treatments-xml/data/03/93/80/03938025FFA0A926A7D8B46DFA9AFBED.xml
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<document id="5CCFC766565244FB9CD23D2EB6317E9A" ID-CLB-Dataset="8556" ID-DOI="10.11646/zootaxa.4843.1.1" ID-GBIF-Dataset="a2aef684-44f1-429d-9dd5-dd394539fa36" ID-ISSN="1175-5326" ID-Zenodo-Dep="4405729" ID-ZooBank="57044928-1950-4525-850E-E381A52599A1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1609419131752" checkinUser="plazi" docAuthor="Salata, Sebastian &amp; Fisher, Brian L." docDate="2020" docId="03938025FFA0A926A7D8B46DFA9AFBED" docLanguage="en" docName="zootaxa.4843.1.1.pdf" docOrigin="Zootaxa 4843 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Pheidole ovalinoda Salata &amp; Fisher 2020, sp. nov." docType="treatment" docVersion="8" lastPageNumber="49" masterDocId="FFAAF85DFF8EA916A74FB608FFC6FFFD" masterDocTitle="The ant genus Pheidole Westwood, 1839 (Hymenoptera: Formicidae) in Madagascar-taxonomic revision of the bessonii species-group" masterLastPageNumber="64" masterPageNumber="1" pageNumber="47" updateTime="1698850390174" updateUser="ExternalLinkService">
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<mods:title id="C4B7F9A336FE2BCB1BF7C6F3D6B71633">The ant genus Pheidole Westwood, 1839 (Hymenoptera: Formicidae) in Madagascar-taxonomic revision of the bessonii species-group</mods:title>
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<mods:name id="DE836D416D48A087C67DDD96B25C642C" type="personal">
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<mods:namePart id="394E2EBBCB88B1583CA2CAE1D16638B9">Salata, Sebastian</mods:namePart>
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<mods:namePart id="C24D81A42102DAC1B7842107D31A4395">Fisher, Brian L.</mods:namePart>
<mods:nameIdentifier id="2128E45953B2DE9F02131D530C38391F" type="ORCID">0000-0002-4653-3270</mods:nameIdentifier>
<mods:nameIdentifier id="076BDBBA7D94EEA1583FCCA61F81C882" type="email">bfisher@calacademy.org</mods:nameIdentifier>
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<mods:title id="3E7032EA0412E9687D95651A0AAAE50C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="4A4D23A31BA718FE0624057D1D2A1BD6">
<mods:date id="8A5D4271310D041CE5DC10E75946EB68">2020</mods:date>
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<mods:number id="EEBE7372580195C60C63D79E0929D59F">2020-08-31</mods:number>
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<mods:number id="A46CEB036732C37C0A7DC4F086FAA992">4843</mods:number>
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<mods:classification id="7CAE5EE96FFF6D07D939930C32DA1A62">journal article</mods:classification>
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<mods:identifier id="08CDE7C5CEBA075D8AB9F5A4BA8ED788" type="DOI">10.11646/zootaxa.4843.1.1</mods:identifier>
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<treatment id="03938025FFA0A926A7D8B46DFA9AFBED" ID-DOI="http://doi.org/10.5281/zenodo.4488399" ID-GBIF-Taxon="177202121" ID-Zenodo-Dep="4488399" LSID="urn:lsid:plazi:treatment:03938025FFA0A926A7D8B46DFA9AFBED" httpUri="http://treatment.plazi.org/id/03938025FFA0A926A7D8B46DFA9AFBED" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47">
<subSubSection id="C32062B8FFA0A938A7D8B46DFE22FD82" box="[151,484,613,640]" pageId="46" pageNumber="47" type="nomenclature">
<paragraph id="8B853133FFA0A938A7D8B46DFE22FD82" blockId="46.[151,484,613,675]" box="[151,484,613,640]" pageId="46" pageNumber="47">
<heading id="D0CD865FFFA0A938A7D8B46DFE22FD82" bold="true" box="[151,484,613,640]" fontSize="11" level="2" pageId="46" pageNumber="47" reason="4">
<emphasis id="B94EED21FFA0A938A7D8B46DFE22FD82" bold="true" box="[151,484,613,640]" pageId="46" pageNumber="47">
<taxonomicName id="4C3A4AB0FFA0A938A7D8B46DFEB9FD7D" authority="Salata &amp; Fisher, 2020" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[151,383,613,640]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="ovalinoda" status="sp. nov.">
<emphasis id="B94EED21FFA0A938A7D8B46DFEB9FD7D" bold="true" box="[151,383,613,640]" italics="true" pageId="46" pageNumber="47">Pheidole ovalinoda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D505AFFA0A938A6C8B46DFE22FD82" box="[391,484,613,639]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FFA0A938A7D8B481FE59FD5E" box="[151,415,649,675]" pageId="46" pageNumber="47" type="description">
<paragraph id="8B853133FFA0A938A7D8B481FE59FD5E" blockId="46.[151,484,613,675]" box="[151,415,649,675]" pageId="46" pageNumber="47">
<figureCitation id="13012DB6FFA0A938A7D8B481FF2EFD5E" box="[151,232,649,675]" captionStart="FIGURE21" captionStartId="47.[151,250,1931,1955]" captionTargetBox="[162,1426,417,1905]" captionTargetId="figure-120@47.[162,1426,417,1908]" captionTargetPageId="47" captionText="FIGURE21.Pheidoleovalinoda,full-faceview(A),profile(C),anddorsalview(E)ofparatypeminorworker(CASENT0923277) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0808080)." figureDoi="http://doi.org/10.5281/zenodo.4405787" httpUri="https://zenodo.org/record/4405787/files/figure.png" pageId="46" pageNumber="47">Figs 21</figureCitation>
AF, 27M, 28M
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FFA0A938A7D8B4C4FBC5FC43" pageId="46" pageNumber="47" type="materials_examined">
<paragraph id="8B853133FFA0A938A7D8B4C4FF3CFCAF" blockId="46.[151,1437,716,2039]" pageId="46" pageNumber="47">
<materialsCitation id="3B523B6EFFA0A938A7D8B4C4FD6BFCD3" ID-GBIF-Occurrence="3027954782" accessionNumber="BLF40902" collectingDate="2018-02-09" collectionCode="CASC" collectorName="NE Andapa &amp; B. L. Fisher" country="Madagascar" county="Sava Region" elevation="850" latitude="-14.43462" location="Manantenina River" longLatPrecision="1" longitude="49.75853" municipality="Parc National de Marojejy" pageId="46" pageNumber="47" specimenCount="1" stateProvince="Antsiranana" typeStatus="holotype">
<typeStatus id="54818F91FFA0A938A7D8B4C4FEE9FD1B" box="[151,303,716,742]" pageId="46" pageNumber="47" type="holotype">HOLOTYPE</typeStatus>
: 1s.,
<collectingCountry id="F32D71A3FFA0A938A621B4C4FE31FD1B" box="[366,503,716,742]" name="Madagascar" pageId="46" pageNumber="47">Madagascar</collectingCountry>
,
<collectingRegion id="49FEFFD1FFA0A938A54BB4C4FD48FD1B" box="[516,654,716,742]" country="Madagascar" name="Antsiranana" pageId="46" pageNumber="47">Antsiranana</collectingRegion>
,
<collectingCounty id="62E449BFFFA0A938A5D3B4C4FCF4FD1B" box="[668,818,716,742]" pageId="46" pageNumber="47">Sava Region</collectingCounty>
:
<collectingMunicipality id="6BE1AB49FFA0A938A40FB4C5FBB5FD1B" box="[832,1139,716,743]" pageId="46" pageNumber="47">Parc National de Marojejy</collectingMunicipality>
,
<location id="8EE567E8FFA0A938A3CEB4C4FA9BFD1B" LSID="urn:lsid:plazi:treatment:03938025FFA0A926A7D8B46DFA9AFBED:8EE567E8FFA0A938A3CEB4C4FA9BFD1B" box="[1153,1373,716,742]" country="Madagascar" county="Sava Region" latitude="-14.43462" longLatPrecision="1" longitude="49.75853" municipality="Parc National de Marojejy" name="Manantenina River" pageId="46" pageNumber="47" stateProvince="Antsiranana">Manantenina River</location>
,
<quantity id="4CC29CD6FFA0A938A224B4C4FF7DFCF7" metricMagnitude="4" metricUnit="m" metricValue="2.79" pageId="46" pageNumber="47" unit="km" value="27.9">27.9 km</quantity>
24.3°
<collectorName id="26CF54E5FFA0A938A649B4F9FE4AFCF7" box="[262,396,752,779]" pageId="46" pageNumber="47">NE Andapa</collectorName>
,
<geoCoordinate id="EE0E57F4FFA0A938A6D8B4F8FDCCFCF7" box="[407,522,752,778]" degrees="14.43462" direction="south" orientation="latitude" pageId="46" pageNumber="47" precision="1" value="-14.43462">-14.43462</geoCoordinate>
<geoCoordinate id="EE0E57F4FFA0A938A55EB4F8FDBBFCF6" box="[529,637,752,779]" degrees="49.75853" direction="east" orientation="longitude" pageId="46" pageNumber="47" precision="1" value="49.75853">49.75853</geoCoordinate>
,
<quantity id="4CC29CD6FFA0A938A5C6B4F8FD14FCF6" box="[649,722,752,779]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="46" pageNumber="47" unit="m" value="850.0">
<elevation id="0017D600FFA0A938A5C6B4F8FD14FCF6" box="[649,722,752,779]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="46" pageNumber="47" unit="m" value="850.0">850 m</elevation>
</quantity>
,
<date id="FF8417F3FFA0A938A590B4F8FCA2FCF7" box="[735,868,752,778]" pageId="46" pageNumber="47" value="2018-02-09">
<collectingDate id="EFC0EE1BFFA0A938A590B4F8FCA2FCF7" box="[735,868,752,778]" pageId="46" pageNumber="47" value="2018-02-09">9-Feb-2018</collectingDate>
</date>
, rainforest, ex tunelle on clay bark,
<collectorName id="26CF54E5FFA0A938A3B5B4F9FA46FCF7" box="[1274,1408,752,779]" pageId="46" pageNumber="47">B. L. Fisher</collectorName>
<emphasis id="B94EED21FFA0A938A2C7B4F9FF6BFCD3" italics="true" pageId="46" pageNumber="47">et al</emphasis>
. leg. BLF40902, CASENT0808080 (
<collectionCode id="ED2BA9F6FFA0A938A51AB51CFD62FCD3" box="[597,676,788,814]" pageId="46" pageNumber="47">CASC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B523B6EFFA0A938A5F7B51CFF30FCAF" ID-GBIF-Occurrence="3027954565" accessionNumber="BLF40902" collectingDate="2018-02-09" collectionCode="CASC" collectorName="NE Andapa &amp; B. L. Fisher" country="Madagascar" county="Sava Region" elevation="850" latitude="-14.43462" location="Manantenina River" longLatPrecision="1" longitude="49.75853" municipality="Parc National de Marojejy" pageId="46" pageNumber="47" specimenCount="1" stateProvince="Antsiranana" typeStatus="paratype">
<typeStatus id="54818F91FFA0A938A5F7B51CFC91FCD3" box="[696,855,788,814]" pageId="46" pageNumber="47" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="9D3CFABAFFA0A938A42CB51CFC4AFCD3" box="[867,908,788,814]" pageId="46" pageNumber="47" type="generic">1w.</specimenCount>
, the same data as holotype, CASENT0923277 (
<collectionCode id="ED2BA9F6FFA0A938A7D0B530FF28FCAF" box="[159,238,824,850]" pageId="46" pageNumber="47">CASC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B853133FFA0A938A788B554FB55FC67" blockId="46.[151,1437,716,2039]" pageId="46" pageNumber="47">
<emphasis id="B94EED21FFA0A938A788B554FE42FC8B" bold="true" box="[199,388,860,886]" pageId="46" pageNumber="47">Other material.</emphasis>
<materialsCitation id="3B523B6EFFA0A938A6C1B554FB49FC67" ID-GBIF-Occurrence="3027954505" collectionCode="CASC" collectorName="NE Andapa &amp; B. L. Fisher" country="Madagascar" county="Sava Region" elevation="850" latitude="-14.43462" location="Manantenina River" longLatPrecision="1" longitude="49.75853" municipality="Parc National de Marojejy" pageId="46" pageNumber="47" specimenCount="3" stateProvince="Antsiranana">
<collectingCountry id="F32D71A3FFA0A938A6C1B554FDD1FC8B" box="[398,535,860,886]" name="Madagascar" pageId="46" pageNumber="47">Madagascar</collectingCountry>
.
<collectingRegion id="49FEFFD1FFA0A938A56BB554FD76FC8B" box="[548,688,860,886]" country="Madagascar" name="Antsiranana" pageId="46" pageNumber="47">Antsiranana</collectingRegion>
:
<specimenCount id="9D3CFABAFFA0A938A5F2B554FD20FC8B" box="[701,742,860,886]" pageId="46" pageNumber="47" type="generic">3w.</specimenCount>
, 2s.,
<collectingCounty id="62E449BFFFA0A938A469B554FC7AFC8B" box="[806,956,860,886]" pageId="46" pageNumber="47">Sava Region</collectingCounty>
:
<collectingMunicipality id="6BE1AB49FFA0A938A486B555FB3AFC8B" box="[969,1276,860,887]" pageId="46" pageNumber="47">Parc National de Marojejy</collectingMunicipality>
,
<location id="8EE567E8FFA0A938A245B554FF11FC67" LSID="urn:lsid:plazi:treatment:03938025FFA0A926A7D8B46DFA9AFBED:8EE567E8FFA0A938A245B554FF11FC67" country="Madagascar" county="Sava Region" latitude="-14.43462" longLatPrecision="1" longitude="49.75853" municipality="Parc National de Marojejy" name="Manantenina River" pageId="46" pageNumber="47" stateProvince="Antsiranana">Manantenina River</location>
,
<quantity id="4CC29CD6FFA0A938A7ADB588FEF8FC67" box="[226,318,896,922]" metricMagnitude="4" metricUnit="m" metricValue="2.79" pageId="46" pageNumber="47" unit="km" value="27.9">27.9 km</quantity>
24.3°
<collectorName id="26CF54E5FFA0A938A6C7B589FDCBFC67" box="[392,525,896,923]" pageId="46" pageNumber="47">NE Andapa</collectorName>
,
<geoCoordinate id="EE0E57F4FFA0A938A557B588FD4CFC67" box="[536,650,896,922]" degrees="14.43462" direction="south" orientation="latitude" pageId="46" pageNumber="47" precision="1" value="-14.43462">-14.43462</geoCoordinate>
<geoCoordinate id="EE0E57F4FFA0A938A5DEB588FD3BFC66" box="[657,765,896,923]" degrees="49.75853" direction="east" orientation="longitude" pageId="46" pageNumber="47" precision="1" value="49.75853">49.75853</geoCoordinate>
,
<quantity id="4CC29CD6FFA0A938A447B588FC97FC66" box="[776,849,896,923]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="46" pageNumber="47" unit="m" value="850.0">
<elevation id="0017D600FFA0A938A447B588FC97FC66" box="[776,849,896,923]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="46" pageNumber="47" unit="m" value="850.0">850 m</elevation>
</quantity>
,
<collectorName id="26CF54E5FFA0A938A412B589FC25FC67" box="[861,995,896,923]" pageId="46" pageNumber="47">B. L. Fisher</collectorName>
<emphasis id="B94EED21FFA0A938A4A5B589FBDDFC67" box="[1002,1051,896,922]" italics="true" pageId="46" pageNumber="47">et al</emphasis>
., (
<collectionCode id="ED2BA9F6FFA0A938A377B588FB41FC67" box="[1080,1159,896,922]" pageId="46" pageNumber="47">CASC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B853133FFA0A938A788B5ACFBC5FC43" blockId="46.[151,1437,716,2039]" box="[199,1027,932,959]" pageId="46" pageNumber="47">
<emphasis id="B94EED21FFA0A938A788B5ACFE60FC43" bold="true" box="[199,422,932,958]" pageId="46" pageNumber="47">Geographic range.</emphasis>
<collectingCountry id="F32D71A3FFA0A938A6E2B5ACFDF0FC43" box="[429,566,932,958]" name="Madagascar" pageId="46" pageNumber="47">Madagascar</collectingCountry>
,
<collectingRegion id="49FEFFD1FFA0A938A570B5ACFD0FFC43" box="[575,713,932,958]" country="Madagascar" name="Antsiranana" pageId="46" pageNumber="47">Antsiranana</collectingRegion>
, Parc National de Marojejy.
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FFA0A938A788B5C0FAF5FA4B" pageId="46" pageNumber="47" type="diagnosis">
<paragraph id="8B853133FFA0A938A788B5C0FAF5FA4B" blockId="46.[151,1437,716,2039]" pageId="46" pageNumber="47">
<emphasis id="B94EED21FFA0A938A788B5C0FDC5FC1F" bold="true" box="[199,515,968,994]" pageId="46" pageNumber="47">Diagnosis. Major workers.</emphasis>
Head in full-face view sub-rectangular, not widening posteriorly, with lateral margins relatively straight; side of head with dense, short, decumbent to suberect pilosity; anteromedial frons with moderately dense and thick costulae; interspaces between costulae indistinctly rugulate; posteromedial frons rugulate; interspaces between rugulae more distinctly rugulate; frons laterally with thick and moderately dense rugae; interspaces between rugae indistinctly rugulate; occipital lobes with thick and moderately dense rugae; interspaces between rugae mostly smooth; scape, when laid back, exceeding the midlength of head by one-fifth of its length; inner hypostomal tooth distinct and bulge-like; outer hypostomal tooth absent to very indistinct; median tooth absent; promesonotum short, angular, and moderately low; posterior mesonotum moderately steep; mesonotal process distinct; mesosoma microreticulate, lateral sides of pronotum, propodeum, and katepisternum with additional thin rugae; pronotal dorsum with additional rugae; gaster shagreened; body brown.
<emphasis id="B94EED21FFA0A938A35CB304FB09FADB" bold="true" box="[1043,1231,1292,1318]" pageId="46" pageNumber="47">Minor workers.</emphasis>
Occiput not elongated; nuchal collar indistinct; head sculpture smooth; scape, when laid back, exceeding the posterior head margin by two-fifths of its length; promesonotum moderately low, moderately long, and arched; promesonotal groove present; metanotal groove distinct; propodeal spines very small and thin; promesonotum smooth; anepisternum, katepisternum, and propodeum punctate; body yellowish brown; petiolar node high, triangular, and thick.
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FFA0A926A788B3C8FE69FD9C" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47" type="description">
<paragraph id="8B853133FFA0A938A788B3C8FBADF9BB" blockId="46.[151,1437,716,2039]" pageId="46" pageNumber="47">
<emphasis id="B94EED21FFA0A938A788B3C8FDDDFA27" bold="true" box="[199,539,1472,1498]" pageId="46" pageNumber="47">Description. Major workers.</emphasis>
Measurements (n=4): HL: 1.351.5 (1.41); HW: 1.31.43 (1.36); SL: 0.920.98 (0.95); EL: 0.180.19 (0.18); WL: 1.251.35 (1.29); PSL: 0.20.23 (0.22); MTL: 0.951.02 (0.99); PNW: 0.53 0.57 (0.56); PTW: 0.170.19 (0.18); PPW: 0.330.42 (0.38); CI: 101.9105.3 (103.9); SI: 68.571.1 (70.2); PSLI: 14.816.6 (15.7); PPI: 43.350.5 (46.3); PNI: 39.941.7 (41.0); MTI: 71.474.3 (73.1).
</paragraph>
<paragraph id="8B853133FFA0A939A788B058FC55FE74" blockId="46.[151,1437,716,2039]" lastBlockId="47.[151,1437,151,393]" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
<emphasis id="B94EED21FFA0A938A788B058FEC8F997" bold="true" box="[199,270,1616,1642]" pageId="46" pageNumber="47">Head.</emphasis>
In full-face view sub-rectangular, not widening posteriorly, with lateral sides relatively straight (
<figureCitation id="13012DB6FFA0A938A220B058FF0FF973" captionStart="FIGURE21" captionStartId="47.[151,250,1931,1955]" captionTargetBox="[162,1426,417,1905]" captionTargetId="figure-120@47.[162,1426,417,1908]" captionTargetPageId="47" captionText="FIGURE21.Pheidoleovalinoda,full-faceview(A),profile(C),anddorsalview(E)ofparatypeminorworker(CASENT0923277) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0808080)." figureDoi="http://doi.org/10.5281/zenodo.4405787" httpUri="https://zenodo.org/record/4405787/files/figure.png" pageId="46" pageNumber="47">Fig. 21B</figureCitation>
). In lateral view sub-oval, not depressed posteriorly; ventral and dorsal margins convex; inner hypostomal tooth visible. Side of head with dense, short, decumbent to suberect pilosity; head dorsum with relatively dense, long, suberect to erect pilosity. Anteromedial frons with moderately dense and thick costulae; interspaces between costulae indistinctly rugulate; posteromedial frons rugulate; interspaces between rugulae more distinctly rugulate. Frons laterally with thick and moderately dense rugae; interspaces between rugae indistinctly rugulate. Occipital lobes with thick and moderately dense rugae; interspaces between rugae mostly smooth. Gena with dense and moderately thick costulae; interspaces between costulae mostly punctate. Sides posterolateral from eyes with thick rugulae; interspaces between rugulae distinctly rugulate. Center of clypeus shiny with weakly developed puncta, lateral sides with distinct rugulae; median notch present, moderately wide and shallow; median longitudinal carina present; lateral longitudinal carinae present. Scape, when laid back, exceeding the midlength of head by one-fifth of its length; pilosity subdecumbent to erect (
<figureCitation id="13012DB6FFA0A938A51EB1D4FD1CF80A" box="[593,730,2012,2039]" captionStart="FIGURE21" captionStartId="47.[151,250,1931,1955]" captionTargetBox="[162,1426,417,1905]" captionTargetId="figure-120@47.[162,1426,417,1908]" captionTargetPageId="47" captionText="FIGURE21.Pheidoleovalinoda,full-faceview(A),profile(C),anddorsalview(E)ofparatypeminorworker(CASENT0923277) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0808080)." figureDoi="http://doi.org/10.5281/zenodo.4405787" httpUri="https://zenodo.org/record/4405787/files/figure.png" pageId="46" pageNumber="47">Fig. 21B, D</figureCitation>
). Inner hypostomal tooth distinct, bulge-like; outer hypostomal tooth absent to very indistinct; median tooth absent (
<figureCitation id="13012DB6FFA1A939A5A3B69FFC9CFF4C" box="[748,858,151,177]" captionStart="FIGURE21" captionStartId="47.[151,250,1931,1955]" captionTargetBox="[162,1426,417,1905]" captionTargetId="figure-120@47.[162,1426,417,1908]" captionTargetPageId="47" captionText="FIGURE21.Pheidoleovalinoda,full-faceview(A),profile(C),anddorsalview(E)ofparatypeminorworker(CASENT0923277) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0808080)." figureDoi="http://doi.org/10.5281/zenodo.4405787" httpUri="https://zenodo.org/record/4405787/files/figure.png" pageId="47" pageNumber="48">Fig. 21M</figureCitation>
).
<emphasis id="B94EED21FFA1A939A420B69FFC32FF4C" bold="true" box="[879,1012,151,177]" pageId="47" pageNumber="48">Mesosoma.</emphasis>
In lateral view, promesonotum short, angular, and moderately low; posterior mesonotum moderately steep; mesonotal process distinct and tubercle-like; promesonotal groove absent; metanotal groove indistinct; propodeal spines moderately long with moderately wide base and acute top; humeral tubercle weakly produced (
<figureCitation id="13012DB6FFA1A939A440B70BFCBFFEE0" box="[783,889,259,285]" captionStart="FIGURE21" captionStartId="47.[151,250,1931,1955]" captionTargetBox="[162,1426,417,1905]" captionTargetId="figure-120@47.[162,1426,417,1908]" captionTargetPageId="47" captionText="FIGURE21.Pheidoleovalinoda,full-faceview(A),profile(C),anddorsalview(E)ofparatypeminorworker(CASENT0923277) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0808080)." figureDoi="http://doi.org/10.5281/zenodo.4405787" httpUri="https://zenodo.org/record/4405787/files/figure.png" pageId="47" pageNumber="48">Fig. 21D</figureCitation>
). Surface microreticulate; lateral sides of pronotum, propodeum, and katepisternum with additional thin rugae; pronotal dorsum with additional rugae. Pilosity moderately dense, long, and erect (
<figureCitation id="13012DB6FFA1A939A569B743FD69FE98" box="[550,687,331,357]" captionStart="FIGURE21" captionStartId="47.[151,250,1931,1955]" captionTargetBox="[162,1426,417,1905]" captionTargetId="figure-120@47.[162,1426,417,1908]" captionTargetPageId="47" captionText="FIGURE21.Pheidoleovalinoda,full-faceview(A),profile(C),anddorsalview(E)ofparatypeminorworker(CASENT0923277) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0808080)." figureDoi="http://doi.org/10.5281/zenodo.4405787" httpUri="https://zenodo.org/record/4405787/files/figure.png" pageId="47" pageNumber="48">Fig. 21D, F</figureCitation>
).
<emphasis id="B94EED21FFA1A939A58BB742FCDCFE99" bold="true" box="[708,794,330,356]" pageId="47" pageNumber="48">Gaster.</emphasis>
Shiny and shagreened; pilosity moderately sparse, long, and erect (
<figureCitation id="13012DB6FFA1A939A643B767FE54FE74" box="[268,402,367,393]" captionStart="FIGURE21" captionStartId="47.[151,250,1931,1955]" captionTargetBox="[162,1426,417,1905]" captionTargetId="figure-120@47.[162,1426,417,1908]" captionTargetPageId="47" captionText="FIGURE21.Pheidoleovalinoda,full-faceview(A),profile(C),anddorsalview(E)ofparatypeminorworker(CASENT0923277) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0808080)." figureDoi="http://doi.org/10.5281/zenodo.4405787" httpUri="https://zenodo.org/record/4405787/files/figure.png" pageId="47" pageNumber="48">Fig. 21D, F</figureCitation>
).
<emphasis id="B94EED21FFA1A939A6E9B766FE29FE75" bold="true" box="[422,495,366,392]" pageId="47" pageNumber="48">Color.</emphasis>
Brown, legs yellowish (
<figureCitation id="13012DB6FFA1A939A44FB767FC40FE74" box="[768,902,367,393]" captionStart="FIGURE21" captionStartId="47.[151,250,1931,1955]" captionTargetBox="[162,1426,417,1905]" captionTargetId="figure-120@47.[162,1426,417,1908]" captionTargetPageId="47" captionText="FIGURE21.Pheidoleovalinoda,full-faceview(A),profile(C),anddorsalview(E)ofparatypeminorworker(CASENT0923277) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0808080)." figureDoi="http://doi.org/10.5281/zenodo.4405787" httpUri="https://zenodo.org/record/4405787/files/figure.png" pageId="47" pageNumber="48">Fig. 21D, F</figureCitation>
).
</paragraph>
<caption id="DF4561BBFFA1A939A7D8B183FB61F835" ID-DOI="http://doi.org/10.5281/zenodo.4405787" ID-Zenodo-Dep="4405787" httpUri="https://zenodo.org/record/4405787/files/figure.png" pageId="47" pageNumber="48" startId="47.[151,250,1931,1955]" targetBox="[162,1426,417,1905]" targetPageId="47">
<paragraph id="8B853133FFA1A939A7D8B183FB61F835" blockId="47.[151,1436,1931,1992]" pageId="47" pageNumber="48">
<emphasis id="B94EED21FFA1A939A7D8B183FEDAF859" bold="true" box="[151,284,1931,1956]" pageId="47" pageNumber="48">FIGURE 21.</emphasis>
<taxonomicName id="4C3A4AB0FFA1A939A650B184FE27F859" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[287,481,1932,1956]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="ovalinoda">
<emphasis id="B94EED21FFA1A939A650B184FE27F859" box="[287,481,1932,1956]" italics="true" pageId="47" pageNumber="48">Pheidole ovalinoda</emphasis>
</taxonomicName>
, full-face view (A), profile (C), and dorsal view (E)of paratype minor worker(CASENT0923277) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0808080).
</paragraph>
</caption>
<paragraph id="8B853133FFBEA926A788B69EFB53FEE1" blockId="48.[151,1437,150,1041]" pageId="48" pageNumber="49">
<emphasis id="B94EED21FFBEA926A788B69EFDDDFF4D" bold="true" box="[199,539,150,176]" pageId="48" pageNumber="49">Description. Minor workers.</emphasis>
Measurements (n=4): HL: 0.60.64 (0.62); HW: 0.460.49 (0.47); SL: 0.770.8 (0.79); EL: 0.110.13 (0.13); WL: 0.810.87 (0.84); PSL: 0.090.1 (0.09); MTL: 0.640.67 (0.65); PNW: 0.30.35 (0.33); PTW: 0.080.11 (0.09); PPW: 0.120.14 (0.13); CI: 127.6134.7 (130.5); SI: 162.9170.7 (166.1); PSLI: 14.115.7 (14.8); PPI: 59.479.9 (69.6); PNI: 64.473.8 (70.0); MTI: 134.5138.6 (137.3).
</paragraph>
<paragraph id="8B853133FFBEA926A788B72EFE69FD9C" blockId="48.[151,1437,150,1041]" pageId="48" pageNumber="49">
<emphasis id="B94EED21FFBEA926A788B72EFEC8FEBD" bold="true" box="[199,270,294,320]" pageId="48" pageNumber="49">Head.</emphasis>
In full-face view oval, posterior region not elongated; nuchal collar weakly developed (
<figureCitation id="13012DB6FFBEA926A3AFB72EFA8EFEBD" box="[1248,1352,294,320]" captionStart="FIGURE21" captionStartId="47.[151,250,1931,1955]" captionTargetBox="[162,1426,417,1905]" captionTargetId="figure-120@47.[162,1426,417,1908]" captionTargetPageId="47" captionText="FIGURE21.Pheidoleovalinoda,full-faceview(A),profile(C),anddorsalview(E)ofparatypeminorworker(CASENT0923277) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0808080)." figureDoi="http://doi.org/10.5281/zenodo.4405787" httpUri="https://zenodo.org/record/4405787/files/figure.png" pageId="48" pageNumber="49">Fig. 21A</figureCitation>
). Pilosity relatively sparse, moderately short, subdecumbent to erect. Sculpture mostly smooth; anterolateral frons punctate. Clypeus with median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, exceeding the posterior head margin by two-fifths of its length; pilosity dense, subdecumbent to erect (
<figureCitation id="13012DB6FFBEA926A260B79AFF6BFE2D" captionStart="FIGURE21" captionStartId="47.[151,250,1931,1955]" captionTargetBox="[162,1426,417,1905]" captionTargetId="figure-120@47.[162,1426,417,1908]" captionTargetPageId="47" captionText="FIGURE21.Pheidoleovalinoda,full-faceview(A),profile(C),anddorsalview(E)ofparatypeminorworker(CASENT0923277) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0808080)." figureDoi="http://doi.org/10.5281/zenodo.4405787" httpUri="https://zenodo.org/record/4405787/files/figure.png" pageId="48" pageNumber="49">Fig. 21A, C</figureCitation>
).
<emphasis id="B94EED21FFBEA926A78EB7BEFE80FE2D" bold="true" box="[193,326,438,464]" pageId="48" pageNumber="49">Mesosoma.</emphasis>
In lateral view, promesonotum moderately low, moderately long, and arched; promesonotal groove present; metanotal groove distinct; propodeal spines very small, thin, and triangular (
<figureCitation id="13012DB6FFBEA926A33DB7D2FB1AFE09" box="[1138,1244,474,500]" captionStart="FIGURE21" captionStartId="47.[151,250,1931,1955]" captionTargetBox="[162,1426,417,1905]" captionTargetId="figure-120@47.[162,1426,417,1908]" captionTargetPageId="47" captionText="FIGURE21.Pheidoleovalinoda,full-faceview(A),profile(C),anddorsalview(E)ofparatypeminorworker(CASENT0923277) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0808080)." figureDoi="http://doi.org/10.5281/zenodo.4405787" httpUri="https://zenodo.org/record/4405787/files/figure.png" pageId="48" pageNumber="49">Fig. 21C</figureCitation>
). Promesonotum smooth; anepisternum, katepisternum, and propodeum punctate. Pilosity sparse, moderately long, and erect (
<figureCitation id="13012DB6FFBEA926A220B7F6FF2FFDC0" captionStart="FIGURE21" captionStartId="47.[151,250,1931,1955]" captionTargetBox="[162,1426,417,1905]" captionTargetId="figure-120@47.[162,1426,417,1908]" captionTargetPageId="47" captionText="FIGURE21.Pheidoleovalinoda,full-faceview(A),profile(C),anddorsalview(E)ofparatypeminorworker(CASENT0923277) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0808080)." figureDoi="http://doi.org/10.5281/zenodo.4405787" httpUri="https://zenodo.org/record/4405787/files/figure.png" pageId="48" pageNumber="49">Fig. 21C, E</figureCitation>
).
<emphasis id="B94EED21FFBEA926A7B3B42AFE94FDC1" bold="true" box="[252,338,546,572]" pageId="48" pageNumber="49">Gaster.</emphasis>
With sparse and erect pilosity (
<figureCitation id="13012DB6FFBEA926A5FCB42AFCFFFDC0" box="[691,825,546,573]" captionStart="FIGURE21" captionStartId="47.[151,250,1931,1955]" captionTargetBox="[162,1426,417,1905]" captionTargetId="figure-120@47.[162,1426,417,1908]" captionTargetPageId="47" captionText="FIGURE21.Pheidoleovalinoda,full-faceview(A),profile(C),anddorsalview(E)ofparatypeminorworker(CASENT0923277) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0808080)." figureDoi="http://doi.org/10.5281/zenodo.4405787" httpUri="https://zenodo.org/record/4405787/files/figure.png" pageId="48" pageNumber="49">Fig. 21C, E</figureCitation>
).
<emphasis id="B94EED21FFBEA926A403B42AFC53FDC1" bold="true" box="[844,917,546,572]" pageId="48" pageNumber="49">Color.</emphasis>
Yellowish brown, gaster and head darker than mesosoma (
<figureCitation id="13012DB6FFBEA926A652B44EFE65FD9C" box="[285,419,582,609]" captionStart="FIGURE21" captionStartId="47.[151,250,1931,1955]" captionTargetBox="[162,1426,417,1905]" captionTargetId="figure-120@47.[162,1426,417,1908]" captionTargetPageId="47" captionText="FIGURE21.Pheidoleovalinoda,full-faceview(A),profile(C),anddorsalview(E)ofparatypeminorworker(CASENT0923277) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0808080)." figureDoi="http://doi.org/10.5281/zenodo.4405787" httpUri="https://zenodo.org/record/4405787/files/figure.png" pageId="48" pageNumber="49">Fig. 21C, E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FFBEA926A788B462FF26FD55" pageId="48" pageNumber="49" type="biology_ecology">
<paragraph id="8B853133FFBEA926A788B462FF26FD55" blockId="48.[151,1437,150,1041]" pageId="48" pageNumber="49">
<emphasis id="B94EED21FFBEA926A788B462FEE1FD79" bold="true" box="[199,295,618,644]" pageId="48" pageNumber="49">Biology.</emphasis>
The species was collected at
<quantity id="4CC29CD6FFBEA926A53DB462FD7FFD78" box="[626,697,618,645]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="48" pageNumber="49" unit="m" value="850.0">850 m</quantity>
elevation, in rainforest. Nests were located in the soil and on clay banks.
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FFBEA926A788B4BAFAE4FC11" pageId="48" pageNumber="49" type="discussion">
<paragraph id="8B853133FFBEA926A788B4BAFAE4FC11" blockId="48.[151,1437,150,1041]" pageId="48" pageNumber="49">
<emphasis id="B94EED21FFBEA926A788B4BAFE89FD31" bold="true" box="[199,335,690,716]" pageId="48" pageNumber="49">Comments.</emphasis>
<taxonomicName id="4C3A4AB0FFBEA926A61AB4BAFDE9FD31" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[341,559,690,716]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="ovalinoda">
<emphasis id="B94EED21FFBEA926A61AB4BAFDE9FD31" box="[341,559,690,716]" italics="true" pageId="48" pageNumber="49">Pheidole ovalinoda</emphasis>
</taxonomicName>
is known only from Parc National de Marojejy in
<collectingRegion id="49FEFFD1FFBEA926A314B4BAFB20FD31" box="[1115,1254,690,716]" country="Madagascar" name="Antsiranana" pageId="48" pageNumber="49">Antsiranana</collectingRegion>
. Its major workers can be confused with parapatric
<taxonomicName id="4C3A4AB0FFBEA926A56BB4DFFD59FD0D" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[548,671,726,752]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="flammea">
<emphasis id="B94EED21FFBEA926A56BB4DFFD59FD0D" box="[548,671,726,752]" italics="true" pageId="48" pageNumber="49">P. flammea</emphasis>
</taxonomicName>
, recorded only from Galoko Mountain. Majors of
<taxonomicName id="4C3A4AB0FFBEA926A381B4DFFA9FFD0D" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[1230,1369,726,752]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="ovalinoda">
<emphasis id="B94EED21FFBEA926A381B4DFFA9FFD0D" box="[1230,1369,726,752]" italics="true" pageId="48" pageNumber="49">P. ovalinoda</emphasis>
</taxonomicName>
differ from
<taxonomicName id="4C3A4AB0FFBEA926A79AB4F3FE97FCE9" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[213,337,762,788]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="flammea">
<emphasis id="B94EED21FFBEA926A79AB4F3FE97FCE9" box="[213,337,762,788]" italics="true" pageId="48" pageNumber="49">P. flammea</emphasis>
</taxonomicName>
in the presence of dense, short and decumbent to suberect setae on the sides of the head, strongly reduced outer hypostomal tooth, and more distinctly sculptured interspaces between rugae on head. Majors of
<taxonomicName id="4C3A4AB0FFBEA926A2C7B517FF31FCA1" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="flammea">
<emphasis id="B94EED21FFBEA926A2C7B517FF31FCA1" italics="true" pageId="48" pageNumber="49">P. flammea</emphasis>
</taxonomicName>
have moderately dense, long and suberect to erect pilosity on the side of the head, large outer hypostomal tooth, and mostly smooth interspaces between rugae on head. Minors of
<taxonomicName id="4C3A4AB0FFBEA926A4F6B56FFB85FC7D" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[953,1091,870,896]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="ovalinoda">
<emphasis id="B94EED21FFBEA926A4F6B56FFB85FC7D" box="[953,1091,870,896]" italics="true" pageId="48" pageNumber="49">P. ovalinoda</emphasis>
</taxonomicName>
are easy to separate from other
<taxonomicName id="4C3A4AB0FFBEA926A7D8B582FF3DFC59" authorityName="Westwood" authorityYear="1839" box="[151,251,906,932]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B94EED21FFBEA926A7D8B582FF3DFC59" box="[151,251,906,932]" italics="true" pageId="48" pageNumber="49">Pheidole</emphasis>
</taxonomicName>
known from the island based on the combination of the following characters: petiolar node high, triangular, and thick; postpetiole short and moderately high; yellowish brown body coloration; neck absent, and nuchal collar indistinct; head and promesonotum mostly smooth; promesonotal and metanotal grooves distinct.
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FFBEA926A788B5FEFA9AFBED" box="[199,1372,1014,1041]" pageId="48" pageNumber="49" type="etymology">
<paragraph id="8B853133FFBEA926A788B5FEFA9AFBED" blockId="48.[151,1437,150,1041]" box="[199,1372,1014,1041]" pageId="48" pageNumber="49">
<emphasis id="B94EED21FFBEA926A788B5FEFE88FBED" bold="true" box="[199,334,1014,1040]" pageId="48" pageNumber="49">Etymology.</emphasis>
Latin for oval and node, in reference to the high and spherical postpetiole of minor workers.
</paragraph>
</subSubSection>
</treatment>
</document>