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<document id="232F68B3BA48B31FB39F09758FA73AFB" ID-DOI="10.5281/zenodo.5341630" ID-ISSN="2345-7600" ID-Zenodo-Dep="5341630" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630191096889" checkinUser="diego" docAuthor="Anker, Arthur &amp; Marin, Ivan N." docDate="2009" docId="03AAE9259F16FFE7FC3FFEB16BC34CB4" docLanguage="en" docName="57rbz091-107.pdf" docOrigin="Raffles Bulletin of Zoology 57 (1)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Leptalpheus dworschaki Anker &amp; Marin 2009, new species" docType="treatment" docVersion="3" lastPageNumber="105" masterDocId="FF93915D9F1FFFE9FFA9FFCA6B294E67" masterDocTitle="The Alpheid Shrimp Genus Leptalpheus Williams, 1965, In The Tropical Western Pacific, With Descriptions Of Two New Species (Crustacea: Decapoda: Caridea)" masterLastPageNumber="107" masterPageNumber="91" pageNumber="100" updateTime="1699181539165" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="2AFD714B709E319652D8D19EF65BF8E6">The Alpheid Shrimp Genus Leptalpheus Williams, 1965, In The Tropical Western Pacific, With Descriptions Of Two New Species (Crustacea: Decapoda: Caridea)</mods:title>
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<mods:namePart id="ABBAA1A4AC0CF82007E9ED8E1FE850AA">Anker, Arthur</mods:namePart>
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<mods:namePart id="094EAD4560881D8BB34AD1BD7401823D">Marin, Ivan N.</mods:namePart>
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<mods:title id="AB0F93293D1363A9F27A8D99C90B2ECE">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="4CA142BA9E520CB840A550B28938093E">2009</mods:date>
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<treatment id="03AAE9259F16FFE7FC3FFEB16BC34CB4" LSID="urn:lsid:plazi:treatment:03AAE9259F16FFE7FC3FFEB16BC34CB4" httpUri="http://treatment.plazi.org/id/03AAE9259F16FFE7FC3FFEB16BC34CB4" lastPageId="14" lastPageNumber="105" pageId="9" pageNumber="100">
<subSubSection id="C3190BB89F16FFE0FC3FFEB16E104FF4" box="[918,1337,379,403]" pageId="9" pageNumber="100" type="nomenclature">
<paragraph id="8BBC58339F16FFE0FC3FFEB16E104FF4" blockId="9.[918,1337,379,434]" box="[918,1337,379,403]" pageId="9" pageNumber="100">
<heading id="D0F4EF5F9F16FFE0FC3FFEB16E104FF4" bold="true" box="[918,1337,379,403]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="100" reason="2">
<emphasis id="B97784219F16FFE0FC3FFEB16E104FF4" bold="true" box="[918,1337,379,403]" pageId="9" pageNumber="100">
<taxonomicName id="4C0323B09F16FFE0FC3FFEB16F8D4FF4" ID-CoL="85TJJ" authority="Anker &amp; Marin, 2009" authorityName="Anker &amp; Marin" authorityYear="2009" box="[918,1188,379,403]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="100" phylum="Arthropoda" rank="species" species="dworschaki" status="sp. nov.">
<emphasis id="B97784219F16FFE0FC3FFEB16F8D4FF4" bold="true" box="[918,1188,379,403]" italics="true" pageId="9" pageNumber="100">Leptalpheus dworschaki</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A244395A9F16FFE0FB1AFEB16E104FF4" box="[1203,1337,379,403]" pageId="9" pageNumber="100" rank="species">new species</taxonomicNameLabel>
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<subSubSection id="C3190BB89F16FFE0FB8CFE516EA94F88" pageId="9" pageNumber="100" type="description">
<paragraph id="8BBC58339F16FFE0FB8CFE516F834FD5" blockId="9.[918,1337,379,434]" box="[1061,1194,411,434]" pageId="9" pageNumber="100">
(
<figureCitation id="133844B69F16FFE0FB84FE516F8B4FD5" box="[1069,1186,411,434]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="12.[132,167,2002,2023]" captionStartId-1="13.[109,144,2030,2051]" captionStartId-2="14.[132,167,2057,2078]" captionStartId-3="15.[109,144,2031,2052]" captionTargetBox-0="[143,1460,196,1957]" captionTargetBox-1="[112,1445,292,1926]" captionTargetBox-2="[139,1461,837,2001]" captionTargetBox-3="[131,1426,195,1973]" captionTargetId-0="figure-92@12.[113,1491,165,1979]" captionTargetId-1="figure-64@13.[78,1476,258,1959]" captionTargetId-2="figure-49@14.[101,1499,802,2027]" captionTargetId-3="figure-61@15.[101,1455,149,2005]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionTargetPageId-2="14" captionTargetPageId-3="15" captionText-0="Fig. 9. Leptalpheus dworschaki, new species, male holotype (NMCR 27540): a, frontal region, dorsal view; b, same, lateral view; c, frontal margin of carapace, lateral view; d, first segment of right antennular peduncle, lateral view; e, lateral flagellum of right antennule, lateral view; f, first pleopod, mesial view; g, second pleopod, mesial view; h, same, appendix masculina and appendix interna; i, right uropod, dorsal view; j, telson, dorsal view; female paratype (ZRC 2005.0085): k, frontal margin, dorsal view. Scale bars = 1 mm." captionText-1="Fig. 10. Leptalpheus dworschaki, new species, female paratype (ZRC 2005.0085): a, habitus; male holotype (NMCR 27540): b, mandible, mesial view; c, same, dorsolateral view; d, maxillule, lateral view; e, maxilla, lateral view; f, first maxilliped, lateral view; g, second maxilliped, lateral view; h, third maxilliped, lateral view. Scale bar = 1 mm." captionText-2="Fig. 11. Leptalpheus dworschaki, new species, male holotype (NMCR 27540): a, major (left) cheliped, mesial view; b, same, lateral view; c, same, chela, carpus and distal merus, ventrolateral view; d, same, distal chela, ventromesial view. Scale bars = 1 mm." captionText-3="Fig. 12. Leptalpheus dworschaki, new species, female paratype (ZRC 2005.0085): a, minor (right) cheliped, lateral view; b, same, chela, mesial view; male holotype (NMCR 27540): c, second pereiopod, lateral view; d, third pereiopod, lateral view; e, fourth pereiopod, lateral view; f, fifth pereiopod, lateral view. Scale bars = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5341648" figureDoi-1="http://doi.org/10.5281/zenodo.5341650" figureDoi-2="http://doi.org/10.5281/zenodo.5341652" figureDoi-3="http://doi.org/10.5281/zenodo.5341654" httpUri-0="https://zenodo.org/record/5341648/files/figure.png" httpUri-1="https://zenodo.org/record/5341650/files/figure.png" httpUri-2="https://zenodo.org/record/5341652/files/figure.png" httpUri-3="https://zenodo.org/record/5341654/files/figure.png" pageId="9" pageNumber="100">Figs. 912</figureCitation>
)
</paragraph>
<paragraph id="8BBC58339F16FFE0FC8EFE106EA94F88" blockId="9.[807,1408,474,495]" box="[807,1408,474,495]" pageId="9" pageNumber="100">
<taxonomicName id="4C0323B09F16FFE0FC8EFE1068FD4F88" box="[807,980,474,495]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="100" phylum="Arthropoda" rank="species" species="undefined-7">
<emphasis id="B97784219F16FFE0FC8EFE1068B54F88" box="[807,924,474,495]" italics="true" pageId="9" pageNumber="100">Leptalpheus</emphasis>
sp. 7
</taxonomicName>
<taxonomicName id="4C0323B09F16FFE0FC72FE106F704F88" baseAuthorityName="Banner &amp; Banner" baseAuthorityYear="1974" box="[987,1113,474,495]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="100" phylum="Arthropoda" rank="species" species="pacificus">
aff.
<emphasis id="B97784219F16FFE0FBADFE106F704F88" box="[1028,1113,474,495]" italics="true" pageId="9" pageNumber="100">pacificus</emphasis>
</taxonomicName>
<bibRefCitation id="EF9225C29F16FFE0FBDCFE106E0C4F88" author="Anker, A" box="[1141,1317,474,495]" pageId="9" pageNumber="100" pagination="683 - 702" refId="ref7571" refString="Anker, A., J. A. Vera Caripe &amp; C. Lira, 2006. Description of a new species of commensal alpheid shrimp (Crustacea, Decapoda) from the southern Caribbean Sea. Zoosystema, 28: 683 - 702" type="journal article" year="2006">Anker et al, 2006</bibRefCitation>
: Table 1.
</paragraph>
</subSubSection>
<subSubSection id="C3190BB89F16FFE0FC8EFDD968D64CB2" pageId="9" pageNumber="100" type="materials_examined">
<paragraph id="8BBC58339F16FFE0FC8EFDD968D64CB2" blockId="9.[807,1448,531,725]" pageId="9" pageNumber="100">
<emphasis id="B97784219F16FFE0FC8EFDD968E74C4F" bold="true" box="[807,974,531,552]" pageId="9" pageNumber="100">
<emphasis id="B97784219F16FFE0FC8EFDD968904C4F" bold="true" box="[807,953,531,552]" italics="true" pageId="9" pageNumber="100">Type material.</emphasis>
</emphasis>
<materialsCitation id="3B6B526E9F16FFE0FC7EFDD96FF44CFC" collectingDate="2004-06-07" collectionCode="NMCR" collectorName="Sta. M &amp; P. C. Dworschak" country="Philippines" latitude="9.638333" location="Panglao Island" longLatPrecision="129" longitude="123.82667" municipality="Sungcolan Bay" pageId="9" pageNumber="100" specimenCode="NMCR 27540" specimenCount="1" typeStatus="holotype">
<emphasis id="B97784219F16FFE0FC7EFDD96F634C4F" bold="true" box="[983,1098,531,552]" pageId="9" pageNumber="100">
<collectingCountry id="F31418A39F16FFE0FC7EFDD96F634C4F" box="[983,1098,531,552]" name="Philippines" pageId="9" pageNumber="100">Philippines</collectingCountry>
</emphasis>
:
<typeStatus id="54B8E6919F16FFE0FBF3FDD96F914C4F" box="[1114,1208,531,552]" pageId="9" pageNumber="100" type="holotype">Holotype</typeStatus>
: male, CL 7.7, TL 21.9 (
<specimenCode id="DBA5F0489F16FFE0FC84FDFA68904C22" box="[813,953,560,581]" collectionCode="NMCR" country="USA" lsid="urn:lsid:biocol.org:col:14936" name="New Mexico State University" pageId="9" pageNumber="100" type="Herbarium">NMCR 27540</specimenCode>
),
<location id="8EDC0EE89F16FFE0FC6EFDFA6F7A4C22" LSID="urn:lsid:plazi:treatment:03AAE9259F16FFE7FC3FFEB16BC34CB4:8EDC0EE89F16FFE0FC6EFDFA6F7A4C22" box="[967,1107,560,581]" country="Philippines" latitude="9.638333" longLatPrecision="129" longitude="123.82667" municipality="Sungcolan Bay" name="Panglao Island" pageId="9" pageNumber="100">Panglao Island</location>
,
<collectingMunicipality id="6BD8C2499F16FFE0FBF2FDFA6FC24C22" box="[1115,1259,560,581]" pageId="9" pageNumber="100">Sungcolan Bay</collectingMunicipality>
,
<collectorName id="26F63DE59F16FFE0FB5DFDFA6E1D4C22" box="[1268,1332,560,581]" pageId="9" pageNumber="100">Sta. M</collectorName>
11,
<geoCoordinate id="EE373EF49F16FFE0FAFDFDFA6E814C22" box="[1364,1448,560,581]" degrees="9" direction="north" minutes="38.3" orientation="latitude" pageId="9" pageNumber="100" precision="92" value="9.638333">9°38.3'N</geoCoordinate>
<geoCoordinate id="EE373EF49F16FFE0FC8EFD8768BD4C05" box="[807,916,589,610]" degrees="123" direction="east" minutes="49.6" orientation="longitude" pageId="9" pageNumber="100" precision="92" value="123.82667">123°49.6'E</geoCoordinate>
, intertidal flat, fringed with mangrove and seagrass, bait suction pump, from burrow of
<taxonomicName id="4C0323B09F16FFE0FBE1FDA06FE34C18" box="[1096,1226,618,639]" class="Malacostraca" family="Callianassidae" genus="Glypturus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="100" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B97784219F16FFE0FBE1FDA06F814C18" box="[1096,1192,618,639]" italics="true" pageId="9" pageNumber="100">Glypturus</emphasis>
sp.
</taxonomicName>
, coll.
<collectorName id="26F63DE59F16FFE0FAABFDA06E8D4C18" box="[1282,1444,618,639]" pageId="9" pageNumber="100">P. C. Dworschak</collectorName>
,
<date id="FFBD7EF39F16FFE0FC8EFD4C68BA4CFC" box="[807,915,646,667]" pageId="9" pageNumber="100" value="2004-06-07">
<collectingDate id="EFF9871B9F16FFE0FC8EFD4C68BA4CFC" box="[807,915,646,667]" pageId="9" pageNumber="100" value="2004-06-07">7 Jun.2004</collectingDate>
</date>
[fcn PD-37; specimen dissected]
</materialsCitation>
.
<materialsCitation id="3B6B526E9F16FFE0FB40FD4C68D24CB2" collectingDate="2004-06-07" collectionCode="ZRC" collectorName="Sta. M &amp; P. C. Dworschak" country="Philippines" latitude="9.638333" location="Panglao Island" longLatPrecision="129" longitude="123.82667" municipality="Sungcolan Bay" pageId="9" pageNumber="100" specimenCode="ZRC 2005.0085" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="54B8E6919F16FFE0FB40FD4C6E684CFC" box="[1257,1345,646,667]" pageId="9" pageNumber="100" type="paratype">Paratype</typeStatus>
:
<specimenCount id="9D0593BA9F16FFE0FAE5FD4C6E8D4CFC" box="[1356,1444,646,667]" pageId="9" pageNumber="100" type="female">1 female</specimenCount>
, CL 7.7, TL 23.9 (
<specimenCode id="DBA5F0489F16FFE0FC77FD696FAA4CDF" box="[990,1155,675,696]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="100">ZRC 2005.0085</specimenCode>
), same collection data as for holotype [fcn PD-36]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3190BB89F16FFE2FC8EFD3468F54A14" lastPageId="11" lastPageNumber="102" pageId="9" pageNumber="100" type="description">
<paragraph id="8BBC58339F16FFE0FC8EFD34689A4A12" blockId="9.[807,1448,766,1141]" pageId="9" pageNumber="100">
<emphasis id="B97784219F16FFE0FC8EFD3468ED4D71" bold="true" box="[807,964,766,790]" pageId="9" pageNumber="100">
<emphasis id="B97784219F16FFE0FC8EFD3468844D71" bold="true" box="[807,941,766,790]" italics="true" pageId="9" pageNumber="100">Description.</emphasis>
</emphasis>
Body not particularly slender or stout (
<figureCitation id="133844B69F16FFE0FAD5FD3468794D52" captionStart="Fig" captionStartId="13.[109,144,2030,2051]" captionTargetBox="[112,1445,292,1926]" captionTargetId="figure-64@13.[78,1476,258,1959]" captionTargetPageId="13" captionText="Fig. 10. Leptalpheus dworschaki, new species, female paratype (ZRC 2005.0085): a, habitus; male holotype (NMCR 27540): b, mandible, mesial view; c, same, dorsolateral view; d, maxillule, lateral view; e, maxilla, lateral view; f, first maxilliped, lateral view; g, second maxilliped, lateral view; h, third maxilliped, lateral view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341650" httpUri="https://zenodo.org/record/5341650/files/figure.png" pageId="9" pageNumber="100">Fig. 10a</figureCitation>
), carapace and abdomen slightly compressed laterally, glabrous. Carapace with inconspicuous suture proximal to base of antenna (
<figureCitation id="133844B69F16FFE0FC49FC946F634D12" box="[992,1098,862,885]" captionStart="Fig" captionStartId="12.[132,167,2002,2023]" captionTargetBox="[143,1460,196,1957]" captionTargetId="figure-92@12.[113,1491,165,1979]" captionTargetPageId="12" captionText="Fig. 9. Leptalpheus dworschaki, new species, male holotype (NMCR 27540): a, frontal region, dorsal view; b, same, lateral view; c, frontal margin of carapace, lateral view; d, first segment of right antennular peduncle, lateral view; e, lateral flagellum of right antennule, lateral view; f, first pleopod, mesial view; g, second pleopod, mesial view; h, same, appendix masculina and appendix interna; i, right uropod, dorsal view; j, telson, dorsal view; female paratype (ZRC 2005.0085): k, frontal margin, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341648" httpUri="https://zenodo.org/record/5341648/files/figure.png" pageId="9" pageNumber="100">Fig. 9b, c</figureCitation>
). Frontal margin protruding and broadly rounded, without rostral projection or orbital teeth, without orbital crests (
<figureCitation id="133844B69F16FFE0FB89FC546F844DD2" box="[1056,1197,926,949]" captionStart="Fig" captionStartId="12.[132,167,2002,2023]" captionTargetBox="[143,1460,196,1957]" captionTargetId="figure-92@12.[113,1491,165,1979]" captionTargetPageId="12" captionText="Fig. 9. Leptalpheus dworschaki, new species, male holotype (NMCR 27540): a, frontal region, dorsal view; b, same, lateral view; c, frontal margin of carapace, lateral view; d, first segment of right antennular peduncle, lateral view; e, lateral flagellum of right antennule, lateral view; f, first pleopod, mesial view; g, second pleopod, mesial view; h, same, appendix masculina and appendix interna; i, right uropod, dorsal view; j, telson, dorsal view; female paratype (ZRC 2005.0085): k, frontal margin, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341648" httpUri="https://zenodo.org/record/5341648/files/figure.png" pageId="9" pageNumber="100">Fig. 9a, b, k</figureCitation>
). Pterygostomial angle rounded (
<figureCitation id="133844B69F16FFE0FC39FC7468C94DB2" box="[912,992,958,981]" captionStart="Fig" captionStartId="12.[132,167,2002,2023]" captionTargetBox="[143,1460,196,1957]" captionTargetId="figure-92@12.[113,1491,165,1979]" captionTargetPageId="12" captionText="Fig. 9. Leptalpheus dworschaki, new species, male holotype (NMCR 27540): a, frontal region, dorsal view; b, same, lateral view; c, frontal margin of carapace, lateral view; d, first segment of right antennular peduncle, lateral view; e, lateral flagellum of right antennule, lateral view; f, first pleopod, mesial view; g, second pleopod, mesial view; h, same, appendix masculina and appendix interna; i, right uropod, dorsal view; j, telson, dorsal view; female paratype (ZRC 2005.0085): k, frontal margin, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341648" httpUri="https://zenodo.org/record/5341648/files/figure.png" pageId="9" pageNumber="100">Fig. 9c</figureCitation>
); branchiostegial region with pronounced “lip” anteriorly; cardiac notch fairly deep (
<figureCitation id="133844B69F16FFE0FAAEFC146E4B4D92" box="[1287,1378,990,1013]" captionStart="Fig" captionStartId="12.[132,167,2002,2023]" captionTargetBox="[143,1460,196,1957]" captionTargetId="figure-92@12.[113,1491,165,1979]" captionTargetPageId="12" captionText="Fig. 9. Leptalpheus dworschaki, new species, male holotype (NMCR 27540): a, frontal region, dorsal view; b, same, lateral view; c, frontal margin of carapace, lateral view; d, first segment of right antennular peduncle, lateral view; e, lateral flagellum of right antennule, lateral view; f, first pleopod, mesial view; g, second pleopod, mesial view; h, same, appendix masculina and appendix interna; i, right uropod, dorsal view; j, telson, dorsal view; female paratype (ZRC 2005.0085): k, frontal margin, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341648" httpUri="https://zenodo.org/record/5341648/files/figure.png" pageId="9" pageNumber="100">Figs. 9c</figureCitation>
,
<figureCitation id="133844B69F16FFE0FAD8FC146EB24D92" box="[1393,1435,990,1013]" captionStart="Fig" captionStartId="13.[109,144,2030,2051]" captionTargetBox="[112,1445,292,1926]" captionTargetId="figure-64@13.[78,1476,258,1959]" captionTargetPageId="13" captionText="Fig. 10. Leptalpheus dworschaki, new species, female paratype (ZRC 2005.0085): a, habitus; male holotype (NMCR 27540): b, mandible, mesial view; c, same, dorsolateral view; d, maxillule, lateral view; e, maxilla, lateral view; f, first maxilliped, lateral view; g, second maxilliped, lateral view; h, third maxilliped, lateral view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341650" httpUri="https://zenodo.org/record/5341650/files/figure.png" pageId="9" pageNumber="100">10a</figureCitation>
). Eyes not visible in dorsal view, anterior portion visible in lateral view (
<figureCitation id="133844B69F16FFE0FC49FBD46F7B4A52" box="[992,1106,1054,1077]" captionStart="Fig" captionStartId="12.[132,167,2002,2023]" captionTargetBox="[143,1460,196,1957]" captionTargetId="figure-92@12.[113,1491,165,1979]" captionTargetPageId="12" captionText="Fig. 9. Leptalpheus dworschaki, new species, male holotype (NMCR 27540): a, frontal region, dorsal view; b, same, lateral view; c, frontal margin of carapace, lateral view; d, first segment of right antennular peduncle, lateral view; e, lateral flagellum of right antennule, lateral view; f, first pleopod, mesial view; g, second pleopod, mesial view; h, same, appendix masculina and appendix interna; i, right uropod, dorsal view; j, telson, dorsal view; female paratype (ZRC 2005.0085): k, frontal margin, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341648" httpUri="https://zenodo.org/record/5341648/files/figure.png" pageId="9" pageNumber="100">Fig. 9a, c</figureCitation>
); anteromesial process feebly marked; cornea small, lateral, pigmented. Ocellar beak not conspicuous.
</paragraph>
<paragraph id="8BBC58339F16FFE0FC8EFB546E314B92" blockId="9.[807,1448,1182,1525]" pageId="9" pageNumber="100">
Antennular peduncle relatively stout, flattened dorsoventrally; second segment about twice as long as broad, about as long as dorsally visible portion of first segment; stylocerite not reaching distal margin of first segment, subacute distally (
<figureCitation id="133844B69F16FFE0FAD4FB34685D4B52" captionStart="Fig" captionStartId="12.[132,167,2002,2023]" captionTargetBox="[143,1460,196,1957]" captionTargetId="figure-92@12.[113,1491,165,1979]" captionTargetPageId="12" captionText="Fig. 9. Leptalpheus dworschaki, new species, male holotype (NMCR 27540): a, frontal region, dorsal view; b, same, lateral view; c, frontal margin of carapace, lateral view; d, first segment of right antennular peduncle, lateral view; e, lateral flagellum of right antennule, lateral view; f, first pleopod, mesial view; g, second pleopod, mesial view; h, same, appendix masculina and appendix interna; i, right uropod, dorsal view; j, telson, dorsal view; female paratype (ZRC 2005.0085): k, frontal margin, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341648" httpUri="https://zenodo.org/record/5341648/files/figure.png" pageId="9" pageNumber="100">Fig. 9a, b, k</figureCitation>
); ventromesial carina of first segment with strong tooth (
<figureCitation id="133844B69F16FFE0FC86FAF468544B32" box="[815,893,1342,1365]" captionStart="Fig" captionStartId="12.[132,167,2002,2023]" captionTargetBox="[143,1460,196,1957]" captionTargetId="figure-92@12.[113,1491,165,1979]" captionTargetPageId="12" captionText="Fig. 9. Leptalpheus dworschaki, new species, male holotype (NMCR 27540): a, frontal region, dorsal view; b, same, lateral view; c, frontal margin of carapace, lateral view; d, first segment of right antennular peduncle, lateral view; e, lateral flagellum of right antennule, lateral view; f, first pleopod, mesial view; g, second pleopod, mesial view; h, same, appendix masculina and appendix interna; i, right uropod, dorsal view; j, telson, dorsal view; female paratype (ZRC 2005.0085): k, frontal margin, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341648" httpUri="https://zenodo.org/record/5341648/files/figure.png" pageId="9" pageNumber="100">Fig. 9d</figureCitation>
); lateral flagellum biramous, with shorter ramus well developed, situated at third segment. Antenna with basicerite bearing strong ventrolateral tooth (
<figureCitation id="133844B69F16FFE0FB1BFAB46E2E4BF2" box="[1202,1287,1406,1429]" captionStart="Fig" captionStartId="12.[132,167,2002,2023]" captionTargetBox="[143,1460,196,1957]" captionTargetId="figure-92@12.[113,1491,165,1979]" captionTargetPageId="12" captionText="Fig. 9. Leptalpheus dworschaki, new species, male holotype (NMCR 27540): a, frontal region, dorsal view; b, same, lateral view; c, frontal margin of carapace, lateral view; d, first segment of right antennular peduncle, lateral view; e, lateral flagellum of right antennule, lateral view; f, first pleopod, mesial view; g, second pleopod, mesial view; h, same, appendix masculina and appendix interna; i, right uropod, dorsal view; j, telson, dorsal view; female paratype (ZRC 2005.0085): k, frontal margin, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341648" httpUri="https://zenodo.org/record/5341648/files/figure.png" pageId="9" pageNumber="100">Fig. 9b</figureCitation>
); scaphocerite broadly ovate, anterior margin of blade convex but not protruding beyond distolateral tooth (
<figureCitation id="133844B69F16FFE0FB6EFA746E304BB2" box="[1223,1305,1470,1493]" captionStart="Fig" captionStartId="12.[132,167,2002,2023]" captionTargetBox="[143,1460,196,1957]" captionTargetId="figure-92@12.[113,1491,165,1979]" captionTargetPageId="12" captionText="Fig. 9. Leptalpheus dworschaki, new species, male holotype (NMCR 27540): a, frontal region, dorsal view; b, same, lateral view; c, frontal margin of carapace, lateral view; d, first segment of right antennular peduncle, lateral view; e, lateral flagellum of right antennule, lateral view; f, first pleopod, mesial view; g, second pleopod, mesial view; h, same, appendix masculina and appendix interna; i, right uropod, dorsal view; j, telson, dorsal view; female paratype (ZRC 2005.0085): k, frontal margin, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341648" httpUri="https://zenodo.org/record/5341648/files/figure.png" pageId="9" pageNumber="100">Fig. 9a</figureCitation>
); carpocerite long, stout, reaching far beyond scaphocerite.
</paragraph>
<paragraph id="8BBC58339F16FFE0FC8EF9D46FB54832" blockId="9.[807,1448,1566,1621]" pageId="9" pageNumber="100">
Mouthparts and third maxilliped as in previous species (
<figureCitation id="133844B69F16FFE0FAD5F9D468444832" captionStart="Fig" captionStartId="13.[109,144,2030,2051]" captionTargetBox="[112,1445,292,1926]" captionTargetId="figure-64@13.[78,1476,258,1959]" captionTargetPageId="13" captionText="Fig. 10. Leptalpheus dworschaki, new species, female paratype (ZRC 2005.0085): a, habitus; male holotype (NMCR 27540): b, mandible, mesial view; c, same, dorsolateral view; d, maxillule, lateral view; e, maxilla, lateral view; f, first maxilliped, lateral view; g, second maxilliped, lateral view; h, third maxilliped, lateral view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341650" httpUri="https://zenodo.org/record/5341650/files/figure.png" pageId="9" pageNumber="100">Fig. 10bh</figureCitation>
; for description see above).
</paragraph>
<paragraph id="8BBC58339F16FFE2FC8EF9B46A964F94" blockId="9.[807,1448,1662,2101]" lastBlockId="11.[109,748,188,499]" lastPageId="11" lastPageNumber="102" pageId="9" pageNumber="100">
Chelipeds strongly asymmetrical in shape, unequal in size (
<figureCitation id="133844B69F16FFE0FC99F95468B948D2" box="[816,912,1694,1717]" captionStart="Fig" captionStartId="14.[132,167,2057,2078]" captionTargetBox="[139,1461,837,2001]" captionTargetId="figure-49@14.[101,1499,802,2027]" captionTargetPageId="14" captionText="Fig. 11. Leptalpheus dworschaki, new species, male holotype (NMCR 27540): a, major (left) cheliped, mesial view; b, same, lateral view; c, same, chela, carpus and distal merus, ventrolateral view; d, same, distal chela, ventromesial view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341652" httpUri="https://zenodo.org/record/5341652/files/figure.png" pageId="9" pageNumber="100">Figs. 11</figureCitation>
,
<figureCitation id="133844B69F16FFE0FC09F95468C348D2" box="[928,1002,1694,1717]" captionStart="Fig" captionStartId="15.[109,144,2031,2052]" captionTargetBox="[131,1426,195,1973]" captionTargetId="figure-61@15.[101,1455,149,2005]" captionTargetPageId="15" captionText="Fig. 12. Leptalpheus dworschaki, new species, female paratype (ZRC 2005.0085): a, minor (right) cheliped, lateral view; b, same, chela, mesial view; male holotype (NMCR 27540): c, second pereiopod, lateral view; d, third pereiopod, lateral view; e, fourth pereiopod, lateral view; f, fifth pereiopod, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341654" httpUri="https://zenodo.org/record/5341654/files/figure.png" pageId="9" pageNumber="100">12a, b</figureCitation>
), carried folded when not in use (
<figureCitation id="133844B69F16FFE0FAD3F954687848B2" captionStart="Fig" captionStartId="13.[109,144,2030,2051]" captionTargetBox="[112,1445,292,1926]" captionTargetId="figure-64@13.[78,1476,258,1959]" captionTargetPageId="13" captionText="Fig. 10. Leptalpheus dworschaki, new species, female paratype (ZRC 2005.0085): a, habitus; male holotype (NMCR 27540): b, mandible, mesial view; c, same, dorsolateral view; d, maxillule, lateral view; e, maxilla, lateral view; f, first maxilliped, lateral view; g, second maxilliped, lateral view; h, third maxilliped, lateral view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341650" httpUri="https://zenodo.org/record/5341650/files/figure.png" pageId="9" pageNumber="100">Fig. 10a</figureCitation>
). Major cheliped (on either left or right side) enlarged, elongate (
<figureCitation id="133844B69F16FFE0FC33F91468D84892" box="[922,1009,1758,1781]" captionStart="Fig" captionStartId="14.[132,167,2057,2078]" captionTargetBox="[139,1461,837,2001]" captionTargetId="figure-49@14.[101,1499,802,2027]" captionTargetPageId="14" captionText="Fig. 11. Leptalpheus dworschaki, new species, male holotype (NMCR 27540): a, major (left) cheliped, mesial view; b, same, lateral view; c, same, chela, carpus and distal merus, ventrolateral view; d, same, distal chela, ventromesial view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341652" httpUri="https://zenodo.org/record/5341652/files/figure.png" pageId="9" pageNumber="100">Fig. 11</figureCitation>
); ischium short, ventromesial margin without subtriangular tooth; merus long, slender, with smooth margins, distally not widening, ventrally flattened, distally more depressed, distal margin with blunt lobes; carpus short, cup-shaped, with blunt distal process; chela subcylindrical, palm smooth, ventromesially excavated, about 2.5 times as long as high (
<figureCitation id="133844B69F16FFE0FC13F8546F3C49D2" box="[954,1045,1950,1973]" captionStart="Fig" captionStartId="14.[132,167,2057,2078]" captionTargetBox="[139,1461,837,2001]" captionTargetId="figure-49@14.[101,1499,802,2027]" captionTargetPageId="14" captionText="Fig. 11. Leptalpheus dworschaki, new species, male holotype (NMCR 27540): a, major (left) cheliped, mesial view; b, same, lateral view; c, same, chela, carpus and distal merus, ventrolateral view; d, same, distal chela, ventromesial view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341652" httpUri="https://zenodo.org/record/5341652/files/figure.png" pageId="9" pageNumber="100">Fig. 11c</figureCitation>
); adhesive discs well developed (
<figureCitation id="133844B69F16FFE0FAD5F854684049B2" captionStart="Fig" captionStartId="14.[132,167,2057,2078]" captionTargetBox="[139,1461,837,2001]" captionTargetId="figure-49@14.[101,1499,802,2027]" captionTargetPageId="14" captionText="Fig. 11. Leptalpheus dworschaki, new species, male holotype (NMCR 27540): a, major (left) cheliped, mesial view; b, same, lateral view; c, same, chela, carpus and distal merus, ventrolateral view; d, same, distal chela, ventromesial view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341652" httpUri="https://zenodo.org/record/5341652/files/figure.png" pageId="9" pageNumber="100">Fig. 11b, c</figureCitation>
); fingers about 1/3 length of palm; dactylus moderately curved distally, with subacute tip, cutting edge with one median subtriangular-rounded tooth, about 1/2 as high as long, without proximal tooth; pollex shorter than dactylus, abruptly ending, with one blunt terminal tooth, cutting edge with two teeth: one much larger subtriangular proximal tooth and one smaller truncate distal tooth (
<figureCitation id="133844B69F14FFE2FDBCFF3669BB4F74" box="[533,658,252,275]" captionStart="Fig" captionStartId="14.[132,167,2057,2078]" captionTargetBox="[139,1461,837,2001]" captionTargetId="figure-49@14.[101,1499,802,2027]" captionTargetPageId="14" captionText="Fig. 11. Leptalpheus dworschaki, new species, male holotype (NMCR 27540): a, major (left) cheliped, mesial view; b, same, lateral view; c, same, chela, carpus and distal merus, ventrolateral view; d, same, distal chela, ventromesial view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341652" httpUri="https://zenodo.org/record/5341652/files/figure.png" pageId="11" pageNumber="102">Fig. 11c, d</figureCitation>
). Minor cheliped shorter and weaker than major cheliped (
<figureCitation id="133844B69F14FFE2FD2AFED669F64F54" box="[643,735,284,307]" captionStart="Fig" captionStartId="15.[109,144,2031,2052]" captionTargetBox="[131,1426,195,1973]" captionTargetId="figure-61@15.[101,1455,149,2005]" captionTargetPageId="15" captionText="Fig. 12. Leptalpheus dworschaki, new species, female paratype (ZRC 2005.0085): a, minor (right) cheliped, lateral view; b, same, chela, mesial view; male holotype (NMCR 27540): c, second pereiopod, lateral view; d, third pereiopod, lateral view; e, fourth pereiopod, lateral view; f, fifth pereiopod, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341654" httpUri="https://zenodo.org/record/5341654/files/figure.png" pageId="11" pageNumber="102">Fig. 12a</figureCitation>
); ischium short, unarmed; merus slender, ventrally depressed; carpus very short, cup-shaped; chela smooth, subcylindrical, with fingers about 1.2 times as long as palm, tips crossing when chela closed; cutting edges of dactylus and pollex with small curved teeth proximally and two larger opposing teeth at about mid-length (
<figureCitation id="133844B69F14FFE2FEFAFE166A9B4F94" box="[339,434,476,499]" captionStart="Fig" captionStartId="15.[109,144,2031,2052]" captionTargetBox="[131,1426,195,1973]" captionTargetId="figure-61@15.[101,1455,149,2005]" captionTargetPageId="15" captionText="Fig. 12. Leptalpheus dworschaki, new species, female paratype (ZRC 2005.0085): a, minor (right) cheliped, lateral view; b, same, chela, mesial view; male holotype (NMCR 27540): c, second pereiopod, lateral view; d, third pereiopod, lateral view; e, fourth pereiopod, lateral view; f, fifth pereiopod, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341654" httpUri="https://zenodo.org/record/5341654/files/figure.png" pageId="11" pageNumber="102">Fig. 12b</figureCitation>
).
</paragraph>
<caption id="DF7C08BB9F15FFE3FF2DF8796819465C" ID-DOI="http://doi.org/10.5281/zenodo.5341644" ID-Zenodo-Dep="5341644" httpUri="https://zenodo.org/record/5341644/files/figure.png" pageId="10" pageNumber="101" startId="10.[132,167,1971,1992]" targetBox="[144,1460,227,1901]" targetPageId="10">
<paragraph id="8BBC58339F15FFE3FF2DF8796819465C" blockId="10.[132,1468,1971,2107]" pageId="10" pageNumber="101">
Fig. 7.
<taxonomicName id="4C0323B09F15FFE3FF62F8796A9449AF" authority="Anker &amp; Marin, 2009" authorityName="Anker &amp; Marin" authorityYear="2009" box="[203,445,1971,1992]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="denticulatus" status="sp. nov.">
<emphasis id="B97784219F15FFE3FF62F8796A9449AF" box="[203,445,1971,1992]" italics="true" pageId="10" pageNumber="101">Leptalpheus denticulatus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A244395A9F15FFE3FE62F879696949AF" box="[459,576,1971,1992]" pageId="10" pageNumber="101" rank="species">new species</taxonomicNameLabel>
(a, b), male from Panglao, Philippines (ZRC 2005.0086): a, frontal region, dorsal view; b, major (left) cheliped, carpus and chela, lateral view.
<taxonomicName id="4C0323B09F15FFE3FD39F81A6F7E4982" authority="Banner &amp; Banner, 1974" authorityName="Banner &amp; Banner" authorityYear="1974" box="[656,1111,2000,2021]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B97784219F15FFE3FD39F81A684B4982" box="[656,866,2000,2021]" italics="true" pageId="10" pageNumber="101">Leptalpheus pacificus</emphasis>
<bibRefCitation id="EF9225C29F15FFE3FCC2F81A6F7E4982" author="Banner, A" box="[875,1111,2000,2021]" pageId="10" pageNumber="101" pagination="423 - 437" refId="ref7616" refString="Banner, A. H. &amp; D. M Banner, 1974. Contributions to the knowledge of the alpheid shrimp of the Pacific Ocean, part XVII: additional notes on the Hawaiian alpheids: new species, subspecies and some nomenclatorial changes. Pacific Science, 28: 423 - 437." type="journal article" year="1974">Banner &amp; Banner, 1974</bibRefCitation>
</taxonomicName>
(ch), female from Hawaii (USNM 205907): c, major (right) cheliped, ventrolateral view; d, same, detail of fingers; breeding female from Hawaii (FLMNH UF Arthropoda 12473): e, frontal region, dorsal view; f, second pereiopod, lateral view; g, right uropod and telson, dorsolateral view, setae omitted; h, left uropod, detail of protopod. Scale bars: ag = 1 mm; h = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BBC58339F14FFE2FFC4FDD66A4A4A34" blockId="11.[109,749,540,1107]" pageId="11" pageNumber="102">
Second pereiopod slender; ischium about 3/4 length of merus; carpus five-segmented, segments with ratio approximately equal to: 4/1/1/1/2.5 (
<figureCitation id="133844B69F14FFE2FEC5FD966AF94C14" box="[364,464,604,627]" captionStart="Fig" captionStartId="15.[109,144,2031,2052]" captionTargetBox="[131,1426,195,1973]" captionTargetId="figure-61@15.[101,1455,149,2005]" captionTargetPageId="15" captionText="Fig. 12. Leptalpheus dworschaki, new species, female paratype (ZRC 2005.0085): a, minor (right) cheliped, lateral view; b, same, chela, mesial view; male holotype (NMCR 27540): c, second pereiopod, lateral view; d, third pereiopod, lateral view; e, fourth pereiopod, lateral view; f, fifth pereiopod, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341654" httpUri="https://zenodo.org/record/5341654/files/figure.png" pageId="11" pageNumber="102">Fig. 12c</figureCitation>
); chela simple, about as long as first carpal segment; fingers as long as palm. Third pereiopod moderately slender (
<figureCitation id="133844B69F14FFE2FE13FD5669314CD4" box="[442,536,668,691]" captionStart="Fig" captionStartId="15.[109,144,2031,2052]" captionTargetBox="[131,1426,195,1973]" captionTargetId="figure-61@15.[101,1455,149,2005]" captionTargetPageId="15" captionText="Fig. 12. Leptalpheus dworschaki, new species, female paratype (ZRC 2005.0085): a, minor (right) cheliped, lateral view; b, same, chela, mesial view; male holotype (NMCR 27540): c, second pereiopod, lateral view; d, third pereiopod, lateral view; e, fourth pereiopod, lateral view; f, fifth pereiopod, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341654" httpUri="https://zenodo.org/record/5341654/files/figure.png" pageId="11" pageNumber="102">Fig. 12d</figureCitation>
); ischium unarmed; merus flattened mesially, about 2.5 times as long as ischium, about five times as long as wide; carpus less than 1/2 length of merus, with distoventral spiniform seta; propodus longer than carpus, with three ventral spiniform setae and one distoventral spinifom seta proximal to dactylus; dactylus simple, conical, about 2/5 length of propodus, curved (
<figureCitation id="133844B69F14FFE2FD68FC966BB04DF4" captionStart="Fig" captionStartId="15.[109,144,2031,2052]" captionTargetBox="[131,1426,195,1973]" captionTargetId="figure-61@15.[101,1455,149,2005]" captionTargetPageId="15" captionText="Fig. 12. Leptalpheus dworschaki, new species, female paratype (ZRC 2005.0085): a, minor (right) cheliped, lateral view; b, same, chela, mesial view; male holotype (NMCR 27540): c, second pereiopod, lateral view; d, third pereiopod, lateral view; e, fourth pereiopod, lateral view; f, fifth pereiopod, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341654" httpUri="https://zenodo.org/record/5341654/files/figure.png" pageId="11" pageNumber="102">Fig. 12d</figureCitation>
). Fourth pereiopod (
<figureCitation id="133844B69F14FFE2FEDAFCB66AE44DF4" box="[371,461,892,915]" captionStart="Fig" captionStartId="15.[109,144,2031,2052]" captionTargetBox="[131,1426,195,1973]" captionTargetId="figure-61@15.[101,1455,149,2005]" captionTargetPageId="15" captionText="Fig. 12. Leptalpheus dworschaki, new species, female paratype (ZRC 2005.0085): a, minor (right) cheliped, lateral view; b, same, chela, mesial view; male holotype (NMCR 27540): c, second pereiopod, lateral view; d, third pereiopod, lateral view; e, fourth pereiopod, lateral view; f, fifth pereiopod, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341654" httpUri="https://zenodo.org/record/5341654/files/figure.png" pageId="11" pageNumber="102">Fig. 12e</figureCitation>
) generally similar to third. Fifth pereiopod (
<figureCitation id="133844B69F14FFE2FE88FC566A5E4DD4" box="[289,375,924,947]" captionStart="Fig" captionStartId="15.[109,144,2031,2052]" captionTargetBox="[131,1426,195,1973]" captionTargetId="figure-61@15.[101,1455,149,2005]" captionTargetPageId="15" captionText="Fig. 12. Leptalpheus dworschaki, new species, female paratype (ZRC 2005.0085): a, minor (right) cheliped, lateral view; b, same, chela, mesial view; male holotype (NMCR 27540): c, second pereiopod, lateral view; d, third pereiopod, lateral view; e, fourth pereiopod, lateral view; f, fifth pereiopod, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341654" httpUri="https://zenodo.org/record/5341654/files/figure.png" pageId="11" pageNumber="102">Fig. 12f</figureCitation>
) much more slender than third and fourth pereiopods; ischium and merus not flattened mesially, unarmed; carpus without distal spiniform seta; propodus as long as merus, without spiniform setae, distally with at least seven rows of setae; dactylus similar to that of third and fourth pereiopods.
</paragraph>
<paragraph id="8BBC58339F14FFE2FFC4FBB66F414F54" blockId="11.[109,748,1148,1267]" lastBlockId="11.[807,1448,188,307]" pageId="11" pageNumber="102">
First to fifth pleomeres with minute pits on surface; posteroventral angles rounded; sixth pleonite with large articulated plate posteroventrally (
<figureCitation id="133844B69F14FFE2FE4FFB76696D4AB4" box="[486,580,1212,1235]" captionStart="Fig" captionStartId="13.[109,144,2030,2051]" captionTargetBox="[112,1445,292,1926]" captionTargetId="figure-64@13.[78,1476,258,1959]" captionTargetPageId="13" captionText="Fig. 10. Leptalpheus dworschaki, new species, female paratype (ZRC 2005.0085): a, habitus; male holotype (NMCR 27540): b, mandible, mesial view; c, same, dorsolateral view; d, maxillule, lateral view; e, maxilla, lateral view; f, first maxilliped, lateral view; g, second maxilliped, lateral view; h, third maxilliped, lateral view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341650" httpUri="https://zenodo.org/record/5341650/files/figure.png" pageId="11" pageNumber="102">Fig. 10a</figureCitation>
). First pleopod woth very short endopod, latter fringed with setae apically (
<figureCitation id="133844B69F14FFE2FC86FF7668AB4EB4" box="[815,898,188,211]" captionStart="Fig" captionStartId="12.[132,167,2002,2023]" captionTargetBox="[143,1460,196,1957]" captionTargetId="figure-92@12.[113,1491,165,1979]" captionTargetPageId="12" captionText="Fig. 9. Leptalpheus dworschaki, new species, male holotype (NMCR 27540): a, frontal region, dorsal view; b, same, lateral view; c, frontal margin of carapace, lateral view; d, first segment of right antennular peduncle, lateral view; e, lateral flagellum of right antennule, lateral view; f, first pleopod, mesial view; g, second pleopod, mesial view; h, same, appendix masculina and appendix interna; i, right uropod, dorsal view; j, telson, dorsal view; female paratype (ZRC 2005.0085): k, frontal margin, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341648" httpUri="https://zenodo.org/record/5341648/files/figure.png" pageId="11" pageNumber="102">Fig. 9f</figureCitation>
). Male second pleopod (
<figureCitation id="133844B69F14FFE2FB09FF766FDA4EB4" box="[1184,1267,188,211]" captionStart="Fig" captionStartId="12.[132,167,2002,2023]" captionTargetBox="[143,1460,196,1957]" captionTargetId="figure-92@12.[113,1491,165,1979]" captionTargetPageId="12" captionText="Fig. 9. Leptalpheus dworschaki, new species, male holotype (NMCR 27540): a, frontal region, dorsal view; b, same, lateral view; c, frontal margin of carapace, lateral view; d, first segment of right antennular peduncle, lateral view; e, lateral flagellum of right antennule, lateral view; f, first pleopod, mesial view; g, second pleopod, mesial view; h, same, appendix masculina and appendix interna; i, right uropod, dorsal view; j, telson, dorsal view; female paratype (ZRC 2005.0085): k, frontal margin, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341648" httpUri="https://zenodo.org/record/5341648/files/figure.png" pageId="11" pageNumber="102">
Fig.
<quantity id="4CFBF5D69F14FFE2FB7EFF766FDA4EB4" box="[1239,1267,188,211]" metricMagnitude="-3" metricUnit="kg" metricValue="9.0" pageId="11" pageNumber="102" unit="g" value="9.0">9g</quantity>
</figureCitation>
) with appendix interna and appendix masculina, latter almost twice as long as former, and bearing numerous slender, spiniform apical and subapical setae (
<figureCitation id="133844B69F14FFE2FBA3FED66F754F54" box="[1034,1116,284,307]" captionStart="Fig" captionStartId="12.[132,167,2002,2023]" captionTargetBox="[143,1460,196,1957]" captionTargetId="figure-92@12.[113,1491,165,1979]" captionTargetPageId="12" captionText="Fig. 9. Leptalpheus dworschaki, new species, male holotype (NMCR 27540): a, frontal region, dorsal view; b, same, lateral view; c, frontal margin of carapace, lateral view; d, first segment of right antennular peduncle, lateral view; e, lateral flagellum of right antennule, lateral view; f, first pleopod, mesial view; g, second pleopod, mesial view; h, same, appendix masculina and appendix interna; i, right uropod, dorsal view; j, telson, dorsal view; female paratype (ZRC 2005.0085): k, frontal margin, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341648" httpUri="https://zenodo.org/record/5341648/files/figure.png" pageId="11" pageNumber="102">Fig. 9h</figureCitation>
).
</paragraph>
<paragraph id="8BBC58339F14FFE2FC8EFE966F034C34" blockId="11.[807,1447,348,595]" pageId="11" pageNumber="102">
Uropod with lateral lobe of protopod bearing two small acute teeth distally (
<figureCitation id="133844B69F14FFE2FBBAFEB66F4A4FF4" box="[1043,1123,380,403]" captionStart="Fig" captionStartId="12.[132,167,2002,2023]" captionTargetBox="[143,1460,196,1957]" captionTargetId="figure-92@12.[113,1491,165,1979]" captionTargetPageId="12" captionText="Fig. 9. Leptalpheus dworschaki, new species, male holotype (NMCR 27540): a, frontal region, dorsal view; b, same, lateral view; c, frontal margin of carapace, lateral view; d, first segment of right antennular peduncle, lateral view; e, lateral flagellum of right antennule, lateral view; f, first pleopod, mesial view; g, second pleopod, mesial view; h, same, appendix masculina and appendix interna; i, right uropod, dorsal view; j, telson, dorsal view; female paratype (ZRC 2005.0085): k, frontal margin, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341648" httpUri="https://zenodo.org/record/5341648/files/figure.png" pageId="11" pageNumber="102">Fig. 9i</figureCitation>
); endopod somewhat longer than exopod, without specific features; exopod with truncate posterior margin and without distinct distolateral tooth adjacent to distolateral spine; lateral portion of diaeresis straight, not curved into tooth centrally; mesial portion deeply incised forming large triangular tooth proximal to mesial margin (
<figureCitation id="133844B69F14FFE2FC7BFDF66F344C34" box="[978,1053,572,595]" captionStart="Fig" captionStartId="12.[132,167,2002,2023]" captionTargetBox="[143,1460,196,1957]" captionTargetId="figure-92@12.[113,1491,165,1979]" captionTargetPageId="12" captionText="Fig. 9. Leptalpheus dworschaki, new species, male holotype (NMCR 27540): a, frontal region, dorsal view; b, same, lateral view; c, frontal margin of carapace, lateral view; d, first segment of right antennular peduncle, lateral view; e, lateral flagellum of right antennule, lateral view; f, first pleopod, mesial view; g, second pleopod, mesial view; h, same, appendix masculina and appendix interna; i, right uropod, dorsal view; j, telson, dorsal view; female paratype (ZRC 2005.0085): k, frontal margin, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341648" httpUri="https://zenodo.org/record/5341648/files/figure.png" pageId="11" pageNumber="102">Fig. 9i</figureCitation>
).
</paragraph>
<paragraph id="8BBC58339F14FFE2FC8EFDB66E544D34" blockId="11.[807,1448,636,851]" pageId="11" pageNumber="102">
Telson moderately slender, more than twice as long as wide proximally (
<figureCitation id="133844B69F14FFE2FC02FD5668DC4CD4" box="[939,1013,668,691]" captionStart="Fig" captionStartId="12.[132,167,2002,2023]" captionTargetBox="[143,1460,196,1957]" captionTargetId="figure-92@12.[113,1491,165,1979]" captionTargetPageId="12" captionText="Fig. 9. Leptalpheus dworschaki, new species, male holotype (NMCR 27540): a, frontal region, dorsal view; b, same, lateral view; c, frontal margin of carapace, lateral view; d, first segment of right antennular peduncle, lateral view; e, lateral flagellum of right antennule, lateral view; f, first pleopod, mesial view; g, second pleopod, mesial view; h, same, appendix masculina and appendix interna; i, right uropod, dorsal view; j, telson, dorsal view; female paratype (ZRC 2005.0085): k, frontal margin, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341648" httpUri="https://zenodo.org/record/5341648/files/figure.png" pageId="11" pageNumber="102">Fig. 9j</figureCitation>
); dorsal surface pitted, with two pairs of robust spiniform setae inserted at short distance from lateral margin, at about 1/3 and 2/3 length of telson, respectively; posterior margin feebly rounded, posterolateral angles each with pair of spiniform setae, mesial at least four times as long as lateral; anal tubercles small, feebly sclerotised.
</paragraph>
<paragraph id="8BBC58339F14FFE2FC8EFCB66EAF4DF4" blockId="11.[807,1414,892,915]" box="[807,1414,892,915]" pageId="11" pageNumber="102">Gill/exopod formula as in previous species (see above).</paragraph>
<paragraph id="8BBC58339F14FFE2FC8EFC7668D34D94" blockId="11.[807,1448,956,1011]" pageId="11" pageNumber="102">
<emphasis id="B97784219F14FFE2FC8EFC7668F74DB3" bold="true" box="[807,990,956,980]" pageId="11" pageNumber="102">
<emphasis id="B97784219F14FFE2FC8EFC7668E54DB3" bold="true" box="[807,972,956,980]" italics="true" pageId="11" pageNumber="102">Colour pattern.</emphasis>
</emphasis>
Field notes indicated: “Translucent, slightly pink, ovary green”.
</paragraph>
<paragraph id="8BBC58339F14FFE2FC8EFBD668F54A14" blockId="11.[807,1448,1052,1139]" pageId="11" pageNumber="102">
<emphasis id="B97784219F14FFE2FC8EFBD668474A53" bold="true" box="[807,878,1052,1076]" pageId="11" pageNumber="102">
<emphasis id="B97784219F14FFE2FC8EFBD6687A4A53" bold="true" box="[807,851,1052,1076]" italics="true" pageId="11" pageNumber="102">Size</emphasis>
.
</emphasis>
The CL of both specimens is
<quantity id="4CFBF5D69F14FFE2FB1BFBD66E2E4A54" box="[1202,1287,1052,1075]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="11" pageNumber="102" unit="mm" value="7.7">7.7 mm</quantity>
; the TL ranges from
<quantity id="4CFBF5D69F14FFE2FCC1FBF668E74A34" box="[872,974,1084,1107]" metricMagnitude="-2" metricUnit="m" metricValue="2.19" pageId="11" pageNumber="102" unit="mm" value="21.9">21.9 mm</quantity>
in the male
<typeStatus id="54B8E6919F14FFE2FBCEFBF66FE14A34" box="[1127,1224,1084,1107]" pageId="11" pageNumber="102" type="holotype">holotype</typeStatus>
to
<quantity id="4CFBF5D69F14FFE2FB5DFBF66E734A34" box="[1268,1370,1084,1107]" metricMagnitude="-2" metricUnit="m" metricValue="2.3899999999999997" pageId="11" pageNumber="102" unit="mm" value="23.9">23.9 mm</quantity>
in the female
<typeStatus id="54B8E6919F14FFE2FCD1FB9668FE4A14" box="[888,983,1116,1139]" pageId="11" pageNumber="102" type="paratype">paratype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C3190BB89F14FFE2FC8EFB5668C24B54" pageId="11" pageNumber="102" type="etymology">
<paragraph id="8BBC58339F14FFE2FC8EFB5668C24B54" blockId="11.[807,1447,1180,1331]" pageId="11" pageNumber="102">
<emphasis id="B97784219F14FFE2FC8EFB5668E84AD3" bold="true" box="[807,961,1180,1204]" pageId="11" pageNumber="102">
<emphasis id="B97784219F14FFE2FC8EFB5668884AD3" bold="true" box="[807,929,1180,1204]" italics="true" pageId="11" pageNumber="102">Etymology</emphasis>
.
</emphasis>
The new species is named after Peter C. Dworschak (Naturhistorisches Museum in
<collectingRegion id="49C796D19F14FFE2FAAEFB766E6D4AB4" box="[1287,1348,1212,1235]" country="Austria" name="Wien" pageId="11" pageNumber="102">Wien</collectingRegion>
,
<collectingRegion id="49C796D19F14FFE2FAFBFB766E8A4AB4" box="[1362,1443,1212,1235]" country="Austria" name="Wien" pageId="11" pageNumber="102">Vienna</collectingRegion>
,
<collectingCountry id="F31418A39F14FFE2FC8EFB1668504A94" box="[807,889,1244,1267]" name="Austria" pageId="11" pageNumber="102">Austria</collectingCountry>
), the collector of the present specimens, our estimated friend and colleague, and also a well known specialist of the Thalassinidea.
</paragraph>
</subSubSection>
<subSubSection id="C3190BB89F14FFE2FC8EFA96688E4BF4" pageId="11" pageNumber="102" type="materials_examined">
<paragraph id="8BBC58339F14FFE2FC8EFA96688E4BF4" blockId="11.[807,1447,1372,1427]" pageId="11" pageNumber="102">
<materialsCitation id="3B6B526E9F14FFE2FC8EFA96688A4BF4" country="Philippines" location="Panglao Island" pageId="11" pageNumber="102" specimenCount="1" stateProvince="Bohol" typeStatus="holotype">
<emphasis id="B97784219F14FFE2FC8EFA9668FE4B13" bold="true" box="[807,983,1372,1396]" pageId="11" pageNumber="102">
<emphasis id="B97784219F14FFE2FC8EFA9668964B13" bold="true" box="[807,959,1372,1396]" italics="true" pageId="11" pageNumber="102">
<typeStatus id="54B8E6919F14FFE2FC8EFA9668754B13" box="[807,860,1372,1396]" pageId="11" pageNumber="102">Type</typeStatus>
locality.
</emphasis>
</emphasis>
<location id="8EDC0EE89F14FFE2FC4BFA966FA34B14" LSID="urn:lsid:plazi:treatment:03AAE9259F16FFE7FC3FFEB16BC34CB4:8EDC0EE89F14FFE2FC4BFA966FA34B14" box="[994,1162,1372,1395]" country="Philippines" name="Panglao Island" pageId="11" pageNumber="102" stateProvince="Bohol">Panglao Island</location>
, southwest of
<collectingRegion id="49C796D19F14FFE2FA9BFA966E5E4B14" box="[1330,1399,1372,1395]" country="Philippines" name="Bohol" pageId="11" pageNumber="102">Bohol</collectingRegion>
, the
<collectingCountry id="F31418A39F14FFE2FC8EFAB6688A4BF4" box="[807,931,1404,1427]" name="Philippines" pageId="11" pageNumber="102">Philippines</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3190BB89F14FFE2FC8EFA766F7C4854" pageId="11" pageNumber="102" type="biology_ecology">
<paragraph id="8BBC58339F14FFE2FC8EFA766F7C4854" blockId="11.[807,1448,1468,1587]" pageId="11" pageNumber="102">
<emphasis id="B97784219F14FFE2FC8EFA7668BC4BB3" bold="true" box="[807,917,1468,1492]" pageId="11" pageNumber="102">
<emphasis id="B97784219F14FFE2FC8EFA7668544BB3" bold="true" box="[807,893,1468,1492]" italics="true" pageId="11" pageNumber="102">Ecology</emphasis>
.
</emphasis>
Both specimens were collected from a
<taxonomicName id="4C0323B09F14FFE2FA88FA766EA04BB4" authorityName="Stimpson" authorityYear="1866" box="[1313,1417,1468,1491]" class="Malacostraca" family="Callianassidae" genus="Glypturus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="102" phylum="Arthropoda" rank="genus">
<emphasis id="B97784219F14FFE2FA88FA766EA04BB4" box="[1313,1417,1468,1491]" italics="true" pageId="11" pageNumber="102">Glypturus</emphasis>
</taxonomicName>
cf.
<emphasis id="B97784219F14FFE2FC8EFA1668544B94" box="[807,893,1500,1523]" italics="true" pageId="11" pageNumber="102">armatus</emphasis>
burrow, together with gastropods (
<taxonomicName id="4C0323B09F14FFE2FB48FA166E894B94" authorityName="Pilsbry" authorityYear="1895" box="[1249,1440,1500,1523]" class="Gastropoda" family="Phenacolepadidae" kingdom="Animalia" order="Cycloneritida" pageId="11" pageNumber="102" phylum="Mollusca" rank="family">Phenacolepadidae</taxonomicName>
) and sentinel crabs,
<taxonomicName id="4C0323B09F14FFE2FC5DFA366E414874" box="[1012,1384,1532,1555]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="102" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B97784219F14FFE2FC5DFA366F804874" box="[1012,1193,1532,1555]" italics="true" pageId="11" pageNumber="102">Macrophthalmus</emphasis>
sp. (Ocypodidae)
</taxonomicName>
[field notes by P. C. Dworschak].
</paragraph>
</subSubSection>
<subSubSection id="C3190BB89F14FFE2FC8EF9966F8F48F4" pageId="11" pageNumber="102" type="distribution">
<paragraph id="8BBC58339F14FFE2FC8EF9966F8F48F4" blockId="11.[807,1447,1628,1683]" pageId="11" pageNumber="102">
<emphasis id="B97784219F14FFE2FC8EF99668EE4813" bold="true" box="[807,967,1628,1652]" pageId="11" pageNumber="102">
<emphasis id="B97784219F14FFE2FC8EF99668854813" bold="true" box="[807,940,1628,1652]" italics="true" pageId="11" pageNumber="102">Distribution</emphasis>
.
</emphasis>
Western Pacific: presently known only from the
<typeStatus id="54B8E6919F14FFE2FCF9F9B6685748F4" box="[848,894,1660,1683]" pageId="11" pageNumber="102">type</typeStatus>
locality in the
<collectingCountry id="F31418A39F14FFE2FB8FF9B66F8B48F4" box="[1062,1186,1660,1683]" name="Philippines" pageId="11" pageNumber="102">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3190BB89F14FFE7FC8EF9766BC34CB4" lastPageId="14" lastPageNumber="105" pageId="11" pageNumber="102" type="discussion">
<paragraph id="8BBC58339F14FFE7FC8EF9766A944FF4" blockId="11.[807,1448,1724,2099]" lastBlockId="14.[132,771,188,403]" lastPageId="14" lastPageNumber="105" pageId="11" pageNumber="102">
<emphasis id="B97784219F14FFE2FC8EF9766F8548B4" box="[807,1196,1724,1748]" italics="true" pageId="11" pageNumber="102">
<emphasis id="B97784219F14FFE2FC8EF976688A48B3" bold="true" box="[807,931,1724,1748]" italics="true" pageId="11" pageNumber="102">Remarks. </emphasis>
<taxonomicName id="4C0323B09F14FFE2FC00F9766F8548B4" authority="Anker &amp; Marin, 2009" authorityName="Anker &amp; Marin" authorityYear="2009" box="[937,1196,1724,1747]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="102" phylum="Arthropoda" rank="species" species="dworschaki" status="sp. nov.">Leptalpheus dworschaki</taxonomicName>
</emphasis>
,
<taxonomicNameLabel id="A244395A9F14FFE2FB11F9766E1248B4" box="[1208,1339,1724,1747]" pageId="11" pageNumber="102" rank="species">new species</taxonomicNameLabel>
, is closely related to
<taxonomicName id="4C0323B09F14FFE2FC25F9166F2A4894" authorityName="Banner &amp; Banner" authorityYear="1974" box="[908,1027,1756,1779]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="102" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B97784219F14FFE2FC25F9166F2A4894" box="[908,1027,1756,1779]" italics="true" pageId="11" pageNumber="102">L. pacificus</emphasis>
</taxonomicName>
and the above-described
<taxonomicName id="4C0323B09F14FFE2FAAEF9166E894894" authority="Anker &amp; Marin, 2009" authorityName="Anker &amp; Marin" authorityYear="2009" box="[1287,1440,1756,1779]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="102" phylum="Arthropoda" rank="species" species="denticulatus" status="sp. nov.">
<emphasis id="B97784219F14FFE2FAAEF9166E894894" box="[1287,1440,1756,1779]" italics="true" pageId="11" pageNumber="102">L. denticulatus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A244395A9F14FFE2FC8EF93668854974" box="[807,940,1788,1811]" pageId="11" pageNumber="102" rank="species">new species</taxonomicNameLabel>
, but can be separated from both species by the armature on the major chela fingers, in particular by the presence of a distinct blunt tooth on the dactylus (absent in
<taxonomicName id="4C0323B09F14FFE2FC8EF89668884914" authorityName="Banner &amp; Banner" authorityYear="1974" box="[807,929,1884,1907]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="102" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B97784219F14FFE2FC8EF89668884914" box="[807,929,1884,1907]" italics="true" pageId="11" pageNumber="102">L. pacificus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0323B09F14FFE2FC7BF8966F464914" authority="Anker &amp; Marin, 2009" authorityName="Anker &amp; Marin" authorityYear="2009" box="[978,1135,1884,1907]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="102" phylum="Arthropoda" rank="species" species="denticulatus" status="sp. nov.">
<emphasis id="B97784219F14FFE2FC7BF8966F464914" box="[978,1135,1884,1907]" italics="true" pageId="11" pageNumber="102">L. denticulatus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A244395A9F14FFE2FBD2F8966FD44914" box="[1147,1277,1884,1907]" pageId="11" pageNumber="102" rank="species">new species</taxonomicNameLabel>
, although larger individuals may have a broad bulge at the same position); the absence of a small proximal tooth on the dactylus (present in
<taxonomicName id="4C0323B09F14FFE2FCEFF87668EF49B4" authorityName="Banner &amp; Banner" authorityYear="1974" box="[838,966,1980,2003]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="102" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B97784219F14FFE2FCEFF87668EF49B4" box="[838,966,1980,2003]" italics="true" pageId="11" pageNumber="102">L. pacificus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0323B09F14FFE2FBA8F8766F8C49B4" authority="Anker &amp; Marin, 2009" authorityName="Anker &amp; Marin" authorityYear="2009" box="[1025,1189,1980,2003]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="102" phylum="Arthropoda" rank="species" species="denticulatus" status="sp. nov.">
<emphasis id="B97784219F14FFE2FBA8F8766F8C49B4" box="[1025,1189,1980,2003]" italics="true" pageId="11" pageNumber="102">L. denticulatus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A244395A9F14FFE2FB1FF8766E6949B4" box="[1206,1344,1980,2003]" pageId="11" pageNumber="102" rank="species">new species</taxonomicNameLabel>
); and the shorter hiatus between the larger proximal and smaller distal teeth on the pollex (this hiatus is almost twice as long in
<taxonomicName id="4C0323B09F14FFE2FC8EF7D668834654" authorityName="Banner &amp; Banner" authorityYear="1974" box="[807,938,2076,2099]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="102" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B97784219F14FFE2FC8EF7D668834654" box="[807,938,2076,2099]" italics="true" pageId="11" pageNumber="102">L. pacificus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0323B09F14FFE2FC41F7D66FB84654" authority="Anker &amp; Marin, 2009" authorityName="Anker &amp; Marin" authorityYear="2009" box="[1000,1169,2076,2099]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="102" phylum="Arthropoda" rank="species" species="denticulatus" status="sp. nov.">
<emphasis id="B97784219F14FFE2FC41F7D66FB84654" box="[1000,1169,2076,2099]" italics="true" pageId="11" pageNumber="102">L. denticulatus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A244395A9F14FFE2FB0AF7D66E044654" box="[1187,1325,2076,2099]" pageId="11" pageNumber="102" rank="species">new species</taxonomicNameLabel>
) (compare
<figureCitation id="133844B69F11FFE7FF2DFF766A224EB4" box="[132,267,188,211]" captionStart="Fig" captionStartId="14.[132,167,2057,2078]" captionTargetBox="[139,1461,837,2001]" captionTargetId="figure-49@14.[101,1499,802,2027]" captionTargetPageId="14" captionText="Fig. 11. Leptalpheus dworschaki, new species, male holotype (NMCR 27540): a, major (left) cheliped, mesial view; b, same, lateral view; c, same, chela, carpus and distal merus, ventrolateral view; d, same, distal chela, ventromesial view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341652" httpUri="https://zenodo.org/record/5341652/files/figure.png" pageId="14" pageNumber="105">Figs. 11c, d</figureCitation>
and
<figureCitation id="133844B69F11FFE7FEE1FF766AED4EB4" box="[328,452,188,211]" captionStart="Fig" captionStartId="5.[109,144,2030,2051]" captionTargetBox="[109,1446,278,1916]" captionTargetId="figure-68@5.[97,1458,266,1929]" captionTargetPageId="5" captionText="Fig. 3. Leptalpheus denticulatus, new species, female paratype from Nhatrang Bay, Vietnam (USNM 1110002): a, major (left) cheliped, lateral view; b, same, mesial view; c, same, chela, carpus and distal merus, ventrolateral view; d, same, distal portion of chela, mesial view; e, minor (right) cheliped, lateral view; f, same, chela, ventrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341636" httpUri="https://zenodo.org/record/5341636/files/figure.png" pageId="14" pageNumber="105">Figs. 3c, d</figureCitation>
;
<figureCitation id="133844B69F11FFE7FE7DFF7669304EB4" box="[468,537,188,211]" captionStart="Fig" captionStartId="6.[132,167,2058,2079]" captionTargetBox="[139,1452,203,1991]" captionTargetId="figure-49@6.[85,1483,171,2023]" captionTargetPageId="6" captionText="Fig. 4. Leptalpheus denticulatus, new species, largest female paratype from Nhatrang Bay, Vietnam (MNHN-Na 17065): a, major (left) cheliped, distal merus, carpus and chela, ventrolateral view; b, same, detail of fingers; c, same, ventromesial view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341638" httpUri="https://zenodo.org/record/5341638/files/figure.png" pageId="14" pageNumber="105">4 b, c</figureCitation>
;
<figureCitation id="133844B69F11FFE7FD80FF76694C4EB4" box="[553,613,188,211]" captionStart="Fig" captionStartId="10.[132,167,1971,1992]" captionTargetBox="[144,1460,227,1901]" captionTargetId="figure-104@10.[114,1490,209,1920]" captionTargetPageId="10" captionText="Fig. 7. Leptalpheus denticulatus, new species (a, b), male from Panglao, Philippines (ZRC 2005.0086): a, frontal region, dorsal view; b, major (left) cheliped, carpus and chela, lateral view. Leptalpheus pacificus Banner &amp; Banner, 1974 (ch), female from Hawaii (USNM 205907): c, major (right) cheliped, ventrolateral view; d, same, detail of fingers; breeding female from Hawaii (FLMNH UF Arthropoda 12473): e, frontal region, dorsal view; f, second pereiopod, lateral view; g, right uropod and telson, dorsolateral view, setae omitted; h, left uropod, detail of protopod. Scale bars: ag = 1 mm; h = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5341644" httpUri="https://zenodo.org/record/5341644/files/figure.png" pageId="14" pageNumber="105">7c, d</figureCitation>
).
<taxonomicName id="4C0323B09F11FFE7FDD4FF766A2A4E94" authority="Anker &amp; Marin, 2009" authorityName="Anker &amp; Marin" authorityYear="2009" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="105" phylum="Arthropoda" rank="species" species="dworschaki" status="sp. nov.">
<emphasis id="B97784219F11FFE7FDD4FF766A2A4E94" italics="true" pageId="14" pageNumber="105">Leptalpheus dworschaki</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A244395A9F11FFE7FEBDFF166AB74E94" box="[276,414,220,243]" pageId="14" pageNumber="105" rank="species">new species</taxonomicNameLabel>
, can be also separated from
<taxonomicName id="4C0323B09F11FFE7FD44FF166A2F4F74" authority="Anker &amp; Marin, 2009" authorityName="Anker &amp; Marin" authorityYear="2009" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="105" phylum="Arthropoda" rank="species" species="denticulatus" status="sp. nov.">
<emphasis id="B97784219F11FFE7FD44FF166A2F4F74" italics="true" pageId="14" pageNumber="105">L. denticulatus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A244395A9F11FFE7FEBFFF366AB54F74" box="[278,412,252,275]" pageId="14" pageNumber="105" rank="species">new species</taxonomicNameLabel>
, by the absence of a small tooth on the central portion of the uropodal diaeresis. We were unable to find any other obvious morphological differences between
<taxonomicName id="4C0323B09F11FFE7FF4FFE966AA94F14" authority="Anker &amp; Marin, 2009" authorityName="Anker &amp; Marin" authorityYear="2009" box="[230,384,348,371]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="105" phylum="Arthropoda" rank="species" species="dworschaki" status="sp. nov.">
<emphasis id="B97784219F11FFE7FF4FFE966AA94F14" box="[230,384,348,371]" italics="true" pageId="14" pageNumber="105">L. dworschaki</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A244395A9F11FFE7FE38FE96693E4F14" box="[401,535,348,371]" pageId="14" pageNumber="105" rank="species">new species</taxonomicNameLabel>
, and
<taxonomicName id="4C0323B09F11FFE7FDFCFE9669FA4F14" authorityName="Banner &amp; Banner" authorityYear="1974" box="[597,723,348,371]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="105" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B97784219F11FFE7FDFCFE9669FA4F14" box="[597,723,348,371]" italics="true" pageId="14" pageNumber="105">L. pacificus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0323B09F11FFE7FF2DFEB66A0D4FF4" authority="Anker &amp; Marin, 2009" authorityName="Anker &amp; Marin" authorityYear="2009" box="[132,292,380,403]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="105" phylum="Arthropoda" rank="species" species="denticulatus" status="sp. nov.">
<emphasis id="B97784219F11FFE7FF2DFEB66A0D4FF4" box="[132,292,380,403]" italics="true" pageId="14" pageNumber="105">L. denticulatus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A244395A9F11FFE7FE9DFEB66A904FF4" box="[308,441,380,403]" pageId="14" pageNumber="105" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
<caption id="DF7C08BB9F14FFE2FFC4F8796A38465B" ID-DOI="http://doi.org/10.5281/zenodo.5341646" ID-Zenodo-Dep="5341646" httpUri="https://zenodo.org/record/5341646/files/figure.png" pageId="11" pageNumber="102" startId="11.[109,144,1971,1992]" targetBox="[109,747,1330,1930]" targetPageId="11">
<paragraph id="8BBC58339F14FFE2FFC4F8796A38465B" blockId="11.[109,748,1971,2108]" pageId="11" pageNumber="102">
Fig. 8.
<taxonomicName id="4C0323B09F14FFE2FF11F87E6A8649AE" authority="Anker &amp; Marin, 2009" authorityName="Anker &amp; Marin" authorityYear="2009" box="[184,431,1972,1993]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="102" phylum="Arthropoda" rank="species" species="denticulatus" status="sp. nov.">
<emphasis id="B97784219F14FFE2FF11F87E6A8649AE" box="[184,431,1972,1993]" italics="true" pageId="11" pageNumber="102">Leptalpheus denticulatus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A244395A9F14FFE2FE17F879691049AF" box="[446,569,1971,1992]" pageId="11" pageNumber="102" rank="species">new species</taxonomicNameLabel>
, colour pattern in life: a, non-type male from Yaqara Bay, Viti Levu, Fiji (MNHN- Na 15776); b, large female paratype from Nhatrang Bay, Vietnam (MNHN-Na 17065), major (left) cheliped detached (photograph by O. Savinkin).
</paragraph>
</caption>
<caption id="DF7C08BB9F13FFE5FF2DF8186E18465A" ID-DOI="http://doi.org/10.5281/zenodo.5341648" ID-Zenodo-Dep="5341648" httpUri="https://zenodo.org/record/5341648/files/figure.png" pageId="12" pageNumber="103" startId="12.[132,167,2002,2023]" targetBox="[143,1460,196,1957]" targetPageId="12">
<paragraph id="8BBC58339F13FFE5FF2DF8186E18465A" blockId="12.[132,1470,2002,2109]" pageId="12" pageNumber="103">
Fig. 9.
<taxonomicName id="4C0323B09F13FFE5FF61F8186A9B4980" authority="Anker &amp; Marin, 2009" authorityName="Anker &amp; Marin" authorityYear="2009" box="[200,434,2002,2023]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="103" phylum="Arthropoda" rank="species" species="dworschaki" status="sp. nov.">
<emphasis id="B97784219F13FFE5FF61F8186A9B4980" box="[200,434,2002,2023]" italics="true" pageId="12" pageNumber="103">Leptalpheus dworschaki</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A244395A9F13FFE5FE17F818691D4980" box="[446,564,2002,2023]" pageId="12" pageNumber="103" rank="species">new species</taxonomicNameLabel>
, male holotype (NMCR 27540): a, frontal region, dorsal view; b, same, lateral view; c, frontal margin of carapace, lateral view; d, first segment of right antennular peduncle, lateral view; e, lateral flagellum of right antennule, lateral view; f, first pleopod, mesial view; g, second pleopod, mesial view; h, same, appendix masculina and appendix interna; i, right uropod, dorsal view; j, telson, dorsal view; female paratype (ZRC 2005.0085): k, frontal margin, dorsal view. Scale bars = 1 mm.
</paragraph>
</caption>
<caption id="DF7C08BB9F12FFE4FFC4F8246877465A" ID-DOI="http://doi.org/10.5281/zenodo.5341650" ID-Zenodo-Dep="5341650" httpUri="https://zenodo.org/record/5341650/files/figure.png" pageId="13" pageNumber="104" startId="13.[109,144,2030,2051]" targetBox="[112,1445,292,1926]" targetPageId="13">
<paragraph id="8BBC58339F12FFE4FFC4F8246877465A" blockId="13.[109,1448,2030,2109]" pageId="13" pageNumber="104">
Fig. 10.
<taxonomicName id="4C0323B09F12FFE4FF17F8256A804663" authority="Anker &amp; Marin, 2009" authorityName="Anker &amp; Marin" authorityYear="2009" box="[190,425,2031,2052]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="104" phylum="Arthropoda" rank="species" species="dworschaki" status="sp. nov.">
<emphasis id="B97784219F12FFE4FF17F8256A804663" box="[190,425,2031,2052]" italics="true" pageId="13" pageNumber="104">Leptalpheus dworschaki</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A244395A9F12FFE4FE1FF82469054664" box="[438,556,2030,2051]" pageId="13" pageNumber="104" rank="species">new species</taxonomicNameLabel>
, female paratype (ZRC 2005.0085): a, habitus; male holotype (NMCR 27540): b, mandible, mesial view; c, same, dorsolateral view; d, maxillule, lateral view; e, maxilla, lateral view; f, first maxilliped, lateral view; g, second maxilliped, lateral view; h, third maxilliped, lateral view. Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph id="8BBC58339F11FFE7FF2DFE766BC34CB4" blockId="14.[132,771,444,723]" pageId="14" pageNumber="105">
The length of the second segment of the antennular peduncle and the shape of the frontal margin appear to be somewhat variable in both
<taxonomicName id="4C0323B09F11FFE7FE99FE366AEE4C74" authority="Anker &amp; Marin, 2009" authorityName="Anker &amp; Marin" authorityYear="2009" box="[304,455,508,531]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="105" phylum="Arthropoda" rank="species" species="dworschaki" status="sp. nov.">
<emphasis id="B97784219F11FFE7FE99FE366AEE4C74" box="[304,455,508,531]" italics="true" pageId="14" pageNumber="105">L. dworschaki</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A244395A9F11FFE7FE7DFE36697D4C74" box="[468,596,508,531]" pageId="14" pageNumber="105" rank="species">new species</taxonomicNameLabel>
(
<figureCitation id="133844B69F11FFE7FDCBFE3669E04C74" box="[610,713,508,531]" captionStart="Fig" captionStartId="12.[132,167,2002,2023]" captionTargetBox="[143,1460,196,1957]" captionTargetId="figure-92@12.[113,1491,165,1979]" captionTargetPageId="12" captionText="Fig. 9. Leptalpheus dworschaki, new species, male holotype (NMCR 27540): a, frontal region, dorsal view; b, same, lateral view; c, frontal margin of carapace, lateral view; d, first segment of right antennular peduncle, lateral view; e, lateral flagellum of right antennule, lateral view; f, first pleopod, mesial view; g, second pleopod, mesial view; h, same, appendix masculina and appendix interna; i, right uropod, dorsal view; j, telson, dorsal view; female paratype (ZRC 2005.0085): k, frontal margin, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341648" httpUri="https://zenodo.org/record/5341648/files/figure.png" pageId="14" pageNumber="105">Fig. 9a, k</figureCitation>
), and
<taxonomicName id="4C0323B09F11FFE7FF2DFDD66A0B4C54" authority="Anker &amp; Marin, 2009" authorityName="Anker &amp; Marin" authorityYear="2009" box="[132,290,540,563]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="105" phylum="Arthropoda" rank="species" species="denticulatus" status="sp. nov.">
<emphasis id="B97784219F11FFE7FF2DFDD66A0B4C54" box="[132,290,540,563]" italics="true" pageId="14" pageNumber="105">L. denticulatus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A244395A9F11FFE7FE99FDD66A994C54" box="[304,432,540,563]" pageId="14" pageNumber="105" rank="species">new species</taxonomicNameLabel>
(
<figureCitation id="133844B69F11FFE7FE17FDD6693C4C54" box="[446,533,540,563]" captionStart="Fig" captionStartId="3.[109,144,1973,1994]" captionTargetBox="[113,1447,260,1891]" captionTargetId="figure-10@3.[87,1467,239,1908]" captionTargetPageId="3" captionText="Fig. 1. Leptalpheus denticulatus, new species, female paratype from Nhatrang Bay, Vietnam (USNM 1110002): a, frontal region, dorsal view; b, same, lateral view; c, frontal margin of carapace, lateral view; d, carapace, lateral view; e, right eye, ventrolateral view; f, first segment of right antennular peduncle, lateral view; g, lateral flagellum of right antennule, lateral view; h, right uropod, dorsal view; i, same, detail of exopod, setae omitted; j, telson, dorsal view; k, same, detail of posterior half; l, sixth pleonite and tail fan, setae omitted, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341632" httpUri="https://zenodo.org/record/5341632/files/figure.png" pageId="14" pageNumber="105">Figs. 1a</figureCitation>
,
<figureCitation id="133844B69F11FFE7FD88FDD669144C54" box="[545,573,540,563]" captionStart="Fig" captionStartId="7.[109,144,1944,1965]" captionTargetBox="[124,1456,222,1846]" captionTargetId="figure-138@7.[88,1486,191,1884]" captionTargetPageId="7" captionText="Fig. 5. Leptalpheus denticulatus, new species, male from Yaqara Bay, Viti Levu, Fiji (LACM CR 2005-007.1): a, frontal region, dorsal view; b, same, lateral view; c, first segment of right antennular peduncle, lateral view; d, mandible, mesial view; e, maxillule, lateral view; f, maxilla, lateral view; g, first maxilliped, lateral view; h, second maxilliped, lateral view; i, third maxilliped, lateral view; j, posterolateral margin of carapace, detail of cardiac notch; k, endopod of second pleopod; l, same, detail of appendix masculina; m, sixth pleonite and tail fan, setae omitted, lateral view; n, left uropod, dorsal view; o, right uropod, detail of exopod, setae omitted, dorsal view; p, telson, dorsal view; q, same, ventral view (showing anal tubercles). Scale bars = 1 mm; l drawn without scale." figureDoi="http://doi.org/10.5281/zenodo.5341640" httpUri="https://zenodo.org/record/5341640/files/figure.png" pageId="14" pageNumber="105">5a</figureCitation>
); the latter species is also variable in the proportions of the merus of the third pereiopod (
<figureCitation id="133844B69F11FFE7FEA8FD966A704C14" box="[257,345,604,627]" captionStart="Fig" captionStartId="4.[132,167,1973,1994]" captionTargetBox="[142,1466,267,1886]" captionTargetId="figure-112@4.[116,1507,248,1919]" captionTargetPageId="4" captionText="Fig. 2. Leptalpheus denticulatus, new species, ak: female paratype from Nhatrang Bay, Vietnam (USNM 1110002); l, m: male holotype from Nhatrang Bay (MNHN-Na 17064): a, mandible, mesial view; b, same, molar process; c, maxillule, lateral view; d, maxilla, lateral view; e, first maxilliped, lateral view; f, second maxilliped, lateral view; g, third maxilliped, lateral view; h, second pereiopod, lateral view; i, third pereiopod, lateral view; j, same, distal carpus, propodus and dactylus; k, fifth pereiopod, lateral view; l, second pleopod, lateral view; m, same, detail of appendix masculina and appendix interna. Scale bars = 1 mm; m drawn without scale." figureDoi="http://doi.org/10.5281/zenodo.5341634" httpUri="https://zenodo.org/record/5341634/files/figure.png" pageId="14" pageNumber="105">Figs. 2i</figureCitation>
,
<figureCitation id="133844B69F11FFE7FEC1FD966AAF4C14" box="[360,390,604,627]" captionStart="Fig" captionStartId="8.[132,167,2003,2024]" captionTargetBox="[127,1466,286,1915]" captionTargetId="figure-78@8.[108,1486,239,1944]" captionTargetPageId="8" captionText="Fig. 6. Leptalpheus denticulatus, new species, male from Yaqara Bay, Viti Levu, Fiji (LACM CR 2005-007.1): a, major (left) cheliped, lateral view; b, same, ventrolateral view; c, same, distal chela, ventromesial view; d, same, ventrolateral view; e, minor (right) cheliped, lateral view; f, same, chela, ventrolateral view. g, second pereiopod, lateral view; h, third pereiopod, lateral view; i, fifth pereiopod, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341642" httpUri="https://zenodo.org/record/5341642/files/figure.png" pageId="14" pageNumber="105">6h</figureCitation>
; see also above). Therefore, these features cannot be reliably used to distinguish
<taxonomicName id="4C0323B09F11FFE7FDCEFDB669D54CF4" authority="Anker &amp; Marin, 2009" authorityName="Anker &amp; Marin" authorityYear="2009" box="[615,764,636,659]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="105" phylum="Arthropoda" rank="species" species="dworschaki" status="sp. nov.">
<emphasis id="B97784219F11FFE7FDCEFDB669D54CF4" box="[615,764,636,659]" italics="true" pageId="14" pageNumber="105">L. dworschaki</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A244395A9F11FFE7FF2DFD566A2F4CD4" box="[132,262,668,691]" pageId="14" pageNumber="105" rank="species">new species</taxonomicNameLabel>
from
<taxonomicName id="4C0323B09F11FFE7FEE4FD566AC74CD4" authority="Anker &amp; Marin, 2009" authorityName="Anker &amp; Marin" authorityYear="2009" box="[333,494,668,691]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="105" phylum="Arthropoda" rank="species" species="denticulatus" status="sp. nov.">
<emphasis id="B97784219F11FFE7FEE4FD566AC74CD4" box="[333,494,668,691]" italics="true" pageId="14" pageNumber="105">L. denticulatus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A244395A9F11FFE7FE57FD5669AD4CD4" box="[510,644,668,691]" pageId="14" pageNumber="105" rank="species">new species</taxonomicNameLabel>
, or from
<taxonomicName id="4C0323B09F11FFE7FD44FD566BCA4CB4" authorityName="Banner &amp; Banner" authorityYear="1974" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="105" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B97784219F11FFE7FD44FD566BCA4CB4" italics="true" pageId="14" pageNumber="105">L. pacificus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>