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<document id="AE7EEB4C4EB327112597C7A8A9F1E22B" ID-CLB-Dataset="148081" ID-DOI="10.11646/zootaxa.5188.6.2" ID-GBIF-Dataset="ecb5de27-f5fe-4b16-903b-633548186b0a" ID-ISSN="1175-5326" ID-Zenodo-Dep="7103618" ID-ZooBank="BF01B6B4-3415-41A1-86A0-F4B187541A55" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1663829136880" checkinUser="plazi" docAuthor="Brooks, Scott E. &amp; Cumming, Jeffrey M." docDate="2022" docId="038D87B8C8172000EBC3E2EA9C4AA953" docLanguage="en" docName="zootaxa.5186.6.2.pdf" docOrigin="Zootaxa 5188 (6)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chimerothalassius riparius Brooks &amp; Cumming 2022, sp. nov." docType="treatment" docUuid="CD5E6F66-2F72-4B6B-8235-30B3DDBE6D23" docUuidSource="ZooBank" docVersion="6" lastPageNumber="530" masterDocId="FFB4FFC0C8102009EB54E27D9F48AB33" masterDocTitle="New Australasian Parathalassiinae (Diptera: Dolichopodidae sensu lato)" masterLastPageNumber="543" masterPageNumber="521" pageNumber="528" updateTime="1699383858559" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="67A3C91F33B45EB3A7A8F3687F12F1B9">New Australasian Parathalassiinae (Diptera: Dolichopodidae sensu lato)</mods:title>
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<mods:namePart id="92790D84F5E5B65A6135E331FC2E6BEC">Brooks, Scott E.</mods:namePart>
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<mods:affiliation id="06FA692FD74748C87AEA7EE77AADB9D2">Diptera Unit, Canadian National Collection of Insects, Agriculture and Agri-Food Canada, K. W. Neatby Building, 960 Carling Avenue, Ottawa, Ontario, K 1 A 0 C 6, CANADA &amp; scott. brooks @ agr. gc. ca; https: // orcid. org / 0000 - 0001 - 7915 - 1941</mods:affiliation>
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<mods:namePart id="26B3EF8A8ABD46A0F2058C25EA27A096">Cumming, Jeffrey M.</mods:namePart>
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<mods:affiliation id="E64602BCA7CBFF277653F9828B073C1B">Diptera Unit, Canadian National Collection of Insects, Agriculture and Agri-Food Canada, K. W. Neatby Building, 960 Carling Avenue, Ottawa, Ontario, K 1 A 0 C 6, CANADA &amp; jeff. cumming @ agr. gc. ca; https: // orcid. org / 0000 - 0002 - 3619 - 3700</mods:affiliation>
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<mods:title id="E26F72DB36B51BE4E92D8EAD30A47D1E">Zootaxa</mods:title>
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<mods:date id="CC5378D4C0534A36044D9D64FB089E64">2022</mods:date>
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<treatment id="038D87B8C8172000EBC3E2EA9C4AA953" ID-DOI="http://doi.org/10.5281/zenodo.7103622" ID-GBIF-Taxon="201105648" ID-Zenodo-Dep="7103622" LSID="urn:lsid:zoobank.org:act:CD5E6F66-2F72-4B6B-8235-30B3DDBE6D23" httpUri="http://treatment.plazi.org/id/038D87B8C8172000EBC3E2EA9C4AA953" lastPageId="9" lastPageNumber="530" pageId="7" pageNumber="528">
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<emphasis id="B950EABCC817200EEBC3E2EA9D0EAB82" bold="true" box="[151,582,151,178]" pageId="7" pageNumber="528">
<taxonomicName id="4C244D2DC817200EEBC3E2EA9EA9AB82" ID-CoL="9J5QT" authority="Brooks &amp; Cumming, 2022" authorityName="Brooks &amp; Cumming" authorityYear="2022" box="[151,481,151,178]" class="Insecta" family="Dolichopodidae" genus="Chimerothalassius" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="528" phylum="Arthropoda" rank="species" species="riparius" status="sp. nov.">
<emphasis id="B950EABCC817200EEBC3E2EA9EA9AB82" bold="true" box="[151,481,151,178]" italics="true" pageId="7" pageNumber="528">Chimerothalassius riparius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357C7C817200EEABDE2EA9D0EAB82" box="[489,582,151,177]" pageId="7" pageNumber="528" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C33E6525C817200EEBC3E2C69C86ABC9" pageId="7" pageNumber="528" type="description">
<paragraph id="8B9B36AEC817200EEBC3E2C69E25ABE5" blockId="7.[151,974,151,250]" box="[151,365,187,214]" pageId="7" pageNumber="528">
(
<figureCitation id="131F2A2BC817200EEBCBE2C69E5AABE6" box="[159,274,187,213]" captionStart="FIGURES 811" captionStartId="2.[151,264,1908,1932]" captionTargetBox="[167,1420,842,1878]" captionTargetId="figure-39@2.[167,1420,833,1884]" captionTargetPageId="2" captionText="FIGURES 811. Known geographical distribution and collection localities of Chimerothalassius and Eothalassius from New Caledonia. 8. Distribution of C. riparius sp. nov., C. sinclairi sp. nov. and Eothalassius sp. 910. Rivère Bleue, type locality of C. riparius sp. nov. and C. sinclairi sp. nov. (photos by Brad Sinclair). 11. Plage de Poé, collection locality of Eothalassius sp. (photo by Brad Sinclair)." figureDoi="http://doi.org/10.5281/zenodo.7103626" httpUri="https://zenodo.org/record/7103626/files/figure.png" pageId="7" pageNumber="528">Figs 810</figureCitation>
,
<figureCitation id="131F2A2BC817200EEA4AE2C69E2DABE5" box="[286,357,187,214]" captionStart-0="FIGURES 1922" captionStart-1="FIGURES 2325" captionStartId-0="8.[151,264,1937,1961]" captionStartId-1="9.[151,264,1559,1583]" captionTargetBox-0="[219,1369,1001,1909]" captionTargetBox-1="[151,1426,641,1526]" captionTargetId-0="figure-389@8.[219,1369,993,1914]" captionTargetId-1="figure-221@9.[151,1436,632,1536]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURES 1922. Chimerothalassius riparius sp. nov. 19. Head of male holotype (CNC1155797), anterior view. 20. Head of male holotype, showing left antenna in medial view. 21. Abdomen of female paratype, lateral view. 22. Right wing of male holotype, dorsal view. Abbreviations: cerc—cercus; M1, M4—medial veins; plp—palpus; R1, R2+3, R 4+5 —radial veins; tg—tergite." captionText-1="FIGURES 2325. Hypopygium of Chimerothalassius riparius sp. nov., holotype (CNC1155797). 23. Hypopygium, left lateral view. 24. Hypopygium, right lateral view. 25. Cerci and hypoproct, dorsal view. Abbreviations: cerc—cercus; d sur—dorsal lobe of surstylus; epand—epandrium; hypd—hypandrium; hyprct—hypoproct; (L)—left; pgt lb—postgonite lobe; ph—phallus; (R)—right; v epand proc—ventral epandrial process; v sur—ventral lobe of surstylus." figureDoi-0="http://doi.org/10.5281/zenodo.7103634" figureDoi-1="http://doi.org/10.5281/zenodo.7103638" httpUri-0="https://zenodo.org/record/7103634/files/figure.png" httpUri-1="https://zenodo.org/record/7103638/files/figure.png" pageId="7" pageNumber="528">1925</figureCitation>
)
</paragraph>
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urn:lsid:zoobank.org:act:
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</paragraph>
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<subSubSection id="C33E6525C817200EEBC3E35F9BF0AA9B" pageId="7" pageNumber="528" type="materials_examined">
<paragraph id="8B9B36AEC817200EEBC3E35F9BF0AA9B" blockId="7.[151,1437,289,2038]" pageId="7" pageNumber="528">
<emphasis id="B950EABCC817200EEBC3E35F9E01AA08" bold="true" box="[151,329,289,316]" pageId="7" pageNumber="528">Type material.</emphasis>
<materialsCitation id="3B4C3CF3C817200EEA07E35C9C9CAAB7" ID-GBIF-Occurrence="3912959316" collectionCode="NEW, CALEDONIA, R, MNHN" collectorName="B. J. Sinclair" latitude="-22.097223" longLatPrecision="21" longitude="166.63805" pageId="7" pageNumber="528" specimenCode="CNC/ 1155797" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B950EABCC817200EEA07E35C9EBAAA08" bold="true" box="[339,498,289,315]" pageId="7" pageNumber="528">
<typeStatus id="549F880CC817200EEA07E35C9EBAAA08" box="[339,498,289,315]" pageId="7" pageNumber="528" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
<specimenCount id="9D22FD27C817200EEAA8E35F9D59AA0F" box="[508,529,290,316]" pageId="7" pageNumber="528" type="male"></specimenCount>
labelled: “
<collectionCode id="ED35AE6BC817200EE9C5E35F9D9BAA0F" box="[657,723,290,316]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="7" pageNumber="528">NEW</collectionCode>
<collectionCode id="ED35AE6BC817200EE989E35F9CCDAA0F" box="[733,901,290,316]" pageId="7" pageNumber="528">CALEDONIA</collectionCode>
: Parc/ Rivève[
<emphasis id="B950EABCC817200EEF64E35E9B07AA0F" box="[1072,1103,291,316]" italics="true" pageId="7" pageNumber="528">sic</emphasis>
] Bleue;
<collectionCode id="ED35AE6BC817200EEFE3E35F9B82AA0F" box="[1207,1226,290,316]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="528">R</collectionCode>
.Bleue/ nr. Refuge [
<emphasis id="B950EABCC817200EEBF5E33A9FF3AA53" box="[161,187,327,352]" italics="true" pageId="7" pageNumber="528">ca</emphasis>
<geoCoordinate id="EE105069C817200EEB97E33A9E0AAA51" box="[195,322,327,354]" degrees="22" direction="south" minutes="05" orientation="latitude" pageId="7" pageNumber="528" precision="15" seconds="50" value="-22.097223">22°05ʹ50ʺS</geoCoordinate>
<geoCoordinate id="EE105069C817200EEA1EE33A9E96AA51" box="[330,478,327,354]" degrees="166" direction="east" minutes="38" orientation="longitude" pageId="7" pageNumber="528" precision="15" seconds="17" value="166.63805">166°38ʹ17ʺE</geoCoordinate>
],
<date id="FF9A106EC817200EEAA5E33B9DE5AA52" box="[497,685,326,353]" pageId="7" pageNumber="528" value="1995-07-20" valueMax="1995-07-21" valueMin="1995-07-20">2021.vii./ 1995</date>
;
<collectorName id="26D15378C817200EE9EEE33B9C0BAA53" box="[698,835,326,352]" pageId="7" pageNumber="528">B.J. Sinclair</collectorName>
/ ex. yellow pans”; “
<specimenCode id="DB829ED5C817200EEF7FE33B9B96AA52" box="[1067,1246,326,353]" collectionCode="CNC" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="7" pageNumber="528">CNC/ 1155797</specimenCode>
”; “
<typeStatus id="549F880CC817200EEFAAE33B9ADCAA53" box="[1278,1428,326,352]" pageId="7" pageNumber="528" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="4C244D2DC817200EEBC3E3179E85AAB7" authorityName="Brooks &amp; Cumming" authorityYear="2022" box="[151,461,362,388]" class="Insecta" family="Dolichopodidae" genus="Chimerothalassius" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="528" phylum="Arthropoda" rank="species" species="riparius">
<emphasis id="B950EABCC817200EEBC3E3179E85AAB7" box="[151,461,362,388]" italics="true" pageId="7" pageNumber="528">Chimerothalassius riparius</emphasis>
</taxonomicName>
/ Brooks &amp; Cumming [red label]” (
<collectionCode id="ED35AE6BC817200EE824E3179C84AAB7" box="[880,972,362,388]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="7" pageNumber="528">MNHN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4C3CF3C817200EE8B6E3149D5DAA9B" ID-GBIF-Occurrence="3912959312" collectionCode="CNC" collectorName="B. J. Sinclair" country="New Caledonia" latitude="-22.097223" longLatPrecision="21" longitude="166.63805" pageId="7" pageNumber="528" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<emphasis id="B950EABCC817200EE8B6E3149AD4AAB0" bold="true" box="[994,1436,361,388]" pageId="7" pageNumber="528">
<typeStatus id="549F880CC817200EE8B6E3149BC2AAB0" box="[994,1162,361,387]" pageId="7" pageNumber="528" type="paratype">PARATYPES</typeStatus>
:
<collectingCountry id="F333763EC817200EEFCEE3179AD0AAB0" box="[1178,1432,361,388]" name="New Caledonia" pageId="7" pageNumber="528">NEW CALEDONIA</collectingCountry>
:
</emphasis>
same data as holotype (
<specimenCount id="9D22FD27C817200EEAF5E3F39E8AAA9B" box="[417,450,398,424]" pageId="7" pageNumber="528" type="female">1♀</specimenCount>
,
<collectionCode id="ED35AE6BC817200EEA9BE3F39D45AA9B" box="[463,525,398,424]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="7" pageNumber="528">CNC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4C3CF3C817200EE975E3F39BFCAA9B" ID-GBIF-Occurrence="3912959318" box="[545,1204,398,425]" collectionCode="MNHN" collectorName="B. J. Sinclair" country="New Caledonia" latitude="-22.097223" longLatPrecision="21" longitude="166.63805" pageId="7" pageNumber="528" specimenCode="CNC1155799" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
same data as holotype except,
<specimenCode id="DB829ED5C817200EE821E3F39B58AA9A" box="[885,1040,398,425]" collectionCode="CNC" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="7" pageNumber="528">CNC1155799</specimenCode>
(
<specimenCount id="9D22FD27C817200EEF77E3F39B0CAA9B" box="[1059,1092,398,424]" pageId="7" pageNumber="528" type="female">1♀</specimenCount>
,
<collectionCode id="ED35AE6BC817200EEF05E3F39BE4AA9B" box="[1105,1196,398,424]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="7" pageNumber="528">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33E6525C817200EEB93E3CC9BB3A9B3" pageId="7" pageNumber="528" type="diagnosis">
<paragraph id="8B9B36AEC817200EEB93E3CC9BB3A9B3" blockId="7.[151,1437,289,2038]" pageId="7" pageNumber="528">
<emphasis id="B950EABCC817200EEB93E3CC9E0AAAF8" bold="true" box="[199,322,433,459]" pageId="7" pageNumber="528">Diagnosis.</emphasis>
<taxonomicName id="4C244D2DC817200EEA1DE3CF9D34AAFF" authority="Brooks &amp; Cumming, 2022" authorityName="Brooks &amp; Cumming" authorityYear="2022" box="[329,636,434,460]" class="Insecta" family="Dolichopodidae" genus="Chimerothalassius" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="528" phylum="Arthropoda" rank="species" species="riparius" status="sp. nov.">
<emphasis id="B950EABCC817200EEA1DE3CF9D34AAFF" box="[329,636,434,460]" italics="true" pageId="7" pageNumber="528">Chimerothalassius riparius</emphasis>
</taxonomicName>
<emphasis id="B950EABCC817200EE9D0E3CF9D96AAFF" bold="true" box="[644,734,434,460]" pageId="7" pageNumber="528">
<taxonomicNameLabel id="A26357C7C817200EE9D0E3CF9D96AAFF" box="[644,734,434,460]" pageId="7" pageNumber="528" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is distinguished from the other known Australasian species of the genus by its brown, capitate palpus (
<figureCitation id="131F2A2BC817200EE933E3AB9DF5AAC3" box="[615,701,470,496]" captionStart="FIGURES 1922" captionStartId="8.[151,264,1937,1961]" captionTargetBox="[219,1369,1001,1909]" captionTargetId="figure-389@8.[219,1369,993,1914]" captionTargetPageId="8" captionText="FIGURES 1922. Chimerothalassius riparius sp. nov. 19. Head of male holotype (CNC1155797), anterior view. 20. Head of male holotype, showing left antenna in medial view. 21. Abdomen of female paratype, lateral view. 22. Right wing of male holotype, dorsal view. Abbreviations: cerc—cercus; M1, M4—medial veins; plp—palpus; R1, R2+3, R 4+5 —radial veins; tg—tergite." figureDoi="http://doi.org/10.5281/zenodo.7103634" httpUri="https://zenodo.org/record/7103634/files/figure.png" pageId="7" pageNumber="528">Fig. 19</figureCitation>
). Males are further distinguished by several hypopygial features (
<figureCitation id="131F2A2BC817200EEBCBE3879E68A926" box="[159,288,506,533]" captionStart="FIGURES 2325" captionStartId="9.[151,264,1559,1583]" captionTargetBox="[151,1426,641,1526]" captionTargetId="figure-221@9.[151,1436,632,1536]" captionTargetPageId="9" captionText="FIGURES 2325. Hypopygium of Chimerothalassius riparius sp. nov., holotype (CNC1155797). 23. Hypopygium, left lateral view. 24. Hypopygium, right lateral view. 25. Cerci and hypoproct, dorsal view. Abbreviations: cerc—cercus; d sur—dorsal lobe of surstylus; epand—epandrium; hypd—hypandrium; hyprct—hypoproct; (L)—left; pgt lb—postgonite lobe; ph—phallus; (R)—right; v epand proc—ventral epandrial process; v sur—ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.7103638" httpUri="https://zenodo.org/record/7103638/files/figure.png" pageId="7" pageNumber="528">Figs 2325</figureCitation>
) including: left ventral epandrial process with tip weakly bent; left ventral surstylus long and narrow with short setae; right ventral epandrial process strongly upcurved; phallus with tip not expanded and lacking process near midlength; hypoproct bifid with narrow dorsal lobe and longer broader ventral lobe. Females are further characterized by their terminalia with acanthophorous spines and pointed, upturned cercus (
<figureCitation id="131F2A2BC817200EEFCEE01B9BA6A9B3" box="[1178,1262,614,640]" captionStart="FIGURES 1922" captionStartId="8.[151,264,1937,1961]" captionTargetBox="[219,1369,1001,1909]" captionTargetId="figure-389@8.[219,1369,993,1914]" captionTargetPageId="8" captionText="FIGURES 1922. Chimerothalassius riparius sp. nov. 19. Head of male holotype (CNC1155797), anterior view. 20. Head of male holotype, showing left antenna in medial view. 21. Abdomen of female paratype, lateral view. 22. Right wing of male holotype, dorsal view. Abbreviations: cerc—cercus; M1, M4—medial veins; plp—palpus; R1, R2+3, R 4+5 —radial veins; tg—tergite." figureDoi="http://doi.org/10.5281/zenodo.7103634" httpUri="https://zenodo.org/record/7103634/files/figure.png" pageId="7" pageNumber="528">Fig. 21</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33E6525C8172000EB93E0F49B77AA73" lastPageId="9" lastPageNumber="530" pageId="7" pageNumber="528" type="description">
<paragraph id="8B9B36AEC8172001EB93E0F49EFCA8FA" blockId="7.[151,1437,289,2038]" lastBlockId="8.[151,1437,150,969]" lastPageId="8" lastPageNumber="529" pageId="7" pageNumber="528">
<emphasis id="B950EABCC817200EEB93E0F49ED5A997" bold="true" box="[199,413,649,676]" pageId="7" pageNumber="528">Description. Male</emphasis>
(
<figureCitation id="131F2A2BC817200EEAF9E0F79DCFA996" box="[429,647,650,677]" captionStart-0="FIGURES 1922" captionStart-1="FIGURES 2325" captionStartId-0="8.[151,264,1937,1961]" captionStartId-1="9.[151,264,1559,1583]" captionTargetBox-0="[219,1369,1001,1909]" captionTargetBox-1="[151,1426,641,1526]" captionTargetId-0="figure-389@8.[219,1369,993,1914]" captionTargetId-1="figure-221@9.[151,1436,632,1536]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURES 1922. Chimerothalassius riparius sp. nov. 19. Head of male holotype (CNC1155797), anterior view. 20. Head of male holotype, showing left antenna in medial view. 21. Abdomen of female paratype, lateral view. 22. Right wing of male holotype, dorsal view. Abbreviations: cerc—cercus; M1, M4—medial veins; plp—palpus; R1, R2+3, R 4+5 —radial veins; tg—tergite." captionText-1="FIGURES 2325. Hypopygium of Chimerothalassius riparius sp. nov., holotype (CNC1155797). 23. Hypopygium, left lateral view. 24. Hypopygium, right lateral view. 25. Cerci and hypoproct, dorsal view. Abbreviations: cerc—cercus; d sur—dorsal lobe of surstylus; epand—epandrium; hypd—hypandrium; hyprct—hypoproct; (L)—left; pgt lb—postgonite lobe; ph—phallus; (R)—right; v epand proc—ventral epandrial process; v sur—ventral lobe of surstylus." figureDoi-0="http://doi.org/10.5281/zenodo.7103634" figureDoi-1="http://doi.org/10.5281/zenodo.7103638" httpUri-0="https://zenodo.org/record/7103634/files/figure.png" httpUri-1="https://zenodo.org/record/7103638/files/figure.png" pageId="7" pageNumber="528">Figs 19, 20, 2225</figureCitation>
): Wing length 1.10 mm.
<emphasis id="B950EABCC817200EE8F4E0F79CA8A997" bold="true" box="[928,992,650,676]" pageId="7" pageNumber="528">Head</emphasis>
(
<figureCitation id="131F2A2BC817200EE8A4E0F79B3DA997" box="[1008,1141,650,676]" captionStart="FIGURES 1922" captionStartId="8.[151,264,1937,1961]" captionTargetBox="[219,1369,1001,1909]" captionTargetId="figure-389@8.[219,1369,993,1914]" captionTargetPageId="8" captionText="FIGURES 1922. Chimerothalassius riparius sp. nov. 19. Head of male holotype (CNC1155797), anterior view. 20. Head of male holotype, showing left antenna in medial view. 21. Abdomen of female paratype, lateral view. 22. Right wing of male holotype, dorsal view. Abbreviations: cerc—cercus; M1, M4—medial veins; plp—palpus; R1, R2+3, R 4+5 —radial veins; tg—tergite." figureDoi="http://doi.org/10.5281/zenodo.7103634" httpUri="https://zenodo.org/record/7103634/files/figure.png" pageId="7" pageNumber="528">Figs 19, 20</figureCitation>
): Dark grey pruinose with green and bronze reflections dorsally; broader than thorax in dorsal view; ovoid in lateral view (higher than broad); about as broad as high in anterior view; larger setae black. Ocellar triangle conspicuous. Occiput concave on upper median part. Eyes covered with short ommatrichia; medial edge of eye with distinct emargination adjacent to antenna; ommatidia progressively smaller anterodorsally. Frons 2.3X broader than high, widening above. Face narrow, with eyes nearly contiguous, concolorous with rest of head. Clypeus not separated from face, small and triangular, weakly produced medially, brownish medially, grey laterally. Setae of head well differentiated: 1 pair of weakly inclinate fronto-orbitals about midway between base of antennae and ocellar triangle; 1 pair of lateroclinate anterior ocellars; 1 pair of small posterior ocellars; 1 pair of weakly inclinate inner verticals (sometimes referred to as postocellars); 2 pairs of lateroclinate outer verticals; postoculars short fine and pale, upper setae uniserial, lower setae scattered, lowermost seta longer. Antenna entirely dark brown, inserted slightly above middle of head in profile; scape short, funnel-shaped; pedicel subequal in length to scape, spheroidal with subapical circlet of setulae; postpedicel 1.1X longer than wide, bulb-shaped with very short narrow tip, clothed in fine setulae; arista-like stylus 3X length of postpedicel, with minute hairs. Palpus brown, narrow basally, abruptly capitate apically, with short hairs. Proboscis dark brown, short. Gena narrow.
<emphasis id="B950EABCC817200EE9E8E6FF9C56AFAF" bold="true" box="[700,798,1154,1180]" pageId="7" pageNumber="528">Thorax:</emphasis>
Dark grey pruinose with green and bronze reflections especially dorsally, setae black. Mesoscutum moderately arched, prescutellar depression present. Proepisternum with minute seta. Postpronotal lobe with 2 minute setae. Mesonotum longer than wide. Acrostichal setae absent; other thoracic setae well differentiated, each side of mesonotum with: 4 dorsocentrals, 1 presutural supra-alar (posthumeral), 1 postsutural supra-alar, 2 notopleurals, 1 postalar. Scutellum broadly subtriangular with 1 long, strong, dorsally projected seta per side. Mesopleuron bare. Halter pale brown.
<emphasis id="B950EABCC817200EE810E7489CCDAE7C" bold="true" box="[836,901,1333,1359]" pageId="7" pageNumber="528">Legs:</emphasis>
Evenly brown, except apex of fore coxa yellow; with mostly short setae; tarsomeres 14 of all legs progressively shorter apically with tarsomere 5 slightly longer than 4; tarsomere 5 of all legs with very weakly developed dorsomedial process; tarsal claws, pulvilli and empodium normally developed on all legs.
<emphasis id="B950EABCC817200EE951E7DD9D15AE89" box="[517,605,1440,1466]" italics="true" pageId="7" pageNumber="528">Foreleg</emphasis>
: Coxa apparently with short, sparse pale setae on anterior surface, apical margin with setae longer; femur, tibia and tarsus subequal in length; tarsomere 1 subequal to combined length of tarsomeres 24.
<emphasis id="B950EABCC817200EEA06E79A9EE9AD32" box="[338,417,1511,1537]" italics="true" pageId="7" pageNumber="528">Midleg</emphasis>
: Coxa with a few pale setae; femur subequal in length to tibia, with row of anteroventral setae (subequal in length to femur width); tarsus slightly longer than tibia; tarsomere 1 partially obscured from view.
<emphasis id="B950EABCC817200EEBC3E4539FB9AD7B" box="[151,241,1582,1608]" italics="true" pageId="7" pageNumber="528">Hindleg</emphasis>
: Coxa with 2 pale setae on lateral surface; femur, tibia and tarsus subequal in length; tarsomere 1 subequal to combined length of tarsomeres 23.
<emphasis id="B950EABCC817200EE904E42F9DD9AD5F" bold="true" box="[592,657,1618,1644]" pageId="7" pageNumber="528">Wing</emphasis>
(
<figureCitation id="131F2A2BC817200EE9F5E42F9DBEAD5F" box="[673,758,1618,1644]" captionStart="FIGURES 1922" captionStartId="8.[151,264,1937,1961]" captionTargetBox="[219,1369,1001,1909]" captionTargetId="figure-389@8.[219,1369,993,1914]" captionTargetPageId="8" captionText="FIGURES 1922. Chimerothalassius riparius sp. nov. 19. Head of male holotype (CNC1155797), anterior view. 20. Head of male holotype, showing left antenna in medial view. 21. Abdomen of female paratype, lateral view. 22. Right wing of male holotype, dorsal view. Abbreviations: cerc—cercus; M1, M4—medial veins; plp—palpus; R1, R2+3, R 4+5 —radial veins; tg—tergite." figureDoi="http://doi.org/10.5281/zenodo.7103634" httpUri="https://zenodo.org/record/7103634/files/figure.png" pageId="7" pageNumber="528">Fig. 22</figureCitation>
): With brownish tinge, veins dark brown, about 2.5X longer than wide. Pterostigma absent, membrane entirely covered with minute microtrichia, alula absent.
<collectingRegion id="49E0F84CC817200EEFA8E40B9A74ADA3" box="[1276,1340,1654,1680]" country="New Zealand" name="West Coast" pageId="7" pageNumber="528">Costa</collectingRegion>
circumambient. Extreme anterior base of costa apparently with 1 strong anterodorsal seta (broken in unique
<typeStatus id="549F880CC817200EEE65E4E79ADDAD87" box="[1329,1429,1690,1716]" pageId="7" pageNumber="528" type="holotype">holotype</typeStatus>
) and at least 1 shorter proximal seta. Anterior section of costa (between base and R
<subScript id="17A034EBC817200EEF78E4AD9B0DADED" attach="left" box="[1068,1093,1744,1758]" fontSize="6" pageId="7" pageNumber="528">2+3</subScript>
) with double row of relatively widely spaced spine-like setae. Posterior section of costa (beyond R
<subScript id="17A034EBC817200EE8CBE4899CF0AC31" attach="left" box="[927,952,1780,1794]" fontSize="6" pageId="7" pageNumber="528">2+3</subScript>
) with setae finer, longer and more closely spaced. Radial and medial veins M
<subScript id="17A034EBC817200EE978E56A9D7CAC16" attach="left" box="[556,564,1815,1829]" fontSize="6" pageId="7" pageNumber="528">1</subScript>
and M
<subScript id="17A034EBC817200EE9DCE56A9DD8AC16" attach="left" box="[648,656,1815,1829]" fontSize="6" pageId="7" pageNumber="528">4</subScript>
complete and reaching wing margin, M
<subScript id="17A034EBC817200EEF34E56A9B20AC16" attach="left" box="[1120,1128,1815,1829]" fontSize="6" pageId="7" pageNumber="528">2</subScript>
and CuA+CuP (anal vein) absent, Sc faint. R
<subScript id="17A034EBC817200EEA3CE5469E38AC7A" attach="left" box="[360,368,1851,1865]" fontSize="6" pageId="7" pageNumber="528">1</subScript>
short, terminating near basal 1/3 of wing. Base of Rs originating opposite humeral crossvein. R
<subScript id="17A034EBC817200EEBFEE5229F8BAC5E" attach="left" box="[170,195,1887,1901]" fontSize="6" pageId="7" pageNumber="528">2+3</subScript>
subparallel with R
<quantity id="4CDC9B4BC817200EEACEE5229E88AC55" box="[410,448,1868,1901]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="7" pageNumber="528" unit="in" value="1.0">
<subScript id="17A034EBC817200EEACEE5229EEAAC5E" attach="left" box="[410,418,1887,1901]" fontSize="6" pageId="7" pageNumber="528">1</subScript>
in
</quantity>
basal part, subparallel with R
<subScript id="17A034EBC817200EE846E5229C64AC5E" attach="left" box="[786,812,1887,1901]" fontSize="6" pageId="7" pageNumber="528">
4+
<quantity id="4CDC9B4BC817200EE870E5229C64AC5E" box="[804,812,1887,1901]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="7" pageNumber="528" unit="in" value="5.0">5</quantity>
</subScript>
in distal part. R
<subScript id="17A034EBC817200EE8B6E5229CB4AC5E" attach="left" box="[994,1020,1887,1901]" fontSize="6" pageId="7" pageNumber="528">4+5</subScript>
weakly sinuous. M
<subScript id="17A034EBC817200EEF8DE5229BA9AC5E" attach="left" box="[1241,1249,1887,1901]" fontSize="6" pageId="7" pageNumber="528">1</subScript>
straight in basal half, sinuous in distal half and curving posteriorly to costa apically. M
<subScript id="17A034EBC817200EE8F7E5FE9CE3ACA2" attach="left" box="[931,939,1923,1937]" fontSize="6" pageId="7" pageNumber="528">4</subScript>
mostly straight with posterior curve to costa apically. CuA rounded. Short r-m crossvein present in basal portion of wing, distal to base of R
<subScript id="17A034EBC817200EEFEFE5DB9B9DAC87" attach="left" box="[1211,1237,1958,1972]" fontSize="6" pageId="7" pageNumber="528">4+5</subScript>
. Crossvein bm-m incomplete. Cell dm, base of M
<subScript id="17A034EBC817200EEAADE5B79D49ACEB" attach="left" box="[505,513,1994,2008]" fontSize="6" pageId="7" pageNumber="528">2</subScript>
and dm-m crossvein absent. Cells br, bm and cua in basal fourth of wing. Cell cua closed, ovoid. Anal lobe not developed. Calypter with fine setae.
<emphasis id="B950EABCC817200EE820E5A69CBAACC6" bold="true" box="[884,1010,2011,2037]" pageId="7" pageNumber="528">Abdomen:</emphasis>
Brown with small dark setae (weaker and pale on sternites); sternite 6 mainly bare, segment 7 bare. Segments 57 narrowed and laterally compressed to form cavity on right side for hypopygium. Sternite 5 lacking pregenitalic process. Sternite 8 ovoid, with short setae; tergite 8 indistinct.
<emphasis id="B950EABCC8182001EA21E2A29D4CABCB" box="[373,516,223,248]" italics="true" pageId="8" pageNumber="529">Hypopygium</emphasis>
(
<figureCitation id="131F2A2BC8182001E941E2A29DD3ABCA" box="[533,667,222,249]" captionStart="FIGURES 2325" captionStartId="9.[151,264,1559,1583]" captionTargetBox="[151,1426,641,1526]" captionTargetId="figure-221@9.[151,1436,632,1536]" captionTargetPageId="9" captionText="FIGURES 2325. Hypopygium of Chimerothalassius riparius sp. nov., holotype (CNC1155797). 23. Hypopygium, left lateral view. 24. Hypopygium, right lateral view. 25. Cerci and hypoproct, dorsal view. Abbreviations: cerc—cercus; d sur—dorsal lobe of surstylus; epand—epandrium; hypd—hypandrium; hyprct—hypoproct; (L)—left; pgt lb—postgonite lobe; ph—phallus; (R)—right; v epand proc—ventral epandrial process; v sur—ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.7103638" httpUri="https://zenodo.org/record/7103638/files/figure.png" pageId="8" pageNumber="529">Figs 2325</figureCitation>
): Concolorous with pregenitalic abdominal sclerites; lateroflexed to right; inverted with posterior end directed anteriorly; small and compact, about 1/4 length of abdomen; asymmetrical; foramen not formed. Epandrium divided into left and right lamellae. Left epandrial lamella (
<figureCitation id="131F2A2BC8182001EF88E35A9A67AA72" box="[1244,1327,295,321]" captionStart="FIGURES 2325" captionStartId="9.[151,264,1559,1583]" captionTargetBox="[151,1426,641,1526]" captionTargetId="figure-221@9.[151,1436,632,1536]" captionTargetPageId="9" captionText="FIGURES 2325. Hypopygium of Chimerothalassius riparius sp. nov., holotype (CNC1155797). 23. Hypopygium, left lateral view. 24. Hypopygium, right lateral view. 25. Cerci and hypoproct, dorsal view. Abbreviations: cerc—cercus; d sur—dorsal lobe of surstylus; epand—epandrium; hypd—hypandrium; hyprct—hypoproct; (L)—left; pgt lb—postgonite lobe; ph—phallus; (R)—right; v epand proc—ventral epandrial process; v sur—ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.7103638" httpUri="https://zenodo.org/record/7103638/files/figure.png" pageId="8" pageNumber="529">Fig. 23</figureCitation>
) partially overlapping left side of hypandrium, posterior margin trifurcate, ventrally fused with hypandrium but margin distinct; ventral epandrial process apparently not articulated at base, long and slender, projected dorsally, tip with weak weakly ventral bend. Left surstylus bilobed, dorsal and ventral lobes separated by U-shaped cleft through which left postgonite lobe protrudes. Dorsal lobe of left surstylus with short apical seta, long preapical dorsal seta, and short dorsal seta, lacking prensiseta. Ventral lobe of left surstylus longer than dorsal lobe, narrow with relatively short setae ventrally. Right epandrial lamella (
<figureCitation id="131F2A2BC8182001E9F8E3829C49A92A" box="[684,769,511,537]" captionStart="FIGURES 2325" captionStartId="9.[151,264,1559,1583]" captionTargetBox="[151,1426,641,1526]" captionTargetId="figure-221@9.[151,1436,632,1536]" captionTargetPageId="9" captionText="FIGURES 2325. Hypopygium of Chimerothalassius riparius sp. nov., holotype (CNC1155797). 23. Hypopygium, left lateral view. 24. Hypopygium, right lateral view. 25. Cerci and hypoproct, dorsal view. Abbreviations: cerc—cercus; d sur—dorsal lobe of surstylus; epand—epandrium; hypd—hypandrium; hyprct—hypoproct; (L)—left; pgt lb—postgonite lobe; ph—phallus; (R)—right; v epand proc—ventral epandrial process; v sur—ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.7103638" httpUri="https://zenodo.org/record/7103638/files/figure.png" pageId="8" pageNumber="529">Fig. 24</figureCitation>
) partially overlapping right side of hypandrium, ventrally fused with hypandrium but margin distinct, slightly longer than hypandrium, with prominent seta near middle of posterior margin; ventral epandrial process present, narrow and strongly upcurved. Right surstylus bilobed, dorsal and ventral lobes separated by U-shaped cleft through which right postgonite lobe protrudes. Dorsal lobe of right surstylus broad with short apical seta, long preapical dorsal seta and short dorsal seta, lacking prensiseta. Ventral lobe of right surstylus about 3X longer than dorsal lobe, with 1 ventral seta near middle and 1 preapical medial seta. Hypandrium bowl-shaped, slightly shorter than epandrium in lateral view. Left postgonite lobe large, deeply bilobate, with thumb-like dorsomedial lobe and longer ventral lobe (
<figureCitation id="131F2A2BC8182001E8FEE0869B48A826" box="[938,1024,763,789]" captionStart="FIGURES 2325" captionStartId="9.[151,264,1559,1583]" captionTargetBox="[151,1426,641,1526]" captionTargetId="figure-221@9.[151,1436,632,1536]" captionTargetPageId="9" captionText="FIGURES 2325. Hypopygium of Chimerothalassius riparius sp. nov., holotype (CNC1155797). 23. Hypopygium, left lateral view. 24. Hypopygium, right lateral view. 25. Cerci and hypoproct, dorsal view. Abbreviations: cerc—cercus; d sur—dorsal lobe of surstylus; epand—epandrium; hypd—hypandrium; hyprct—hypoproct; (L)—left; pgt lb—postgonite lobe; ph—phallus; (R)—right; v epand proc—ventral epandrial process; v sur—ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.7103638" httpUri="https://zenodo.org/record/7103638/files/figure.png" pageId="8" pageNumber="529">Fig. 23</figureCitation>
). Right postgonite lobe shorter than left lobe, apically bilobate with ventral lobe about as long as dorsomedial lobe (
<figureCitation id="131F2A2BC8182001EF41E1629B22A80A" box="[1045,1130,799,825]" captionStart="FIGURES 2325" captionStartId="9.[151,264,1559,1583]" captionTargetBox="[151,1426,641,1526]" captionTargetId="figure-221@9.[151,1436,632,1536]" captionTargetPageId="9" captionText="FIGURES 2325. Hypopygium of Chimerothalassius riparius sp. nov., holotype (CNC1155797). 23. Hypopygium, left lateral view. 24. Hypopygium, right lateral view. 25. Cerci and hypoproct, dorsal view. Abbreviations: cerc—cercus; d sur—dorsal lobe of surstylus; epand—epandrium; hypd—hypandrium; hyprct—hypoproct; (L)—left; pgt lb—postgonite lobe; ph—phallus; (R)—right; v epand proc—ventral epandrial process; v sur—ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.7103638" httpUri="https://zenodo.org/record/7103638/files/figure.png" pageId="8" pageNumber="529">Fig. 24</figureCitation>
). Phallus tubular, J-shaped, projected dorsally, with flange-like ventral projection near base, with tip not expanded. Ejaculatory apodeme keellike. Hypoproct bifid with narrow dorsal lobe and longer broader ventral lobe, left and right sides symmetrical (
<figureCitation id="131F2A2BC8182001EE3BE11A9FFFA896" captionStart="FIGURES 2325" captionStartId="9.[151,264,1559,1583]" captionTargetBox="[151,1426,641,1526]" captionTargetId="figure-221@9.[151,1436,632,1536]" captionTargetPageId="9" captionText="FIGURES 2325. Hypopygium of Chimerothalassius riparius sp. nov., holotype (CNC1155797). 23. Hypopygium, left lateral view. 24. Hypopygium, right lateral view. 25. Cerci and hypoproct, dorsal view. Abbreviations: cerc—cercus; d sur—dorsal lobe of surstylus; epand—epandrium; hypd—hypandrium; hyprct—hypoproct; (L)—left; pgt lb—postgonite lobe; ph—phallus; (R)—right; v epand proc—ventral epandrial process; v sur—ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.7103638" httpUri="https://zenodo.org/record/7103638/files/figure.png" pageId="8" pageNumber="529">Fig. 25</figureCitation>
). Cercus subtriangular in dorsal view (
<figureCitation id="131F2A2BC8182001E938E1F69D88A896" box="[620,704,906,933]" captionStart="FIGURES 2325" captionStartId="9.[151,264,1559,1583]" captionTargetBox="[151,1426,641,1526]" captionTargetId="figure-221@9.[151,1436,632,1536]" captionTargetPageId="9" captionText="FIGURES 2325. Hypopygium of Chimerothalassius riparius sp. nov., holotype (CNC1155797). 23. Hypopygium, left lateral view. 24. Hypopygium, right lateral view. 25. Cerci and hypoproct, dorsal view. Abbreviations: cerc—cercus; d sur—dorsal lobe of surstylus; epand—epandrium; hypd—hypandrium; hyprct—hypoproct; (L)—left; pgt lb—postgonite lobe; ph—phallus; (R)—right; v epand proc—ventral epandrial process; v sur—ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.7103638" httpUri="https://zenodo.org/record/7103638/files/figure.png" pageId="8" pageNumber="529">Fig. 25</figureCitation>
), with small preapical seta and 2 prominent dorsal setae, left and right cercus symmetrical.
</paragraph>
<caption id="DF5B6626C8182001EBC3E5EC9AC1ACC2" ID-DOI="http://doi.org/10.5281/zenodo.7103634" ID-Zenodo-Dep="7103634" httpUri="https://zenodo.org/record/7103634/files/figure.png" pageId="8" pageNumber="529" startId="8.[151,264,1937,1961]" targetBox="[219,1369,1001,1909]" targetPageId="8">
<paragraph id="8B9B36AEC8182001EBC3E5EC9AC1ACC2" blockId="8.[151,1436,1937,2040]" pageId="8" pageNumber="529">
<emphasis id="B950EABCC8182001EBC3E5EC9E1DAC9A" bold="true" box="[151,341,1937,1961]" pageId="8" pageNumber="529">FIGURES 1922.</emphasis>
<taxonomicName id="4C244D2DC8182001EA0FE5EC9D39AC9A" authority="Brooks &amp; Cumming, 2022" authorityName="Brooks &amp; Cumming" authorityYear="2022" box="[347,625,1937,1961]" class="Insecta" family="Dolichopodidae" genus="Chimerothalassius" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" species="riparius" status="sp. nov.">
<emphasis id="B950EABCC8182001EA0FE5EC9D39AC9A" box="[347,625,1937,1961]" italics="true" pageId="8" pageNumber="529">Chimerothalassius riparius</emphasis>
</taxonomicName>
<emphasis id="B950EABCC8182001E92CE5EC9DB8AC9A" bold="true" box="[632,752,1937,1961]" pageId="8" pageNumber="529">
<taxonomicNameLabel id="A26357C7C8182001E92CE5EC9D81AC9A" box="[632,713,1937,1961]" pageId="8" pageNumber="529" rank="species">sp. nov.</taxonomicNameLabel>
19.
</emphasis>
Head of male holotype (CNC1155797), anterior view.
<emphasis id="B950EABCC8182001EE70E5EC9A0CAC9A" bold="true" box="[1316,1348,1937,1961]" pageId="8" pageNumber="529">20.</emphasis>
Head of male holotype, showing left antenna in medial view.
<emphasis id="B950EABCC8182001E9F3E5C89D8FACFE" bold="true" box="[679,711,1973,1997]" pageId="8" pageNumber="529">21.</emphasis>
Abdomen of female paratype, lateral view.
<emphasis id="B950EABCC8182001EF2FE5C89BD3ACFE" bold="true" box="[1147,1179,1973,1997]" pageId="8" pageNumber="529">22.</emphasis>
Right wing of male holotype, dorsal view. Abbreviations: cerc—cercus; M
<subScript id="17A034EBC8182001E9C0E5949DD3ACC4" attach="right" box="[660,667,2025,2039]" fontSize="6" pageId="8" pageNumber="529">1</subScript>
, M
<subScript id="17A034EBC8182001E9EAE5949D8DACC4" attach="left" box="[702,709,2025,2039]" fontSize="6" pageId="8" pageNumber="529">4</subScript>
—medial veins; plp—palpus; R
<subScript id="17A034EBC8182001EF53E5949B46ACC4" attach="left" box="[1031,1038,2025,2039]" fontSize="6" pageId="8" pageNumber="529">1</subScript>
, R
<subScript id="17A034EBC8182001EF78E5949B0BACC4" attach="left" box="[1068,1091,2025,2039]" fontSize="6" pageId="8" pageNumber="529">2+3</subScript>
, R
<subScript id="17A034EBC8182001EF35E5979B30ACCB" attach="left" box="[1121,1144,2026,2040]" fontSize="6" pageId="8" pageNumber="529">4+5</subScript>
—radial veins; tg—tergite.
</paragraph>
</caption>
<paragraph id="8B9B36AEC8192000EB93E2EB9B77AA73" blockId="9.[151,1437,150,609]" pageId="9" pageNumber="530">
<emphasis id="B950EABCC8192000EB93E2EB9E60AB83" bold="true" box="[199,296,150,176]" pageId="9" pageNumber="530">Female:</emphasis>
Body length 1.30 mm, wing length 1.40 mm. Similar to male except as follows:
<emphasis id="B950EABCC8192000EFE2E2EB9A63AB83" bold="true" box="[1206,1323,150,176]" pageId="9" pageNumber="530">Abdomen</emphasis>
(
<figureCitation id="131F2A2BC8192000EE6FE2EB9AC7AB83" box="[1339,1423,150,176]" captionStart="FIGURES 1922" captionStartId="8.[151,264,1937,1961]" captionTargetBox="[219,1369,1001,1909]" captionTargetId="figure-389@8.[219,1369,993,1914]" captionTargetPageId="8" captionText="FIGURES 1922. Chimerothalassius riparius sp. nov. 19. Head of male holotype (CNC1155797), anterior view. 20. Head of male holotype, showing left antenna in medial view. 21. Abdomen of female paratype, lateral view. 22. Right wing of male holotype, dorsal view. Abbreviations: cerc—cercus; M1, M4—medial veins; plp—palpus; R1, R2+3, R 4+5 —radial veins; tg—tergite." figureDoi="http://doi.org/10.5281/zenodo.7103634" httpUri="https://zenodo.org/record/7103634/files/figure.png" pageId="9" pageNumber="530">Fig. 21</figureCitation>
): Tapering posteriorly, apical segments retracted into segment 5.
<emphasis id="B950EABCC8192000E801E2C79C87ABE7" box="[853,975,186,212]" italics="true" pageId="9" pageNumber="530">Terminalia</emphasis>
with tergite 8 medially divided, narrowly fused with sternite 8 anterolaterally; syntergite 9+10 medially divided with 4 acanthophorous spines on each side; cercus with narrow, upturned and pointed tip, with short ventral setulae; spermathecal duct an unsclerotized tube with broad ridged sperm pump in basal part (terminal part lost in dissected female).
</paragraph>
</subSubSection>
<subSubSection id="C33E6525C8192000EB93E3379DA5AABB" pageId="9" pageNumber="530" type="distribution">
<paragraph id="8B9B36AEC8192000EB93E3379DA5AABB" blockId="9.[151,1437,150,609]" pageId="9" pageNumber="530">
<emphasis id="B950EABCC8192000EB93E3379E29AA57" bold="true" box="[199,353,330,356]" pageId="9" pageNumber="530">Distribution.</emphasis>
This new species is known only from the
<typeStatus id="549F880CC8192000E86DE3369C21AA56" box="[825,873,331,357]" pageId="9" pageNumber="530">type</typeStatus>
locality along Rivère Bleue in the Yaté Commune of the
<collectingRegion id="49E0F84CC8192000EB8BE3139EC3AABB" box="[223,395,366,392]" country="New Caledonia" name="Province Sud" pageId="9" pageNumber="530">South Province</collectingRegion>
of
<collectingCountry id="F333763EC8192000EAE4E3129D16AABB" box="[432,606,366,393]" name="New Caledonia" pageId="9" pageNumber="530">New Caledonia</collectingCountry>
(
<figureCitation id="131F2A2BC8192000E938E3139DA9AABB" box="[620,737,366,392]" captionStart="FIGURES 811" captionStartId="2.[151,264,1908,1932]" captionTargetBox="[167,1420,842,1878]" captionTargetId="figure-39@2.[167,1420,833,1884]" captionTargetPageId="2" captionText="FIGURES 811. Known geographical distribution and collection localities of Chimerothalassius and Eothalassius from New Caledonia. 8. Distribution of C. riparius sp. nov., C. sinclairi sp. nov. and Eothalassius sp. 910. Rivère Bleue, type locality of C. riparius sp. nov. and C. sinclairi sp. nov. (photos by Brad Sinclair). 11. Plage de Poé, collection locality of Eothalassius sp. (photo by Brad Sinclair)." figureDoi="http://doi.org/10.5281/zenodo.7103626" httpUri="https://zenodo.org/record/7103626/files/figure.png" pageId="9" pageNumber="530">Figs 810</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33E6525C8192000EB93E3EF9B88AA9F" box="[199,1216,402,429]" pageId="9" pageNumber="530" type="etymology">
<paragraph id="8B9B36AEC8192000EB93E3EF9B88AA9F" blockId="9.[151,1437,150,609]" box="[199,1216,402,429]" pageId="9" pageNumber="530">
<emphasis id="B950EABCC8192000EB93E3EF9E06AA9F" bold="true" box="[199,334,402,428]" pageId="9" pageNumber="530">Etymology.</emphasis>
The specific epithet refers to the riparian habit of the new species (
<figureCitation id="131F2A2BC8192000EF6BE3EF9BFAAA9F" box="[1087,1202,402,428]" captionStart="FIGURES 811" captionStartId="2.[151,264,1908,1932]" captionTargetBox="[167,1420,842,1878]" captionTargetId="figure-39@2.[167,1420,833,1884]" captionTargetPageId="2" captionText="FIGURES 811. Known geographical distribution and collection localities of Chimerothalassius and Eothalassius from New Caledonia. 8. Distribution of C. riparius sp. nov., C. sinclairi sp. nov. and Eothalassius sp. 910. Rivère Bleue, type locality of C. riparius sp. nov. and C. sinclairi sp. nov. (photos by Brad Sinclair). 11. Plage de Poé, collection locality of Eothalassius sp. (photo by Brad Sinclair)." figureDoi="http://doi.org/10.5281/zenodo.7103626" httpUri="https://zenodo.org/record/7103626/files/figure.png" pageId="9" pageNumber="530">Figs 9, 10</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33E6525C8192000EB93E3CB9C4AA953" pageId="9" pageNumber="530" type="discussion">
<paragraph id="8B9B36AEC8192000EB93E3CB9C4AA953" blockId="9.[151,1437,150,609]" pageId="9" pageNumber="530">
<emphasis id="B950EABCC8192000EB93E3CB9E73AAE3" bold="true" box="[199,315,438,464]" pageId="9" pageNumber="530">Remarks.</emphasis>
Formerly,
<bibRefCitation id="EFB54B5FC8192000EAE2E3CB9DA4AAE3" author="Cumming, J. M. &amp; Brooks, S. E." box="[438,748,438,464]" pageId="9" pageNumber="530" pagination="111 - 129" refId="ref15062" refString="Cumming, J. M. &amp; Brooks, S. E. (2019) Phylogenetic analysis and preliminary classification of the Parathalassiinae (Empidoidea: Dolichopodidae sensu lato). Zootaxa, 4648 (1), 111 - 129. https: // doi. org / 10.11646 / zootaxa. 4648.1.5" type="journal article" year="2019">Cumming &amp; Brooks (2019)</bibRefCitation>
included this exemplar species (
<emphasis id="B950EABCC8192000EF0CE3CA9B33AAE3" box="[1112,1147,439,464]" italics="true" pageId="9" pageNumber="530">i.e.</emphasis>
, “
<collectingCountry id="F333763EC8192000EFC6E3CB9A0AAAE3" box="[1170,1346,438,464]" name="New Caledonia" pageId="9" pageNumber="530">New Caledonia</collectingCountry>
sp.”) as a member of “Undescribed genus [
<collectingCountry id="F333763EC8192000E948E3A69D9AAAC7" box="[540,722,474,501]" name="New Caledonia" pageId="9" pageNumber="530">New Caledonia</collectingCountry>
]” in their phylogenetic analysis of the
<taxonomicName id="4C244D2DC8192000EFD5E3A79A70AAC7" baseAuthorityName="Cumming &amp; Brooks" baseAuthorityYear="2019" box="[1153,1336,474,500]" class="Insecta" family="Dolichopodidae" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="530" phylum="Arthropoda" rank="subFamily" subFamily="Parathalassiinae">Parathalassiinae</taxonomicName>
, because presence of female terminalia with acanthophorous spines was not considered typical of
<taxonomicName id="4C244D2DC8192000EF2AE3839A07A92B" authorityName="Shamshev &amp; Grootaert" authorityYear="2003" box="[1150,1359,510,536]" class="Insecta" family="Dolichopodidae" genus="Chimerothalassius" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="530" phylum="Arthropoda" rank="genus">
<emphasis id="B950EABCC8192000EF2AE3839A07A92B" box="[1150,1359,510,536]" italics="true" pageId="9" pageNumber="530">Chimerothalassius</emphasis>
</taxonomicName>
at that time. The type series of
<taxonomicName id="4C244D2DC8192000EAF0E05E9D57A90F" authority="Brooks &amp; Cumming, 2022" authorityName="Brooks &amp; Cumming" authorityYear="2022" box="[420,543,547,573]" class="Insecta" family="Dolichopodidae" genus="Chimerothalassius" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="530" phylum="Arthropoda" rank="species" species="riparius" status="sp. nov.">
<emphasis id="B950EABCC8192000EAF0E05E9D57A90F" box="[420,543,547,573]" italics="true" pageId="9" pageNumber="530">C. riparius</emphasis>
</taxonomicName>
<emphasis id="B950EABCC8192000E972E05F9D37A90F" bold="true" box="[550,639,546,572]" pageId="9" pageNumber="530">
<taxonomicNameLabel id="A26357C7C8192000E972E05F9D37A90F" box="[550,639,546,572]" pageId="9" pageNumber="530" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was collected together with the
<typeStatus id="549F880CC8192000E8BCE05F9B02A90F" box="[1000,1098,546,572]" pageId="9" pageNumber="530" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C244D2DC8192000EF3BE05E9BA5A90F" authority="Brooks &amp; Cumming, 2022" authorityName="Brooks &amp; Cumming" authorityYear="2022" box="[1135,1261,546,573]" class="Insecta" family="Dolichopodidae" genus="Chimerothalassius" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="530" phylum="Arthropoda" rank="species" species="sinclairi" status="sp. nov.">
<emphasis id="B950EABCC8192000EF3BE05E9BA5A90F" box="[1135,1261,546,573]" italics="true" pageId="9" pageNumber="530">C. sinclairi</emphasis>
</taxonomicName>
<emphasis id="B950EABCC8192000EFA0E05F9A05A90F" bold="true" box="[1268,1357,546,572]" pageId="9" pageNumber="530">
<taxonomicNameLabel id="A26357C7C8192000EFA0E05F9A05A90F" box="[1268,1357,546,572]" pageId="9" pageNumber="530" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in yellow pan traps placed along the margin of Rivère Bleue.
</paragraph>
</subSubSection>
</treatment>
</document>