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<document id="2D9AE71691FF5E28658311F9386A6315" ID-CLB-Dataset="53812" ID-DOI="10.11646/zootaxa.5250.1.1" ID-GBIF-Dataset="da8acfed-df61-403e-9a49-7f856ce56196" ID-ISSN="1175­5334" ID-Zenodo-Dep="7705657" ID-ZooBank="4729EAF3-8B78-4AAD-B373-9179D008F239" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="felipe" checkinTime="1678178160677" checkinUser="plazi" docAuthor="WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI &amp; HUANG, JIA" docDate="2023" docId="0393914C2A7F8726C282FA316CDEF9C7" docLanguage="en" docName="zootaxa.5250.1.1.pdf" docOrigin="Zootaxa 5250 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5250.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Stegana (Steganina) sciophila Li &amp; Chen, sp. nov." docType="treatment" docVersion="10" lastPageNumber="95" masterDocId="FFAAE9342A228778C215FF896958FB43" masterDocTitle="Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae)" masterLastPageNumber="109" masterPageNumber="1" pageNumber="94" updateTime="1699476226253" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2E5AFD108932EFE0C472106D12295C19">Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae)</mods:title>
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<mods:namePart id="94B62C2B72E031EA5CCDDE1EC80A46E7">LI, BING-XUE</mods:namePart>
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<mods:namePart id="D940826B5281CFB8D1B231849E71F48A">CUI, MENG-DI</mods:namePart>
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<mods:namePart id="ECE3F84E225EE7B3B55D1E9590F9FACD">ZHANG, YUAN</mods:namePart>
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<mods:namePart id="E471778982EA38FAD2F552636BBB4E2B">WANG, LE</mods:namePart>
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<mods:namePart id="A9CD31FE8C9A74BED2372D8AF10982F2">ZHANG, CAI-HONG</mods:namePart>
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<mods:namePart id="C12BEADC4BF93DF5CEDE44F1C49CDD93">TSAUR, SHUN-CHERN</mods:namePart>
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<mods:namePart id="D750D4B0AEB845FF4900C77CD7CB4FE6">CHEN, HONG-WEI</mods:namePart>
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<mods:namePart id="98D7848961C30651EE7448C9D6BAE37B">HUANG, JIA</mods:namePart>
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<mods:title id="52737A092053971FBD3070EC168D10DE">Zootaxa</mods:title>
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<treatment id="0393914C2A7F8726C282FA316CDEF9C7" ID-DOI="http://doi.org/10.5281/zenodo.7705618" ID-GBIF-Taxon="209579556" ID-Zenodo-Dep="7705618" LSID="urn:lsid:plazi:treatment:0393914C2A7F8726C282FA316CDEF9C7" httpUri="http://treatment.plazi.org/id/0393914C2A7F8726C282FA316CDEF9C7" lastPageId="94" lastPageNumber="95" pageId="93" pageNumber="94">
<subSubSection id="C32073D12A7F8725C282FA316BA4FE91" box="[151,764,1464,1491]" pageId="93" pageNumber="94" type="nomenclature">
<paragraph id="8B85205A2A7F8725C282FA316BA4FE91" blockId="93.[151,764,1464,1527]" box="[151,764,1464,1491]" pageId="93" pageNumber="94">
<heading id="D0CD97362A7F8725C282FA316BA4FE91" bold="true" box="[151,764,1464,1491]" fontSize="11" level="1" pageId="93" pageNumber="94" reason="1">
<emphasis id="B94EFC482A7F8725C282FA316BA4FE91" bold="true" box="[151,764,1464,1491]" pageId="93" pageNumber="94">
<taxonomicName id="4C3A5BD92A7F8725C282FA316BC9FE91" ID-CoL="BDMSP" authority="Li &amp; Chen" authorityName="Li &amp; Chen" box="[151,657,1464,1491]" class="Insecta" family="Drosophilidae" genus="Stegana" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="sciophila" status="sp. nov." subGenus="Steganina">
<emphasis id="B94EFC482A7F8725C282FA3169A1FE91" bold="true" box="[151,249,1464,1490]" italics="true" pageId="93" pageNumber="94">Stegana</emphasis>
(
<emphasis id="B94EFC482A7F8725C31FFA3168DCFE91" bold="true" box="[266,388,1464,1490]" italics="true" pageId="93" pageNumber="94">Steganina</emphasis>
)
<emphasis id="B94EFC482A7F8725C380FA316B5AFE90" bold="true" box="[405,514,1464,1491]" italics="true" pageId="93" pageNumber="94">sciophila</emphasis>
Li &amp; Chen
</taxonomicName>
,
<taxonomicNameLabel id="A27D41332A7F8725C08BFA316BA4FE91" box="[670,764,1464,1490]" pageId="93" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C32073D12A7F8725C282FA556827FEB4" box="[151,383,1500,1527]" pageId="93" pageNumber="94" type="description">
<paragraph id="8B85205A2A7F8725C282FA556827FEB4" blockId="93.[151,764,1464,1527]" box="[151,383,1500,1527]" pageId="93" pageNumber="94">
(
<figureCitation id="13013CDF2A7F8725C28AFA556816FEB5" box="[159,334,1500,1526]" captionStart="FIGURE 13" captionStartId="36.[151,250,1943,1967]" captionTargetBox="[186,1415,181,1919]" captionTargetId="figure-30@36.[173,1415,181,1919]" captionTargetPageId="36" captionText="FIGURE 13. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) rava Cui &amp; Chen, sp. nov.; C, D. S. (S.) sciophila Li &amp; Chen, sp. nov.; E, F. S. (S.) septencolorata Li &amp; Chen, sp. nov.; G, H. S. (S.) silvestrella Zhang &amp; Chen, sp. nov.; I, J. S. (S.) simola Cui &amp; Chen, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7705693" httpUri="https://zenodo.org/record/7705693/files/figure.png" pageId="93" pageNumber="94">Figs. 13C, 13D</figureCitation>
,
<figureCitation id="13013CDF2A7F8725C34FFA55682FFEB4" box="[346,375,1500,1527]" captionStart="FIGURE 59" captionStartId="94.[151,250,1540,1564]" captionTargetBox="[159,1430,675,1511]" captionTargetId="figure-250@94.[151,1436,668,1516]" captionTargetPageId="94" captionText="FIGURE 59. Stegana (Steganina) sciophila Li &amp; Chen, sp. nov., male terminalia.A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F. Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705812" httpUri="https://zenodo.org/record/7705812/files/figure.png" pageId="93" pageNumber="94">59</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32073D12A7F8725C282F9AD689EFDE8" pageId="93" pageNumber="94" type="diagnosis">
<paragraph id="8B85205A2A7F8725C282F9AD689EFDE8" blockId="93.[151,1437,1572,1923]" pageId="93" pageNumber="94">
<emphasis id="B94EFC482A7F8725C282F9AD684AFD7D" bold="true" box="[151,274,1572,1598]" pageId="93" pageNumber="94">Diagnosis.</emphasis>
This species is similar to
<emphasis id="B94EFC482A7F8725C020F9AC6B1BFD7C" box="[565,579,1573,1599]" italics="true" pageId="93" pageNumber="94">S</emphasis>
.
<taxonomicName id="4C3A5BD92A7F8725C044F9AD6D4AFD7D" authority="Wang, Gao, &amp; Chen, 2013" authorityName="Wang, Gao, &amp; Chen" authorityYear="2013" box="[593,1042,1572,1599]" class="Insecta" family="Drosophilidae" genus="Stegana" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="hirticeps" subGenus="Steganina">
(
<emphasis id="B94EFC482A7F8725C04FF9AC6B30FD7C" box="[602,616,1573,1599]" italics="true" pageId="93" pageNumber="94">S</emphasis>
.)
<emphasis id="B94EFC482A7F8725C095F9AD6BBAFD7D" box="[640,738,1572,1598]" italics="true" pageId="93" pageNumber="94">hirticeps</emphasis>
Wang, Gao, &amp; Chen,
<quantity id="4CC28DBF2A7F8725C1CFF9AD6D4AFD7D" box="[986,1042,1572,1598]" metricMagnitude="1" metricUnit="m" metricValue="5.11302" pageId="93" pageNumber="94" unit="in" value="2013.0">2013</quantity>
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in the shape of male terminalia (as fig.
<quantity id="4CC28DBF2A7F8725C2D1F9C169A8FD21" box="[196,240,1608,1635]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="93" pageNumber="94" unit="in" value="5.0">5 in</quantity>
<bibRefCitation id="EFAB5DAB2A7F8725C2E3F9C068E0FD21" author="Wang, L. &amp; Gao, J. J. &amp; Chen, H. W." box="[246,440,1608,1635]" pageId="93" pageNumber="94" pagination="1993 - 2013" refId="ref74574" refString="Wang, L., Gao, J. J. &amp; Chen, H. W. (2013) Ten new species of the subgenus Steganina (Diptera: Drosophilidae: Stegana) from southern China. Journal of Natural History, 47, 1993 - 2013. https: // doi. org / 10.1080 / 00222933.2012.763055" type="journal article" year="2013">
Wang
<emphasis id="B94EFC482A7F8725C32BF9C0682EFD21" box="[318,374,1608,1634]" italics="true" pageId="93" pageNumber="94">et al.</emphasis>
2013
</bibRefCitation>
,
<figureCitation id="13013CDF2A7F8725C3D6F9C16B4DFD20" box="[451,533,1608,1635]" captionStart="FIGURE 59" captionStartId="94.[151,250,1540,1564]" captionTargetBox="[159,1430,675,1511]" captionTargetId="figure-250@94.[151,1436,668,1516]" captionTargetPageId="94" captionText="FIGURE 59. Stegana (Steganina) sciophila Li &amp; Chen, sp. nov., male terminalia.A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F. Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705812" httpUri="https://zenodo.org/record/7705812/files/figure.png" pageId="93" pageNumber="94">Fig. 59</figureCitation>
), but can be distinguished from the latter by having the surstylus with 1 peg-like tooth near distal margin (
<figureCitation id="13013CDF2A7F8725C3ACF9E56B1CFDC5" box="[441,580,1644,1671]" captionStart="FIGURE 59" captionStartId="94.[151,250,1540,1564]" captionTargetBox="[159,1430,675,1511]" captionTargetId="figure-250@94.[151,1436,668,1516]" captionTargetPageId="94" captionText="FIGURE 59. Stegana (Steganina) sciophila Li &amp; Chen, sp. nov., male terminalia.A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F. Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705812" httpUri="https://zenodo.org/record/7705812/files/figure.png" pageId="93" pageNumber="94">Fig. 59A, B</figureCitation>
); the subepandrial sclerite vaulted, bifurcated basally per side, roundly expanded distally (
<figureCitation id="13013CDF2A7F8725C344F91968E2FDE8" box="[337,442,1680,1707]" captionStart="FIGURE 59" captionStartId="94.[151,250,1540,1564]" captionTargetBox="[159,1430,675,1511]" captionTargetId="figure-250@94.[151,1436,668,1516]" captionTargetPageId="94" captionText="FIGURE 59. Stegana (Steganina) sciophila Li &amp; Chen, sp. nov., male terminalia.A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F. Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705812" httpUri="https://zenodo.org/record/7705812/files/figure.png" pageId="93" pageNumber="94">Fig. 59G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32073D12A7F8726C2D2F93D68FFFAEE" lastPageId="94" lastPageNumber="95" pageId="93" pageNumber="94" type="description">
<paragraph id="8B85205A2A7F8725C2D2F93D6C5BFCAD" blockId="93.[151,1437,1572,1923]" lastBlockId="93.[151,1437,1932,2031]" pageId="93" pageNumber="94">
<emphasis id="B94EFC482A7F8725C2D2F93D6801FD8D" bold="true" box="[199,345,1716,1742]" pageId="93" pageNumber="94">Description.</emphasis>
Male and female: First flagellomere brown. Clypeus yellow. Mesonotum dark brown medially, brownish laterally (
<figureCitation id="13013CDF2A7F8725C37AF951688CFDB1" box="[367,468,1752,1778]" captionStart="FIGURE 13" captionStartId="36.[151,250,1943,1967]" captionTargetBox="[186,1415,181,1919]" captionTargetId="figure-30@36.[173,1415,181,1919]" captionTargetPageId="36" captionText="FIGURE 13. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) rava Cui &amp; Chen, sp. nov.; C, D. S. (S.) sciophila Li &amp; Chen, sp. nov.; E, F. S. (S.) septencolorata Li &amp; Chen, sp. nov.; G, H. S. (S.) silvestrella Zhang &amp; Chen, sp. nov.; I, J. S. (S.) simola Cui &amp; Chen, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7705693" httpUri="https://zenodo.org/record/7705693/files/figure.png" pageId="93" pageNumber="94">Fig. 13C</figureCitation>
). Katepisternum yellowish (
<figureCitation id="13013CDF2A7F8725C118F9516A2BFDB1" box="[781,883,1752,1778]" captionStart="FIGURE 13" captionStartId="36.[151,250,1943,1967]" captionTargetBox="[186,1415,181,1919]" captionTargetId="figure-30@36.[173,1415,181,1919]" captionTargetPageId="36" captionText="FIGURE 13. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) rava Cui &amp; Chen, sp. nov.; C, D. S. (S.) sciophila Li &amp; Chen, sp. nov.; E, F. S. (S.) septencolorata Li &amp; Chen, sp. nov.; G, H. S. (S.) silvestrella Zhang &amp; Chen, sp. nov.; I, J. S. (S.) simola Cui &amp; Chen, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7705693" httpUri="https://zenodo.org/record/7705693/files/figure.png" pageId="93" pageNumber="94">Fig. 13D</figureCitation>
). Scutellum mostly yellow, brown anteromedially (
<figureCitation id="13013CDF2A7F8725C28AF975685DFC55" box="[159,261,1788,1814]" captionStart="FIGURE 13" captionStartId="36.[151,250,1943,1967]" captionTargetBox="[186,1415,181,1919]" captionTargetId="figure-30@36.[173,1415,181,1919]" captionTargetPageId="36" captionText="FIGURE 13. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) rava Cui &amp; Chen, sp. nov.; C, D. S. (S.) sciophila Li &amp; Chen, sp. nov.; E, F. S. (S.) septencolorata Li &amp; Chen, sp. nov.; G, H. S. (S.) silvestrella Zhang &amp; Chen, sp. nov.; I, J. S. (S.) simola Cui &amp; Chen, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7705693" httpUri="https://zenodo.org/record/7705693/files/figure.png" pageId="93" pageNumber="94">Fig. 13C</figureCitation>
). Halter mostly brownish (
<figureCitation id="13013CDF2A7F8725C039F9756BCBFC55" box="[556,659,1788,1814]" captionStart="FIGURE 13" captionStartId="36.[151,250,1943,1967]" captionTargetBox="[186,1415,181,1919]" captionTargetId="figure-30@36.[173,1415,181,1919]" captionTargetPageId="36" captionText="FIGURE 13. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) rava Cui &amp; Chen, sp. nov.; C, D. S. (S.) sciophila Li &amp; Chen, sp. nov.; E, F. S. (S.) septencolorata Li &amp; Chen, sp. nov.; G, H. S. (S.) silvestrella Zhang &amp; Chen, sp. nov.; I, J. S. (S.) simola Cui &amp; Chen, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7705693" httpUri="https://zenodo.org/record/7705693/files/figure.png" pageId="93" pageNumber="94">Fig. 13D</figureCitation>
). Fore femur with a row of 4 long setae on distal part of posteroventral surface. Abdominal tergites dark brown to black (
<figureCitation id="13013CDF2A7F8725C0AAF8A96A1DFC79" box="[703,837,1824,1850]" captionStart="FIGURE 13" captionStartId="36.[151,250,1943,1967]" captionTargetBox="[186,1415,181,1919]" captionTargetId="figure-30@36.[173,1415,181,1919]" captionTargetPageId="36" captionText="FIGURE 13. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) rava Cui &amp; Chen, sp. nov.; C, D. S. (S.) sciophila Li &amp; Chen, sp. nov.; E, F. S. (S.) septencolorata Li &amp; Chen, sp. nov.; G, H. S. (S.) silvestrella Zhang &amp; Chen, sp. nov.; I, J. S. (S.) simola Cui &amp; Chen, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7705693" httpUri="https://zenodo.org/record/7705693/files/figure.png" pageId="93" pageNumber="94">Fig. 13C, D</figureCitation>
); sternites brown in male, yellow in female, about 2.2 times as wide as long. Male terminalia (
<figureCitation id="13013CDF2A7F8725C047F8CD6BFEFC1C" box="[594,678,1860,1887]" captionStart="FIGURE 59" captionStartId="94.[151,250,1540,1564]" captionTargetBox="[159,1430,675,1511]" captionTargetId="figure-250@94.[151,1436,668,1516]" captionTargetPageId="94" captionText="FIGURE 59. Stegana (Steganina) sciophila Li &amp; Chen, sp. nov., male terminalia.A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F. Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705812" httpUri="https://zenodo.org/record/7705812/files/figure.png" pageId="93" pageNumber="94">Fig. 59</figureCitation>
): Epandrium with
<emphasis id="B94EFC482A7F8725C160F8CC6AD7FC1D" box="[885,911,1861,1886]" italics="true" pageId="93" pageNumber="94">ca</emphasis>
. 8 setae on dorsal to posterior portions per side (
<figureCitation id="13013CDF2A7F8725C28AF8E16850FCC0" box="[159,264,1896,1923]" captionStart="FIGURE 59" captionStartId="94.[151,250,1540,1564]" captionTargetBox="[159,1430,675,1511]" captionTargetId="figure-250@94.[151,1436,668,1516]" captionTargetPageId="94" captionText="FIGURE 59. Stegana (Steganina) sciophila Li &amp; Chen, sp. nov., male terminalia.A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F. Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705812" httpUri="https://zenodo.org/record/7705812/files/figure.png" pageId="93" pageNumber="94">Fig. 59A</figureCitation>
). Surstylus with some minute, serrated processes on outer surface and setulae near distal margin, lacking pubescence (
<figureCitation id="13013CDF2A7F8725C332F80568D6FCE4" box="[295,398,1932,1959]" captionStart="FIGURE 59" captionStartId="94.[151,250,1540,1564]" captionTargetBox="[159,1430,675,1511]" captionTargetId="figure-250@94.[151,1436,668,1516]" captionTargetPageId="94" captionText="FIGURE 59. Stegana (Steganina) sciophila Li &amp; Chen, sp. nov., male terminalia.A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F. Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705812" httpUri="https://zenodo.org/record/7705812/files/figure.png" pageId="93" pageNumber="94">Fig. 59A</figureCitation>
). Cercus lacking pubescence (
<figureCitation id="13013CDF2A7F8725C0C9F8056A1CFCE4" box="[732,836,1932,1959]" captionStart="FIGURE 59" captionStartId="94.[151,250,1540,1564]" captionTargetBox="[159,1430,675,1511]" captionTargetId="figure-250@94.[151,1436,668,1516]" captionTargetPageId="94" captionText="FIGURE 59. Stegana (Steganina) sciophila Li &amp; Chen, sp. nov., male terminalia.A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F. Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705812" httpUri="https://zenodo.org/record/7705812/files/figure.png" pageId="93" pageNumber="94">Fig. 59A</figureCitation>
). Hypandrium roundly expanded anteromedially (
<figureCitation id="13013CDF2A7F8725C77AF80569B4FC89" captionStart="FIGURE 59" captionStartId="94.[151,250,1540,1564]" captionTargetBox="[159,1430,675,1511]" captionTargetId="figure-250@94.[151,1436,668,1516]" captionTargetPageId="94" captionText="FIGURE 59. Stegana (Steganina) sciophila Li &amp; Chen, sp. nov., male terminalia.A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F. Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705812" httpUri="https://zenodo.org/record/7705812/files/figure.png" pageId="93" pageNumber="94">Fig. 59C, D</figureCitation>
). Pregonite small, with 2 sensilla (
<figureCitation id="13013CDF2A7F8725C065F8396BA2FC89" box="[624,762,1968,1995]" captionStart="FIGURE 59" captionStartId="94.[151,250,1540,1564]" captionTargetBox="[159,1430,675,1511]" captionTargetId="figure-250@94.[151,1436,668,1516]" captionTargetPageId="94" captionText="FIGURE 59. Stegana (Steganina) sciophila Li &amp; Chen, sp. nov., male terminalia.A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F. Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705812" httpUri="https://zenodo.org/record/7705812/files/figure.png" pageId="93" pageNumber="94">Fig. 59C, D</figureCitation>
). Aedeagus with much pubescence distomedially (
<figureCitation id="13013CDF2A7F8725C727F83969F7FCAD" captionStart="FIGURE 59" captionStartId="94.[151,250,1540,1564]" captionTargetBox="[159,1430,675,1511]" captionTargetId="figure-250@94.[151,1436,668,1516]" captionTargetPageId="94" captionText="FIGURE 59. Stegana (Steganina) sciophila Li &amp; Chen, sp. nov., male terminalia.A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F. Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705812" httpUri="https://zenodo.org/record/7705812/files/figure.png" pageId="93" pageNumber="94">Fig. 59C, D</figureCitation>
). Aedeagal sheath distally dorsad curved and with several minute, serrated processes (
<figureCitation id="13013CDF2A7F8725C666F85D6DAEFCAD" box="[1139,1270,2004,2031]" captionStart="FIGURE 59" captionStartId="94.[151,250,1540,1564]" captionTargetBox="[159,1430,675,1511]" captionTargetId="figure-250@94.[151,1436,668,1516]" captionTargetPageId="94" captionText="FIGURE 59. Stegana (Steganina) sciophila Li &amp; Chen, sp. nov., male terminalia.A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F. Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705812" httpUri="https://zenodo.org/record/7705812/files/figure.png" pageId="93" pageNumber="94">Fig. 59E, F</figureCitation>
).
</paragraph>
<paragraph id="8B85205A2A7C8726C2D2FF1F68FFFAEE" blockId="94.[151,1437,150,645]" pageId="94" pageNumber="95">
<emphasis id="B94EFC482A7C8726C2D2FF1F6B57FBF3" bold="true" box="[199,527,150,176]" pageId="94" pageNumber="95">Measurements and indices.</emphasis>
BL =
<quantity id="4CC28DBF2A7C8726C04EFF1F6B99FBF2" box="[603,705,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="3.07" pageId="94" pageNumber="95" unit="mm" value="3.07">3.07 mm</quantity>
in
<typeStatus id="54819EF82A7C8726C0FDFF1F6A12FBF3" box="[744,842,150,176]" pageId="94" pageNumber="95" type="holotype">holotype</typeStatus>
(
<quantity id="4CC28DBF2A7C8726C14FFF1F6AB3FBF3" box="[858,1003,150,177]" metricMagnitude="-2" metricUnit="m" metricValue="6.8834" metricValueMax="7.3152" metricValueMin="6.4516" pageId="94" pageNumber="95" unit="in" value="2.71" valueMax="2.88" valueMin="2.54">2.54–2.88 in</quantity>
4♂,
<quantity id="4CC28DBF2A7C8726C633FF1F6DEDFBF3" box="[1062,1205,150,176]" metricMagnitude="-2" metricUnit="m" metricValue="8.0518" metricValueMax="8.1534" metricValueMin="7.9502" pageId="94" pageNumber="95" unit="in" value="3.17" valueMax="3.21" valueMin="3.13">3.13–3.21 in</quantity>
<specimenCount id="9D3CEBD32A7C8726C6A8FF1F6DB8FBF3" box="[1213,1248,150,176]" pageId="94" pageNumber="95" type="female">3♀</specimenCount>
<typeStatus id="54819EF82A7C8726C6FCFF1F6C01FBF3" box="[1257,1369,150,176]" pageId="94" pageNumber="95" type="paratype">paratypes</typeStatus>
), ThL =
<quantity id="4CC28DBF2A7C8726C2BAFF33684CFB96" box="[175,276,186,213]" metricMagnitude="-3" metricUnit="m" metricValue="1.48" pageId="94" pageNumber="95" unit="mm" value="1.48">1.48 mm</quantity>
(
<quantity id="4CC28DBF2A7C8726C331FF3368ECFB97" box="[292,436,186,212]" metricMagnitude="-2" metricUnit="m" metricValue="3.2639" metricValueMax="3.4544" metricValueMin="3.0734" pageId="94" pageNumber="95" unit="in" value="1.285" valueMax="1.36" valueMin="1.21">1.21–1.36 in</quantity>
♂,
<quantity id="4CC28DBF2A7C8726C3F5FF336B36FB97" box="[480,622,186,212]" metricMagnitude="-2" metricUnit="m" metricValue="3.6195" metricValueMax="3.7846" metricValueMin="3.4544" pageId="94" pageNumber="95" unit="in" value="1.425" valueMax="1.49" valueMin="1.36">1.36–1.49 in</quantity>
<specimenCount id="9D3CEBD32A7C8726C062FF336BD2FB97" box="[631,650,186,212]" pageId="94" pageNumber="95" type="female">♀</specimenCount>
), WL =
<quantity id="4CC28DBF2A7C8726C0FBFF336A0CFB96" box="[750,852,186,213]" metricMagnitude="-3" metricUnit="m" metricValue="2.97" pageId="94" pageNumber="95" unit="mm" value="2.97">2.97 mm</quantity>
(
<quantity id="4CC28DBF2A7C8726C176FF336AABFB97" box="[867,1011,186,212]" metricMagnitude="-2" metricUnit="m" metricValue="6.489699999999999" metricValueMax="7.0611999999999995" metricValueMin="5.918200000000001" pageId="94" pageNumber="95" unit="in" value="2.5549999999999997" valueMax="2.78" valueMin="2.33">2.33–2.78 in</quantity>
♂,
<quantity id="4CC28DBF2A7C8726C60AFF336DF6FB97" box="[1055,1198,186,212]" metricMagnitude="-2" metricUnit="m" metricValue="7.048500000000001" metricValueMax="7.391400000000001" metricValueMin="6.7056000000000004" pageId="94" pageNumber="95" unit="in" value="2.775" valueMax="2.91" valueMin="2.64">2.64–2.91 in</quantity>
<specimenCount id="9D3CEBD32A7C8726C6A3FF336D91FB97" box="[1206,1225,186,212]" pageId="94" pageNumber="95" type="female">♀</specimenCount>
), WW =
<quantity id="4CC28DBF2A7C8726C722FF336CC5FB96" box="[1335,1437,186,213]" metricMagnitude="-3" metricUnit="m" metricValue="1.23" pageId="94" pageNumber="95" unit="mm" value="1.23">1.23 mm</quantity>
(
<quantity id="4CC28DBF2A7C8726C28AFF576876FBBB" box="[159,302,222,248]" metricMagnitude="-2" metricUnit="m" metricValue="2.4892" metricValueMax="2.6416" metricValueMin="2.3368" pageId="94" pageNumber="95" unit="in" value="0.98" valueMax="1.04" valueMin="0.92">0.92–1.04 in</quantity>
♂,
<quantity id="4CC28DBF2A7C8726C34FFF5768BFFBBB" box="[346,487,222,248]" metricMagnitude="-2" metricUnit="m" metricValue="2.9464" metricValueMax="3.3274" metricValueMin="2.5654" pageId="94" pageNumber="95" unit="in" value="1.16" valueMax="1.31" valueMin="1.01">1.01–1.31 in</quantity>
<specimenCount id="9D3CEBD32A7C8726C3FAFF576B5AFBBB" box="[495,514,222,248]" pageId="94" pageNumber="95" type="female">♀</specimenCount>
), arb = 7/5 (5–7/4–5), avd = 0.76 (0.73–0.91), adf = 1.44 (0.96–1.30), flw = 2.05 (1.53–2.21), FW/HW = 0.34 (0.33–0.39), ch/o = 0.19 (0.26–0.34), prorb = 1.09 (1.07–1.46), rcorb = 0.78 (0.69– 1.04), vb = 0.87 (0.60–0.82), dcl = 0.43 (0.49–0.82), presctl = 0.49 (0.55–0.83), sctl = 2.33 (1.66–3.04), sterno = 0.81 (0.69–1.45), orbito = 1.43 (1.56–2.00), dcp = 0.31 (0.31–0.37), sctlp = 1.73 (1.14–1.92), C = 1.82 (1.60–1.95), 4c = 1.19 (1.02–1.22), 4v = 1.84 (1.51–1.69), 5x = 1.94 (1.33–1.69), ac = 9.33 (9.91–14.00), M = 0.66 (0.51–0.60), C3F = 0.62 (0.55–0.64).
</paragraph>
</subSubSection>
<subSubSection id="C32073D12A7C8726C2D2FE3F6C45F97F" pageId="94" pageNumber="95" type="materials_examined">
<paragraph id="8B85205A2A7C8726C2D2FE3F6C45F97F" blockId="94.[151,1437,150,645]" pageId="94" pageNumber="95">
<emphasis id="B94EFC482A7C8726C2D2FE3F68DFFA93" bold="true" box="[199,391,438,464]" pageId="94" pageNumber="95">Type materials.</emphasis>
<materialsCitation id="3B522A072A7C8726C380FE3F6A98FAB7" ID-GBIF-Occurrence="4097216381" collectingDate="2017-10-25" collectionCode="SCAU" collectorName="H. W. Chen" country="China" elevation="940" latitude="22.324722" location="Menglian" longLatPrecision="21" longitude="99.583336" municipality="Jinshan" pageId="94" pageNumber="95" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54819EF82A7C8726C380FE3F68A5FA93" box="[405,509,438,464]" pageId="94" pageNumber="95" type="holotype">Holotype</typeStatus>
♂ (
<collectionCode id="ED2BB89F2A7C8726C022FE3F6BDEFA93" box="[567,646,438,464]" pageId="94" pageNumber="95">SCAU</collectionCode>
, No. 125717, DNA #2667),
<collectingCountry id="F32D60CA2A7C8726C1FDFE3F6D1DFA93" box="[1000,1093,438,464]" name="China" pageId="94" pageNumber="95">CHINA</collectingCountry>
:
<collectingMunicipality id="6BE1BA202A7C8726C642FE3F6DF6FA93" box="[1111,1198,438,464]" pageId="94" pageNumber="95">Jinshan</collectingMunicipality>
,
<location id="8EE576812A7C8726C6D5FE3F6C76FA93" LSID="urn:lsid:plazi:treatment:0393914C2A7F8726C282FA316CDEF9C7:8EE576812A7C8726C6D5FE3F6C76FA93" box="[1216,1326,438,464]" country="China" latitude="22.324722" longLatPrecision="21" longitude="99.583336" municipality="Jinshan" name="Menglian" pageId="94" pageNumber="95" stateProvince="Yunnan">Menglian</location>
,
<collectingRegion id="49FEEEB82A7C8726C755FE3E6CC1FA92" box="[1344,1433,439,465]" country="China" name="Yunnan" pageId="94" pageNumber="95">Yunnan</collectingRegion>
,
<geoCoordinate id="EE0E469D2A7C8726C282FE536844FAB7" box="[151,284,474,500]" degrees="22" direction="north" minutes="19" orientation="latitude" pageId="94" pageNumber="95" precision="15" seconds="29" value="22.324722">22°19'29''N</geoCoordinate>
,
<geoCoordinate id="EE0E469D2A7C8726C332FE5368F1FAB6" box="[295,425,474,501]" degrees="99" direction="east" minutes="35" orientation="longitude" pageId="94" pageNumber="95" precision="15" seconds="00" value="99.583336">99°35'00''E</geoCoordinate>
, alt.
<quantity id="4CC28DBF2A7C8726C3C8FE536B78FAB7" box="[477,544,474,500]" metricMagnitude="2" metricUnit="m" metricValue="9.4" pageId="94" pageNumber="95" unit="m" value="940.0">
<elevation id="0017C7692A7C8726C3C8FE536B78FAB7" box="[477,544,474,500]" metricMagnitude="2" metricUnit="m" metricValue="9.4" pageId="94" pageNumber="95" unit="m" value="940.0">940m</elevation>
</quantity>
,
<date id="FF84069A2A7C8726C03FFE536BC5FAB6" box="[554,669,474,501]" pageId="94" pageNumber="95" value="2017-10-25">
<collectingDate id="EFC0FF722A7C8726C03FFE536BC5FAB6" box="[554,669,474,501]" pageId="94" pageNumber="95" value="2017-10-25">25.x.2017</collectingDate>
</date>
, ex tree trunk,
<collectorName id="26CF458C2A7C8726C154FE526A98FAB7" box="[833,960,474,501]" pageId="94" pageNumber="95">H.W. Chen</collectorName>
</materialsCitation>
.
<materialsCitation id="3B522A072A7C8726C1DFFE526B82F95B" ID-GBIF-Occurrence="4097216406" collectingDate="2017-10-25" collectionCode="SCAU" collectorName="L. Gong &amp; B. X. Li" country="China" elevation="940" latitude="22.324722" location="Menglian" longLatPrecision="21" longitude="99.583336" municipality="Jinshan" pageId="94" pageNumber="95" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="54819EF82A7C8726C1DFFE526D63FAB6" box="[970,1083,475,501]" pageId="94" pageNumber="95" type="paratype">Paratypes</typeStatus>
: 3♂,
<specimenCount id="9D3CEBD32A7C8726C660FE536DC0FAB7" box="[1141,1176,474,500]" pageId="94" pageNumber="95" type="female">1♀</specimenCount>
(
<collectionCode id="ED2BB89F2A7C8726C6B3FE536DADFAB7" box="[1190,1269,474,500]" pageId="94" pageNumber="95">SCAU</collectionCode>
, Nos. 125718– 21),
<collectorName id="26CF458C2A7C8726C2DEFE766874F95B" box="[203,300,510,537]" pageId="94" pageNumber="95">L. Gong</collectorName>
,
<collectorName id="26CF458C2A7C8726C32DFE7668C8F95B" box="[312,400,510,537]" pageId="94" pageNumber="95">B.X. Li</collectorName>
, other data same as holotype
</materialsCitation>
;
<materialsCitation id="3B522A072A7C8726C0F0FD896C45F97F" ID-GBIF-Occurrence="4097216314" collectingDate="2018-04-16" collectionCode="SCAU" collectorName="H. W. Chen &amp; L. Gong" country="China" elevation="1100" latitude="22.623056" location="Ximeng" longLatPrecision="21" longitude="99.60305" municipality="Muyiji Park" pageId="94" pageNumber="95" specimenCount="2" specimenCount-female="2" stateProvince="Yunnan" typeStatus="paratype">
1♂,
<specimenCount id="9D3CEBD32A7C8726C102FE776A62F95B" box="[791,826,510,536]" pageId="94" pageNumber="95" type="female">2♀</specimenCount>
(
<collectionCode id="ED2BB89F2A7C8726C15FFE776AC1F95B" box="[842,921,510,536]" pageId="94" pageNumber="95">SCAU</collectionCode>
, Nos. 125722–24, DNA #2668),
<collectingMunicipality id="6BE1BA202A7C8726C71BFE776CC0F95B" box="[1294,1432,510,536]" pageId="94" pageNumber="95">Muyiji Park</collectingMunicipality>
,
<location id="8EE576812A7C8726C282FDAB69ACF97F" LSID="urn:lsid:plazi:treatment:0393914C2A7F8726C282FA316CDEF9C7:8EE576812A7C8726C282FDAB69ACF97F" box="[151,244,546,572]" country="China" latitude="22.623056" longLatPrecision="21" longitude="99.60305" municipality="Muyiji Park" name="Ximeng" pageId="94" pageNumber="95" stateProvince="Yunnan">Ximeng</location>
,
<collectingRegion id="49FEEEB82A7C8726C2EBFDAA680FF97E" box="[254,343,547,573]" country="China" name="Yunnan" pageId="94" pageNumber="95">Yunnan</collectingRegion>
,
<geoCoordinate id="EE0E469D2A7C8726C377FDAB68BFF97F" box="[354,487,546,572]" degrees="22" direction="north" minutes="37" orientation="latitude" pageId="94" pageNumber="95" precision="15" seconds="23" value="22.623056">22°37'23''N</geoCoordinate>
,
<geoCoordinate id="EE0E469D2A7C8726C3E7FDAB6B2BF97F" box="[498,627,546,572]" degrees="99" direction="east" minutes="36" orientation="longitude" pageId="94" pageNumber="95" precision="15" seconds="11" value="99.60305">99°36'11''E</geoCoordinate>
, alt.
<quantity id="4CC28DBF2A7C8726C0BCFDAB6BA0F97F" box="[681,760,546,572]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="94" pageNumber="95" unit="m" value="1100.0">
<elevation id="0017C7692A7C8726C0BCFDAB6BA0F97F" box="[681,760,546,572]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="94" pageNumber="95" unit="m" value="1100.0">1100m</elevation>
</quantity>
,
<date id="FF84069A2A7C8726C111FDAB6A24F97F" box="[772,892,546,572]" pageId="94" pageNumber="95" value="2018-04-16">
<collectingDate id="EFC0FF722A7C8726C111FDAB6A24F97F" box="[772,892,546,572]" pageId="94" pageNumber="95" value="2018-04-16">16.iv.2018</collectingDate>
</date>
, ex tree trunks,
<collectorName id="26CF458C2A7C8726C63BFDAA6DF5F97F" box="[1070,1197,546,573]" pageId="94" pageNumber="95">H.W. Chen</collectorName>
,
<collectorName id="26CF458C2A7C8726C6ADFDAA6C41F97F" box="[1208,1305,546,573]" pageId="94" pageNumber="95">L. Gong</collectorName>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32073D12A7C8726C2D2FDCF6B79F923" box="[199,545,582,608]" pageId="94" pageNumber="95" type="distribution">
<paragraph id="8B85205A2A7C8726C2D2FDCF6B79F923" blockId="94.[151,1437,150,645]" box="[199,545,582,608]" pageId="94" pageNumber="95">
<emphasis id="B94EFC482A7C8726C2D2FDCF6839F923" bold="true" box="[199,353,582,608]" pageId="94" pageNumber="95">Distribution.</emphasis>
<collectingCountry id="F32D60CA2A7C8726C372FDCF68F2F923" box="[359,426,582,608]" name="China" pageId="94" pageNumber="95">China</collectingCountry>
(
<collectingRegion id="49FEEEB82A7C8726C3ACFDCF6B4DF923" box="[441,533,582,608]" country="China" name="Yunnan" pageId="94" pageNumber="95">Yunnan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C32073D12A7C8726C2D2FDE36CDEF9C7" box="[199,1414,618,645]" pageId="94" pageNumber="95" type="etymology">
<paragraph id="8B85205A2A7C8726C2D2FDE36CDEF9C7" blockId="94.[151,1437,150,645]" box="[199,1414,618,645]" pageId="94" pageNumber="95">
<emphasis id="B94EFC482A7C8726C2D2FDE36816F9C7" bold="true" box="[199,334,618,644]" pageId="94" pageNumber="95">Etymology.</emphasis>
From the Greek word &quot;
<emphasis id="B94EFC482A7C8726C04CFDE36B81F9C7" box="[601,729,618,644]" italics="true" pageId="94" pageNumber="95">sciophilous</emphasis>
&quot; (= sciophilous), referring to the sciophilous
<taxonomicName id="4C3A5BD92A7C8726C6C2FDE26C1FF9C6" authorityName="Wheeler" authorityYear="1960" box="[1239,1351,619,645]" class="Insecta" family="Drosophilidae" genus="Stegana" kingdom="Animalia" order="Diptera" pageId="94" pageNumber="109" phylum="Arthropoda" rank="subGenus" subGenus="Steganina">
<emphasis id="B94EFC482A7C8726C6C2FDE26C1FF9C6" box="[1239,1351,619,645]" italics="true" pageId="94" pageNumber="95">Steganina</emphasis>
</taxonomicName>
flies.
</paragraph>
</subSubSection>
</treatment>
</document>
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