" box="[329,759,1959,1986]" class="Insecta" family="Scarabaeidae" genus="Tetraserica" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="yucheni" status="sp. nov.">
<emphasis id="B96A5C6DBE782746D83A0DB1FDDDF810" box="[329,551,1959,1985]" italics="true" pageId="11" pageNumber="462">Tetraserica yucheni</emphasis>
Liu, Li
<emphasis id="B96A5C6DBE782746DBF60DB1FD61F813" box="[645,667,1960,1986]" italics="true" pageId="11" pageNumber="462">&amp;</emphasis>
Ahrens
</taxonomicName>
,
<taxonomicNameLabel id="A259E116BE782746DA700DB1FC70F810" box="[771,906,1959,1986]" pageId="11" pageNumber="462" rank="species">new species</taxonomicNameLabel>
differs from
<taxonomicName id="4C1EFBFCBE782746DD520DB1FA66F810" box="[1057,1436,1959,1985]" pageId="11" pageNumber="462">
<emphasis id="B96A5C6DBE782746DD520DB1FB23F810" box="[1057,1241,1959,1985]" italics="true" pageId="11" pageNumber="462">T. bolavensensis</emphasis>
<bibRefCitation id="EF8FFD8EBE782746DD920DBEFA66F810" author="Fabrizi, S. &amp; Dalstein, V. &amp; Ahrens, D." box="[1249,1436,1959,1985]" pageId="11" pageNumber="462" pagination="1 - 155" refId="ref44465" refString="Fabrizi, S., Dalstein, V. &amp; Ahrens, D. (2019) A monograph on the genus Tetraserica from the Indochinese region (Coleoptera, Scarabaeidae, Sericini). ZooKeys, 837, 1 - 155. https: // doi. org / 10.3897 / zookeys. 837.32057" type="journal article" year="2019">Fabrizi, Dalstein</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BA1807FBE782741D9E40DD5FE9CFF04" blockId="11.[151,1436,1959,2022]" lastBlockId="12.[151,1436,151,357]" lastPageId="12" lastPageNumber="463" pageId="11" pageNumber="462">
&amp; Ahrens, 2019 by the short ventral lobe of right paramere which is only as long as the slightly longer dorsal one and about one third of the length of the right paramere (in
<taxonomicName id="4C1EFBFCBE7F2741DA530A8EFC2DFF60" authorityName="Fabrizi, Dalstein &amp; Ahrens" authorityYear="2019" box="[800,983,151,177]" class="Insecta" family="Scarabaeidae" genus="Tetraserica" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="bolavensensis">
<emphasis id="B96A5C6DBE7F2741DA530A8EFC2DFF60" box="[800,983,151,177]" italics="true" pageId="12" pageNumber="463">T. bolavensensis</emphasis>
</taxonomicName>
the ventral lobe is almost half as long as the left paramere).
</paragraph>
</subSubSection>
<subSubSection id="C304D3F4BE7F2741D9B40AC7FF3EFE90" pageId="12" pageNumber="463" type="description">
<paragraph id="8BA1807FBE7F2741D9B40AC7FF3EFE90" blockId="12.[151,1436,151,357]" pageId="12" pageNumber="463">
<emphasis id="B96A5C6DBE7F2741D9B40AC7FEC5FF29" bold="true" box="[199,319,222,248]" pageId="12" pageNumber="463">Variation.</emphasis>
Length:
<quantity id="4CE62D9ABE7F2741D8D20AC6FDCDFF28" box="[417,567,223,249]" metricMagnitude="-3" metricUnit="m" metricValue="9.399999999999999" metricValueMax="10.1" metricValueMin="8.7" pageId="12" pageNumber="463" unit="mm" value="9.399999999999999" valueMax="10.1" valueMin="8.7">8.7–10.1 mm</quantity>
, length of elytra:
<quantity id="4CE62D9ABE7F2741DB8B0AC6FC7AFF28" box="[760,896,223,249]" metricMagnitude="-3" metricUnit="m" metricValue="7.35" metricValueMax="7.9" metricValueMin="6.8" pageId="12" pageNumber="463" unit="mm" value="7.35" valueMax="7.9" valueMin="6.8">6.8–7.9 mm</quantity>
, width: 5.2–6.0 mm. Female: Antennal club with three antennomeres, as long as remaining antennomeres combined. Eyes slightly smaller than in male. Pygidium flat.
</paragraph>
</subSubSection>
<subSubSection id="C304D3F4BE7F2741D9B40B52FA82FEB4" box="[199,1400,331,357]" pageId="12" pageNumber="463" type="etymology">
<paragraph id="8BA1807FBE7F2741D9B40B52FA82FEB4" blockId="12.[151,1436,151,357]" box="[199,1400,331,357]" pageId="12" pageNumber="463">
<emphasis id="B96A5C6DBE7F2741D9B40B52FEB4FEB4" bold="true" box="[199,334,331,357]" pageId="12" pageNumber="463">Etymology.</emphasis>
The new species is named after of its collectors, Zhao Yuchen (noun in genitive singular case).
</paragraph>
</subSubSection>
</treatment>
</document>
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