239 lines
31 KiB
XML
239 lines
31 KiB
XML
<document id="CE441DF32252BC37817267BFDDE68991" ID="10.11646/zootaxa.4263.2.10SLASH10996" ID-CLB-Dataset="33081" ID-DOI="10.11646/zootaxa.4263.2.10" ID-GBIF-Dataset="c4c23fc2-2586-49bc-b6ca-5ed8cb867965" ID-ISSN="1175-5326" ID-Zenodo-Dep="573188" ID-ZooBank="23426070-2520-4F06-B0CF-643FD9680A75" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1494321999588" checkinUser="plazi" docAuthor="Cranston, Peter S." docDate="2017" docId="03AD9627A93D5E70E588FD04FB6FFBD8" docLanguage="en" docName="zootaxa.4263.2.10SLASH10996.pdf" docOrigin="Zootaxa 4263 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Yarrhpelopia Cranston, 2017, gen. nov." docType="treatment" docUuid="EE6A29B3-13BD-4D8D-8635-11177CE7C610" docUuidSource="ZooBank" docVersion="8" lastPageNumber="371" masterDocId="FF94EE5FA93C5E72E51FFF88FFF7FFE5" masterDocTitle="A new genus and species of Australian Tanypodinae (Diptera: Chironomidae) tolerant to mine waste" masterLastPageNumber="377" masterPageNumber="369" pageNumber="370" updateTime="1698439406747" updateUser="ExternalLinkService">
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<mods:title id="8E72558BF301CBAEC2A7B0517C6FF3C3">A new genus and species of Australian Tanypodinae (Diptera: Chironomidae) tolerant to mine waste</mods:title>
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<mods:namePart id="D792C2ECF9FF97B59F54E04321EBB192">Cranston, Peter S.</mods:namePart>
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<mods:date id="3B6D9542D1F51703F655F403DABB098E">2017</mods:date>
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<treatment id="03AD9627A93D5E70E588FD04FB6FFBD8" ID-DOI="http://doi.org/10.5281/zenodo.6044855" ID-GBIF-Taxon="130509199" ID-Zenodo-Dep="6044855" LSID="urn:lsid:zoobank.org:act:EE6A29B3-13BD-4D8D-8635-11177CE7C610" httpUri="http://treatment.plazi.org/id/03AD9627A93D5E70E588FD04FB6FFBD8" lastPageId="2" lastPageNumber="371" pageId="1" pageNumber="370">
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<subSubSection id="C31E74BAA93D5E73E588FD04FEE6FD2D" pageId="1" pageNumber="370" type="nomenclature">
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<paragraph id="8BBB2731A93D5E73E588FD04FE5EFD42" blockId="1.[151,960,652,749]" box="[151,425,652,679]" pageId="1" pageNumber="370">
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<heading id="D0F3905DA93D5E73E588FD04FE5EFD42" bold="true" box="[151,425,652,679]" fontSize="11" level="1" pageId="1" pageNumber="370" reason="1">
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<emphasis id="B970FB23A93D5E73E588FD04FE5EFD42" bold="true" box="[151,425,652,679]" pageId="1" pageNumber="370">
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<taxonomicName id="4C045CB2A93D5E73E588FD04FEC3FD43" ID-CoL="89RQZ" ID-ENA="2781158" box="[151,308,652,678]" class="Insecta" family="Chironomidae" genus="Yarrhpelopia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B970FB23A93D5E73E588FD04FEC3FD43" bold="true" box="[151,308,652,678]" italics="true" pageId="1" pageNumber="370">Yarrhpelopia</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2434658A93D5E73E424FD05FE5EFD42" box="[315,425,653,679]" pageId="1" pageNumber="370" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BBB2731A93D5E73E588FD27FEE6FD2D" blockId="1.[151,960,652,749]" box="[151,273,687,712]" pageId="1" pageNumber="370">
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(
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<figureCitation id="133F3BB4A93D5E73E580FD27FEFEFD2D" box="[159,265,687,712]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,250,1427,1449]" captionStartId-1="5.[151,250,1936,1958]" captionStartId-2="6.[151,250,1760,1782]" captionTargetBox-0="[165,1427,536,1399]" captionTargetBox-1="[170,1415,206,1899]" captionTargetBox-2="[168,1417,204,1733]" captionTargetId-0="figure@3.[151,1436,519,1403]" captionTargetId-1="figure@5.[151,1436,193,1915]" captionTargetId-2="figure@6.[151,1437,193,1739]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURE 1. Yarrhpelopia norrisi gen. n. sp. n. Adult male. A. Antennal apex; B. Wing; C. Tibial apices, TI – TIII L to R; D. Hypopygium, left side dorsal, right side stylised ventral / internal." captionText-1="FIGURE 2. Yarrhpelopia norrisi gen. n. sp. n. Pupa. A. Thoracic horn and thoracic comb; B. Dorsal abdomen; C. Tergite IV spinulation (' shagreen'), detail; D. Anal lobe, left half." captionText-2="FIGURE 3. Yarrhpelopia norrisi gen. n. sp. n. Larva. A. Head, left ventral, right dorsal: Abbreviations: S 5 - S 10 cephalic setae, DP—dorsal pit, VP—ventral pit: B. Segments 2 – 4 of antenna; C. Antennal apex; D. Mandible; E. Ligula; F. Palp; G. Submentum; H. Posterior parapod claws (selected)." httpUri-0="https://zenodo.org/record/573189/files/figure.png" httpUri-1="https://zenodo.org/record/573190/files/figure.png" httpUri-2="https://zenodo.org/record/573191/files/figure.png" pageId="1" pageNumber="370">Figs. 1–3</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C31E74BAA93D5E73E588FD5CFD13FCD7" pageId="1" pageNumber="370" type="description">
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<paragraph id="8BBB2731A93D5E73E588FD5CFC37FD08" blockId="1.[151,960,652,749]" box="[151,960,724,749]" pageId="1" pageNumber="370">rn:lsid:zoobank.org:act: EE6A29B3-13BD-4D8D-8635-11177CE7C610</paragraph>
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<paragraph id="8BBB2731A93D5E73E588FC94FD13FCD7" blockId="1.[151,740,796,818]" box="[151,740,796,818]" pageId="1" pageNumber="370">
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<taxonomicName id="4C045CB2A93D5E73E588FC94FEE0FCD7" box="[151,279,796,818]" class="Insecta" family="Chironomidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="370" phylum="Arthropoda" rank="tribe" tribe="Pentaneurini">Pentaneurini</taxonomicName>
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genus A (
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<bibRefCitation id="EF955AC0A93D5E73E461FC94FDE1FCD7" author="Cranston" box="[382,534,796,818]" pageId="1" pageNumber="370" refString="Cranston, P. S. (1996) Identification Guide to the Chironomidae of New South Wales. AWT Identification Guide Number 1. Australian Water Technologies Pty Ltd., West Ryde, NSW," type="book" year="1996">Cranston 1996</bibRefCitation>
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, Krosch
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<emphasis id="B970FB23A93D5E73E76FFC95FD6BFCD7" box="[624,668,796,818]" italics="true" pageId="1" pageNumber="370">et al</emphasis>
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. 2007)
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</paragraph>
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</subSubSection>
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<subSubSection id="C31E74BAA93D5E73E588FCD6FBE9FA9D" pageId="1" pageNumber="370" type="diagnosis">
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<paragraph id="8BBB2731A93D5E73E588FCD6FA69FB99" blockId="1.[151,1438,862,2015]" pageId="1" pageNumber="370">
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<emphasis id="B970FB23A93D5E73E588FCD6FE6DFC92" bold="true" box="[151,410,862,887]" pageId="1" pageNumber="370">Diagnosis. Adult male</emphasis>
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. Pedicel setose, flagellomeres with dark plumes; terminal segment of male short, darkened, with apical fine seta (
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<figureCitation id="133F3BB4A93D5E73E499FC0AFE33FC7E" box="[390,452,898,923]" captionStart="FIGURE 1" captionStartId="3.[151,250,1427,1449]" captionTargetBox="[165,1427,536,1399]" captionTargetId="figure@3.[151,1436,519,1403]" captionTargetPageId="3" captionText="FIGURE 1. Yarrhpelopia norrisi gen. n. sp. n. Adult male. A. Antennal apex; B. Wing; C. Tibial apices, TI – TIII L to R; D. Hypopygium, left side dorsal, right side stylised ventral / internal." httpUri="https://zenodo.org/record/573189/files/figure.png" pageId="1" pageNumber="370">Fig. 1</figureCitation>
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A). Wing (
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<figureCitation id="133F3BB4A93D5E73E75FFC0AFD89FC7E" box="[576,638,898,923]" captionStart="FIGURE 1" captionStartId="3.[151,250,1427,1449]" captionTargetBox="[165,1427,536,1399]" captionTargetId="figure@3.[151,1436,519,1403]" captionTargetPageId="3" captionText="FIGURE 1. Yarrhpelopia norrisi gen. n. sp. n. Adult male. A. Antennal apex; B. Wing; C. Tibial apices, TI – TIII L to R; D. Hypopygium, left side dorsal, right side stylised ventral / internal." httpUri="https://zenodo.org/record/573189/files/figure.png" pageId="1" pageNumber="370">Fig. 1</figureCitation>
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B) translucent, veins including crossveins pale, membrane and all veins densely setose; costa (C) extending to apex of R4+5 prior to wing apex, and proximal to end of M3+4; R1 and R4+5 narrowly separated such that R2+3 is poorly visible, but a faint R2 seems to connect to apex of R1 and costa; R4+5 runs close to and terminates in costa. Squama setose, uniserial. Tibial spurs (
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<figureCitation id="133F3BB4A93D5E73E6A3FC7CFC0CFBE8" box="[956,1019,1012,1037]" captionStart="FIGURE 1" captionStartId="3.[151,250,1427,1449]" captionTargetBox="[165,1427,536,1399]" captionTargetId="figure@3.[151,1436,519,1403]" captionTargetPageId="3" captionText="FIGURE 1. Yarrhpelopia norrisi gen. n. sp. n. Adult male. A. Antennal apex; B. Wing; C. Tibial apices, TI – TIII L to R; D. Hypopygium, left side dorsal, right side stylised ventral / internal." httpUri="https://zenodo.org/record/573189/files/figure.png" pageId="1" pageNumber="370">Fig. 1</figureCitation>
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C) 1, 2, 2: elongate with several side teeth, inner spur on t2 and t3 50–60% length of outer, tibial comb of 3 spines on apex of hind tibia; pulvilli absent; claws very slender, pointed. Posterior margin of TIX with 5–6 long setae. Hypopygium (
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<figureCitation id="133F3BB4A93D5E73E162FBB6FB4BFBB2" box="[1149,1212,1086,1111]" captionStart="FIGURE 1" captionStartId="3.[151,250,1427,1449]" captionTargetBox="[165,1427,536,1399]" captionTargetId="figure@3.[151,1436,519,1403]" captionTargetPageId="3" captionText="FIGURE 1. Yarrhpelopia norrisi gen. n. sp. n. Adult male. A. Antennal apex; B. Wing; C. Tibial apices, TI – TIII L to R; D. Hypopygium, left side dorsal, right side stylised ventral / internal." httpUri="https://zenodo.org/record/573189/files/figure.png" pageId="1" pageNumber="370">Fig. 1</figureCitation>
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D) with mediobasal spinose lobe, possibly representing volsella. Parameres strong, long, and dark within cleared gonocoxite (
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<figureCitation id="133F3BB4A93D5E73E010FBEBFABCFB9E" box="[1295,1355,1123,1148]" captionStart="FIGURE 1" captionStartId="3.[151,250,1427,1449]" captionTargetBox="[165,1427,536,1399]" captionTargetId="figure@3.[151,1436,519,1403]" captionTargetPageId="3" captionText="FIGURE 1. Yarrhpelopia norrisi gen. n. sp. n. Adult male. A. Antennal apex; B. Wing; C. Tibial apices, TI – TIII L to R; D. Hypopygium, left side dorsal, right side stylised ventral / internal." httpUri="https://zenodo.org/record/573189/files/figure.png" pageId="1" pageNumber="370">Fig. 1</figureCitation>
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D left).
