192 lines
20 KiB
XML
192 lines
20 KiB
XML
<document ID-DOI="10.5733/afin.056.0101" ID-GBIF-Dataset="41a28e23-93d6-4070-a683-157c38cc0413" ID-ISSN="2305-2562" ID-Zenodo-Dep="7670381" checkinTime="1676640526494" checkinUser="felipe" docAuthor="Gerstmeier, Roland" docDate="2015" docId="03F5D02DA552A020FD835FC8FCF8FA97" docLanguage="en" docName="AfrInvertebr.56.1.19-24.pdf" docOrigin="African Invertebrates 56 (1)" docSource="http://www.bioone.org/doi/10.5733/afin.056.0101" docStyle="DocumentStyle:B78C63EC8CEBBB75364A82B35BB1F83D.2:AfrInvertebr.2008-2015.journal_article.0cover" docStyleId="B78C63EC8CEBBB75364A82B35BB1F83D" docStyleName="AfrInvertebr.2008-2015.journal_article.0cover" docStyleVersion="2" docTitle="Nonalatus epigaeus Gerstmeier 2015, sp. n." docType="treatment" docVersion="3" lastPageNumber="23" masterDocId="FFCCA855A553A024FF835A41FFF0FFF0" masterDocTitle="Nonalatus epigaeus sp. n., a new apterous species of Cleridae from Tanzania (Coleoptera: Cleroidea)" masterLastPageNumber="24" masterPageNumber="19" pageNumber="20" updateTime="1677158216855" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Nonalatus epigaeus sp. n., a new apterous species of Cleridae from Tanzania (Coleoptera: Cleroidea)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Gerstmeier, Roland</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>African Invertebrates</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2015</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2015-01-21</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>56</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>19</mods:start>
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<mods:end>24</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://www.bioone.org/doi/10.5733/afin.056.0101</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.5733/afin.056.0101</mods:identifier>
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<mods:identifier type="GBIF-Dataset">41a28e23-93d6-4070-a683-157c38cc0413</mods:identifier>
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<mods:identifier type="ISSN">2305-2562</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7670381</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7670450" ID-Zenodo-Dep="7670450" LSID="urn:lsid:plazi:treatment:03F5D02DA552A020FD835FC8FCF8FA97" httpUri="http://treatment.plazi.org/id/03F5D02DA552A020FD835FC8FCF8FA97" lastPageId="4" lastPageNumber="23" pageId="1" pageNumber="20">
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<subSubSection box="[512,810,1417,1443]" pageId="1" pageNumber="20" type="nomenclature">
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<paragraph blockId="1.[181,1142,1417,1676]" box="[512,810,1417,1443]" pageId="1" pageNumber="20">
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<heading box="[512,810,1417,1443]" centered="true" fontSize="11" level="2" pageId="1" pageNumber="20" reason="2">
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<taxonomicName authority="Gerstmeier, 2015" authorityName="Gerstmeier" authorityYear="2015" box="[512,744,1417,1443]" class="Insecta" family="Cleridae" genus="Nonalatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="20" phylum="Arthropoda" rank="species" species="epigaeus" status="sp. n.">
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<emphasis bold="true" box="[512,744,1417,1443]" pageId="1" pageNumber="20">Nonalatus epigaeus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[751,810,1417,1443]" pageId="1" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="4" lastPageNumber="23" pageId="1" pageNumber="20" type="description">
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<paragraph blockId="1.[181,1142,1417,1676]" box="[612,710,1457,1483]" pageId="1" pageNumber="20">
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<figureCitation box="[612,677,1457,1483]" captionStart="Fig" captionStartId="2.[432,465,1137,1159]" captionTargetBox="[369,878,209,1116]" captionTargetId="figure-186@2.[354,892,201,1125]" captionTargetPageId="2" captionText="Fig. 1. Nonalatus epigaeus sp. n., habitus." figureDoi="http://doi.org/10.5281/zenodo.7670383" httpUri="https://zenodo.org/record/7670383/files/figure.png" pageId="1" pageNumber="20">Figs 1</figureCitation>
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–9
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</paragraph>
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<paragraph blockId="1.[181,1142,1417,1676]" pageId="1" pageNumber="20">
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Etymology: From the Latin adjective
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<taxonomicName authorityName="Gerstmeier" authorityYear="2015" box="[607,707,1498,1523]" class="Insecta" family="Cleridae" genus="Nonalatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="20" phylum="Arthropoda" rank="species" species="epigaeus">
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<emphasis box="[607,707,1498,1523]" italics="true" pageId="1" pageNumber="20">epigaeus</emphasis>
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</taxonomicName>
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, referring to its presumable biology in forest litter.
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</paragraph>
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<paragraph blockId="1.[181,1142,1417,1676]" box="[181,321,1569,1595]" pageId="1" pageNumber="20">Description:</paragraph>
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<paragraph blockId="1.[181,1142,1417,1676]" box="[181,247,1609,1635]" pageId="1" pageNumber="20">
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<emphasis box="[181,247,1609,1635]" italics="true" pageId="1" pageNumber="20">Male.</emphasis>
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</paragraph>
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<paragraph blockId="1.[181,1142,1417,1676]" box="[181,532,1649,1676]" pageId="1" pageNumber="20">
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<emphasis box="[181,226,1649,1675]" italics="true" pageId="1" pageNumber="20">Size</emphasis>
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: Body length
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<quantity box="[386,527,1649,1676]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" metricValueMax="6.6" metricValueMin="5.2" pageId="1" pageNumber="20" unit="mm" value="5.9" valueMax="6.6" valueMin="5.2">5.2–6.6 mm</quantity>
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.
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</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.7670383" ID-Zenodo-Dep="7670383" box="[432,815,1137,1159]" httpUri="https://zenodo.org/record/7670383/files/figure.png" pageId="2" pageNumber="21" startId="2.[432,465,1137,1159]" targetBox="[369,878,209,1116]" targetPageId="2">
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<paragraph blockId="2.[432,815,1137,1159]" box="[432,815,1137,1159]" pageId="2" pageNumber="21">
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Fig. 1.
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<taxonomicName authority="Gerstmeier, 2015" authorityName="Gerstmeier" authorityYear="2015" box="[496,676,1137,1159]" class="Insecta" family="Cleridae" genus="Nonalatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="21" phylum="Arthropoda" rank="species" species="epigaeus" status="sp. n.">
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<emphasis box="[496,676,1137,1159]" italics="true" pageId="2" pageNumber="21">Nonalatus epigaeus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[683,733,1138,1159]" pageId="2" pageNumber="21" rank="species">sp. n.</taxonomicNameLabel>
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, habitus.
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</paragraph>
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</caption>
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<paragraph blockId="2.[144,1104,1220,1677]" lastBlockId="3.[181,1142,1193,1677]" lastPageId="3" lastPageNumber="22" pageId="2" pageNumber="21">
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<emphasis box="[144,205,1220,1246]" italics="true" pageId="2" pageNumber="21">Head</emphasis>
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: Red-brown to darK brown, glossy; with dense, fine, regular punctation. Labrum brown, bilobed; anterior part of clypeus yellow, glossy; mouth parts more or less yellowbrown to light yellow, except the blacK mandibles; terminal labial palpomeres securiform, terminal maxillary palpomeres with sides parallel, constricted apically. Gula long, light red-brown, glossy, gular sutures apically slightly divergent, gular process of medium size. Head including eyes more or less as broad as anterior width of pronotum, vested with black and white setae, behind middle with a broadly Y-shaped pattern of white, depressed setae. Eyes small, coarsely facetted, margined, conspicuously emarginate at antennal insertion, with a brush of yellow setae; interocular space (frons) almost three eye widths.Antennae with 11 antennomeres, long, stout, reaching base of pronotum when laid back, brown, from A8 onwards becoming darker, A11 only basally narrowly dark, remainder light brown; A1 very long, slightly bent; A2 short, cylindrical; A3 twice as long as A2;A4 slightly shorter than A3, slightly dilated distally; from A5 onwards dilated distally, becoming thicker and more compact;A4–A10 progressively becoming slightly shorter; A11 elongate, evenly pointed; antennae with long, erect, dark brown setae, from A8 onwards increasingly tomentose. Lower surface of head dark brown, glossy.
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<emphasis box="[181,262,1265,1291]" italics="true" pageId="3" pageNumber="22">Thorax</emphasis>
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: Prothorax compact, pronotum wider than long (l =
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<quantity box="[853,1020,1265,1292]" metricMagnitude="-3" metricUnit="m" metricValue="1.445" metricValueMax="1.58" metricValueMin="1.31" pageId="3" pageNumber="22" unit="mm" value="1.445" valueMax="1.58" valueMin="1.31">1.31–1.58 mm</quantity>
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, w=
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<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.82" metricValueMax="2.02" metricValueMin="1.62" pageId="3" pageNumber="22" unit="mm" value="1.82" valueMax="2.02" valueMin="1.62">1.62– 2.02mm</quantity>
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, length to width ratio 1:0.81), widest in the middle, constricted conspicuously towards base; anterior transverse impression conspicuous; surface red-brown, towards sides dark brown to black; punctation conspicuous but not deep, transverse impression with slight transverse wrinkles in the middle; sides in the middle with slight, grooveliKe depression; procoxal cavities broadly open posteriorly, pro-intercoxal process very narrow, long. Pronotum with short, white, more or less depressed setae, sides with strong, erect, black setae. Lower surface dark brown, glossy, sides smooth, black; proepimeron broadly triangular.
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</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.7670385" ID-Zenodo-Dep="7670385" httpUri="https://zenodo.org/record/7670385/files/figure.png" pageId="3" pageNumber="22" startId="3.[183,222,1081,1103]" targetBox="[337,989,207,1059]" targetPageId="3">
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<paragraph blockId="3.[183,1144,1081,1151]" pageId="3" pageNumber="22">
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Figs 2–8.
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<taxonomicName authority="Gerstmeier, 2015" authorityName="Gerstmeier" authorityYear="2015" box="[272,451,1081,1103]" class="Insecta" family="Cleridae" genus="Nonalatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="22" phylum="Arthropoda" rank="species" species="epigaeus" status="sp. n.">
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<emphasis box="[272,451,1081,1103]" italics="true" pageId="3" pageNumber="22">Nonalatus epigaeus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[455,503,1081,1102]" pageId="3" pageNumber="22" rank="species">sp. n.</taxonomicNameLabel>
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(2) aedeagus ventral; (3) phallus ventral; (4) phallus lateral; (5) tegmen ventral; (6) spicular forK; (7) aedeagus tip (Scale bar 0.1 mm); (8) pygidium; (9) sixth ventrite. Scale bars = 0.5 mm.
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</paragraph>
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</caption>
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<paragraph blockId="3.[181,1142,1193,1677]" lastBlockId="4.[144,1104,204,905]" lastPageId="4" lastPageNumber="23" pageId="3" pageNumber="22">
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Elytra compact, broad, widest in the middle or shortly behind, base not margined (l =
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<quantity box="[223,389,1587,1613]" metricMagnitude="-3" metricUnit="m" metricValue="3.295" metricValueMax="3.72" metricValueMin="2.87" pageId="3" pageNumber="22" unit="mm" value="3.295" valueMax="3.72" valueMin="2.87">2.87–3.72 mm</quantity>
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, w =
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<quantity box="[446,615,1586,1613]" metricMagnitude="-3" metricUnit="m" metricValue="2.285" metricValueMax="2.52" metricValueMin="2.05" pageId="3" pageNumber="22" unit="mm" value="2.285" valueMax="2.52" valueMin="2.05">2.05–2.52 mm</quantity>
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, length to width ratio =1:1.42); apices slightly dehiscent, each broadly rounded. Elytral ground colour yellow-brown (apex) to red-brown (shoulder) or more or less entirely yellow-brown, with a variable pattern of black fasciae (
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<figureCitation box="[343,411,204,230]" captionStart="Fig" captionStartId="2.[432,465,1137,1159]" captionTargetBox="[369,878,209,1116]" captionTargetId="figure-186@2.[354,892,201,1125]" captionTargetPageId="2" captionText="Fig. 1. Nonalatus epigaeus sp. n., habitus." figureDoi="http://doi.org/10.5281/zenodo.7670383" httpUri="https://zenodo.org/record/7670383/files/figure.png" pageId="4" pageNumber="23">Fig. 1</figureCitation>
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). Elytral base, within the part of the ground colour with 10 rows of punctation, actually only row five to six conspicuous, diameter of punctation conspicuously smaller than interstices, apex with very fine punctation. In the area of the ground colour with short, white, mostly depressed setae; pilosity of the black fasciae black, partly erect, partly depressed.
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</paragraph>
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<paragraph blockId="4.[144,1104,204,905]" box="[170,860,362,389]" pageId="4" pageNumber="23">Scutellum transverse ovate, yellow-brown to dark red-brown.</paragraph>
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<paragraph blockId="4.[144,1104,204,905]" pageId="4" pageNumber="23">Mesoventrite red-brown to black-brown, with dense, coarse, shallow punctation; posterior mesoventral process long, depressed, connected with the anterior, long, slender metaventral process.</paragraph>
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<paragraph blockId="4.[144,1104,204,905]" pageId="4" pageNumber="23">Legs stout, relatively long; basal half of femora yellow-brown, then black, distal tip narrowly red-brown; tibiae straight, basally red-brown to black-brown, then yellowbrown shortly behind the middle, distal third black-brown. Tarsi brown (T1–T3) to yellow-brown (T4, T5), pretarsus long, claws with an acute basal denticle; legs predominantly with slightly erect, short, white setae, only the outer margin of the black part of front tibiae with black setae; the black part of all tibiae with isolated black setae. Tarsal pad formula 4–4–2, T1 and T2 of front and middle tarsi with straight to slightly emarginate euplantulae, T3 and T4 deeply emarginate, hind tarsi T3 and T4 deeply emarginate; tibial spur formula 1–2–2.</paragraph>
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<paragraph blockId="4.[144,1104,204,905]" pageId="4" pageNumber="23">
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<emphasis box="[144,249,783,809]" italics="true" pageId="4" pageNumber="23">Abdomen</emphasis>
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: Dark brown, glossy, posterior margin of abdominal ventrites 1–3 with a broad light-yellow margin, abdominal ventrite 4 with a narrow yellow margin; scattered with oblique posteriorly directed, short, white setae; abdominal ventrite 6 yellow-brown, pygidium brown. Male terminalia, see
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<figureCitation box="[578,649,878,904]" captionStart="Figs 2–8" captionStartId="3.[183,222,1081,1103]" captionTargetBox="[337,989,207,1059]" captionTargetId="figure-188@3.[325,998,204,1065]" captionTargetPageId="3" captionText="Figs 2–8. Nonalatus epigaeus sp. n. (2) aedeagus ventral; (3) phallus ventral; (4) phallus lateral; (5) tegmen ventral; (6) spicular forK;(7) aedeagus tip (Scale bar 0.1 mm); (8) pygidium; (9) sixth ventrite.Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7670385" httpUri="https://zenodo.org/record/7670385/files/figure.png" pageId="4" pageNumber="23">Figs 2</figureCitation>
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–9.
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</paragraph>
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<paragraph blockId="4.[144,1105,922,1032]" pageId="4" pageNumber="23">
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<materialsCitation collectingDate="2012-12-15" collectionCode="V, RGCM" collectorName="Mt. Hanang, NE & Grebennikov" country="Tanzania" elevation="2366" latitude="-4.43175" location="Mt. Hanang" longLatPrecision="1" longitude="35.41357" pageId="4" pageNumber="23" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<typeStatus box="[144,232,922,944]" pageId="4" pageNumber="23">Holotype</typeStatus>
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:
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<specimenCount box="[240,257,922,943]" pageId="4" pageNumber="23" type="male">♂</specimenCount>
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<collectingCountry box="[262,381,922,943]" name="Tanzania" pageId="4" pageNumber="23">TANZANIA</collectingCountry>
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:
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<collectorName box="[389,538,922,943]" pageId="4" pageNumber="23">
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<location LSID="urn:lsid:plazi:treatment:03F5D02DA552A020FD835FC8FCF8FA97:8E8337E0A557A020FE0659DBFE03FC5F" box="[389,499,922,943]" country="Tanzania" latitude="-4.43175" longLatPrecision="1" longitude="35.41357" name="Mt. Hanang" pageId="4" pageNumber="23">Mt. Hanang</location>
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, NE
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</collectorName>
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slope,
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<geoCoordinate box="[603,698,922,944]" degrees="4.43175" direction="south" orientation="latitude" pageId="4" pageNumber="23" precision="1" value="-4.43175">4.43175°S</geoCoordinate>
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<geoCoordinate box="[704,814,922,944]" degrees="35.41357" direction="east" orientation="longitude" pageId="4" pageNumber="23" precision="1" value="35.41357">35.41357°E</geoCoordinate>
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,
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<quantity box="[823,893,922,944]" metricMagnitude="3" metricUnit="m" metricValue="2.366" pageId="4" pageNumber="23" unit="m" value="2366.0">
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<elevation box="[823,893,922,944]" metricMagnitude="3" metricUnit="m" metricValue="2.366" pageId="4" pageNumber="23" unit="m" value="2366.0">2366 m</elevation>
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</quantity>
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,
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<date box="[902,1007,922,944]" pageId="4" pageNumber="23" value="2012-12-15">
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<collectingDate box="[902,1007,922,944]" pageId="4" pageNumber="23" value="2012-12-15">15.xii.2012</collectingDate>
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</date>
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, sift 21.
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<collectionCode box="[1084,1100,922,943]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="23" type="Museum">V</collectionCode>
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.
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<collectorName box="[144,262,946,968]" pageId="4" pageNumber="23">Grebennikov</collectorName>
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leg. (
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<collectionCode box="[313,385,946,968]" pageId="4" pageNumber="23">RGCM</collectionCode>
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||
).
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</materialsCitation>
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</paragraph>
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<paragraph blockId="4.[144,1105,922,1032]" pageId="4" pageNumber="23">
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<materialsCitation collectingDate="2012-12-15" collectionCode="RGCM, NMSA" collectorName="Mt. Hanang, NE & Grebennikov" country="Tanzania" elevation="2366" latitude="-4.43175" location="Mt. Hanang" longLatPrecision="1" longitude="35.41357" pageId="4" pageNumber="23" specimenCount="11" typeStatus="paratype">
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<typeStatus box="[144,236,978,999]" pageId="4" pageNumber="23">Paratypes</typeStatus>
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:
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<specimenCount box="[245,357,978,1000]" pageId="4" pageNumber="23" type="generic">9 specimens</specimenCount>
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(not sexed), same data as holotype, (
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<specimenCount box="[703,815,978,1000]" pageId="4" pageNumber="23" type="generic">2 specimens</specimenCount>
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in absolute alcohol) (8
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<collectionCode box="[1037,1104,978,999]" pageId="4" pageNumber="23">RGCM</collectionCode>
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& 1
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<collectionCode box="[184,251,1010,1032]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="4" pageNumber="23" type="Museum">NMSA</collectionCode>
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)
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</materialsCitation>
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</paragraph>
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<paragraph blockId="4.[144,1034,1061,1383]" box="[144,290,1061,1087]" pageId="4" pageNumber="23">
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<emphasis box="[144,282,1061,1087]" italics="true" pageId="4" pageNumber="23">Comparison</emphasis>
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:
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</paragraph>
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<paragraph blockId="4.[144,1034,1061,1383]" box="[144,759,1100,1126]" pageId="4" pageNumber="23">
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<emphasis box="[144,692,1100,1126]" italics="true" pageId="4" pageNumber="23">
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N. brevis
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<taxonomicName authority="Gerstmeier, 2015" authorityName="Gerstmeier" authorityYear="2015" box="[560,692,1100,1125]" class="Insecta" family="Cleridae" genus="Nonalatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="epigaeus" status="sp. n.">N. epigaeus</taxonomicName>
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</emphasis>
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<taxonomicNameLabel box="[699,759,1100,1126]" pageId="4" pageNumber="23" rank="species">sp. n.</taxonomicNameLabel>
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</paragraph>
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<paragraph blockId="4.[144,1034,1061,1383]" box="[144,732,1138,1165]" pageId="4" pageNumber="23">
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Size
|
||
<quantity box="[199,338,1138,1165]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" metricValueMax="4.5" metricValueMin="3.7" pageId="4" pageNumber="23" unit="mm" value="4.1" valueMax="4.5" valueMin="3.7">3.7–4.5 mm</quantity>
|
||
Size
|
||
<quantity box="[615,732,1138,1165]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" metricValueMax="6.0" metricValueMin="5.2" pageId="4" pageNumber="23" unit="mm" value="5.6" valueMax="6.0" valueMin="5.2">5.2–6 mm</quantity>
|
||
</paragraph>
|
||
<paragraph blockId="4.[144,1034,1061,1383]" box="[144,743,1177,1203]" pageId="4" pageNumber="23">Head black Head red-brown</paragraph>
|
||
<paragraph blockId="4.[144,1034,1061,1383]" pageId="4" pageNumber="23">Pronotum without a conspicuous Pronotum without a conspicuous transverse impression transverse impression</paragraph>
|
||
<paragraph blockId="4.[144,1034,1061,1383]" pageId="4" pageNumber="23">Elytral punctation irregular Elytral punctation arranged into striae, but only rows 5 and 6 conspicuous</paragraph>
|
||
<paragraph blockId="4.[144,1034,1061,1383]" box="[144,776,1357,1383]" pageId="4" pageNumber="23">Hind tibiae slightly curved Hind tibiae straight</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |