124 lines
24 KiB
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124 lines
24 KiB
XML
<document id="F80588A70238D44F969DE144B20B3393" ID-DOI="10.1111/j.1096-3642.2011.00725.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5441985" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630452374775" checkinUser="valdenar" docAuthor="Clarke, Dave J." docDate="2011" docId="039B414F195CFFDBFEC2FDB54829FB79" docLanguage="en" docName="j.1096-3642.2011.00725.x.pdf" docOrigin="Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 163 (3)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2011.00725.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Creophilus Leach 1819" docType="treatment" docVersion="5" lastPageNumber="755" masterDocId="FFA23937197CFFFBFFE7FFFA4A24FFAB" masterDocTitle="Testing the phylogenetic utility of morphological character systems, with a revision of Creophilus Leach (Coleoptera: Staphylinidae)" masterLastPageNumber="812" masterPageNumber="723" pageNumber="755" updateTime="1699275617143" updateUser="plazi">
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<mods:titleInfo id="1ED23477B91E9673DF891F3A7F77A20F">
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<mods:title id="025B54A752B00935FD9AC6C8E4076DE9">Testing the phylogenetic utility of morphological character systems, with a revision of Creophilus Leach (Coleoptera: Staphylinidae)</mods:title>
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<mods:namePart id="601312A337003688CDAC8959A5D23771">Clarke, Dave J.</mods:namePart>
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<mods:title id="B0FCDA88EC707FE6595DFA0CC1308910">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="B16105282CF06DF4487155C653BB3FDF">2011</mods:date>
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<mods:title id="EBC5F0427EBD627B861DF2FEDB62D59D">Zool. J. Linn. Soc.</mods:title>
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<mods:number id="2BF7BCB6117212DC88EA09C6137C37CC">2011-10-25</mods:number>
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<mods:number id="C8D1B9ACCEE741BABB2F04519379076A">163</mods:number>
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<mods:number id="CDC9ABD14661AA50ADF783B1B6481E94">3</mods:number>
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<mods:identifier id="90AD2FE953970B5ABB107C71106EEF76" type="ISSN">0024-4082</mods:identifier>
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<treatment id="039B414F195CFFDBFEC2FDB54829FB79" ID-DOI="http://doi.org/10.5281/zenodo.5492127" ID-Zenodo-Dep="5492127" LSID="urn:lsid:plazi:treatment:039B414F195CFFDBFEC2FDB54829FB79" httpUri="http://treatment.plazi.org/id/039B414F195CFFDBFEC2FDB54829FB79" lastPageNumber="755" pageId="32" pageNumber="755">
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<subSubSection id="C328A3D2195CFFDBFEC2FDB54845FDC8" box="[293,609,590,612]" pageId="32" pageNumber="755" type="nomenclature">
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<paragraph id="8B8DF059195CFFDBFEC2FDB54845FDC8" blockId="32.[293,609,590,612]" box="[293,609,590,612]" pageId="32" pageNumber="755">
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<heading id="D0C54735195CFFDBFEC2FDB54845FDC8" box="[293,609,590,612]" centered="true" fontSize="8" level="2" pageId="32" pageNumber="755" reason="8">
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THE
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<taxonomicName id="4C328BDA195CFFDBFEB8FDB44BCBFDC9" ID-CoL="8VRLQ" authorityName="Leach" authorityYear="1819" box="[351,495,590,611]" class="Insecta" family="Staphylinidae" genus="Creophilus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="755" phylum="Arthropoda" rank="genus">
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<emphasis id="B9462C4B195CFFDBFEB8FDB44BCBFDC9" box="[351,495,590,611]" italics="true" pageId="32" pageNumber="755">CREOPHILUS</emphasis>
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</taxonomicName>
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COMPLEX
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C328A3D2195CFFDBFF77FD8F4829FB79" pageId="32" pageNumber="755" type="diagnosis">
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<paragraph id="8B8DF059195CFFDBFF77FD8F4B9AFB9D" blockId="32.[144,759,629,1079]" pageId="32" pageNumber="755">
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<emphasis id="B9462C4B195CFFDBFF77FD8F4B6BFD21" box="[144,335,629,650]" italics="true" pageId="32" pageNumber="755">Adult diagnosis:</emphasis>
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Species of the
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<taxonomicName id="4C328BDA195CFFDBFDF6FD8F48A8FD21" authorityName="Leach" authorityYear="1819" box="[529,652,629,650]" class="Insecta" family="Staphylinidae" genus="Creophilus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="755" phylum="Arthropoda" rank="genus">
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<emphasis id="B9462C4B195CFFDBFDF6FD8F48A8FD21" box="[529,652,629,650]" italics="true" pageId="32" pageNumber="755">Creophilus</emphasis>
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</taxonomicName>
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complex can be recognized by the combination of: disc of head and pronotum glabrous; ventroscapal macrosetae absent or reduced; superior marginal pronotal carina separate from inferior marginal carina along entire length, effaced apically (
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<figureCitation id="1309ECDC195CFFDBFE56FCF44822FC88" box="[433,518,782,804]" captionStart="Figure 2" captionStartId="10.[144,223,198,217]" captionText="Figure 2. Creophilus, details of morphology (setae omitted except in H–K). C. erythrocephalus (A-B, L, large male); C. variegatus (C, D, large male); C. maxillosus maxillosus (E–G); C. lanio (H–J); C. incanus (K). A, head, ventral (mouthparts removed); B, head, dorsal (antennae removed); C, left mandible, lateral and ventral (prostheca omitted); D, mandibles, dorsal (prostheca omitted); E, prothorax, ventral (small male); F, pterothorax, ventral; G, pronotum, dorsal (large male); H, left maxilla, ventral; I, labium, ventral; J, abdominal tergite II, dorsal; K, right protibia, anterior (smaller setae/bristles omitted); L, abdominal segments III–V, dorsal. Abbreviations: af, anterolateral flange of abdominal tergite II; atf, antennal fossa; bgi, basal gular impression; bpd, basolateral pronotal denticle; btf, basal transverse fold; dbl, dorsal basolateral ridge; dbr, dorsal basal ridge; f, muricate fossa of abdominal tergite II; HL, head length; il, inferior pronotal line; io, infraorbital ridge; mac, posterior marginal carina of mesocoxal acetabulum; mb, mandibular blade; mip, mesoventral intercoxal projection; ML, mandible length; mt, medial tumescence of abdominal tergite II; nr, nuchal ridge; pf, posteromedian furcasternal fovea; pg, postgenal ridge; plf, posterolateral fossa; pmg, premetacoxal groove; pos, postoccipital suture; ppp, pronotal postcoxal process; ps (fused) parasclerites of abdominal segment II; ptm, mesothoracic peritreme; sbr, secondary basal ridge; sl, superior pronotal line; spi, spiracle; sr, sternacoxal ridge; T1, T2, T3, mandibular teeth; td, transverse depression of abdominal tergite II; vb, ventral basal ridge; vbr, ventral basolateral ridge of left mandible; vl, ventrolateral carina. Scale bars = 1 mm." pageId="32" pageNumber="755">Fig. 2E</figureCitation>
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,
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<emphasis id="B9462C4B195CFFDBFDF1FCF44802FC88" box="[534,550,782,803]" italics="true" pageId="32" pageNumber="755">il</emphasis>
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,
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<emphasis id="B9462C4B195CFFDBFDDFFCF4486FFC88" box="[568,587,782,803]" italics="true" pageId="32" pageNumber="755">sl</emphasis>
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,
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<figureCitation id="1309ECDC195CFFDBFDBAFCF44889FC88" box="[605,685,782,804]" captionStart="Figure 9" captionStartId="20.[144,223,198,217]" captionText="Figure 9. Scanning electron micrographs of Creophilus imitator (A, B), C. maxillosus villosus (C–F, L, M), C. oculatus (J, K), Hadrotes crassus (G), Xanthopygus xanthopygus (H, I) and Staphylinus caesareus (N, O). A, vertex showing micropunctate microsculpture and shallow depressions (ch4-2, circle); B, same, detail of micropunctate microsculpture (ch5-2) and two shallow depressions each with associated pore and microseta (ch4-2, arrows); C, vertex showing small abrupt pits (ch4-1, arrow); D, left procoxa and femur of large male with basal femoral spine (ch36-1, arrow), ventrolateral; E, left protibia with mesal pair of protibial spines (ch37-1, arrows), posterior; F, prothorax with arcuate sternacoxal ridge (ch23-1, upper arrow) and medially extended mesothoracic peritremes (ch25-1, lower arrow), ventral; G, prothorax with anteromedially projecting sternacoxal ridge (ch23-0, arrow), ventral; H, prothorax showing anterior (left arrow), inferior (middle arrow), and superior (right arrow) pronotal marginal lines, ventral-oblique; I, pterothorax showing two distinct ridges of mesepimeron (ch27-0, arrows), lateral; J, prothorax showing anterior (left arrow), inferior (middle arrow), and superior (right arrow) pronotal marginal lines, lateral-oblique; K, pterothorax showing single distinct ridge of mesepimeron (ch27-1, arrow), lateral; L, left side of abdominal segment II showing position of one muricate fossa (ch41-1, arrow), dorsal; M, surface detail of vaginal plate; N, female internal genitalia showing sclerotized plate at opening of bursa copulatrix (ch87-1, left arrow), ciliose vaginal plate (ch85-0, middle arrow), and extensive muricate region (vaginal fold) distal to vaginal plate (right arrow); O, surface detail of K. Scale bars: B = 0.01 mm; A, C, L–O = 0.1 mm; F–K = 0.5 mm; D = 1 mm." pageId="32" pageNumber="755">Fig. 9J</figureCitation>
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, right and middle arrows); inferior marginal pronotal carina continuous with anterior marginal carina (
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<figureCitation id="1309ECDC195CFFDBFD46FCB148D5FCCB" box="[673,753,843,865]" captionStart="Figure 9" captionStartId="20.[144,223,198,217]" captionText="Figure 9. Scanning electron micrographs of Creophilus imitator (A, B), C. maxillosus villosus (C–F, L, M), C. oculatus (J, K), Hadrotes crassus (G), Xanthopygus xanthopygus (H, I) and Staphylinus caesareus (N, O). A, vertex showing micropunctate microsculpture and shallow depressions (ch4-2, circle); B, same, detail of micropunctate microsculpture (ch5-2) and two shallow depressions each with associated pore and microseta (ch4-2, arrows); C, vertex showing small abrupt pits (ch4-1, arrow); D, left procoxa and femur of large male with basal femoral spine (ch36-1, arrow), ventrolateral; E, left protibia with mesal pair of protibial spines (ch37-1, arrows), posterior; F, prothorax with arcuate sternacoxal ridge (ch23-1, upper arrow) and medially extended mesothoracic peritremes (ch25-1, lower arrow), ventral; G, prothorax with anteromedially projecting sternacoxal ridge (ch23-0, arrow), ventral; H, prothorax showing anterior (left arrow), inferior (middle arrow), and superior (right arrow) pronotal marginal lines, ventral-oblique; I, pterothorax showing two distinct ridges of mesepimeron (ch27-0, arrows), lateral; J, prothorax showing anterior (left arrow), inferior (middle arrow), and superior (right arrow) pronotal marginal lines, lateral-oblique; K, pterothorax showing single distinct ridge of mesepimeron (ch27-1, arrow), lateral; L, left side of abdominal segment II showing position of one muricate fossa (ch41-1, arrow), dorsal; M, surface detail of vaginal plate; N, female internal genitalia showing sclerotized plate at opening of bursa copulatrix (ch87-1, left arrow), ciliose vaginal plate (ch85-0, middle arrow), and extensive muricate region (vaginal fold) distal to vaginal plate (right arrow); O, surface detail of K. Scale bars: B = 0.01 mm; A, C, L–O = 0.1 mm; F–K = 0.5 mm; D = 1 mm." pageId="32" pageNumber="755">Fig. 9J</figureCitation>
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, left and middle arrows); mesepimeron with single marginal ridge (
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<figureCitation id="1309ECDC195CFFDBFEAEFC724BBBFC36" box="[329,415,904,926]" captionStart="Figure 9" captionStartId="20.[144,223,198,217]" captionText="Figure 9. Scanning electron micrographs of Creophilus imitator (A, B), C. maxillosus villosus (C–F, L, M), C. oculatus (J, K), Hadrotes crassus (G), Xanthopygus xanthopygus (H, I) and Staphylinus caesareus (N, O). A, vertex showing micropunctate microsculpture and shallow depressions (ch4-2, circle); B, same, detail of micropunctate microsculpture (ch5-2) and two shallow depressions each with associated pore and microseta (ch4-2, arrows); C, vertex showing small abrupt pits (ch4-1, arrow); D, left procoxa and femur of large male with basal femoral spine (ch36-1, arrow), ventrolateral; E, left protibia with mesal pair of protibial spines (ch37-1, arrows), posterior; F, prothorax with arcuate sternacoxal ridge (ch23-1, upper arrow) and medially extended mesothoracic peritremes (ch25-1, lower arrow), ventral; G, prothorax with anteromedially projecting sternacoxal ridge (ch23-0, arrow), ventral; H, prothorax showing anterior (left arrow), inferior (middle arrow), and superior (right arrow) pronotal marginal lines, ventral-oblique; I, pterothorax showing two distinct ridges of mesepimeron (ch27-0, arrows), lateral; J, prothorax showing anterior (left arrow), inferior (middle arrow), and superior (right arrow) pronotal marginal lines, lateral-oblique; K, pterothorax showing single distinct ridge of mesepimeron (ch27-1, arrow), lateral; L, left side of abdominal segment II showing position of one muricate fossa (ch41-1, arrow), dorsal; M, surface detail of vaginal plate; N, female internal genitalia showing sclerotized plate at opening of bursa copulatrix (ch87-1, left arrow), ciliose vaginal plate (ch85-0, middle arrow), and extensive muricate region (vaginal fold) distal to vaginal plate (right arrow); O, surface detail of K. Scale bars: B = 0.01 mm; A, C, L–O = 0.1 mm; F–K = 0.5 mm; D = 1 mm." pageId="32" pageNumber="755">Fig. 9K</figureCitation>
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, arrow); male sternite IX subtruncate to shallowly emarginate apically, produced into elongate process basally (
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<figureCitation id="1309ECDC195CFFDBFE01FC3F4818FC70" box="[486,572,965,987]" captionStart="Figure 3" captionStartId="12.[144,223,198,217]" captionText="Figure 3. Male genitalia, details of morphology. Creophilus erythrocephalus (A, B, G, H, O); C. lanio (Q); C. maxillosus maxillosus (C, D, I, P); C. oculatus (J); Hadropinus fossor (E, F); Staphylinus caesareus (K, L); Hadrotes wakefieldi (M); Dinothenarus badipes (N). A, aedeagus with details of inverted internal sac, ventral; B, same, right lateral (parameral setae omitted); C, median lobe with fully everted internal sac, ventral – arrow indicates membranous region; D, apical part of median lobe with fully everted internal sac, right lateral; E, aedeagus with fully everted internal sac, right lateral (parameral setae omitted), note: bit has become separated from cp in this specimen; F, copulatory piece and appendix of internal sac, dorsal; G, ventral sclerite of internal sac, ventral; H, apex of median lobe with fully everted internal sac, right lateral; I, ventral sclerite of internal sac, ventral (lower) and apical (upper); J, apical part of median lobe with partially everted internal sac, ventral; K, apical part of median lobe showing part of everted internal sac, right lateral; L–N, paramere, dorsal; O–P, median lobe apex, ventral; Q, male genital segment (setae omitted). Abbreviations: app, appendix; as, apicolateral sclerite of median lobe; bc, basal cap; bit, basal inner tube; bms, basomedian sclerite of internal sac; ce, carinate edge of median orifice; cp, copulatory piece; dr, dorsal receptacle for copulatory piece/appendix; is, internal sac; lit, lateral inflexed tooth of median lobe; ml, membranous lobe; mlb, median lobe of aedeagus; stm, subtriangular membranous region; vp, ventral process; vr, ventromedian spiculose strip; vs, ventral sclerite; sh, spinose sheath surrounding copulatory piece; sr, one of pair of basoventral spiculose regions. Scale bars: G and I = 0.1 mm; rest = 1 mm. ·" pageId="32" pageNumber="755">Fig. 3Q</figureCitation>
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); median lobe of aedeagus with large apicolateral sclerites (
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<figureCitation id="1309ECDC195CFFDBFD99FC1E48E4FC52" box="[638,704,996,1018]" captionStart="Figure 3" captionStartId="12.[144,223,198,217]" captionText="Figure 3. Male genitalia, details of morphology. Creophilus erythrocephalus (A, B, G, H, O); C. lanio (Q); C. maxillosus maxillosus (C, D, I, P); C. oculatus (J); Hadropinus fossor (E, F); Staphylinus caesareus (K, L); Hadrotes wakefieldi (M); Dinothenarus badipes (N). A, aedeagus with details of inverted internal sac, ventral; B, same, right lateral (parameral setae omitted); C, median lobe with fully everted internal sac, ventral – arrow indicates membranous region; D, apical part of median lobe with fully everted internal sac, right lateral; E, aedeagus with fully everted internal sac, right lateral (parameral setae omitted), note: bit has become separated from cp in this specimen; F, copulatory piece and appendix of internal sac, dorsal; G, ventral sclerite of internal sac, ventral; H, apex of median lobe with fully everted internal sac, right lateral; I, ventral sclerite of internal sac, ventral (lower) and apical (upper); J, apical part of median lobe with partially everted internal sac, ventral; K, apical part of median lobe showing part of everted internal sac, right lateral; L–N, paramere, dorsal; O–P, median lobe apex, ventral; Q, male genital segment (setae omitted). Abbreviations: app, appendix; as, apicolateral sclerite of median lobe; bc, basal cap; bit, basal inner tube; bms, basomedian sclerite of internal sac; ce, carinate edge of median orifice; cp, copulatory piece; dr, dorsal receptacle for copulatory piece/appendix; is, internal sac; lit, lateral inflexed tooth of median lobe; ml, membranous lobe; mlb, median lobe of aedeagus; stm, subtriangular membranous region; vp, ventral process; vr, ventromedian spiculose strip; vs, ventral sclerite; sh, spinose sheath surrounding copulatory piece; sr, one of pair of basoventral spiculose regions. Scale bars: G and I = 0.1 mm; rest = 1 mm. ·" pageId="32" pageNumber="755">Fig. 3</figureCitation>
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,
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<emphasis id="B9462C4B195CFFDBFD29FC1E48C3FC52" box="[718,743,996,1017]" italics="true" pageId="32" pageNumber="755">as</emphasis>
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); internal sac without ventral basomedian sclerotized strip (as in
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<figureCitation id="1309ECDC195CFFDBFEFCFBDB4B55FB9D" box="[283,369,1057,1079]" captionStart="Figure 3" captionStartId="12.[144,223,198,217]" captionText="Figure 3. Male genitalia, details of morphology. Creophilus erythrocephalus (A, B, G, H, O); C. lanio (Q); C. maxillosus maxillosus (C, D, I, P); C. oculatus (J); Hadropinus fossor (E, F); Staphylinus caesareus (K, L); Hadrotes wakefieldi (M); Dinothenarus badipes (N). A, aedeagus with details of inverted internal sac, ventral; B, same, right lateral (parameral setae omitted); C, median lobe with fully everted internal sac, ventral – arrow indicates membranous region; D, apical part of median lobe with fully everted internal sac, right lateral; E, aedeagus with fully everted internal sac, right lateral (parameral setae omitted), note: bit has become separated from cp in this specimen; F, copulatory piece and appendix of internal sac, dorsal; G, ventral sclerite of internal sac, ventral; H, apex of median lobe with fully everted internal sac, right lateral; I, ventral sclerite of internal sac, ventral (lower) and apical (upper); J, apical part of median lobe with partially everted internal sac, ventral; K, apical part of median lobe showing part of everted internal sac, right lateral; L–N, paramere, dorsal; O–P, median lobe apex, ventral; Q, male genital segment (setae omitted). Abbreviations: app, appendix; as, apicolateral sclerite of median lobe; bc, basal cap; bit, basal inner tube; bms, basomedian sclerite of internal sac; ce, carinate edge of median orifice; cp, copulatory piece; dr, dorsal receptacle for copulatory piece/appendix; is, internal sac; lit, lateral inflexed tooth of median lobe; ml, membranous lobe; mlb, median lobe of aedeagus; stm, subtriangular membranous region; vp, ventral process; vr, ventromedian spiculose strip; vs, ventral sclerite; sh, spinose sheath surrounding copulatory piece; sr, one of pair of basoventral spiculose regions. Scale bars: G and I = 0.1 mm; rest = 1 mm. ·" pageId="32" pageNumber="755">Fig. 3K</figureCitation>
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,
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<emphasis id="B9462C4B195CFFDBFE67FBDB4B8AFB9D" box="[384,430,1057,1078]" italics="true" pageId="32" pageNumber="755">bms</emphasis>
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).
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</paragraph>
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<paragraph id="8B8DF059195CFFDBFF77FB9A4829FB79" blockId="32.[144,759,1120,1234]" pageId="32" pageNumber="755">
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<emphasis id="B9462C4B195CFFDBFF77FB9A4ADFFBDE" box="[144,251,1120,1141]" italics="true" pageId="32" pageNumber="755">Remarks:</emphasis>
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The following taxa are here included in the
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<taxonomicName id="4C328BDA195CFFDBFF77FB844B2FFB38" authorityName="Leach" authorityYear="1819" box="[144,267,1150,1171]" class="Insecta" family="Staphylinidae" genus="Creophilus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="755" phylum="Arthropoda" rank="genus">
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<emphasis id="B9462C4B195CFFDBFF77FB844B2FFB38" box="[144,267,1150,1171]" italics="true" pageId="32" pageNumber="755">Creophilus</emphasis>
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</taxonomicName>
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complex:
|
||
<taxonomicName id="4C328BDA195CFFDBFE9BFB844BD3FB38" authorityName="Leach" authorityYear="1819" box="[380,503,1150,1171]" class="Insecta" family="Staphylinidae" genus="Creophilus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="755" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9462C4B195CFFDBFE9BFB844BD3FB38" box="[380,503,1150,1171]" italics="true" pageId="32" pageNumber="755">Creophilus</emphasis>
|
||
</taxonomicName>
|
||
(12 species),
|
||
<taxonomicName id="4C328BDA195CFFDBFD68FB844AC0FB18" baseAuthorityName="Moore" baseAuthorityYear="1964" class="Insecta" family="Staphylinidae" genus="Hadrotes" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="755" phylum="Arthropoda" rank="species" species="crassus">
|
||
<emphasis id="B9462C4B195CFFDBFD68FB844AC0FB18" italics="true" pageId="32" pageNumber="755">Hadrotes crassus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C328BDA195CFFDBFF1CFB644B86FB19" authorityName="Cameron" authorityYear="1945" box="[251,418,1181,1203]" class="Insecta" family="Staphylinidae" genus="Hadrotes" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="755" phylum="Arthropoda" rank="species" species="wakefieldi">
|
||
<emphasis id="B9462C4B195CFFDBFF1CFB644B86FB19" box="[251,418,1181,1203]" italics="true" pageId="32" pageNumber="755">Ht. wakefieldi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C328BDA195CFFDBFE5DFB6748AEFB19" box="[442,650,1181,1202]" class="Insecta" family="Staphylinidae" genus="Hadropinus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="755" phylum="Arthropoda" rank="species" species="fosso">
|
||
<emphasis id="B9462C4B195CFFDBFE5DFB6748AEFB19" box="[442,650,1181,1202]" italics="true" pageId="32" pageNumber="755">Hadropinus fosso</emphasis>
|
||
</taxonomicName>
|
||
r,
|
||
<taxonomicName id="4C328BDA195CFFDBFD4DFB6448D3FB18" authorityName="Sharp" authorityYear="1889" box="[682,759,1182,1203]" class="Insecta" family="Staphylinidae" genus="Liusus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="755" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9462C4B195CFFDBFD4DFB6448D3FB18" box="[682,759,1182,1203]" italics="true" pageId="32" pageNumber="755">Liusus</emphasis>
|
||
</taxonomicName>
|
||
(two species),
|
||
<taxonomicName id="4C328BDA195CFFDBFED3FB46482CFB79" baseAuthorityName="Boving & Craighead" baseAuthorityYear="1931" box="[308,520,1212,1234]" class="Insecta" family="Staphylinidae" genus="Thinopinus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="811" phylum="Arthropoda" rank="species" species="pictus">
|
||
<emphasis id="B9462C4B195CFFDBFED3FB464B93FB7A" box="[308,439,1212,1233]" italics="true" pageId="32" pageNumber="755">Thinopinus</emphasis>
|
||
pictus
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |