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<document id="688EC41FC3B5D3E11897ED8BB69611DF" ID-CLB-Dataset="21603" ID-DOI="10.5852/ejt.2020.673" ID-GBIF-Dataset="de4f2e41-d1c5-47c8-98af-548b7f9842bb" ID-ISSN="2118-9773" ID-Zenodo-Dep="3908345" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1593097361916" checkinUser="valdenar" docAuthor="Sankaran, Pradeep M., Caleb, John T. D. &amp; Sebastian, Pothalil A." docDate="2020" docId="03B5D875805DFFEF9594FA49FC97F969" docLanguage="en" docName="ejt-673_sankaran_caleb_sebastian.pdf.imf" docOrigin="European Journal of Taxonomy 673" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cryptodrassus khajuriai Sankaran &amp; Caleb &amp; Sebastian 2020, comb. nov." docType="treatment" docVersion="5" lastPageNumber="5" masterDocId="FF8CA00D805CFFEB943AFFD7FFD5FF8D" masterDocTitle="On the taxonomic validity of Indian ground spiders: II. Genera Drassyllus Chamberlin, 1922 and Nodocion Chamberlin, 1922 (Araneae: Gnaphosidae)" masterLastPageNumber="14" masterPageNumber="1" pageNumber="2" updateTime="1698842909708" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="ABFA5033D826513F24D76B0561D3E991">On the taxonomic validity of Indian ground spiders: II. Genera Drassyllus Chamberlin, 1922 and Nodocion Chamberlin, 1922 (Araneae: Gnaphosidae)</mods:title>
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<mods:namePart id="93AAB5A63647E45866A5B3FB37C60AAC">Sankaran, Pradeep M.</mods:namePart>
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<mods:affiliation id="C2E109090364CE42608C3D9978D4A4C5">Division of Arachnology, Department of Zoology, Sacred Heart College, Thevara, Cochin, Kerala 682 013, India. &amp; urn: lsid: zoobank. org: author: 0 AEC 69 AA- 7 E 17 - 401 F-B 83 B- 280 C 2 F 04 AC 6 E &amp; Corresponding author: pradeepmspala @ rediffmail. com</mods:affiliation>
<mods:nameIdentifier id="82AB94556546739221167860E23E94DC" type="email">pradeepmspala@rediffmail.com</mods:nameIdentifier>
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<mods:namePart id="DE0A37519563FAF8AEE477197AD35548">Caleb, John T. D.</mods:namePart>
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<mods:affiliation id="D84AB1C5C7A94B17C120FADB0D7716D4">Zoological Survey of India, Prani Vigyan Bhawan, M-Block, New Alipore, Kolkata, West Bengal 700 053, India. &amp; Email: caleb 87 woodgate @ gmail. com &amp; urn: lsid: zoobank. org: author: 2 B 3052 EB- 5000 - 4 A 94 - 8 E 38 - F 88 CC 308 ECAD</mods:affiliation>
<mods:nameIdentifier id="BC37C10601ABE611072A34B02D548875" type="email">caleb87woodgate@gmail.com</mods:nameIdentifier>
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<mods:namePart id="5770F5A78E7A81C797799986E0410731">Sebastian, Pothalil A.</mods:namePart>
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<mods:affiliation id="5FA7745BD0F2A5657BA5A5B8B26A3C5C">Division of Arachnology, Department of Zoology, Sacred Heart College, Thevara, Cochin, Kerala 682 013, India. &amp; Email: drpothalil @ rediffmail. com &amp; urn: lsid: zoobank. org: author: C 22767 DA- 4 C 75 - 4685 - B 77 E- 2 AB 058 C 1091 D</mods:affiliation>
<mods:nameIdentifier id="A06A9DB6C4BFD6E3BAC30D1682690888" type="email">drpothalil@rediffmail.com</mods:nameIdentifier>
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<mods:title id="318AE134BEB91D89032B767AF7A74C0A">European Journal of Taxonomy</mods:title>
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<mods:date id="E6E5B6F6BEF24304E4FCFD8BB63494F2">2020</mods:date>
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<treatment id="03B5D875805DFFEF9594FA49FC97F969" ID-GBIF-Taxon="164743194" LSID="urn:lsid:plazi:treatment:03B5D875805DFFEF9594FA49FC97F969" httpUri="http://treatment.plazi.org/id/03B5D875805DFFEF9594FA49FC97F969" lastPageId="4" lastPageNumber="5" pageId="1" pageNumber="2">
<subSubSection id="C3063AE8805DFFEA9594FA49FB53FA37" box="[430,1158,1438,1466]" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="8BA36963805DFFEA9594FA49FB53FA37" blockId="1.[430,1158,1438,1503]" box="[430,1158,1438,1466]" pageId="1" pageNumber="2">
<heading id="D0EBDE0F805DFFEA9594FA49FB53FA37" box="[430,1158,1438,1466]" centered="true" fontSize="11" level="2" pageId="1" pageNumber="2" reason="2">
<taxonomicName id="4C1C12E0805DFFEA9594FA49FC29FA37" ID-CoL="98ZKX" authority="(Tikader &amp; Gajbe, 1976)" authorityName="Sankaran &amp; Caleb &amp; Sebastian" authorityYear="2020" baseAuthorityName="Tikader &amp; Gajbe" baseAuthorityYear="1976" box="[430,1020,1438,1466]" class="Arachnida" family="Gnaphosidae" genus="Cryptodrassus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="khajuriai" status="comb. nov.">
<emphasis id="B968B571805DFFEA9594FA49FD07FA34" bold="true" box="[430,722,1438,1465]" italics="true" pageId="1" pageNumber="2">Cryptodrassus khajuriai</emphasis>
(
<bibRefCitation id="EF8D1492805DFFEA96D8FA48FC21FA37" author="Tikader B. K. &amp; Gajbe U. A." box="[738,1012,1439,1466]" pageId="1" pageNumber="2" pagination="431 - 434" refId="ref5325" refString="Tikader B. K. &amp; Gajbe U. A. 1976. New spiders of Drassyllus from India (Gnaphosidae). Oriental Insects 10: 431 - 434." type="journal article" year="1976">Tikader &amp; Gajbe, 1976</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A25B080A805DFFEA903EFA48FB53FA37" box="[1028,1158,1439,1466]" pageId="1" pageNumber="2" rank="species">comb. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3063AE8805DFFEA96DDFA12FC98FA52" box="[743,845,1477,1503]" pageId="1" pageNumber="2" type="description">
<paragraph id="8BA36963805DFFEA96DDFA12FC98FA52" blockId="1.[430,1158,1438,1503]" box="[743,845,1477,1503]" pageId="1" pageNumber="2">
<figureCitation id="132775E6805DFFEA96DDFA12FC98FA52" box="[743,845,1477,1503]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="2.[189,232,1897,1923]" captionStartId-1="3.[189,232,1894,1920]" captionTargetBox-0="[198,1388,467,1853]" captionTargetBox-1="[198,1387,518,1852]" captionTargetId-0="figure@2.[189,1398,456,1863]" captionTargetId-1="figure@3.[189,1398,511,1859]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Fig. 1. Cryptodrassus khajuriai (Tikader &amp; Gajbe, 1976) comb. nov., ♀, holotype of Drassyllus khajuriai Tikader &amp; Gajbe, 1976 (NZC-ZSI-5043/18). A. Habitus, dorsal view. B. Eyes of the same, dorsal view. C. Epigyne, ventral view. D. Same, dorsal view. E. Label from type bottle. Scale bars: A = 1 mm; B = 0.5 mm; CD = 0.2 mm." captionText-1="Fig. 2. Cryptodrassus khajuriai (Tikader &amp; Gajbe, 1976) comb. nov., ♀, holotype of Drassyllus jabalpurensis Gajbe, 2005 (NZC-ZSI-5452/18). A. Habitus, dorsal view. B. Eyes of the same, dorsal view. C. Epigyne, ventral view. D. Same, dorsal view. E. Label from type bottle. Scale bars: A = 1 mm; B = 0.5 mm; CD = 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3908347" figureDoi-1="http://doi.org/10.5281/zenodo.3908349" httpUri-0="https://zenodo.org/record/3908347/files/figure.png" httpUri-1="https://zenodo.org/record/3908349/files/figure.png" pageId="1" pageNumber="2">Figs 12</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3063AE8805DFFEA9487F9DDFB3AF902" pageId="1" pageNumber="2" type="reference_group">
