255 lines
30 KiB
XML
255 lines
30 KiB
XML
<document id="045EA727F229EBD6F950ADC0F9A2D29E" ID-DOI="10.5281/zenodo.10396362" ID-ISSN="1942-1354" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1707168658757" checkinUser="felipe" docAuthor="Zhang, Jing, Cong, Qian & Grishin, Nick V." docDate="2023" docId="03810139FFF9BB77C0CAFC6DE2F3B312" docLanguage="en" docName="InsectaMundi.2023.1026.1-115.pdf" docOrigin="Insecta Mundi 2023 (26)" docSource="http://dx.doi.org/10.5281/zenodo.10396362" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.3:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="3" docTitle="Heliopetes (Heliopetes) lana Zhang & Cong & Grishin 2023, new species" docType="treatment" docVersion="1" lastPageNumber="38" masterDocId="FFB87941FFDFBB50C00AFF56E355B262" masterDocTitle="Supplementary Materials and Appendix" masterLastPageNumber="115" masterPageNumber="1" pageNumber="37" updateTime="1707169288913" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-NC-3.0">
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<mods:mods id="ECE88063015AD6A34ABB4E3E016D76A5" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="AB7179BC8575F36206749AC8744C6DEF">
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<mods:title id="2FCC2EFBE441251B9529A8B70A462F07">Supplementary Materials and Appendix</mods:title>
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</mods:titleInfo>
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<mods:name id="091EDA7737BE949110759B94C0463F3F" type="personal">
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<mods:role id="DB583A4FA8CFCD336721136332AA42D7">
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<mods:roleTerm id="4649E547D93D33F10E530B3FBECA7003">Author</mods:roleTerm>
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<mods:namePart id="47F14E7AFDEB644E1057F01CF8883ABD">Zhang, Jing</mods:namePart>
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<mods:affiliation id="03D9B248C4678D14EDEB26A7DF143067">McDermott Center for Human Growth and Development and Department of Biophysics University of Texas Southwestern Medical Center 5323 Harry Hines Blvd., Dallas, TX, 75390 - 8816 USA</mods:affiliation>
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<mods:name id="81854A23A12FDB69B3494EA8A27D03B9" type="personal">
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<mods:roleTerm id="D763CF0FD459A17FF00BAF5B7F6BC78F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E8365477917934D825B792F988E39ECC">Cong, Qian</mods:namePart>
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<mods:affiliation id="535562AC55A358F4269DE0DB732CF586">McDermott Center for Human Growth and Development and Department of Biophysics University of Texas Southwestern Medical Center 5323 Harry Hines Blvd., Dallas, TX, 75390 - 8816 USA</mods:affiliation>
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<mods:roleTerm id="276663E209A9A80F182015C5B4596644">Author</mods:roleTerm>
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<mods:namePart id="AEA558E6583714EAFB7651E247D4573A">Grishin, Nick V.</mods:namePart>
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<mods:affiliation id="33645A70F500F0F520278396890CDF01">Departments of Biophysics and Biochemistry University of Texas Southwestern Medical Center 5323 Harry Hines Blvd., Dallas, TX, 75390 - 9050 USA</mods:affiliation>
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<mods:typeOfResource id="CFC7E0DA7BEA0F6EAB9DE38917CC3C5D">text</mods:typeOfResource>
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<mods:titleInfo id="EB9DC2EB477AA7C9329A3447A22AA233">
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<mods:title id="E44B82E850A15CF80B4ECFA9F56CF8BB">Insecta Mundi</mods:title>
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</mods:titleInfo>
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<mods:part id="9F5E2B3D9FD40CE5001ECFAD24682037">
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<mods:date id="A02D59D7DA6D3274874D30BB43F03AA8">2023</mods:date>
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<mods:number id="DA505E8B0A68978CCA08F3A5936BF086">2023-12-29</mods:number>
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<mods:detail id="162CA0E20352EFBE134AA32BAD63361D" type="volume">
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<mods:number id="C5833911F2C30511CF9E1131A0FCB12A">2023</mods:number>
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<mods:number id="FB2C85DCD4944E61A0B3113B21344A13">26</mods:number>
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<mods:start id="124F2F3A2982FC730D0B02515D8B69A1">1</mods:start>
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<mods:url id="DA13D15758594700918DB94DA3F957CB">http://dx.doi.org/10.5281/zenodo.10396362</mods:url>
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<mods:identifier id="F12CD8D44BF05D95AA191CB94C116E73" type="DOI">10.5281/zenodo.10396362</mods:identifier>
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<mods:identifier id="6F86291596516585DD8EA14EDB310393" type="ISSN">1942-1354</mods:identifier>
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<treatment id="03810139FFF9BB77C0CAFC6DE2F3B312" LSID="urn:lsid:plazi:treatment:03810139FFF9BB77C0CAFC6DE2F3B312" httpUri="http://treatment.plazi.org/id/03810139FFF9BB77C0CAFC6DE2F3B312" lastPageId="39" lastPageNumber="38" pageId="38" pageNumber="37">
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<subSubSection id="C332E3A4FFF9BB76C0CAFC6DE0E5B118" pageId="38" pageNumber="37" type="nomenclature">
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<paragraph id="8B97B02FFFF9BB76C0CAFC6DE07EB13A" blockId="38.[192,944,827,924]" box="[192,811,827,856]" pageId="38" pageNumber="37">
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<heading id="D0DF0743FFF9BB76C0CAFC6DE07EB13A" box="[192,811,827,856]" fontSize="12" level="2" pageId="38" pageNumber="37" reason="2">
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<emphasis id="B95C6C3DFFF9BB76C0CAFC6DE07EB13A" bold="true" box="[192,811,827,856]" pageId="38" pageNumber="37">
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<taxonomicName id="4C28CBACFFF9BB76C0CAFC6DE1DEB13A" authority="Grishin" authorityName="Zhang & Cong & Grishin" authorityYear="2023" box="[192,651,827,856]" class="Insecta" family="Hesperiidae" genus="Heliopetes" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="37" phylum="Arthropoda" rank="species" species="lana" status="sp. nov." subGenus="Heliopetes">
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<emphasis id="B95C6C3DFFF9BB76C0CAFC6DE217B13A" bold="true" box="[192,322,827,856]" italics="true" pageId="38" pageNumber="37">Heliopetes</emphasis>
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(
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<emphasis id="B95C6C3DFFF9BB76C159FC6DE280B13A" bold="true" box="[339,469,827,856]" italics="true" pageId="38" pageNumber="37">Heliopetes</emphasis>
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)
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<emphasis id="B95C6C3DFFF9BB76C1ECFC6DE14AB13A" bold="true" box="[486,543,827,856]" italics="true" pageId="38" pageNumber="37">lana</emphasis>
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Grishin
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</taxonomicName>
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,
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<taxonomicNameLabel id="A26FD146FFF9BB76C29DFC6DE07EB13A" box="[663,811,827,856]" pageId="38" pageNumber="37" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="8B97B02FFFF9BB76C0CAFC09E0E5B118" blockId="38.[192,944,827,924]" box="[192,944,863,890]" pageId="38" pageNumber="37">
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https://zoobank.org/
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<uuid id="FF8E8AFAFFF9BB76C1ABFC09E0E5B118" box="[417,944,863,890]" pageId="38" pageNumber="37">2E6992E1-9CAE-4FF2-AD84-B73CB2C57F00</uuid>
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</paragraph>
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</subSubSection>
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<subSubSection id="C332E3A4FFF9BB76C0CAFCD7E157B1F9" box="[192,514,897,924]" pageId="38" pageNumber="37" type="description">
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<paragraph id="8B97B02FFFF9BB76C0CAFCD7E157B1F9" blockId="38.[192,944,827,924]" box="[192,514,897,924]" pageId="38" pageNumber="37">
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(
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<figureCitation id="1313ACAAFFF9BB76C0C2FCD7E252B1F9" box="[200,263,897,923]" captionStart="Figure 3" captionStartId="33.[192,261,1387,1411]" captionTargetBox="[193,1438,244,1339]" captionTargetId="figure-11@33.[192,1440,242,1340]" captionTargetPageId="33" captionText="Figure 3. Phylogenetic trees of remaining Pyrginae (excluding Achlyodini) inferred from protein-coding regions in a) the Z chromosome and b) the mitochondrial genome (in segments). See Fig. 1 legend for other notations." pageId="38" pageNumber="37">Fig. 3</figureCitation>
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part, 83–84, 302–303)
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</paragraph>
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</subSubSection>
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<subSubSection id="C332E3A4FFF9BB76C0CAFCFDE7D8B401" pageId="38" pageNumber="37" type="diagnosis">
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<paragraph id="8B97B02FFFF9BB76C0CAFCFDE299B7E0" blockId="38.[192,1442,939,1410]" pageId="38" pageNumber="37">
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<emphasis id="B95C6C3DFFF9BB76C0CAFCFDE2A1B1A7" bold="true" box="[192,500,939,965]" pageId="38" pageNumber="37">Definition and diagnosis.</emphasis>
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Phylogenetic analysis reveals that specimens from North America identified as
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<taxonomicName id="4C28CBACFFF9BB76C0CAFC9BE178B185" authority="(Reakirt, 1868)" baseAuthorityName="Reakirt" baseAuthorityYear="1868" box="[192,557,973,999]" class="Insecta" family="Hesperiidae" genus="Heliopetes" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="37" phylum="Arthropoda" rank="species" species="alana">
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<emphasis id="B95C6C3DFFF9BB76C0CAFC9BE22DB185" box="[192,376,973,999]" italics="true" pageId="38" pageNumber="37">Heliopetes alana</emphasis>
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(Reakirt, 1868)
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</taxonomicName>
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(type locality in
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<collectingCountry id="F33FF0BFFFF9BB76C2FEFC9BE033B185" box="[756,870,973,999]" name="Colombia" pageId="38" pageNumber="37">Colombia</collectingCountry>
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,
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<typeStatus id="54930E8DFFF9BB76C379FC98E09FB18A" box="[883,970,974,1000]" pageId="38" pageNumber="37" type="syntype">syntype</typeStatus>
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sequenced as NVG-15039C04, sequence completeness insufficient to add it to the Z chromosome tree) are not monophyletic with it and instead are sister to both
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<taxonomicName id="4C28CBACFFF9BB76C151FB45E2E9B64E" baseAuthorityName="Reakirt" baseAuthorityYear="1868" box="[347,444,1042,1068]" class="Insecta" family="Hesperiidae" genus="Heliopetes" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="37" phylum="Arthropoda" rank="species" species="alana">
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<emphasis id="B95C6C3DFFF9BB76C151FB45E2E9B64E" box="[347,444,1042,1068]" italics="true" pageId="38" pageNumber="37">H. alana</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C28CBACFFF9BB76C1F9FB44E013B64F" authority="Evans, 1953" authorityName="Evans" authorityYear="1953" box="[499,838,1042,1069]" class="Insecta" family="Hesperiidae" genus="Heliopetes" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="37" phylum="Arthropoda" rank="species" species="chimbo">
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<emphasis id="B95C6C3DFFF9BB76C1F9FB44E1EFB64E" box="[499,698,1042,1068]" italics="true" pageId="38" pageNumber="37">Heliopetes chimbo</emphasis>
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<bibRefCitation id="EFB9CDDEFFF9BB76C2CBFB44E013B64F" author="Evans WH" box="[705,838,1042,1069]" pageId="38" pageNumber="37" pagination="26 - 53" refId="ref77110" refString="Evans WH. 1953. A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part III. Pyrginae. Section 2. The Trustees of the British Museum (Natural History); London. v + 246 p., pl. 26 - 53." type="book chapter" year="1953">Evans, 1953</bibRefCitation>
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</taxonomicName>
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(type locality in
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<collectingCountry id="F33FF0BFFFF9BB76C40CFB44E73FB64E" box="[1030,1130,1042,1068]" name="Ecuador" pageId="38" pageNumber="37">Ecuador</collectingCountry>
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), exhibiting notable genetic differentiation (
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<figureCitation id="1313ACAAFFF9BB76C178FB62E2E1B62C" box="[370,436,1076,1102]" captionStart="Figure 3" captionStartId="33.[192,261,1387,1411]" captionTargetBox="[193,1438,244,1339]" captionTargetId="figure-11@33.[192,1440,242,1340]" captionTargetPageId="33" captionText="Figure 3. Phylogenetic trees of remaining Pyrginae (excluding Achlyodini) inferred from protein-coding regions in a) the Z chromosome and b) the mitochondrial genome (in segments). See Fig. 1 legend for other notations." pageId="38" pageNumber="37">Fig. 3</figureCitation>
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): e.g., COI barcodes of a specimen from
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<collectingCountry id="F33FF0BFFFF9BB76C380FB62E751B62C" box="[906,1028,1076,1102]" name="Guatemala" pageId="38" pageNumber="37">Guatemala</collectingCountry>
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and
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<collectingCountry id="F33FF0BFFFF9BB76C434FB62E7FBB62C" box="[1086,1198,1076,1102]" name="Colombia" pageId="38" pageNumber="37">Colombia</collectingCountry>
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differ by 2% (14 bp). All available names in synonymy with
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<taxonomicName id="4C28CBACFFF9BB76C289FB01E1B2B612" baseAuthorityName="Reakirt" baseAuthorityYear="1868" box="[643,743,1110,1136]" class="Insecta" family="Hesperiidae" genus="Heliopetes" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="37" phylum="Arthropoda" rank="species" species="alana">
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<emphasis id="B95C6C3DFFF9BB76C289FB01E1B2B612" box="[643,743,1110,1136]" italics="true" pageId="38" pageNumber="37">H. alana</emphasis>
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</taxonomicName>
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have type localities in South America. Therefore, the North American populations are a new species. This new species keys to
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<taxonomicName id="4C28CBACFFF9BB76C3ADFB2FE75DB6F0" baseAuthorityName="Reakirt" baseAuthorityYear="1868" box="[935,1032,1144,1170]" class="Insecta" family="Hesperiidae" genus="Heliopetes" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="37" phylum="Arthropoda" rank="species" species="alana">
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<emphasis id="B95C6C3DFFF9BB76C3ADFB2FE75DB6F0" box="[935,1032,1144,1170]" italics="true" pageId="38" pageNumber="37">H. alana</emphasis>
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</taxonomicName>
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(G.2.12) in
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<bibRefCitation id="EFB9CDDEFFF9BB76C485FB2FE677B6F1" author="Evans WH" box="[1167,1314,1144,1171]" pageId="38" pageNumber="37" pagination="26 - 53" refId="ref77110" refString="Evans WH. 1953. A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part III. Pyrginae. Section 2. The Trustees of the British Museum (Natural History); London. v + 246 p., pl. 26 - 53." type="book chapter" year="1953">Evans (1953)</bibRefCitation>
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and differs from its relatives by the following combination of characters: typically whiter (i.e., less yellow), especially on ventral hindwing (
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<figureCitation id="1313ACAAFFF9BB76C19FFBEAE2B3B6B5" box="[405,486,1212,1239]" captionStart="Figures 63–84" captionStartId="98.[192,271,1810,1834]" captionTargetBox="[196,1434,233,1782]" captionTargetId="figure-99@98.[192,1440,229,1789]" captionTargetPageId="98" captionText="Figures 63–84. Holotypes, data in text. 63–64) Myrinia aragua sp. n. 65–66) Myrinia maculosa sp. n. 67–68) Myrinia manchada sp. n. 69–70) Polyctor (Fenops) lamperus sp. n. 71–72) Nisoniades (Nisoniades) lutum sp. n. 73–74) Bolla (Stolla) vena sp. n. 75–76) Staphylus (Vulga) vula sp. n. 77–78) Staphylus (Vulga) vulga sp. n. 79–80) Staphylus (Staphylus) rotundalus sp. n. 81–82) Staphylus (Staphylus) yucatanus sp. n. 83–84) Heliopetes (Heliopetes) lana sp. n." pageId="38" pageNumber="37">Fig. 84</figureCitation>
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, but due to extensive variation hardly identifiable by facies), rounder in dorsal view anterior margin of tegumen, more extended harpe and, correspondingly, longer expansion of ampulla (
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<figureCitation id="1313ACAAFFF9BB76C57FFB89E270B779" captionStart="Figures 285–311" captionStartId="110.[192,271,1719,1743]" captionTargetBox="[205,1420,253,1687]" captionTargetId="figure-150@110.[192,1440,229,1702]" captionTargetPageId="110" captionText="Figures 285–311. Genitalia of holotypes (unless indicated), data in text.285–287) Polyctor (Fenops) lamperus sp. n.288–290) Nisoniades (Nisoniades) lutum sp. n. 291–292) Bolla (Stolla) vena sp. n. 293–294) Staphylus (Vulga) vula sp. n. 295–297) Staphylus (Vulga) vulga sp. n. 298–299) Staphylus (Staphylus) rotundalus sp. n. 300–301) Staphylus (Staphylus) yucatanus sp. n. paratype ♂ NVG-20063A08. 302–303) Heliopetes (Heliopetes) lana sp. n. paratype ♂ NVG-20062H06. 304–306) Canesia ella sp. n. 307–309) Paches (Paches) loxeca sp. n. 310–311) Clito congruens sp. n. Views: dorsal (285, 288, 291, 293, 295, 298, 300, 302, 305, 307, 310), left lateral (287, 290, 292, 294, 297, 299, 301, 303, 304 aedeagus, 306, 308, 309 aedeagus, 311), right lateral (286, 289, 296)." pageId="38" pageNumber="37">Fig. 302–303</figureCitation>
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). Due to the cryptic nature of this species, most reliable identification is achieved by DNA and a combination of the following base pairs is diagnostic in the nuclear genome: aly1079.16.3:T70C, aly173.27.12:C85T, aly173.27.12:A144G, aly276665.11.2:T224G, aly276665.11.2:A302T, and COI barcode: A229G, C235T, A289A, T397C, A477G, T557C.
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</paragraph>
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<paragraph id="8B97B02FFFF9BB76C0CAFAC7E7D8B401" blockId="38.[192,1440,1425,1635]" pageId="38" pageNumber="37">
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||
<emphasis id="B95C6C3DFFF9BB76C0CAFAC7E11BB7C9" bold="true" box="[192,590,1425,1451]" pageId="38" pageNumber="37">
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Barcode sequence of the
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<typeStatus id="54930E8DFFF9BB76C1EAFAC7E11CB7C9" box="[480,585,1425,1451]" pageId="38" pageNumber="37" type="holotype">holotype</typeStatus>
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.
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</emphasis>
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Sample NVG-17109G07, GenBank OR837660, 658 base pairs: AACTTTATATTTTATTTTTGGAATTTGAGCAGGAATAGTAGGTACTTCTTTAAGTTTATTAATTCGAACTGAATTAGGAAATCCAGGATCTTTAATT GGAGATGATCAAATTTATAATACTATTGTTACAGCCCATGCTTTTATTATAATTTTTTTCATAGTAATACCAATTATAATTGGAGGATTTGGAAATT GATTAGTACCTTTAATATTAGGAGCCCCAGATATGGCATTTCCTCGTATAAATAATATAAGATTTTGACTTTTACCCCCATCCCTAACATTATTAAT TTCAAGAAGTGTAGTAGAAAATGGAGCAGGAACTGGTTGAACAGTTTACCCCCCTCTCTCGGCTAATATTGCCCATCAAGGATCTTCTGTTGATTTA GCTATTTTCTCTTTACATTTAGCTGGAATTTCATCTATTTTAGGAGCTATTAATTTTATTACAACTATTATTAATATACGTATTAGAAGTATATCAT TTGATCAAATACCTTTATTTGTATGAGCGGTAGGAATTACTGCTTTATTACTACTATTATCATTACCTGTTCTAGCAGGTGCCATTACAATATTATT AACAGATCGAAATTTAAATACATCATTCTTCGATCCTGCTGGAGGAGGAGATCCTATTTTATATCAACATTTATTC
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C332E3A4FFF9BB77C0CAF923E14FB343" lastPageId="39" lastPageNumber="38" pageId="38" pageNumber="37" type="materials_examined">
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<paragraph id="8B97B02FFFF9BB76C0CAF923E265B53E" blockId="38.[191,1441,1653,1884]" pageId="38" pageNumber="37">
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||
<emphasis id="B95C6C3DFFF9BB76C0CAF923E23DB4ED" bold="true" box="[192,360,1653,1679]" pageId="38" pageNumber="37">Type material.</emphasis>
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<materialsCitation id="3B40BA72FFF9BB76C164F923E20EB57A" collectingDate="2003-06-05" collectingDateMax="2003-06-10" collectingDateMin="2003-06-05" collectionCode="CA, LACM, V" collectorName="Ron Leuschner. & Nick & Grishin" country="Guatemala" county="Los Angeles County Museum of Natural History" elevation="518" latitude="16.303888" location="Finca Ixobel" longLatPrecision="21" longitude="-89.42222" municipality="Los Angeles" pageId="38" pageNumber="37" specimenCount="2" specimenCount-male="2" stateProvince="Peten" typeStatus="holotype">
|
||
<emphasis id="B95C6C3DFFF9BB76C164F923E2B7B4ED" bold="true" box="[366,482,1653,1679]" pageId="38" pageNumber="37">
|
||
<typeStatus id="54930E8DFFF9BB76C164F923E288B4ED" box="[366,477,1653,1679]" pageId="38" pageNumber="37" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D2E7BA6FFF9BB76C1E2F920E2A8B4F0" box="[488,509,1654,1682]" count="1" pageId="38" pageNumber="37" type="male">♂</specimenCount>
|
||
deposited in the
|
||
<collectingCounty id="62F6C8A3FFF9BB76C2B6F920E782B4ED" box="[700,1239,1653,1680]" pageId="38" pageNumber="37">Los Angeles County Museum of Natural History</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6BF32A55FFF9BB76C4EBF920E63FB4ED" box="[1249,1386,1653,1680]" pageId="38" pageNumber="37">Los Angeles</collectingMunicipality>
|
||
,
|
||
<collectionCode id="ED3928EAFFF9BB76C57EF923E6C9B4ED" box="[1396,1436,1653,1679]" country="USA" name="Chicago Academy of Sciences" pageId="38" pageNumber="37" type="Museum">CA</collectionCode>
|
||
, USA (
|
||
<collectionCode id="ED3928EAFFF9BB76C108F9C1E203B4D3" box="[258,342,1687,1713]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="38" pageNumber="37" type="Museum">LACM</collectionCode>
|
||
), illustrated in
|
||
<figureCitation id="1313ACAAFFF9BB76C1F6F9C1E127B4D3" box="[508,626,1687,1713]" captionStart="Figures 63–84" captionStartId="98.[192,271,1810,1834]" captionTargetBox="[196,1434,233,1782]" captionTargetId="figure-99@98.[192,1440,229,1789]" captionTargetPageId="98" captionText="Figures 63–84. Holotypes, data in text. 63–64) Myrinia aragua sp. n. 65–66) Myrinia maculosa sp. n. 67–68) Myrinia manchada sp. n. 69–70) Polyctor (Fenops) lamperus sp. n. 71–72) Nisoniades (Nisoniades) lutum sp. n. 73–74) Bolla (Stolla) vena sp. n. 75–76) Staphylus (Vulga) vula sp. n. 77–78) Staphylus (Vulga) vulga sp. n. 79–80) Staphylus (Staphylus) rotundalus sp. n. 81–82) Staphylus (Staphylus) yucatanus sp. n. 83–84) Heliopetes (Heliopetes) lana sp. n." pageId="38" pageNumber="37">Fig. 83–84</figureCitation>
|
||
, bears the following three rectangular labels, two white: [
|
||
<collectingCountry id="F33FF0BFFFF9BB76C4FBF9C1E6C9B4D3" box="[1265,1436,1687,1713]" name="Guatemala" pageId="38" pageNumber="37">GUATEMALA</collectingCountry>
|
||
.
|
||
<collectingRegion id="49EC7ECDFFF9BB76C0CAF9ECE20DB4B1" box="[192,344,1721,1748]" country="Guatemala" name="Peten" pageId="38" pageNumber="37">Peten District</collectingRegion>
|
||
|
|
||
<location id="8EF7E6F4FFF9BB76C161F9EFE2A1B4B1" LSID="urn:lsid:plazi:treatment:03810139FFF9BB77C0CAFC6DE2F3B312:8EF7E6F4FFF9BB76C161F9EFE2A1B4B1" box="[363,500,1721,1747]" country="Guatemala" county="Los Angeles County Museum of Natural History" latitude="16.303888" longLatPrecision="21" longitude="-89.42222" municipality="Los Angeles" name="Finca Ixobel" pageId="38" pageNumber="37" stateProvince="Peten">Finca Ixobel</location>
|
||
<locationDeviation id="9A25A90DFFF9BB76C1F3F9EFE12CB4B6" box="[505,633,1721,1748]" location="Poptun" pageId="38" pageNumber="37">S of Poptun</locationDeviation>
|
||
|
|
||
<geoCoordinate id="EE1CD6E8FFF9BB76C280F9EFE048B4B6" box="[650,797,1721,1748]" degrees="16" direction="north" minutes="18" orientation="latitude" pageId="38" pageNumber="37" precision="15" seconds="14" value="16.303888">16° 18’ 14” N</geoCoordinate>
|
||
:
|
||
<geoCoordinate id="EE1CD6E8FFF9BB76C32CF9EFE0E8B4B6" box="[806,957,1721,1748]" degrees="89" direction="west" minutes="25" orientation="longitude" pageId="38" pageNumber="37" precision="15" seconds="20" value="-89.42222">89° 25’ 20” W</geoCoordinate>
|
||
|
|
||
<date id="FF9696EFFFF9BB76C3C5F9EFE7D2B4B1" box="[975,1159,1721,1748]" pageId="38" pageNumber="37" value="2003-06-05" valueMax="2003-06-10" valueMin="2003-06-05">
|
||
<collectingDate id="EFD26F07FFF9BB76C3C5F9EFE7D2B4B1" box="[975,1159,1721,1748]" pageId="38" pageNumber="37" value="2003-06-05" valueMax="2003-06-10" valueMin="2003-06-05">5-10 JUNE 2003</collectingDate>
|
||
</date>
|
||
:
|
||
<quantity id="4CD01DCAFFF9BB76C485F9EFE7B7B4B1" box="[1167,1250,1721,1747]" metricMagnitude="2" metricUnit="m" metricValue="5.1816" pageId="38" pageNumber="37" unit="ft" value="1700.0">
|
||
<elevation id="0005571CFFF9BB76C485F9EFE7B7B4B1" box="[1167,1250,1721,1747]" metricMagnitude="2" metricUnit="m" metricValue="5.1816" pageId="38" pageNumber="37" unit="ft" value="1700.0">1700 ft.</elevation>
|
||
</quantity>
|
||
|
|
||
<collectorName id="26DDD5F9FFF9BB76C4F9F9ECE6F4B4B1" box="[1267,1441,1721,1748]" pageId="38" pageNumber="37">Ron Leuschner.</collectorName>
|
||
coll.], [DNA sample ID: | NVG-17109G07 | c/o
|
||
<collectorName id="26DDD5F9FFF9BB76C2E0F98DE04AB497" box="[746,799,1755,1781]" pageId="38" pageNumber="37">Nick</collectorName>
|
||
<collectionCode id="ED3928EAFFF9BB76C32DF98AE06FB494" box="[807,826,1756,1782]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="38" pageNumber="37" type="Museum">V</collectionCode>
|
||
.
|
||
<collectorName id="26DDD5F9FFF9BB76C34EF98DE0CFB497" box="[836,922,1755,1781]" pageId="38" pageNumber="37">Grishin</collectorName>
|
||
], and one red [
|
||
<typeStatus id="54930E8DFFF9BB76C452F98AE7BCB494" box="[1112,1257,1756,1782]" pageId="38" pageNumber="37" type="holotype">HOLOTYPE</typeStatus>
|
||
<specimenCount id="9D2E7BA6FFF9BB76C4FAF98BE650B49B" box="[1264,1285,1757,1785]" count="1" pageId="38" pageNumber="37" type="male">♂</specimenCount>
|
||
|
|
||
<taxonomicName id="4C28CBACFFF9BB76C511F98DE6C4B497" authorityName="Billberg" authorityYear="1820" box="[1307,1425,1755,1781]" class="Insecta" family="Hesperiidae" genus="Heliopetes" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="37" phylum="Arthropoda" rank="genus">Heliopetes</taxonomicName>
|
||
|
|
||
<taxonomicName id="4C28CBACFFF9BB76C0CBF9A8E21EB57A" authority="Grishin" authorityName="Zhang & Cong & Grishin" authorityYear="2023" box="[193,331,1790,1816]" class="Insecta" family="Hesperiidae" genus="Heliopetes" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="37" phylum="Arthropoda" rank="species" species="lana">lana Grishin</taxonomicName>
|
||
]
|
||
</materialsCitation>
|
||
.
|
||
<emphasis id="B95C6C3DFFF9BB76C16FF9A8E28AB57A" bold="true" box="[357,479,1790,1816]" pageId="38" pageNumber="37">
|
||
<typeStatus id="54930E8DFFF9BB76C16FF9A8E28FB57A" box="[357,474,1790,1816]" pageId="38" pageNumber="37" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D2E7BA6FFF9BB76C1EFF9A8E149B579" box="[485,540,1790,1819]" count="2" pageId="38" pageNumber="37" type="male">2♂♂</specimenCount>
|
||
: 11-BOA-13383A02 the same data as the
|
||
<typeStatus id="54930E8DFFF9BB76C3EDF9A8E71CB57A" box="[999,1097,1790,1816]" pageId="38" pageNumber="37" type="holotype">holotype</typeStatus>
|
||
[
|
||
<collectionCode id="ED3928EAFFF9BB76C45CF9A8E7F9B57A" box="[1110,1196,1790,1816]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="38" pageNumber="37" type="Museum">USNM</collectionCode>
|
||
] and NVG-20062H06
|
||
<collectingCountry id="F33FF0BFFFF9BB76C0B5F876E247B558" box="[191,274,1824,1850]" name="Mexico" pageId="38" pageNumber="37">Mexico</collectingCountry>
|
||
:
|
||
<collectingRegion id="49EC7ECDFFF9BB76C12AF876E2F7B558" box="[288,418,1824,1850]" country="Mexico" name="Tamaulipas" pageId="38" pageNumber="37">Tamaulipas</collectingRegion>
|
||
,
|
||
<quantity id="4CD01DCAFFF9BB76C1A6F876E154B558" box="[428,513,1824,1850]" metricMagnitude="3" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="38" pageNumber="37" unit="km" value="2.5" valueMax="4.0" valueMin="1.0">1–4 km</quantity>
|
||
N of Gomez Farias,
|
||
<quantity id="4CD01DCAFFF9BB76C2ECF876E07BB558" box="[742,814,1824,1851]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="38" pageNumber="37" unit="m" value="350.0">350 m</quantity>
|
||
,
|
||
<date id="FF9696EFFFF9BB76C333F876E09BB558" box="[825,974,1824,1850]" pageId="38" pageNumber="37" value="1972-08-17">17-Aug-1972</date>
|
||
, C. J. Durden leg. (genitalia
|
||
<figureCitation id="1313ACAAFFF9BB76C50DF876E6CCB558" box="[1287,1433,1824,1850]" captionStart="Figures 285–311" captionStartId="110.[192,271,1719,1743]" captionTargetBox="[205,1420,253,1687]" captionTargetId="figure-150@110.[192,1440,229,1702]" captionTargetPageId="110" captionText="Figures 285–311. Genitalia of holotypes (unless indicated), data in text.285–287) Polyctor (Fenops) lamperus sp. n.288–290) Nisoniades (Nisoniades) lutum sp. n. 291–292) Bolla (Stolla) vena sp. n. 293–294) Staphylus (Vulga) vula sp. n. 295–297) Staphylus (Vulga) vulga sp. n. 298–299) Staphylus (Staphylus) rotundalus sp. n. 300–301) Staphylus (Staphylus) yucatanus sp. n. paratype ♂ NVG-20063A08. 302–303) Heliopetes (Heliopetes) lana sp. n. paratype ♂ NVG-20062H06. 304–306) Canesia ella sp. n. 307–309) Paches (Paches) loxeca sp. n. 310–311) Clito congruens sp. n. Views: dorsal (285, 288, 291, 293, 295, 298, 300, 302, 305, 307, 310), left lateral (287, 290, 292, 294, 297, 299, 301, 303, 304 aedeagus, 306, 308, 309 aedeagus, 311), right lateral (286, 289, 296)." pageId="38" pageNumber="37">Fig. 302–303</figureCitation>
|
||
) [
|
||
<collectionCode id="ED3928EAFFF9BB76C0CDF814E270B53E" box="[199,293,1858,1884]" country="USA" httpUri="http://grbio.org/cool/5vn9-fr8a" name="Texas Memorial Museum" pageId="38" pageNumber="37">TMMC</collectionCode>
|
||
].
|
||
</paragraph>
|
||
<paragraph id="8B97B02FFFF9BB77C0CAF83AE14FB343" blockId="38.[192,1403,1900,1927]" lastBlockId="39.[192,1440,230,290]" lastPageId="39" lastPageNumber="38" pageId="38" pageNumber="37">
|
||
<emphasis id="B95C6C3DFFF9BB76C0CAF83AE20EB5E4" bold="true" box="[192,347,1900,1926]" pageId="38" pageNumber="37">
|
||
<typeStatus id="54930E8DFFF9BB76C0CAF83AE3ACB5E4" box="[192,249,1900,1926]" pageId="38" pageNumber="37">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<collectingCountry id="F33FF0BFFFF9BB76C16BF83AE288B5E4" box="[353,477,1900,1926]" name="Guatemala" pageId="38" pageNumber="37">Guatemala</collectingCountry>
|
||
:
|
||
<collectingRegion id="49EC7ECDFFF9BB76C1EDF83BE1D1B5E4" box="[487,644,1900,1927]" country="Guatemala" name="Peten" pageId="38" pageNumber="37">Peten District</collectingRegion>
|
||
, Finca Ixobel S of Poptún, elevation
|
||
<quantity id="4CD01DCAFFF9BB76C413F83AE73DB5E4" box="[1049,1128,1900,1926]" metricMagnitude="2" metricUnit="m" metricValue="5.1816" pageId="38" pageNumber="37" unit="ft" value="1700.0">1700 ft</quantity>
|
||
,
|
||
<collectionCode id="ED3928EAFFF9BB76C478F83AE7F6B5E4" box="[1138,1187,1900,1926]" pageId="38" pageNumber="37">GPS</collectionCode>
|
||
<geoCoordinate id="EE1CD6E8FFF9BB76C4A3F83AE657B5E4" box="[1193,1282,1900,1926]" degrees="16.3039" direction="north" orientation="latitude" pageId="38" pageNumber="37" precision="5" value="16.3039">16.3039</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1CD6E8FFF9BB76C507F83AE62DB5E4" box="[1293,1400,1900,1926]" degrees="89.4222" direction="west" orientation="longitude" pageId="38" pageNumber="37" precision="5" value="-89.4222">−89.4222</geoCoordinate>
|
||
.
|
||
<emphasis id="B95C6C3DFFF8BB77C0CAFFB0E211B362" bold="true" box="[192,324,230,256]" pageId="39" pageNumber="38">Etymology.</emphasis>
|
||
The name removes a negating “a” from the name of its more southern sister species
|
||
<taxonomicName id="4C28CBACFFF8BB77C4FDFFB1E602B362" baseAuthorityName="Reakirt" baseAuthorityYear="1868" box="[1271,1367,230,256]" class="Insecta" family="Hesperiidae" genus="Heliopetes" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="alana">
|
||
<emphasis id="B95C6C3DFFF8BB77C4FDFFB1E602B362" box="[1271,1367,230,256]" italics="true" pageId="39" pageNumber="38">H. alana</emphasis>
|
||
</taxonomicName>
|
||
and is a feminine noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C332E3A4FFF8BB77C0CAFE62E2F3B312" pageId="39" pageNumber="38" type="distribution">
|
||
<paragraph id="8B97B02FFFF8BB77C0CAFE62E2F3B312" blockId="39.[192,1440,308,368]" pageId="39" pageNumber="38">
|
||
<emphasis id="B95C6C3DFFF8BB77C0CAFE62E20EB32C" bold="true" box="[192,347,308,334]" pageId="39" pageNumber="38">Distribution.</emphasis>
|
||
This species is currently known from
|
||
<collectingCountry id="F33FF0BFFFF8BB77C301FE62E00BB32C" box="[779,862,308,334]" name="Mexico" pageId="39" pageNumber="38">Mexico</collectingCountry>
|
||
and
|
||
<collectingCountry id="F33FF0BFFFF8BB77C392FE62E744B32C" box="[920,1041,308,334]" name="Guatemala" pageId="39" pageNumber="38">Guatemala</collectingCountry>
|
||
but is expected from other Central American countries.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |