treatments-xml/data/03/81/40/0381406C3614484CFF67032FE7A9FA82.xml

<document id="457E612C5AA1349018F4CA618186D290" ID-DOI="10.5281/zenodo.19105" ID-GBIF-Dataset="6c1840f0-100f-42d2-9c1a-c28eee124991" checkinTime="1435582000948" checkinUser="donat" docAuthor="Fernando AMOR &amp; Patrocinio ORTEGA" docDate="2014" docId="0381406C3614484CFF67032FE7A9FA82" docLanguage="en" docName="27641.pdf" docOrigin="Myrmecological News 19" docTitle="Cataglyphis tartessica AMOR &amp; ORTEGA, 2014, sp.n." docType="treatment" docVersion="6" lastPageNumber="130" masterDocId="FFB838143615484AFFF00460E550FFE3" masterDocTitle="Cataglyphis tartessica sp. n., a new ant species (Hymenoptera: Formicidae) in southwestern Spain" masterLastPageNumber="132" masterPageNumber="125" pageNumber="126" updateTime="1695866497708" updateUser="admin">
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<mods:title id="DAA35597635ACA7B95F095F7C5D3BA35">Cataglyphis tartessica sp. n., a new ant species (Hymenoptera: Formicidae) in southwestern Spain</mods:title>
</mods:titleInfo>
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<mods:namePart id="1B90C96E3C1C0E36278498DDAADFDDF2">Fernando AMOR</mods:namePart>
</mods:name>
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<mods:namePart id="2ED3D84B66816F39340B17E749BC8EB9">Patrocinio ORTEGA</mods:namePart>
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<mods:title id="1E369EE40F4C63CB36C6AA502E091AE5">Myrmecological News</mods:title>
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<mods:part id="C122BA1AA9746E8D8BC7E08666B2E3A1">
<mods:date id="FBE2D11EB5B545DF304378CF375BFF8D">2014</mods:date>
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<treatment id="0381406C3614484CFF67032FE7A9FA82" ID-DOI="http://doi.org/10.5281/zenodo.6134611" ID-GBIF-Taxon="114923734" ID-Zenodo-Dep="6134611" LSID="urn:lsid:plazi:treatment:0381406C3614484CFF67032FE7A9FA82" httpUri="http://treatment.plazi.org/id/0381406C3614484CFF67032FE7A9FA82" lastPageId="6" lastPageNumber="130" pageId="1" pageNumber="126">
<subSubSection id="C332A2F13614484BFF67032FE493F884" box="[151,451,1871,1895]" pageId="1" pageNumber="126" type="nomenclature">
<paragraph id="8B97F17A3614484BFF67032FE493F884" box="[151,451,1871,1895]" pageId="1" pageNumber="126">
<heading id="D0DF46163614484BFF67032FE493F884" bold="true" box="[151,451,1871,1895]" fontSize="10" level="1" pageId="1" pageNumber="126" reason="6">
<emphasis id="B95C2D683614484BFF67032FE493F884" bold="true" box="[151,451,1871,1895]" pageId="1" pageNumber="126">
<taxonomicName id="4C288AF93614484BFF67032FE4D7F884" box="[151,391,1871,1895]" class="Insecta" family="Formicidae" genus="Cataglyphis" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="126" phylum="Arthropoda" rank="species" species="tartessica" status="sp.n.">
<emphasis id="B95C2D683614484BFF67032FE4D7F884" bold="true" box="[151,391,1871,1895]" italics="true" pageId="1" pageNumber="126">Cataglyphis tartessica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F90133614484BFE7E032FE493F884" box="[398,451,1871,1895]" pageId="1" pageNumber="126">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C332A2F13614484BFF67031AE03DFA3A" pageId="1" pageNumber="126" type="materials_examined">
<paragraph id="8B97F17A3614484BFF67031AE03DFA3A" pageId="1" pageNumber="126">
<emphasis id="708C893F2077534A4723F7F7768FD400" bold="true" box="[151,435,1914,1938]" pageId="1" pageNumber="126">Type material examined:</emphasis>
<materialsCitation id="183176349CB90873D57D3F068470DE78" ID-GBIF-Occurrence="1226907227" collectingDate="2014-01-01" collectingDateMax="2014-12-31" collectingDateMin="2014-01-01" collectionCode="MNCN" country="Spain" latitude="36.99401" location="Doñana Biological Reserve" longLatPrecision="1" longitude="-6.4465947" pageId="1" pageNumber="126" specimenCode="Ent75407" specimenCount="1" specimenCount-worker="1" stateProvince="Huelva" typeStatus="holotype">
<typeStatus id="54934FD83614484BFE4B031AE714F872" box="[443,580,1913,1937]" pageId="1" pageNumber="126">Holotype</typeStatus>
(
<specimenCount id="9D2E3AF33614484BFDAA0319E7FAF872" box="[602,682,1913,1937]" pageId="1" pageNumber="126" type="worker">worker</specimenCount>
): 
<collectingCountry id="F33FB1EA3614484BFD4D0319E7ABF872" box="[701,763,1913,1937]" name="Spain" pageId="1" pageNumber="126" value="Spain">Spain</collectingCountry>
, 
<collectingRegion id="0D32BBC3989FBF2A8F761D4DF43E0339">Huelva</collectingRegion>
, 
<location id="02E857EA89F7D1B41E5B5E8903A02A7B" LSID="urn:lsid:plazi:treatment:0381406C3614484CFF67032FE7A9FA82:02E857EA89F7D1B41E5B5E8903A02A7B" country="Spain" latitude="36.99401" longLatPrecision="1" longitude="-6.4465947" name="Doñana Biological Reserve" stateProvince="Huelva">Doñana Biological Reserve</location>
<geoCoordinate id="EE1C97BD3614484BFDFC03F7E7E9F84C" box="[524,697,1943,1967]" direction="north" orientation="latitude" pageId="1" pageNumber="126" precision="1" value="36.99401">36° 59' 38.44&quot; N</geoCoordinate>
, 
<geoCoordinate id="EE1C97BD3614484BFD3403F7E451F82F" direction="west" orientation="longitude" pageId="1" pageNumber="126" precision="1" value="-6.4465947">6° 26' 47.74&quot; W</geoCoordinate>
, 
<collectingDate id="EFD22E523614484BFEFC03D4E4C0F82F" box="[268,400,1972,1996]" pageId="1" pageNumber="126">
<date id="CEEE09D68D851981A2B28E6424F17510" value="2009-03-07">2009-07-03</date>
;
</collectingDate>
coll. Museo Nacional de Ciencias Naturales de Madrid (
<collectionCode id="ED3969BF3614484BFE7403B1E487F80A" box="[388,471,2001,2025]" collectionName="MNCN" pageId="1" pageNumber="126">MNCN</collectionCode>
- 
<specimenCode id="DB8E59013614484BFE1503B1E71EF80A" box="[485,590,2001,2025]" pageId="1" pageNumber="126">Ent75407</specimenCode>
)
</materialsCitation>
. 
<materialsCitation id="3B40FB273614484BFD9003B2E039FA3A" ID-GBIF-Occurrence="1226907228" collectingDate="2008" collectingDateMax="2010" collectingDateMin="2008" collectionCode="DBR, ALC, AZN, UTR, BRN" country="Spain" pageId="1" pageNumber="126" specimenCount="355" specimenCount-male="30" specimenCount-queen="72" specimenCount-worker="181" stateProvince="Madrid" typeStatus="paratype">
<typeStatus id="54934FD83614484BFD9003B2E7BEF80A" box="[608,750,2002,2025]" pageId="1" pageNumber="126">Paratypes</typeStatus>
: 
<specimenCount id="9D2E3AF33614484BFCC401E7E6F0FA7C" box="[820,928,1415,1439]" pageId="1" pageNumber="126" type="queen">72 queens</specimenCount>
(
<specimenCount id="9D2E3AF33614484BFC4101E6E138FA7C" box="[945,1128,1414,1439]" pageId="1" pageNumber="126">25 brachypterous</specimenCount>
and 
<specimenCount id="9D2E3AF33614484BFB5101E7E076FA7C" box="[1185,1318,1414,1439]" pageId="1" pageNumber="126">47 ergatoids</specimenCount>
; 
<collectionCode id="ED3969BF3614484BFAC401E6E03DFA7C" box="[1332,1389,1414,1439]" collectionName="DBR" pageId="1" pageNumber="126">DBR</collectionCode>
), 
<specimenCount id="9D2E3AF33614484BFA7201E7E624FA5F" pageId="1" pageNumber="126" type="male">30 males</specimenCount>
(
<collectionCode id="ED3969BF3614484BFC7501C4E6EEFA5F" box="[901,958,1444,1468]" collectionName="DBR" pageId="1" pageNumber="126">DBR</collectionCode>
and 
<collectionCode id="ED3969BF3614484BFC0801C4E162FA5F" box="[1016,1074,1444,1468]" collectionName="ALC" pageId="1" pageNumber="126">ALC</collectionCode>
), and 
<specimenCount id="9D2E3AF33614484BFB8801C4E053FA5F" box="[1144,1283,1444,1468]" pageId="1" pageNumber="126" type="worker">181 workers</specimenCount>
(
<collectionCode id="ED3969BF3614484BFAE401C4E000FA5F" box="[1300,1360,1444,1468]" collectionName="DBR" pageId="1" pageNumber="126">DBR</collectionCode>
, 
<collectionCode id="ED3969BF3614484BFAAD01C4E0C8FA5F" box="[1373,1432,1444,1468]" collectionName="AZN" pageId="1" pageNumber="126">AZN</collectionCode>
, 
<collectionCode id="ED3969BF3614484BFCC401A2E63CFA3A" box="[820,876,1474,1497]" collectionName="ALC" pageId="1" pageNumber="126">ALC</collectionCode>
, 
<collectionCode id="ED3969BF3614484BFC8701A2E6FCFA3A" box="[887,940,1474,1497]" collectionName="Russia, Tomsk, University of Tomsk" pageId="1" pageNumber="126">UTR</collectionCode>
, 
<collectionCode id="ED3969BF3614484BFC4901A1E6A2FA3A" box="[953,1010,1473,1497]" collectionName="BRN" pageId="1" pageNumber="126">BRN</collectionCode>
) collected between 
<collectingDate id="EFD22E523614484BFB3601A2E039FA3A" box="[1222,1385,1473,1497]" pageId="1" pageNumber="126" value="2008" valueMax="2010">
<date id="FF96D7BA3614484BFB3601A2E039FA3A" box="[1222,1385,1473,1497]" pageId="1" pageNumber="126" value="2008" valueMax="2010">2008 and 2010</date>
</collectingDate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C332A2F13614484EFCAA01BFE419F812" lastPageId="4" lastPageNumber="129" pageId="1" pageNumber="126" type="description">
<paragraph id="8B97F17A36144848FCAA01BFE7AFFA41" lastPageId="2" lastPageNumber="127" pageId="1" pageNumber="126">
<emphasis id="B95C2D683614484BFCAA01BFE192FA14" bold="true" box="[858,1218,1503,1527]" pageId="1" pageNumber="126">
Description of 
<typeStatus id="54934FD83614484BFBF001BFE130FA14" box="[1024,1120,1503,1527]" pageId="1" pageNumber="126">holotype</typeStatus>
worker:
</emphasis>
HL: 
<quantity id="4CD05C9F3614484BFB0C01BFE00DFA15" box="[1276,1373,1503,1526]" metricMagnitude="-1" metricUnit="m" metricValue="1.17" pageId="1" pageNumber="126" unit="mm" value="117.0">1.17 mm</quantity>
, HW: 
<quantity id="4CD05C9F3614484BFCC4019CE6C6F9F7" box="[820,918,1532,1556]" metricMagnitude="-1" metricUnit="m" metricValue="1.13" pageId="1" pageNumber="126" unit="mm" value="113.0">1.13 mm</quantity>
, SL: 
<quantity id="4CD05C9F3614484BFC3E019CE160F9F7" box="[974,1072,1532,1556]" metricMagnitude="-1" metricUnit="m" metricValue="1.42" pageId="1" pageNumber="126" unit="mm" value="142.0">1.42 mm</quantity>
, ML: 
<quantity id="4CD05C9F3614484BFB82019CE184F9F7" box="[1138,1236,1532,1556]" metricMagnitude="-1" metricUnit="m" metricValue="1.9" pageId="1" pageNumber="126" unit="mm" value="190.0">1.90 mm</quantity>
, FL: 
<quantity id="4CD05C9F3614484BFAFD019CE03FF9F7" box="[1293,1391,1532,1556]" metricMagnitude="-1" metricUnit="m" metricValue="1.85" pageId="1" pageNumber="126" unit="mm" value="185.0">1.85 mm</quantity>
, CI: 96.58, SI: 125.66. Bicoloured: shiny black gaster, black eyes and ocelli, rest of body amber. Integument densely microsculptured, matte, except on slightly shiny gaster. Abundant pubescence not obscuring integument texture. Head rectangular; vertex smooth and rounded, with two to three suberect hairs. Compound eyes large and ovoid located in upper half of face sides; ocelli normal. Mandibles triangular and slightly striated. Pilosity particularly present on outer face and on or near margins; long and posteriorly curved or J-shaped hairs in external margin particularly notable. Masticatory margin armed with 6 teeth; apical tooth largest and most pointed, preapical tooth of intermediate length, and remaining teeth small and irregular. Clypeus with ridge in centre and 7 stout setae along front edge; one pair of setae straddling posterior clypeus ridge. Posterior (basal) area of maxillae with six long and anteriorly curved Tab. 1: Measurements in millimetres of 
<taxonomicName id="4C288AF936174848FDB201EAE796FA41" box="[578,710,1418,1442]" class="Insecta" family="Formicidae" genus="Cataglyphis" genus.bestMatchDistance="11" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="species" species="tartessica" status="sp.n.">
<emphasis id="B95C2D6836174848FDB201EAE796FA41" box="[578,710,1418,1442]" italics="true" pageId="2" pageNumber="127">C. tartessica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F901336174848FD3E01EAE7AFFA41" box="[718,767,1418,1442]" pageId="2" pageNumber="127">sp.n.</taxonomicNameLabel>
</paragraph>
<caption id="DF57A1F236174848FBB80081E0CBFAB1" pageId="2" pageNumber="127">
<paragraph id="8B97F17A36174848FBB80081E0CBFAB1" pageId="2" pageNumber="127">
Fig. 2: 
<taxonomicName id="4C288AF936174848FB620081E028FB19" box="[1170,1400,1249,1274]" class="Insecta" family="Formicidae" genus="Cataglyphis" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="species" species="tartessica" status="sp. n.">
<emphasis id="B95C2D6836174848FB620081E028FB19" box="[1170,1400,1249,1274]" italics="true" pageId="2" pageNumber="127">Cataglyphis tartessica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F901336174848FA8F0082E10CFAF4" pageId="2" pageNumber="127" rank="species">sp. n.</taxonomicNameLabel>
(a) Brachypterous queen. (b) Ergatoid queen (minute wing insertion indicated with an arrow).
</paragraph>
</caption>
<paragraph id="8B97F17A36174848FF6701C6E40EFA5C" box="[151,350,1446,1471]" pageId="2" pageNumber="127">workers (n = 181).</paragraph>
<paragraph id="8B97F17A36174848FF5401BDE4A9FA17" box="[164,505,1501,1524]" pageId="2" pageNumber="127">
<emphasis id="B95C2D6836174848FF5401BDE4A9FA17" bold="true" box="[164,505,1501,1524]" pageId="2" pageNumber="127">
<taxonomicName id="4C288AF936174848FF5401BDE4C5FA17" box="[164,405,1501,1524]" class="Insecta" family="Formicidae" genus="Cataglyphis" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="species" species="tartessica">
<emphasis id="B95C2D6836174848FF5401BDE4C5FA17" bold="true" box="[164,405,1501,1524]" italics="true" pageId="2" pageNumber="127">Cataglyphis tartessica</emphasis>
</taxonomicName>
workers
</emphasis>
</paragraph>
<paragraph id="8B97F17A36174848FF540269E792F8BD" pageId="2" pageNumber="127">
<table id="F92803DA36174848FF540269E792F8BD" box="[164,706,1545,1886]" pageId="2" pageNumber="127">
<tr id="FFB8366A36174848FF540269E7FDF9C2" box="[164,706,1545,1569]" pageId="2" pageNumber="127">
<td id="FFB8366436174848FF540269E450F9C2" box="[164,256,1545,1569]" pageId="2" pageNumber="127">Variable</td>
<td id="FFB8366436174848FDC6026AE7FDF9C2" box="[566,685,1546,1569]" pageId="2" pageNumber="127">mean ± SD</td>
</tr>
<tr id="FFB8366A36174848FF540256E7E5F9AC" box="[164,706,1590,1615]" pageId="2" pageNumber="127">
<td id="FFB8366436174848FF540256E476F9AC" box="[164,294,1590,1615]" pageId="2" pageNumber="127">Head length</td>
<td id="FFB8366436174848FDDE0257E7E5F9AC" box="[558,693,1591,1615]" pageId="2" pageNumber="127">11.23 ± 0.12</td>
</tr>
<tr id="FFB8366A36174848FF540204E7E5F99F" box="[164,706,1636,1660]" pageId="2" pageNumber="127">
<td id="FFB8366436174848FF540204E470F99F" box="[164,288,1636,1660]" pageId="2" pageNumber="127">Head width</td>
<td id="FFB8366436174848FDDE0204E7E5F99F" box="[558,693,1636,1660]" pageId="2" pageNumber="127">11.15 ± 0.12</td>
</tr>
<tr id="FFB8366A36174848FF5402F1E7E5F94A" box="[164,706,1681,1705]" pageId="2" pageNumber="127">
<td id="FFB8366436174848FF5402F1E47DF94A" box="[164,301,1681,1705]" pageId="2" pageNumber="127">Scape length</td>
<td id="FFB8366436174848FDDE02F1E7E5F94A" box="[558,693,1681,1705]" pageId="2" pageNumber="127">11.47 ± 0.12</td>
</tr>
<tr id="FFB8366A36174848FF5402DFE7E5F935" box="[164,706,1726,1750]" pageId="2" pageNumber="127">
<td id="FFB8366436174848FF5402DFE431F935" box="[164,353,1726,1750]" pageId="2" pageNumber="127">Mesosoma length</td>
<td id="FFB8366436174848FDDE02DEE7E5F935" box="[558,693,1726,1750]" pageId="2" pageNumber="127">11.94 ± 0.16</td>
</tr>
<tr id="FFB8366A36174848FF54028CE7E5F8E0" box="[164,706,1771,1795]" pageId="2" pageNumber="127">
<td id="FFB8366436174848FF54028CE463F8E0" box="[164,307,1771,1795]" pageId="2" pageNumber="127">Femur length</td>
<td id="FFB8366436174848FDDE028CE7E5F8E0" box="[558,693,1772,1795]" pageId="2" pageNumber="127">12.03 ± 0.14</td>
</tr>
<tr id="FFB8366A36174848FF540379E792F8D2" box="[164,706,1817,1841]" pageId="2" pageNumber="127">
<td id="FFB8366436174848FF540379E415F8D2" box="[164,325,1817,1841]" pageId="2" pageNumber="127">Cephalic index</td>
<td id="FFB8366436174848FDD10379E792F8D2" box="[545,706,1817,1841]" pageId="2" pageNumber="127">0 93.60 ± 3.940</td>
</tr>
<tr id="FFB8366A36174848FF540326E792F8BD" box="[164,706,1862,1886]" pageId="2" pageNumber="127">
<td id="FFB8366436174848FF540326E476F8BD" box="[164,294,1862,1886]" pageId="2" pageNumber="127">Scape index</td>
<td id="FFB8366436174848FDD10326E792F8BD" box="[545,706,1862,1886]" pageId="2" pageNumber="127">128.10 ± 7.660</td>
</tr>
</table>
</paragraph>
<paragraph id="8B97F17A36174848FF6703FCE754F80D" pageId="2" pageNumber="127">or J-shaped hairs. Sparse pubescence on genae and ventral cephalic capsule. Pubescence more abundant in occipital area between and behind the eyes.</paragraph>
<paragraph id="8B97F17A36174848FCAA01C7E6ADF98D" pageId="2" pageNumber="127">Mesosoma typical for this genus; propodeum evenly convex, long legs. Propodeal spiracle a long, narrow ellipse. Metapleural gland just above hind coxa; orifice protected by tuft of guard hairs. Scale-like petiole thick, with rounded apex. Sparse pubescence on mesonotum; abundant silver pubescence on pronotum, pleura, propodeum, petiole, and coxae.</paragraph>
<paragraph id="8B97F17A36174848FCAA0214E192F8E2" pageId="2" pageNumber="127">Gaster shiny due to a very lightly microsculptured integument and very sparse pubescence. Apex of gaster with circular acidopore (nozzle-shaped) and fringed with setae. Long setae on gastral sternites; only sparse setae present on dorsal side of most apical tergites.</paragraph>
<paragraph id="8B97F17A36174848FCAA0367E62EF80F" pageId="2" pageNumber="127">
<emphasis id="B95C2D6836174848FCAA0367E1E9F8FD" bold="true" box="[858,1209,1799,1823]" pageId="2" pageNumber="127">
Description of 
<typeStatus id="54934FD836174848FC090367E10AF8FC" box="[1017,1114,1799,1823]" pageId="2" pageNumber="127">paratype</typeStatus>
workers
</emphasis>
(Tab. 1, 
<figureCitation id="1313EDFF36174848FAE60366E000F8FD" box="[1302,1360,1798,1822]" captionStart="Fig. 1" captionStartId="1.[151,189,1236,1260]" captionText="Fig. 1: Cataglyphis tartessica sp. n., worker. (a) Body in profile. (b) Head in full-face view." pageId="2" pageNumber="127">Fig. 1</figureCitation>
): Workers from different sampling locations showed morphometric variation. The biggest difference in body size occurred between workers from 
<collectionCode id="ED3969BF36174848FB87033FE1FDF895" box="[1143,1197,1887,1910]" collectionName="BRN" pageId="2" pageNumber="127">BRN</collectionCode>
and 
<collectionCode id="ED3969BF36174848FB12033FE048F895" box="[1250,1304,1887,1910]" collectionName="AZN" pageId="2" pageNumber="127">AZN</collectionCode>
populations (
<collectionCode id="ED3969BF36174848FCCC031CE639F877" box="[828,873,1916,1940]" pageId="2" pageNumber="127">ML</collectionCode>
: 2.13 ± 0.2 vs. 1.77 ± 0.11, n = 35, from nest 5, and n = 30, from nest 6, respectively). Workers also differed, generally, in their number of setae and mandibular teeth (5 - 7).
</paragraph>
<paragraph id="8B97F17A36164849FF6704D1E1A4FF2A" box="[151,1268,177,202]" pageId="3" pageNumber="128">
Tab. 2: Measurements in millimetres of 
<taxonomicName id="4C288AF936164849FDB404D1E663FF29" box="[580,819,177,202]" class="Insecta" family="Formicidae" genus="Cataglyphis" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="tartessica" status="sp.n.">
<emphasis id="B95C2D6836164849FDB404D1E663FF29" box="[580,819,177,202]" italics="true" pageId="3" pageNumber="128">Cataglyphis tartessica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F901336164849FCCA04D2E63EFF2A" box="[826,878,178,201]" pageId="3" pageNumber="128">sp.n.</taxonomicNameLabel>
, brachypterous and ergatoid queens.
</paragraph>
<paragraph id="8B97F17A36164849FF520487E03FFD75" pageId="3" pageNumber="128">
<table id="F92803DA36164849FF520487E03FFD75" box="[162,1391,231,662]" pageId="3" pageNumber="128">
<tr id="FFB8366A36164849FF520487E05CFF1C" box="[162,1391,231,255]" pageId="3" pageNumber="128">
<emphasis id="B95C2D6836164849FF520487E05CFF1C" bold="true" box="[162,1292,231,255]" pageId="3" pageNumber="128">
<td id="FFB8366436164849FF520487E4B8FF1C" box="[162,488,231,255]" pageId="3" pageNumber="128">
<taxonomicName id="4C288AF936164849FF520487E4C3FF1C" box="[162,403,231,255]" class="Insecta" family="Formicidae" genus="Cataglyphis" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="tartessica">
<emphasis id="B95C2D6836164849FF520487E4C3FF1C" bold="true" box="[162,403,231,255]" italics="true" pageId="3" pageNumber="128">Cataglyphis tartessica</emphasis>
</taxonomicName>
queens
</td>
<td id="FFB8366436164849FD0A0488E6F3FF1C" box="[762,931,232,255]" pageId="3" pageNumber="128">Brachypterous</td>
<td id="FFB8366436164849FB570487E05CFF1C" box="[1191,1292,231,255]" pageId="3" pageNumber="128">Ergatoid</td>
</emphasis>
</tr>
<tr id="FFB8366A36164849FF520574E038FECF" box="[162,1391,276,300]" pageId="3" pageNumber="128">
<td id="FFB8366436164849FF520574E5AEFECF" box="[162,254,276,300]" pageId="3" pageNumber="128">Variable</td>
<td id="FFB8366436164849FD240575E61CFECF" box="[724,844,276,300]" pageId="3" pageNumber="128">mean ± SD</td>
<td id="FFB8366436164849FC230575E6B0FECF" box="[979,992,277,300]" pageId="3" pageNumber="128">n</td>
<td id="FFB8366436164849FBAB0575E182FECF" box="[1115,1234,276,300]" pageId="3" pageNumber="128">mean ± SD</td>
<td id="FFB8366436164849FAAA0575E038FECF" box="[1370,1384,277,300]" pageId="3" pageNumber="128">n</td>
</tr>
<tr id="FFB8366A36164849FF520521E03FFEBA" box="[162,1391,321,345]" pageId="3" pageNumber="128">
<td id="FFB8366436164849FF520521E474FEBA" box="[162,292,321,345]" pageId="3" pageNumber="128">Head length</td>
<td id="FFB8366436164849FD340522E609FEBA" box="[708,857,322,345]" pageId="3" pageNumber="128">111.26 ± 0.14</td>
<td id="FFB8366436164849FC2B0522E6B8FEBA" box="[987,1000,322,345]" pageId="3" pageNumber="128">7</td>
<td id="FFB8366436164849FBA70522E18EFEBA" box="[1111,1246,322,345]" pageId="3" pageNumber="128">11.20 ± 0.11</td>
<td id="FFB8366436164849FAA40522E03FFEBA" box="[1364,1391,322,345]" pageId="3" pageNumber="128">15</td>
</tr>
<tr id="FFB8366A36164849FF52050EE03FFE64" box="[162,1391,366,391]" pageId="3" pageNumber="128">
<td id="FFB8366436164849FF52050EE44EFE64" box="[162,286,366,391]" pageId="3" pageNumber="128">Head width</td>
<td id="FFB8366436164849FD34050FE609FE64" box="[708,857,367,391]" pageId="3" pageNumber="128">111.33 ± 0.09</td>
<td id="FFB8366436164849FC3E050FE6B9FE64" box="[974,1001,367,391]" pageId="3" pageNumber="128">24</td>
<td id="FFB8366436164849FBA7050FE18EFE64" box="[1111,1246,367,391]" pageId="3" pageNumber="128">11.13 ± 0.05</td>
<td id="FFB8366436164849FAA4050FE03FFE64" box="[1364,1391,367,391]" pageId="3" pageNumber="128">46</td>
</tr>
<tr id="FFB8366A36164849FF5205FCE03FFE57" box="[162,1391,411,436]" pageId="3" pageNumber="128">
<td id="FFB8366436164849FF5205FCE47BFE57" box="[162,299,411,436]" pageId="3" pageNumber="128">Scape length</td>
<td id="FFB8366436164849FD3405FCE609FE57" box="[708,857,412,436]" pageId="3" pageNumber="128">111.25 ± 0.07</td>
<td id="FFB8366436164849FC3E05FCE6B9FE57" box="[974,1001,412,436]" pageId="3" pageNumber="128">10</td>
<td id="FFB8366436164849FBA705FCE18EFE57" box="[1111,1246,412,436]" pageId="3" pageNumber="128">11.30 ± 0.04</td>
<td id="FFB8366436164849FAA405FCE03FFE57" box="[1364,1391,412,436]" pageId="3" pageNumber="128">15</td>
</tr>
<tr id="FFB8366A36164849FF5205A9E03FFE02" box="[162,1391,457,481]" pageId="3" pageNumber="128">
<td id="FFB8366436164849FF5205A9E40FFE02" box="[162,351,457,481]" pageId="3" pageNumber="128">Mesosoma length</td>
<td id="FFB8366436164849FD3405A9E609FE02" box="[708,857,457,481]" pageId="3" pageNumber="128">112.25 ± 0.16</td>
<td id="FFB8366436164849FC3E05A9E6B9FE02" box="[974,1001,457,481]" pageId="3" pageNumber="128">25</td>
<td id="FFB8366436164849FBA705A9E18EFE02" box="[1111,1246,457,481]" pageId="3" pageNumber="128">11.90 ± 0.11</td>
<td id="FFB8366436164849FAA405A9E03FFE02" box="[1364,1391,457,481]" pageId="3" pageNumber="128">47</td>
</tr>
<tr id="FFB8366A36164849FF520597E03FFDED" box="[162,1391,502,526]" pageId="3" pageNumber="128">
<td id="FFB8366436164849FF520597E435FDED" box="[162,357,502,526]" pageId="3" pageNumber="128">Mesonotum width</td>
<td id="FFB8366436164849FD340596E609FDED" box="[708,857,502,526]" pageId="3" pageNumber="128">110.86 ± 0.08</td>
<td id="FFB8366436164849FC3E0596E6B9FDED" box="[974,1001,502,526]" pageId="3" pageNumber="128">25</td>
<td id="FFB8366436164849FBA70596E18EFDED" box="[1111,1246,502,526]" pageId="3" pageNumber="128">10.52 ± 0.03</td>
<td id="FFB8366436164849FAA40596E03FFDED" box="[1364,1391,502,526]" pageId="3" pageNumber="128">47</td>
</tr>
<tr id="FFB8366A36164849FF520644E03FFDD8" box="[162,1391,547,571]" pageId="3" pageNumber="128">
<td id="FFB8366436164849FF520644E461FDD8" box="[162,305,547,571]" pageId="3" pageNumber="128">Femur length</td>
<td id="FFB8366436164849FD340644E609FDD8" box="[708,857,548,571]" pageId="3" pageNumber="128">111.64 ± 0.09</td>
<td id="FFB8366436164849FC3E0644E6B9FDD8" box="[974,1001,548,571]" pageId="3" pageNumber="128">23</td>
<td id="FFB8366436164849FBA70644E18EFDD8" box="[1111,1246,548,571]" pageId="3" pageNumber="128">11.75 ± 0.10</td>
<td id="FFB8366436164849FAA40644E03FFDD8" box="[1364,1391,548,571]" pageId="3" pageNumber="128">46</td>
</tr>
<tr id="FFB8366A36164849FF520630E03FFD8B" box="[162,1391,592,616]" pageId="3" pageNumber="128">
<td id="FFB8366436164849FF520630E413FD8B" box="[162,323,592,616]" pageId="3" pageNumber="128">Cephalic index</td>
<td id="FFB8366436164849FD340631E609FD8B" box="[708,857,593,616]" pageId="3" pageNumber="128">104.45 ± 13.4</td>
<td id="FFB8366436164849FC2B0631E6B8FD8B" box="[987,1000,593,616]" pageId="3" pageNumber="128">7</td>
<td id="FFB8366436164849FBB90631E1BBFD8B" box="[1097,1259,593,616]" pageId="3" pageNumber="128">194.89 ± 12.00</td>
<td id="FFB8366436164849FAA40631E03FFD8B" box="[1364,1391,593,616]" pageId="3" pageNumber="128">15</td>
</tr>
<tr id="FFB8366A36164849FF52061EE03FFD75" box="[162,1391,637,662]" pageId="3" pageNumber="128">
<td id="FFB8366436164849FF52061EE474FD75" box="[162,292,637,662]" pageId="3" pageNumber="128">Scape index</td>
<td id="FFB8366436164849FD34061EE608FD75" box="[708,856,638,662]" pageId="3" pageNumber="128">196.23 ± 5.60</td>
<td id="FFB8366436164849FC2B061EE6B8FD75" box="[987,1000,638,662]" pageId="3" pageNumber="128">7</td>
<td id="FFB8366436164849FBB9061EE1BBFD75" box="[1097,1259,638,662]" pageId="3" pageNumber="128">115.58 ± 4.700</td>
<td id="FFB8366436164849FAA4061EE03FFD75" box="[1364,1391,638,662]" pageId="3" pageNumber="128">15</td>
</tr>
</table>
</paragraph>
<paragraph id="8B97F17A36164849FF4D068CE7F9FCB8" pageId="3" pageNumber="128">
<emphasis id="B95C2D6836164849FF4D068CE491FCE0" bold="true" box="[189,449,748,771]" pageId="3" pageNumber="128">Description of queens</emphasis>
(Tab. 2, 
<figureCitation id="1313EDFF36164849FDC0068BE727FCE0" box="[560,631,747,771]" captionStart="Fig. 2" captionStartId="2.[1096,1133,1249,1274]" captionText="Fig. 2: Cataglyphis tartessica sp. n. (a) Brachypterous queen. (b) Ergatoid queen (minute wing insertion indicated with an arrow)." pageId="3" pageNumber="128">Fig. 2</figureCitation>
): A notable feature of 
<taxonomicName id="4C288AF936164849FEF80769E4C0FCC2" box="[264,400,777,801]" class="Insecta" family="Formicidae" genus="Cataglyphis" genus.bestMatchDistance="333" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="tartessica">
<emphasis id="B95C2D6836164849FEF80769E4C0FCC2" box="[264,400,777,801]" italics="true" pageId="3" pageNumber="128">C. tartessica</emphasis>
</taxonomicName>
is the presence of two morphs of queens, brachypterous (short-winged queen) and ergatoid (wingless queen), which are described separately.
</paragraph>
<paragraph id="8B97F17A36164849FF4D0701E499FAF0" pageId="3" pageNumber="128">
<emphasis id="B95C2D6836164849FF4D0701E729FC9A" bold="true" box="[189,633,865,889]" pageId="3" pageNumber="128">Description of brachypterous queens:</emphasis>
Same body colouration as worker. but with bigger head, mesosoma, and gaster; head squarer, scape shorter. Compound eyes and ocelli normal. Strong, slightly striated mandibles; apical tooth more pointed and developed than in worker, other teeth small and short. Mesosoma (scutum, scutellum, and metanotum) less developed than those seen in species with macropterous queens, but very different from those of ergatoids and workers (
<collectionCode id="ED3969BF36164849FE84002BE4E9FB80" box="[372,441,1099,1123]" pageId="3" pageNumber="128">AMOR</collectionCode>
&amp; al. 2011a). Short wings with weak venation; tegulae (scale-like sclerite covering forewing base) and wing stumps present and distinct. Legs shorter than those of ergatoids and workers (
<collectionCode id="ED3969BF36164849FDCD00C3E70FFB58" box="[573,607,1187,1211]" pageId="3" pageNumber="128">FL</collectionCode>
: 1.64, 1.75 and 2.03 respectively). Scale-like petiole large and thick, with rounded apex. Gaster similar but larger than that of worker. Pubescence very sparse.
</paragraph>
<paragraph id="8B97F17A36164849FCB1070BE009F9CA" pageId="3" pageNumber="128">
<table id="F92803DA36164849FCB1070BE009F9CA" box="[833,1396,875,1577]" pageId="3" pageNumber="128">
<tr id="FFB8366A36164849FCB1070BE00DFC60" box="[833,1396,875,899]" pageId="3" pageNumber="128">
<td id="FFB8366436164849FCB1070BE6CDFC60" box="[833,925,875,899]" pageId="3" pageNumber="128">Variable</td>
<td id="FFB8366436164849FB2B070CE00DFC60" box="[1243,1373,876,899]" pageId="3" pageNumber="128">means ± SD</td>
</tr>
<tr id="FFB8366A36164849FCB107F9E009FC52" box="[833,1396,921,945]" pageId="3" pageNumber="128">
<td id="FFB8366436164849FCB107F9E693FC52" box="[833,963,921,945]" pageId="3" pageNumber="128">Head length</td>
<td id="FFB8366436164849FB1007F9E009FC52" box="[1248,1369,921,945]" pageId="3" pageNumber="128">1.18 ± 0.07</td>
</tr>
<tr id="FFB8366A36164849FCB107A6E009FC3D" box="[833,1396,966,990]" pageId="3" pageNumber="128">
<td id="FFB8366436164849FCB107A6E6EDFC3D" box="[833,957,966,990]" pageId="3" pageNumber="128">Head width</td>
<td id="FFB8366436164849FB1007A6E009FC3D" box="[1248,1369,966,990]" pageId="3" pageNumber="128">1.29 ± 0.15</td>
</tr>
<tr id="FFB8366A36164849FCB10793E009FBE8" box="[833,1396,1011,1035]" pageId="3" pageNumber="128">
<td id="FFB8366436164849FCB10793E69AFBE8" box="[833,970,1011,1035]" pageId="3" pageNumber="128">Scape length</td>
<td id="FFB8366436164849FB100793E009FBE8" box="[1248,1369,1011,1035]" pageId="3" pageNumber="128">1.43 ± 0.12</td>
</tr>
<tr id="FFB8366A36164849FCB10041E009FBDB" box="[833,1396,1056,1080]" pageId="3" pageNumber="128">
<td id="FFB8366436164849FCB10041E6AEFBDB" box="[833,1022,1056,1080]" pageId="3" pageNumber="128">Mesosoma length</td>
<td id="FFB8366436164849FB100040E009FBDB" box="[1248,1369,1056,1080]" pageId="3" pageNumber="128">2.63 ± 0.17</td>
</tr>
<tr id="FFB8366A36164849FCB1002EE009FB85" box="[833,1396,1101,1126]" pageId="3" pageNumber="128">
<td id="FFB8366436164849FCB1002EE154FB85" box="[833,1028,1101,1126]" pageId="3" pageNumber="128">Mesonotum width</td>
<td id="FFB8366436164849FB10002EE009FB85" box="[1248,1369,1102,1126]" pageId="3" pageNumber="128">1.39 ± 0.15</td>
</tr>
<tr id="FFB8366A36164849FCB1001BE009FB70" box="[833,1396,1147,1171]" pageId="3" pageNumber="128">
<td id="FFB8366436164849FCB1001BE680FB70" box="[833,976,1147,1171]" pageId="3" pageNumber="128">Femur length</td>
<td id="FFB8366436164849FB10001BE009FB70" box="[1248,1369,1147,1171]" pageId="3" pageNumber="128">2.04 ± 0.10</td>
</tr>
<tr id="FFB8366A36164849FCB100C8E009FB23" box="[833,1396,1192,1216]" pageId="3" pageNumber="128">
<td id="FFB8366436164849FCB100C8E695FB23" box="[833,965,1192,1216]" pageId="3" pageNumber="128">Wing length</td>
<td id="FFB8366436164849FB1000C8E009FB23" box="[1248,1369,1192,1216]" pageId="3" pageNumber="128">4.77 ± 0.28</td>
</tr>
<tr id="FFB8366A36164849FCB100B5E024FB0E" box="[833,1396,1237,1261]" pageId="3" pageNumber="128">
<td id="FFB8366436164849FCB100B5E6B2FB0E" box="[833,994,1237,1261]" pageId="3" pageNumber="128">Cephalic index</td>
<td id="FFB8366436164849FB3500B5E024FB0E" box="[1221,1396,1237,1261]" pageId="3" pageNumber="128">109.23 ± 10.240</td>
</tr>
<tr id="FFB8366A36164849FCB10163E024FAF9" box="[833,1396,1282,1306]" pageId="3" pageNumber="128">
<td id="FFB8366436164849FCB10163E693FAF9" box="[833,963,1282,1306]" pageId="3" pageNumber="128">Scape index</td>
<td id="FFB8366436164849FB350163E024FAF9" box="[1221,1396,1283,1306]" pageId="3" pageNumber="128">111.39 ± 8.1400</td>
</tr>
<tr id="FFB8366A36164849FCB1014FE152FAAB" box="[833,1396,1327,1352]" pageId="3" pageNumber="128">
<td id="FFB8366436164849FCB1014FE152FAAB" box="[833,1026,1327,1352]" pageId="3" pageNumber="128">Copulatory pieces</td>
</tr>
<tr id="FFB8366A36164849FCB1013CE009FA96" box="[833,1396,1372,1397]" pageId="3" pageNumber="128">
<td id="FFB8366436164849FCB1013CE161FA96" box="[833,1073,1372,1397]" pageId="3" pageNumber="128">Subgenital plate width</td>
<td id="FFB8366436164849FB10013DE009FA96" box="[1248,1369,1373,1397]" pageId="3" pageNumber="128">0.59 ± 0.07</td>
</tr>
<tr id="FFB8366A36164849FCB101EAE009FA41" box="[833,1396,1418,1442]" pageId="3" pageNumber="128">
<td id="FFB8366436164849FCB101EAE6A8FA41" box="[833,1016,1418,1442]" pageId="3" pageNumber="128">Penisvalva width</td>
<td id="FFB8366436164849FB1001EAE009FA41" box="[1248,1369,1418,1442]" pageId="3" pageNumber="128">0.55 ± 0.04</td>
</tr>
<tr id="FFB8366A36164849FCB101D7E009FA2C" box="[833,1396,1463,1487]" pageId="3" pageNumber="128">
<td id="FFB8366436164849FCB101D7E6AEFA2C" box="[833,1022,1463,1487]" pageId="3" pageNumber="128">Penisvalva length</td>
<td id="FFB8366436164849FB1001D7E009FA2C" box="[1248,1369,1463,1487]" pageId="3" pageNumber="128">0.96 ± 0.05</td>
</tr>
<tr id="FFB8366A36164849FCB10185E009FA1F" box="[833,1396,1508,1532]" pageId="3" pageNumber="128">
<td id="FFB8366436164849FCB10185E6BFFA1F" box="[833,1007,1508,1532]" pageId="3" pageNumber="128">Paramere length</td>
<td id="FFB8366436164849FB100185E009FA1F" box="[1248,1369,1509,1532]" pageId="3" pageNumber="128">0.89 ± 0.04</td>
</tr>
<tr id="FFB8366A36164849FCB10271E009F9CA" box="[833,1396,1553,1577]" pageId="3" pageNumber="128">
<td id="FFB8366436164849FCB10271E114F9CA" box="[833,1092,1553,1577]" pageId="3" pageNumber="128">Volsella (digitus) length</td>
<td id="FFB8366436164849FB100272E009F9CA" box="[1248,1369,1554,1577]" pageId="3" pageNumber="128">0.74 ± 0.05</td>
</tr>
</table>
</paragraph>
<paragraph id="8B97F17A36164849FF4D0179E486FAAD" pageId="3" pageNumber="128">Variability: Little differences in mesonotum size and degree of thoracic development.</paragraph>
<paragraph id="8B97F17A36164849FF4D0134E590F990" pageId="3" pageNumber="128">
<emphasis id="B95C2D6836164849FF4D0134E48DFA88" bold="true" box="[189,477,1364,1387]" pageId="3" pageNumber="128">Description of ergatoids:</emphasis>
Looks like a worker; hard to distinguish between the two using the naked eye. Scape length intermediate to those of brachypterous queens and worker. Mesosoma only slightly developed, with very small scutum, scutellum, and metanotum, in profile view appearing slightly elevated; scutum-scutellum suture very slightly impressed or disappeared. Metanotum plate arcuate and wrinkled. Wing insertion points extremely reduced (
<figureCitation id="1313EDFF36164849FD440240E7A2F9DA" box="[692,754,1568,1593]" captionStart="Fig. 2" captionStartId="2.[1096,1133,1249,1274]" captionText="Fig. 2: Cataglyphis tartessica sp. n. (a) Brachypterous queen. (b) Ergatoid queen (minute wing insertion indicated with an arrow)." pageId="3" pageNumber="128">Fig. 2</figureCitation>
). Leg length intermediate to those of brachypterous and worker.
</paragraph>
<paragraph id="8B97F17A36164849FF4D0218E7E8F928" pageId="3" pageNumber="128">Variability: At a glance, some ergatoids without wing insertion points; they can only be distinguished from workers by their physogastric abdomen and behaviour.</paragraph>
<paragraph id="8B97F17A36164849FF4D02B1E490F8A2" pageId="3" pageNumber="128">
<emphasis id="B95C2D6836164849FF4D02B1E4FFF90A" bold="true" box="[189,431,1745,1769]" pageId="3" pageNumber="128">Description of males:</emphasis>
External morphology (Tab. 3): Head, scape, mesosoma, petiole, and coxae dull black; funiculus, legs, and gaster dull amber. Large, light brown wings with strong venation.
</paragraph>
<paragraph id="8B97F17A36164849FF4D0325E0CBFCE0" pageId="3" pageNumber="128">
Head ovoid and small. Prominent large ovoid compound eyes located in mid to upper half of face sides; prominent large ocelli also present. Cephalic capsule with a slight blister in the occipital margin behind the ocelli. Scoop-shaped mandibles, small, fragile, and without striations, that taper into a small apical tooth. Antennae with Tab. 3: Measurements in millimetres of males of 
<emphasis id="B95C2D6836164849FAAD068BE0CBFCE0" box="[1373,1435,747,771]" italics="true" pageId="3" pageNumber="128">Cata-</emphasis>
</paragraph>
<paragraph id="8B97F17A36164849FCC40768E104FCC3" box="[820,1108,776,801]" pageId="3" pageNumber="128">
<emphasis id="B95C2D6836164849FCC40768E6BFFCC2" box="[820,1007,776,801]" italics="true" pageId="3" pageNumber="128">
glyphis 
<taxonomicName id="4C288AF936164849FC780769E6BFFCC2" box="[904,1007,777,801]" class="Insecta" family="Formicidae" genus="Cataglyphis" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="tartessica">tartessica</taxonomicName>
</emphasis>
(n = 30).
</paragraph>
<paragraph id="8B97F17A36164849FCB1075FE187FCB5" box="[833,1239,830,854]" pageId="3" pageNumber="128">
<emphasis id="B95C2D6836164849FCB1075FE12AFCB5" bold="true" box="[833,1146,831,854]" pageId="3" pageNumber="128">
<taxonomicName id="4C288AF936164849FCB1075FE162FCB5" box="[833,1074,831,854]" class="Insecta" family="Formicidae" genus="Cataglyphis" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="tartessica">
<emphasis id="B95C2D6836164849FCB1075FE162FCB5" bold="true" box="[833,1074,831,854]" italics="true" pageId="3" pageNumber="128">Cataglyphis tartessica</emphasis>
</taxonomicName>
males
</emphasis>
(n = 30)
</paragraph>
<paragraph id="8B97F17A36164849FCC40219E10DF94D" pageId="3" pageNumber="128">13 segments; the two basalmost funicular segments somewhat longer than the others.</paragraph>
<paragraph id="8B97F17A36164849FCAA02D4E11DF87B" pageId="3" pageNumber="128">Mesosoma massive, with well-developed scutum and scutellum; metanotum small. Pleural sclerites slightly striated next to wing insertions. Petiole scale-like, large, and thick, with bilobed or heart-shaped apex due to a slight, longitudinal constriction. Scutum, scutellum, and metanotum with very sparse pubescence. Scattered setae on scutum. Abundant pubescence on pronotum, propodeum, pleura, coxae, and petiole.</paragraph>
<paragraph id="8B97F17A36164849FCAA03FEE1DBF813" pageId="3" pageNumber="128">Gaster cylindrical densely microsculptured and with medium pubescence. Sternites with long and numerous setae, and tergites lacking setae.</paragraph>
<caption id="DF57A1F23611484EFF67010FE1EFFA22" pageId="4" pageNumber="129" targetBox="[151,1434,178,1370]" targetPageId="4">
<paragraph id="8B97F17A3611484EFF67010FE1EFFA22" pageId="4" pageNumber="129">
Fig. 3: Views in scanning electron microscopy of the copulatory appendages of (a - d) 
<taxonomicName id="4C288AF93611484EFBE2010FE1C4FA64" box="[1042,1172,1391,1415]" class="Insecta" family="Formicidae" genus="Cataglyphis" genus.bestMatchDistance="312" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="tartessica" status="sp.n.">
<emphasis id="B95C2D683611484EFBE2010FE1C4FA64" box="[1042,1172,1391,1415]" italics="true" pageId="4" pageNumber="129">C. tartessica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F90133611484EFB6A0110E19BFA64" box="[1178,1227,1392,1415]" pageId="4" pageNumber="129">sp.n.</taxonomicNameLabel>
and (e) 
<taxonomicName id="4C288AF93611484EFAEF010FE0C5FA64" box="[1311,1429,1391,1415]" class="Insecta" family="Formicidae" genus="Cataglyphis" genus.bestMatchDistance="790" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="floricola">
<emphasis id="B95C2D683611484EFAEF010FE0C5FA64" box="[1311,1429,1391,1415]" italics="true" pageId="4" pageNumber="129">C. floricola</emphasis>
</taxonomicName>
: (a) paramere; (b) penisvalva; (c) volsella, with digitus (the largest) and cuspis; (d) subgenital plate. Note the differences in size and shape between the subgenital plates. Measurements are indicated with double-headed arrows.
</paragraph>
</caption>
<paragraph id="8B97F17A3611484EFF4D0186E414F8E5" pageId="4" pageNumber="129">
Genital morphology (
<figureCitation id="1313EDFF3611484EFE550186E4B4FA1C" box="[421,484,1510,1535]" captionStart="Fig. 3" captionStartId="4.[151,189,1391,1415]" captionTargetBox="[151,1434,178,1370]" captionTargetId="figure@4.[151,1436,178,1370]" captionTargetPageId="4" captionText="Fig. 3: Views in scanning electron microscopy of the copulatory appendages of (a - d) C. tartessica sp. n. and (e) C. floricola: (a) paramere; (b) penisvalva; (c) volsella, with digitus (the largest) and cuspis; (d) subgenital plate. Note the differences in size and shape between the subgenital plates. Measurements are indicated with double-headed arrows." pageId="4" pageNumber="129">Fig. 3</figureCitation>
): Paramere elongated and oval, with abundant long setae, and externally bearing a small finger-like process at its base next to cuspis. Volsella with digitus and cuspis straight and parallel, digitus larger than cuspis. Penisvalva twice as long as wide, with 7 (6 - 9) minute teeth in its internal arcuate margin, and a flat peak in its terminal end. Subgenital plate concave and bilobed posterolaterally, with rounded notch in centre of depression, at posterior margin; numerous setae present on its exterior face.
</paragraph>
<paragraph id="8B97F17A3611484EFF4D036CE419F812" pageId="4" pageNumber="129">Variability: The extent of black colouration can vary among males; for example, the scape, which is usually black, can also be amber. The first funicular and gastral segments can be black in some males. The genitalia are very uniform; the number of penisvalva teeth can differ, and the posterior margin of the subgenital plate can be continuous (unbroken in the middle) or sometimes present a slight convexity.</paragraph>
</subSubSection>
<subSubSection id="C332A2F13611484EFCAA0187E62AF94C" pageId="4" pageNumber="129" type="etymology">
<paragraph id="8B97F17A3611484EFCAA0187E62AF94C" pageId="4" pageNumber="129">
<emphasis id="B95C2D683611484EFCAA0187E68FFA1D" bold="true" box="[858,991,1511,1534]" pageId="4" pageNumber="129">Etymology:</emphasis>
The name of this species comes from its current geographic distribution in southwestern Andalusia. It is distributed across the area thought to have been occupied by the ancient Tartessian civilisation, which existed three thousand years ago in the southwestern Iberian Peninsula (LÓPEZ-DAVALILLO 1999, GONZÁLEZ 
<collectionCode id="ED3969BF3611484EFAE3021DE07FF973" box="[1299,1327,1661,1680]" pageId="4" pageNumber="129">DE</collectionCode>
CANALES 2004).
</paragraph>
</subSubSection>
<subSubSection id="C332A2F13611484FFCAA02D4E4ABF9AA" lastPageId="5" lastPageNumber="130" pageId="4" pageNumber="129" type="discussion">
<paragraph id="8B97F17A3611484FFCAA02D4E4ABF9AA" lastPageId="5" lastPageNumber="130" pageId="4" pageNumber="129">
<emphasis id="B95C2D683611484EFCAA02D4E061F92F" bold="true" box="[858,1329,1716,1740]" pageId="4" pageNumber="129">
Relationship to other 
<taxonomicName id="4C288AF93611484EFBA102D4E183F92F" box="[1105,1235,1716,1740]" class="Insecta" family="Formicidae" genus="Cataglyphis" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis id="B95C2D683611484EFBA102D4E183F92F" bold="true" box="[1105,1235,1716,1740]" italics="true" pageId="4" pageNumber="129">Cataglyphis</emphasis>
</taxonomicName>
species:
</emphasis>
As previously noted, 
<taxonomicName id="4C288AF93611484EFC3102B1E1E6F90A" box="[961,1206,1745,1770]" class="Insecta" family="Formicidae" genus="Cataglyphis" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="tartessica" status="sp.n.">
<emphasis id="B95C2D683611484EFC3102B1E1E6F90A" box="[961,1206,1745,1770]" italics="true" pageId="4" pageNumber="129">Cataglyphis tartessica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F90133611484EFB4E02B2E1A2F90A" box="[1214,1266,1746,1769]" pageId="4" pageNumber="129">sp.n.</taxonomicNameLabel>
is phylogenetically closely related to 
<taxonomicName id="4C288AF93611484EFBDC028FE1F8F8E4" box="[1068,1192,1774,1799]" class="Insecta" family="Formicidae" genus="Cataglyphis" genus.bestMatchDistance="478" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="floricola">
<emphasis id="B95C2D683611484EFBDC028FE1F8F8E4" box="[1068,1192,1774,1799]" italics="true" pageId="4" pageNumber="129">C. floricola</emphasis>
</taxonomicName>
(see 
<collectionCode id="ED3969BF3611484EFB11028FE075F8E4" box="[1249,1317,1775,1799]" pageId="4" pageNumber="129">AMOR</collectionCode>
2011), and belongs, as 
<taxonomicName id="4C288AF93611484EFC45036CE163F8C7" box="[949,1075,1804,1828]" class="Insecta" family="Formicidae" genus="Cataglyphis" genus.bestMatchDistance="465" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="floricola">
<emphasis id="B95C2D683611484EFC45036CE163F8C7" box="[949,1075,1804,1828]" italics="true" pageId="4" pageNumber="129">C. floricola</emphasis>
</taxonomicName>
(see TINAUT 1993), to the 
<taxonomicName id="4C288AF93611484EFAAF036CE680F8A1" class="Insecta" family="Formicidae" genus="Cataglyphis" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="emmae">
<emphasis id="B95C2D683611484EFAAF036CE680F8A1" italics="true" pageId="4" pageNumber="129">Cataglyphis emmae</emphasis>
</taxonomicName>
group (AGOSTI 1990). Apart from their different colouration, the two species are most notably distinguished by differences in the size of their brachypterous queens and males (brachypterous 
<collectionCode id="ED3969BF3611484EFB5F03E2E18CF87A" box="[1199,1244,1922,1945]" pageId="4" pageNumber="129">ML</collectionCode>
: 2.25 ± 0.16, n = 25 vs. 1.97 ± 0.08, n = 16, 
<taxonomicName id="4C288AF93611484EFB7A03FFE04BF854" box="[1162,1307,1951,1975]" class="Insecta" family="Formicidae" genus="Cataglyphis" genus.bestMatchDistance="137" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="tartessica" status="sp.n.">
<emphasis id="B95C2D683611484EFB7A03FFE04BF854" box="[1162,1307,1951,1975]" italics="true" pageId="4" pageNumber="129">C. tartessica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F90133611484EFAD803FFE00CF854" box="[1320,1372,1951,1975]" pageId="4" pageNumber="129">sp.n.</taxonomicNameLabel>
– 
<taxonomicName id="4C288AF93611484EFA7303FFE6C0F837" class="Insecta" family="Formicidae" genus="Cataglyphis" genus.bestMatchDistance="393" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="floricola">
<emphasis id="B95C2D683611484EFA7303FFE6C0F837" italics="true" pageId="4" pageNumber="129">C. floricola</emphasis>
</taxonomicName>
, respectively; male 
<collectionCode id="ED3969BF3611484EFB9B03DDE1C7F837" box="[1131,1175,1981,2004]" pageId="4" pageNumber="129">ML</collectionCode>
: 2.63 ± 0.17, n = 29 vs. 2.20 ± 0.13, n = 23, 
<emphasis id="B95C2D683611484EFBE303BAE069F812" box="[1043,1337,2009,2033]" italics="true" pageId="4" pageNumber="129">
<taxonomicName id="4C288AF93611484EFBE303BAE1CDF812" box="[1043,1181,2010,2033]" class="Insecta" family="Formicidae" genus="Cataglyphis" genus.bestMatchDistance="103" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="tartessica">C. tartessica</taxonomicName>
– 
<taxonomicName id="4C288AF93611484EFB4A03BAE069F812" box="[1210,1337,2009,2033]" class="Insecta" family="Formicidae" genus="Cataglyphis" genus.bestMatchDistance="363" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="floricola">C. floricola</taxonomicName>
</emphasis>
, respectively. Student's t-test: t = 6.33 and t = 9.71, respectively; 
<emphasis id="B95C2D683610484FFF6701BAE5F7FA12" box="[151,167,1498,1521]" italics="true" pageId="5" pageNumber="130">
<collectionCode id="ED3969BF3610484FFF6701BAE5F7FA12" box="[151,167,1498,1521]" pageId="5" pageNumber="130">P</collectionCode>
</emphasis>
&lt;10- 
<quantity id="4CD05C9F3610484FFF1701B4E45BFA12" box="[231,267,1492,1521]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="5" pageNumber="130" unit="in" value="6.0">6 in</quantity>
both cases; 
<figureCitation id="1313EDFF3610484FFE6301B9E486FA12" box="[403,470,1497,1521]" captionStart="Fig. 4" captionStartId="5.[151,190,617,641]" captionTargetBox="[154,763,180,590]" captionTargetId="figure@5.[151,763,176,591]" captionTargetPageId="5" captionText="Fig. 4: Box (25 % - 75 % of data; with the median position) and Whisker (not outlier data range) graph showing the mesosoma length differences between Cataglyphis tartessica sp. n. and C. floricola sexuals (brachypterous females and males). Different letters denote significant differences." pageId="5" pageNumber="130">Fig. 4</figureCitation>
). Both species are ecologically very similar; they exhibit similar nesting and feeding preferences as well as comparable social structures and dispersal modes (F. Amor, unpubl.).
</paragraph>
</subSubSection>
<caption id="DF57A1F23610484FFF670609E5A7FCF7" pageId="5" pageNumber="130" targetBox="[154,763,180,590]" targetPageId="5">
<paragraph id="8B97F17A3610484FFF670609E5A7FCF7" pageId="5" pageNumber="130">
Fig. 4: Box (25% - 75% of data; with the median position) and Whisker (not outlier data range) graph showing the mesosoma length differences between 
<taxonomicName id="4C288AF93610484FFDA306C4E5B1FD3A" class="Insecta" family="Formicidae" genus="Cataglyphis" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="tartessica" status="sp.n.">
<emphasis id="B95C2D683610484FFDA306C4E5B1FD3A" italics="true" pageId="5" pageNumber="130">Cataglyphis tartessica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F90133610484FFF1A06A2E44EFD3A" box="[234,286,706,729]" pageId="5" pageNumber="130">sp.n.</taxonomicNameLabel>
and 
<taxonomicName id="4C288AF93610484FFEA506A2E484FD39" box="[341,468,705,730]" class="Insecta" family="Formicidae" genus="Cataglyphis" genus.bestMatchDistance="257" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="floricola">
<emphasis id="B95C2D683610484FFEA506A2E484FD39" box="[341,468,705,730]" italics="true" pageId="5" pageNumber="130">C. floricola</emphasis>
</taxonomicName>
sexuals (brachypterous females and males). Different letters denote significant differences.
</paragraph>
</caption>
<caption id="DF57A1F23610484FFF670149E402FA7A" pageId="5" pageNumber="130" targetBox="[151,763,815,1298]" targetPageId="5">
<paragraph id="8B97F17A3610484FFF670149E402FA7A" pageId="5" pageNumber="130">
Fig. 5: Geographic distribution of 
<taxonomicName id="4C288AF93610484FFDFE0149E7AFFAA1" box="[526,767,1321,1346]" class="Insecta" family="Formicidae" genus="Cataglyphis" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="tartessica" status="sp.n.">
<emphasis id="B95C2D683610484FFDFE0149E7AFFAA1" box="[526,767,1321,1346]" italics="true" pageId="5" pageNumber="130">Cataglyphis tartessica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F90133610484FFF670127E59AFABC" box="[151,202,1351,1375]" pageId="5" pageNumber="130">sp.n.</taxonomicNameLabel>
(red dots) and 
<taxonomicName id="4C288AF93610484FFE9E0127E4B9FABC" box="[366,489,1350,1375]" class="Insecta" family="Formicidae" genus="Cataglyphis" genus.bestMatchDistance="222" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="floricola">
<emphasis id="B95C2D683610484FFE9E0127E4B9FABC" box="[366,489,1350,1375]" italics="true" pageId="5" pageNumber="130">C. floricola</emphasis>
</taxonomicName>
(black dots) in southwestern Spain. Note the parapatric relationship between the two distributions.
</paragraph>
</caption>
<subSubSection id="C332A2F13610484CFF4D022FE7A9FA82" lastPageId="6" lastPageNumber="131" pageId="5" pageNumber="130" type="biology_ecology">
<paragraph id="8B97F17A3610484FFF4D022FE195FEC2" pageId="5" pageNumber="130">
<emphasis id="B95C2D683610484FFF4D022FE4E7F984" bold="true" box="[189,439,1615,1639]" pageId="5" pageNumber="130">Natural history notes:</emphasis>
<taxonomicName id="4C288AF93610484FFE4E022EE7FDF984" box="[446,685,1614,1639]" class="Insecta" family="Formicidae" genus="Cataglyphis" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="tartessica" status="sp.n.">
<emphasis id="B95C2D683610484FFE4E022EE7FDF984" box="[446,685,1614,1639]" italics="true" pageId="5" pageNumber="130">Cataglyphis tartessica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F90133610484FFD44022FE7B7F984" box="[692,743,1615,1639]" pageId="5" pageNumber="130">sp.n.</taxonomicNameLabel>
is endemic to Spain. It occurs in habitats with sandy soils and has a small range. Its populations are located in southwestern Spain, in an arc of about 
<quantity id="4CD05C9F3610484FFE4802C7E756F95D" box="[440,518,1702,1726]" metricMagnitude="5" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="130" unit="km" value="200.0">200 km</quantity>
around the lower stretch of the Guadalquivir River; its distribution is parapatric to that of 
<taxonomicName id="4C288AF93610484FFF110282E409F91A" box="[225,345,1761,1785]" class="Insecta" family="Formicidae" genus="Cataglyphis" genus.bestMatchDistance="143" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="floricola">
<emphasis id="B95C2D683610484FFF110282E409F91A" box="[225,345,1761,1785]" italics="true" pageId="5" pageNumber="130">C. floricola</emphasis>
</taxonomicName>
(
<figureCitation id="1313EDFF3610484FFE990281E4F7F91A" box="[361,423,1761,1785]" captionStart="Fig. 5" captionStartId="5.[151,190,1321,1345]" captionTargetBox="[151,763,815,1298]" captionTargetId="figure@5.[151,763,815,1298]" captionTargetPageId="5" captionText="Fig. 5: Geographic distribution of Cataglyphis tartessica sp. n. (red dots) and C. floricola (black dots) in southwestern Spain. Note the parapatric relationship between the two distributions." pageId="5" pageNumber="130">Fig. 5</figureCitation>
). The Guadalquivir river estuary underwent very strong changes throughout the Quaternary (SALVANY &amp; CUSTODIO 1995, RODRÍGUEZ-RAMÍREZ &amp; al. 1996, 
<collectionCode id="ED3969BF3610484FFF0F035AE46BF8B2" box="[255,315,1850,1873]" pageId="5" pageNumber="130">ZAZO</collectionCode>
&amp; al. 2005, SALVANY &amp; al. 2010); these changes may have generated barriers that resulted in isolation and speciation. Thus, the current parapatric distribution of the two taxa could be interpreted as their first meeting since the Quaternary: now that the barriers have disappeared, their ranges have spread. The apparent stable boundary between their populations – 
<taxonomicName id="4C288AF93610484FFB7E04D2E054FF2A" box="[1166,1284,177,201]" class="Insecta" family="Formicidae" genus="Cataglyphis" genus.bestMatchDistance="44" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="floricola">
<emphasis id="B95C2D683610484FFB7E04D2E054FF2A" box="[1166,1284,177,201]" italics="true" pageId="5" pageNumber="130">C. floricola</emphasis>
</taxonomicName>
occurs largely on the coast and 
<taxonomicName id="4C288AF93610484FFC1904AFE13DFF04" box="[1001,1133,207,231]" class="Insecta" family="Formicidae" genus="Cataglyphis" genus.bestMatchDistance="140" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="tartessica" status="sp.n.">
<emphasis id="B95C2D683610484FFC1904AFE13DFF04" box="[1001,1133,207,231]" italics="true" pageId="5" pageNumber="130">C. tartessica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F90133610484FFB8504AFE1F7FF05" box="[1141,1191,207,230]" pageId="5" pageNumber="130">sp.n.</taxonomicNameLabel>
is found mostly inland – could be a consequence of each species being more fit under different ecological conditions.
</paragraph>
<paragraph id="8B97F17A3610484FFCAA0546E6BBFD62" pageId="5" pageNumber="130">
Before the species' distributions were well characterised, 
<taxonomicName id="4C288AF93610484FFCC40524E142FEBF" box="[820,1042,324,348]" class="Insecta" family="Formicidae" genus="Cataglyphis" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="floricola">
<emphasis id="B95C2D683610484FFCC40524E142FEBF" box="[820,1042,324,348]" italics="true" pageId="5" pageNumber="130">Cataglyphis floricola</emphasis>
</taxonomicName>
(the black morph) was thought to occupy environments with more vegetative cover than 
<taxonomicName id="4C288AF93610484FFAA70502E62CFE74" class="Insecta" family="Formicidae" genus="Cataglyphis" genus.bestMatchDistance="88" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="tartessica" status="sp.n.">
<emphasis id="B95C2D683610484FFAA70502E62CFE74" italics="true" pageId="5" pageNumber="130">C. tartessica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F90133610484FFC75051FE6E8FE75" box="[901,952,383,406]" pageId="5" pageNumber="130">sp.n.</taxonomicNameLabel>
(previously identified as the 
<taxonomicName id="4C288AF93610484FFB09051FE026FE74" box="[1273,1398,382,407]" class="Insecta" family="Formicidae" genus="Cataglyphis" genus.bestMatchDistance="87" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="floricola">
<emphasis id="B95C2D683610484FFB09051FE026FE74" box="[1273,1398,382,407]" italics="true" pageId="5" pageNumber="130">C. floricola</emphasis>
</taxonomicName>
bicolour morph). Thus, the difference in their colouration was proposed to be related to different habitat preferences, which were linked to different environmental temperatures: Because 
<taxonomicName id="4C288AF93610484FFCC40594E6E4FDEF" box="[820,948,500,524]" class="Insecta" family="Formicidae" genus="Cataglyphis" genus.bestMatchDistance="186" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="floricola">
<emphasis id="B95C2D683610484FFCC40594E6E4FDEF" box="[820,948,500,524]" italics="true" pageId="5" pageNumber="130">C. floricola</emphasis>
</taxonomicName>
nested in areas characterised by lower temperatures, its colouration was black (CARPINTERO &amp; al. 2002). However, current knowledge of their distribution patterns suggests that they do not prefer different levels of vegetative cover.
</paragraph>
<paragraph id="8B97F17A3610484FFCAA06E6E006FC44" pageId="5" pageNumber="130">
<taxonomicName id="4C288AF93610484FFCAA06E6E16DFD7C" box="[858,1085,646,671]" class="Insecta" family="Formicidae" genus="Cataglyphis" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="tartessica" status="sp.n.">
<emphasis id="B95C2D683610484FFCAA06E6E16DFD7C" box="[858,1085,646,671]" italics="true" pageId="5" pageNumber="130">Cataglyphis tartessica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F90133610484FFBB406E7E124FD7C" box="[1092,1140,647,671]" pageId="5" pageNumber="130">sp.n.</taxonomicNameLabel>
is a monomorphic, mediumsized ant (total body size: 4.51 ± 
<quantity id="4CD05C9F3610484FFB6F06C4E052FD5F" box="[1183,1282,676,700]" metricMagnitude="-2" metricUnit="m" metricValue="4.1" pageId="5" pageNumber="130" unit="mm" value="41.0">0.41 mm</quantity>
; n = 35), very similar in size and appearance to other 
<taxonomicName id="4C288AF93610484FFB3C06A1E01AFD3A" box="[1228,1354,705,729]" class="Insecta" family="Formicidae" genus="Cataglyphis" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="genus">
<emphasis id="B95C2D683610484FFB3C06A1E01AFD3A" box="[1228,1354,705,729]" italics="true" pageId="5" pageNumber="130">Cataglyphis</emphasis>
</taxonomicName>
species of the southern Iberian Peninsula (
<figureCitation id="1313EDFF3610484FFB5C06BEE1BDFD15" box="[1196,1261,734,759]" captionStart="Fig. 6" captionStartId="6.[151,189,1052,1076]" captionTargetBox="[151,765,176,1029]" captionTargetId="figure@6.[152,764,176,1028]" captionTargetPageId="6" captionText="Fig. 6: Comparative view among workers of four Cataglyphis species. Apart from the colour, note the similarity in size and design. (a) C. tartessica sp. n.; (b) C. floricola; (c) C. rosenhaueri SANTSCHI, 1925; (d) C. iberica EMERY, 1906." pageId="5" pageNumber="130">Fig. 6</figureCitation>
). The species is monodomous (each colony occupies one single nest) and monogynous (each colony is headed by a single queen), with a mean number of workers per nest of 182.8 ± 130.6 (n = 106). Although the species range is rather small, local nest density is high, with about 6 nests per 
<quantity id="4CD05C9F3610484FFB010712E076FC6A" box="[1265,1318,882,905]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="130" unit="m" value="10.0">10 m</quantity>
2 that are in close proximity to each other (0.81 ± 0.35; n = 202).
</paragraph>
<paragraph id="8B97F17A3610484FFCAA07CCE052FA9F" pageId="5" pageNumber="130">
Colonies of 
<taxonomicName id="4C288AF93610484FFC1207CCE13EFC27" box="[994,1134,940,964]" class="Insecta" family="Formicidae" genus="Cataglyphis" genus.bestMatchDistance="319" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="tartessica" status="sp.n.">
<emphasis id="B95C2D683610484FFC1207CCE13EFC27" box="[994,1134,940,964]" italics="true" pageId="5" pageNumber="130">C. tartessica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F90133610484FFB8607CDE1FAFC27" box="[1142,1194,941,964]" pageId="5" pageNumber="130">sp.n.</taxonomicNameLabel>
reproduce by fission. This process is described in detail by AMOR (2011). Around the summer solstice, a third of the workers and cocoons of a reproducing colony are transported, one by one, by a small number of &quot;transporter&quot; ants; the distance from the mother nest to the new nest is 
<quantity id="4CD05C9F3610484FFB8B005FE1F4FBB4" box="[1147,1188,1087,1111]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="130" unit="m" value="8.0">8 m</quantity>
on average. The gynes are transported while still in their cocoons, and the queen of the colony remains in the mother nest. The presence in the same population of two queen morphs, brachypterous and ergatoid, that differ in reproductive potential (that of the brachypterous being greater) could be a consequence of a conflict of interest within the colony. So, ergatoids may compete with the brachypterous for the colony leadership and escape policing activity through their morphological similarity to the workers. Supernumerary gynes are reduced by worker aggression (
<collectionCode id="ED3969BF3610484FFBD60104E13DFA9F" box="[1062,1133,1380,1404]" pageId="5" pageNumber="130">AMOR</collectionCode>
&amp; al. 2011a).
</paragraph>
<paragraph id="8B97F17A3610484FFCAA01E1E1C2F91A" pageId="5" pageNumber="130">
The diet of 
<taxonomicName id="4C288AF93610484FFC2D01E2E138FA79" box="[989,1128,1410,1434]" class="Insecta" family="Formicidae" genus="Cataglyphis" genus.bestMatchDistance="141" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="tartessica" status="sp.n.">
<emphasis id="B95C2D683610484FFC2D01E2E138FA79" box="[989,1128,1410,1434]" italics="true" pageId="5" pageNumber="130">C. tartessica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F90133610484FFB8101E2E1F4FA7A" box="[1137,1188,1410,1433]" pageId="5" pageNumber="130">sp.n.</taxonomicNameLabel>
mainly consists of arthropod, which is typical of the 
<taxonomicName id="4C288AF93610484FFB7101FEE1AFFA54" box="[1153,1279,1438,1463]" class="Insecta" family="Formicidae" genus="Cataglyphis" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="genus">
<emphasis id="B95C2D683610484FFB7101FEE1AFFA54" box="[1153,1279,1438,1463]" italics="true" pageId="5" pageNumber="130">Cataglyphis</emphasis>
</taxonomicName>
genus (
<collectionCode id="ED3969BF3610484FFABF01FDE0CCFA54" box="[1359,1436,1437,1463]" pageId="5" pageNumber="130">CERDÁ</collectionCode>
&amp; al. 1989). However, the species, which is mostly a scavenger, also opportunistically harvests other available food resources, a characteristic that is an adaptation to resourcepoor environments. Thus, depending on location, we observe large differences in diet composition. Some diets contain a large quantity of plant material: in 
<collectionCode id="ED3969BF3610484FFAE0022FE016F985" box="[1296,1350,1615,1638]" collectionName="DBR" pageId="5" pageNumber="130">DBR</collectionCode>
in July, 80% of the diet consisted of 
<taxonomicName id="4C288AF93610484FFB93020CE008F967" box="[1123,1368,1644,1668]" class="Magnoliopsida" family="Cistaceae" genus="Halimium" higherTaxonomySource="CoL" kingdom="Plantae" order="Malvales" pageId="5" pageNumber="130" phylum="Magnoliophyta" rank="species" species="halimifolium">
<emphasis id="B95C2D683610484FFB93020CE008F967" box="[1123,1368,1644,1668]" italics="true" pageId="5" pageNumber="130">Halimium halimifolium</emphasis>
</taxonomicName>
petals (
<collectionCode id="ED3969BF3610484FFCCA02E8E6D8F941" box="[826,904,1672,1698]" pageId="5" pageNumber="130">CERDÁ</collectionCode>
&amp; al. 1992, 1996). Others demonstrate a clear preponderance of arthropod corpses where 
<taxonomicName id="4C288AF93610484FFB1F02C7E0CCF95C" box="[1263,1436,1702,1727]" class="Magnoliopsida" family="Cistaceae" genus="Halimium" genus.bestMatchDistance="22" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Plantae" order="Malvales" pageId="5" pageNumber="130" phylum="Magnoliophyta" rank="species" species="halimifolium">
<emphasis id="B95C2D683610484FFB1F02C7E0CCF95C" box="[1263,1436,1702,1727]" italics="true" pageId="5" pageNumber="130">H. halimifolium</emphasis>
</taxonomicName>
is absent: in 
<collectionCode id="ED3969BF3610484FFC3002A4E6A8F93F" box="[960,1016,1732,1756]" collectionName="ALC" pageId="5" pageNumber="130">ALC</collectionCode>
, also in July, 90% of diet consisted of arthropod corpses (
<collectionCode id="ED3969BF3610484FFBF00281E116F91A" box="[1024,1094,1761,1785]" pageId="5" pageNumber="130">AMOR</collectionCode>
2011).
</paragraph>
<paragraph id="8B97F17A3610484CFCAA029FE7A9FA82" lastPageId="6" lastPageNumber="131" pageId="5" pageNumber="130">
In 
<taxonomicName id="4C288AF93610484FFC87029EE131F8F4" box="[887,1121,1790,1815]" class="Insecta" family="Formicidae" genus="Cataglyphis" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="tartessica" status="sp.n.">
<emphasis id="B95C2D683610484FFC87029EE131F8F4" box="[887,1121,1790,1815]" italics="true" pageId="5" pageNumber="130">Cataglyphis tartessica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F90133610484FFB98029FE1C9F8F4" box="[1128,1177,1791,1815]" pageId="5" pageNumber="130">sp.n.</taxonomicNameLabel>
, foraging for food items is solitary, with individual ants searching for resources, as in all 
<taxonomicName id="4C288AF93610484FFC800359E6BDF8B1" box="[880,1005,1849,1874]" class="Insecta" family="Formicidae" genus="Cataglyphis" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="genus">
<emphasis id="B95C2D683610484FFC800359E6BDF8B1" box="[880,1005,1849,1874]" italics="true" pageId="5" pageNumber="130">Cataglyphis</emphasis>
</taxonomicName>
species (WEHNER &amp; al. 1983). However, although it is widely thought that the genus 
<taxonomicName id="4C288AF93610484FFB070336E024F88C" box="[1271,1396,1878,1903]" class="Insecta" family="Formicidae" genus="Cataglyphis" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="genus">
<emphasis id="B95C2D683610484FFB070336E024F88C" box="[1271,1396,1878,1903]" italics="true" pageId="5" pageNumber="130">Cataglyphis</emphasis>
</taxonomicName>
has lost its ability to recruit as a result of selection imposed by high surface temperatures (
<collectionCode id="ED3969BF3610484FFB9303F2E1E4F84A" box="[1123,1204,1938,1961]" pageId="5" pageNumber="130">RUANO</collectionCode>
&amp; al. 2000), the foragers of 
<taxonomicName id="4C288AF93610484FFC6603CFE172F824" box="[918,1058,1967,1991]" class="Insecta" family="Formicidae" genus="Cataglyphis" genus.bestMatchDistance="216" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="tartessica">
<emphasis id="B95C2D683610484FFC6603CFE172F824" box="[918,1058,1967,1991]" italics="true" pageId="5" pageNumber="130">C. tartessica</emphasis>
</taxonomicName>
are able to carry out rudimentary but very efficient recruitment, without indicating the food location, when they find a large prey item near their nest. The result is cooperative prey-retrieval for large or heavy items that could not be exploited otherwise (
<collectionCode id="ED3969BF3613484CFD8C0092E790FAEA" box="[636,704,1266,1289]" pageId="6" pageNumber="131">AMOR</collectionCode>
&amp; al. 2010). A similar, short-distance recruitment behaviour, used to exploit rich food patches, has already been observed in 
<taxonomicName id="4C288AF93613484CFD18014CE5B2FA82" class="Insecta" family="Formicidae" genus="Cataglyphis" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="131" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B95C2D683613484CFD18014CE5B2FA82" italics="true" pageId="6" pageNumber="131">C. bicolor</emphasis>
</taxonomicName>
(FABRICIUS, 1793) in Tunisia (WEHNER 1987: 33).
</paragraph>
</subSubSection>
</treatment>
</document>