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</paragraph>
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<paragraph id="8BBB2731A93D5E73E5D8FB0EFBE9FA9D" blockId="1.[151,1438,862,2015]" pageId="1" pageNumber="370">
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<emphasis id="B970FB23A93D5E73E5D8FB0EFEFBFB7A" bold="true" box="[199,268,1158,1183]" pageId="1" pageNumber="370">Pupa.</emphasis>
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With elongate flat-tubular thoracic horn, about 3–4× as long as broad with horn sac near filling lumen, contracted strongly in distal 1/4 with narrow neck connecting to laterally orientated ovoid plastron plate, filling half of surrounding corona; corona oval, faintly defined, occupying c. 25% of length of horn. Thoracic horn (except corona) with complete dense cover of pointed scales. Thoracic comb modestly developed, uniserial row of 5–7 short, apically rounded tubercles. Basal lobe dome-shaped. Thorax smooth. Tergite I with pigmented scar. Shagreen of small tubercle /spinules, dense and aligned in rows of 2–4; L(ateral) setae taeniate on VII (4) and VIII (5), anal lobe outer margin with very fine spines, inner bare, anal setae adhesive.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31E74BAA93D5E73E5D8FA0BFBA2F993" pageId="1" pageNumber="370" type="description">
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<paragraph id="8BBB2731A93D5E73E5D8FA0BFBA2F993" blockId="1.[151,1438,862,2015]" pageId="1" pageNumber="370">
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<emphasis id="B970FB23A93D5E73E5D8FA0BFEE0FA79" bold="true" box="[199,279,1411,1436]" pageId="1" pageNumber="370">Larva.</emphasis>
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Head index c. 0.5; ligula 5-toothed with weakly concave tooth row, inner teeth weakly curved outward and mid-tooth slightly recessed; 2nd antennal segment smooth, AR c. 3, ring organ at c. 60% from base. Lauterborn organs short and apex does not resemble 'tuning fork'. Basal palp segment undivided, ring organ at 75% from base; b sensillum of two sections, basal section about 3 times length of apical section. Mandible with seta subdentalis arising on strong distal molar 'tooth', posterior to well-defined, rounded inner tooth. SSm retracted,
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<date id="FFBA01F1A93D5E73E00BF99EFACFF9CB" box="[1300,1336,1558,1582]" pageId="1" pageNumber="370">V9</date>
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, VP and
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<date id="FFBA01F1A93D5E73E588F9B2FF30F9B7" box="[151,199,1594,1618]" pageId="1" pageNumber="370" value="2010-05">V10</date>
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near aligned 45° to longitudinal axis, with ventral pit (VP) slightly posterior to
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<date id="FFBA01F1A93D5E73E164F9B2FB29F9B7" box="[1147,1246,1594,1618]" pageId="1" pageNumber="370">V9–V10</date>
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; dorsal pit (DP) present, S7 separated from S8. Body without swim hairs. Parapod claws simple, pale.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31E74BAA93D5E73E5D8F909FD7AF97F" box="[199,653,1665,1690]" pageId="1" pageNumber="370" type="materials_examined">
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<paragraph id="8BBB2731A93D5E73E5D8F909FD7AF97F" blockId="1.[151,1438,862,2015]" box="[199,653,1665,1690]" pageId="1" pageNumber="370">
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<emphasis id="B970FB23A93D5E73E5D8F909FEACF97F" bold="true" box="[199,347,1665,1690]" pageId="1" pageNumber="370">Type species</emphasis>
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.
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<taxonomicName id="4C045CB2A93D5E73E476F90AFDB9F97C" box="[361,590,1666,1690]" class="Insecta" family="Chironomidae" genus="Yarrhpelopia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="370" phylum="Arthropoda" rank="species" species="norrisi" status="sp. nov.">
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<emphasis id="B970FB23A93D5E73E476F90AFDB9F97C" box="[361,590,1666,1690]" italics="true" pageId="1" pageNumber="370">Yarrhpelopia norrisi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2434658A93D5E73E74AF909FD7AF97F" box="[597,653,1665,1690]" pageId="1" pageNumber="370" rank="species">
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<emphasis id="B970FB23A93D5E73E74AF909FD7AF97F" bold="true" box="[597,653,1665,1690]" pageId="1" pageNumber="370">sp.n.</emphasis>
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</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31E74BAA93D5E73E5D8F92DFC32F8E3" pageId="1" pageNumber="370" type="etymology">
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<paragraph id="8BBB2731A93D5E73E5D8F92DFC32F8E3" blockId="1.[151,1438,862,2015]" pageId="1" pageNumber="370">
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<emphasis id="B970FB23A93D5E73E5D8F92DFEB1F95B" bold="true" box="[199,326,1701,1726]" pageId="1" pageNumber="370">Etymology</emphasis>
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.
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<emphasis id="B970FB23A93D5E73E444F92EFE6EF95B" box="[347,409,1702,1726]" italics="true" pageId="1" pageNumber="370">Yarrh</emphasis>
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(pronounced with silent ‘h) derives from the southern
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<collectingRegion id="49C0E9D3A93D5E73E121F92FFAE2F95B" box="[1086,1301,1701,1726]" country="Australia" name="New South Wales" pageId="1" pageNumber="370">
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New South
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<collectingCountry id="F31367A1A93D5E73E1CEF92DFAE2F95B" box="[1233,1301,1701,1726]" name="United Kingdom" pageId="1" pageNumber="370">Wales</collectingCountry>
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</collectingRegion>
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Aboriginal Ngunnawal language meaning 'running water', recognising the ecology of the genus and its centre of distribution, and—
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<emphasis id="B970FB23A93D5E73E5C4F966FEC1F8E3" box="[219,310,1774,1798]" italics="true" pageId="1" pageNumber="370">pelopia,</emphasis>
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a common suffix in
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<taxonomicName id="4C045CB2A93D5E73E701F965FD46F8E3" box="[542,689,1773,1798]" class="Insecta" family="Chironomidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="370" phylum="Arthropoda" rank="subFamily" subFamily="Tanypodinae">Tanypodinae</taxonomicName>
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, the suppressed
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<emphasis id="B970FB23A93D5E73E678F966FC49F8E3" box="[871,958,1774,1798]" italics="true" pageId="1" pageNumber="370">Pelopia</emphasis>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31E74BAA93D5E73E5D8F899FC5EF8AB" pageId="1" pageNumber="370" type="distribution">
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<paragraph id="8BBB2731A93D5E73E5D8F899FC5EF8AB" blockId="1.[151,1438,862,2015]" pageId="1" pageNumber="370">
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<emphasis id="B970FB23A93D5E73E5D8F899FEAEF8CF" bold="true" box="[199,345,1809,1834]" pageId="1" pageNumber="370">Distribution</emphasis>
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. Australian endemic, probably monotypic, distributed along the eastern margin of
|
||
<collectingCountry id="F31367A1A93D5E73E1EEF899FAACF8CF" box="[1265,1371,1809,1834]" name="Australia" pageId="1" pageNumber="370">Australia</collectingCountry>
|
||
, from 30°S to 42°S. Apparently absent from northern and
|
||
<collectingRegion id="49C0E9D3A93D5E73E7C3F8BFFC53F8AB" box="[732,932,1845,1870]" country="Australia" name="Western Australia" pageId="1" pageNumber="370">
|
||
western
|
||
<collectingCountry id="F31367A1A93D5E73E625F8BDFC53F8AB" box="[826,932,1845,1870]" name="Australia" pageId="1" pageNumber="370">Australia</collectingCountry>
|
||
</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31E74BAA93D5E70E5D8F8D1FB6FFBD8" lastPageId="2" lastPageNumber="371" pageId="1" pageNumber="370" type="discussion">
|
||
<paragraph id="8BBB2731A93D5E70E5D8F8D1FD53FE81" blockId="1.[151,1438,862,2015]" lastBlockId="2.[151,1437,151,1085]" lastPageId="2" lastPageNumber="371" pageId="1" pageNumber="370">
|
||
<emphasis id="B970FB23A93D5E73E5D8F8D1FEC3F897" bold="true" box="[199,308,1881,1906]" pageId="1" pageNumber="370">Remarks</emphasis>
|
||
. All stages of
|
||
<taxonomicName id="4C045CB2A93D5E73E4F4F8D2FD8CF897" box="[491,635,1882,1906]" class="Insecta" family="Chironomidae" genus="Yarrhpelopia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B970FB23A93D5E73E4F4F8D2FD8CF897" box="[491,635,1882,1906]" italics="true" pageId="1" pageNumber="370">Yarrhpelopia</emphasis>
|
||
</taxonomicName>
|
||
conform to diagnoses of the tribe
|
||
<taxonomicName id="4C045CB2A93D5E73E13DF8D1FB45F897" box="[1058,1202,1881,1906]" class="Insecta" family="Chironomidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="370" phylum="Arthropoda" rank="tribe" tribe="Pentaneurini">Pentaneurini</taxonomicName>
|
||
, but each differs in generic identity according to respective stage keyed. In the key to larval
|
||
<taxonomicName id="4C045CB2A93D5E73E6E9F8F5FB71F873" box="[1014,1158,1917,1942]" class="Insecta" family="Chironomidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="370" phylum="Arthropoda" rank="subFamily" subFamily="Tanypodinae">Tanypodinae</taxonomicName>
|
||
of the Holarctic region (
|
||
<bibRefCitation id="EF955AC0A93D5E73E581F82AFE49F85F" author="Cranston" box="[158,446,1953,1978]" pageId="1" pageNumber="370" refString="Cranston, P. S. & Epler, J. (2013) 5. The larvae of Tanypodinae (Diptera: Chironomidae) of the Holarctic region. Keys and diagnoses. In: Andersen, T., Cranston, P. S. & Epler, J. H. (Eds.), Chironomidae of the Holarctic Region: Keys and diagnoses. Part 1. Larvae. Insect Systematics and Evolution Supplements, 66, pp. 39 - 136." type="journal article" year="2013">Cranston & Epler, 2013</bibRefCitation>
|
||
) the simple palpal base, slightly recessed central ligula tooth, body lacking fringe, antennal peg sensilla absent, strong molar tooth, conventionally short posterior parapods, and smooth head and antennal length 3.5× mandible and medially located ring organ leads to couplet 33. The presence of a dorsal pit and simple small posterior parapod claws conform to
|
||
<taxonomicName id="4C045CB2A93E5E70E7DDFF35FCFCFF31" box="[706,779,189,212]" class="Insecta" family="Chironomidae" genus="Larsia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B970FB23A93E5E70E7DDFF35FCFCFF31" box="[706,779,189,212]" italics="true" pageId="2" pageNumber="371">Larsia</emphasis>
|
||
</taxonomicName>
|
||
whereas the ligula shape and A.R. of less than 3.5 suggest
|
||
<taxonomicName id="4C045CB2A93E5E70E588FF68FEEAFF1D" box="[151,285,224,248]" class="Insecta" family="Chironomidae" genus="Zavrelimyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B970FB23A93E5E70E588FF68FEEAFF1D" box="[151,285,224,248]" italics="true" pageId="2" pageNumber="371">Zavrelimyia</emphasis>
|
||
</taxonomicName>
|
||
. In
|
||
<bibRefCitation id="EF955AC0A93E5E70E451FF68FDFDFF1D" author="Cranston" box="[334,522,223,248]" pageId="2" pageNumber="371" refString="Cranston, P. S. (1996) Identification Guide to the Chironomidae of New South Wales. AWT Identification Guide Number 1. Australian Water Technologies Pty Ltd., West Ryde, NSW," type="book" year="1996">Cranston (1996)</bibRefCitation>
|
||
, the larva is keyed as 'genus A' as distinct from other Australian genera on the basis of the distribution of cephalic setae and pits, and on the sculpturing ('creasing') of the postmentum and gula. However Cranston's (loc. cit.) interpretation of the inner teeth of the ligula as directed anteriorly is erroneous as unworn teeth have an outwardly directed apex.
|
||
</paragraph>
|
||
<paragraph id="8BBB2731A93E5E70E5D8FEE7FBABFD42" blockId="2.[151,1437,151,1085]" pageId="2" pageNumber="371">
|
||
Regarding the pupa, in the key of
|
||
<bibRefCitation id="EF955AC0A93E5E70E76BFEE7FC56FE6D" author="Fittkau" box="[628,929,367,392]" pageId="2" pageNumber="371" refString="Fittkau E. J. & Murray, D. A. (1986) The pupae of Tanypodinae (Diptera: Chironomidae) of the Holarctic region. Keys and diagnoses. Entomologica scandinavica Supplement, 28, 31 - 113." type="journal article" year="1986">Fittkau & Murray (1986)</bibRefCitation>
|
||
the unfringed, narrow anal lobe indicates
|
||
<taxonomicName id="4C045CB2A93E5E70E588FE1CFED0FE48" box="[151,295,404,429]" class="Insecta" family="Chironomidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="tribe" tribe="Pentaneurini">Pentaneurini</taxonomicName>
|
||
, within which the dense simple spines of the abdominal shagreen preclude the Thienemannimyia group of genera. The presence of 4 LS on segment VII and adhesive sheaths on anal macrosetae, distinct thoracic comb, corona on thoracic horn and male genital sacs extending to 70% of length of the anal lobe that lacks spines on the inner margin leads to couplet 39.
|
||
<taxonomicName id="4C045CB2A93E5E70E749FD88FD11FDFD" box="[598,742,512,536]" class="Insecta" family="Chironomidae" genus="Yarrhpelopia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="370" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B970FB23A93E5E70E749FD88FD11FDFD" box="[598,742,512,536]" italics="true" pageId="2" pageNumber="371">Yarrhpelopia</emphasis>
|
||
</taxonomicName>
|
||
has both transverse serial spinule (shagreen) groups of
|
||
<taxonomicName id="4C045CB2A93E5E70E04CFD88FA6BFDF2" box="[1363,1436,512,535]" class="Insecta" family="Chironomidae" genus="Larsia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B970FB23A93E5E70E04CFD88FA6BFDF2" box="[1363,1436,512,535]" italics="true" pageId="2" pageNumber="371">Larsia</emphasis>
|
||
</taxonomicName>
|
||
and simple spinules of
|
||
<taxonomicName id="4C045CB2A93E5E70E480FDACFD3DFDD9" box="[415,714,548,572]" class="Insecta" family="Chironomidae" genus="Zavrelimyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B970FB23A93E5E70E480FDACFDD2FDD9" box="[415,549,548,572]" italics="true" pageId="2" pageNumber="371">Zavrelimyia</emphasis>
|
||
(
|
||
<emphasis id="B970FB23A93E5E70E728FDACFD4AFDD9" box="[567,701,548,572]" italics="true" pageId="2" pageNumber="371">Zavrelimyia</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
: the thoracic horn lacks the characteristically lobate respiratory atrium of
|
||
<taxonomicName id="4C045CB2A93E5E70E41BFDC0FEBAFDBA" box="[260,333,584,607]" class="Insecta" family="Chironomidae" genus="Larsia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B970FB23A93E5E70E41BFDC0FEBAFDBA" box="[260,333,584,607]" italics="true" pageId="2" pageNumber="371">Larsia</emphasis>
|
||
</taxonomicName>
|
||
. Keyed as 'genus A' (
|
||
<bibRefCitation id="EF955AC0A93E5E70E723FDC0FD1EFD85" author="Cranston" box="[572,745,583,608]" pageId="2" pageNumber="371" refString="Cranston, P. S. (1996) Identification Guide to the Chironomidae of New South Wales. AWT Identification Guide Number 1. Australian Water Technologies Pty Ltd., West Ryde, NSW," type="book" year="1996">Cranston, 1996</bibRefCitation>
|
||
) it is close to
|
||
<emphasis id="B970FB23A93E5E70E69CFDC0FBEEFDBA" box="[899,1049,584,607]" italics="true" pageId="2" pageNumber="371">
|
||
<taxonomicName id="4C045CB2A93E5E70E69CFDC0FBFDFDBA" box="[899,1034,584,607]" class="Insecta" family="Chironomidae" genus="Paramerina" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="genus">Paramerina</taxonomicName>
|
||
(
|
||
</emphasis>
|
||
now
|
||
<taxonomicName id="4C045CB2A93E5E70E14FFDC0FB21FD85" box="[1104,1238,584,608]" class="Insecta" family="Chironomidae" genus="Zavrelimyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B970FB23A93E5E70E14FFDC0FB21FD85" box="[1104,1238,584,608]" italics="true" pageId="2" pageNumber="371">Zavrelimyia</emphasis>
|
||
</taxonomicName>
|
||
) based largely on the thoracic horn (tubular, non-lobate atrium, corona present, plastron plate apical with coarse surface), but differs from
|
||
<taxonomicName id="4C045CB2A93E5E70E5CBFD18FE0CFD4D" authority="(Paramerina)" baseAuthorityName="Paramerina" box="[212,507,656,680]" class="Insecta" family="Chironomidae" genus="Zavrelimyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B970FB23A93E5E70E5CBFD18FEADFD4D" box="[212,346,656,680]" italics="true" pageId="2" pageNumber="371">Zavrelimyia</emphasis>
|
||
(
|
||
<emphasis id="B970FB23A93E5E70E474FD18FE05FD42" box="[363,498,656,679]" italics="true" pageId="2" pageNumber="371">Paramerina</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
in the horn being spinose distally as far as the corona.
