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<document ID-DOI="10.15553/c2021v762a3" ID-ISSN="2235-3658" approvalRequired="43" approvalRequired_for_textStreams="43" checkinTime="1665435377359" checkinUser="felipe" docAuthor="Schatz, George E., Lowry Ii, Porter P., Rakouth, Hasina N. &amp; Randrianaivo, Richard" docDate="2021" docId="03F5091DFFBC5F11FF94FABC9B9C01F9" docLanguage="en" docName="Candollea.76.2.201-236.pdf" docOrigin="Candollea 76 (2)" docSource="http://dx.doi.org/10.15553/c2021v762a3" docStyle="DocumentStyle:1E5CC43F38D0DBC531513809D1267F7B.2:Candollea.2015-.journal_article" docStyleId="1E5CC43F38D0DBC531513809D1267F7B" docStyleName="Candollea.2015-.journal_article" docStyleVersion="2" docTitle="Diospyros lowryi G. E. Schatz. A. Branch 2021, sp. nov." docType="treatment" docVersion="1" lastPageNumber="222" masterDocId="FFCC7165FFAF5F04FFE5FFAB99750247" masterDocTitle="Taxonomic studies of Diospyros (Ebenaceae) from the Malagasy region. VI. New species of large trees from Madagascar" masterLastPageNumber="236" masterPageNumber="201" pageNumber="220" updateTime="1665435621213" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title>Taxonomic studies of Diospyros (Ebenaceae) from the Malagasy region. VI. New species of large trees from Madagascar</mods:title>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Schatz, George E.</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Lowry Ii, Porter P.</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Rakouth, Hasina N.</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Randrianaivo, Richard</mods:namePart>
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<mods:title>Candollea</mods:title>
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<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-07-07</mods:number>
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<mods:detail type="volume">
<mods:number>76</mods:number>
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<mods:identifier type="ISSN">2235-3658</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:03F5091DFFBC5F11FF94FABC9B9C01F9" httpUri="http://treatment.plazi.org/id/03F5091DFFBC5F11FF94FABC9B9C01F9" lastPageId="21" lastPageNumber="222" pageId="19" pageNumber="220">
<subSubSection pageId="19" pageNumber="220" type="nomenclature">
<paragraph blockId="19.[113,628,1302,1327]" box="[113,628,1302,1327]" pageId="19" pageNumber="220">
<heading box="[113,628,1302,1327]" centered="true" fontSize="10" level="2" pageId="19" pageNumber="220" reason="2">
<taxonomicName authority="G.E. Schatz" authorityName="G. E. Schatz. A. Branch" authorityYear="2021" box="[113,432,1302,1327]" class="Magnoliopsida" family="Ebenaceae" genus="Diospyros" kingdom="Plantae" order="Ericales" pageId="19" pageNumber="220" phylum="Tracheophyta" rank="species" species="lowryi" status="sp. nov.">
<emphasis bold="true" box="[113,293,1302,1327]" italics="true" pageId="19" pageNumber="220">Diospyros lowryi</emphasis>
G.E. Schatz
</taxonomicName>
,
<emphasis bold="true" box="[442,526,1303,1327]" pageId="19" pageNumber="220">
<taxonomicNameLabel box="[442,526,1303,1327]" pageId="19" pageNumber="220" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation box="[540,616,1303,1327]" captionStart="Fig" captionStartId="22.[136,165,1915,1935]" captionText="Fig. 11. Diospyros lowryi G.E. Schatz. A. Branch in fruit; B. Fruit; C. Detail of leaf (adaxial surface). [AC: Schatz &amp; Lowry 2617, P] [Drawing:Alain Jouy]" pageId="19" pageNumber="220">Fig. 11</figureCitation>
).
</heading>
</paragraph>
<paragraph blockId="19.[151,764,1345,1466]" pageId="19" pageNumber="220">
<materialsCitation collectingDate="1989-02-21" collectionCode="K, P, TAN, WAG" collectorName="Schatz &amp; Lowry" country="Madagascar" location="Masoala Peninsula" pageId="19" pageNumber="220" specimenCode="P03975074, TAN001945" specimenCount="1" stateProvince="Toamasina" typeStatus="holotype">
<emphasis bold="true" pageId="19" pageNumber="220">
<typeStatus box="[151,280,1346,1370]" pageId="19" pageNumber="220">Holotypus</typeStatus>
:
<collectingCountry box="[293,458,1345,1369]" name="Madagascar" pageId="19" pageNumber="220">MADAGASCAR</collectingCountry>
. Reg. Analanjirofo [Prov.
<collectingRegion box="[151,280,1377,1401]" country="Madagascar" name="Toamasina" pageId="19" pageNumber="220">Toamasina</collectingRegion>
]:
</emphasis>
<location LSID="urn:lsid:plazi:treatment:03F5091DFFBC5F11FF94FABC9B9C01F9:8E83EED0FFBC5F17FECEFAC99883073D" box="[299,502,1378,1402]" country="Madagascar" name="Masoala Peninsula" pageId="19" pageNumber="220" stateProvince="Toamasina">Masoala Peninsula</location>
,
<date box="[513,621,1378,1402]" pageId="19" pageNumber="220" value="1989-02-21">
<collectingDate box="[513,621,1378,1402]" pageId="19" pageNumber="220" value="1989-02-21">21.II.1989</collectingDate>
</date>
, fr.,
<emphasis italics="true" pageId="19" pageNumber="220">
<collectorName box="[667,734,1378,1402]" pageId="19" pageNumber="220">Schatz</collectorName>
&amp;
<collectorName box="[151,219,1410,1433]" pageId="19" pageNumber="220">Lowry</collectorName>
2617
</emphasis>
(MO-3706473!; iso-:
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,
<collectionCode box="[534,550,1410,1433]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="19" pageNumber="220" type="Herbarium">P</collectionCode>
[
<specimenCode box="[563,688,1409,1433]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="19" pageNumber="220" type="Herbarium">P03975074</specimenCode>
]!,
<collectionCode box="[707,764,1410,1434]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" lsid="urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="19" pageNumber="220" type="Herbarium">TAN</collectionCode>
[
<specimenCode box="[157,301,1441,1465]" collectionCode="TAN" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" lsid="urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="19" pageNumber="220" type="Herbarium">TAN001945</specimenCode>
]!,
<collectionCode box="[321,386,1442,1466]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15578" lsid="urn:lsid:biocol.org:col:15578" name="Wageningen University" pageId="19" pageNumber="220" type="Herbarium">WAG</collectionCode>
[
<collectionCode box="[399,466,1441,1465]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15578" lsid="urn:lsid:biocol.org:col:15578" name="Wageningen University" pageId="19" pageNumber="220" type="Herbarium">WAG</collectionCode>
.1180486] image!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="21" lastPageNumber="222" pageId="19" pageNumber="220" type="description">
<paragraph blockId="19.[151,764,1484,1700]" pageId="19" pageNumber="220">
<emphasis italics="true" pageId="19" pageNumber="220">
<taxonomicName authority="G. E. Schatz" authorityName="G. E. Schatz. A. Branch" authorityYear="2021" box="[151,470,1484,1508]" class="Magnoliopsida" family="Ebenaceae" genus="Diospyros" kingdom="Plantae" order="Ericales" pageId="19" pageNumber="220" phylum="Tracheophyta" rank="species" species="lowryi">Diospyros lowryi G.E. Schatz</taxonomicName>
is distinguished from other members of the genus in
<collectingCountry box="[404,530,1516,1540]" name="Madagascar" pageId="19" pageNumber="220">Madagascar</collectingCountry>
by its ovate to elliptic, glabrous lamina with an abruptly short acuminate apex and its paired axillary infrutescences, each with 3 6 fruits, the ovoid to elliptic fruit densely covered with short, appressed, beige trichomes
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long, rendering the fruit beige in vivo, light brown in sicco.
</emphasis>
</paragraph>
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<emphasis box="[151,196,1741,1764]" italics="true" pageId="19" pageNumber="220">Tree</emphasis>
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tall,
<quantity box="[346,407,1740,1764]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" pageId="19" pageNumber="220" unit="cm" value="25.0">25 cm</quantity>
DBH.
<emphasis box="[486,605,1741,1764]" italics="true" pageId="19" pageNumber="220">Young stems</emphasis>
terete, sparsely covered with minute, erect, white trichomes
<quantity box="[616,701,1772,1796]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="220" unit="mm" value="0.1">0.1 mm</quantity>
long.
<emphasis box="[113,183,1805,1828]" italics="true" pageId="19" pageNumber="220">Leaves</emphasis>
distichous, lamina 2.76 ×
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, ovate to elliptic, chartaceous, glabrous on both surfaces, base rounded to cuneate to acuminate, margin flat to slightly revolute, apex abruptly short acuminate, acumen
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long, the very tip rounded, midvein very slightly impressed above, raised below, venation brochidodromous, with 57 secondary veins per side, indistinct above, slightly raised below, tertiary venation reticulate; petiole
<quantity box="[1051,1142,342,367]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="19" pageNumber="220" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">35 mm</quantity>
long,
<quantity box="[1211,1273,343,367]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="220" unit="mm" value="1.0">1 mm</quantity>
in diam., rather sparsely covered with short, appressed trichomes c.
