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<document id="98F8B0D2EA9D6968AEB2842A00734BED" ID-CLB-Dataset="2598" ID-DOI="10.5852/ejt.2022.788.1625" ID-GBIF-Dataset="d6e6119f-4100-4cfc-9cc3-0d6951ee2c28" ID-ISSN="2118-9773" ID-Zenodo-Dep="5846636" ID-ZooBank="46C3CFCA-0CED-4432-AFD8-F4CFC1E0E1E7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1642092502222" checkinUser="felipe" docAuthor="Castillo, Stephanie, Rédei, Dávid &amp; Weirauch, Christiane" docDate="2022" docId="015D8E70FFC9FFC7FE09063DFDEEBDA0" docLanguage="en" docName="ejt-788_1-95.pdf" docOrigin="European Journal of Taxonomy 788 (1)" docSource="http://dx.doi.org/10.5852/ejt.2022.788.1625" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Voconia motoensis Castillo &amp; Rédei &amp; Weirauch 2022, comb. nov." docType="treatment" docVersion="10" lastPageNumber="68" masterDocId="FD64F608FF8BFF84FFE7031FFFACBA3D" masterDocTitle="Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs" masterLastPageNumber="95" masterPageNumber="1" pageNumber="67" updateTime="1699302197121" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="1F9C10BAC465C673E71C1E0A91FD9808">Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs</mods:title>
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<mods:affiliation id="1E406E8FCD33D4425F30635434D68EC3">University of California, Riverside, 900 University Avenue, Riverside, CA 92521, USA.</mods:affiliation>
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<mods:namePart id="17AA99991AD6C1EB34A161E6E3CE9607">Rédei, Dávid</mods:namePart>
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<mods:affiliation id="9525B7D01DD0B0A50D4706B9F7A680E4">National Chung Hsing University, 145 Xingda Rd., 40227 Taichung, Taiwan.</mods:affiliation>
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<mods:namePart id="5CC31E9609BF6B6BABCEBD00FA9FC1F3">Weirauch, Christiane</mods:namePart>
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<mods:affiliation id="06EBF8A50B6251882C985BE9E4471680">University of California, Riverside, 900 University Avenue, Riverside, CA 92521, USA.</mods:affiliation>
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<paragraph id="894B3F66FFC9FFC6FE09063DFBEABF01" blockId="66.[494,1094,1314,1375]" box="[494,1094,1314,1340]" pageId="66" pageNumber="67">
<heading id="D203880AFFC9FFC6FE09063DFBEABF01" box="[494,1094,1314,1340]" centered="true" fontSize="11" level="2" pageId="66" pageNumber="67" reason="2">
<taxonomicName id="4EF444E5FFC9FFC6FE09063DFC10BF01" authority="(Schouteden, 1929)" authorityName="Castillo &amp; Rédei &amp; Weirauch" authorityYear="2022" baseAuthorityName="Schouteden" baseAuthorityYear="1929" box="[494,956,1314,1340]" class="Insecta" family="Reduviidae" genus="Voconia" kingdom="Animalia" order="Hemiptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="motoensis" status="comb. nov.">
<emphasis id="BB80E374FFC9FFC6FE09063DFD7CBF01" bold="true" box="[494,720,1314,1340]" italics="true" pageId="66" pageNumber="67">Voconia motoensis</emphasis>
(
<bibRefCitation id="ED654297FFC9FFC6FD38063DFC18BF01" author="Schouteden H." box="[735,948,1314,1340]" pageId="66" pageNumber="67" pagination="250 - 262" refId="ref52018" refString="Schouteden H. 1929. Reduviides africains nouveaux. Revue de Zoologie et de Botanique africaines 18 (2): 250 - 262." type="journal article" year="1929">Schouteden, 1929</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A0B35E0FFFC9FFC6FC23063DFBEABF01" box="[964,1094,1314,1340]" pageId="66" pageNumber="67" rank="species">comb. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C1EE6CEDFFC9FFC6FD4B065AFC2BBF62" box="[684,903,1349,1375]" pageId="66" pageNumber="67" type="description">
<paragraph id="894B3F66FFC9FFC6FD4B065AFC2BBF62" blockId="66.[494,1094,1314,1375]" box="[684,903,1349,1375]" pageId="66" pageNumber="67">
<figureCitation id="11CF23E3FFC9FFC6FD4B065AFCB8BF62" box="[684,788,1349,1375]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="8.[190,233,1950,1976]" captionStartId-1="9.[189,232,1883,1909]" captionTargetBox-0="[204,1371,808,1898]" captionTargetBox-1="[199,1395,651,1845]" captionTargetId-0="figure-168@8.[189,1398,793,1908]" captionTargetId-1="figure-137@9.[189,1398,644,1845]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="Fig. 1. Strict consensus tree resulting from equal weighting analysis. Jackknife values over 51 are reported for tree one of eight." captionText-1="Fig. 2. Tree resulting from implied weighting analysis using K = 12 with characters unambiguously optimized. Symmetric resampling values over 51 are represented for trees obtained with K = 3, K = 6, K = 9, and K = 12 analyses. Characters and character states are described in pp. 1122. Specimens not to scale." figureDoi-0="http://doi.org/10.5281/zenodo.5846638" figureDoi-1="http://doi.org/10.5281/zenodo.5846640" httpUri-0="https://zenodo.org/record/5846638/files/figure.png" httpUri-1="https://zenodo.org/record/5846640/files/figure.png" pageId="66" pageNumber="67">Figs 12</figureCitation>
,
<figureCitation id="11CF23E3FFC9FFC6FCC7065AFC9CBF62" box="[800,816,1349,1375]" captionStart="Fig" captionStartId="33.[189,231,1954,1980]" captionTargetBox="[189,1398,597,1914]" captionTargetId="figure-95@33.[189,1398,597,1914]" captionTargetPageId="33" captionText="Fig. 8. Dorsal habitus of holotypes of Voconia Stål, 1866 species with distribution: purple = Afrotropical region, green = Australasian region, blue = Neotropical region, pink = Oriental region. Scale bars = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.5846652" httpUri="https://zenodo.org/record/5846652/files/figure.png" pageId="66" pageNumber="67">8</figureCitation>
,
<figureCitation id="11CF23E3FFC9FFC6FCD9065AFCF2BF62" box="[830,862,1349,1375]" captionStart="Fig" captionStartId="39.[188,231,1955,1981]" captionTargetBox="[189,1398,882,1916]" captionTargetId="figure-204@39.[189,1399,882,1916]" captionTargetPageId="39" captionText="Fig. 10. Lateral habitus of species of Voconia Stål, 1866. Scale bars = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.5846656" httpUri="https://zenodo.org/record/5846656/files/figure.png" pageId="66" pageNumber="67">10</figureCitation>
,
<figureCitation id="11CF23E3FFC9FFC6FC8D065AFC2BBF62" box="[874,903,1349,1375]" captionStart="Fig" captionStartId="85.[189,232,1988,2014]" captionTargetBox="[189,1398,872,1948]" captionTargetId="figure-209@85.[189,1398,872,1948]" captionTargetPageId="85" captionText="Fig. 17. The distribution of species of Voconia Stål, 1866 in Africa." figureDoi="http://doi.org/10.5281/zenodo.5846674" httpUri="https://zenodo.org/record/5846674/files/figure.png" pageId="66" pageNumber="67">17</figureCitation>
</paragraph>
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<subSubSection id="C1EE6CEDFFC9FFC6FF5A0698FB7FBFF9" pageId="66" pageNumber="67" type="reference_group">
<paragraph id="894B3F66FFC9FFC6FF5A0698FD72BF9C" blockId="66.[189,1235,1414,1476]" box="[189,734,1414,1441]" pageId="66" pageNumber="67">
<treatmentCitationGroup id="A9E41848FFC9FFC6FF5A0698FD72BF9C" box="[189,734,1414,1441]" pageId="66" pageNumber="67">
<treatmentCitation id="08551977FFC9FFC6FF5A0698FD76BF9C" author="Schouteden H." box="[189,730,1414,1441]" page="252" pageId="66" pageNumber="67" year="1929">
<taxonomicName id="4EF444E5FFC9FFC6FF5A0698FD76BF9C" ID-CoL="439L5" authority="Schouteden, 1929: 252" authorityName="Schouteden" authorityPageNumber="252" authorityYear="1929" box="[189,730,1414,1441]" class="Insecta" family="Reduviidae" genus="Microvarus" kingdom="Animalia" order="Hemiptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="motoensis">
<emphasis id="BB80E374FFC9FFC6FF5A0698FE6DBF9D" box="[189,449,1415,1440]" italics="true" pageId="66" pageNumber="67">Microvarus motoensis</emphasis>
<bibRefCitation id="ED654297FFC9FFC6FE2E0699FD76BF9C" author="Schouteden H." box="[457,730,1414,1441]" pageId="66" pageNumber="67" pagination="250 - 262" refId="ref52018" refString="Schouteden H. 1929. Reduviides africains nouveaux. Revue de Zoologie et de Botanique africaines 18 (2): 250 - 262." type="journal article" year="1929">Schouteden, 1929: 252</bibRefCitation>
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.
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</paragraph>
<paragraph id="894B3F66FFC9FFC6FF5A06B5FB7FBFF9" blockId="66.[189,1235,1414,1476]" box="[189,1235,1450,1476]" pageId="66" pageNumber="67">
<treatmentCitationGroup id="A9E41848FFC9FFC6FF5A06B5FB7FBFF9" box="[189,1235,1450,1476]" pageId="66" pageNumber="67">
<treatmentCitation id="08551977FFC9FFC6FF5A06B5FD1DBFF9" author="Villiers A." box="[189,689,1450,1476]" page="107" pageId="66" pageNumber="67" year="1964">
<taxonomicName id="4EF444E5FFC9FFC6FF5A06B5FD1DBFF9" authority="Villiers, 1964: 107" authorityName="Villiers" authorityPageNumber="107" authorityYear="1964" box="[189,689,1450,1476]" class="Insecta" family="Reduviidae" genus="Microvarus" kingdom="Animalia" order="Hemiptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="inermiceps">
<emphasis id="BB80E374FFC9FFC6FF5A06B5FE61BFFE" box="[189,461,1450,1475]" italics="true" pageId="66" pageNumber="67">Microvarus inermiceps</emphasis>
<bibRefCitation id="ED654297FFC9FFC6FE3306B5FD1DBFF9" author="Villiers A." box="[468,689,1450,1476]" pageId="66" pageNumber="67" refId="ref52304" refString="Villiers A. 1964. Reduviidae (Hemiptera Heteroptera). In: Exploration du Parc National de la Garamba. Mission H. de Saeger, en collaboration avec P. Baert, G. Demoulin, I. Denisoff, J. Martin, M. Micha, A. Noirfalise, P. Schoemaker, G. Troupin et J. Verschuren (1949 - 1952), Fasc. 43. Institut des Parcs Nationaux du Congo et du Rwanda, Bruxelles." type="book" year="1964">Villiers, 1964: 107</bibRefCitation>
</taxonomicName>
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, fig. 27. Synonymized by
<treatmentCitation id="08551977FFC9FFC6FC0306B5FB62BFF9" author="Villiers A." box="[996,1230,1450,1476]" page="171" pageId="66" pageNumber="67" year="1976">
<bibRefCitation id="ED654297FFC9FFC6FC0306B5FB62BFF9" author="Villiers A." box="[996,1230,1450,1476]" pageId="66" pageNumber="67" pagination="162 - 176" refId="ref52391" refString="Villiers A. 1976. Descriptions, localisations et synonymies d'Hemipteres Reduviidae du Musee Royal de l'Afrique Centrale. Revue de Zoologie africaine 90 (1): 162 - 176." type="journal article" year="1976">Villiers (1976: 171)</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="C1EE6CEDFFC9FFC6FF5A06F4FBA4BCD9" pageId="66" pageNumber="67" type="diagnosis">
<paragraph id="894B3F66FFC9FFC6FF5A06F4FE36BC38" blockId="66.[189,410,1515,1541]" box="[189,410,1515,1541]" pageId="66" pageNumber="67">
<heading id="D203880AFFC9FFC6FF5A06F4FE36BC38" bold="true" box="[189,410,1515,1541]" fontSize="11" level="3" pageId="66" pageNumber="67" reason="3">
<emphasis id="BB80E374FFC9FFC6FF5A06F4FE36BC38" bold="true" box="[189,410,1515,1541]" pageId="66" pageNumber="67">Revised diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="894B3F66FFC9FFC6FF5A0505FBA4BCD9" blockId="66.[188,1398,1562,1764]" pageId="66" pageNumber="67">
Distinguished from other species in
<taxonomicName id="4EF444E5FFC9FFC6FDBA0505FD10BC0E" authorityName="Stal" authorityYear="1866" box="[605,700,1562,1587]" class="Insecta" family="Reduviidae" genus="Voconia" kingdom="Animalia" order="Hemiptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="BB80E374FFC9FFC6FDBA0505FD10BC0E" box="[605,700,1562,1587]" italics="true" pageId="66" pageNumber="67">Voconia</emphasis>
</taxonomicName>
by its finely granulose head; short labial segment I that does not reach posteroventral eye margin; anterior margin of buccula with dense, long macrosetae; and stout body. This species most closely resembles
<taxonomicName id="4EF444E5FFC9FFC6FD54057EFC9FBC47" authorityName="Jeannel" authorityYear="1917" box="[691,819,1632,1658]" class="Insecta" family="Reduviidae" genus="Microvarus" kingdom="Animalia" order="Hemiptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="conradti">
<emphasis id="BB80E374FFC9FFC6FD54057EFC9FBC47" box="[691,819,1632,1658]" italics="true" pageId="66" pageNumber="67">V. conradti</emphasis>
</taxonomicName>
and
<taxonomicName id="4EF444E5FFC9FFC6FC8A057EFC49BC47" authority="Castillo &amp; Rédei &amp; Weirauch, 2022" authorityName="Castillo &amp; Rédei &amp; Weirauch" authorityYear="2022" box="[877,997,1632,1658]" class="Insecta" family="Reduviidae" genus="Voconia" kingdom="Animalia" order="Hemiptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="smithae" status="sp. nov.">
<emphasis id="BB80E374FFC9FFC6FC8A057EFC49BC47" box="[877,997,1632,1658]" italics="true" pageId="66" pageNumber="67">V. smithae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A0B35E0FFFC9FFC6FC0B057EFBEBBC46" box="[1004,1095,1633,1659]" pageId="66" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
in its stout body plan and Afrotropical distribution. It is easily recognized from these two species by the absent or vestigial fossula spongiosa on the midleg and its distinct coloration: uniformly brown scutellum, abducted corium with an anteromedial yellow spot, and uniformly brown dorsal laterotergites.
</paragraph>
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<heading id="D203880AFFC9FFC6FF5A0414FEC0BD18" bold="true" box="[189,364,1803,1829]" fontSize="11" level="3" pageId="66" pageNumber="67" reason="3">
<emphasis id="BB80E374FFC9FFC6FF5A0414FEC0BD18" bold="true" box="[189,364,1803,1829]" pageId="66" pageNumber="67">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="894B3F66FFC9FFC6FF0D0425FB43BDA7" blockId="66.[189,1398,1850,1947]" pageId="66" pageNumber="67">
<materialsCitation id="399C353BFFC9FFC6FF0D0425FB43BDA7" ID-GBIF-Occurrence="3436573353" collectionCode="UCR, RMCA" collectorName="L. Burgeon" country="Democratic Republic of the Congo" elevation="898" latitude="2.90556" location="Haut-Uele" longLatPrecision="55" longitude="29.415" municipality="Moto" pageId="66" pageNumber="67" specimenCode="UCR_ENT 00070525, RMCA ENT 000018004" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
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<typeStatus id="564F81C4FFC9FFC6FF0D0425FEF0BD69" box="[234,348,1850,1876]" pageId="66" pageNumber="67" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F1E37FF6FFC9FFC6FF5A0442FCAABD4A" box="[189,774,1885,1911]" name="Democratic Republic of the Congo" pageId="66" pageNumber="67">DEMOCRATIC REPUBLIC OF THE CONGO</collectingCountry>
<specimenCount id="9FF2F4EFFFC9FFC6FCCB0442FCEFBD4A" box="[812,835,1885,1911]" pageId="66" pageNumber="67" type="female"></specimenCount>
;
<location id="8C2B69BDFFC9FFC6FCB00442FC78BD4A" LSID="urn:lsid:plazi:treatment:015D8E70FFC9FFC7FE09063DFDEEBDA0:8C2B69BDFFC9FFC6FCB00442FC78BD4A" box="[855,980,1885,1911]" country="Democratic Republic of the Congo" latitude="2.90556" longLatPrecision="55" longitude="29.415" municipality="Moto" name="Haut-Uele" pageId="66" pageNumber="67">Haut-Uélé</location>
,
<collectingMunicipality id="692FA51CFFC9FFC6FC010442FB85BD4A" box="[998,1065,1885,1911]" pageId="66" pageNumber="67">Moto</collectingMunicipality>
; [
<geoCoordinate id="ECC059A1FFC9FFC6FBA50442FB7ABD4A" box="[1090,1238,1885,1912]" degrees="2.90556" direction="north" orientation="latitude" pageId="66" pageNumber="67" precision="1" value="2.90556">
2.90556
<emphasis id="BB80E374FFC9FFC6FB420442FB1DBD4A" bold="true" box="[1189,1201,1885,1911]" pageId="66" pageNumber="67">°</emphasis>
N
</geoCoordinate>
,
<geoCoordinate id="ECC059A1FFC9FFC6FB0F0442FAC4BD4A" box="[1256,1384,1884,1912]" degrees="29.415" direction="east" orientation="longitude" pageId="66" pageNumber="67" precision="55" value="29.415">
29.415
<emphasis id="BB80E374FFC9FFC6FADE0442FAE9BD4A" bold="true" box="[1337,1349,1885,1911]" pageId="66" pageNumber="67">°</emphasis>
E
</geoCoordinate>
]; [elev.
<quantity id="4E0C9283FFC9FFC6FEE3049FFEF8BDA7" box="[260,340,1919,1946]" metricMagnitude="2" metricUnit="m" metricValue="8.98" pageId="66" pageNumber="67" unit="m" value="898.0">
<elevation id="02D9D855FFC9FFC6FEE3049FFEF8BDA7" box="[260,340,1919,1946]" metricMagnitude="2" metricUnit="m" metricValue="8.98" pageId="66" pageNumber="67" unit="m" value="898.0">898 m</elevation>
</quantity>
]; 1920;
<collectorName id="24015AB0FFC9FFC6FE55049EFD9ABDA6" box="[434,566,1921,1947]" pageId="66" pageNumber="67">L. Burgeon</collectorName>
leg.; USI:
<specimenCode id="D952971DFFC9FFC6FD5F049FFC17BDA6" box="[696,955,1920,1947]" pageId="66" pageNumber="67">UCR_ENT 00070525</specimenCode>
;
<specimenCode id="D952971DFFC9FFC6FC20049FFB47BDA7" box="[967,1259,1920,1947]" pageId="66" pageNumber="67">RMCA ENT 000018004</specimenCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C1EE6CEDFFC8FFC7FF5A0217FE95BC6E" pageId="67" pageNumber="68" type="description">
<paragraph id="894B3F66FFC8FFC7FF5A0217FEC7BB1F" blockId="67.[189,363,264,290]" box="[189,363,264,290]" pageId="67" pageNumber="68">
<heading id="D203880AFFC8FFC7FF5A0217FEC7BB1F" bold="true" box="[189,363,264,290]" fontSize="11" level="3" pageId="67" pageNumber="68" reason="3">
<emphasis id="BB80E374FFC8FFC7FF5A0217FEC7BB1F" bold="true" box="[189,363,264,290]" pageId="67" pageNumber="68">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="894B3F66FFC8FFC7FF0D0228FE79BB6C" blockId="67.[189,724,311,373]" box="[234,469,311,337]" pageId="67" pageNumber="68">
<emphasis id="BB80E374FFC8FFC7FF0D0228FEE9BB6C" bold="true" box="[234,325,311,337]" pageId="67" pageNumber="68">Female</emphasis>
(
<figureCitation id="11CF23E3FFC8FFC7FEB20228FE0DBB6C" box="[341,417,311,337]" captionStart="Fig" captionStartId="33.[189,231,1954,1980]" captionTargetBox="[189,1398,597,1914]" captionTargetId="figure-95@33.[189,1398,597,1914]" captionTargetPageId="33" captionText="Fig. 8. Dorsal habitus of holotypes of Voconia Stål, 1866 species with distribution: purple = Afrotropical region, green = Australasian region, blue = Neotropical region, pink = Oriental region. Scale bars = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.5846652" httpUri="https://zenodo.org/record/5846652/files/figure.png" pageId="67" pageNumber="68">Figs 8</figureCitation>
,
<figureCitation id="11CF23E3FFC8FFC7FE490228FE60BB6C" box="[430,460,311,337]" captionStart="Fig" captionStartId="39.[188,231,1955,1981]" captionTargetBox="[189,1398,882,1916]" captionTargetId="figure-204@39.[189,1399,882,1916]" captionTargetPageId="39" captionText="Fig. 10. Lateral habitus of species of Voconia Stål, 1866. Scale bars = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.5846656" httpUri="https://zenodo.org/record/5846656/files/figure.png" pageId="67" pageNumber="68">10</figureCitation>
)
</paragraph>
<paragraph id="894B3F66FFC8FFC7FF5A0244FD78BB48" blockId="67.[189,724,311,373]" box="[189,724,346,373]" pageId="67" pageNumber="68">
BODY LENGTH. About
<quantity id="4E0C9283FFC8FFC7FE510245FD8DBB48" box="[438,545,346,373]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" pageId="67" pageNumber="68" unit="mm" value="11.6">11.6 mm</quantity>
; macropterous.
</paragraph>
<paragraph id="894B3F66FFC8FFC7FF5A0283FC7CB85B" blockId="67.[189,1399,412,614]" pageId="67" pageNumber="68">COLORATION. Head: dark brown; labium nearly as dark as head. Thorax: coloration as head; posterior pronotal lobe with posterior margin lighter; scutellum uniformly dark brown. Hemelytron (abducted): clavus dark reddish-brown with distal yellow stripe; corium dark reddish-brown with anteromedial and posteromedial yellow spots; membrane dark with pale V-shaped marking along R and M veins; membranal veins proximally pale. Legs: yellowish-brown with distal half of femora dark brown. Abdomen: dark brown; dorsal laterotergites uniformly dark brown.</paragraph>
<paragraph id="894B3F66FFC8FFC7FF5A0191FE39B96A" blockId="67.[188,1399,653,855]" pageId="67" pageNumber="68">INTEGUMENT AND VESTITURE. Head and pronotum: finely granulose with sparse, short macrosetae interspersed among dense pubescence; antennifer with macroseta, base not protruded; morphologically ventral surface of labium with dense, short macrosetae. Thorax: anterolateral angles of pronotal collar with macrosetae. Hemelytron: corium with sparse long setation. Legs: posterior row of protuberances on mid and hind femora with three large spines on distal half, middle spine much longer. Abdomen: ventral surface pubescent.</paragraph>
<paragraph id="894B3F66FFC8FFC7FF5A0060FE19BFD2" blockId="67.[187,1401,894,1519]" pageId="67" pageNumber="68">STRUCTURE. Head: elongate, about 1.2 times as long as wide; anteocular region about one quarter of head length, shorter than postocular region (measured to anterior margin of neck); postocular region in dorsal view shorter than eye, lateral margins gently rounded; pedicel about 1.5 times length of head width; maxillary plates ellipsoidal, adjacent to and longer than clypeus; apices of maxillary plates in dorsal view directed straight; clypeus in dorsal view wider than maxillary plates; clypeal apex round, not narrowed; interocular glabrous markings anterolaterally curved, joined at interocular sulcus paramedially; interocular sulcus in dorsal view bent anteromedially, subtriangular; eye width in dorsal view wider than synthlipsis, eye not reaching ventral head margin in lateral view; ventrolateral swelling of buccula without lateral protrusion that surpasses buccular margin, flat margin; labial segment I in lateral view straight, not reaching posteroventral eye margin; morphologically dorsal surface of labial segment II nearly straight, about 0.7 times length of segment I. Thorax: pronotal collar wide medially with anterolateral angles short, projected forward; anterior pronotal lobe about 0.7 times length of posterior pronotal lobe, lateral margins almost straight; glabrous markings on pronotum thin and not deeply depressed; median apodeme depression of pronotum deep, circular; scutellar spine long, raised; anteriad-directed process of prosternum smoothly rounded, without paramedial lobes; anterior margin of stridulitrum not elongated into protuberance; proepimeron with smoothly rounded posteroventral margin. Legs: fossula spongiosa present on fore leg, absent or vestigial on mid leg.</paragraph>
<paragraph id="894B3F66FFC8FFC7FF0D0509FE86BC0D" blockId="67.[189,313,1558,1619]" box="[234,298,1558,1584]" pageId="67" pageNumber="68">
<emphasis id="BB80E374FFC8FFC7FF0D0509FE86BC0D" bold="true" box="[234,298,1558,1584]" pageId="67" pageNumber="68">Male</emphasis>
</paragraph>
<paragraph id="894B3F66FFC8FFC7FF5A0526FE95BC6E" blockId="67.[189,313,1558,1619]" box="[189,313,1593,1619]" pageId="67" pageNumber="68">Unknown.</paragraph>
</subSubSection>
<subSubSection id="C1EE6CEDFFC8FFC7FF5A0564FDCDBCDA" pageId="67" pageNumber="68" type="distribution">
<paragraph id="894B3F66FFC8FFC7FF5A0564FE68BCA8" blockId="67.[189,452,1659,1685]" box="[189,452,1659,1685]" pageId="67" pageNumber="68">
<emphasis id="BB80E374FFC8FFC7FF5A0564FEFABCA8" bold="true" box="[189,342,1659,1685]" pageId="67" pageNumber="68">Distribution</emphasis>
(
<figureCitation id="11CF23E3FFC8FFC7FE810564FE17BCA8" box="[358,443,1659,1685]" captionStart="Fig" captionStartId="85.[189,232,1988,2014]" captionTargetBox="[189,1398,872,1948]" captionTargetId="figure-209@85.[189,1398,872,1948]" captionTargetPageId="85" captionText="Fig. 17. The distribution of species of Voconia Stål, 1866 in Africa." figureDoi="http://doi.org/10.5281/zenodo.5846674" httpUri="https://zenodo.org/record/5846674/files/figure.png" pageId="67" pageNumber="68">Fig. 17</figureCitation>
)
</paragraph>
<paragraph id="894B3F66FFC8FFC7FF5A05B6FDCDBCDA" blockId="67.[189,1399,1705,1767]" pageId="67" pageNumber="68">
Only known from Moto and Garamba National Park (
<bibRefCitation id="ED654297FFC8FFC7FCA505B6FC73BCF9" author="Villiers A." box="[834,991,1705,1732]" pageId="67" pageNumber="68" refId="ref52304" refString="Villiers A. 1964. Reduviidae (Hemiptera Heteroptera). In: Exploration du Parc National de la Garamba. Mission H. de Saeger, en collaboration avec P. Baert, G. Demoulin, I. Denisoff, J. Martin, M. Micha, A. Noirfalise, P. Schoemaker, G. Troupin et J. Verschuren (1949 - 1952), Fasc. 43. Institut des Parcs Nationaux du Congo et du Rwanda, Bruxelles." type="book" year="1964">Villiers 1964</bibRefCitation>
) in the Haut-Uélé province of
<collectingCountry id="F1E37FF6FFC8FFC7FAB405B6FDF1BCDA" name="Democratic Republic of the Congo" pageId="67" pageNumber="68">the Democratic Republic of the Congo</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C1EE6CEDFFC8FFC7FF5A0411FDEEBDA0" pageId="67" pageNumber="68" type="discussion">
<paragraph id="894B3F66FFC8FFC7FF5A0411FE83BD15" blockId="67.[189,303,1806,1832]" box="[189,303,1806,1832]" pageId="67" pageNumber="68">
<heading id="D203880AFFC8FFC7FF5A0411FE83BD15" bold="true" box="[189,303,1806,1832]" fontSize="11" level="3" pageId="67" pageNumber="68" reason="3">
<emphasis id="BB80E374FFC8FFC7FF5A0411FE83BD15" bold="true" box="[189,303,1806,1832]" pageId="67" pageNumber="68">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="894B3F66FFC8FFC7FF5A0422FDEEBDA0" blockId="67.[189,1399,1853,1950]" pageId="67" pageNumber="68">
The carinules on the dorsal surface of the abdomen were not examined to prevent damage to the
<typeStatus id="564F81C4FFC8FFC7FAEF0422FADFBD6A" box="[1288,1395,1853,1879]" pageId="67" pageNumber="68" type="holotype">holotype</typeStatus>
. The synonymy of
<taxonomicName id="4EF444E5FFC8FFC7FE70047EFD0BBD47" authorityName="Villiers" authorityYear="1964" box="[407,679,1889,1914]" class="Insecta" family="Reduviidae" genus="Microvarus" kingdom="Animalia" order="Hemiptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="inermiceps">
<emphasis id="BB80E374FFC8FFC7FE70047EFD0BBD47" box="[407,679,1889,1914]" italics="true" pageId="67" pageNumber="68">Microvarus inermiceps</emphasis>
</taxonomicName>
proposed by
<bibRefCitation id="ED654297FFC8FFC7FCAF047FFB9EBD47" author="Villiers A." box="[840,1074,1888,1914]" pageId="67" pageNumber="68" pagination="162 - 176" refId="ref52391" refString="Villiers A. 1976. Descriptions, localisations et synonymies d'Hemipteres Reduviidae du Musee Royal de l'Afrique Centrale. Revue de Zoologie africaine 90 (1): 162 - 176." type="journal article" year="1976">Villiers (1976: 171)</bibRefCitation>
was accepted without a reexamination of the type material.
</paragraph>
</subSubSection>
</treatment>
</document>