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<document id="9282A8365E2788E7CD119080B7B5AD98" ID-DOI="10.11646/phytotaxa.48.1.7" ID-ISSN="1179-3163" ID-Zenodo-Dep="4776766" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1621533780868" checkinUser="felipe" docAuthor="Snow, Neil, Rabenantoandro, Johny, Randriatafika, Faly, Rabehevitra, David, Razafimamonjy, N. Darie &amp; Cable, Stuart" docDate="2012" docId="0381A35C006EFFD26099F829FC257DB8" docLanguage="en" docName="Phytotaxa.48.39-60.pdf" docOrigin="Phytotaxa 48 (1)" docSource="http://biotaxa.org/Phytotaxa/article/view/phytotaxa.48.1.7" docStyle="DocumentStyle:13F189B31CB4CCCE9197361A4E100104.3:Phytotaxa.2011-2013.journal_article" docStyleId="13F189B31CB4CCCE9197361A4E100104" docStyleName="Phytotaxa.2011-2013.journal_article" docStyleVersion="3" docTitle="Eugenia petrikensis N. Snow &amp; Randriat. 2012, sp. nov." docType="treatment" docVersion="3" lastPageNumber="49" masterDocId="FFB8DB240069FFD8600EFFAEFF917F10" masterDocTitle="Studies of Malagasy Eugenia (Myrtaceae) - III: Seven new species of high conservation concern from the eastern littoral forests" masterLastPageNumber="60" masterPageNumber="39" pageNumber="46" updateTime="1698924875650" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="9C9AE2B2EFBB98B33992888FF3ED3BB9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4CF23727587E9FD7FFED81095E2B469C">
<mods:title id="FC6E0C9811194FBFE0F4EB970A6B46B5">Studies of Malagasy Eugenia (Myrtaceae) - III: Seven new species of high conservation concern from the eastern littoral forests</mods:title>
</mods:titleInfo>
<mods:name id="1325B04E8015E024E7A27A9034392E69" type="personal">
<mods:role id="BAFC99CD6809BC94022B735273322565">
<mods:roleTerm id="27C96B31D96DB82A92068D4E51716B80">Author</mods:roleTerm>
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<mods:namePart id="B2CB7906E3D01F38036D96D0234C2614">Snow, Neil</mods:namePart>
<mods:affiliation id="367042C623AA66FA5FE65AFBB5208709">Herbarium Pacificum, B. P. Bishop Museum, 1525 Bernice Street, Honolulu, Hawaii 96825, USA</mods:affiliation>
<mods:nameIdentifier id="EEC96B90130F6BAFDADB8E878133B0F6" type="email">nsnow2@mt.gov</mods:nameIdentifier>
</mods:name>
<mods:name id="A22270CEB4E1259A8A4DDB33A4D5F818" type="personal">
<mods:role id="FC9792D7CB647DC5D9237C05B944A40B">
<mods:roleTerm id="D8A421C44C407398C3B93EBA0A092273">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A605A7092D5614979A8F290C5FCBAF0C">Rabenantoandro, Johny</mods:namePart>
<mods:affiliation id="3F6A327BF701DFD5DA4F72ABBBCFD1D9">Rio Tinto / QIT Madagascar Minerals S. A., Villa 3 H Lot J, 169 Antananarivo, 101, MADAGASCAR; E-mails: johny. rabenantoandro @ riotinto. com;</mods:affiliation>
<mods:nameIdentifier id="46C667E7C1960EEEA38EFF0FEDA3B851" type="email">johny.rabenantoandro@riotinto.com</mods:nameIdentifier>
</mods:name>
<mods:name id="8F4CC85A6540E6A07FF40483E1A55AD2" type="personal">
<mods:role id="335718B5AC99A14E6DB0180FE46E93C1">
<mods:roleTerm id="B3DDB5404B92BE069D6B6818DC4CCA01">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E617184935827ED4EA1789E6A75594CA">Randriatafika, Faly</mods:namePart>
<mods:affiliation id="8388D807AE812978D557E06C61BBCAF6">Rio Tinto / QIT Madagascar Minerals S. A., Villa 3 H Lot J, 169 Antananarivo, 101, MADAGASCAR; E-mails: johny. rabenantoandro @ riotinto. com;</mods:affiliation>
<mods:nameIdentifier id="E797E64A1D5E2DD6BE554CEC556AC452" type="email">johny.rabenantoandro@riotinto.com</mods:nameIdentifier>
</mods:name>
<mods:name id="E552EAB334D1ADA61C3E8C6CB9262F8B" type="personal">
<mods:role id="A71C268583BB0143C0C95F063297C3BA">
<mods:roleTerm id="39586F3ADD9EBDA28A75588C441840C0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B8557374558D6FC9796604146C20A5D7">Rabehevitra, David</mods:namePart>
<mods:affiliation id="1902ABFCB74DBA4FCD0BD64DC19F421D">Rio Tinto / QIT Madagascar Minerals S. A., Villa 3 H Lot J, 169 Antananarivo, 101, MADAGASCAR; E-mails: johny. rabenantoandro @ riotinto. com;</mods:affiliation>
<mods:nameIdentifier id="2B4510DD8367C1C4ABF744CA45B946F9" type="email">johny.rabenantoandro@riotinto.com</mods:nameIdentifier>
</mods:name>
<mods:name id="C3E3E8D47452DD07DA9D3A9FF891D099" type="personal">
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<mods:roleTerm id="CE555DD4FAC5668FD1260433B5DD05FF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="92E9B38DA855C4B0A0E789D6EACF3820">Razafimamonjy, N. Darie</mods:namePart>
<mods:affiliation id="E6C284A64BD2AB4C21C10AA54BF989E1">Department of Botany, University of Antananarivo, MADAGASCAR; E-mail: dariejosmail @ yahoo. com</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="A44B78FBD5267826C5666A87141C4C64">Cable, Stuart</mods:namePart>
<mods:affiliation id="2944DEA7453068C6ED268E37C90FB151">Royal Botanic Gardens, Kew, Richmond, Surrey, UNITED KINGDOM; E-mail: s. cable @ kew. org</mods:affiliation>
</mods:name>
<mods:typeOfResource id="BA1D57008B9146B60B6348ED547B2890">text</mods:typeOfResource>
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<mods:title id="7EE764CCB50178EA985A692F17A680D2">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="395E23B5E5E75FD91A3B950D11D6286C">
<mods:date id="30981527C2F30E17273CBB10CBA5439E">2012</mods:date>
<mods:detail id="831F5CA72DA82D97088A33D2B1796605" type="pubDate">
<mods:number id="B68BFAF40FB4591DDBB89994D4C2C0C4">2012-03-15</mods:number>
</mods:detail>
<mods:detail id="3C6DD2C00FF1CBAF939E0310ECE16AD1" type="volume">
<mods:number id="6B1AF23381F3B3D81A36465BB57BE68C">48</mods:number>
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<mods:number id="23683485222C8F6DB9936ECAB2ED41EC">1</mods:number>
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<mods:start id="E7C5BB7F3ED9116184D2A58BDEDD08B4">39</mods:start>
<mods:end id="4BC4F796FAFFF33F251CA4E8F492320F">60</mods:end>
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<mods:url id="C65F6F41E1EA3EBF460B48DD4B8F8EC6">http://biotaxa.org/Phytotaxa/article/view/phytotaxa.48.1.7</mods:url>
</mods:location>
<mods:classification id="8B3488937D986E6267FA2CEC8CD6AE5E">journal article</mods:classification>
<mods:identifier id="53ACFE550D7BEADF8E9C4BAB41AB5951" type="DOI">10.11646/phytotaxa.48.1.7</mods:identifier>
<mods:identifier id="10AF74975298FB0A5A23C8699011EF3C" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="BD090CFC6851A19ECEDFF7E8924D124B" type="Zenodo-Dep">4776766</mods:identifier>
</mods:mods>
<treatment id="0381A35C006EFFD26099F829FC257DB8" LSID="urn:lsid:plazi:treatment:0381A35C006EFFD26099F829FC257DB8" httpUri="http://treatment.plazi.org/id/0381A35C006EFFD26099F829FC257DB8" lastPageId="10" lastPageNumber="49" pageId="7" pageNumber="46">
<subSubSection id="C33241C1006EFFDF6099F829FA0D78F9" pageId="7" pageNumber="46" type="nomenclature">
<paragraph id="8B97124A006EFFDF6099F829FC4678B4" blockId="7.[151,983,1927,1956]" box="[151,983,1927,1956]" pageId="7" pageNumber="46">
<heading id="D0DFA526006EFFDF6099F829FC4678B4" bold="true" box="[151,983,1927,1956]" fontSize="12" level="1" pageId="7" pageNumber="46" reason="1">
<emphasis id="B95CCE58006EFFDF6099F829FC4678B4" bold="true" box="[151,983,1927,1956]" pageId="7" pageNumber="46">
<taxonomicName id="4C2869C9006EFFDF6099F829FD5978B4" authority="N. Snow &amp; Randriat." authorityName="N. Snow &amp; Randriat." authorityYear="2012" box="[151,712,1927,1956]" class="Insecta" family="Tachinidae" genus="Eugenia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="46" phylum="Arthropoda" rank="species" species="petrikensis" status="sp. nov.">
<emphasis id="B95CCE58006EFFDF6099F829FE0C78B3" bold="true" box="[151,413,1927,1955]" italics="true" pageId="7" pageNumber="46">Eugenia petrikensis</emphasis>
N. Snow &amp; Randriat.
</taxonomicName>
,
<emphasis id="B95CCE58006EFFDF62D8F829FCA878B3" bold="true" box="[726,825,1927,1955]" italics="true" pageId="7" pageNumber="46">
<taxonomicNameLabel id="A26F7323006EFFDF62D8F829FCA878B3" box="[726,825,1927,1955]" pageId="7" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13130ECF006EFFDF6344F829FC3478B4" box="[842,933,1927,1956]" captionStart="FIGURE 2" captionStartId="2.[151,244,1785,1806]" captionTargetBox="[166,1430,868,1771]" captionTargetId="figure-397@2.[166,1431,868,1771]" captionTargetPageId="2" captionText="FIGURE 2. Distribution of species and selected specimens of Eugenia in southeastern Madagascar based on vouchered herbarium specimens and recently captured (but non-vouchered) digital images. Remaining forested areas (Petriky, Mandena, Sainte-Luce) along the coast are indicated in gray. In cases where the icons largely or completely overlapped based on geo-coordinates they have been separated slightly for easier viewing (e.g., E. ardyceae, E. stictophylla and E. cloiselii along the coast west of Tolagnaro). Many additional specimens of Eugenia from the region await identification or description and cannot be mapped confidently at the present time." figureDoi="http://doi.org/10.5281/zenodo.4776772" httpUri="https://zenodo.org/record/4776772/files/figure.png" pageId="7" pageNumber="46">Figs. 2</figureCitation>
,
<figureCitation id="13130ECF006EFFDF63BAF829FC5778B4" box="[948,966,1927,1956]" captionStart="FIGURE 5" captionStartId="9.[151,244,1644,1665]" captionTargetBox="[151,1436,181,1622]" captionTargetId="figure-162@9.[151,1436,181,1622]" captionTargetPageId="9" captionText="FIGURE 5. Eugenia petrikensis. Upper left: bark of main bole. Upper right: sessile to subsessile inflorescences. Lower left: emerging leaves with whitish indumentum. Lower right: mature leaves; abaxial (upper) and adaxial (lower) surfaces. (No voucher; photos by D. Rabehevitra)." figureDoi="http://doi.org/10.5281/zenodo.4776784" httpUri="https://zenodo.org/record/4776784/files/figure.png" pageId="7" pageNumber="46">5</figureCitation>
).
</emphasis>
</heading>
</paragraph>
<paragraph id="8B97124A006EFFDF6099F87CFA0D78F9" blockId="7.[151,1436,2002,2025]" box="[151,1436,2002,2025]" pageId="7" pageNumber="46">
<emphasis id="B95CCE58006EFFDF6099F87CFA0D78F9" box="[151,1436,2002,2025]" italics="true" pageId="7" pageNumber="46">
A congeneribis madagascariensibus combinatione characterum sequentium distinguitur: frutex
<quantity id="4CD0BFAF006EFFDF64A1F87CFA9978F9" box="[1199,1288,2002,2025]" metricMagnitude="0" metricUnit="m" metricValue="1.15" metricValueMax="2.0" metricValueMin="0.3" pageId="7" pageNumber="46" unit="m" value="1.15" valueMax="2.0" valueMin="0.3">0.3–2 m</quantity>
altus; petioli
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33241C10061FFD060C9FF36FC877FDF" pageId="8" pageNumber="47" type="description">
<paragraph id="8B97124A0061FFD060C9FF36FC877FDF" blockId="8.[199,1436,152,207]" pageId="8" pageNumber="47">
<emphasis id="B95CCE580061FFD060C9FF36FC877FDF" italics="true" pageId="8" pageNumber="47">
<quantity id="4CD0BFAF0061FFD060C9FF36FED47FBF" box="[199,325,152,175]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.5" metricValueMin="3.5" pageId="8" pageNumber="47" unit="mm" value="4.0" valueMax="4.5" valueMin="3.5">3.5–4.5 mm</quantity>
longi; folia
<quantity id="4CD0BFAF0061FFD061DEFF36FDC77FBF" box="[464,598,152,175]" metricMagnitude="-2" metricUnit="m" metricValue="2.325" metricValueMax="3.0" metricValueMin="1.65" pageId="8" pageNumber="47" unit="mm" value="23.25" valueMax="30.0" valueMin="16.5">16.5–30 mm</quantity>
longa,
<quantity id="4CD0BFAF0061FFD062A4FF36FCA17FBF" box="[682,816,152,175]" metricMagnitude="-2" metricUnit="m" metricValue="1.375" metricValueMax="1.75" metricValueMin="1.0" pageId="8" pageNumber="47" unit="mm" value="13.75" valueMax="17.5" valueMin="10.0">10–17.5 mm</quantity>
lata, elliptica, basi cuneata vel rotundata, apice obtuso; petala magentea ad marginem hyalina; stamina 20–30.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33241C10061FFD06099FF59FE7A7ABC" pageId="8" pageNumber="47" type="materials_examined">
<paragraph id="8B97124A0061FFD06099FF59FBD37E3F" blockId="8.[151,1437,247,303]" pageId="8" pageNumber="47">
<emphasis id="B95CCE580061FFD06099FF59FF497E1F" bold="true" box="[151,216,247,271]" pageId="8" pageNumber="47">Type:</emphasis>
―
<materialsCitation id="3B4018170061FFD060FCFF56FBAF7E3F" collectingDate="1989-10-26" collectionCode="R" collectorName="de Petriky &amp; N. Dumetz &amp; G. McPherson &amp; Rabevohitra" country="Madagascar" latitude="-25.083334" location="District Tolagnaro" longLatPrecision="1248" longitude="46.866665" pageId="8" pageNumber="47" specimenCount="1" stateProvince="Toliara" typeStatus="holotype">
<collectingCountry id="F33F52DA0061FFD060FCFF56FE227E1F" box="[242,435,248,271]" name="Madagascar" pageId="8" pageNumber="47">MADAGASCAR</collectingCountry>
. Prov.
<collectingRegion id="49ECDCA80061FFD0620EFF56FDDE7E1F" box="[512,591,248,271]" country="Madagascar" name="Toliara" pageId="8" pageNumber="47">Toliara</collectingRegion>
:
<location id="8EF744910061FFD06252FF56FCB57E1F" LSID="urn:lsid:plazi:treatment:0381A35C006EFFD26099F829FC257DB8:8EF744910061FFD06252FF56FCB57E1F" box="[604,804,248,271]" country="Madagascar" latitude="-25.083334" longLatPrecision="1248" longitude="46.866665" name="District Tolagnaro" pageId="8" pageNumber="47" stateProvince="Toliara">District Tolagnaro</location>
; forêt
<collectorName id="26DD779C0061FFD06363FF56FC4C7E1F" box="[877,989,248,271]" pageId="8" pageNumber="47">de Petriky</collectorName>
,
<geoCoordinate id="EE1C748D0061FFD063E5FF56FBD57E1F" box="[1003,1092,248,271]" degrees="25" direction="south" minutes="05" orientation="latitude" pageId="8" pageNumber="47" precision="925" value="-25.083334">25°05’S</geoCoordinate>
,
<geoCoordinate id="EE1C748D0061FFD0645CFF56FB3C7E1F" box="[1106,1197,248,271]" degrees="46" direction="east" minutes="52" orientation="longitude" pageId="8" pageNumber="47" precision="925" value="46.866665">46°52’E</geoCoordinate>
,
<date id="FF96348A0061FFD064B5FF56FAE77E1F" box="[1211,1398,248,271]" pageId="8" pageNumber="47" value="1989-10-26">
<collectingDate id="EFD2CD620061FFD064B5FF56FAE77E1F" box="[1211,1398,248,271]" pageId="8" pageNumber="47" value="1989-10-26">26 October 1989</collectingDate>
</date>
,
<emphasis id="B95CCE580061FFD0658AFF56FD217E3F" italics="true" pageId="8" pageNumber="47">
<collectorName id="26DD779C0061FFD0658AFF56FE897E3F" pageId="8" pageNumber="47">N. Dumetz</collectorName>
812,
<collectorName id="26DD779C0061FFD0615AFEB6FE7D7E3F" box="[340,492,280,303]" pageId="8" pageNumber="47">G. McPherson</collectorName>
&amp;
<collectionCode id="ED398A8F0061FFD06200FEB6FDB07E3F" box="[526,545,280,303]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="8" pageNumber="47">R</collectionCode>
.
<collectorName id="26DD779C0061FFD06222FEB6FD217E3F" box="[556,688,280,303]" pageId="8" pageNumber="47">Rabevohitra</collectorName>
</emphasis>
(
<typeStatus id="5493ACE80061FFD062B0FEB6FC8C7E3F" box="[702,797,280,303]" pageId="8" pageNumber="47" type="holotype">holotype</typeStatus>
MO! [sheet no. 3778658])
</materialsCitation>
.
</paragraph>
<paragraph id="8B97124A0061FFD06099FEF6FD667A28" blockId="8.[151,1436,344,1994]" pageId="8" pageNumber="47">
<emphasis id="B95CCE580061FFD06099FEF6FF7B7E61" box="[151,234,344,369]" italics="true" pageId="8" pageNumber="47">Shrubs</emphasis>
0.3–2 meters at flowering.
<emphasis id="B95CCE580061FFD06239FEF6FD297E61" box="[567,696,344,369]" italics="true" pageId="8" pageNumber="47">Branchlets</emphasis>
moderately to densely sericeous on emergence but becoming sparsely so to nearly glabrous (trichomes dibrachiate, dense, clear to ferrugineous that become whitish brown); oil glands not visible or only faint (becoming more prominent in age), occasional; internodes ca. 1.0–
<quantity id="4CD0BFAF0061FFD06099FE62FF797EF6" box="[151,232,460,486]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="8" pageNumber="47" unit="cm" value="1.2">1.2 cm</quantity>
long.
<emphasis id="B95CCE580061FFD0613AFE62FE177EF5" box="[308,390,460,485]" italics="true" pageId="8" pageNumber="47">Leaves</emphasis>
coriaceous, evenly distributed along branchlets, slightly discolorous, emerald green and slightly glossy above, citrine and matte below; venation brochidodromous. Stipules absent.
<emphasis id="B95CCE580061FFD064D2FE5CFAAA7D1B" box="[1244,1339,498,523]" italics="true" pageId="8" pageNumber="47">Petioles</emphasis>
3.0–
<quantity id="4CD0BFAF0061FFD06579FE5CFF587D23" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="8" pageNumber="47" unit="mm" value="4.5">4.5 mm</quantity>
, slightly sulcate above, eglandular but rugose-wrinkled throughout, glabrous, yellowish-green.
<emphasis id="B95CCE580061FFD06568FDB7FF757D49" italics="true" pageId="8" pageNumber="47">Leaf blades</emphasis>
1.6–3.0 x 1.0–
<quantity id="4CD0BFAF0061FFD06197FDEEFE7D7D4A" box="[409,492,576,602]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="8" pageNumber="47" unit="cm" value="1.7">1.7 cm</quantity>
, elliptic, base cuneate to rounded, margin somewhat revolute except near apex, laminar surface flat, apex and tip obtuse; adaxial surface sparsely sericeous towards base, oil glands absent, midvein slightly sulcate proximally but becoming flush distally; abaxial surface glabrous, oil glands absent, secondary veins invisible or faint with 4–5 veins arising on each side of midvein, sometimes slightly raised (dried material), intramarginal vein faint,
<quantity id="4CD0BFAF0061FFD06273FD74FC9B7DE4" box="[637,778,730,756]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="8" pageNumber="47" unit="mm" value="0.35" valueMax="0.5" valueMin="0.2">0.2–0.5 mm</quantity>
from margin at laminar midpoint.
<emphasis id="B95CCE580061FFD06491FD74FAAD7DE3" box="[1183,1340,730,755]" italics="true" pageId="8" pageNumber="47">Inflorescence</emphasis>
axillary or ramiflorous, consisting of monads or 3–flowered cymes, solitary, paired or generally in fascicles (sometimes densely so).
<emphasis id="B95CCE580061FFD061B4FC86FDA37C51" box="[442,562,808,833]" italics="true" pageId="8" pageNumber="47">Peduncles</emphasis>
(or pedicels, if flowers solitary)
<quantity id="4CD0BFAF0061FFD063B6FC86FB897C52" box="[952,1048,808,834]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="8" pageNumber="47" unit="mm" value="3.5" valueMax="5.0" valueMin="2.0">2–5 mm</quantity>
long, stiff, terete to compressed, medium green, glabrous or with a few scattered hairs. Anthopodia absent; metaxphylls mostly absent (present on one flower).
<emphasis id="B95CCE580061FFD0615FFCDBFE5F7C9E" box="[337,462,885,910]" italics="true" pageId="8" pageNumber="47">Bracteoles</emphasis>
<quantity id="4CD0BFAF0061FFD061DBFCDBFDF07C9F" box="[469,609,885,911]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="8" pageNumber="47" unit="mm" value="0.6" valueMax="0.7" valueMin="0.5">0.5–0.7 mm</quantity>
long, ca.
<quantity id="4CD0BFAF0061FFD062DDFCDBFCBC7C9F" box="[723,813,885,911]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="47" unit="mm" value="0.3">0.3 mm</quantity>
wide at base, narrowly triangular to triangular, scalelike, spreading, stiff, sparsely hairy, persistent in flower.
<emphasis id="B95CCE580061FFD06334FC32FC5A7CA5" box="[826,971,924,949]" italics="true" pageId="8" pageNumber="47">Hypanthium</emphasis>
1.0–
<quantity id="4CD0BFAF0061FFD06407FC32FBF27CA6" box="[1033,1123,924,950]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="47" unit="mm" value="1.5">1.5 mm</quantity>
long,
<quantity id="4CD0BFAF0061FFD064A1FC32FAAC7CA6" box="[1199,1341,924,950]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="8" pageNumber="47" unit="mm" value="1.4" valueMax="1.5" valueMin="1.3">1.3–1.5 mm</quantity>
wide at apex at base of calyx lobes, obconic, glabrous, whitish to ivory or stramineous, oil glands common but small and inconspicuous, surface smooth, apex of ovary glabrous at base of style but villous towards staminal ring.
<emphasis id="B95CCE580061FFD06099FBBEFEB77B39" box="[151,294,1040,1065]" italics="true" pageId="8" pageNumber="47">Calyx lobes</emphasis>
4,
<quantity id="4CD0BFAF0061FFD06140FBBEFE4E7B3A" box="[334,479,1040,1066]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.8" metricValueMin="1.4" pageId="8" pageNumber="47" unit="mm" value="1.6" valueMax="1.8" valueMin="1.4">1.4–1.8 mm</quantity>
long, rounded to oblong, apex broadly rounded, glabrous, upper surface often magenta or cerise during anthesis and hyaline along margin, lower surface greenish.
<emphasis id="B95CCE580061FFD0648CFB99FB5A7B40" box="[1154,1227,1079,1104]" italics="true" pageId="8" pageNumber="47">Petals</emphasis>
4, 3.0–4.1 x
<quantity id="4CD0BFAF0061FFD06566FB98FF677B67" metricMagnitude="-3" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="8" pageNumber="47" unit="mm" value="2.15" valueMax="2.2" valueMin="2.1">2.1– 2.2 mm</quantity>
, ovate, oil glands sparse, magenta but hyaline on margins. Staminal ring villous, diameter
<quantity id="4CD0BFAF0061FFD0654CFBF3FF587B8E" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="8" pageNumber="47" unit="mm" value="1.65" valueMax="1.8" valueMin="1.5">1.5–1.8 mm</quantity>
.
<emphasis id="B95CCE580061FFD060D8FB2AFEAB7B8D" box="[214,314,1156,1181]" italics="true" pageId="8" pageNumber="47">Stamens</emphasis>
20–30, in 1 or 2 series; filaments ca.
<quantity id="4CD0BFAF0061FFD0631EFB2AFCE47B8E" box="[784,885,1156,1182]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="8" pageNumber="47" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
long, whitish; anther sacs
<quantity id="4CD0BFAF0061FFD064CCFB2AFAC47B8E" box="[1218,1365,1156,1182]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="8" pageNumber="47" unit="mm" value="0.8" valueMax="0.9" valueMin="0.7">0.7–0.9 mm</quantity>
long, globose to cylindrical, dorsifixed, bearing a single apical gland, cream colored.
<emphasis id="B95CCE580061FFD0644BFB05FBEC7BD4" box="[1093,1149,1195,1220]" italics="true" pageId="8" pageNumber="47">Style</emphasis>
ca.
<quantity id="4CD0BFAF0061FFD064A0FB05FB637BD5" box="[1198,1266,1195,1221]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="47" unit="mm" value="3.0">3 mm</quantity>
long, whitish, glabrous; stigma punctiform (scarcely if at all capitate). Locules 2; placentation axile, capitate, ovules 5–7 per placenta and radiating irregularly from placenta.
<emphasis id="B95CCE580061FFD062D3FB56FC8B7A01" box="[733,794,1272,1297]" italics="true" pageId="8" pageNumber="47">Fruit</emphasis>
(dimensions not measured) globose, glabrous, yellow at maturity, apex crowned by persistent calyx lobes.
</paragraph>
<paragraph id="8B97124A0061FFD060C8FAEBFE7A7ABC" blockId="8.[151,1436,344,1994]" pageId="8" pageNumber="47">
<emphasis id="B95CCE580061FFD060C8FAEBFE0A7A4F" bold="true" box="[198,411,1349,1375]" pageId="8" pageNumber="47">Field characters:</emphasis>
―Shrubs up to 2 meters tall; flowers often ramiflorous in clusters and relatively small; hypanthium glabrous; calyx lobes sometimes with three relatively prominent veins; petals magenta; stamens relatively 30 or less. (
<figureCitation id="13130ECF0061FFD0619AFA3CFE4D7ABC" box="[404,476,1426,1452]" captionStart="FIGURE 5" captionStartId="9.[151,244,1644,1665]" captionTargetBox="[151,1436,181,1622]" captionTargetId="figure-162@9.[151,1436,181,1622]" captionTargetPageId="9" captionText="FIGURE 5. Eugenia petrikensis. Upper left: bark of main bole. Upper right: sessile to subsessile inflorescences. Lower left: emerging leaves with whitish indumentum. Lower right: mature leaves; abaxial (upper) and adaxial (lower) surfaces. (No voucher; photos by D. Rabehevitra)." figureDoi="http://doi.org/10.5281/zenodo.4776784" httpUri="https://zenodo.org/record/4776784/files/figure.png" pageId="8" pageNumber="47">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33241C10061FFD060C8FA16FB3E79AB" pageId="8" pageNumber="47" type="distribution">
<paragraph id="8B97124A0061FFD060C8FA16FD5C7957" blockId="8.[151,1436,344,1994]" pageId="8" pageNumber="47">
<emphasis id="B95CCE580061FFD060C8FA16FEF87AC2" bold="true" box="[198,361,1464,1490]" pageId="8" pageNumber="47">Distribution:</emphasis>
―Known only from the eastern part of Petriky Forest (
<figureCitation id="13130ECF0061FFD06409FA17FBDE7AC3" box="[1031,1103,1465,1491]" captionStart="FIGURE 2" captionStartId="2.[151,244,1785,1806]" captionTargetBox="[166,1430,868,1771]" captionTargetId="figure-397@2.[166,1431,868,1771]" captionTargetPageId="2" captionText="FIGURE 2. Distribution of species and selected specimens of Eugenia in southeastern Madagascar based on vouchered herbarium specimens and recently captured (but non-vouchered) digital images. Remaining forested areas (Petriky, Mandena, Sainte-Luce) along the coast are indicated in gray. In cases where the icons largely or completely overlapped based on geo-coordinates they have been separated slightly for easier viewing (e.g., E. ardyceae, E. stictophylla and E. cloiselii along the coast west of Tolagnaro). Many additional specimens of Eugenia from the region await identification or description and cannot be mapped confidently at the present time." figureDoi="http://doi.org/10.5281/zenodo.4776772" httpUri="https://zenodo.org/record/4776772/files/figure.png" pageId="8" pageNumber="47">Fig. 2</figureCitation>
) and Ambinanibe Forest in
<collectingRegion id="49ECDCA80061FFD06099FA4EFECF7AEA" box="[151,350,1504,1530]" country="Madagascar" name="Toliara" pageId="8" pageNumber="47">Toliara Province</collectingRegion>
in southeastern
<collectingCountry id="F33F52DA0061FFD0622BFA4EFD297AEA" box="[549,696,1504,1530]" name="Madagascar" pageId="8" pageNumber="47">Madagascar</collectingCountry>
. Ambinanibe Forest (not labeled in
<figureCitation id="13130ECF0061FFD06465FA4EFB257AEA" box="[1131,1204,1504,1530]" captionStart="FIGURE 2" captionStartId="2.[151,244,1785,1806]" captionTargetBox="[166,1430,868,1771]" captionTargetId="figure-397@2.[166,1431,868,1771]" captionTargetPageId="2" captionText="FIGURE 2. Distribution of species and selected specimens of Eugenia in southeastern Madagascar based on vouchered herbarium specimens and recently captured (but non-vouchered) digital images. Remaining forested areas (Petriky, Mandena, Sainte-Luce) along the coast are indicated in gray. In cases where the icons largely or completely overlapped based on geo-coordinates they have been separated slightly for easier viewing (e.g., E. ardyceae, E. stictophylla and E. cloiselii along the coast west of Tolagnaro). Many additional specimens of Eugenia from the region await identification or description and cannot be mapped confidently at the present time." figureDoi="http://doi.org/10.5281/zenodo.4776772" httpUri="https://zenodo.org/record/4776772/files/figure.png" pageId="8" pageNumber="47">Fig. 2</figureCitation>
), only infrequently indicated on maps and from digital sources, is the highly fragmented area west of Tolagnaro, north of Lac Andriambe, an area of active mining operations.
</paragraph>
<paragraph id="8B97124A0061FFD060C8F9FDFE307984" blockId="8.[151,1436,344,1994]" pageId="8" pageNumber="47">
<emphasis id="B95CCE580061FFD060C8F9FDFE5A797D" bold="true" box="[198,459,1619,1645]" pageId="8" pageNumber="47">Habitat and ecology:</emphasis>
―Growing in full sun (or more rarely in shade) in low forest on littoral sands from 0–10 meters elevation.
</paragraph>
<paragraph id="8B97124A0061FFD060C8F90EFB3E79AB" blockId="8.[151,1436,344,1994]" box="[198,1199,1696,1723]" pageId="8" pageNumber="47">
<emphasis id="B95CCE580061FFD060C8F90EFEC379AA" bold="true" box="[198,338,1696,1722]" pageId="8" pageNumber="47">Phenology:</emphasis>
―Flowering late October to February; fruiting from December to March.
</paragraph>
</subSubSection>
<subSubSection id="C33241C10061FFD160C8F969FACF78E0" lastPageId="9" lastPageNumber="48" pageId="8" pageNumber="47" type="etymology">
<paragraph id="8B97124A0061FFD060C8F969FC6979F2" blockId="8.[151,1436,344,1994]" box="[198,1016,1735,1762]" pageId="8" pageNumber="47">
<emphasis id="B95CCE580061FFD060C8F969FEC679F1" bold="true" box="[198,343,1735,1761]" pageId="8" pageNumber="47">Etymology:</emphasis>
―The specific epithet is based on Petriky Forest (
<figureCitation id="13130ECF0061FFD063AFF966FC7879F2" box="[929,1001,1736,1762]" captionStart="FIGURE 5" captionStartId="9.[151,244,1644,1665]" captionTargetBox="[151,1436,181,1622]" captionTargetId="figure-162@9.[151,1436,181,1622]" captionTargetPageId="9" captionText="FIGURE 5. Eugenia petrikensis. Upper left: bark of main bole. Upper right: sessile to subsessile inflorescences. Lower left: emerging leaves with whitish indumentum. Lower right: mature leaves; abaxial (upper) and adaxial (lower) surfaces. (No voucher; photos by D. Rabehevitra)." figureDoi="http://doi.org/10.5281/zenodo.4776784" httpUri="https://zenodo.org/record/4776784/files/figure.png" pageId="8" pageNumber="47">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="8B97124A0061FFD060C8F940FEE378DA" blockId="8.[151,1436,344,1994]" pageId="8" pageNumber="47">
<emphasis id="B95CCE580061FFD060C8F940FE577818" bold="true" box="[198,454,1774,1800]" pageId="8" pageNumber="47">Vernacular name:—</emphasis>
<emphasis id="B95CCE580061FFD061C8F941FDFD7818" box="[454,620,1775,1800]" italics="true" pageId="8" pageNumber="47">Ropasy lahiny</emphasis>
.
<emphasis id="B95CCE580061FFD06272F941FD437818" box="[636,722,1775,1800]" italics="true" pageId="8" pageNumber="47">Ropasy</emphasis>
is a contraction of
<emphasis id="B95CCE580061FFD063B6F941FC637818" box="[952,1010,1775,1800]" italics="true" pageId="8" pageNumber="47">rotry</emphasis>
and
<emphasis id="B95CCE580061FFD06422F941FBFF7818" box="[1068,1134,1775,1800]" italics="true" pageId="8" pageNumber="47">fasiky</emphasis>
, the former of which is a generic vernacular name used for species of
<taxonomicName id="4C2869C90061FFD062A6F8BBFC9B783E" box="[680,778,1813,1838]" class="Insecta" family="Tachinidae" genus="Eugenia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="B95CCE580061FFD062A6F8BBFC9B783E" box="[680,778,1813,1838]" italics="true" pageId="8" pageNumber="47">Eugenia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2869C90061FFD0634BF8BBFC25783E" box="[837,948,1813,1838]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" higherTaxonomySource="IPNI" kingdom="Plantae" order="Myrtales" pageId="8" pageNumber="47" phylum="Magnoliophyta" rank="genus">
<emphasis id="B95CCE580061FFD0634BF8BBFC25783E" box="[837,948,1813,1838]" italics="true" pageId="8" pageNumber="47">Syzygium</emphasis>
</taxonomicName>
around Tolagnaro.
<emphasis id="B95CCE580061FFD06493F8BBFB7D783E" box="[1181,1260,1813,1838]" italics="true" pageId="8" pageNumber="47">Fasiky</emphasis>
refers to sand; thus
<emphasis id="B95CCE580061FFD060DFF892FEB37845" box="[209,290,1852,1877]" italics="true" pageId="8" pageNumber="47">ropasy</emphasis>
means “
<taxonomicName id="4C2869C90061FFD06185F892FE7F7845" box="[395,494,1852,1877]" class="Insecta" family="Tachinidae" genus="Eugenia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="B95CCE580061FFD06185F892FE7F7845" box="[395,494,1852,1877]" italics="true" pageId="8" pageNumber="47">Eugenia</emphasis>
</taxonomicName>
that grows on sand”. When more than one species of a genus occurs locally a second name often is added for distinction, such as
<emphasis id="B95CCE580061FFD062FAF8CDFCAC786C" box="[756,829,1891,1916]" italics="true" pageId="8" pageNumber="47">lahiny</emphasis>
(male) or
<emphasis id="B95CCE580061FFD063B7F8CDFB97786C" box="[953,1030,1891,1916]" italics="true" pageId="8" pageNumber="47">vaviny</emphasis>
(female). The name
<emphasis id="B95CCE580061FFD064F5F8CDFADB786C" box="[1275,1354,1891,1916]" italics="true" pageId="8" pageNumber="47">ropasy</emphasis>
is also used in Mandena (north of Tolagnaro) for
<taxonomicName id="4C2869C90061FFD0628DF827FC4778B3" authority="H. Perrier" authorityName="H. Perrier" box="[643,982,1929,1955]" class="Insecta" family="Tachinidae" genus="Eugenia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="arenicola">
<emphasis id="B95CCE580061FFD0628DF827FCCB78B2" box="[643,858,1929,1954]" italics="true" pageId="8" pageNumber="47">Eugenia arenicola</emphasis>
H. Perrier
</taxonomicName>
, another species from littoral forests in the general region.
</paragraph>
<caption id="DF5742C20060FFD16099F9C2FE8B79AC" ID-DOI="http://doi.org/10.5281/zenodo.4776784" ID-Zenodo-Dep="4776784" httpUri="https://zenodo.org/record/4776784/files/figure.png" pageId="9" pageNumber="48" startId="9.[151,244,1644,1665]" targetBox="[151,1436,181,1622]" targetPageId="9">
<paragraph id="8B97124A0060FFD16099F9C2FE8B79AC" blockId="9.[151,1436,1644,1724]" pageId="9" pageNumber="48">
<emphasis id="B95CCE580060FFD16099F9C2FE9D7991" bold="true" box="[151,268,1644,1665]" pageId="9" pageNumber="48">FIGURE 5.</emphasis>
<taxonomicName id="4C2869C90060FFD1611FF9C2FE5E7991" box="[273,463,1644,1665]" class="Insecta" family="Tachinidae" genus="Eugenia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="petrikensis">
<emphasis id="B95CCE580060FFD1611FF9C2FE5E7991" box="[273,463,1644,1665]" italics="true" pageId="9" pageNumber="48">Eugenia petrikensis</emphasis>
</taxonomicName>
. Upper left: bark of main bole. Upper right: sessile to subsessile inflorescences. Lower left: emerging leaves with whitish indumentum. Lower right: mature leaves; abaxial (upper) and adaxial (lower) surfaces. (No voucher; photos by D. Rabehevitra).
</paragraph>
</caption>
<paragraph id="8B97124A0060FFD160C8F8BFFACF78E0" blockId="9.[151,1436,1809,2032]" pageId="9" pageNumber="48">
<emphasis id="B95CCE580060FFD160C8F8BFFE5C783B" bold="true" box="[198,461,1809,1835]" pageId="9" pageNumber="48">Conservation status:</emphasis>
―The type gathering is in or near the eastern part of conservation zone P
<quantity id="4CD0BFAF0060FFD16563F8BCFA0D783C" box="[1389,1436,1810,1836]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="9" pageNumber="48" unit="in" value="1.0">1 in</quantity>
Petriky Forest (
<figureCitation id="13130ECF0060FFD16145F896FE057842" box="[331,404,1848,1874]" captionStart="FIGURE 2" captionStartId="2.[151,244,1785,1806]" captionTargetBox="[166,1430,868,1771]" captionTargetId="figure-397@2.[166,1431,868,1771]" captionTargetPageId="2" captionText="FIGURE 2. Distribution of species and selected specimens of Eugenia in southeastern Madagascar based on vouchered herbarium specimens and recently captured (but non-vouchered) digital images. Remaining forested areas (Petriky, Mandena, Sainte-Luce) along the coast are indicated in gray. In cases where the icons largely or completely overlapped based on geo-coordinates they have been separated slightly for easier viewing (e.g., E. ardyceae, E. stictophylla and E. cloiselii along the coast west of Tolagnaro). Many additional specimens of Eugenia from the region await identification or description and cannot be mapped confidently at the present time." figureDoi="http://doi.org/10.5281/zenodo.4776772" httpUri="https://zenodo.org/record/4776772/files/figure.png" pageId="9" pageNumber="48">Fig. 2</figureCitation>
; see also
<bibRefCitation id="EFB96FBB0060FFD1620DF896FD487842" author="Lowry, P. P. II &amp; Randriatafika, F. &amp; Rabenantoandro, J." box="[515,729,1848,1874]" pageId="9" pageNumber="48" pagination="55 - 63" refId="ref11601" refString="Lowry, P. P. II, Randriatafika, F., &amp; Rabenantoandro, J. (2008) Conservation status of vascular plant species from the QMM / Rio Tinto mining area at Mandena, Tolagnaro (Fort Dauphin) region, southeast Madagascar. Madagascar Conservation and Development 3: 55 - 63." type="journal article" year="2008">
Lowry
<emphasis id="B95CCE580060FFD16256F897FD027842" box="[600,659,1849,1874]" italics="true" pageId="9" pageNumber="48">et al.</emphasis>
2008
</bibRefCitation>
). The
<typeStatus id="5493ACE80060FFD1632CF896FC167842" box="[802,903,1848,1874]" pageId="9" pageNumber="48" type="paratype">paratype</typeStatus>
(
<figureCitation id="13130ECF0060FFD16398F896FC4D7842" box="[918,988,1848,1874]" captionStart="FIGURE 5" captionStartId="9.[151,244,1644,1665]" captionTargetBox="[151,1436,181,1622]" captionTargetId="figure-162@9.[151,1436,181,1622]" captionTargetPageId="9" captionText="FIGURE 5. Eugenia petrikensis. Upper left: bark of main bole. Upper right: sessile to subsessile inflorescences. Lower left: emerging leaves with whitish indumentum. Lower right: mature leaves; abaxial (upper) and adaxial (lower) surfaces. (No voucher; photos by D. Rabehevitra)." figureDoi="http://doi.org/10.5281/zenodo.4776784" httpUri="https://zenodo.org/record/4776784/files/figure.png" pageId="9" pageNumber="48">Fig. 5</figureCitation>
) was from a non-protected and highly disturbed area. Although herbarium material of
<taxonomicName id="4C2869C90060FFD162D4F8F1FC567868" box="[730,967,1887,1912]" class="Insecta" family="Tachinidae" genus="Eugenia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="petrikensis">
<emphasis id="B95CCE580060FFD162D4F8F1FC567868" box="[730,967,1887,1912]" italics="true" pageId="9" pageNumber="48">Eugenia petrikensis</emphasis>
</taxonomicName>
currently is known only from the type gathering, the species has been observed and photographed recently at Ambinanibe Forest (25°02’35’S,
<geoCoordinate id="EE1C748D0060FFD16099F801FEB478D9" box="[151,293,1967,1993]" degrees="46" direction="east" minutes="56" orientation="longitude" pageId="9" pageNumber="48" precision="15" seconds="04" value="46.934444">46°56’04”E</geoCoordinate>
) growing in full sun or rarely in shade. With an AOO of
<quantity id="4CD0BFAF0060FFD163C1F801FB8878D9" box="[975,1049,1967,1993]" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="9" pageNumber="48" unit="km" value="18.0">18 km</quantity>
<superScript id="7C5DBF020060FFD16414F802FBB378AA" attach="right" box="[1050,1058,1964,1978]" fontSize="6" pageId="9" pageNumber="48">2</superScript>
, and 2 subpopulations, one in a protected area (Petriky),
<taxonomicName id="4C2869C90060FFD161B4F878FDCB78FF" box="[442,602,2006,2032]" class="Insecta" family="Tachinidae" genus="Eugenia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="petrikensis">
<emphasis id="B95CCE580060FFD161B4F878FE5D78FF" box="[442,460,2006,2031]" italics="true" pageId="9" pageNumber="48">E</emphasis>
.
<emphasis id="B95CCE580060FFD161D5F878FDCB78FF" box="[475,602,2006,2031]" italics="true" pageId="9" pageNumber="48">petrikensis</emphasis>
</taxonomicName>
is assigned a preliminary status of “Endangered” (EN: B2ab[iii]).
</paragraph>
</subSubSection>
<subSubSection id="C33241C10063FFD260C8FF39FC257DB8" pageId="10" pageNumber="49" type="discussion">
<paragraph id="8B97124A0063FFD260C8FF39FEF07D92" blockId="10.[151,1436,151,680]" pageId="10" pageNumber="49">
<emphasis id="B95CCE580063FFD260C8FF39FEC57FA1" bold="true" box="[198,340,151,177]" pageId="10" pageNumber="49">Discussion:</emphasis>
―
<taxonomicName id="4C2869C90063FFD26171FF36FDFC7FA1" box="[383,621,152,177]" class="Insecta" family="Tachinidae" genus="Eugenia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="petrikensis">
<emphasis id="B95CCE580063FFD26171FF36FDFC7FA1" box="[383,621,152,177]" italics="true" pageId="10" pageNumber="49">Eugenia petrikensis</emphasis>
</taxonomicName>
may be more common than suggested by the known number of herbarium specimens. Its vegetative and floral morphologies most closely resemble those of
<taxonomicName id="4C2869C90063FFD264E4FF11FEEA7FEF" authority="N. Snow" authorityName="N. Snow" authorityYear="2011" class="Insecta" family="Tachinidae" genus="Eugenia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="ardyceae">
<emphasis id="B95CCE580063FFD264E4FF11FAE67FC8" box="[1258,1399,191,216]" italics="true" pageId="10" pageNumber="49">E. ardyceae</emphasis>
N. Snow (
<bibRefCitation id="EFB96FBB0063FFD260E7FF4BFEE37FEF" author="Snow, N." box="[233,370,229,255]" pageId="10" pageNumber="49" pagination="677 - 689" refId="ref12071" refString="Snow, N. (2011) Studies of Malagasy Eugenia (Myrtaceae) - II: Four new species, including one eaten by black lemurs on Nosy Be. Systematic Botany 36 (3): 677 - 689." type="journal article" year="2011">Snow 2011</bibRefCitation>
)
</taxonomicName>
, which also occurs in the Tolagnaro region (
<figureCitation id="13130ECF0063FFD2638AFF4BFC5D7FEF" box="[900,972,229,255]" captionStart="FIGURE 2" captionStartId="2.[151,244,1785,1806]" captionTargetBox="[166,1430,868,1771]" captionTargetId="figure-397@2.[166,1431,868,1771]" captionTargetPageId="2" captionText="FIGURE 2. Distribution of species and selected specimens of Eugenia in southeastern Madagascar based on vouchered herbarium specimens and recently captured (but non-vouchered) digital images. Remaining forested areas (Petriky, Mandena, Sainte-Luce) along the coast are indicated in gray. In cases where the icons largely or completely overlapped based on geo-coordinates they have been separated slightly for easier viewing (e.g., E. ardyceae, E. stictophylla and E. cloiselii along the coast west of Tolagnaro). Many additional specimens of Eugenia from the region await identification or description and cannot be mapped confidently at the present time." figureDoi="http://doi.org/10.5281/zenodo.4776772" httpUri="https://zenodo.org/record/4776772/files/figure.png" pageId="10" pageNumber="49">Fig. 2</figureCitation>
). However, the pedicillate flowers with glabrous hypanthia of
<emphasis id="B95CCE580063FFD261A5FEA2FDC07E35" box="[427,593,268,293]" italics="true" pageId="10" pageNumber="49">
R.
<taxonomicName id="4C2869C90063FFD261C3FEA2FDC07E35" box="[461,593,268,293]" class="Insecta" family="Tachinidae" genus="Eugenia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="petrikensis">petrikensis</taxonomicName>
</emphasis>
contrast with the sessile to subsessile flowers with sericeous-villous hypanthia of
<taxonomicName id="4C2869C90063FFD26148FE9CFE717E5B" authorityName="N. Snow" authorityYear="2011" box="[326,480,306,331]" class="Insecta" family="Tachinidae" genus="Eugenia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="ardyceae">
<emphasis id="B95CCE580063FFD26148FE9CFE717E5B" box="[326,480,306,331]" italics="true" pageId="10" pageNumber="49">E. ardyceae</emphasis>
</taxonomicName>
. The dibrachiate trichomes of
<taxonomicName id="4C2869C90063FFD26374FE9CFB857E5B" authorityName="N. Snow" authorityYear="2011" box="[890,1044,306,331]" class="Insecta" family="Tachinidae" genus="Eugenia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="ardyceae">
<emphasis id="B95CCE580063FFD26374FE9CFB857E5B" box="[890,1044,306,331]" italics="true" pageId="10" pageNumber="49">E. ardyceae</emphasis>
</taxonomicName>
on younger leaves are highly asymmetrical (i.e., arms of differing lengths [
<bibRefCitation id="EFB96FBB0063FFD262DAFEF7FCF87E63" author="Snow, N." box="[724,873,345,371]" pageId="10" pageNumber="49" pagination="677 - 689" refId="ref12071" refString="Snow, N. (2011) Studies of Malagasy Eugenia (Myrtaceae) - II: Four new species, including one eaten by black lemurs on Nosy Be. Systematic Botany 36 (3): 677 - 689." type="journal article" year="2011">Snow 2011</bibRefCitation>
]), which contrast with the more symmetrical trichomes in the emerging foliage of
<taxonomicName id="4C2869C90063FFD26258FE2EFD687E89" box="[598,761,384,409]" class="Insecta" family="Tachinidae" genus="Eugenia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="petrikensis">
<emphasis id="B95CCE580063FFD26258FE2EFD687E89" box="[598,761,384,409]" italics="true" pageId="10" pageNumber="49">E. petrikensis</emphasis>
</taxonomicName>
. The calyx lobes on the
<typeStatus id="5493ACE80063FFD2642DFE2EFBC77E8A" box="[1059,1110,384,410]" pageId="10" pageNumber="49">type</typeStatus>
specimen of
<taxonomicName id="4C2869C90063FFD264F7FE2EFA0D7E89" box="[1273,1436,384,409]" class="Insecta" family="Tachinidae" genus="Eugenia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="petrikensis">
<emphasis id="B95CCE580063FFD264F7FE2EFA0D7E89" box="[1273,1436,384,409]" italics="true" pageId="10" pageNumber="49">E. petrikensis</emphasis>
</taxonomicName>
have 3 straight and equally-spaced vascular bundles that converge apically slightly inside of the margin (magnification required), but it is uncertain if this trait occurs in all specimens.The leaves, inflorescences, and flowers of
<taxonomicName id="4C2869C90063FFD26112FE5AFD727D1D" box="[284,739,500,526]" class="Insecta" family="Tachinidae" genus="Eugenia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="49" phylum="Arthropoda" rank="variety" species="urschaiana" variety="urschiana">
<emphasis id="B95CCE580063FFD26112FE5AFE547D1D" box="[284,453,500,525]" italics="true" pageId="10" pageNumber="49">E. urschaiana</emphasis>
H. Perr. var.
<emphasis id="B95CCE580063FFD26265FE5AFD727D1D" box="[619,739,500,525]" italics="true" pageId="10" pageNumber="49">urschiana</emphasis>
</taxonomicName>
also somewhat resemble those of
<taxonomicName id="4C2869C90063FFD26489FE5AFABC7D1D" box="[1159,1325,500,525]" class="Insecta" family="Tachinidae" genus="Eugenia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="petrikensis">
<emphasis id="B95CCE580063FFD26489FE5AFABC7D1D" box="[1159,1325,500,525]" italics="true" pageId="10" pageNumber="49">E. petrikensis</emphasis>
</taxonomicName>
(e.g.,
<emphasis id="B95CCE580063FFD26572FE5AFE987D23" italics="true" pageId="10" pageNumber="49">M. Ursch 17</emphasis>
, holo: P! [image seen]). However,
<taxonomicName id="4C2869C90063FFD262C8FDB4FBA07D23" box="[710,1073,538,564]" class="Insecta" family="Tachinidae" genus="Eugenia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="49" phylum="Arthropoda" rank="variety" species="urschaiana" variety="urschiana">
<emphasis id="B95CCE580063FFD262C8FDB4FCE47D23" box="[710,885,538,563]" italics="true" pageId="10" pageNumber="49">E. urschaiana</emphasis>
var.
<emphasis id="B95CCE580063FFD263BBFDB4FBA07D23" box="[949,1073,538,563]" italics="true" pageId="10" pageNumber="49">urschiana</emphasis>
</taxonomicName>
occurs ca.
<quantity id="4CD0BFAF0063FFD264CFFDB4FABF7D24" box="[1217,1326,538,564]" metricMagnitude="6" metricUnit="m" metricValue="1.3" pageId="10" pageNumber="49" unit="km" value="1300.0">1300 km</quantity>
north of Tolagnaro at ca.
<specimenCount id="9D2ED9C30063FFD26155FDEFFE3C7D4B" box="[347,429,577,603]" pageId="10" pageNumber="49" type="generic">
<quantity id="4CD0BFAF0063FFD26155FDEFFE397D4B" box="[347,424,577,603]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="49" unit="m" value="250.0">250 m</quantity>
.
</specimenCount>
elevation in forests at or near Analamazaotra (
<bibRefCitation id="EFB96FBB0063FFD263DDFDEFFBE97D4B" author="Perrier de la Bathie, H." box="[979,1144,577,603]" pageId="10" pageNumber="49" pagination="161 - 202" refId="ref11754" refString="Perrier de la Bathie, H. (1953 a) Les Myrtacees de Madagascar et des Comores, revision, diagnoses et biologie. Memoires de L' Institut Scientifiique de Madagascar, B, 4 (2): 161 - 202." type="journal article" year="1953">Perrier 1953a</bibRefCitation>
, b) (&quot;Analamazoatra&quot; on herbarium label).
</paragraph>
<paragraph id="8B97124A0063FFD260C8FD20FC257DB8" blockId="10.[151,1436,151,680]" box="[198,948,654,680]" pageId="10" pageNumber="49">More detailed field surveys are urgently needed for this species.</paragraph>
</subSubSection>
</treatment>
</document>
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