treatments-xml/data/03/93/05/03930553FFCE3018FEC1FC111EE2FE4D.xml
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<document id="961F3411A02AED4468F9C832C06C696F" ID-DOI="10.5281/zenodo.273223" ID-GBIF-Dataset="5ddffa85-25af-4c28-b891-100cdfec2c23" ID-ISSN="1175­5326" ID-Zenodo-Dep="273223" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459716340439" checkinUser="plazi" docAuthor="Perna, Rafael La" docDate="2005" docId="03930553FFCE3018FEC1FC111EE2FE4D" docLanguage="en" docName="zt00881.pdf" docOrigin="Zootaxa 881" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Nucinella boucheti Perna, 2005, n.sp." docType="treatment" docVersion="8" lastPageNumber="6" masterDocId="FFAA7D2BFFCC301EFFC9FFEA1A72FFCB" masterDocTitle="A gigantic deep­sea Nucinellidae from the tropical West Pacific (Bivalvia: Protobranchia)" masterLastPageNumber="10" masterPageNumber="1" pageNumber="3" updateTime="1698208037436" updateUser="plazi">
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<mods:title id="ADE609270CF4114407DF0CD15C6698FA">A gigantic deep­sea Nucinellidae from the tropical West Pacific (Bivalvia: Protobranchia)</mods:title>
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<mods:namePart id="2EA00C4CB98FBC9FEC18276107C770D4">Perna, Rafael La</mods:namePart>
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<mods:date id="F0D4E0C35B37FB7EDF80FB7ECCFC691B">2005</mods:date>
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<treatment id="03930553FFCE3018FEC1FC111EE2FE4D" ID-DOI="http://doi.org/10.5281/zenodo.6264975" ID-GBIF-Taxon="119334694" ID-Zenodo-Dep="6264975" LSID="urn:lsid:plazi:treatment:03930553FFCE3018FEC1FC111EE2FE4D" httpUri="http://treatment.plazi.org/id/03930553FFCE3018FEC1FC111EE2FE4D" lastPageId="6" lastPageNumber="6" pageId="2" pageNumber="3">
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<paragraph id="8B85B445FFCE301CFEC1FC111843FBDE" blockId="2.[264,561,1019,1080]" box="[264,561,1019,1045]" pageId="2" pageNumber="3">
<heading id="D0CD0329FFCE301CFEC1FC111843FBDE" bold="true" box="[264,561,1019,1045]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
<emphasis id="B94E6857FFCE301CFEC1FC111843FBDE" bold="true" box="[264,561,1019,1045]" pageId="2" pageNumber="3">
<taxonomicName id="4C3ACFC6FFCE301CFEC1FC111B9CFBDE" box="[264,494,1019,1045]" class="Bivalvia" family="Manzanellidae" genus="Nucinella" kingdom="Animalia" order="Solemyoida" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="boucheti" status="sp. nov.">
<emphasis id="B94E6857FFCE301CFEC1FC111B9CFBDE" bold="true" box="[264,494,1019,1045]" italics="true" pageId="2" pageNumber="3">Nucinella boucheti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27DD52CFFCE301CFE3FFC111843FBDE" box="[502,561,1019,1045]" pageId="2" pageNumber="3" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B85B445FFCE301CFEC1FBF41B07FBF3" blockId="2.[264,561,1019,1080]" box="[264,373,1054,1080]" pageId="2" pageNumber="3">
<figureCitation id="1301A8C0FFCE301CFEC1FBF41B28FBF3" box="[264,346,1054,1080]" captionStart="FIGURE 2" captionStartId="4.[264,367,1748,1772]" captionTargetBox="[269,1316,449,1722]" captionTargetId="figure@4.[269,1317,449,1722]" captionTargetPageId="4" captionText="FIGURE 2. Nucinella boucheti n. sp. A, B. Holotype, height 24 mm. C, D. Paratype 1, height 18 mm. E. Paratype 2, height 21 mm. F. Paratype 3, height 20 mm. G. Paratype 4, height 15 mm." httpUri="https://zenodo.org/record/273225/files/figure.png" pageId="2" pageNumber="3">Figs. 2</figureCitation>
,
<figureCitation id="1301A8C0FFCE301CFEAFFBF41B07FBF3" box="[358,373,1054,1080]" captionStart="FIGURE 3" captionStartId="5.[264,368,1558,1582]" captionTargetBox="[269,1316,283,1550]" captionTargetId="figure@5.[269,1317,281,1554]" captionTargetPageId="5" captionText="FIGURE 3. Nucinella boucheti n. sp. A C. Holotype, A. hinge (arrow: ligament pit), B. posterior teeth, C. anterior teeth. D, E. Paratype 2, hinge. F, G. Paratype 4, hinge. H. Paratype 3, ligament (arrows: periostracum). Scale bars: 1 mm." httpUri="https://zenodo.org/record/273226/files/figure.png" pageId="2" pageNumber="3">3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C320E7CEFFCE301CFEC1FB8418DAFA9B" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="8B85B445FFCE301CFEC1FB841EBAFB7B" blockId="2.[264,1324,1134,1880]" pageId="2" pageNumber="3">
<emphasis id="B94E6857FFCE301CFEC1FB841BCFFB43" bold="true" box="[264,445,1134,1160]" pageId="2" pageNumber="3">
<typeStatus id="54810AE7FFCE301CFEC1FB841B37FB43" box="[264,325,1134,1160]" pageId="2" pageNumber="3">Type</typeStatus>
locality.
</emphasis>
MUSORSTOM 2 st. CP 42,
<geoCoordinate id="EE0ED282FFCE301CFCBCFB8419E0FB43" box="[885,914,1134,1160]" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="925" value="13.083333">13</geoCoordinate>
°05N
<geoCoordinate id="EE0ED282FFCE301CFC23FB841E29FB43" box="[1002,1115,1134,1160]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="925" value="122.416664">122°25E</geoCoordinate>
, 1610
<quantity id="4CC219A0FFCE301CFB08FB841F58FB43" box="[1217,1322,1134,1160]" metricMagnitude="3" metricUnit="m" metricValue="1.58" pageId="2" pageNumber="3" unit="m" value="1580.0">1580 m</quantity>
(
<date id="FF849285FFCE301CFED9FB7C1BEBFB7B" box="[272,409,1174,1200]" pageId="2" pageNumber="3" value="1980-11-25">25.11.1980</date>
), southwest off Bondoc Peninsula, Luzon Island,
<collectingCountry id="F32DF4D5FFCE301CFC17FB7C1E10FB7B" box="[990,1122,1174,1200]" name="Philippines" pageId="2" pageNumber="3">Philippines</collectingCountry>
(
<figureCitation id="1301A8C0FFCE301CFBB8FB7C1EC4FB7B" box="[1137,1206,1174,1200]" captionStart="FIGURE 1" captionStartId="2.[264,367,901,925]" captionTargetBox="[273,1301,417,867]" captionTargetId="figure@2.[267,1310,412,881]" captionTargetPageId="2" captionText="FIGURE 1. Location of the MUSORSTOM 2 station CP 42." httpUri="https://zenodo.org/record/273224/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
).
</paragraph>
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<emphasis id="B94E6857FFCE301CFEF1FB541B99FB13" bold="true" box="[312,491,1214,1240]" pageId="2" pageNumber="3">
<typeStatus id="54810AE7FFCE301CFEF1FB541B01FB13" box="[312,371,1214,1240]" pageId="2" pageNumber="3">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54810AE7FFCE301CFE3CFB541810FB13" box="[501,610,1214,1240]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
(right valve) and 18
<typeStatus id="54810AE7FFCE301CFC91FB5419BAFB13" box="[856,968,1214,1240]" pageId="2" pageNumber="3" type="paratype">paratypes</typeStatus>
(
<collectionCode id="ED2B2C80FFCE301CFC1EFB541E47FB13" LSID="urn:lsid:biocol.org:col:34988" box="[983,1077,1214,1240]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="3">MNHN</collectionCode>
, 4411, 4412).
</paragraph>
<paragraph id="8B85B445FFCE301CFEF1FB0C18DAFA9B" blockId="2.[264,1324,1134,1880]" pageId="2" pageNumber="3">
<emphasis id="B94E6857FFCE301CFEF1FB0C185EFACB" bold="true" box="[312,556,1254,1280]" pageId="2" pageNumber="3">Material examined.</emphasis>
Only the
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material, consisting of 18 valves and a single livecollected specimen. The valves surface is somewhat corroded, particularly on the umbo, and the periostracum is mostly lost.
</paragraph>
</subSubSection>
<subSubSection id="C320E7CEFFCE301CFEF1FAB419C8FA6B" pageId="2" pageNumber="3" type="etymology">
<paragraph id="8B85B445FFCE301CFEF1FAB419C8FA6B" blockId="2.[264,1324,1134,1880]" pageId="2" pageNumber="3">
<emphasis id="B94E6857FFCE301CFEF1FAB41BB6FAB3" bold="true" box="[312,452,1374,1400]" pageId="2" pageNumber="3">Etymology.</emphasis>
Named after the French malacologist Philippe Bouchet, in honor of his intense research activity on tropical West Pacific molluscs.
</paragraph>
</subSubSection>
<subSubSection id="C320E7CEFFCE301CFEF1FA44190EFA03" box="[312,892,1454,1480]" pageId="2" pageNumber="3" type="distribution">
<paragraph id="8B85B445FFCE301CFEF1FA44190EFA03" blockId="2.[264,1324,1134,1880]" box="[312,892,1454,1480]" pageId="2" pageNumber="3">
<emphasis id="B94E6857FFCE301CFEF1FA441BA3FA03" bold="true" box="[312,465,1454,1480]" pageId="2" pageNumber="3">Distribution</emphasis>
. Only known from the
<typeStatus id="54810AE7FFCE301CFD2CFA441965FA03" box="[741,791,1454,1480]" pageId="2" pageNumber="3">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C320E7CEFFCE301CFEF1FA3C19E9F9D3" pageId="2" pageNumber="3" type="diagnosis">
<paragraph id="8B85B445FFCE301CFEF1FA3C19E9F9D3" blockId="2.[264,1324,1134,1880]" pageId="2" pageNumber="3">
<emphasis id="B94E6857FFCE301CFEF1FA3C1BC2FA3B" bold="true" box="[312,432,1494,1520]" pageId="2" pageNumber="3">Diagnosis</emphasis>
. An unusually large species for the family, up to
<quantity id="4CC219A0FFCE301CFBC1FA3C1E2FFA3B" box="[1032,1117,1494,1520]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="3" unit="mm" value="25.0">25 mm</quantity>
in height, with a markedly convex, nuculiform shell and a delicate hinge.
</paragraph>
</subSubSection>
<subSubSection id="C320E7CEFFCE301DFEF1F9CC1807FC45" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="description">
<paragraph id="8B85B445FFCE301DFEF1F9CC1B88FCF5" blockId="2.[264,1324,1134,1880]" lastBlockId="3.[264,1324,284,1870]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
<emphasis id="B94E6857FFCE301CFEF1F9CC1BBBF98B" bold="true" box="[312,457,1574,1600]" pageId="2" pageNumber="3">Description</emphasis>
. Shell equivalve, strongly inequilateral, obliquely ovate, nuculiform, markedly convex. Shell wall thin, fragile. Umbo large, projecting well beyond dorsal margin, opisthogyrous. Antero­dorsal margin short, convex, obtusely curving to a slightly convex anterior margin. Ventral margin strongly convex, smoothly curving to a long, barely convex postero­ventral margin. Postero­dorsal margin short, slightly concave, forming a smooth, narrow curve when joining to posterior margin. Junction between postero­dorsal and postero­ventral margin as an obtuse smooth angulation. Hinge plate not strong, wide, extending from postero­dorsal margin to about 2/3 of anterior side length, with a wide, flat, somewhat triangular area below dentition. Subumbonal teeth of different shape, size and orientation, in a single, arched series, numbering about
<quantity id="4CC219A0FFCF301DFC0EFEAE1E71FE95" box="[967,1027,324,350]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" pageId="3" pageNumber="4" unit="in" value="13.0">13 in</quantity>
<typeStatus id="54810AE7FFCF301DFBC2FEAE1E07FE95" box="[1035,1141,324,350]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
. Posterior teeth mainly crescent­shaped, the last one is a blunt tubercle. Below beak, few irregularly shaped to lamellar teeth. Anterior teeth mainly chevron­shaped. Lateral dentition consisting of a rather strong, lamellar, grossly triangular tooth, continuous with hinge plate margin, with rather deep longitudinal grooves, and a wide submarginal socket with 23 small ridge­like teeth inside. Ligament pit external, wide, immediately posterior to beak, occupying all postero­dorsal margin. Ligament pit wide, deep; pit margins with a deep groove for ligament insertion. Ligament thick, strongly bulging, partly covered by a thick periostracum. Inner valve surface white, dull. Pallial line fairly well distinct, no sinus. Anterior muscle scar large, ovate, marginal in position, below shell height midpoint; no posterior scar. Outer surface with shallow, ill defined, concentric riblets and thin growth striae. Periostracum rather thick, shining, varnish­like, greenish­browinsh, mostly not preserved. Prodissoconch not seen.
</paragraph>
<paragraph id="8B85B445FFCF301DFEF1FCA61807FC45" blockId="3.[264,1324,284,1870]" pageId="3" pageNumber="4">
<emphasis id="B94E6857FFCF301DFEF1FCA61BBBFCAD" bold="true" box="[312,457,844,870]" pageId="3" pageNumber="4">Dimensions</emphasis>
.
<typeStatus id="54810AE7FFCF301DFE17FCA61839FCAD" box="[478,587,844,870]" pageId="3" pageNumber="4" type="holotype">Holotype</typeStatus>
<quantity id="4CC219A0FFCF301DFD93FCA618C6FCAD" box="[602,692,844,870]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="3" pageNumber="4" unit="mm" value="24.0">24 mm</quantity>
in height,
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in length,
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in thickness.
<typeStatus id="54810AE7FFCF301DFEC1FC9E1B08FC45" box="[264,378,884,910]" pageId="3" pageNumber="4" type="paratype">Paratypes</typeStatus>
<quantity id="4CC219A0FFCF301DFE48FC9E1B8DFC45" box="[385,511,884,910]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="3" pageNumber="4" unit="mm" value="20.0" valueMax="25.0" valueMin="15.0">1525 mm</quantity>
in height.
</paragraph>
</subSubSection>
<subSubSection id="C320E7CEFFCF3018FEF1FC761EE2FE4D" lastPageId="6" lastPageNumber="7" pageId="3" pageNumber="4" type="discussion">
<paragraph id="8B85B445FFCF301DFEF1FC761ECAFAD5" blockId="3.[264,1324,284,1870]" pageId="3" pageNumber="4">
<emphasis id="B94E6857FFCF301DFEF1FC761BD8FC7D" bold="true" box="[312,426,924,950]" pageId="3" pageNumber="4">Remarks</emphasis>
. Because of its large size and distinct hinge (see below),
<taxonomicName id="4C3ACFC6FFCF301DFB8DFC761EB9FC7E" box="[1092,1227,924,949]" class="Bivalvia" family="Manzanellidae" genus="Nucinella" kingdom="Animalia" order="Solemyoida" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="boucheti">
<emphasis id="B94E6857FFCF301DFB8DFC761EB9FC7E" box="[1092,1227,924,949]" italics="true" pageId="3" pageNumber="4">N. boucheti</emphasis>
</taxonomicName>
is strikingly different from all the other species in the genus. The sole species which may deserve some comparisons with the present one, is
<taxonomicName id="4C3ACFC6FFCF301DFCCDFC061EBAFBCD" authority="Thiele and Jaeckel, 1931" authorityName="Thiele and Jaeckel" authorityYear="1931" box="[772,1224,1004,1030]" class="Bivalvia" family="Manzanellidae" genus="Nucinella" kingdom="Animalia" order="Solemyoida" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="maxima">
<emphasis id="B94E6857FFCF301DFCCDFC0619F6FBCE" box="[772,900,1004,1029]" italics="true" pageId="3" pageNumber="4">N. maxima</emphasis>
(Thiele and Jaeckel, 1931)
</taxonomicName>
, a deepsea species from
<collectingCountry id="F32DF4D5FFCF301DFE1EFBFE184DFBE5" box="[471,575,1044,1070]" name="Tanzania" pageId="3" pageNumber="4">Zanzibar</collectingCountry>
Strait. This species is notably larger (12.5 mm) than the usual size range of the family and it actually was the largest known species before the present description. Under this name,
<bibRefCitation id="EFABC9B4FFCF301DFDAEFB8E1997FBB5" author="Kusnetsov" box="[615,997,1124,1150]" pageId="3" pageNumber="10" refString="Kusnetsov, A. P. &amp; Schileyko, A. A. (1984) On the gutless Protobranchia (Bivalvia). Biologicheskie nauki, 2, 39 - 49 (in Russian)." type="journal article" year="1984">Kusnetsov and Schileyko (1984)</bibRefCitation>
reported a deep­sea species from the Gulf of Aden, but their species is much smaller (about
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) and roughly rectangular in shape. Another particularly large species is
<taxonomicName id="4C3ACFC6FFCF301DFCB6FB5E1F58FB05" authority="Matsukuma et al., 1982" authorityName="Matsukuma et al." authorityYear="1982" box="[895,1322,1204,1230]" class="Bivalvia" family="Manzanellidae" genus="Nucinella" kingdom="Animalia" order="Solemyoida" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="viridis">
<emphasis id="B94E6857FFCF301DFCB6FB5E199DFB06" box="[895,1007,1204,1229]" italics="true" pageId="3" pageNumber="4">N. viridis</emphasis>
(
<bibRefCitation id="EFABC9B4FFCF301DFBC8FB5E1F50FB05" author="Matsukuma" box="[1025,1314,1204,1230]" pageId="3" pageNumber="10" refString="Matsukuma, A., Okutani, T. &amp; Tsuchi, R. (1982) Three new species of the Nucinellidae (Bivalvia: Protobranchia) from the Pacific coast of Japan. Ven us, 40, 177 - 186." type="journal article" year="1982">
Matsukuma
<emphasis id="B94E6857FFCF301DFB5EFB5E1EA7FB06" box="[1175,1237,1204,1229]" italics="true" pageId="3" pageNumber="4">et al.</emphasis>
, 1982
</bibRefCitation>
)
</taxonomicName>
(about
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), from the Central East
<collectingCountry id="F32DF4D5FFCF301DFD07FB361963FB3D" box="[718,785,1244,1270]" name="Japan" pageId="3" pageNumber="4">Japan</collectingCountry>
in
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<quantity id="4CC219A0FFCF301DFCFEFB3619F4FB3D" box="[823,902,1244,1270]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="4" unit="m" value="500.0">500 m</quantity>
.
</specimenCount>
Like the present species,
<taxonomicName id="4C3ACFC6FFCF301DFB75FB361F59FB3E" box="[1212,1323,1244,1269]" class="Bivalvia" family="Manzanellidae" genus="Nucinella" kingdom="Animalia" order="Solemyoida" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="viridis">
<emphasis id="B94E6857FFCF301DFB75FB361F59FB3E" box="[1212,1323,1244,1269]" italics="true" pageId="3" pageNumber="4">N. viridis</emphasis>
</taxonomicName>
has a marked nuculiform shape, but its hinge consists of few, rather coarse teeth.
</paragraph>
<paragraph id="8B85B445FFCF301AFEF1FAC61BAEFE95" blockId="3.[264,1324,284,1870]" lastBlockId="4.[264,1323,284,350]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
The hinge of
<taxonomicName id="4C3ACFC6FFCF301DFE1EFAC6182DFA8E" box="[471,607,1324,1349]" class="Bivalvia" family="Manzanellidae" genus="Nucinella" kingdom="Animalia" order="Solemyoida" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="boucheti">
<emphasis id="B94E6857FFCF301DFE1EFAC6182DFA8E" box="[471,607,1324,1349]" italics="true" pageId="3" pageNumber="4">N. boucheti</emphasis>
</taxonomicName>
contrasts with the typical hinge of the genus (Bernard 1987;
<bibRefCitation id="EFABC9B4FFCF301DFEC1FABE1BE7FAA5" author="Vokes" box="[264,405,1364,1390]" pageId="3" pageNumber="10" refString="Vokes, H. E. (1956) Notes on the Nucinellidae (Pelecypoda) with description of a new species from the Eocene of Oregon. Journal of Paleontology, 30, 652 - 671." type="journal article" year="1956">Vokes 1956</bibRefCitation>
;
<bibRefCitation id="EFABC9B4FFCF301DFE68FABE18C7FAA5" author="Matsukuma" box="[417,693,1364,1390]" pageId="3" pageNumber="10" refString="Matsukuma, A., Okutani, T. &amp; Tsuchi, R. (1982) Three new species of the Nucinellidae (Bivalvia: Protobranchia) from the Pacific coast of Japan. Ven us, 40, 177 - 186." type="journal article" year="1982">
Matsukuma
<emphasis id="B94E6857FFCF301DFDFDFABE181AFAA6" box="[564,616,1364,1389]" italics="true" pageId="3" pageNumber="4">et al</emphasis>
. 1982
</bibRefCitation>
;
<bibRefCitation id="EFABC9B4FFCF301DFD0BFABE1923FAA5" author="Pojeta" box="[706,849,1364,1390]" pageId="3" pageNumber="10" refString="Pojeta, J. Jr. (1988) The origin and Paleozoic diversification of solemyoid pelecypods. In Wolberg D. L. (Ed.) Contributions to Paleozoic paleontology and stratigraphy in honor of Rousseau H. Flower, New Mexico Bureau of Mines &amp; Mineral Resources, Memoirs, 44, 201 - 222." type="journal article" year="1988">Pojeta 1988</bibRefCitation>
;
<bibRefCitation id="EFABC9B4FFCF301DFC97FABE1E10FAA5" author="Beu" box="[862,1122,1364,1390]" pageId="3" pageNumber="9" refString="Beu, A. G. &amp; Maxwell, P. A. (1990) Cenozoic Mollusca of New Zealand. New Zealand Geological Survey Paleontological Bulletin, 58, 1 - 518." type="journal article" year="1990">Beu &amp; Maxwell 1990</bibRefCitation>
;
<bibRefCitation id="EFABC9B4FFCF301DFBA6FABE1F6DFAA5" author="La" box="[1135,1311,1364,1390]" pageId="3" pageNumber="10" refString="La Perna, R. (2004) Nucinella alibrandi (Conti, 1864) and N. seguenzae (Dall, 1898), the last European nucinellids (Bivalvia, Protobranchia). Rivista Italiana di Paleontologia e Stratigrafia, 110, 571 - 577." type="journal article" year="2004">La Perna 2004</bibRefCitation>
), which has few subumbonal teeth (generally not exceeding six), somewhat different in size and orientation, rather stout (“peg­like”) and coarse, giving a rather strong appearance. In spite of the large shell size, the present species has small and thin subumbonal teeth, giving a delicate appearance to the hinge, and they also exhibit some unusual shapes. The posterior teeth are oddly crescent­shaped, with the convex side towards the beak (
<figureCitation id="1301A8C0FFCF301DFB08F9F61F74F9FD" box="[1217,1286,1564,1590]" captionStart="FIGURE 3" captionStartId="5.[264,368,1558,1582]" captionTargetBox="[269,1316,283,1550]" captionTargetId="figure@5.[269,1317,281,1554]" captionTargetPageId="5" captionText="FIGURE 3. Nucinella boucheti n. sp. A C. Holotype, A. hinge (arrow: ligament pit), B. posterior teeth, C. anterior teeth. D, E. Paratype 2, hinge. F, G. Paratype 4, hinge. H. Paratype 3, ligament (arrows: periostracum). Scale bars: 1 mm." httpUri="https://zenodo.org/record/273226/files/figure.png" pageId="3" pageNumber="4">Fig. 3</figureCitation>
B), whereas the last posterior one is stout, roundish to elliptical, or with a strange circular shape (
<figureCitation id="1301A8C0FFCF301DFEA9F9861BD8F94D" box="[352,426,1644,1670]" captionStart="FIGURE 3" captionStartId="5.[264,368,1558,1582]" captionTargetBox="[269,1316,283,1550]" captionTargetId="figure@5.[269,1317,281,1554]" captionTargetPageId="5" captionText="FIGURE 3. Nucinella boucheti n. sp. A C. Holotype, A. hinge (arrow: ligament pit), B. posterior teeth, C. anterior teeth. D, E. Paratype 2, hinge. F, G. Paratype 4, hinge. H. Paratype 3, ligament (arrows: periostracum). Scale bars: 1 mm." httpUri="https://zenodo.org/record/273226/files/figure.png" pageId="3" pageNumber="4">Fig. 3</figureCitation>
G). The central teeth are thin, lamellar or irregularly shaped (resorbed?) (
<figureCitation id="1301A8C0FFCF301DFED8F97E1B11F965" box="[273,355,1684,1710]" captionStart="FIGURE 3" captionStartId="5.[264,368,1558,1582]" captionTargetBox="[269,1316,283,1550]" captionTargetId="figure@5.[269,1317,281,1554]" captionTargetPageId="5" captionText="FIGURE 3. Nucinella boucheti n. sp. A C. Holotype, A. hinge (arrow: ligament pit), B. posterior teeth, C. anterior teeth. D, E. Paratype 2, hinge. F, G. Paratype 4, hinge. H. Paratype 3, ligament (arrows: periostracum). Scale bars: 1 mm." httpUri="https://zenodo.org/record/273226/files/figure.png" pageId="3" pageNumber="4">Figs. 3</figureCitation>
B, E, G). The anterior teeth are slightly stronger, asymmetrically chevron­shaped, with a short dorsal branch parallel to shell margin (
<figureCitation id="1301A8C0FFCF301DFCA6F95619B3F91D" box="[879,961,1724,1750]" captionStart="FIGURE 3" captionStartId="5.[264,368,1558,1582]" captionTargetBox="[269,1316,283,1550]" captionTargetId="figure@5.[269,1317,281,1554]" captionTargetPageId="5" captionText="FIGURE 3. Nucinella boucheti n. sp. A C. Holotype, A. hinge (arrow: ligament pit), B. posterior teeth, C. anterior teeth. D, E. Paratype 2, hinge. F, G. Paratype 4, hinge. H. Paratype 3, ligament (arrows: periostracum). Scale bars: 1 mm." httpUri="https://zenodo.org/record/273226/files/figure.png" pageId="3" pageNumber="4">Figs. 3</figureCitation>
E, C), whereas the most distal ones are elliptical or elongated (
<figureCitation id="1301A8C0FFCF301DFD41F90E18A8F935" box="[648,730,1764,1790]" captionStart="FIGURE 3" captionStartId="5.[264,368,1558,1582]" captionTargetBox="[269,1316,283,1550]" captionTargetId="figure@5.[269,1317,281,1554]" captionTargetPageId="5" captionText="FIGURE 3. Nucinella boucheti n. sp. A C. Holotype, A. hinge (arrow: ligament pit), B. posterior teeth, C. anterior teeth. D, E. Paratype 2, hinge. F, G. Paratype 4, hinge. H. Paratype 3, ligament (arrows: periostracum). Scale bars: 1 mm." httpUri="https://zenodo.org/record/273226/files/figure.png" pageId="3" pageNumber="4">Figs. 3</figureCitation>
A, C). Another difference consists in the arrangement of the subumbonal teeth, which do not extend all through the hinge plate height, whereas in the other species most part of hinge plate is occupied by teeth. The lateral dentition is similar to that of the other species, but the large lamellar tooth has longitudinal grooves (
<figureCitation id="1301A8C0FFC8301AFEBCFEAE1BC9FE95" box="[373,443,324,350]" captionStart="FIGURE 3" captionStartId="5.[264,368,1558,1582]" captionTargetBox="[269,1316,283,1550]" captionTargetId="figure@5.[269,1317,281,1554]" captionTargetPageId="5" captionText="FIGURE 3. Nucinella boucheti n. sp. A C. Holotype, A. hinge (arrow: ligament pit), B. posterior teeth, C. anterior teeth. D, E. Paratype 2, hinge. F, G. Paratype 4, hinge. H. Paratype 3, ligament (arrows: periostracum). Scale bars: 1 mm." httpUri="https://zenodo.org/record/273226/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
F).
</paragraph>
<caption id="DF45E4CDFFC8301AFEC1F93E1E9FF8C5" httpUri="https://zenodo.org/record/273225/files/figure.png" pageId="4" pageNumber="5" targetBox="[269,1316,449,1722]" targetPageId="4">
<paragraph id="8B85B445FFC8301AFEC1F93E1E9FF8C5" blockId="4.[264,1323,1748,1806]" pageId="4" pageNumber="5">
<emphasis id="B94E6857FFC8301AFEC1F93E1BFEF927" bold="true" box="[264,396,1748,1772]" pageId="4" pageNumber="5">FIGURE 2.</emphasis>
<taxonomicName id="4C3ACFC6FFC8301AFE5DF93F1812F927" box="[404,608,1749,1772]" class="Bivalvia" family="Manzanellidae" genus="Nucinella" kingdom="Animalia" order="Solemyoida" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="boucheti" status="sp. nov.">
<emphasis id="B94E6857FFC8301AFE5DF93F1812F927" box="[404,608,1749,1772]" italics="true" pageId="4" pageNumber="5">Nucinella boucheti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27DD52CFFC8301AFDA0F93F18E9F927" box="[617,667,1749,1772]" pageId="4" pageNumber="5" rank="species">n.sp.</taxonomicNameLabel>
A, B. Holotype, height 24 mm. C, D. Paratype 1, height 18 mm. E. Paratype 2, height 21 mm. F. Paratype 3, height 20 mm. G. Paratype 4, height 15 mm.
</paragraph>
</caption>
<caption id="DF45E4CDFFC9301BFEC1F9FC18BAF9B9" httpUri="https://zenodo.org/record/273226/files/figure.png" pageId="5" pageNumber="6" targetBox="[269,1316,283,1550]" targetPageId="5">
<paragraph id="8B85B445FFC9301BFEC1F9FC18BAF9B9" blockId="5.[264,1323,1558,1650]" pageId="5" pageNumber="6">
<emphasis id="B94E6857FFC9301BFEC1F9FC1BFEF9E5" bold="true" box="[264,396,1558,1582]" pageId="5" pageNumber="6">FIGURE 3.</emphasis>
<taxonomicName id="4C3ACFC6FFC9301BFE5DF9FC1812F9E6" box="[404,608,1558,1581]" class="Bivalvia" family="Manzanellidae" genus="Nucinella" kingdom="Animalia" order="Solemyoida" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="boucheti" status="sp. nov.">
<emphasis id="B94E6857FFC9301BFE5DF9FC1812F9E6" box="[404,608,1558,1581]" italics="true" pageId="5" pageNumber="6">Nucinella boucheti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27DD52CFFC9301BFDA1F9FC18E8F9E6" box="[616,666,1558,1581]" pageId="5" pageNumber="6" rank="species">n.sp.</taxonomicNameLabel>
AC. Holotype, A. hinge (arrow: ligament pit), B. posterior teeth, C. anterior teeth. D, E. Paratype 2, hinge. F, G. Paratype 4, hinge. H. Paratype 3, ligament (arrows: periostracum). Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="8B85B445FFC93018FEF1F94C1EE2FE4D" blockId="5.[264,1323,1702,1888]" lastBlockId="6.[264,1323,284,390]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
The ligament is thick, strongly bulging and partly covered by a rather thick periostracum (
<figureCitation id="1301A8C0FFC9301BFEA5F9241BC1F923" box="[364,435,1742,1768]" captionStart="FIGURE 3" captionStartId="5.[264,368,1558,1582]" captionTargetBox="[269,1316,283,1550]" captionTargetId="figure@5.[269,1317,281,1554]" captionTargetPageId="5" captionText="FIGURE 3. Nucinella boucheti n. sp. A C. Holotype, A. hinge (arrow: ligament pit), B. posterior teeth, C. anterior teeth. D, E. Paratype 2, hinge. F, G. Paratype 4, hinge. H. Paratype 3, ligament (arrows: periostracum). Scale bars: 1 mm." httpUri="https://zenodo.org/record/273226/files/figure.png" pageId="5" pageNumber="6">Fig. 3</figureCitation>
H), within a wide and deep attachment area, whose margins forms a deep groove in the shell wall for the ligament insertion (
<figureCitation id="1301A8C0FFC9301BFCA4F91C19CDF8DB" box="[877,959,1782,1808]" captionStart="FIGURE 3" captionStartId="5.[264,368,1558,1582]" captionTargetBox="[269,1316,283,1550]" captionTargetId="figure@5.[269,1317,281,1554]" captionTargetPageId="5" captionText="FIGURE 3. Nucinella boucheti n. sp. A C. Holotype, A. hinge (arrow: ligament pit), B. posterior teeth, C. anterior teeth. D, E. Paratype 2, hinge. F, G. Paratype 4, hinge. H. Paratype 3, ligament (arrows: periostracum). Scale bars: 1 mm." httpUri="https://zenodo.org/record/273226/files/figure.png" pageId="5" pageNumber="6">Figs. 3</figureCitation>
A, D). There is a thin external lamellar layer, and a thicker, partly mineralised, coarsely fibrous internal layer. The ligament attachment area of
<taxonomicName id="4C3ACFC6FFC9301BFDFBF8AC18C9F894" box="[562,699,1862,1887]" class="Bivalvia" family="Manzanellidae" genus="Nucinella" kingdom="Animalia" order="Solemyoida" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="boucheti">
<emphasis id="B94E6857FFC9301BFDFBF8AC18C9F894" box="[562,699,1862,1887]" italics="true" pageId="5" pageNumber="6">N. boucheti</emphasis>
</taxonomicName>
is not different from that of the other species in the genus (e.g.,
<bibRefCitation id="EFABC9B4FFCA3018FE5CFEF61856FEFD" author="Pojeta" box="[405,548,284,310]" pageId="6" pageNumber="10" refString="Pojeta, J. Jr. (1988) The origin and Paleozoic diversification of solemyoid pelecypods. In Wolberg D. L. (Ed.) Contributions to Paleozoic paleontology and stratigraphy in honor of Rousseau H. Flower, New Mexico Bureau of Mines &amp; Mineral Resources, Memoirs, 44, 201 - 222." type="journal article" year="1988">Pojeta 1988</bibRefCitation>
: pl. 5, fig. 3; pl. 7 fig. 6;
<bibRefCitation id="EFABC9B4FFCA3018FC8BFEF61E27FEFD" author="Beu" box="[834,1109,284,310]" pageId="6" pageNumber="9" refString="Beu, A. G. &amp; Maxwell, P. A. (1990) Cenozoic Mollusca of New Zealand. New Zealand Geological Survey Paleontological Bulletin, 58, 1 - 518." type="journal article" year="1990">Beu and Maxwell 1990</bibRefCitation>
, pl. 50, fig. b), but the large size allows the observations by
<bibRefCitation id="EFABC9B4FFCA3018FD2EFEAE19F6FE95" author="Waller" box="[743,900,324,350]" pageId="6" pageNumber="10" refString="Waller, T. R. (1990) The evolution of ligament systems in the Bivalvia. In: Morton, B. (Ed.), The Bivalvia. Proceedings of a Memorial Symposium in Honour of Sir Charles Maurice Yonge, Edinburgh, 1986, Hong Kong University Press, Hong Kong, pp. 49 - 71." type="journal article" year="1990">Waller (1990</bibRefCitation>
;
<bibRefCitation id="EFABC9B4FFCA3018FC59FEAE19BEFE95" author="Waller" box="[912,972,324,350]" pageId="6" pageNumber="10" refString="Waller, T. R. (1998) Origin of the molluscan class Bivalvia and a phylogeny of major groups. In Johnston, P. A. &amp; Haggart, J. W. (Eds) Bivalves: An Eon of Evolution, Paleobiological Studies Honoring Norman D. Newell, University of Calgary Press, Calgary, 1 - 45." type="book chapter" year="1998">1998</bibRefCitation>
) to be confirmed,
<emphasis id="B94E6857FFCA3018FB6DFEAE1EBAFE96" box="[1188,1224,324,349]" italics="true" pageId="6" pageNumber="7">i.e.</emphasis>
the lack of nymphs in nucinellids (which are instead present in the other solemyoids).
</paragraph>
</subSubSection>
</treatment>
</document>