|
||
</paragraph>
|
||
<paragraph id="8BBB2731A93E5E70E5D8FD3DFD52FC11" blockId="2.[151,1437,151,1085]" pageId="2" pageNumber="371">
|
||
In the adult male if the setose lobe on the basal median gonocoxite is considered to be a volsella (in the sense of the volsella of the Thienemannimyia group in
|
||
<bibRefCitation id="EF955AC0A93E5E70E7DFFD50FC27FD15" author="Murray" box="[704,976,727,752]" pageId="2" pageNumber="371" refString="Murray, D. A. & Fittkau, E. J. (1989) The adult males of Tanypodinae (Diptera: Chironomidae) of the Holarctic region. Keys and diagnoses. In: Wiederholm, T. (Ed.), Chironomidae of the Holarctic region. Keys and diagnoses. Part 3. Adult males. Entomologica Scandinavica Supplement, 34, pp. 37 - 123." type="journal article" year="1989">Murray & Fittkau, 1989</bibRefCitation>
|
||
) it would be the least developed volsella in the group. Treating
|
||
<taxonomicName id="4C045CB2A93E5E70E489FD74FDDEFCF1" box="[406,553,764,788]" class="Insecta" family="Chironomidae" genus="Yarrhpelopia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="370" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B970FB23A93E5E70E489FD74FDDEFCF1" box="[406,553,764,788]" italics="true" pageId="2" pageNumber="371">Yarrhpelopia</emphasis>
|
||
</taxonomicName>
|
||
as excluded from the Thienemannimyia group, the tapering gonostylus, plain wing, bare eye, costa ending clearly before M1+2, absence of scutal tubercle and paired spurs present with outer half length of inner, leads to a cluster of genera in which wing venation is critical in further keying. The anterior wing venation is difficult to interpret due to weakly-indicated veins and dense setosity. However R1 and R4+5 are approximated especially in the male, and with R2+3 compacted between them and with R2 faintly detectable. Although these keys do not strongly follow phylogeny, and each stage differs in morphological proximity, adjacency to
|
||
<taxonomicName id="4C045CB2A93E5E70E434FC54FDA6FC10" authority="(Paramerina)" baseAuthorityName="Paramerina" box="[299,593,988,1013]" class="Insecta" family="Chironomidae" genus="Zavrelimyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B970FB23A93E5E70E434FC54FE46FC11" box="[299,433,988,1012]" italics="true" pageId="2" pageNumber="371">Zavrelimyia</emphasis>
|
||
(
|
||
<emphasis id="B970FB23A93E5E70E4DEFC55FDBFFC11" box="[449,584,989,1012]" italics="true" pageId="2" pageNumber="371">Paramerina</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
recurs.
|
||
</paragraph>
|
||
<paragraph id="8BBB2731A93E5E70E5D8FB88FB6FFBD8" blockId="2.[151,1437,151,1085]" pageId="2" pageNumber="371">
|
||
The following description includes features of taxonomic significance at generic level. Until further species are collected, the description of
|
||
<taxonomicName id="4C045CB2A93E5E70E4CAFBACFD4DFBD9" box="[469,698,1060,1084]" class="Insecta" family="Chironomidae" genus="Yarrhpelopia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="norrisi" status="sp. nov.">
|
||
<emphasis id="B970FB23A93E5E70E4CAFBACFD4DFBD9" box="[469,698,1060,1084]" italics="true" pageId="2" pageNumber="371">Yarrhpelopia norrisi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2434658A93E5E70E7DEFBADFD01FBD9" box="[705,758,1061,1084]" pageId="2" pageNumber="371" rank="species">sp.n.</taxonomicNameLabel>
|
||
summarises features of a new genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |