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<mods:title id="6B366F8D849985CCC1E4E5C054F397F5">A new species of Pleurothallis (Orchidaceae: Pleurothallidinae) from Zamora in the Province of Zamora Chinchipe, Ecuador</mods:title>
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<mods:namePart id="2436758432A1F048220EE93BDB2AF589">Jiménez, Marco M.</mods:namePart>
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<mods:affiliation id="01E3F17E2222204F7B5D01F50F2A88F8">Vivero de Conservación La Paphinia, Avenida del Ejército y Juan Izquierdo, Zamora, Zamora Chinchipe, 190102, Ecuador. &amp; Grupo Científico Calaway Dodson: Investigación y Conservación de Orquídeas del Ecuador, Quito, 170510, Pichincha, Ecuador. &amp; Associazione Orchidofili Centro Italia, Treglio (Ch), Abruzzo, Italia. &amp; mmjimenez 473 @ gmail. com; https: // orcid. org / 0000 - 0002 - 9502 - 5651</mods:affiliation>
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<mods:namePart id="6638F58BC0D2C2FE1D0C3FA942B829AB">Horna, Luis Ocupa</mods:namePart>
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<mods:affiliation id="A5A00DD73937075C394842871C953702">Departamento de Orquideología, Centro de Investigación en Biología Tropical y Conservación - CINBIOTYC, Piura, Perú. &amp; Grupo Científico Calaway Dodson: Investigación y Conservación de Orquídeas del Ecuador, Quito, 170510, Pichincha, Ecuador. &amp; Departamento de Orquideología, Instituto de Ciencias Antonio Brack, Lima, Perú. &amp; luisocupa. horna @ gmail. com; https: // orcid. org / 0000 - 0002 - 4904 - 1060</mods:affiliation>
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<mods:namePart id="D5930F628A522705C0B37CFBE4CDA1B9">Vélez-Abarca, Leisberth</mods:namePart>
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<mods:affiliation id="032B5EAAA995C2C9A4E06F3020FD1AD1">Carrera de Biología, Universidad Estatal Amazónica (Sede El Pangui), Luis Imaicela entre Azuay y Mayor Rene Ulloa, El Pangui, Zamora Chinchipe, Ecuador. &amp; Grupo Científico Calaway Dodson: Investigación y Conservación de Orquídeas del Ecuador, Quito, 170510, Pichincha, Ecuador. &amp; Associazione Orchidofili Centro Italia, Treglio (Ch), Abruzzo, Italia. &amp; leis. alexis 92 @ gmail. com; https: // orcid. org / 0000 - 0003 - 3764 - 9682</mods:affiliation>
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<taxonomicName id="4C0A66FB7E3BFF99FF2DF898FC85AD21" ID-CoL="9V337" authority="M.M.Jimenez, Ocupa &amp; Velez-Abarca" authorityName="M. M. Jimenez, Ocupa &amp; Velez-Abarca" authorityYear="2021" box="[136,852,1808,1835]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="0" pageNumber="79" phylum="Tracheophyta" rank="species" species="bilobulata" status="sp. nov.">
<emphasis id="B97EC16A7E3BFF99FF2DF898FE48AD20" bold="true" box="[136,409,1808,1835]" italics="true" pageId="0" pageNumber="79">Pleurothallis bilobulata</emphasis>
M.M.Jiménez, Ocupa &amp; Vélez-Abarca
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<figureCitation id="133101FD7E3BFF99FC1EF898FBF7AD21" box="[955,1062,1808,1834]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="1.[136,229,1936,1958]" captionStartId-1="2.[136,229,1877,1899]" captionStartId-2="3.[136,229,906,928]" captionStartId-3="3.[136,229,2028,2050]" captionStartId-4="4.[136,229,1271,1293]" captionStartId-5="5.[136,229,1793,1815]" captionTargetBox-0="[166,1436,191,1869]" captionTargetBox-1="[151,1436,190,1853]" captionTargetBox-2="[291,1296,197,874]" captionTargetBox-3="[289,1299,959,1998]" captionTargetBox-4="[286,1302,196,1241]" captionTargetBox-5="[351,1234,473,1755]" captionTargetId-0="figure-18@1.[151,1436,188,1912]" captionTargetId-1="figure-22@2.[151,1436,190,1853]" captionTargetId-2="figure-18@3.[285,1302,190,881]" captionTargetId-3="figure-41@3.[283,1305,953,2004]" captionTargetId-4="figure-22@4.[280,1308,190,1247]" captionTargetId-5="figure-154@5.[340,1247,460,1768]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="3" captionTargetPageId-4="4" captionTargetPageId-5="5" captionText-0="FIGURE 1. Line drawing of P. bilobulata. A. Habit B. Flower C. Dissected perianth D. Column, lip and ovary, lateral view. E. Lip, adaxial and abaxial view. F. Anther cap and pollinarium. By L. Vélez-Abarca based on the the plant that served as type." captionText-1="FIGURE 2. Lankester Composite Dissection Plate (LCDP) of P. bilobulata. A. Habit B. Flower C. Dissected perianth. D. Close-up of the shortly dentate-ciliate petal. E. Column, lip and ovary, lateral view. F. Lip, adaxial view. G. Close up of the diffusely cellular verrucose lip. H. Anther cap and pollinarium. By L. Ocupa Horna and M.M. Jiménez based on the the plant that served as type." captionText-2="FIGURE 3. Detail of the flower. A. Pleurothallis bilobulata. B. Pleurothallis lanigera. Photos by M. Jiménez." captionText-3="FIGURE 4. Lip, column and ovary of P. bilobulata (left) and P. lanigera (right). A. Frontal view of the lip. B. Lateral view of the column, lip and ovary. Photos by M. Jiménez." captionText-4="FIGURE 5. Column, anther and pollinarium of P. bilobulata (left) and P. lanigera (right). A. Frontal view of the column. B. Anther and pollinarium. Photos by M. Jiménez." captionText-5="FIGURE 6. Distribution map of Pleurothallis bilobulata. Prepared by Eugenio Restrepo using ArcGIS." figureDoi-0="http://doi.org/10.5281/zenodo.5447961" figureDoi-1="http://doi.org/10.5281/zenodo.5447963" figureDoi-2="http://doi.org/10.5281/zenodo.5447965" figureDoi-3="http://doi.org/10.5281/zenodo.5447967" figureDoi-4="http://doi.org/10.5281/zenodo.5447969" figureDoi-5="http://doi.org/10.5281/zenodo.5447971" httpUri-0="https://zenodo.org/record/5447961/files/figure.png" httpUri-1="https://zenodo.org/record/5447963/files/figure.png" httpUri-2="https://zenodo.org/record/5447965/files/figure.png" httpUri-3="https://zenodo.org/record/5447967/files/figure.png" httpUri-4="https://zenodo.org/record/5447969/files/figure.png" httpUri-5="https://zenodo.org/record/5447971/files/figure.png" pageId="0" pageNumber="79">Figs. 1–6</figureCitation>
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<paragraph id="8BB51D787E3BFF99FF2DF82AFBE7A22D" blockId="0.[136,1452,1954,2086]" pageId="0" pageNumber="79">
Similar to 
<taxonomicName id="4C0A66FB7E3BFF99FF58F82BFDDAADB1" authority="Luer &amp; Hirtz" authorityName="Luer &amp; Hirtz" box="[253,523,1954,1978]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="0" pageNumber="79" phylum="Tracheophyta" rank="species" species="lanigera">
<emphasis id="B97EC16A7E3BFF99FF58F82BFEA2ADB1" box="[253,371,1954,1978]" italics="true" pageId="0" pageNumber="79">P. lanigera</emphasis>
Luer &amp; Hirtz
</taxonomicName>
in 
<bibRefCitation id="EF9B60897E3BFF99FD94F82AFD3AADB1" author="Luer, C. A." box="[561,747,1954,1978]" pageId="0" pageNumber="79" pagination="133 - 149" refId="ref2721" refString="Luer, C. A. (1988) A revision of some sections of subgenus Pleurothallis. Lindleyana 3: 133 - 149." type="journal article" year="1988">Luer (1988: 146)</bibRefCitation>
from which it is distinguished by the narrowly ovate leaves (
<emphasis id="B97EC16A7E3BFF99FA2BF82BFA7DADB1" box="[1422,1452,1955,1978]" italics="true" pageId="0" pageNumber="79">vs.</emphasis>
cordate-ovate), the straight, subverrucose ovary (
<emphasis id="B97EC16A7E3BFF99FD2BF84FFD7AADD5" box="[654,683,1991,2014]" italics="true" pageId="0" pageNumber="79">vs.</emphasis>
curved, minutely papillose), the broadly elliptic dorsal sepal (
<emphasis id="B97EC16A7E3BFF99FA92F84FFA84ADD5" box="[1335,1365,1991,2014]" italics="true" pageId="0" pageNumber="79">vs.</emphasis>
broadly ovate), the shortly acuminate, narrowly lanceolate, papillose-verrucose, 1-veined petals (
<emphasis id="B97EC16A7E3BFF99FBF7F863FBA1A209" box="[1106,1136,2027,2050]" italics="true" pageId="0" pageNumber="79">vs.</emphasis>
acute, elliptic, pubescent, 3- veined) and the lip with a basal, bilobed glenion (
<emphasis id="B97EC16A7E3BFF99FD3CF787FD67A22D" box="[665,694,2063,2086]" italics="true" pageId="0" pageNumber="79">vs.</emphasis>
lip with a central, circular glenion).
</paragraph>
<caption id="DF754DF07E3AFF98FF2DF818FAD5ADC2" ID-DOI="http://doi.org/10.5281/zenodo.5447961" ID-Zenodo-Dep="5447961" httpUri="https://zenodo.org/record/5447961/files/figure.png" pageId="1" pageNumber="80" startId="1.[136,229,1936,1958]" targetBox="[166,1436,191,1869]" targetPageId="1">
<paragraph id="8BB51D787E3AFF98FF2DF818FAD5ADC2" blockId="1.[136,1452,1936,1994]" pageId="1" pageNumber="80">
<emphasis id="B97EC16A7E3AFF98FF2DF818FED1ADAE" bold="true" box="[136,256,1936,1958]" pageId="1" pageNumber="80">FIGURE 1.</emphasis>
Line drawing of 
<emphasis id="B97EC16A7E3AFF98FE0AF818FDE3ADAE" box="[431,562,1936,1957]" italics="true" pageId="1" pageNumber="80">
<taxonomicName id="4C0A66FB7E3AFF98FE0AF818FDFFADAE" authorityName="M. M. Jimenez, Ocupa &amp; Velez-Abarca" authorityYear="2021" box="[431,558,1936,1957]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="80" phylum="Tracheophyta" rank="species" species="bilobulata">P. bilobulata</taxonomicName>
.
</emphasis>
<emphasis id="B97EC16A7E3AFF98FD9CF818FD80ADAE" bold="true" box="[569,593,1936,1957]" pageId="1" pageNumber="80">A.</emphasis>
Habit B. Flower 
<emphasis id="B97EC16A7E3AFF98FCA5F818FCC9ADAD" bold="true" box="[768,792,1936,1958]" pageId="1" pageNumber="80">C.</emphasis>
Dissected perianth 
<emphasis id="B97EC16A7E3AFF98FC79F818FC25ADAE" bold="true" box="[988,1012,1936,1957]" pageId="1" pageNumber="80">D.</emphasis>
Column, lip and ovary, lateral view. 
<emphasis id="B97EC16A7E3AFF98FAC3F818FAADADAE" bold="true" box="[1382,1404,1936,1957]" pageId="1" pageNumber="80">E.</emphasis>
Lip, adaxial and abaxial view. 
<emphasis id="B97EC16A7E3AFF98FE25F83CFE42ADC2" bold="true" box="[384,403,1972,1993]" pageId="1" pageNumber="80">F.</emphasis>
Anther cap and pollinarium. By L. Vélez-Abarca based on the the plant that served as type.
</paragraph>
</caption>
<caption id="DF754DF07E39FF9BFF2DF8DDFB19ADB8" ID-DOI="http://doi.org/10.5281/zenodo.5447963" ID-Zenodo-Dep="5447963" httpUri="https://zenodo.org/record/5447963/files/figure.png" pageId="2" pageNumber="81" startId="2.[136,229,1877,1899]" targetBox="[151,1436,190,1853]" targetPageId="2">
<paragraph id="8BB51D787E39FF9BFF2DF8DDFB19ADB8" blockId="2.[136,1452,1877,1971]" pageId="2" pageNumber="81">
<emphasis id="B97EC16A7E39FF9BFF2DF8DDFF2CAD61" bold="true" box="[136,253,1877,1899]" pageId="2" pageNumber="81">FIGURE 2.</emphasis>
Lankester Composite Dissection Plate (LCDP) of 
<taxonomicName id="4C0A66FB7E39FF9BFD45F8DEFC8BAD61" authorityName="M. M. Jimenez, Ocupa &amp; Velez-Abarca" authorityYear="2021" box="[736,858,1877,1899]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="81" phylum="Tracheophyta" rank="species" species="bilobulata">
<emphasis id="B97EC16A7E39FF9BFD45F8DEFC8BAD61" box="[736,858,1877,1899]" italics="true" pageId="2" pageNumber="81">P. bilobulata</emphasis>
</taxonomicName>
. 
<emphasis id="B97EC16A7E39FF9BFCC3F8DDFCACAD61" bold="true" box="[870,893,1877,1898]" pageId="2" pageNumber="81">A.</emphasis>
Habit 
<emphasis id="B97EC16A7E39FF9BFC18F8DEFC02AD60" bold="true" box="[957,979,1878,1899]" pageId="2" pageNumber="81">B.</emphasis>
Flower 
<emphasis id="B97EC16A7E39FF9BFB87F8DDFBE8AD60" bold="true" box="[1058,1081,1877,1899]" pageId="2" pageNumber="81">C.</emphasis>
Dissected perianth. 
<emphasis id="B97EC16A7E39FF9BFB59F8DEFAC2AD60" bold="true" box="[1276,1299,1878,1899]" pageId="2" pageNumber="81">D.</emphasis>
Close-up of the shortly dentate-ciliate petal. 
<emphasis id="B97EC16A7E39FF9BFE32F8F2FE7CAD84" bold="true" box="[407,429,1914,1935]" pageId="2" pageNumber="81">E.</emphasis>
Column, lip and ovary, lateral view. 
<emphasis id="B97EC16A7E39FF9BFCAFF8F2FCCDAD84" bold="true" box="[778,796,1914,1935]" pageId="2" pageNumber="81">F.</emphasis>
Lip, adaxial view. 
<emphasis id="B97EC16A7E39FF9BFC75F8F1FC38AD84" bold="true" box="[976,1001,1913,1935]" pageId="2" pageNumber="81">G.</emphasis>
Close up of the diffusely cellular verrucose lip. 
<emphasis id="B97EC16A7E39FF9BFF2DF816FF70ADB8" bold="true" box="[136,161,1950,1971]" pageId="2" pageNumber="81">H.</emphasis>
Anther cap and pollinarium. By L. Ocupa Horna and M.M. Jiménez based on the the plant that served as type.
</paragraph>
</caption>
<caption id="DF754DF07E38FF9AFF2DFC02FB60A9AB" ID-DOI="http://doi.org/10.5281/zenodo.5447965" ID-Zenodo-Dep="5447965" box="[136,1201,906,928]" httpUri="https://zenodo.org/record/5447965/files/figure.png" pageId="3" pageNumber="82" startId="3.[136,229,906,928]" targetBox="[291,1296,197,874]" targetPageId="3">
<paragraph id="8BB51D787E38FF9AFF2DFC02FB60A9AB" blockId="3.[136,1201,906,928]" box="[136,1201,906,928]" pageId="3" pageNumber="82">
<emphasis id="B97EC16A7E38FF9AFF2DFC02FF2FA994" bold="true" box="[136,254,906,928]" pageId="3" pageNumber="82">FIGURE 3.</emphasis>
Detail of the flower. 
<emphasis id="B97EC16A7E38FF9AFE6EFC02FE0DA994" bold="true" box="[459,476,906,927]" pageId="3" pageNumber="82">A</emphasis>
. 
<taxonomicName id="4C0A66FB7E38FF9AFE4DFC02FD1CA994" authorityName="M. M. Jimenez, Ocupa &amp; Velez-Abarca" authorityYear="2021" box="[488,717,906,927]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="82" phylum="Tracheophyta" rank="species" species="bilobulata">
<emphasis id="B97EC16A7E38FF9AFE4DFC02FD1CA994" box="[488,717,906,927]" italics="true" pageId="3" pageNumber="82">Pleurothallis bilobulata</emphasis>
</taxonomicName>
. 
<emphasis id="B97EC16A7E38FF9AFD7CFC02FD38A994" bold="true" box="[729,745,906,927]" pageId="3" pageNumber="82">B</emphasis>
. 
<taxonomicName id="4C0A66FB7E38FF9AFD50FC02FC18A994" box="[757,969,906,927]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="82" phylum="Tracheophyta" rank="species" species="lanigera">
<emphasis id="B97EC16A7E38FF9AFD50FC02FC18A994" box="[757,969,906,927]" italics="true" pageId="3" pageNumber="82">Pleurothallis lanigera</emphasis>
</taxonomicName>
. Photos by M. Jiménez.
</paragraph>
</caption>
<caption id="DF754DF07E38FF9AFF2DF864FE3FA22C" ID-DOI="http://doi.org/10.5281/zenodo.5447967" ID-Zenodo-Dep="5447967" httpUri="https://zenodo.org/record/5447967/files/figure.png" pageId="3" pageNumber="82" startId="3.[136,229,2028,2050]" targetBox="[289,1299,959,1998]" targetPageId="3">
<paragraph id="8BB51D787E38FF9AFF2DF864FE3FA22C" blockId="3.[136,1452,2028,2087]" pageId="3" pageNumber="82">
<emphasis id="B97EC16A7E38FF9AFF2DF864FF2CA209" bold="true" box="[136,253,2028,2050]" pageId="3" pageNumber="82">FIGURE 4.</emphasis>
Lip, column and ovary of 
<taxonomicName id="4C0A66FB7E38FF9AFE5FF865FDA5A209" authorityName="M. M. Jimenez, Ocupa &amp; Velez-Abarca" authorityYear="2021" box="[506,628,2029,2050]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="82" phylum="Tracheophyta" rank="species" species="bilobulata">
<emphasis id="B97EC16A7E38FF9AFE5FF865FDA5A209" box="[506,628,2029,2050]" italics="true" pageId="3" pageNumber="82">P. bilobulata</emphasis>
</taxonomicName>
(left) and 
<taxonomicName id="4C0A66FB7E38FF9AFD73F865FCEFA209" box="[726,830,2029,2050]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="82" phylum="Tracheophyta" rank="species" species="lanigera">
<emphasis id="B97EC16A7E38FF9AFD73F865FCEFA209" box="[726,830,2029,2050]" italics="true" pageId="3" pageNumber="82">P. lanigera</emphasis>
</taxonomicName>
(right). 
<emphasis id="B97EC16A7E38FF9AFC29F865FC4CA209" bold="true" box="[908,925,2029,2050]" pageId="3" pageNumber="82">A</emphasis>
. Frontal view of the lip. 
<emphasis id="B97EC16A7E38FF9AFB2CF865FB48A209" bold="true" box="[1161,1177,2029,2050]" pageId="3" pageNumber="82">B</emphasis>
. Lateral view of the column, lip and ovary. Photos by M. Jiménez.
</paragraph>
</caption>
<caption id="DF754DF07E3FFF9DFF2DFB7FFE30AF3A" ID-DOI="http://doi.org/10.5281/zenodo.5447969" ID-Zenodo-Dep="5447969" httpUri="https://zenodo.org/record/5447969/files/figure.png" pageId="4" pageNumber="83" startId="4.[136,229,1271,1293]" targetBox="[286,1302,196,1241]" targetPageId="4">
<paragraph id="8BB51D787E3FFF9DFF2DFB7FFE30AF3A" blockId="4.[136,1452,1271,1329]" pageId="4" pageNumber="83">
<emphasis id="B97EC16A7E3FFF9DFF2DFB7FFF2FAF07" bold="true" box="[136,254,1271,1293]" pageId="4" pageNumber="83">FIGURE 5.</emphasis>
Column, anther and pollinarium of 
<taxonomicName id="4C0A66FB7E3FFF9DFDFFFB70FD04AF07" authorityName="M. M. Jimenez, Ocupa &amp; Velez-Abarca" authorityYear="2021" box="[602,725,1271,1293]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="83" phylum="Tracheophyta" rank="species" species="bilobulata">
<emphasis id="B97EC16A7E3FFF9DFDFFFB70FD04AF07" box="[602,725,1271,1293]" italics="true" pageId="4" pageNumber="83">P. bilobulata</emphasis>
</taxonomicName>
(left) and 
<taxonomicName id="4C0A66FB7E3FFF9DFC9FFB70FC72AF07" box="[826,931,1271,1293]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="83" phylum="Tracheophyta" rank="species" species="lanigera">
<emphasis id="B97EC16A7E3FFF9DFC9FFB70FC72AF07" box="[826,931,1271,1293]" italics="true" pageId="4" pageNumber="83">P. lanigera</emphasis>
</taxonomicName>
(right). 
<emphasis id="B97EC16A7E3FFF9DFC51FB7FFBD4AF07" bold="true" box="[1012,1029,1271,1292]" pageId="4" pageNumber="83">A</emphasis>
. Frontal view of the column. 
<emphasis id="B97EC16A7E3FFF9DFA80FB7FFAE4AF07" bold="true" box="[1317,1333,1271,1292]" pageId="4" pageNumber="83">B</emphasis>
. Anther and pollinarium. Photos by M. Jiménez.
</paragraph>
</caption>
<paragraph id="8BB51D787E3FFF9DFF18FAE9FF28AD9E" blockId="4.[136,1452,1377,2086]" pageId="4" pageNumber="83">
<emphasis id="B97EC16A7E3FFF9DFF18FAE9FF26AF72" box="[189,247,1377,1401]" italics="true" pageId="4" pageNumber="83">Plant</emphasis>
epiphytic, up to 
<quantity id="4CF2B09D7E3FFF9DFE09FAE9FE3EAF72" box="[428,495,1377,1401]" metricMagnitude="-1" metricUnit="m" metricValue="1.6" pageId="4" pageNumber="83" unit="cm" value="16.0">16 cm</quantity>
tall; 
<emphasis id="B97EC16A7E3FFF9DFD83FAEAFD8DAF72" box="[550,604,1378,1401]" italics="true" pageId="4" pageNumber="83">roots</emphasis>
slender, flexuous, 
<quantity id="4CF2B09D7E3FFF9DFC83FAE9FC74AF72" box="[806,933,1377,1401]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" metricValueMax="13.0" metricValueMin="6.0" pageId="4" pageNumber="83" unit="mm" value="0.95" valueMax="1.3" valueMin="0.6">0.6–1.3 mm</quantity>
in diameter. 
<emphasis id="B97EC16A7E3FFF9DFB97FAE9FB72AF72" box="[1074,1187,1377,1401]" italics="true" pageId="4" pageNumber="83">Ramicauls</emphasis>
slender, suberect, terete, 45–112 × 
<quantity id="4CF2B09D7E3FFF9DFF52FA0DFEADAF96" box="[247,380,1413,1437]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="4" pageNumber="83" unit="mm" value="1.0" valueMax="1.2" valueMin="0.8">0.8–1.2 mm</quantity>
, enclosed by papyraceous, tubular sheaths, upper sheath 
<quantity id="4CF2B09D7E3FFF9DFC4EFA0DFB56AF96" box="[1003,1159,1413,1437]" metricMagnitude="-2" metricUnit="m" metricValue="1.5899999999999999" metricValueMax="1.89" metricValueMin="1.29" pageId="4" pageNumber="83" unit="mm" value="15.899999999999999" valueMax="18.9" valueMin="12.9">12.9–18.9 mm</quantity>
long, basal sheath 
<quantity id="4CF2B09D7E3FFF9DFAFEFA0DFF60AFCA" metricMagnitude="-3" metricUnit="m" metricValue="6.95" metricValueMax="7.7" metricValueMin="6.2" pageId="4" pageNumber="83" unit="mm" value="6.95" valueMax="7.7" valueMin="6.2">6.2–7.7 mm</quantity>
long. 
<emphasis id="B97EC16A7E3FFF9DFF59FA22FE97AFCA" box="[252,326,1450,1473]" italics="true" pageId="4" pageNumber="83">Leaves</emphasis>
deflexed, coriaceous, microscopically papillate, narrowly ovate, 4.6–7.5 × 
<quantity id="4CF2B09D7E3FFF9DFB29FA21FAD8AFCA" box="[1164,1289,1449,1473]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" metricValueMax="2.2" metricValueMin="1.4" pageId="4" pageNumber="83" unit="cm" value="1.8" valueMax="2.2" valueMin="1.4">1.4–2.2 cm</quantity>
, acute, shortly acuminate, sessile, cordate at the base, lobes unequal. 
<emphasis id="B97EC16A7E3FFF9DFD77FA45FCB0AFEE" box="[722,865,1485,1509]" italics="true" pageId="4" pageNumber="83">Inflorescence</emphasis>
a fascicle of successive flowers produced from a suberect, scabrous spathaceous bract, 
<quantity id="4CF2B09D7E3FFF9DFE5BFA79FDAEAC02" box="[510,639,1521,1545]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" metricValueMax="7.4" metricValueMin="4.8" pageId="4" pageNumber="83" unit="mm" value="6.1" valueMax="7.4" valueMin="4.8">4.8–7.4 mm</quantity>
long; 
<emphasis id="B97EC16A7E3FFF9DFD62FA79FC92AC02" box="[711,835,1521,1545]" italics="true" pageId="4" pageNumber="83">floral bract</emphasis>
infundibuliform, 
<quantity id="4CF2B09D7E3FFF9DFBA0FA79FB57AC02" box="[1029,1158,1521,1545]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="4.6" metricValueMin="4.4" pageId="4" pageNumber="83" unit="mm" value="4.5" valueMax="4.6" valueMin="4.4">4.4–4.6 mm</quantity>
long; 
<emphasis id="B97EC16A7E3FFF9DFB6BFA79FACAAC02" box="[1230,1307,1521,1545]" italics="true" pageId="4" pageNumber="83">pedicel</emphasis>
subflexuous, 
<quantity id="4CF2B09D7E3FFF9DFF2DF99DFED9AC26" box="[136,264,1557,1581]" metricMagnitude="-3" metricUnit="m" metricValue="2.95" metricValueMax="3.5" metricValueMin="2.4" pageId="4" pageNumber="83" unit="mm" value="2.95" valueMax="3.5" valueMin="2.4">2.4–3.5 mm</quantity>
long; 
<emphasis id="B97EC16A7E3FFF9DFEE9F99EFE58AC26" box="[332,393,1558,1581]" italics="true" pageId="4" pageNumber="83">ovary</emphasis>
subverrucose, straight, 2.6–3.0 × 1.0 mm. 
<emphasis id="B97EC16A7E3FFF9DFCF5F99DFC76AC26" box="[848,935,1557,1581]" italics="true" pageId="4" pageNumber="83">Flowers</emphasis>
resupinate or not; dorsal sepal chestnut, broadly elliptic, acute, 6.9–8.0 × 
<quantity id="4CF2B09D7E3FFF9DFE29F9B1FDDFAC5A" box="[396,526,1593,1617]" metricMagnitude="-3" metricUnit="m" metricValue="4.199999999999999" metricValueMax="4.3" metricValueMin="4.1" pageId="4" pageNumber="83" unit="mm" value="4.199999999999999" valueMax="4.3" valueMin="4.1">4.1–4.3 mm</quantity>
, 5–veined, pubescent-verruculose, glossy externally; the lateral sepals fused in an ovoid synsepal, burgundy-chestnut, acute, 5.7–6.5 × 
<quantity id="4CF2B09D7E3FFF9DFD23F9D5FCDAAC7E" box="[646,779,1629,1653]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.1" metricValueMin="3.9" pageId="4" pageNumber="83" unit="mm" value="4.0" valueMax="4.1" valueMin="3.9">3.9–4.1 mm</quantity>
, concave centrally, 4-veined, verruculose. Petals burgundy to chestnut, narrowly lanceolate, shortly acuminate, 4.7–5.3 × 1.0– 
<quantity id="4CF2B09D7E3FFF9DFCE5F909FC45AC92" box="[832,916,1665,1689]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="83" unit="mm" value="1.2">1.2 mm</quantity>
, 1-veined, microscopically papillose-verruculose, margin shortly dentate-ciliate, constricted and oblique at the base. Lip burgundy chestnut, stout, thick, ovate, obtuse, 2.6–2.8 × 
<quantity id="4CF2B09D7E3FFF9DFF3AF941FEF3ACEA" box="[159,290,1737,1761]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.3" metricValueMin="2.2" pageId="4" pageNumber="83" unit="mm" value="2.25" valueMax="2.3" valueMin="2.2">2.2–2.3 mm</quantity>
, diffusely cellular verrucose, convex, slightly concave at the apex and to either side of the glenion; the base subtruncate, hinged to the column-foot, with short, obtuse angles and a large, deep, obovoid, bilobed glenion occupying the basal third of the lip. Column stout, pale green suffused with purple, 1.3–1.4 × 
<quantity id="4CF2B09D7E3FFF9DFBADF899FB5DAD22" box="[1032,1164,1809,1833]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="4" pageNumber="83" unit="mm" value="1.6" valueMax="1.7" valueMin="1.5">1.5–1.7 mm</quantity>
, papillose, stigma bilobed, ventral, reniform, minutely ciliate on margins, rostellar flap 
<quantity id="4CF2B09D7E3FFF9DFCB0F8BDFCB6AD46" box="[789,871,1845,1869]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="83" unit="mm" value="0.4">0.4 mm</quantity>
long. Anther narrowly deltoid, white, apical, 
<quantity id="4CF2B09D7E3FFF9DFAFDF8BDFA7AAD46" box="[1368,1451,1845,1869]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="83" unit="mm" value="0.6">0.6 mm</quantity>
long, bilobed at the base. Pollinarium with 2 narrowly obovoid, yellow pollinia, 
<quantity id="4CF2B09D7E3FFF9DFC77F8D1FBF3AD7A" box="[978,1058,1881,1905]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="83" unit="mm" value="0.6">0.6 mm</quantity>
long, brought together by a drop-like viscidium.
</paragraph>
</subSubSection>
<subSubSection id="C3104EF37E3FFF9DFF18F829FA4AADB2" box="[189,1435,1953,1977]" pageId="4" pageNumber="83" type="etymology">
<paragraph id="8BB51D787E3FFF9DFF18F829FA4AADB2" blockId="4.[136,1452,1377,2086]" box="[189,1435,1953,1977]" pageId="4" pageNumber="83">
<emphasis id="B97EC16A7E3FFF9DFF18F829FE90ADB2" bold="true" box="[189,321,1953,1977]" pageId="4" pageNumber="83">Etymology:</emphasis>
—From the Latin 
<emphasis id="B97EC16A7E3FFF9DFDA6F829FDC9ADB2" box="[515,536,1953,1977]" italics="true" pageId="4" pageNumber="83">bi</emphasis>
, “two”, and 
<emphasis id="B97EC16A7E3FFF9DFD3AF829FCD3ADB2" box="[671,770,1953,1977]" italics="true" pageId="4" pageNumber="83">lobulatus</emphasis>
, “small lobes”, referring to two lobes in the glenion of the lip.
</paragraph>
</subSubSection>
<subSubSection id="C3104EF37E3FFF9CFF18F84DFCE1ABB8" lastPageId="5" lastPageNumber="84" pageId="4" pageNumber="83" type="distribution">
<paragraph id="8BB51D787E3FFF9CFF18F84DFCE1ABB8" blockId="4.[136,1452,1377,2086]" lastBlockId="5.[136,1452,159,436]" lastPageId="5" lastPageNumber="84" pageId="4" pageNumber="83">
<emphasis id="B97EC16A7E3FFF9DFF18F84DFE0FADD6" bold="true" box="[189,478,1989,2013]" pageId="4" pageNumber="83">Distribution and habitat:</emphasis>
— 
<taxonomicName id="4C0A66FB7E3FFF9DFE5DF84DFD28ADD6" authorityName="M. M. Jimenez, Ocupa &amp; Velez-Abarca" authorityYear="2021" box="[504,761,1989,2013]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="83" phylum="Tracheophyta" rank="species" species="bilobulata">
<emphasis id="B97EC16A7E3FFF9DFE5DF84DFD28ADD6" box="[504,761,1989,2013]" italics="true" pageId="4" pageNumber="83">Pleurothallis bilobulata</emphasis>
</taxonomicName>
is only known from the 
<typeStatus id="54B1A3DA7E3FFF9DFBA0F84EFBE2ADD5" box="[1029,1075,1990,2014]" pageId="4" pageNumber="83">type</typeStatus>
locality 
<typeStatus id="54B1A3DA7E3FFF9DFB36F84EFB10ADD5" box="[1171,1217,1990,2014]" pageId="4" pageNumber="83">type</typeStatus>
on the steep hillsides near Zamora, in the 
<collectingRegion id="49CED39A7E3FFF9DFECCF862FD49A20A" box="[361,664,2025,2049]" country="Ecuador" name="Zamora-Chinchipe" pageId="4" pageNumber="83">Zamora Chinchipe province</collectingRegion>
(
<figureCitation id="133101FD7E3FFF9DFD0CF861FD3CA20A" box="[681,749,2025,2049]" captionStart="FIGURE 6" captionStartId="5.[136,229,1793,1815]" captionTargetBox="[351,1234,473,1755]" captionTargetId="figure-154@5.[340,1247,460,1768]" captionTargetPageId="5" captionText="FIGURE 6. Distribution map of Pleurothallis bilobulata. Prepared by Eugenio Restrepo using ArcGIS." figureDoi="http://doi.org/10.5281/zenodo.5447971" httpUri="https://zenodo.org/record/5447971/files/figure.png" pageId="4" pageNumber="83">Fig. 6</figureCitation>
). The species grows on hill-tops in the premontane rainforest at around 
<quantity id="4CF2B09D7E3FFF9DFF7FF785FEF1A22E" box="[218,288,2061,2085]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="83" unit="m" value="900.0">900 m</quantity>
in elevation, on the Eastern slope of the Cordillera Oriental of the Andes. It is a rare species that grows in primary forests, on understory lianas, together with other orchid species such as 
<taxonomicName id="4C0A66FB7E3EFF9CFBAEFF17FA74AABC" authority="Garay (1978: 71)" authorityName="Garay" authorityPageNumber="71" authorityYear="1978" box="[1035,1445,159,183]" class="Liliopsida" family="Orchidaceae" genus="Elleanthus" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="84" phylum="Tracheophyta" rank="species" species="blatteus">
<emphasis id="B97EC16A7E3EFF9CFBAEFF17FB0DAABC" box="[1035,1244,159,183]" italics="true" pageId="5" pageNumber="84">Elleanthus blatteus</emphasis>
<bibRefCitation id="EF9B60897E3EFF9CFB43FF17FA74AABC" author="Garay, L. A." box="[1254,1445,159,183]" pageId="5" pageNumber="84" refId="ref2411" refString="Garay, L. A. (1978) Orchidaceae. Cypripedioideae Orchidoideae Neottioideae. In: Harling, G. &amp; Sparre, B. (Eds.) Flora of Ecuador. Volume 9. Goteborg University, Riksmuseum, pp. 1 - 305." type="book" year="1978">Garay (1978: 71)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C0A66FB7E3EFF9CFF2DFF4CFC7EAAD0" authority="(Poeppig &amp; Endlicher 1836: 33) Reichenbach (1863: 481)" authorityName="Reichenbach" authorityPageNumber="481" authorityYear="1863" baseAuthorityName="Poeppig &amp; Endlicher" baseAuthorityPageNumber="33" baseAuthorityYear="1836" box="[136,943,195,219]" class="Liliopsida" family="Orchidaceae" genus="Elleanthus" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="84" phylum="Tracheophyta" rank="species" species="oliganthus">
<emphasis id="B97EC16A7E3EFF9CFF2DFF4CFECAAAD0" box="[136,283,195,219]" italics="true" pageId="5" pageNumber="84">E. oliganthus</emphasis>
(
<bibRefCitation id="EF9B60897E3EFF9CFE89FF4BFD54AAD0" author="Poeppig, E. F. &amp; Endlicher, S. F." box="[300,645,195,219]" pageId="5" pageNumber="84" refId="ref2948" refString="Poeppig, E. F. &amp; Endlicher, S. F. (1836) Nova genera ac species plantarum, quas in regno Chilensi Peruviano et in terra Amazonica, vol. 1. Hofmeister, Leipzig, 100 pp." type="book" year="1836">Poeppig &amp; Endlicher 1836: 33</bibRefCitation>
) 
<bibRefCitation id="EF9B60897E3EFF9CFD32FF4BFC7EAAD0" author="Reichenbach, H. G." box="[663,943,195,219]" pageId="5" pageNumber="84" pagination="167 - 930" refId="ref3274" refString="Reichenbach, H. G. (1863) Orchides Juss. Annales Botanices Systematicae 6: 167 - 930." type="journal article" year="1863">Reichenbach (1863: 481)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C0A66FB7E3EFF9CFC1BFF4BFF68AAF4" authority="Sweet &amp; Garay (1966: 377)" authorityName="Sweet &amp; Garay" authorityPageNumber="377" authorityYear="1966" class="Liliopsida" family="Orchidaceae" genus="Masdevallia" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="84" phylum="Tracheophyta" rank="species" species="strobelii">
<emphasis id="B97EC16A7E3EFF9CFC1BFF4BFB75AAD0" box="[958,1188,195,219]" italics="true" pageId="5" pageNumber="84">Masdevallia strobelii</emphasis>
<bibRefCitation id="EF9B60897E3EFF9CFB0AFF4BFF68AAF4" author="Sweet, H. R. &amp; Garay, L. A." pageId="5" pageNumber="84" pagination="377 - 378" refId="ref3502" refString="Sweet, H. R. &amp; Garay, L. A. (1966) A new Masdevallia from Ecuador. American Orchid Society Bulletin 35: 377 - 378." type="journal article" year="1966">Sweet &amp; Garay (1966: 377)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C0A66FB7E3EFF9CFF6DFF60FD12AAF4" authority="Reichenbach (1883: 198)" authorityName="Reichenbach" authorityPageNumber="198" authorityYear="1883" box="[200,707,231,255]" class="Liliopsida" family="Orchidaceae" genus="Peristeria" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="84" phylum="Tracheophyta" rank="species" species="ephippium">
<emphasis id="B97EC16A7E3EFF9CFF6DFF60FE7AAAF4" box="[200,427,231,255]" italics="true" pageId="5" pageNumber="84">Peristeria ephippium</emphasis>
<bibRefCitation id="EF9B60897E3EFF9CFE17FF6FFD12AAF4" author="Reichenbach, H. G." box="[434,707,231,255]" pageId="5" pageNumber="84" pagination="198" refId="ref3295" refString="Reichenbach, H. G. (1883) New garden plants. Gardeners' Chronicle, new series 20: 198." type="book chapter" year="1883">Reichenbach (1883: 198)</bibRefCitation>
</taxonomicName>
, 
<emphasis id="B97EC16A7E3EFF9CFD74FF6FFC14AAF4" box="[721,965,231,255]" italics="true" pageId="5" pageNumber="84">
<taxonomicName id="4C0A66FB7E3EFF9CFD74FF6FFC10AAF4" box="[721,961,231,255]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="84" phylum="Tracheophyta" rank="species" species="lanigera">Pleurothallis lanigera</taxonomicName>
.
</emphasis>
The local flora is dominated by 
<taxonomicName id="4C0A66FB7E3EFF9CFA8DFF6FFDD8AB28" authority="Occhioni (1954: 23)" authorityName="Occhioni" authorityPageNumber="23" authorityYear="1954" class="Magnoliopsida" family="Chloranthaceae" genus="Hedyosmum" kingdom="Plantae" order="Chloranthales" pageId="5" pageNumber="84" phylum="Tracheophyta" rank="species" species="cuatrecazanum">
<emphasis id="B97EC16A7E3EFF9CFA8DFF6FFEFBAB28" italics="true" pageId="5" pageNumber="84">Hedyosmum cuatrecazanum</emphasis>
<bibRefCitation id="EF9B60897E3EFF9CFE94FE83FDD8AB28" author="Occhioni, P." box="[305,521,267,291]" pageId="5" pageNumber="84" refId="ref2828" refString="Occhioni, P. (1954) Contribuicao ao estudo da familia Chloranthaceae com especial referencia ao genero Hedyosmum Sw. Tese (Professor Catedratico). Rio de Janeiro: Universidade do Brasil, 176 pp." type="book" year="1954">Occhioni (1954: 23)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C0A66FB7E3EFF9CFDB3FE83FB1CAB28" authority="(Ruiz &amp; Pavon 1802: 86) Don (1823: 295)" authorityName="Don" authorityPageNumber="295" authorityYear="1823" baseAuthorityName="Ruiz &amp; Pavon" baseAuthorityPageNumber="86" baseAuthorityYear="1802" box="[534,1229,267,291]" class="Magnoliopsida" family="Melastomataceae" genus="Pleroma" kingdom="Plantae" order="Myrtales" pageId="5" pageNumber="84" phylum="Tracheophyta" rank="species" species="ochypetalum">
<emphasis id="B97EC16A7E3EFF9CFDB3FE83FCD1AB28" box="[534,768,267,291]" italics="true" pageId="5" pageNumber="84">Pleroma ochypetalum</emphasis>
(
<bibRefCitation id="EF9B60897E3EFF9CFCABFE83FBDCAB28" author="Ruiz, H. &amp; Pavon, J." box="[782,1037,267,291]" pageId="5" pageNumber="84" refId="ref3360" refString="Ruiz, H. &amp; Pavon, J. (1802) Flora Peruviana, et Chilensis: Tomus III. Madrid: Gabrielis de Sancha." type="book" year="1802">Ruiz &amp; Pavón 1802: 86</bibRefCitation>
) 
<bibRefCitation id="EF9B60897E3EFF9CFBBEFE84FB1CAB28" author="Don, D." box="[1051,1229,267,291]" pageId="5" pageNumber="84" pagination="276 - 329" refId="ref2380" refString="Don, D. (1823) An illustration of the natural family of plants called Melastomaceae. Memoirs of the Wernerian Natural History Society 4: 276 - 329." type="journal article" year="1823">Don (1823: 295)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C0A66FB7E3EFF9CFB7FFE84FDE0AB4C" authority="(Kunth 1818: 35 - 36) Pruski (1996: 97)" authorityName="Pruski" authorityPageNumber="97" authorityYear="1996" baseAuthorityName="Kunth" baseAuthorityPageNumber="35 - 36" baseAuthorityYear="1818" class="Magnoliopsida" family="Asteraceae" genus="Piptocoma" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="84" phylum="Tracheophyta" rank="species" species="discolor">
<emphasis id="B97EC16A7E3EFF9CFB7FFE84FA7AAB28" box="[1242,1451,267,291]" italics="true" pageId="5" pageNumber="84">Piptocoma discolor</emphasis>
(
<bibRefCitation id="EF9B60897E3EFF9CFF2AFEA7FEB5AB4C" author="Kunth, K. S." box="[143,356,303,327]" pageId="5" pageNumber="84" pagination="301 - 412" refId="ref2615" refString="Kunth, K. S. (1818) Nova Genera et Species Plantarum (folio ed.), vol. 4. N. Maze, Paris, 301 - 412." type="book chapter" year="1818">Kunth 1818: 35–36</bibRefCitation>
) 
<bibRefCitation id="EF9B60897E3EFF9CFED1FEA7FDE0AB4C" author="Pruski, J. F." box="[372,561,303,327]" pageId="5" pageNumber="84" pagination="96 - 102" refId="ref3112" refString="Pruski, J. F. (1996) Compositae of the Guayana Highland: X. Reduction of Pollalesta to Piptocoma (Vernonieae: Piptocarphinae) and Consequent Nomenclatural Adjustments. Novon 6: 96 - 102. https: // doi. org / 10.2307 / 3392220" type="journal article" year="1996">Pruski (1996: 97)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C0A66FB7E3EFF9CFDE5FEA7FA75AB4C" authority="(Humboldt ex Mutis 1793: 465) Andersson (1994: 19)" authorityName="Andersson" authorityPageNumber="19" authorityYear="1994" baseAuthorityName="Mutis" baseAuthorityPageNumber="465" baseAuthorityYear="1793" box="[576,1444,303,327]" class="Magnoliopsida" family="Rubiaceae" genus="Ladenbergia" kingdom="Plantae" order="Gentianales" pageId="5" pageNumber="84" phylum="Tracheophyta" rank="species" species="oblongifolia">
<emphasis id="B97EC16A7E3EFF9CFDE5FEA7FC80AB4C" box="[576,849,303,327]" italics="true" pageId="5" pageNumber="84">Ladenbergia oblongifolia</emphasis>
(Humboldt ex 
<bibRefCitation id="EF9B60897E3EFF9CFC52FEA7FB7AAB4C" author="Mutis, J. C." box="[1015,1195,303,327]" pageId="5" pageNumber="84" pagination="461 - 465" refId="ref2800" refString="Mutis, J. C. (1793) El Arcano de la Quina. Papel Periodico de Santa Fe de Bogota 111: 461 - 465." type="journal article" year="1793">Mutis 1793: 465</bibRefCitation>
) 
<bibRefCitation id="EF9B60897E3EFF9CFB1FFEA7FA75AB4C" author="Andersson, L." box="[1210,1444,303,327]" pageId="5" pageNumber="84" pagination="16 - 33" refId="ref2189" refString="Andersson, L. (1994) Rubiaceae: Ladenbergia. In: Harling, G. &amp; Andersson, L. (Eds.) Flora of Ecuador. Volume 50: Goteborg University, Riksmuseum, pp. 16 - 33." type="book chapter" year="1994">Andersson (1994: 19)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C0A66FB7E3EFF9CFF2DFEDCFC89AB60" authority="(Bergius 1772: 131 - 132) Rusby (1907: 447)" authorityName="Rusby" authorityPageNumber="447" authorityYear="1907" baseAuthorityName="Bergius" baseAuthorityPageNumber="131 - 132" baseAuthorityYear="1772" box="[136,856,339,363]" class="Magnoliopsida" family="Lacistemataceae" genus="Lacistema" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="84" phylum="Tracheophyta" rank="species" species="aggregatum">
<emphasis id="B97EC16A7E3EFF9CFF2DFEDCFEADAB60" box="[136,380,340,363]" italics="true" pageId="5" pageNumber="84">Lacistema aggregatum</emphasis>
(
<bibRefCitation id="EF9B60897E3EFF9CFE2CFEDBFD55AB60" author="Bergius, P. J." box="[393,644,339,363]" pageId="5" pageNumber="84" pagination="131 - 132" refId="ref2349" refString="Bergius, P. J. (1772) Piper aggregatum. Acta Helvetica, Physico-Mathematico-Anatomico-Botanico-Medica Figuris Aeneis Illustrata, et in usus publicos exarata 7: 131 - 132." type="journal article" year="1772">Bergius 1772: 131–132</bibRefCitation>
) 
<bibRefCitation id="EF9B60897E3EFF9CFD37FEDBFC89AB60" author="Rusby, H. H." box="[658,856,339,363]" pageId="5" pageNumber="84" pagination="309 - 470" refId="ref3317" refString="Rusby, H. H. (1907) An enumeration of the plants collected by Miguel Bang. Part 4. With descriptions of new genera and species. Bulletin of the New York Botanical Garden 4: 309 - 470." type="journal article" year="1907">Rusby (1907: 447)</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C0A66FB7E3EFF9CFC2EFEDCFF6DAB84" authority="(Swartz 1788: 70) Steudel (1841: 139)" authorityName="Steudel" authorityPageNumber="139" authorityYear="1841" baseAuthorityName="Swartz" baseAuthorityPageNumber="70" baseAuthorityYear="1788" class="Magnoliopsida" family="Melastomataceae" genus="Miconia" kingdom="Plantae" order="Myrtales" pageId="5" pageNumber="84" phylum="Tracheophyta" rank="species" species="albicans">
<emphasis id="B97EC16A7E3EFF9CFC2EFEDCFB95AB60" box="[907,1092,339,363]" italics="true" pageId="5" pageNumber="84">Miconia albicans</emphasis>
(
<bibRefCitation id="EF9B60897E3EFF9CFBF7FEDBFAD3AB60" author="Swartz, O." box="[1106,1282,339,363]" pageId="5" pageNumber="84" pagination="70" refId="ref3484" refString="Swartz, O. (1788) Nova Genera et Species Plantarum. Prodromus 5: 70." type="journal article" year="1788">Swartz 1788: 70</bibRefCitation>
) 
<bibRefCitation id="EF9B60897E3EFF9CFAB5FEDBFF6DAB84" author="Steudel, E. G. von" pageId="5" pageNumber="84" refId="ref3450" refString="Steudel, E. G. von (1841) Nomenclator Botanicus (Vol. 2). Typis et sumptibus J. G. Cottae, Stuttgartiae &amp; Tubingae, 810 pp." type="book" year="1841">Steudel (1841: 139)</bibRefCitation>
</taxonomicName>
. The species may be considered as threatened given the destruction of the habitat where the only individuals have been found. It is not currently known to occur in any protected areas.
</paragraph>
</subSubSection>
<caption id="DF754DF07E3EFF9CFF2DF889FBBCAD1D" ID-DOI="http://doi.org/10.5281/zenodo.5447971" ID-Zenodo-Dep="5447971" box="[136,1133,1793,1815]" httpUri="https://zenodo.org/record/5447971/files/figure.png" pageId="5" pageNumber="84" startId="5.[136,229,1793,1815]" targetBox="[351,1234,473,1755]" targetPageId="5">
<paragraph id="8BB51D787E3EFF9CFF2DF889FBBCAD1D" blockId="5.[136,1133,1793,1815]" box="[136,1133,1793,1815]" pageId="5" pageNumber="84">
<emphasis id="B97EC16A7E3EFF9CFF2DF889FF2CAD1D" bold="true" box="[136,253,1793,1815]" pageId="5" pageNumber="84">FIGURE 6.</emphasis>
Distribution map of 
<emphasis id="B97EC16A7E3EFF9CFE62F889FD62AD1D" box="[455,691,1793,1814]" italics="true" pageId="5" pageNumber="84">
<taxonomicName id="4C0A66FB7E3EFF9CFE62F889FD7EAD1D" authorityName="M. M. Jimenez, Ocupa &amp; Velez-Abarca" authorityYear="2021" box="[455,687,1793,1814]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="84" phylum="Tracheophyta" rank="species" species="bilobulata">Pleurothallis bilobulata</taxonomicName>
.
</emphasis>
Prepared by Eugenio Restrepo using ArcGIS.
</paragraph>
</caption>
<subSubSection id="C3104EF37E3EFF9FFF18F8CFFE77AB28" lastPageId="6" lastPageNumber="85" pageId="5" pageNumber="84" type="discussion">
<paragraph id="8BB51D787E3EFF9CFF18F8CFFDEFA218" blockId="5.[136,1452,1863,2067]" pageId="5" pageNumber="84">
<emphasis id="B97EC16A7E3EFF9CFF18F8CFFE90AD54" bold="true" box="[189,321,1863,1887]" pageId="5" pageNumber="84">Comments:</emphasis>
—The new species is reminiscent of 
<taxonomicName id="4C0A66FB7E3EFF9CFD6FF8C0FCEFAD54" box="[714,830,1863,1887]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="84" phylum="Tracheophyta" rank="species" species="lanigera">
<emphasis id="B97EC16A7E3EFF9CFD6FF8C0FCEFAD54" box="[714,830,1863,1887]" italics="true" pageId="5" pageNumber="84">P. lanigera</emphasis>
</taxonomicName>
(
<figureCitation id="133101FD7E3EFF9CFCE8F8CFFC4FAD54" box="[845,926,1863,1887]" captionStart="FIGURE 3" captionStartId="3.[136,229,906,928]" captionTargetBox="[291,1296,197,874]" captionTargetId="figure-18@3.[285,1302,190,881]" captionTargetPageId="3" captionText="FIGURE 3. Detail of the flower. A. Pleurothallis bilobulata. B. Pleurothallis lanigera. Photos by M. Jiménez." figureDoi="http://doi.org/10.5281/zenodo.5447965" httpUri="https://zenodo.org/record/5447965/files/figure.png" pageId="5" pageNumber="84">Fig. 3B</figureCitation>
) in the pubescent sepals that are similar in length to the petals and the papillose lip. Both occur in southeastern 
<collectingCountry id="F31D5DE87E3EFF9CFC89F8E3FC54AD88" box="[812,901,1899,1923]" name="Ecuador" pageId="5" pageNumber="84">Ecuador</collectingCountry>
and may be found growing in sympatry at around 
<quantity id="4CF2B09D7E3EFF9CFF2DF807FF0AADAC" box="[136,219,1935,1959]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="84" unit="m" value="1000.0">1000 m</quantity>
near Zamora. The lack of intermediate specimens suggests they are reproductively isolated and don’t hybridize. 
<taxonomicName id="4C0A66FB7E3EFF9CFF2DF83BFE57ADC0" authorityName="M. M. Jimenez, Ocupa &amp; Velez-Abarca" authorityYear="2021" box="[136,390,1971,1995]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="84" phylum="Tracheophyta" rank="species" species="bilobulata">
<emphasis id="B97EC16A7E3EFF9CFF2DF83BFE57ADC0" box="[136,390,1971,1995]" italics="true" pageId="5" pageNumber="84">Pleurothallis bilobulata</emphasis>
</taxonomicName>
can be distinguished from 
<taxonomicName id="4C0A66FB7E3EFF9CFD07F83CFCC4ADC0" box="[674,789,1971,1995]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="84" phylum="Tracheophyta" rank="species" species="lanigera">
<emphasis id="B97EC16A7E3EFF9CFD07F83CFCC4ADC0" box="[674,789,1971,1995]" italics="true" pageId="5" pageNumber="84">P. lanigera</emphasis>
</taxonomicName>
by the narrowly ovate leaves, the straight, subverrucose ovary, the broadly elliptic dorsal sepal, the shortly acuminate, narrowly lanceolate, papillose-verrucose, 1-veined petals and the lip with a basal, bilobed glenion (
<figureCitation id="133101FD7E3EFF9CFE6FF873FDC4A218" box="[458,533,2043,2067]" captionStart="FIGURE 4" captionStartId="3.[136,229,2028,2050]" captionTargetBox="[289,1299,959,1998]" captionTargetId="figure-41@3.[283,1305,953,2004]" captionTargetPageId="3" captionText="FIGURE 4. Lip, column and ovary of P. bilobulata (left) and P. lanigera (right). A. Frontal view of the lip. B. Lateral view of the column, lip and ovary. Photos by M. Jiménez." figureDoi="http://doi.org/10.5281/zenodo.5447967" httpUri="https://zenodo.org/record/5447967/files/figure.png" pageId="5" pageNumber="84">Figs. 4</figureCitation>
, 
<figureCitation id="133101FD7E3EFF9CFD84F873FDE1A218" box="[545,560,2043,2067]" captionStart="FIGURE 5" captionStartId="4.[136,229,1271,1293]" captionTargetBox="[286,1302,196,1241]" captionTargetId="figure-22@4.[280,1308,190,1247]" captionTargetPageId="4" captionText="FIGURE 5. Column, anther and pollinarium of P. bilobulata (left) and P. lanigera (right). A. Frontal view of the column. B. Anther and pollinarium. Photos by M. Jiménez." figureDoi="http://doi.org/10.5281/zenodo.5447969" httpUri="https://zenodo.org/record/5447969/files/figure.png" pageId="5" pageNumber="84">5</figureCitation>
).
</paragraph>
<paragraph id="8BB51D787E3DFF9FFF18FF17FE77AB28" blockId="6.[136,1452,159,291]" pageId="6" pageNumber="85">
The bilobed glenion is an uncommon feature among the members of 
<taxonomicName id="4C0A66FB7E3DFF9FFC36FF17FBCCAABC" authorityName=", Lindley" authorityYear="1859" box="[915,1053,159,183]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="85" phylum="Tracheophyta" rank="genus">
<emphasis id="B97EC16A7E3DFF9FFC36FF17FBCCAABC" box="[915,1053,159,183]" italics="true" pageId="6" pageNumber="85">Pleurothallis</emphasis>
</taxonomicName>
subsect. 
<emphasis id="B97EC16A7E3DFF9FFBD8FF17FA75AABC" box="[1149,1444,159,183]" italics="true" pageId="6" pageNumber="85">Macrophyllae-Fasciculatae</emphasis>
. It is only known to occur in the distantly related 
<taxonomicName id="4C0A66FB7E3DFF9FFD19FF4BFB00AAD0" authority="Schlechter (1922: 22)" authorityName="Schlechter" authorityPageNumber="22" authorityYear="1922" box="[700,1233,195,219]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="85" phylum="Tracheophyta" rank="species" species="rhodoglossa">
<emphasis id="B97EC16A7E3DFF9FFD19FF4BFC05AAD0" box="[700,980,195,219]" italics="true" pageId="6" pageNumber="85">Pleurothallis rhodoglossa</emphasis>
<bibRefCitation id="EF9B60897E3DFF9FFC45FF4BFB00AAD0" author="Schlechter, R." box="[992,1233,195,219]" pageId="6" pageNumber="85" pagination="22 - 23" refId="ref3416" refString="Schlechter, R. (1922) Beitrage Zur Orchideenkunde von Zentralamerika. I. Orchidaceae Powellianae Panamenses. Repertorium Specierum Novarum Regni Vegetabilis, Beihefte. Band 17: 22 - 23." type="book chapter" year="1922">Schlechter (1922: 22)</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C0A66FB7E3DFF9FFAA8FF4CFDC2AAF4" authority="M. M. Jimenez, Tobar &amp; Mark Wilson" authorityName="M. M. Jimenez, Tobar &amp; Mark Wilson" authorityYear="2016" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="85" phylum="Tracheophyta" rank="species" species="nangaritzae">
<emphasis id="B97EC16A7E3DFF9FFAA8FF4CFA7AAAD0" box="[1293,1451,196,219]" italics="true" pageId="6" pageNumber="85">P. nangaritzae</emphasis>
M.M.Jiménez, Tobar &amp; Mark Wilson
</taxonomicName>
in 
<bibRefCitation id="EF9B60897E3DFF9FFD96FF6FFCEEAAF4" author="Wilson, M. &amp; Baquero, L. &amp; Dupree, K. &amp; Jimenez, M. M. &amp; LeBlanc, C. M. &amp; Merino, G. &amp; Portilla, J. &amp; Salas Guerrero, M. &amp; Tobar Suarez, F. &amp; Werner, J. D." box="[563,831,231,255]" pageId="6" pageNumber="85" pagination="349 - 350" refId="ref3807" refString="Wilson, M., Baquero, L., Dupree, K., Jimenez, M. M., LeBlanc, C. M., Merino, G., Portilla, J., Salas Guerrero, M., Tobar Suarez, F., Werner, J. D. (2016) Three new species of Pleurothallis (Orchidaceae: Pleurothallidinae) in subsection Macrophyllae-Fasciculatae from northern South America. Lankesteriana 16: 349 - 350. https: // doi. org / 10.15517 / lank. v 16 i 3.27314" type="journal article" year="2016">
Wilson 
<emphasis id="B97EC16A7E3DFF9FFD20FF60FD6AAAF4" box="[645,699,231,255]" italics="true" pageId="6" pageNumber="85">et al.</emphasis>
(2016: 358)
</bibRefCitation>
, and the closely related 
<taxonomicName id="4C0A66FB7E3DFF9FFB9BFF60FF7DAB28" authority="Reichenbach (1854: 26)" authorityName="Reichenbach" authorityPageNumber="26" authorityYear="1854" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="85" phylum="Tracheophyta" rank="species" species="cardiostola">
<emphasis id="B97EC16A7E3DFF9FFB9BFF60FB01AAF4" box="[1086,1232,231,255]" italics="true" pageId="6" pageNumber="85">P. cardiostola</emphasis>
<bibRefCitation id="EF9B60897E3DFF9FFB73FF6FFF7DAB28" author="Reichenbach, H. G." pageId="6" pageNumber="85" pagination="9 - 26" refId="ref3248" refString="Reichenbach, H. G. (1854) Die Wagener'schen Orchideen. Bonplandia, Zeitschrift fur die gesammte Botanik 2: 9 - 26." type="journal article" year="1854">Reichenbach (1854: 26)</bibRefCitation>
</taxonomicName>
(Wilson, 
<emphasis id="B97EC16A7E3DFF9FFEB1FE84FE46AB28" box="[276,407,268,291]" italics="true" pageId="6" pageNumber="85">pers. comm.</emphasis>
).
</paragraph>
</subSubSection>
</treatment>
</document>