treatments-xml/data/03/AA/F7/03AAF74DFF97996C1DF5FA6E61C8FEC7.xml
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<document id="DB0CF969593C64C5C1D7B20CCBEEBF8D" ID-CLB-Dataset="55502" ID-DOI="10.11646/zootaxa.5138.2.4" ID-GBIF-Dataset="48c6ef6a-bd20-4f98-8d6b-7a1b79ab2937" ID-ISSN="1175-5326" ID-Zenodo-Dep="6557093" ID-ZooBank="0B0E6A73-9FC2-44BD-9F30-57AD9B9FE4F0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1652772555109" checkinUser="plazi" docAuthor="Leavengood, John M., Pinkerton, Morgan G. &amp; Rifkind, Jacques" docDate="2022" docId="03AAF74DFF97996C1DF5FA6E61C8FEC7" docLanguage="en" docName="zootaxa.5138.2.4.pdf" docOrigin="Zootaxa 5138 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tarsobaenus letourneauae , Leavengood, Pinkerton and Rifkind 2022, new species" docType="treatment" docUuid="D8B90C8E-903E-4A3C-BEBC-F29E99683E06" docUuidSource="ZooBank" docVersion="7" lastPageNumber="172" masterDocId="FF938F35FF9599691D62FFC86558FFA2" masterDocTitle="Description of the new genus Tarsobaenus and three new species from Costa Rica (Coleoptera: Cleridae: Clerinae: Hydnocerini)" masterLastPageNumber="176" masterPageNumber="167" pageNumber="169" updateTime="1699337758796" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="1834FB420220EF72CD756BB299330842">Description of the new genus Tarsobaenus and three new species from Costa Rica (Coleoptera: Cleridae: Clerinae: Hydnocerini)</mods:title>
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<mods:namePart id="88D59D30139436C2464CD1E35B07197C">Leavengood, John M.</mods:namePart>
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<mods:affiliation id="F30CE07FC3185F2E1F6AF9BDF3B364F9">United States Department of Agriculture, APHIS, PPQ, 9325 Bay Plaza Blvd, Suite 206, Tampa, FL 33619 USA John. M. Leavengood @ usda. gov; https: // orcid. org / 0000 - 0003 - 3223 - 455 X</mods:affiliation>
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<mods:namePart id="6A7C948A7A1DE2B2738CCB9E226C4025">Rifkind, Jacques</mods:namePart>
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<taxonomicName id="4C033DD8FF97996B1DF5FA6E66FBFA62" authority="Leavengood, Pinkerton and Rifkind" authorityName=", Leavengood, Pinkerton and Rifkind" authorityYear="2022" box="[151,931,1446,1473]" class="Insecta" family="Cleridae" genus="Tarsobaenus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="169" phylum="Arthropoda" rank="species" species="letourneauae" status="sp. nov.">
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Leavengood, Pinkerton and Rifkind
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,
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(
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)
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: Male.
<collectingCountry id="F31406CBFF97996B1C3BF9FF6758F9F3" box="[345,512,1591,1617]" name="Costa Rica" pageId="2" pageNumber="169">COSTA RICA</collectingCountry>
:
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:
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,
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N de Cariari
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,
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,
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, L N 286000_567500 #2917 (
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)
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.
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:
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: Estac. Pitilla,
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S Santa Cecilia
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,
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&amp;
<collectorName id="26F6238DFF97996B19D6F9B7605BF93B" box="[1204,1283,1663,1689]" pageId="2" pageNumber="169">P. Rios</collectorName>
, L-N 330200, 380200 (
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, 1 f),
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Est. Pitilla,
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S Santa Cecilia
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,
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&amp;
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, L-N 330200, 380200 (
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,
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), Estac. Pitilla,
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S Santa Cecilia
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,
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trap, 1988, GNP Biod. Survey, L-N 330200, 380200 (
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, 1 f;
<collectionCode id="ED12DE9EFF97996B1C1EF923649DF8A7" box="[380,453,1771,1797]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="2" pageNumber="169">INBio</collectionCode>
, 2 f),
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Estac. Pitilla,
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,
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S Santa Cecilia
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, P. N.
<collectingRegion id="49C788B9FF97996B1978F92361C6F8A7" box="[1050,1182,1771,1797]" country="Costa Rica" name="Guanacaste" pageId="2" pageNumber="169">Guanacaste</collectingRegion>
,
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<collectorName id="26F6238DFF97996B18E0F92365A9F88B" pageId="2" pageNumber="169">C. Moraga</collectorName>
, L-N 330200, 380200 (
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, 1 f).
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<materialsCitation id="3B6B4C06FF97996B1FE3F8C7615EF8EC" ID-GBIF-Occurrence="3771643311" accessionNumber="N 84003" collectingDate="1986-02-29" collectingDateMax="2010-08-25" collectingDateMin="1986-02-29" collectionCode="INBio" country="Costa Rica" county="Malaise" elevation="700" location="La" municipality="La" pageId="2" pageNumber="169" specimenCount="1" stateProvince="Guanacaste" typeStatus="paratype">
<collectingRegion id="49C788B9FF97996B1FE3F8C76611F88B" box="[641,841,1807,1833]" country="Costa Rica" name="Heredia" pageId="2" pageNumber="169">Heredia Province</collectingRegion>
:
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<quantity id="4CFBEBBEFF97996B1E31F8C766CAF88B" box="[851,914,1807,1833]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="169" unit="km" value="11.0">11km</quantity>
ESE La Virgen
</locationDeviation>
,
<quantity id="4CFBEBBEFF97996B192CF8C6619CF88B" box="[1102,1220,1806,1833]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="2" pageNumber="169" unit="m" value="300.0" valueMax="350.0" valueMin="250.0">
<elevation id="002EA168FF97996B192CF8C6619CF88B" box="[1102,1220,1806,1833]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="2" pageNumber="169" unit="m" value="300.0" valueMax="350.0" valueMin="250.0">250-350m</elevation>
</quantity>
,
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N
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W,
<date id="FFBD609BFF97996B1DF5F8FB6443F8EF" box="[151,283,1843,1869]" pageId="2" pageNumber="169" value="2004-04-18">
<collectingDate id="EFF99973FF97996B1DF5F8FB6443F8EF" box="[151,283,1843,1869]" pageId="2" pageNumber="169" value="2004-04-18">18-IV-2004</collectingDate>
</date>
, INBio-OET-ALAS transect,
<collectorName id="26F6238DFF97996B1F12F8FB6793F8EF" box="[624,715,1843,1869]" pageId="2" pageNumber="169">Malaise</collectorName>
, 03/M/07/087 (
<collectionCode id="ED12DE9EFF97996B1E1FF8FB669FF8EF" box="[893,967,1843,1869]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="2" pageNumber="169">INBio</collectionCode>
, 1 f),
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<materialsCitation id="3B6B4C06FF97996B196DF8FB6184F8D0" ID-GBIF-Occurrence="3771643301" accessionNumber="N 84003" collectingDate="1986-02-29" collectingDateMax="2010-08-25" collectingDateMin="1986-02-29" collectionCode="JMLC" collectorName="Gerardo Vega" country="Costa Rica" county="Malaise" elevation="700" location="La" municipality="La" pageId="2" pageNumber="169" specimenCount="1" stateProvince="Guanacaste" typeStatus="paratype">
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<collectingMunicipality id="6BD8DC21FF97996B196DF8FB6175F8EF" box="[1039,1069,1843,1869]" pageId="2" pageNumber="169">La</collectingMunicipality>
Selva
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,
<date id="FFBD609BFF97996B19E7F8FB6044F8EF" box="[1157,1308,1843,1869]" pageId="2" pageNumber="169" value="1998-01-18" valueMax="1998-01-21" valueMin="1998-01-18">
<collectingDate id="EFF99973FF97996B19E7F8FB6044F8EF" box="[1157,1308,1843,1869]" pageId="2" pageNumber="169" value="1998-01-18" valueMax="1998-01-21" valueMin="1998-01-18">18-21-I-1998</collectingDate>
</date>
,
<collectorName id="26F6238DFF97996B1848F8FB60C4F8EF" box="[1322,1436,1843,1869]" pageId="2" pageNumber="169">F. Hovore</collectorName>
(
<collectionCode id="ED12DE9EFF97996B1DFDF89F65B0F8D3" box="[159,232,1879,1905]" pageId="2" pageNumber="169">JNRC</collectionCode>
, 1 f),
<collectorName id="26F6238DFF97996B1C57F89F6480F8D3" box="[309,472,1879,1905]" pageId="2" pageNumber="169">
<location id="8EDC1080FF97996B1C57F89F640BF8D3" LSID="urn:lsid:plazi:treatment:03AAF74DFF97996C1DF5FA6E61C8FEC7:8EDC1080FF97996B1C57F89F640BF8D3" box="[309,339,1879,1905]" country="Costa Rica" county="Malaise" municipality="La" name="La" pageId="2" pageNumber="169" stateProvince="Guanacaste">La</location>
Selva B.S.
</collectorName>
,
<quantity id="4CFBEBBEFF97996B1C8AF89E6708F8D3" box="[488,592,1878,1905]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="2" pageNumber="169" unit="m" value="100.0" valueMax="150.0" valueMin="50.0">
<elevation id="002EA168FF97996B1C8AF89E6708F8D3" box="[488,592,1878,1905]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="2" pageNumber="169" unit="m" value="100.0" valueMax="150.0" valueMin="50.0">50-150m</elevation>
</quantity>
,
<collectorName id="26F6238DFF97996B1F02F89F679AF8D3" box="[608,706,1879,1905]" pageId="2" pageNumber="169">Locality</collectorName>
: STR,
<date id="FFBD609BFF97996B1E77F89F66C9F8D3" box="[789,913,1879,1905]" pageId="2" pageNumber="169" value="1986-02-29">
<collectingDate id="EFF99973FF97996B1E77F89F66C9F8D3" box="[789,913,1879,1905]" pageId="2" pageNumber="169" value="1986-02-29">29-II-1986</collectingDate>
</date>
,
<collectorName id="26F6238DFF97996B1EC2F89F6166F8D3" box="[928,1086,1879,1905]" pageId="2" pageNumber="169">Gerardo Vega</collectorName>
(
<collectionCode id="ED12DE9EFF97996B1933F89F61C2F8D3" box="[1105,1178,1879,1905]" pageId="2" pageNumber="169">JNRC</collectionCode>
, 1 f),
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<materialsCitation id="3B6B4C06FF97996B1985F89F61BAF834" ID-GBIF-Occurrence="3771643313" accessionNumber="N 84003" collectingDate="1986-02-29" collectingDateMax="2010-08-25" collectingDateMin="1986-02-29" collectionCode="JMLC" collectorName="J. M. Leavengood, Jr." country="Costa Rica" county="Malaise" elevation="700" location="La" municipality="La" pageId="2" pageNumber="169" specimenCount="1" stateProvince="Guanacaste" typeStatus="paratype">
<location id="8EDC1080FF97996B1985F89F645DF837" LSID="urn:lsid:plazi:treatment:03AAF74DFF97996C1DF5FA6E61C8FEC7:8EDC1080FF97996B1985F89F645DF837" country="Costa Rica" county="Malaise" municipality="La" name="Puerto Viejo de Sarapiqui" pageId="2" pageNumber="169" stateProvince="Guanacaste">Puerto Viejo de Sarapiqui</location>
,
<collectorName id="26F6238DFF97996B1C70F8B367C3F837" box="[274,667,1915,1941]" pageId="2" pageNumber="169">
<location id="8EDC1080FF97996B1C70F8B36468F837" LSID="urn:lsid:plazi:treatment:03AAF74DFF97996C1DF5FA6E61C8FEC7:8EDC1080FF97996B1C70F8B36468F837" box="[274,304,1915,1941]" country="Costa Rica" county="Malaise" municipality="La" name="La" pageId="2" pageNumber="169" stateProvince="Guanacaste">La</location>
Selva Biological Research Sta.
</collectorName>
,
<date id="FFBD609BFF97996B1FCAF8B2661AF837" box="[680,834,1914,1941]" pageId="2" pageNumber="169" value="2010-08-25">
<collectingDate id="EFF99973FF97996B1FCAF8B2661AF837" box="[680,834,1914,1941]" pageId="2" pageNumber="169" value="2010-08-25">25-VIII-2010</collectingDate>
</date>
,
<collectorName id="26F6238DFF97996B1E2DF8B36178F837" box="[847,1056,1915,1941]" pageId="2" pageNumber="169">J. M. Leavengood</collectorName>
, Jr. (
<collectionCode id="ED12DE9EFF97996B193AF8B361FCF837" box="[1112,1188,1915,1941]" pageId="2" pageNumber="169">JMLC</collectionCode>
, 1 f),
</materialsCitation>
<materialsCitation id="3B6B4C06FF97996B1989F8B36180F81B" ID-GBIF-Occurrence="3771643305" accessionNumber="N 84003" collectingDate="1986-02-29" collectingDateMax="2010-08-25" collectingDateMin="1986-02-29" collectionCode="JMLC" collectorName="J. M. Leavengood, Jr." country="Costa Rica" county="Malaise" elevation="700" location="La" municipality="La" pageId="2" pageNumber="169" specimenCount="1" stateProvince="Guanacaste" typeStatus="paratype">
<location id="8EDC1080FF97996B1989F8B3645DF81B" LSID="urn:lsid:plazi:treatment:03AAF74DFF97996C1DF5FA6E61C8FEC7:8EDC1080FF97996B1989F8B3645DF81B" country="Costa Rica" county="Malaise" municipality="La" name="Puerto Viejo de Sarapiqui" pageId="2" pageNumber="169" stateProvince="Guanacaste">Puerto Viejo de Sarapiqui</location>
,
<collectorName id="26F6238DFF97996B1C73F85767CAF81B" box="[273,658,1951,1977]" pageId="2" pageNumber="169">
<location id="8EDC1080FF97996B1C73F8576477F81B" LSID="urn:lsid:plazi:treatment:03AAF74DFF97996C1DF5FA6E61C8FEC7:8EDC1080FF97996B1C73F8576477F81B" box="[273,303,1951,1977]" country="Costa Rica" county="Malaise" municipality="La" name="La" pageId="2" pageNumber="169" stateProvince="Guanacaste">La</location>
Selva Biological Research Sta.
</collectorName>
,
<date id="FFBD609BFF97996B1FFCF857666FF81B" box="[670,823,1951,1977]" pageId="2" pageNumber="169" value="2010-08-13">
<collectingDate id="EFF99973FF97996B1FFCF857666FF81B" box="[670,823,1951,1977]" pageId="2" pageNumber="169" value="2010-08-13">13-VIII-2010</collectingDate>
</date>
,
<collectorName id="26F6238DFF97996B1E21F8576148F81B" box="[835,1040,1951,1977]" pageId="2" pageNumber="169">J. M. Leavengood</collectorName>
, Jr. (
<collectionCode id="ED12DE9EFF97996B1927F85761C9F81B" box="[1093,1169,1951,1977]" pageId="2" pageNumber="169">JMLC</collectionCode>
,
<quantity id="4CFBEBBEFF97996B19FEF8576194F81B" box="[1180,1228,1951,1977]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="169" unit="m" value="1.0">
<elevation id="002EA168FF97996B19FEF8576194F81B" box="[1180,1228,1951,1977]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="169" unit="m" value="1.0">1 m</elevation>
</quantity>
).
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<materialsCitation id="3B6B4C06FF97996A19BDF8576656FDC0" ID-GBIF-Occurrence="3771643304" accessionNumber="N 84003" collectingDate="1986-02-29" collectingDateMax="2010-08-25" collectingDateMin="1986-02-29" collectionCode="INBio" collectorName="E. Rojas" country="Costa Rica" county="Malaise" elevation="700" lastPageId="3" lastPageNumber="170" location="La" municipality="La" pageId="2" pageNumber="169" specimenCount="1" stateProvince="Guanacaste" typeStatus="paratype">
<collectingRegion id="49C788B9FF97996B19BDF85760C1F81B" box="[1247,1433,1951,1977]" country="Costa Rica" name="Limon" pageId="2" pageNumber="169">Limón Province</collectingRegion>
:
<collectorName id="26F6238DFF97996B1DF5F80B6424F87F" box="[151,380,1987,2013]" pageId="2" pageNumber="169">Sector Cerro Cocori</collectorName>
,
<collectorName id="26F6238DFF97996B1CE5F80B64EFF87F" box="[391,439,1987,2013]" pageId="2" pageNumber="169">Fca.</collectorName>
de
<collectorName id="26F6238DFF97996B1CBDF80B6719F87F" box="[479,577,1987,2013]" pageId="2" pageNumber="169">E. Rojas</collectorName>
,
<quantity id="4CFBEBBEFF97996B1F2EF80A67D7F87F" box="[588,655,1986,2013]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="169" unit="m" value="150.0">
<elevation id="002EA168FF97996B1F2EF80A67D7F87F" box="[588,655,1986,2013]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="169" unit="m" value="150.0">150m</elevation>
</quantity>
,
<date id="FFBD609BFF97996B1FF8F80B6601F87F" box="[666,857,1987,2013]" pageId="2" pageNumber="169" value="1992-08-12" valueMax="1992-08-31" valueMin="1992-08-12">
<collectingDate id="EFF99973FF97996B1FF8F80B6601F87F" box="[666,857,1987,2013]" pageId="2" pageNumber="169" value="1992-08-12" valueMax="1992-08-31" valueMin="1992-08-12">12-31-VIII-1992</collectingDate>
</date>
,
<collectorName id="26F6238DFF97996B1E06F80B669FF87F" box="[868,967,1987,2013]" pageId="2" pageNumber="169">E. Rojas</collectorName>
, L-N 286000, 567500 (
<collectionCode id="ED12DE9EFF97996B19AEF80B604EF87F" box="[1228,1302,1987,2013]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="2" pageNumber="169" type="Museum">FSCA</collectionCode>
, 1 f;
<collectionCode id="ED12DE9EFF97996B182DF80B60C0F87F" box="[1359,1432,1987,2013]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="2" pageNumber="169">INBio</collectionCode>
, 1 f),
<collectorName id="26F6238DFF96996A1DACFF5F64EBFF13" box="[206,435,151,177]" pageId="3" pageNumber="170">Sector Cerro Cocori</collectorName>
,
<collectorName id="26F6238DFF96996A1CDCFF5F64B6FF13" box="[446,494,151,177]" pageId="3" pageNumber="170">Fca.</collectorName>
de
<collectorName id="26F6238DFF96996A1F75FF5F6721FF13" box="[535,633,151,177]" pageId="3" pageNumber="170">E. Rojas</collectorName>
,
<quantity id="4CFBEBBEFF96996A1FE6FF5E679FFF13" box="[644,711,150,177]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="170" unit="m" value="150.0">
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</quantity>
, [??]-I-29-II-1992,
<collectorName id="26F6238DFF96996A1EC0FF5F615CFF13" box="[930,1028,151,177]" pageId="3" pageNumber="170">E. Rojas</collectorName>
, L-N 286000, 567500 (
<collectionCode id="ED12DE9EFF96996A1869FF5F600DFF13" box="[1291,1365,151,177]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="3" pageNumber="170">INBio</collectionCode>
,
<quantity id="4CFBEBBEFF96996A1803FF5F60D7FF13" box="[1377,1423,151,177]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="170" unit="m" value="1.0">
<elevation id="002EA168FF96996A1803FF5F60D7FF13" box="[1377,1423,151,177]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="170" unit="m" value="1.0">1 m</elevation>
</quantity>
),
<collectorName id="26F6238DFF96996A1DF5FF7364D9FF77" box="[151,385,187,213]" pageId="3" pageNumber="170">Sector Cerro Cocori</collectorName>
,
<collectorName id="26F6238DFF96996A1CEDFF7364E7FF77" box="[399,447,187,213]" pageId="3" pageNumber="170">Fca.</collectorName>
de
<collectorName id="26F6238DFF96996A1C8FFF736709FF77" box="[493,593,187,213]" pageId="3" pageNumber="170">E. Rojas</collectorName>
,
<quantity id="4CFBEBBEFF96996A1F3DFF7267FAFF77" box="[607,674,186,213]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="170" unit="m" value="150.0">
<elevation id="002EA168FF96996A1F3DFF7267FAFF77" box="[607,674,186,213]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="170" unit="m" value="150.0">150m</elevation>
</quantity>
,
<date id="FFBD609BFF96996A1FD2FF7366C8FF77" box="[688,912,187,213]" pageId="3" pageNumber="170" value="1992-07-26" valueMax="1992-08-02" valueMin="1992-07-26">
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</date>
,
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, L-N 286000, 567500 (
<collectionCode id="ED12DE9EFF96996A1876FF736005FF77" box="[1300,1373,187,213]" pageId="3" pageNumber="170">JNRC</collectionCode>
, 1 f),
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,
<collectorName id="26F6238DFF96996A1CE8FF1764E2FF5B" box="[394,442,223,249]" pageId="3" pageNumber="170">Fca.</collectorName>
de
<collectorName id="26F6238DFF96996A1C86FF176710FF5B" box="[484,584,223,249]" pageId="3" pageNumber="170">E. Rojas</collectorName>
,
<quantity id="4CFBEBBEFF96996A1F36FF1667CEFF5B" box="[596,662,222,249]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="170" unit="m" value="150.0">
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,
<date id="FFBD609BFF96996A1FC1FF17665CFF5B" box="[675,772,223,249]" pageId="3" pageNumber="170" value="1991-11">
<collectingDate id="EFF99973FF96996A1FC1FF17665CFF5B" box="[675,772,223,249]" pageId="3" pageNumber="170" value="1991-11">XI-1991</collectingDate>
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,
<collectorName id="26F6238DFF96996A1E73FF17662CFF5B" box="[785,884,223,249]" pageId="3" pageNumber="170">E. Rojas</collectorName>
, L-N 286000, 567500 (
<collectionCode id="ED12DE9EFF96996A191CFF176190FF5B" box="[1150,1224,223,249]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="3" pageNumber="170">INBio</collectionCode>
, 1 f),
<collectorName id="26F6238DFF96996A186FFF1765BEFEBF" pageId="3" pageNumber="170">Sector Cerro Cocori</collectorName>
,
<collectorName id="26F6238DFF96996A1D91FECB647BFEBF" box="[243,291,259,285]" pageId="3" pageNumber="170">Fca.</collectorName>
de
<collectorName id="26F6238DFF96996A1C2DFECB64EBFEBF" box="[335,435,259,285]" pageId="3" pageNumber="170">E. Rojas</collectorName>
,
<quantity id="4CFBEBBEFF96996A1CA2FECA675BFEBF" box="[448,515,258,285]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="170" unit="m" value="150.0">
<elevation id="002EA168FF96996A1CA2FECA675BFEBF" box="[448,515,258,285]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="170" unit="m" value="150.0">150m</elevation>
</quantity>
,
<date id="FFBD609BFF96996A1F72FECB6730FEBF" box="[528,616,259,285]" pageId="3" pageNumber="170" value="1991-10">
<collectingDate id="EFF99973FF96996A1F72FECB6730FEBF" box="[528,616,259,285]" pageId="3" pageNumber="170" value="1991-10">X-1991</collectingDate>
</date>
,
<collectorName id="26F6238DFF96996A1F17FECB6781FEBF" box="[629,729,259,285]" pageId="3" pageNumber="170">E. Rojas</collectorName>
, L-N 286000, 567500 (
<collectionCode id="ED12DE9EFF96996A1E84FECB6168FEBF" box="[998,1072,259,285]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="3" pageNumber="170">INBio</collectionCode>
, 1 f),
<collectorName id="26F6238DFF96996A1915FECB6038FEBF" box="[1143,1376,259,285]" pageId="3" pageNumber="170">Sector Cerro Cocori</collectorName>
,
<collectorName id="26F6238DFF96996A180FFECB60C5FEBF" box="[1389,1437,259,285]" pageId="3" pageNumber="170">Fca.</collectorName>
de
<collectorName id="26F6238DFF96996A1DDBFEEF6444FEE3" box="[185,284,295,321]" pageId="3" pageNumber="170">E. Rojas</collectorName>
,
<quantity id="4CFBEBBEFF96996A1C4AFEEE6433FEE3" box="[296,363,294,321]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="170" unit="m" value="150.0">
<elevation id="002EA168FF96996A1C4AFEEE6433FEE3" box="[296,363,294,321]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="170" unit="m" value="150.0">150m</elevation>
</quantity>
,
<date id="FFBD609BFF96996A1C15FEEF670BFEE3" box="[375,595,295,321]" pageId="3" pageNumber="170" value="1992-06-26" valueMax="1992-07-16" valueMin="1992-06-26">
<collectingDate id="EFF99973FF96996A1C15FEEF670BFEE3" box="[375,595,295,321]" pageId="3" pageNumber="170" value="1992-06-26" valueMax="1992-07-16" valueMin="1992-06-26">26-VI-16-VII-1992</collectingDate>
</date>
,
<collectorName id="26F6238DFF96996A1F3DFEEF679AFEE3" box="[607,706,295,321]" pageId="3" pageNumber="170">E. Rojas</collectorName>
, L-N 286000, 567500 (
<collectionCode id="ED12DE9EFF96996A1EA9FEEF614DFEE3" box="[971,1045,295,321]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="3" pageNumber="170">INBio</collectionCode>
, 1 f),
<collectorName id="26F6238DFF96996A193AFEEF6066FEE3" box="[1112,1342,295,321]" pageId="3" pageNumber="170">Sector Cerro Cocori</collectorName>
,
<collectorName id="26F6238DFF96996A1828FEEF6022FEE3" box="[1354,1402,295,321]" pageId="3" pageNumber="170">Fca.</collectorName>
de
<collectorName id="26F6238DFF96996A1DF5FE8365A3FEC7" box="[151,251,331,357]" pageId="3" pageNumber="170">E. Rojas</collectorName>
,
<quantity id="4CFBEBBEFF96996A1C65FE826412FEC7" box="[263,330,330,357]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="170" unit="m" value="150.0">
<elevation id="002EA168FF96996A1C65FE826412FEC7" box="[263,330,330,357]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="170" unit="m" value="150.0">150m</elevation>
</quantity>
,
<date id="FFBD609BFF96996A1C34FE8364FCFEC7" box="[342,420,331,357]" pageId="3" pageNumber="170" value="1992-01">
<collectingDate id="EFF99973FF96996A1C34FE8364FCFEC7" box="[342,420,331,357]" pageId="3" pageNumber="170" value="1992-01">I-1992</collectingDate>
</date>
,
<collectorName id="26F6238DFF96996A1CD2FE83674BFEC7" box="[432,531,331,357]" pageId="3" pageNumber="170">E. Rojas</collectorName>
, L-N 286000, 567500 (
<collectionCode id="ED12DE9EFF96996A1E7CFE836633FEC7" box="[798,875,331,357]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" lsid="urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="3" pageNumber="170" type="Museum">CSCA</collectionCode>
, 1 f;
<collectionCode id="ED12DE9EFF96996A1ECAFE8366A9FEC7" box="[936,1009,331,357]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="3" pageNumber="170">INBio</collectionCode>
, 1 f),
<collectorName id="26F6238DFF96996A1954FE836045FEC7" box="[1078,1309,331,357]" pageId="3" pageNumber="170">Sector Cerro Cocori</collectorName>
,
<collectorName id="26F6238DFF96996A184BFE836001FEC7" box="[1321,1369,331,357]" pageId="3" pageNumber="170">Fca.</collectorName>
de
<collectorName id="26F6238DFF96996A18E6FE836582FE2B" pageId="3" pageNumber="170">E. Rojas</collectorName>
,
<quantity id="4CFBEBBEFF96996A1D87FEA66470FE2B" box="[229,296,366,393]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="170" unit="m" value="150.0">
<elevation id="002EA168FF96996A1D87FEA66470FE2B" box="[229,296,366,393]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="170" unit="m" value="150.0">150m</elevation>
</quantity>
,
<date id="FFBD609BFF96996A1C51FEA764D8FE2B" box="[307,384,367,393]" pageId="3" pageNumber="170" value="1993-01">
<collectingDate id="EFF99973FF96996A1C51FEA764D8FE2B" box="[307,384,367,393]" pageId="3" pageNumber="170" value="1993-01">I-1993</collectingDate>
</date>
,
<collectorName id="26F6238DFF96996A1CEEFEA764B6FE2B" box="[396,494,367,393]" pageId="3" pageNumber="170">E. Rojas</collectorName>
, L-N 286000, 567500 (
<collectionCode id="ED12DE9EFF96996A1F96FEA76665FE2B" box="[756,829,367,393]" pageId="3" pageNumber="170">JNRC</collectionCode>
, 1 f),
<collectorName id="26F6238DFF96996A1E1CFEA7613AFE2B" box="[894,1122,367,393]" pageId="3" pageNumber="170">Sector Cerro Cocori</collectorName>
,
<collectorName id="26F6238DFF96996A190CFEA760C1FE2B" box="[1134,1433,367,393]" pageId="3" pageNumber="170">
<location id="8EDC1080FF96996A190CFEA7600FFE2B" LSID="urn:lsid:plazi:treatment:03AAF74DFF97996C1DF5FA6E61C8FEC7:8EDC1080FF96996A190CFEA7600FFE2B" box="[1134,1367,367,393]" country="Costa Rica" county="Malaise" municipality="La" name="Finca de Elias Rojas" pageId="3" pageNumber="170" stateProvince="Guanacaste">Finca de Elias Rojas</location>
, A. C
</collectorName>
. Tortuguero,
<quantity id="4CFBEBBEFF96996A1C43FE5A643BFE0F" box="[289,355,402,429]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="170" unit="m" value="150.0">
<elevation id="002EA168FF96996A1C43FE5A643BFE0F" box="[289,355,402,429]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="170" unit="m" value="150.0">150m</elevation>
</quantity>
,
<date id="FFBD609BFF96996A1C0FFE5B6497FE0F" box="[365,463,403,429]" pageId="3" pageNumber="170" value="1993-06">
<collectingDate id="EFF99973FF96996A1C0FFE5B6497FE0F" box="[365,463,403,429]" pageId="3" pageNumber="170" value="1993-06">VI-1993</collectingDate>
</date>
,
<collectorName id="26F6238DFF96996A1CB8FE5B6763FE0F" box="[474,571,403,429]" pageId="3" pageNumber="170">E. Rojas</collectorName>
, L N 286000, 567500 #2917 (
<collectionCode id="ED12DE9EFF96996A1EE5FE5B6689FE0F" box="[903,977,403,429]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="3" pageNumber="170">INBio</collectionCode>
, 1 f),
<collectorName id="26F6238DFF96996A1973FE5B61ACFE0F" box="[1041,1268,403,429]" pageId="3" pageNumber="170">Sector Cerro Cocori</collectorName>
,
<location id="8EDC1080FF96996A199DFE5B6582FE73" LSID="urn:lsid:plazi:treatment:03AAF74DFF97996C1DF5FA6E61C8FEC7:8EDC1080FF96996A199DFE5B6582FE73" country="Costa Rica" county="Malaise" municipality="La" name="Finca de Elias Rojas" pageId="3" pageNumber="170" stateProvince="Guanacaste">Finca de Elias Rojas</location>
,
<quantity id="4CFBEBBEFF96996A1D84FE7E6470FE73" box="[230,296,438,465]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="170" unit="m" value="150.0">
<elevation id="002EA168FF96996A1D84FE7E6470FE73" box="[230,296,438,465]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="170" unit="m" value="150.0">150m</elevation>
</quantity>
,
<date id="FFBD609BFF96996A1C56FE7F64CEFE73" box="[308,406,439,465]" pageId="3" pageNumber="170" value="1993-09">
<collectingDate id="EFF99973FF96996A1C56FE7F64CEFE73" box="[308,406,439,465]" pageId="3" pageNumber="170" value="1993-09">IX-1993</collectingDate>
</date>
,
<collectorName id="26F6238DFF96996A1CC0FE7F675CFE73" box="[418,516,439,465]" pageId="3" pageNumber="170">E. Rojas</collectorName>
, L N 286000, 567500 #2347 (
<collectionCode id="ED12DE9EFF96996A1E35FE7F66F9FE73" box="[855,929,439,465]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="3" pageNumber="170" type="Museum">FSCA</collectionCode>
,
<quantity id="4CFBEBBEFF96996A1ECCFE7F6684FE73" box="[942,988,439,465]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="170" unit="m" value="1.0">
<elevation id="002EA168FF96996A1ECCFE7F6684FE73" box="[942,988,439,465]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="170" unit="m" value="1.0">1 m</elevation>
</quantity>
;
<collectionCode id="ED12DE9EFF96996A1E8BFE7F616AFE73" box="[1001,1074,439,465]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="3" pageNumber="170">INBio</collectionCode>
, 1 f),
<collectorName id="26F6238DFF96996A1916FE7F604AFE73" box="[1140,1298,439,465]" pageId="3" pageNumber="170">Sector Cocori</collectorName>
,
<locationDeviation id="9A0E5F79FF96996A187CFE7F65B0FE57" location="Cariari" pageId="3" pageNumber="170">
<quantity id="4CFBEBBEFF96996A187CFE7F6006FE73" box="[1310,1374,439,465]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="170" unit="km" value="30.0">30km</quantity>
N de Cariari
</locationDeviation>
,
<location id="8EDC1080FF96996A1D96FE136487FE57" LSID="urn:lsid:plazi:treatment:03AAF74DFF97996C1DF5FA6E61C8FEC7:8EDC1080FF96996A1D96FE136487FE57" box="[244,479,475,501]" country="Costa Rica" county="Malaise" municipality="La" name="Finca de Elias Rojas" pageId="3" pageNumber="170" stateProvince="Guanacaste">Finca de Elias Rojas</location>
,
<collectorName id="26F6238DFF96996A1C8BFE1367F4FE57" box="[489,684,475,501]" pageId="3" pageNumber="170">A. C. Tortuguero</collectorName>
,
<quantity id="4CFBEBBEFF96996A1FDAFE1267A2FE57" box="[696,762,474,501]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="170" unit="m" value="150.0">
<elevation id="002EA168FF96996A1FDAFE1267A2FE57" box="[696,762,474,501]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="170" unit="m" value="150.0">150m</elevation>
</quantity>
,
<date id="FFBD609BFF96996A1E64FE13660BFE57" box="[774,851,475,501]" pageId="3" pageNumber="170" value="1994-01">
<collectingDate id="EFF99973FF96996A1E64FE13660BFE57" box="[774,851,475,501]" pageId="3" pageNumber="170" value="1994-01">I-1994</collectingDate>
</date>
,
<collectorName id="26F6238DFF96996A1E02FE13669BFE57" box="[864,963,475,501]" pageId="3" pageNumber="170">E. Rojas</collectorName>
, L N 286000, 567500 #2549 (
<collectionCode id="ED12DE9EFF96996A1874FE136038FE57" box="[1302,1376,475,501]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="3" pageNumber="170">INBio</collectionCode>
, 1 f),
<collectorName id="26F6238DFF96996A1DF5FE37646DFDBB" box="[151,309,511,537]" pageId="3" pageNumber="170">Sector Cocori</collectorName>
,
<locationDeviation id="9A0E5F79FF96996A1C23FE37674FFDBB" box="[321,535,511,537]" location="Cariari" pageId="3" pageNumber="170">
<quantity id="4CFBEBBEFF96996A1C23FE3764D9FDBB" box="[321,385,511,537]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="170" unit="km" value="30.0">30km</quantity>
N de Cariari
</locationDeviation>
,
<location id="8EDC1080FF96996A1F40FE376655FDBB" LSID="urn:lsid:plazi:treatment:03AAF74DFF97996C1DF5FA6E61C8FEC7:8EDC1080FF96996A1F40FE376655FDBB" box="[546,781,511,537]" country="Costa Rica" county="Malaise" municipality="La" name="Finca de Elias Rojas" pageId="3" pageNumber="170" stateProvince="Guanacaste">Finca de Elias Rojas</location>
,
<collectorName id="26F6238DFF96996A1E75FE376681FDBB" box="[791,985,511,537]" pageId="3" pageNumber="170">A. C. Tortuguero</collectorName>
,
<quantity id="4CFBEBBEFF96996A1E87FE36617FFDBB" box="[997,1063,510,537]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="170" unit="m" value="150.0">
<elevation id="002EA168FF96996A1E87FE36617FFDBB" box="[997,1063,510,537]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="170" unit="m" value="150.0">150m</elevation>
</quantity>
,
<date id="FFBD609BFF96996A1950FE3761CCFDBB" box="[1074,1172,511,537]" pageId="3" pageNumber="170" value="1994-06">
<collectingDate id="EFF99973FF96996A1950FE3761CCFDBB" box="[1074,1172,511,537]" pageId="3" pageNumber="170" value="1994-06">VI-1994</collectingDate>
</date>
,
<collectorName id="26F6238DFF96996A19C2FE37605BFDBB" box="[1184,1283,511,537]" pageId="3" pageNumber="170">E. Rojas</collectorName>
, L N 286000, 567500 #2549 (
<collectionCode id="ED12DE9EFF96996A1C2BFDEB64CBFD9F" box="[329,403,547,573]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="3" pageNumber="170">INBio</collectionCode>
,
<quantity id="4CFBEBBEFF96996A1CFDFDEB6495FD9F" box="[415,461,547,573]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="170" unit="m" value="1.0">
<elevation id="002EA168FF96996A1CFDFDEB6495FD9F" box="[415,461,547,573]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="170" unit="m" value="1.0">1 m</elevation>
</quantity>
),
<collectorName id="26F6238DFF96996A1C80FDEB67D8FD9F" box="[482,640,547,573]" pageId="3" pageNumber="170">Sector Cocori</collectorName>
,
<locationDeviation id="9A0E5F79FF96996A1FEEFDEB663BFD9F" box="[652,867,547,573]" location="Cariari" pageId="3" pageNumber="170">
<quantity id="4CFBEBBEFF96996A1FEEFDEB6794FD9F" box="[652,716,547,573]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="170" unit="km" value="30.0">30km</quantity>
N de Cariari
</locationDeviation>
,
<location id="8EDC1080FF96996A1E0CFDEB6101FD9F" LSID="urn:lsid:plazi:treatment:03AAF74DFF97996C1DF5FA6E61C8FEC7:8EDC1080FF96996A1E0CFDEB6101FD9F" box="[878,1113,547,573]" country="Costa Rica" county="Malaise" municipality="La" name="Finca de Elias Rojas" pageId="3" pageNumber="170" stateProvince="Guanacaste">Finca de Elias Rojas</location>
,
<collectorName id="26F6238DFF96996A1901FDEB607EFD9F" box="[1123,1318,547,573]" pageId="3" pageNumber="170">A. C. Tortuguero</collectorName>
,
<quantity id="4CFBEBBEFF96996A1850FDEB602CFD9F" box="[1330,1396,547,573]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="170" unit="m" value="100.0">
<elevation id="002EA168FF96996A1850FDEB602CFD9F" box="[1330,1396,547,573]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="170" unit="m" value="100.0">100m</elevation>
</quantity>
,
<date id="FFBD609BFF96996A18E2FDEB658AFDC3" pageId="3" pageNumber="170" value="1994-02">
<collectingDate id="EFF99973FF96996A18E2FDEB658AFDC3" pageId="3" pageNumber="170" value="1994-02">II- 1994</collectingDate>
</date>
,
<collectorName id="26F6238DFF96996A1DBFFD8F6467FDC3" box="[221,319,583,609]" pageId="3" pageNumber="170">E. Rojas</collectorName>
, L N 286000, 567500 #2608 (
<collectionCode id="ED12DE9EFF96996A1FECFD8F6780FDC3" box="[654,728,583,609]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="3" pageNumber="170">INBio</collectionCode>
, 1 f)
</materialsCitation>
.
</paragraph>
<paragraph id="8BBC465BFF96996D1DA5FDA26780FF77" blockId="3.[151,1437,150,2013]" lastBlockId="4.[151,1437,151,323]" lastPageId="4" lastPageNumber="171" pageId="3" pageNumber="170">
<emphasis id="B9779A49FF96996A1DA5FDA26409FD26" bold="true" box="[199,337,618,644]" pageId="3" pageNumber="170">Description</emphasis>
:
<typeStatus id="54B8F8F9FF96996A1C02FDA36490FD27" box="[352,456,619,645]" pageId="3" pageNumber="170" type="holotype">Holotype</typeStatus>
(male). Body length:
<quantity id="4CFBEBBEFF96996A1FDFFDA2667BFD27" box="[701,803,618,645]" metricMagnitude="-3" metricUnit="m" metricValue="5.35" pageId="3" pageNumber="170" unit="mm" value="5.35">5.35 mm</quantity>
. Head, scape and pedicel, mouthparts, thorax, legs, basal tarsomeres, elytra and abdomen bright yellowish-testaceous; apical tarsomeres and antennomeres III-VI somewhat infuscate (darkening apically), mandibles weakly infuscate basally, pale in central portion, darkest at apices. Antennomeres VII-X black; metathoracic macula near metacoxa with a black macula; apical third of elytra gilvous at apices; dorsum with black markings as follows: head with basomedial macula, pronotum with medial longitudinal vitta, posterior margin of pronotal collar with transverse border, entire scutellum, elytra with small macula at humeral angle, elytral suture from scutellum to midelytral fascia, midelytral fascia broad and complete.
<emphasis id="B9779A49FF96996A1994FC8A6066FCFF" bold="true" box="[1270,1342,834,861]" italics="true" pageId="3" pageNumber="170">Head:</emphasis>
Slightly wider than humeri; surface smooth, impunctate, moderately covered in fine, long, pale, erect, semirecumbent setae; setae on eyes and posterolateral margin of head (behind eyes) longer; eyes prominent, width across eyes distinctly wider than at pronotal tubercles, sparsely clothed with fine, long, erect, pale setae. Antennae 10-segmented with distinct single-segment club. Frons with two dorsally convergent elongate depressions. Mouthparts with clypeus longitudinally subdivided; mandibles bifid, right mandible dorsally excavated to receive the left; apical maxillary palpomeres securiform; apical labial palpomeres digitiform.
<emphasis id="B9779A49FF96996A1E2BFBD266C7FB97" bold="true" box="[841,927,1050,1077]" italics="true" pageId="3" pageNumber="170">Thorax</emphasis>
: Pronotum distinctly widest at lateral angles but narrower than width of head, covered in long, erect, pale setae interspersed with short, pale, semirecumbent setae; pronotal disc shiny, sparsely, finely punctate; punctures obsolete on anterior and posterior pronotal collars; disc very convex, anterior collar strongly constricted, upturned at head, transverse impression complete, anterior fringe of short, pale, erect setae projecting over head; posterior collar abruptly curved upward at scutellar margin; posterior collar with fringe of long, pale, erect setae projecting over scutellum; scutellum with long, pale, erect setae projecting over base of elytral suture. Ventral prothorax sparsely covered with long, fine, suberect setae which become shorter and more recumbent towards middle; procoxal cavities open; mesothorax densely covered with white recumbent setae; metepisternum similarly but less densely hairy; metathorax sparely covered with long, pale, erect setae. Metathorax large, somewhat inflated; surface smooth, minutely granulate, shiny.
<emphasis id="B9779A49FF96996A19D4FA9761B4FADB" bold="true" box="[1206,1260,1375,1401]" italics="true" pageId="3" pageNumber="170">Legs</emphasis>
: Surface shiny, covered with long, fine, testaceous, erect setae of varying length; femora expanded, weakly laterally compressed, nearly twice width of tibia, slightly wider distally; metafemora extending well past apex of elytra when projecting posteriorly; tibiae narrow, elongate, covered with long, fine, testaceous, erect setae of varying lengths, distal third with short fine, testaceous, semirecumbent setae, hind legs slightly longer but of similar form; tibial spur formula 1-2-2(?); tarsal claw bifid (ungues present).
<emphasis id="B9779A49FF96996A1FE1F9DB6793F98F" bold="true" box="[643,715,1555,1581]" italics="true" pageId="3" pageNumber="170">Wings</emphasis>
: Elytra parallel to subparallel, widest at humeri, shiny, distinctly punctate with large, widely spaced punctures, barely covering abdomen, disc subflattened dorsoventrally, covered in long, fine, pale, erect, semirecumbent setae of varying lengths; internal margin of plica forming subulate excavation with distinct margin, dorsal margin of excavation black (consistent with midelytral fascia), ventral margin of excavation testaceous, projecting inward more so than dorsal margin; lateral margins weakly serrulate, each serrulate denticle with a single seta, serrulations strongest apically, more feeble anteriorly, obsolete in anterior third; strongly dehiscent in apical third beginning at posterior end of midelytral fascia where plica broadens noticeably, apices distinctly tumid, serrate-spicular, independently subtruncate, interior angles of truncation negligibly serrate. Hindwing dusky-testaceous radiating from venation, apical third entirely cloudy black, not covered by elytra, obscuring apex of abdomen dorsally.
<emphasis id="B9779A49FF96996A1CECF89E64A4F8D3" bold="true" box="[398,508,1878,1905]" italics="true" pageId="3" pageNumber="170">Abdomen</emphasis>
: weakly shiny, sparsely covered with long, erect, pale setae; fifth sternite simple; sixth visible sternite entire, modified with lateral, inward-curved clasper-like appendages fused at middle; apical tergites unmodified.
<emphasis id="B9779A49FF96996A1CE3F85664BBF81B" bold="true" box="[385,483,1950,1977]" italics="true" pageId="3" pageNumber="170">Females</emphasis>
(
<typeStatus id="54B8F8F9FF96996A1C91F857673AF81B" box="[499,610,1951,1977]" pageId="3" pageNumber="170" type="paratype">paratypes</typeStatus>
) with a broad, gradual, smooth emargination on posterior margin of fifth sternite; sixth sternite divided longitudinally into two sclerites with widely-spaced, long, testaceous, subapical setae and dense apical fringe of shorter setae on each sclerite; apical tergite unmodified, but with pale setae of varying length, some subapical setae quite long and curved.
</paragraph>
</subSubSection>
<subSubSection id="C31915D0FF91996D1DA5FF1666E4FEE3" pageId="4" pageNumber="171" type="diagnosis">
<paragraph id="8BBC465BFF91996D1DA5FF1666E4FEE3" blockId="4.[151,1437,151,323]" pageId="4" pageNumber="171">
<emphasis id="B9779A49FF91996D1DA5FF166462FF5A" bold="true" box="[199,314,222,248]" pageId="4" pageNumber="171">Diagnosis</emphasis>
:
<taxonomicName id="4C033DD8FF91996D1C2AFF176728FF5B" authorityName=", Leavengood, Pinkerton and Rifkind" authorityYear="2022" box="[328,624,223,249]" class="Insecta" family="Cleridae" genus="Tarsobaenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="171" phylum="Arthropoda" rank="species" species="letourneauae">
<emphasis id="B9779A49FF91996D1C2AFF176728FF5B" box="[328,624,223,249]" italics="true" pageId="4" pageNumber="171">Tarsobaenus letourneauae</emphasis>
</taxonomicName>
is readily distinguished from its congeners by its broad, black, midelytral fascia (
<figureCitation id="13385ADEFF91996D1D85FECB6470FEBF" box="[231,296,259,285]" captionStart="FIGURES 16" captionStartId="4.[151,264,1584,1608]" captionTargetBox="[151,1436,345,1550]" captionTargetId="figure-102@4.[151,1436,345,1560]" captionTargetPageId="4" captionText="FIGURES 16. Tarsobaenus letourneauae, n. sp. (paratypes): 1—dorsal habitus; 2—head; 3—male, ventral abdomen; 4—female, ventral abdomen; 5—elytral plica; 6—lateral habitus." figureDoi="http://doi.org/10.5281/zenodo.6557095" httpUri="https://zenodo.org/record/6557095/files/figure.png" pageId="4" pageNumber="171">Fig. 1</figureCitation>
).
<taxonomicName id="4C033DD8FF91996D1C5DFECB6775FEBF" authorityName="Leavengood, Pinkerton and Rifkind" authorityYear="2022" box="[319,557,259,285]" class="Insecta" family="Cleridae" genus="Tarsobaenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="171" phylum="Arthropoda" rank="species" species="hansoni">
<emphasis id="B9779A49FF91996D1C5DFECB6775FEBF" box="[319,557,259,285]" italics="true" pageId="4" pageNumber="171">Tarsobaenus hansoni</emphasis>
</taxonomicName>
also has a broad, black, elytral fascia (
<figureCitation id="13385ADEFF91996D1EB8FECB6172FEBF" box="[986,1066,259,285]" captionStart="FIGURES 1216" captionStartId="8.[151,264,1473,1497]" captionTargetBox="[151,1436,237,1443]" captionTargetId="figure-48@8.[151,1436,237,1449]" captionTargetPageId="8" captionText="FIGURES 1216. Tarsobaenus hansoni, n. sp. (holotype, paratype): 12—holotype, dorsal habitus; 13—head; 14—paratype, dorsal habitus; 15—elytral plica; 16—lateral habitus." figureDoi="http://doi.org/10.5281/zenodo.6557101" httpUri="https://zenodo.org/record/6557101/files/figure.png" pageId="4" pageNumber="171">Fig. 12</figureCitation>
) but in this species the band covers most of the anterior elytral ½, rather than being positioned medially.
</paragraph>
</subSubSection>
<caption id="DF7C16D3FF91996D1DF5F9F867ACF9CE" ID-DOI="http://doi.org/10.5281/zenodo.6557095" ID-Zenodo-Dep="6557095" httpUri="https://zenodo.org/record/6557095/files/figure.png" pageId="4" pageNumber="171" startId="4.[151,264,1584,1608]" targetBox="[151,1436,345,1550]" targetPageId="4">
<paragraph id="8BBC465BFF91996D1DF5F9F867ACF9CE" blockId="4.[151,1436,1584,1644]" pageId="4" pageNumber="171">
<emphasis id="B9779A49FF91996D1DF5F9F86462F9EA" bold="true" box="[151,314,1584,1608]" pageId="4" pageNumber="171">FIGURES 16.</emphasis>
<taxonomicName id="4C033DD8FF91996D1C23F9F86716F9EA" authority="Leavengood &amp; Pinkerton &amp; Rifkind, 2022" authorityName="Leavengood &amp; Pinkerton &amp; Rifkind" authorityYear="2022" box="[321,590,1584,1608]" class="Insecta" family="Cleridae" genus="Tarsobaenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="171" phylum="Arthropoda" rank="species" species="letourneauae" status="sp. nov.">
<emphasis id="B9779A49FF91996D1C23F9F86716F9EA" box="[321,590,1584,1608]" italics="true" pageId="4" pageNumber="171">Tarsobaenus letourneauae</emphasis>
</taxonomicName>
,
<emphasis id="B9779A49FF91996D1F39F9F867CCF9EA" bold="true" box="[603,660,1584,1608]" pageId="4" pageNumber="171">
<taxonomicNameLabel id="A2442732FF91996D1F39F9F867CCF9EA" box="[603,660,1584,1608]" pageId="4" pageNumber="171" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(paratypes): 1—dorsal habitus; 2—head; 3—male, ventral abdomen; 4—female, ventral abdomen; 5—elytral plica; 6—lateral habitus.
</paragraph>
</caption>
<subSubSection id="C31915D0FF91996D1DA5F95264B1F8E6" pageId="4" pageNumber="171" type="description">
<paragraph id="8BBC465BFF91996D1DA5F95264B1F8E6" blockId="4.[151,1437,1690,2041]" pageId="4" pageNumber="171">
<emphasis id="B9779A49FF91996D1DA5F9526460F916" bold="true" box="[199,312,1690,1716]" pageId="4" pageNumber="171">Variation</emphasis>
: Head sometimes markedly pale ivory between the eyes; basomedial macula on head sometimes absent or expanded into a vitta; small macula on the elytral humeral angle may be reduced or absent; apical elytral whitening is consistent with increased tumidity resulting in obscured punctation; males with elytra slightly more elongate-parallel, occasionally constricted at middle. The disposition of the abdomen in most specimens obscures measurements of body length.
</paragraph>
</subSubSection>
<subSubSection id="C31915D0FF91996C1DA5F88661FAFF5B" lastPageId="5" lastPageNumber="172" pageId="4" pageNumber="171" type="biology_ecology">
<paragraph id="8BBC465BFF91996C1DA5F88661FAFF5B" blockId="4.[151,1437,1690,2041]" lastBlockId="5.[151,1437,151,357]" lastPageId="5" lastPageNumber="172" pageId="4" pageNumber="171">
<emphasis id="B9779A49FF91996D1DA5F8866479F8CA" bold="true" box="[199,289,1870,1896]" pageId="4" pageNumber="171">Biology</emphasis>
: The larvae of
<taxonomicName id="4C033DD8FF91996D1CB2F88667A2F8CA" authorityName="Leavengood &amp; Pinkerton &amp; Rifkind" authorityYear="2022" box="[464,762,1870,1896]" class="Insecta" family="Cleridae" genus="Tarsobaenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="171" phylum="Arthropoda" rank="species" species="letourneauae">
<emphasis id="B9779A49FF91996D1CB2F88667A2F8CA" box="[464,762,1870,1896]" italics="true" pageId="4" pageNumber="171">Tarsobaenus letourneauae</emphasis>
</taxonomicName>
occupy the petiole chambers of
<emphasis id="B9779A49FF91996D190EF8876062F8CA" box="[1132,1338,1870,1896]" italics="true" pageId="4" pageNumber="171">Piper cenocladum</emphasis>
(
<taxonomicName id="4C033DD8FF91996D182BF8866595F82F" class="Magnoliopsida" family="Piperaceae" kingdom="Plantae" order="Piperales" pageId="4" pageNumber="171" phylum="Tracheophyta" rank="family">Piperaceae</taxonomicName>
), in which they feed on the lipid and protein-rich food bodies produced by the plant. The larvae also feed on adult and immature
<taxonomicName id="4C033DD8FF91996D1C17F85E6723F812" authority="Forel" authorityName="Forel" authorityYear="1899" box="[373,635,1942,1968]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="171" phylum="Arthropoda" rank="species" species="bicornis">
<emphasis id="B9779A49FF91996D1C17F85E6762F812" box="[373,570,1942,1968]" italics="true" pageId="4" pageNumber="171">Pheidole bicornis</emphasis>
Forel
</taxonomicName>
(
<taxonomicName id="4C033DD8FF91996D1FEBF85E6670F812" box="[649,808,1942,1968]" class="Insecta" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="171" phylum="Arthropoda" rank="order">Hymenoptera</taxonomicName>
:
<taxonomicName id="4C033DD8FF91996D1E53F85E66EEF813" box="[817,950,1942,1969]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="171" phylum="Arthropoda" rank="family">Formicidae</taxonomicName>
). They appear to chemically mimic their ant prey to induce food body production in the plant, which is normally a species-specific response to the presence of the ants (
<bibRefCitation id="EF923BAAFF91996D1D9BF816649AF85A" author="Letourneau, D. K." box="[249,450,2014,2040]" pageId="4" pageNumber="171" pagination="215 - 217" refId="ref7504" refString="Letourneau, D. K. (1990) Code of ant-plant mutualism broken by parasite. Science, 248, 215 - 217. https: // doi. org / 10.1126 / science. 248.4952.215" type="journal article" year="1990">Letourneau, 1990</bibRefCitation>
;
<bibRefCitation id="EF923BAAFF91996D1CAFF81767A7F85A" author="Letourneau, D. K. &amp; Dyer, L. A." box="[461,767,2014,2041]" pageId="4" pageNumber="171" pagination="162 - 169" refId="ref7568" refString="Letourneau, D. K. &amp; Dyer, L. A. (1998) Density patterns of Piper ant-plants and associated arthropods: top-predator trophic cascades in a terrestrial system? Biotropica, 30 (2), 162 - 169. https: // doi. org / 10.1111 / j. 1744 - 7429.1998. tb 00051. x" type="journal article" year="1998">Letourneau and Dyer, 1998</bibRefCitation>
;
<bibRefCitation id="EF923BAAFF91996D1E6BF8176164F85A" author="Dyer, L. A. &amp; Letourneau, D. K." box="[777,1084,2014,2041]" pageId="4" pageNumber="171" pagination="5072 - 5076" refId="ref7005" refString="Dyer, L. A. &amp; Letourneau, D. K. (1999) Trophic cascades in a complex terrestrial community. Proceedings of the National Academy of Sciences (USA), 96, 5072 - 5076." type="journal article" year="1999">Dyer and Letourneau, 1999</bibRefCitation>
,
<bibRefCitation id="EF923BAAFF91996D1924F81661D9F85A" author="Dyer, L. A. &amp; Letourneau, D. K." box="[1094,1153,2014,2040]" pageId="4" pageNumber="171" pagination="60 - 68" refId="ref7046" refString="Dyer, L. A. &amp; Letourneau, D. K. (2003) Top-down and bottom-up diversity cascades in detrital vs. living food webs. Ecology Letters, 6, 60 - 68. https: // doi. org / 10.1046 / j. 1461 - 0248.2003.00398. x" type="journal article" year="2003">2003</bibRefCitation>
;
<bibRefCitation id="EF923BAAFF91996D19E9F81760C9F85A" author="Letourneau, D. K. &amp; Dyer, L. A. &amp; Vega C." box="[1163,1425,2014,2041]" pageId="4" pageNumber="171" pagination="2144 - 2152" refId="ref7631" refString="Letourneau, D. K., Dyer, L. A. &amp; Vega C., G. (2004) Indirect effects of a top predator on a rainforest understory plant community. Ecology, 85 (8), 2144 - 2152. https: // doi. org / 10.1890 / 03 - 0525" type="journal article" year="2004">
Letourneau
<emphasis id="B9779A49FF91996D1873F8176018F85A" box="[1297,1344,2014,2040]" italics="true" pageId="4" pageNumber="171">et al</emphasis>
., 2004
</bibRefCitation>
). These beetles have been observed in primary and secondary lowland rain forests at the La Selva Biological Station.
<bibRefCitation id="EF923BAAFF90996C1DF5FF73648FFF77" author="Letourneau, D. K. &amp; Dyer, L. A." box="[151,471,187,213]" pageId="5" pageNumber="172" pagination="162 - 169" refId="ref7568" refString="Letourneau, D. K. &amp; Dyer, L. A. (1998) Density patterns of Piper ant-plants and associated arthropods: top-predator trophic cascades in a terrestrial system? Biotropica, 30 (2), 162 - 169. https: // doi. org / 10.1111 / j. 1744 - 7429.1998. tb 00051. x" type="journal article" year="1998">Letourneau and Dyer (1998)</bibRefCitation>
reported the beetles as being most active in March and April. However, our specimen label data indicate most adult collection records occurred in January, August and September.
</paragraph>
</subSubSection>
<subSubSection id="C31915D0FF90996C1DA5FECA66B5FEBF" box="[199,1005,258,285]" pageId="5" pageNumber="172" type="distribution">
<paragraph id="8BBC465BFF90996C1DA5FECA66B5FEBF" blockId="5.[151,1437,151,357]" box="[199,1005,258,285]" pageId="5" pageNumber="172">
<emphasis id="B9779A49FF90996C1DA5FECA6401FEBE" bold="true" box="[199,345,258,284]" pageId="5" pageNumber="172">Distribution</emphasis>
:
<collectingRegion id="49C788B9FF90996C1C0AFECB64EEFEBF" box="[360,438,259,285]" country="Costa Rica" name="Limon" pageId="5" pageNumber="172">Limón</collectingRegion>
,
<collectingRegion id="49C788B9FF90996C1CA3FECB671AFEBF" box="[449,578,259,285]" country="Costa Rica" name="Guanacaste" pageId="5" pageNumber="172">Guanacaste</collectingRegion>
and
<collectingRegion id="49C788B9FF90996C1F1AFECB6789FEBF" box="[632,721,259,285]" country="Costa Rica" name="Heredia" pageId="5" pageNumber="172">Heredia</collectingRegion>
Provinces of
<collectingCountry id="F31406CBFF90996C1E0EFECB66B2FEBF" box="[876,1002,259,285]" name="Costa Rica" pageId="5" pageNumber="172">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C31915D0FF90996C1DA5FEEF61C8FEC7" pageId="5" pageNumber="172" type="etymology">
<paragraph id="8BBC465BFF90996C1DA5FEEF61C8FEC7" blockId="5.[151,1437,151,357]" pageId="5" pageNumber="172">
<emphasis id="B9779A49FF90996C1DA5FEEF6410FEE3" bold="true" box="[199,328,295,321]" pageId="5" pageNumber="172">Etymology</emphasis>
: The patronym
<taxonomicName id="4C033DD8FF90996C1C95FEEF67D3FEE3" authorityName="Leavengood &amp; Pinkerton &amp; Rifkind" authorityYear="2022" box="[503,651,295,321]" class="Insecta" family="Cleridae" genus="Tarsobaenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="species" species="letourneauae">
<emphasis id="B9779A49FF90996C1C95FEEF67D3FEE3" box="[503,651,295,321]" italics="true" pageId="5" pageNumber="172">letourneauae</emphasis>
</taxonomicName>
honors Dr. Deborah Letourneau (University of California-Santa Cruz), the scientist who first discovered this species and first documented its interesting behavior.
</paragraph>
</subSubSection>
</treatment>
</document>