<paragraph id="8BA36963805DFFEA9487F9DDFCA7F9A8" blockId="1.[189,956,1546,1611]" box="[189,882,1546,1573]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="AB0C4E4D805DFFEA9487F9DDFCA7F9A8" box="[189,882,1546,1573]" pageId="1" pageNumber="2">
<treatmentCitation id="0ABD4F72805DFFEA9487F9DDFCD7F9A9" author="Tikader B. K. &amp; Gajbe U. A." box="[189,770,1546,1572]" page="432" pageId="1" pageNumber="2" year="1976">
<taxonomicName id="4C1C12E0805DFFEA9487F9DDFCD7F9A9" ID-CoL="6DL2P" authority="Tikader &amp; Gajbe, 1976: 432" authorityName="Tikader &amp; Gajbe" authorityPageNumber="432" authorityYear="1976" box="[189,770,1546,1572]" class="Arachnida" family="Gnaphosidae" genus="Drassyllus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="khajuriai">
<emphasis id="B968B571805DFFEA9487F9DDFE7BF9A9" box="[189,430,1546,1572]" italics="true" pageId="1" pageNumber="2">Drassyllus khajuriai</emphasis>
<bibRefCitation id="EF8D1492805DFFEA958EF9DDFCD7F9A9" author="Tikader B. K. &amp; Gajbe U. A." box="[436,770,1546,1572]" pageId="1" pageNumber="2" pagination="431 - 434" refId="ref5325" refString="Tikader B. K. &amp; Gajbe U. A. 1976. New spiders of Drassyllus from India (Gnaphosidae). Oriental Insects 10: 431 - 434." type="journal article" year="1976">Tikader &amp; Gajbe, 1976: 432</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 58.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA36963805DFFEA9487F9E7FC69F9C6" blockId="1.[189,956,1546,1611]" box="[189,956,1584,1611]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="AB0C4E4D805DFFEA9487F9E7FC69F9C6" box="[189,956,1584,1611]" pageId="1" pageNumber="2">
<treatmentCitation id="0ABD4F72805DFFEA9487F9E7FD63F9C7" author="Gajbe U. A." box="[189,694,1584,1611]" page="136" pageId="1" pageNumber="2" year="2005">
<taxonomicName id="4C1C12E0805DFFEA9487F9E7FD63F9C7" authority="Gajbe, 2005: 136" authorityName="Gajbe" authorityPageNumber="136" authorityYear="2005" box="[189,694,1584,1611]" class="Arachnida" family="Gnaphosidae" genus="Drassyllus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="jabalpurensis">
<emphasis id="B968B571805DFFEA9487F9E7FE34F9C7" box="[189,481,1584,1610]" italics="true" pageId="1" pageNumber="2">Drassyllus jabalpurensis</emphasis>
<bibRefCitation id="EF8D1492805DFFEA95D2F9E7FD63F9C7" author="Gajbe U. A." box="[488,694,1584,1611]" pageId="1" pageNumber="2" pagination="111 - 140" refId="ref4969" refString="Gajbe U. A. 2005. Studies on some spiders of the family Gnaphosidae (Araneae: Arachnida) from Madhya Pradesh, India. Records of the Zoological Survey of India 105: 111 - 140." type="journal article" year="2005">Gajbe, 2005: 136</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 2327.
<emphasis id="B968B571805DFFEA9771F9E7FC69F9C6" bold="true" box="[843,956,1584,1611]" pageId="1" pageNumber="2">Syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA36963805DFFEA9487F9A2FB3AF902" blockId="1.[189,1263,1653,1680]" box="[189,1263,1653,1680]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="AB0C4E4D805DFFEA9487F9A2FB3AF902" box="[189,1263,1653,1680]" pageId="1" pageNumber="2">
<taxonomicName id="4C1C12E0805DFFEA9487F9A2FE7BF902" ID-CoL="6DL2P" authorityName="Tikader &amp; Gajbe" authorityYear="1976" box="[189,430,1653,1679]" class="Arachnida" family="Gnaphosidae" genus="Drassyllus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="khajuriai">
<emphasis id="B968B571805DFFEA9487F9A2FE7BF902" box="[189,430,1653,1679]" italics="true" pageId="1" pageNumber="2">Drassyllus khajuriai</emphasis>
</taxonomicName>
<treatmentCitation id="0ABD4F72805DFFEA95F0F9A2FD70F91D" author="Tikader B. K." box="[458,677,1653,1680]" page="512" pageId="1" pageNumber="2" year="1982">
<bibRefCitation id="EF8D1492805DFFEA95F0F9A2FD70F91D" author="Tikader B. K." box="[458,677,1653,1680]" pageId="1" pageNumber="2" pagination="295 - 536" refId="ref5286" refString="Tikader B. K. 1982. Part 2. Family Gnaphosidae. In: The Fauna of India. Spiders: Araneae. Vol. II: 295 - 536. Zoological Survey of India, Calcutta." type="book chapter" year="1982">Tikader 1982: 512</bibRefCitation>
</treatmentCitation>
, figs 490494. —
<treatmentCitation id="0ABD4F72805DFFEA9746F9A2FB96F902" author="Gajbe U. A." box="[892,1091,1653,1679]" page="489" pageId="1" pageNumber="2" year="2007">
<bibRefCitation id="EF8D1492805DFFEA9746F9A2FB96F902" author="Gajbe U. A." box="[892,1091,1653,1679]" pageId="1" pageNumber="2" pagination="419 - 540" refId="ref5008" refString="Gajbe U. A. 2007. Araneae: Arachnida. In: the Director, Zoological Survey of India (ed.) Fauna of Madhya Pradesh (including Chhattisgarh), State Fauna Series 15: 419 - 540. Zoological Survey of India, Kolkata." type="book chapter" year="2007">Gajbe 2007: 489</bibRefCitation>
</treatmentCitation>
, figs 208212.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3063AE8805DFFEA9487F96EFD64F861" pageId="1" pageNumber="2" type="diagnosis">
<paragraph id="8BA36963805DFFEA9487F96EFEE3F95E" blockId="1.[189,310,1721,1747]" box="[189,310,1721,1747]" pageId="1" pageNumber="2">
<heading id="D0EBDE0F805DFFEA9487F96EFEE3F95E" bold="true" box="[189,310,1721,1747]" fontSize="11" level="3" pageId="1" pageNumber="2" reason="3">
<emphasis id="B968B571805DFFEA9487F96EFEE3F95E" bold="true" box="[189,310,1721,1747]" pageId="1" pageNumber="2">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA36963805DFFEA9487F93CFD64F861" blockId="1.[189,1399,1771,2028]" pageId="1" pageNumber="2">
<taxonomicName id="4C1C12E0805DFFEA9487F93CFE34F888" authorityName="Sankaran &amp; Caleb &amp; Sebastian" authorityYear="2020" baseAuthorityName="Tikader &amp; Gajbe" baseAuthorityYear="1976" box="[189,481,1771,1797]" class="Arachnida" family="Gnaphosidae" genus="Cryptodrassus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="khajuriai" status="comb. nov.">
<emphasis id="B968B571805DFFEA9487F93CFE34F888" box="[189,481,1771,1797]" italics="true" pageId="1" pageNumber="2">Cryptodrassus khajuriai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B080A805DFFEA95D4F93CFDA3F888" box="[494,630,1771,1797]" pageId="1" pageNumber="2" rank="species">comb. nov.</taxonomicNameLabel>
seems closely related to
<taxonomicName id="4C1C12E0805DFFEA9782F93CFB06F888" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1872" box="[952,1235,1771,1797]" class="Arachnida" family="Gnaphosidae" genus="Cryptodrassus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="helvolus">
<emphasis id="B968B571805DFFEA9782F93CFB06F888" box="[952,1235,1771,1797]" italics="true" pageId="1" pageNumber="2">Cryptodrassus helvolus</emphasis>
</taxonomicName>
(O. Pickard- Cambridge, 1872), as both have a large epigynal atrium. However, the former species can be distinguished from the latter by the following combination of characters: epigyne with anterior hood (epigyne of
<taxonomicName id="4C1C12E0805DFFEA9487F889FE95F8F5" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1872" box="[189,320,1886,1912]" class="Arachnida" family="Gnaphosidae" genus="Cryptodrassus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="helvolus">
<emphasis id="B968B571805DFFEA9487F889FE95F8F5" box="[189,320,1886,1912]" italics="true" pageId="1" pageNumber="2">C. helvolus</emphasis>
</taxonomicName>
lacks anterior hood), highly coiled copulatory ducts with thick proximal and thin distal parts (copulatory ducts of
<taxonomicName id="4C1C12E0805DFFEA9581F852FD97F813" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1872" box="[443,578,1924,1951]" class="Arachnida" family="Gnaphosidae" genus="Cryptodrassus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="helvolus">
<emphasis id="B968B571805DFFEA9581F852FD97F813" box="[443,578,1924,1951]" italics="true" pageId="1" pageNumber="2">C. helvolus</emphasis>
</taxonomicName>
less coiled and nearly uniform in diameter along the entire length) and globular receptacles (
<taxonomicName id="4C1C12E0805DFFEA95C9F87CFDACF849" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1872" box="[499,633,1962,1989]" class="Arachnida" family="Gnaphosidae" genus="Cryptodrassus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="helvolus">
<emphasis id="B968B571805DFFEA95C9F87CFDACF849" box="[499,633,1962,1989]" italics="true" pageId="1" pageNumber="2">C. helvolus</emphasis>
</taxonomicName>
with nearly oval receptacles) (compare
<figureCitation id="132775E6805DFFEA9065F87DFB70F848" box="[1119,1189,1962,1989]" captionStart="Fig" captionStartId="2.[189,232,1897,1923]" captionTargetBox="[198,1388,467,1853]" captionTargetId="figure@2.[189,1398,456,1863]" captionTargetPageId="2" captionText="Fig. 1. Cryptodrassus khajuriai (Tikader &amp; Gajbe, 1976) comb. nov., ♀, holotype of Drassyllus khajuriai Tikader &amp; Gajbe, 1976 (NZC-ZSI-5043/18). A. Habitus, dorsal view. B. Eyes of the same, dorsal view. C. Epigyne, ventral view. D. Same, dorsal view. E. Label from type bottle. Scale bars: A = 1 mm; B = 0.5 mm; CD = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3908347" httpUri="https://zenodo.org/record/3908347/files/figure.png" pageId="1" pageNumber="2">Figs 1</figureCitation>
CD, 2CD with
<bibRefCitation id="EF8D1492805DFFEA9487F806FDE2F866" author="Chatzaki M. &amp; Russell-Smith A." box="[189,567,2001,2027]" pageId="1" pageNumber="2" pagination="237 - 255" refId="ref4861" refString="Chatzaki M. &amp; Russell-Smith A. 2017. New species and new records of ground spiders (Araneae: Gnaphosidae) from Cyprus. Zootaxa 4329: 237 - 255. https: // doi. org / 10.11646 / zootaxa. 4329.3.3" type="journal article" year="2017">Chatzaki &amp; Russell-Smith 2017</bibRefCitation>
: figs 56).
</paragraph>
</subSubSection>
<subSubSection id="C3063AE8805EFFE89487FEDCFD48FEE1" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="8BA36963805EFFE99487FEDCFE7FFEA8" blockId="2.[189,426,267,293]" box="[189,426,267,293]" pageId="2" pageNumber="3">
<heading id="D0EBDE0F805EFFE99487FEDCFE7FFEA8" bold="true" box="[189,426,267,293]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="B968B571805EFFE99487FEDCFE7FFEA8" bold="true" box="[189,426,267,293]" pageId="2" pageNumber="3">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BA36963805EFFE994D0FEEDFDC2FED9" blockId="2.[189,1398,313,411]" box="[234,535,313,340]" pageId="2" pageNumber="3">
<emphasis id="B968B571805EFFE994D0FEEDFDC2FED9" bold="true" box="[234,535,313,340]" pageId="2" pageNumber="3">
<typeStatus id="54A7D7C1805EFFE994D0FEEDFE89FED9" box="[234,348,314,340]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="4C1C12E0805EFFE995BEFEEDFDC2FED9" authorityName="Sankaran &amp; Caleb &amp; Sebastian" authorityYear="2020" baseAuthorityName="Tikader &amp; Gajbe" baseAuthorityYear="1976" box="[388,535,313,340]" class="Arachnida" family="Gnaphosidae" genus="Cryptodrassus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="khajuriai">
<emphasis id="B968B571805EFFE995BEFEEDFDC2FED9" bold="true" box="[388,535,313,340]" italics="true" pageId="2" pageNumber="3">D. khajuriai</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BA36963805EFFE99487FE8AFE5CFE16" blockId="2.[189,1398,313,411]" pageId="2" pageNumber="3">
<materialsCitation id="3B74633E805EFFE99487FE8AFE5CFE16" ID-GBIF-Occurrence="2643239305" collectingDate="1974-02-28" collectorName="H. Khajuria" country="India" location="rest house in Madla" municipality="Jabalpur" pageId="2" pageNumber="3" specimenCode="5043/18" specimenCount="1" specimenCount-female="1" stateProvince="Madhya Pradesh" typeStatus="holotype">
<collectingCountry id="F30B29F3805EFFE99487FE8AFEDBFEFA" box="[189,270,349,375]" name="India" pageId="2" pageNumber="3">INDIA</collectingCountry>
<specimenCount id="9D1AA2EA805EFFE99512FE8AFEEAFEFA" box="[296,319,349,375]" pageId="2" pageNumber="3" type="female"></specimenCount>
;
<collectingRegion id="49D8A781805EFFE99577FE8AFDC0FEFA" box="[333,533,349,375]" country="India" name="Madhya Pradesh" pageId="2" pageNumber="3">Madhya Pradesh</collectingRegion>
,
<collectingMunicipality id="6BC7F319805EFFE9961AFE8AFD50FEFA" box="[544,645,349,375]" pageId="2" pageNumber="3">Jabalpur</collectingMunicipality>
,
<location id="8EC33FB8805EFFE996ABFE8AFCA9FEFA" LSID="urn:lsid:plazi:treatment:03B5D875805DFFEF9594FA49FC97F969:8EC33FB8805EFFE996ABFE8AFCA9FEFA" box="[657,892,349,375]" country="India" municipality="Jabalpur" name="rest house in Madla" pageId="2" pageNumber="3" stateProvince="Madhya Pradesh">rest house in Madla</location>
;
<date id="FFA24FA3805EFFE997B2FE8AFBF3FEFA" box="[904,1062,349,375]" pageId="2" pageNumber="3" value="1974-02-28">
<collectingDate id="EFE6B64B805EFFE997B2FE8AFBF3FEFA" box="[904,1062,349,375]" pageId="2" pageNumber="3" value="1974-02-28">28 Feb. 1974</collectingDate>
</date>
;
<collectorName id="26E90CB5805EFFE99008FE8AFB68FEFA" box="[1074,1213,349,375]" pageId="2" pageNumber="3">H. Khajuria</collectorName>
leg.; NZC-ZSI, Kolkata
<specimenCode id="DBBAC118805EFFE9951BFE57FE50FE16" box="[289,389,384,411]" pageId="2" pageNumber="3">5043/18</specimenCode>
.
</materialsCitation>
</paragraph>
<caption id="DF6339EB805EFFE99487F8BEFE0BF860" ID-DOI="http://doi.org/10.5281/zenodo.3908347" ID-Zenodo-Dep="3908347" httpUri="https://zenodo.org/record/3908347/files/figure.png" pageId="2" pageNumber="3" startId="2.[189,232,1897,1923]" targetBox="[198,1388,467,1853]" targetPageId="2">
<paragraph id="8BA36963805EFFE99487F8BEFE0BF860" blockId="2.[189,1399,1897,2030]" pageId="2" pageNumber="3">
<emphasis id="B968B571805EFFE99487F8BEFEDDF80E" bold="true" box="[189,264,1897,1923]" pageId="2" pageNumber="3">Fig. 1.</emphasis>
<taxonomicName id="4C1C12E0805EFFE99534F8BEFC9EF809" authority="(Tikader &amp; Gajbe, 1976)" authorityName="Sankaran &amp; Caleb &amp; Sebastian" authorityYear="2020" baseAuthorityName="Tikader &amp; Gajbe" baseAuthorityYear="1976" box="[270,843,1897,1924]" class="Arachnida" family="Gnaphosidae" genus="Cryptodrassus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="khajuriai" status="comb. nov.">
<emphasis id="B968B571805EFFE99534F8BEFDFFF80E" box="[270,554,1897,1923]" italics="true" pageId="2" pageNumber="3">Cryptodrassus khajuriai</emphasis>
(
<bibRefCitation id="EF8D1492805EFFE9960CF8BEFC96F809" author="Tikader B. K. &amp; Gajbe U. A." box="[566,835,1897,1924]" pageId="2" pageNumber="3" pagination="431 - 434" refId="ref5325" refString="Tikader B. K. &amp; Gajbe U. A. 1976. New spiders of Drassyllus from India (Gnaphosidae). Oriental Insects 10: 431 - 434." type="journal article" year="1976">Tikader &amp; Gajbe, 1976</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A25B080A805EFFE9976AF8BEFC1AF809" box="[848,975,1897,1924]" pageId="2" pageNumber="3" rank="species">comb. nov.</taxonomicNameLabel>
, ♀, holotype of
<taxonomicName id="4C1C12E0805EFFE990B2F8BEFE19F82B" authority="Tikader &amp; Gajbe, 1976" authorityName="Tikader &amp; Gajbe" authorityYear="1976" class="Arachnida" family="Gnaphosidae" genus="Drassyllus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="khajuriai">
<emphasis id="B968B571805EFFE990B2F8BEFAA2F80E" box="[1160,1399,1897,1923]" italics="true" pageId="2" pageNumber="3">Drassyllus khajuriai</emphasis>
<bibRefCitation id="EF8D1492805EFFE99487F85BFE19F82B" author="Tikader B. K. &amp; Gajbe U. A." box="[189,460,1932,1959]" pageId="2" pageNumber="3" pagination="431 - 434" refId="ref5325" refString="Tikader B. K. &amp; Gajbe U. A. 1976. New spiders of Drassyllus from India (Gnaphosidae). Oriental Insects 10: 431 - 434." type="journal article" year="1976">Tikader &amp; Gajbe, 1976</bibRefCitation>
</taxonomicName>
(NZC-ZSI-5043/18).
<emphasis id="B968B571805EFFE996F5F85BFD31F82B" bold="true" box="[719,740,1932,1958]" pageId="2" pageNumber="3">A</emphasis>
. Habitus, dorsal view.
<emphasis id="B968B571805EFFE997CBF85AFBD0F82A" bold="true" box="[1009,1029,1933,1959]" pageId="2" pageNumber="3">B</emphasis>
. Eyes of the same, dorsal view.
<emphasis id="B968B571805EFFE99487F867FF07F847" bold="true" box="[189,210,1968,1994]" pageId="2" pageNumber="3">C</emphasis>
. Epigyne, ventral view.
<emphasis id="B968B571805EFFE995C9F867FDDDF847" bold="true" box="[499,520,1968,1994]" pageId="2" pageNumber="3">D</emphasis>
. Same, dorsal view.
<emphasis id="B968B571805EFFE996C4F867FCC7F847" bold="true" box="[766,786,1968,1994]" pageId="2" pageNumber="3">E</emphasis>
. Label from type bottle. Scale bars: A = 1 mm; B = 0.5 mm; CD = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BA36963805FFFE894D0FEDCFD9FFEA8" blockId="3.[189,1398,266,364]" box="[234,586,266,293]" pageId="3" pageNumber="4">
<emphasis id="B968B571805FFFE894D0FEDCFD9FFEA8" bold="true" box="[234,586,266,293]" pageId="3" pageNumber="4">
<typeStatus id="54A7D7C1805FFFE894D0FEDCFE89FEA8" box="[234,348,267,293]" pageId="3" pageNumber="4" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="4C1C12E0805FFFE895BEFEDCFD9FFEA8" authorityName="Gajbe" authorityYear="2005" box="[388,586,266,293]" class="Arachnida" family="Gnaphosidae" genus="Drassyllus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="jabalpurensis">
<emphasis id="B968B571805FFFE895BEFEDCFD9FFEA8" bold="true" box="[388,586,266,293]" italics="true" pageId="3" pageNumber="4">D. jabalpurensis</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BA36963805FFFE89487FEF9FD48FEE1" blockId="3.[189,1398,266,364]" pageId="3" pageNumber="4">
<materialsCitation id="3B74633E805FFFE89487FEF9FD48FEE1" ID-GBIF-Occurrence="2643239301" collectingDate="1970-11-24" collectorName="D. K. Ghosal" country="India" location="Bhirki Village on Jabalpur-Chargawan" municipality="Jabalpur" pageId="3" pageNumber="4" specimenCode="5452/18" specimenCount="1" specimenCount-female="1" stateProvince="Madhya Pradesh" typeStatus="holotype">
<collectingCountry id="F30B29F3805FFFE89487FEF9FEC7FEC5" box="[189,274,302,328]" name="India" pageId="3" pageNumber="4">INDIA</collectingCountry>
<specimenCount id="9D1AA2EA805FFFE8951BFEF8FEEDFEC4" box="[289,312,303,329]" pageId="3" pageNumber="4" type="female"></specimenCount>
;
<collectingRegion id="49D8A781805FFFE8957CFEF9FDD9FEC5" box="[326,524,302,328]" country="India" name="Madhya Pradesh" pageId="3" pageNumber="4">Madhya Pradesh</collectingRegion>
,
<collectingMunicipality id="6BC7F319805FFFE8962DFEF9FDA9FEC5" box="[535,636,302,328]" pageId="3" pageNumber="4">Jabalpur</collectingMunicipality>
,
<location id="8EC33FB8805FFFE896BDFEF9FB9CFEC5" LSID="urn:lsid:plazi:treatment:03B5D875805DFFEF9594FA49FC97F969:8EC33FB8805FFFE896BDFEF9FB9CFEC5" box="[647,1097,302,329]" country="India" municipality="Jabalpur" name="Bhirki Village on Jabalpur-Chargawan" pageId="3" pageNumber="4" stateProvince="Madhya Pradesh">Bhirki Village on Jabalpur-Chargawan</location>
road;
<date id="FFA24FA3805FFFE890A8FEF9FAE7FEC5" box="[1170,1330,302,329]" pageId="3" pageNumber="4" value="1970-11-24">
<collectingDate id="EFE6B64B805FFFE890A8FEF9FAE7FEC5" box="[1170,1330,302,329]" pageId="3" pageNumber="4" value="1970-11-24">24 Nov. 1970</collectingDate>
</date>
;
<collectorName id="26E90CB5805FFFE89107FEF8FEC5FEE6" pageId="3" pageNumber="4">D.K. Ghosal</collectorName>
leg.; NZC-ZSI, Kolkata
<specimenCode id="DBBAC118805FFFE8960FFE86FD4CFEE1" box="[565,665,337,364]" pageId="3" pageNumber="4">5452/18</specimenCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF6339EB805FFFE89487F8B1FDC4F867" ID-DOI="http://doi.org/10.5281/zenodo.3908349" ID-Zenodo-Dep="3908349" httpUri="https://zenodo.org/record/3908349/files/figure.png" pageId="3" pageNumber="4" startId="3.[189,232,1894,1920]" targetBox="[198,1387,518,1852]" targetPageId="3">
<paragraph id="8BA36963805FFFE89487F8B1FDC4F867" blockId="3.[189,1399,1894,2026]" pageId="3" pageNumber="4">
<emphasis id="B968B571805FFFE89487F8B1FEC4F80D" bold="true" box="[189,273,1894,1920]" pageId="3" pageNumber="4">Fig. 2.</emphasis>
<taxonomicName id="4C1C12E0805FFFE89525F8B1FC52F80C" authority="(Tikader &amp; Gajbe, 1976)" authorityName="Sankaran &amp; Caleb &amp; Sebastian" authorityYear="2020" baseAuthorityName="Tikader &amp; Gajbe" baseAuthorityYear="1976" box="[287,903,1894,1921]" class="Arachnida" family="Gnaphosidae" genus="Cryptodrassus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="khajuriai" status="comb. nov.">
<emphasis id="B968B571805FFFE89525F8B1FD91F80D" box="[287,580,1894,1920]" italics="true" pageId="3" pageNumber="4">Cryptodrassus khajuriai</emphasis>
(
<bibRefCitation id="EF8D1492805FFFE89663F8B1FCAAF80C" author="Tikader B. K. &amp; Gajbe U. A." box="[601,895,1894,1921]" pageId="3" pageNumber="4" pagination="431 - 434" refId="ref5325" refString="Tikader B. K. &amp; Gajbe U. A. 1976. New spiders of Drassyllus from India (Gnaphosidae). Oriental Insects 10: 431 - 434." type="journal article" year="1976">Tikader &amp; Gajbe, 1976</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A25B080A805FFFE897ACF8B1FBC8F80C" box="[918,1053,1894,1921]" pageId="3" pageNumber="4" rank="species">comb. nov.</taxonomicNameLabel>
, ♀, holotype of
<taxonomicName id="4C1C12E0805FFFE890C3F8B1FE22F829" authority="Gajbe, 2005" authorityName="Gajbe" authorityYear="2005" class="Arachnida" family="Gnaphosidae" genus="Drassyllus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="jabalpurensis">
<emphasis id="B968B571805FFFE890C3F8B1FE89F82E" italics="true" pageId="3" pageNumber="4">Drassyllus jabalpurensis</emphasis>
<bibRefCitation id="EF8D1492805FFFE8955FF85EFE22F829" author="Gajbe U. A." box="[357,503,1929,1956]" pageId="3" pageNumber="4" pagination="111 - 140" refId="ref4969" refString="Gajbe U. A. 2005. Studies on some spiders of the family Gnaphosidae (Araneae: Arachnida) from Madhya Pradesh, India. Records of the Zoological Survey of India 105: 111 - 140." type="journal article" year="2005">Gajbe, 2005</bibRefCitation>
</taxonomicName>
(NZC-ZSI-5452/18).
<emphasis id="B968B571805FFFE896C5F85EFCC1F82E" bold="true" box="[767,788,1929,1955]" pageId="3" pageNumber="4">A</emphasis>
. Habitus, dorsal view.
<emphasis id="B968B571805FFFE89011F85EFBEAF82E" bold="true" box="[1067,1087,1929,1955]" pageId="3" pageNumber="4">B</emphasis>
. Eyes of the same, dorsal view.
<emphasis id="B968B571805FFFE89539F87BFECDF84B" bold="true" box="[259,280,1964,1990]" pageId="3" pageNumber="4">C</emphasis>
. Epigyne, ventral view.
<emphasis id="B968B571805FFFE8960FF87AFD9FF84A" bold="true" box="[565,586,1965,1991]" pageId="3" pageNumber="4">D</emphasis>
. Same, dorsal view.
<emphasis id="B968B571805FFFE89706F87AFC85F84A" bold="true" box="[828,848,1965,1991]" pageId="3" pageNumber="4">E</emphasis>
. Label from type bottle. Scale bars: A = 1 mm; B = 0.5 mm; CD = 0.2 mm.
</paragraph>
</caption>
<subSubSection id="C3063AE88058FFEF9487FEDCFCE4FA51" pageId="4" pageNumber="5" type="description">
<paragraph id="8BA369638058FFEF9487FEDCFDC4FEA8" blockId="4.[189,529,267,293]" box="[189,529,267,293]" pageId="4" pageNumber="5">
<heading id="D0EBDE0F8058FFEF9487FEDCFDC4FEA8" bold="true" box="[189,529,267,293]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="B968B5718058FFEF9487FEDCFDC4FEA8" bold="true" box="[189,529,267,293]" pageId="4" pageNumber="5">Supplementary description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA369638058FFEF94D0FEEBFDCEFEDB" blockId="4.[189,1399,316,640]" box="[234,539,316,342]" pageId="4" pageNumber="5">
<emphasis id="B968B5718058FFEF94D0FEEBFE90FEDB" bold="true" box="[234,325,316,342]" pageId="4" pageNumber="5">Female</emphasis>
(
<typeStatus id="54A7D7C18058FFEF956FFEEBFE15FEDB" box="[341,448,316,342]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
,
<figureCitation id="132775E68058FFEF95F6FEEBFDDAFEDB" box="[460,527,316,342]" captionStart="Fig" captionStartId="2.[189,232,1897,1923]" captionTargetBox="[198,1388,467,1853]" captionTargetId="figure@2.[189,1398,456,1863]" captionTargetPageId="2" captionText="Fig. 1. Cryptodrassus khajuriai (Tikader &amp; Gajbe, 1976) comb. nov., ♀, holotype of Drassyllus khajuriai Tikader &amp; Gajbe, 1976 (NZC-ZSI-5043/18). A. Habitus, dorsal view. B. Eyes of the same, dorsal view. C. Epigyne, ventral view. D. Same, dorsal view. E. Label from type bottle. Scale bars: A = 1 mm; B = 0.5 mm; CD = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3908347" httpUri="https://zenodo.org/record/3908347/files/figure.png" pageId="4" pageNumber="5">Fig. 1</figureCitation>
)
</paragraph>
<paragraph id="8BA369638058FFEF9487FEB6FD98FD0D" blockId="4.[189,1399,316,640]" pageId="4" pageNumber="5">
Body length 5.38. Prosoma: length 2.18, width 1.76. Opisthosoma: length 3.20, width 1.71. Eye diameters: ALE 0.12, AME 0.13, PLE 0.11, PME 0.19. Eye interdistances: AMEAME 0.05, AME PME 0.09, PMEPLE 0.04. Chelicerae length 0.67. Measurement of leg II 5.27 [1.51, 0.87, 1.03, 1.13, 0.73]. Epigyne (
<typeStatus id="54A7D7C18058FFEF95F3FE06FDE1FE66" box="[457,564,465,491]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
,
<figureCitation id="132775E68058FFEF967EFE06FD5CFE66" box="[580,649,465,491]" captionStart="Fig" captionStartId="2.[189,232,1897,1923]" captionTargetBox="[198,1388,467,1853]" captionTargetId="figure@2.[189,1398,456,1863]" captionTargetPageId="2" captionText="Fig. 1. Cryptodrassus khajuriai (Tikader &amp; Gajbe, 1976) comb. nov., ♀, holotype of Drassyllus khajuriai Tikader &amp; Gajbe, 1976 (NZC-ZSI-5043/18). A. Habitus, dorsal view. B. Eyes of the same, dorsal view. C. Epigyne, ventral view. D. Same, dorsal view. E. Label from type bottle. Scale bars: A = 1 mm; B = 0.5 mm; CD = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3908347" httpUri="https://zenodo.org/record/3908347/files/figure.png" pageId="4" pageNumber="5">Fig. 1</figureCitation>
CD): Epigynal plate moderately sclerotized, triangular, with smoothly triangular atrium with anterior hood (
<figureCitation id="132775E68058FFEF96C0FE21FC8DFD9D" box="[762,856,502,528]" captionStart="Fig" captionStartId="2.[189,232,1897,1923]" captionTargetBox="[198,1388,467,1853]" captionTargetId="figure@2.[189,1398,456,1863]" captionTargetPageId="2" captionText="Fig. 1. Cryptodrassus khajuriai (Tikader &amp; Gajbe, 1976) comb. nov., ♀, holotype of Drassyllus khajuriai Tikader &amp; Gajbe, 1976 (NZC-ZSI-5043/18). A. Habitus, dorsal view. B. Eyes of the same, dorsal view. C. Epigyne, ventral view. D. Same, dorsal view. E. Label from type bottle. Scale bars: A = 1 mm; B = 0.5 mm; CD = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3908347" httpUri="https://zenodo.org/record/3908347/files/figure.png" pageId="4" pageNumber="5">Fig. 1C</figureCitation>
). Copulatory openings indistinct. Copulatory ducts long, tubular, thick proximally and narrowed distally, obliquely twisted (
<figureCitation id="132775E68058FFEF9048FDCCFB07FDB8" box="[1138,1234,539,565]" captionStart="Fig" captionStartId="2.[189,232,1897,1923]" captionTargetBox="[198,1388,467,1853]" captionTargetId="figure@2.[189,1398,456,1863]" captionTargetPageId="2" captionText="Fig. 1. Cryptodrassus khajuriai (Tikader &amp; Gajbe, 1976) comb. nov., ♀, holotype of Drassyllus khajuriai Tikader &amp; Gajbe, 1976 (NZC-ZSI-5043/18). A. Habitus, dorsal view. B. Eyes of the same, dorsal view. C. Epigyne, ventral view. D. Same, dorsal view. E. Label from type bottle. Scale bars: A = 1 mm; B = 0.5 mm; CD = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3908347" httpUri="https://zenodo.org/record/3908347/files/figure.png" pageId="4" pageNumber="5">Fig. 1D</figureCitation>
). Receptacles small, spherical, nearly contiguous, lying adjacent to posterior epigynal margin (
<figureCitation id="132775E68058FFEF9049FD96FB04FDD6" box="[1139,1233,577,603]" captionStart="Fig" captionStartId="2.[189,232,1897,1923]" captionTargetBox="[198,1388,467,1853]" captionTargetId="figure@2.[189,1398,456,1863]" captionTargetPageId="2" captionText="Fig. 1. Cryptodrassus khajuriai (Tikader &amp; Gajbe, 1976) comb. nov., ♀, holotype of Drassyllus khajuriai Tikader &amp; Gajbe, 1976 (NZC-ZSI-5043/18). A. Habitus, dorsal view. B. Eyes of the same, dorsal view. C. Epigyne, ventral view. D. Same, dorsal view. E. Label from type bottle. Scale bars: A = 1 mm; B = 0.5 mm; CD = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3908347" httpUri="https://zenodo.org/record/3908347/files/figure.png" pageId="4" pageNumber="5">Fig. 1D</figureCitation>
). Fertilization ducts narrow, diverging (
<figureCitation id="132775E68058FFEF95D8FDB1FDEAFD0D" box="[482,575,614,640]" captionStart="Fig" captionStartId="2.[189,232,1897,1923]" captionTargetBox="[198,1388,467,1853]" captionTargetId="figure@2.[189,1398,456,1863]" captionTargetPageId="2" captionText="Fig. 1. Cryptodrassus khajuriai (Tikader &amp; Gajbe, 1976) comb. nov., ♀, holotype of Drassyllus khajuriai Tikader &amp; Gajbe, 1976 (NZC-ZSI-5043/18). A. Habitus, dorsal view. B. Eyes of the same, dorsal view. C. Epigyne, ventral view. D. Same, dorsal view. E. Label from type bottle. Scale bars: A = 1 mm; B = 0.5 mm; CD = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3908347" httpUri="https://zenodo.org/record/3908347/files/figure.png" pageId="4" pageNumber="5">Fig. 1D</figureCitation>
).
</paragraph>
<paragraph id="8BA369638058FFEF94D0FD7DFEFFFD49" blockId="4.[189,313,682,745]" box="[234,298,682,708]" pageId="4" pageNumber="5">
<emphasis id="B968B5718058FFEF94D0FD7DFEFFFD49" bold="true" box="[234,298,682,708]" pageId="4" pageNumber="5">Male</emphasis>
</paragraph>
<paragraph id="8BA369638058FFEF9487FD18FEECFD64" blockId="4.[189,313,682,745]" box="[189,313,719,745]" pageId="4" pageNumber="5">Unknown.</paragraph>
<paragraph id="8BA369638058FFEF9487FCC5FDC4FCA1" blockId="4.[189,529,786,813]" box="[189,529,786,813]" pageId="4" pageNumber="5">
<heading id="D0EBDE0F8058FFEF9487FCC5FDC4FCA1" bold="true" box="[189,529,786,813]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="B968B5718058FFEF9487FCC5FDC4FCA1" bold="true" box="[189,529,786,813]" pageId="4" pageNumber="5">Justification of the transfer</emphasis>
</heading>
</paragraph>
<paragraph id="8BA369638058FFEF9487FC94FDF1FB5F" blockId="4.[189,1399,835,1234]" pageId="4" pageNumber="5">
<bibRefCitation id="EF8D14928058FFEF9487FC94FE0CFCD3" author="Tikader B. K. &amp; Gajbe U. A." box="[189,473,835,862]" pageId="4" pageNumber="5" pagination="431 - 434" refId="ref5325" refString="Tikader B. K. &amp; Gajbe U. A. 1976. New spiders of Drassyllus from India (Gnaphosidae). Oriental Insects 10: 431 - 434." type="journal article" year="1976">Tikader &amp; Gajbe (1976)</bibRefCitation>
described
<taxonomicName id="4C1C12E08058FFEF9663FC93FD3CFCD0" authorityName="Sankaran &amp; Caleb &amp; Sebastian" authorityYear="2020" baseAuthorityName="Tikader &amp; Gajbe" baseAuthorityYear="1976" box="[601,745,835,861]" class="Arachnida" family="Gnaphosidae" genus="Cryptodrassus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="khajuriai">
<emphasis id="B968B5718058FFEF9663FC93FDBBFCD0" box="[601,622,836,861]" italics="true" pageId="4" pageNumber="5">D</emphasis>
.
<emphasis id="B968B5718058FFEF9647FC94FD3CFCD0" box="[637,745,835,861]" italics="true" pageId="4" pageNumber="5">khajuriai</emphasis>
</taxonomicName>
on the basis of a female specimen collected in
<collectingRegion id="49D8A7818058FFEF912CFC94FECBFC0F" country="India" name="Madhya Pradesh" pageId="4" pageNumber="5">Madhya Pradesh</collectingRegion>
. It resembles
<taxonomicName id="4C1C12E08058FFEF95F6FCBFFD9CFC0F" authorityName="Chamberlin" authorityYear="1922" box="[460,585,872,898]" class="Arachnida" family="Gnaphosidae" genus="Drassyllus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B968B5718058FFEF95F6FCBFFD9CFC0F" box="[460,585,872,898]" italics="true" pageId="4" pageNumber="5">Drassyllus</emphasis>
</taxonomicName>
spp. only in the posterior median eyes, which are the largest and remaining contiguous in both
<taxonomicName id="4C1C12E08058FFEF961FFC5AFD05FC2A" authorityName="Miller" authorityYear="1943" box="[549,720,909,935]" class="Arachnida" family="Gnaphosidae" genus="Cryptodrassus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B968B5718058FFEF961FFC5AFD05FC2A" box="[549,720,909,935]" italics="true" pageId="4" pageNumber="5">Cryptodrassus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1C12E08058FFEF9737FC5AFC5FFC2A" authorityName="Chamberlin" authorityYear="1922" box="[781,906,909,935]" class="Arachnida" family="Gnaphosidae" genus="Drassyllus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B968B5718058FFEF9737FC5AFC5FFC2A" box="[781,906,909,935]" italics="true" pageId="4" pageNumber="5">Drassyllus</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EF8D14928058FFEF97F6FC5AFBA7FC25" author="Murphy J." box="[972,1138,909,936]" pageId="4" pageNumber="5" refId="ref5125" refString="Murphy J. 2007. Gnaphosid Genera of the World. British Arachnological Society, St Neots, Cambridgeshire." type="book" year="2007">Murphy 2007</bibRefCitation>
: figs 184, 528; herein
<figureCitation id="132775E68058FFEF9487FC64FEC0FC40" box="[189,277,947,973]" captionStart="Fig" captionStartId="2.[189,232,1897,1923]" captionTargetBox="[198,1388,467,1853]" captionTargetId="figure@2.[189,1398,456,1863]" captionTargetPageId="2" captionText="Fig. 1. Cryptodrassus khajuriai (Tikader &amp; Gajbe, 1976) comb. nov., ♀, holotype of Drassyllus khajuriai Tikader &amp; Gajbe, 1976 (NZC-ZSI-5043/18). A. Habitus, dorsal view. B. Eyes of the same, dorsal view. C. Epigyne, ventral view. D. Same, dorsal view. E. Label from type bottle. Scale bars: A = 1 mm; B = 0.5 mm; CD = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3908347" httpUri="https://zenodo.org/record/3908347/files/figure.png" pageId="4" pageNumber="5">Fig. 1B</figureCitation>
). Detailed examination of the
<typeStatus id="54A7D7C18058FFEF9648FC64FD0CFC40" box="[626,729,947,973]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C1C12E08058FFEF96C7FC64FC59FC40" authorityName="Sankaran &amp; Caleb &amp; Sebastian" authorityYear="2020" baseAuthorityName="Tikader &amp; Gajbe" baseAuthorityYear="1976" box="[765,908,947,973]" class="Arachnida" family="Gnaphosidae" genus="Cryptodrassus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="khajuriai">
<emphasis id="B968B5718058FFEF96C7FC64FC59FC40" box="[765,908,947,973]" italics="true" pageId="4" pageNumber="5">D. khajuriai</emphasis>
</taxonomicName>
revealed that it has all of the characteristic features of
<taxonomicName id="4C1C12E08058FFEF9571FC0FFE23FC7F" authorityName="Miller" authorityYear="1943" box="[331,502,984,1010]" class="Arachnida" family="Gnaphosidae" genus="Cryptodrassus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B968B5718058FFEF9571FC0FFE23FC7F" box="[331,502,984,1010]" italics="true" pageId="4" pageNumber="5">Cryptodrassus</emphasis>
</taxonomicName>
spp. as described and illustrated for
<taxonomicName id="4C1C12E08058FFEF97F2FC0FFED1FB95" authority="(Balogh, 1935)" baseAuthorityName="Balogh" baseAuthorityYear="1935" class="Arachnida" family="Gnaphosidae" genus="Cryptodrassus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="hungaricus">
<emphasis id="B968B5718058FFEF97F2FC0FFAD1FC7F" box="[968,1284,984,1010]" italics="true" pageId="4" pageNumber="5">Cryptodrassus hungaricus</emphasis>
(Balogh, 1935)
</taxonomicName>
, the type species of the genus (
<bibRefCitation id="EF8D14928058FFEF965DFC2AFCD8FB9A" author="Murphy J." box="[615,781,1021,1047]" pageId="4" pageNumber="5" refId="ref5125" refString="Murphy J. 2007. Gnaphosid Genera of the World. British Arachnological Society, St Neots, Cambridgeshire." type="book" year="2007">Murphy 2007</bibRefCitation>
: figs 528529;
<bibRefCitation id="EF8D14928058FFEF9786FC2AFB2AFB9A" author="Kovblyuk M. M. &amp; Nadolny A. A." box="[956,1279,1021,1047]" pageId="4" pageNumber="5" pagination="189 - 197" refId="ref5057" refString="Kovblyuk M. M. &amp; Nadolny A. A. 2010. Cryptodrassus hungaricus and Leptodrassex memorialis from Crimea (Aranei: Gnaphosidae). Arthropoda Selecta 19: 189 - 197." type="journal article" year="2010">Kovblyuk &amp; Nadolny 2010</bibRefCitation>
: figs 78),
<taxonomicName id="4C1C12E08058FFEF9487FBF5FCE8FBB0" authority="(O. Pickard-Cambridge, 1872)" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1872" box="[189,829,1058,1085]" class="Arachnida" family="Gnaphosidae" genus="Cryptodrassus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="helvolus">
<emphasis id="B968B5718058FFEF9487FBF5FE04FBB1" box="[189,465,1058,1084]" italics="true" pageId="4" pageNumber="5">Cryptodrassus helvolus</emphasis>
(O. Pickard-Cambridge, 1872)
</taxonomicName>
(
<bibRefCitation id="EF8D14928058FFEF9776FBF4FC05FBB0" author="Levy G." box="[844,976,1059,1085]" pageId="4" pageNumber="5" pagination="93 - 158" refId="ref5091" refString="Levy G. 1998. The ground-spider genera Setaphis, Trachyzelotes, Zelotes, and Drassyllus (Araneae: Gnaphosidae) in Israel. Israel Journal of Zoology 44: 93 - 158." type="journal article" year="1998">Levy 1998</bibRefCitation>
: fig. 126;
<bibRefCitation id="EF8D14928058FFEF9079FBF4FF29FBEF" author="Chatzaki M. &amp; Russell-Smith A." pageId="4" pageNumber="5" pagination="237 - 255" refId="ref4861" refString="Chatzaki M. &amp; Russell-Smith A. 2017. New species and new records of ground spiders (Araneae: Gnaphosidae) from Cyprus. Zootaxa 4329: 237 - 255. https: // doi. org / 10.11646 / zootaxa. 4329.3.3" type="journal article" year="2017">Chatzaki &amp; Russell-Smith 2017</bibRefCitation>
: fig. 5) and
<taxonomicName id="4C1C12E08058FFEF95A8FB9FFCA7FBEF" authority="(Levy, 1998)" baseAuthorityName="Levy" baseAuthorityYear="1998" box="[402,882,1096,1122]" class="Arachnida" family="Gnaphosidae" genus="Cryptodrassus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="helvoloides">
<emphasis id="B968B5718058FFEF95A8FB9FFD1BFBEF" box="[402,718,1096,1122]" italics="true" pageId="4" pageNumber="5">Cryptodrassus helvoloides</emphasis>
(
<bibRefCitation id="EF8D14928058FFEF96DAFB9FFCBFFBEF" author="Levy G." box="[736,874,1096,1122]" pageId="4" pageNumber="5" pagination="93 - 158" refId="ref5091" refString="Levy G. 1998. The ground-spider genera Setaphis, Trachyzelotes, Zelotes, and Drassyllus (Araneae: Gnaphosidae) in Israel. Israel Journal of Zoology 44: 93 - 158." type="journal article" year="1998">Levy, 1998</bibRefCitation>
)
</taxonomicName>
(
<bibRefCitation id="EF8D14928058FFEF97BEFB9FFADCFBEF" author="Chatzaki M. &amp; Russell-Smith A." box="[900,1289,1096,1122]" pageId="4" pageNumber="5" pagination="237 - 255" refId="ref4861" refString="Chatzaki M. &amp; Russell-Smith A. 2017. New species and new records of ground spiders (Araneae: Gnaphosidae) from Cyprus. Zootaxa 4329: 237 - 255. https: // doi. org / 10.11646 / zootaxa. 4329.3.3" type="journal article" year="2017">Chatzaki &amp; Russell-Smith 2017</bibRefCitation>
: fig. 11): PMEs largest, irregular, all other eyes round, cheliceral promargin with three and retromargin with one tooth and epigyne with wide, anterior atrium. Considering these observations, we propose to transfer
<taxonomicName id="4C1C12E08058FFEF9487FB6FFE98FB5F" authorityName="Sankaran &amp; Caleb &amp; Sebastian" authorityYear="2020" baseAuthorityName="Tikader &amp; Gajbe" baseAuthorityYear="1976" box="[189,333,1208,1234]" class="Arachnida" family="Gnaphosidae" genus="Cryptodrassus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="khajuriai">
<emphasis id="B968B5718058FFEF9487FB6FFF07FB5C" box="[189,210,1208,1233]" italics="true" pageId="4" pageNumber="5">D</emphasis>
.
<emphasis id="B968B5718058FFEF94DBFB6FFE98FB5F" box="[225,333,1208,1234]" italics="true" pageId="4" pageNumber="5">khajuriai</emphasis>
</taxonomicName>
to
<taxonomicName id="4C1C12E08058FFEF9548FB6FFDC8FB5F" authorityName="Miller" authorityYear="1943" box="[370,541,1208,1234]" class="Arachnida" family="Gnaphosidae" genus="Cryptodrassus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B968B5718058FFEF9548FB6FFDC8FB5F" box="[370,541,1208,1234]" italics="true" pageId="4" pageNumber="5">Cryptodrassus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA369638058FFEF9487FB2CFDFEFA9B" blockId="4.[189,555,1275,1302]" box="[189,555,1275,1302]" pageId="4" pageNumber="5">
<heading id="D0EBDE0F8058FFEF9487FB2CFDFEFA9B" bold="true" box="[189,555,1275,1302]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="B968B5718058FFEF9487FB2CFDFEFA9B" bold="true" box="[189,555,1275,1302]" pageId="4" pageNumber="5">Justification of the synonymy</emphasis>
</heading>
</paragraph>
<paragraph id="8BA369638058FFEF9487FAFBFCE4FA51" blockId="4.[189,1399,1324,1500]" pageId="4" pageNumber="5">
A detailed examination of the
<typeStatus id="54A7D7C18058FFEF9608FAFBFD4CFACB" box="[562,665,1324,1350]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C1C12E08058FFEF96F2FAFAFC5BFACB" authorityName="Gajbe" authorityYear="2005" box="[712,910,1324,1350]" class="Arachnida" family="Gnaphosidae" genus="Drassyllus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="jabalpurensis">
<emphasis id="B968B5718058FFEF96F2FAFAFC5BFACB" box="[712,910,1324,1350]" italics="true" pageId="4" pageNumber="5">D. jabalpurensis</emphasis>
</taxonomicName>
shows that it has diagnostic features of
<taxonomicName id="4C1C12E08058FFEF9487FA85FE87FAE6" authorityName="Sankaran &amp; Caleb &amp; Sebastian" authorityYear="2020" baseAuthorityName="Tikader &amp; Gajbe" baseAuthorityYear="1976" box="[189,338,1361,1388]" class="Arachnida" family="Gnaphosidae" genus="Cryptodrassus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="khajuriai">
<emphasis id="B968B5718058FFEF9487FA85FF07FAE6" box="[189,210,1362,1387]" italics="true" pageId="4" pageNumber="5">D</emphasis>
.
<emphasis id="B968B5718058FFEF94DCFA86FE87FAE6" box="[230,338,1361,1387]" italics="true" pageId="4" pageNumber="5">khajuriai</emphasis>
</taxonomicName>
: triangular epigynal atrium with single anterior hood, highly coiled obliquely twisted copulatory ducts with thick proximal and narrow distal parts and spherical and contiguous receptacles (compare
<figureCitation id="132775E68058FFEF9507FA4BFE52FA3B" box="[317,391,1436,1462]" captionStart="Fig" captionStartId="2.[189,232,1897,1923]" captionTargetBox="[198,1388,467,1853]" captionTargetId="figure@2.[189,1398,456,1863]" captionTargetPageId="2" captionText="Fig. 1. Cryptodrassus khajuriai (Tikader &amp; Gajbe, 1976) comb. nov., ♀, holotype of Drassyllus khajuriai Tikader &amp; Gajbe, 1976 (NZC-ZSI-5043/18). A. Habitus, dorsal view. B. Eyes of the same, dorsal view. C. Epigyne, ventral view. D. Same, dorsal view. E. Label from type bottle. Scale bars: A = 1 mm; B = 0.5 mm; CD = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3908347" httpUri="https://zenodo.org/record/3908347/files/figure.png" pageId="4" pageNumber="5">Fig. 1</figureCitation>
CD with
<figureCitation id="132775E68058FFEF9621FA4BFDBFFA3B" box="[539,618,1436,1462]" captionStart="Fig" captionStartId="3.[189,232,1894,1920]" captionTargetBox="[198,1387,518,1852]" captionTargetId="figure@3.[189,1398,511,1859]" captionTargetPageId="3" captionText="Fig. 2. Cryptodrassus khajuriai (Tikader &amp; Gajbe, 1976) comb. nov., ♀, holotype of Drassyllus jabalpurensis Gajbe, 2005 (NZC-ZSI-5452/18). A. Habitus, dorsal view. B. Eyes of the same, dorsal view. C. Epigyne, ventral view. D. Same, dorsal view. E. Label from type bottle. Scale bars: A = 1 mm; B = 0.5 mm; CD = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3908349" httpUri="https://zenodo.org/record/3908349/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
CD). Based on these observations, we propose to consider
<taxonomicName id="4C1C12E08058FFEF9487FA15FE55FA56" authorityName="Gajbe" authorityYear="2005" box="[189,384,1473,1499]" class="Arachnida" family="Gnaphosidae" genus="Drassyllus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="jabalpurensis">
<emphasis id="B968B5718058FFEF9487FA15FE55FA56" box="[189,384,1473,1499]" italics="true" pageId="4" pageNumber="5">D. jabalpurensis</emphasis>
</taxonomicName>
as a junior synonym of
<taxonomicName id="4C1C12E08058FFEF96A1FA15FCFEFA56" authorityName="Sankaran &amp; Caleb &amp; Sebastian" authorityYear="2020" baseAuthorityName="Tikader &amp; Gajbe" baseAuthorityYear="1976" box="[667,811,1473,1500]" class="Arachnida" family="Gnaphosidae" genus="Cryptodrassus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="khajuriai">
<emphasis id="B968B5718058FFEF96A1FA15FD65FA56" box="[667,688,1474,1499]" italics="true" pageId="4" pageNumber="5">D</emphasis>
.
<emphasis id="B968B5718058FFEF9685FA16FCFEFA56" box="[703,811,1473,1499]" italics="true" pageId="4" pageNumber="5">khajuriai</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3063AE88058FFEF9487F9D2FC97F969" pageId="4" pageNumber="5" type="discussion">
<paragraph id="8BA369638058FFEF9487F9D2FEFAF992" blockId="4.[189,303,1541,1567]" box="[189,303,1541,1567]" pageId="4" pageNumber="5">
<heading id="D0EBDE0F8058FFEF9487F9D2FEFAF992" bold="true" box="[189,303,1541,1567]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="B968B5718058FFEF9487F9D2FEFAF992" bold="true" box="[189,303,1541,1567]" pageId="4" pageNumber="5">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BA369638058FFEF9487F9E2FC97F969" blockId="4.[189,1399,1589,1765]" pageId="4" pageNumber="5">
The ZSI collection has one glass bottle for
<taxonomicName id="4C1C12E08058FFEF96FBF9E1FC87F9C2" authorityName="Sankaran &amp; Caleb &amp; Sebastian" authorityYear="2020" baseAuthorityName="Tikader &amp; Gajbe" baseAuthorityYear="1976" box="[705,850,1589,1616]" class="Arachnida" family="Gnaphosidae" genus="Cryptodrassus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="khajuriai">
<emphasis id="B968B5718058FFEF96FBF9E1FD03F9C2" box="[705,726,1590,1615]" italics="true" pageId="4" pageNumber="5">D</emphasis>
.
<emphasis id="B968B5718058FFEF96DCF9E2FC87F9C2" box="[742,850,1589,1615]" italics="true" pageId="4" pageNumber="5">khajuriai</emphasis>
</taxonomicName>
, labeled as holotype (5043/18), containing a female specimen in fairly good condition, with only left leg II. The same bottle has a small glass vial containing the dissected epigyne. The ZSI collection has one glass bottle for
<taxonomicName id="4C1C12E08058FFEF9075F957FAC7F917" authorityName="Gajbe" authorityYear="2005" box="[1103,1298,1664,1690]" class="Arachnida" family="Gnaphosidae" genus="Drassyllus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="jabalpurensis">
<emphasis id="B968B5718058FFEF9075F957FAC7F917" box="[1103,1298,1664,1690]" italics="true" pageId="4" pageNumber="5">D. jabalpurensis</emphasis>
</taxonomicName>
, labeled as holotype (with no register number), containing a female specimen in bad condition. The same bottle has a small glass vial containing the dissected epigyne.
</paragraph>
</subSubSection>
</treatment>
</document>