<quantity box="[1367,1451,375,399]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="19" pageNumber="220" unit="mm" value="0.3">0.3 mm</quantity>
long, glabrescent.
<emphasis box="[1002,1140,407,431]" italics="true" pageId="19" pageNumber="220">Male flowers</emphasis>
in axillary cymose inflorescences of 24 flowers,
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long, each flower subtended by a bracteole, the axes and bracteole orange-brown, pedicel c.
<quantity box="[827,888,503,527]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="220" unit="mm" value="1.0">1 mm</quantity>
long, &lt;
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in diam., densely covered with short, appressed, light beige trichomes c.
<quantity box="[1167,1248,535,559]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="19" pageNumber="220" unit="mm" value="0.3">0.3 mm</quantity>
long; calyx broadly urceolate, 1 ×
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, entire, rather densely covered with short, appressed, light beige trichomes c.
<quantity box="[1178,1311,598,623]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="19" pageNumber="220" unit="mm" value="0.25" valueMax="0.3" valueMin="0.2">0.20.3 mm</quantity>
long; corolla obconical in bud; stamens c. 18, immature.
<emphasis box="[1259,1409,631,655]" italics="true" pageId="19" pageNumber="220">Female flowers</emphasis>
not seen.
<emphasis box="[861,925,664,687]" italics="true" pageId="19" pageNumber="220">Fruits</emphasis>
axillary, borne in paired cymose infructescences, each with 3 6 fruits, the axes densely covered with short, appressed, light brown trichomes c.
<quantity box="[1180,1311,726,751]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="19" pageNumber="220" unit="mm" value="0.25" valueMax="0.3" valueMin="0.2">0.20.3 mm</quantity>
long; pedicel in fruit c.
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long, c.
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in diam., densely covered with appressed, light brown trichomes c.
<quantity box="[1227,1309,791,815]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="220" unit="mm" value="0.2">0.2 mm</quantity>
long; fruiting calyx broadly cupuliform, 2 ×
<quantity box="[1112,1201,822,847]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="19" pageNumber="220" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">56 mm</quantity>
, entire, sparsely covered with appressed, light brown trichomes c.
<quantity box="[1231,1313,855,879]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="220" unit="mm" value="0.2">0.2 mm</quantity>
long, margin entire, flat, dark chocolate brown in sicco, pedicel scar
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in diam.; fruits ovoid to ellipsoid, 1215 ×
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, slightly asymmetrical, beige in vivo, light brown in sicco, densely covered with short, appressed, beige trichomes c.
<quantity box="[1320,1451,982,1007]" metricMagnitude="-4" metricUnit="m" metricValue="3" metricValueMax="4.0" metricValueMin="2.0" pageId="19" pageNumber="220" unit="mm" value="0.3" valueMax="0.4" valueMin="0.2">0.20.4 mm</quantity>
long, apex acute, shortly apiculate, apiculum
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long.
<emphasis box="[1400,1451,1015,1039]" italics="true" pageId="19" pageNumber="220">Seeds</emphasis>
13, ovoid to ellipsoid, 1012 ×
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, glabrous, black, shiny.
</paragraph>
<paragraph blockId="19.[801,1451,1111,1295]" pageId="19" pageNumber="220">
<emphasis box="[839,959,1111,1135]" italics="true" pageId="19" pageNumber="220">Etymology.</emphasis>
The specific epithet honors my colleague Porter P. Lowry II, who has led the Missouri Botanical Gardens program in
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since 1986. Although he is a specialist in
<taxonomicName box="[939,1046,1207,1231]" class="Magnoliopsida" family="Araliaceae" kingdom="Plantae" order="Apiales" pageId="19" pageNumber="220" phylum="Tracheophyta" rank="family">
<emphasis box="[939,1046,1207,1231]" italics="true" pageId="19" pageNumber="220">Araliaceae</emphasis>
</taxonomicName>
, in 2009 he agreed to work together with me on a revision of Malagasy
<taxonomicName box="[1177,1281,1239,1263]" class="Magnoliopsida" family="Ebenaceae" kingdom="Plantae" order="Ericales" pageId="19" pageNumber="220" phylum="Tracheophyta" rank="family">
<emphasis box="[1177,1281,1239,1263]" italics="true" pageId="19" pageNumber="220">Ebenaceae</emphasis>
</taxonomicName>
, a collaboration that has surpassed our wildest expectations.
</paragraph>
<paragraph blockId="19.[801,1452,1335,1391]" pageId="19" pageNumber="220">
<emphasis box="[839,1016,1335,1359]" italics="true" pageId="19" pageNumber="220">Vernacular names.</emphasis>
“Maintipototra” (
<emphasis box="[1214,1437,1335,1359]" italics="true" pageId="19" pageNumber="220">Service Forestier 12978</emphasis>
); “Maintimpototra à petites feuilles” (
<emphasis box="[1174,1400,1367,1391]" italics="true" pageId="19" pageNumber="220">Service Forestier 15365</emphasis>
).
</paragraph>
<paragraph blockId="19.[801,1451,1431,1615]" pageId="19" pageNumber="220">
<emphasis box="[839,1313,1431,1455]" italics="true" pageId="19" pageNumber="220">
Distribution and ecology.
<taxonomicName authorityName="G. E. Schatz. A. Branch" authorityYear="2021" box="[1135,1313,1431,1455]" class="Magnoliopsida" family="Ebenaceae" genus="Diospyros" kingdom="Plantae" order="Ericales" pageId="19" pageNumber="220" phylum="Tracheophyta" rank="species" species="lowryi">Diospyros lowryi</taxonomicName>
</emphasis>
is restricted to the area around the Baie dAntongil, where it has been collected at just two sites, near Ambanizana, situated just outside Masoala National Park, and at the Farankaraina Forestry Station, to the east of Maroantsetra (
<bibRefCitation author="MADAGASCAR CATALOGUE" pageId="19" pageNumber="220" refId="ref25178" refString="MADAGASCAR CATALOGUE (2021). Catalogue of the Plants of Madagascar. Missouri Botanical Garden, St. Louis &amp; Antananarivo. [http: // www. tropicos. org / Project / Madagascar]" type="url" year="2021">
<collectingCountry box="[1299,1451,1559,1583]" name="Madagascar" pageId="19" pageNumber="220">MADAGASCAR</collectingCountry>
CATALOGUE, 2021
</bibRefCitation>
). It occurs in low-elevation humid forest.
</paragraph>
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<emphasis box="[839,955,1655,1679]" italics="true" pageId="19" pageNumber="220">Phenology.</emphasis>
Flowering material has been collected in
</paragraph>
<paragraph blockId="19.[801,1450,1655,1711]" box="[801,1289,1687,1711]" pageId="19" pageNumber="220">November, and fruits in February and March.</paragraph>
<paragraph blockId="19.[801,1452,1751,1935]" pageId="19" pageNumber="220">
<emphasis box="[839,1259,1751,1775]" italics="true" pageId="19" pageNumber="220">
Conservation status.
<taxonomicName authorityName="G. E. Schatz. A. Branch" authorityYear="2021" box="[1082,1259,1751,1775]" class="Magnoliopsida" family="Ebenaceae" genus="Diospyros" kingdom="Plantae" order="Ericales" pageId="19" pageNumber="220" phylum="Tracheophyta" rank="species" species="lowryi">Diospyros lowryi</taxonomicName>
</emphasis>
has an extremely restricted geographic range in the form of an Area of Occupancy of
<quantity box="[902,957,1815,1839]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="19" pageNumber="220" unit="km" value="8.0">8 km</quantity>
². It is known from just outside the protected area of Masoala, where it is threatened by exploitation of trees for firewood and house construction material, which will result in continuing decline. With respect to the most serious
</paragraph>
<paragraph blockId="20.[1315,1366,540,560]" lastBlockId="20.[136,1418,1888,1960]" pageId="20" pageNumber="221">
mm [
<emphasis bold="true" box="[142,190,1939,1960]" pageId="20" pageNumber="221">AD:</emphasis>
<emphasis box="[195,423,1938,1960]" italics="true" pageId="20" pageNumber="221">Réserves Naturelles 9731</emphasis>
, P;
<emphasis bold="true" box="[453,472,1940,1960]" pageId="20" pageNumber="221">E:</emphasis>
<emphasis box="[476,627,1938,1960]" italics="true" pageId="20" pageNumber="221">Lowry et al. 7482</emphasis>
, from photo] [Drawing: Alain Jouy]
</paragraph>
<caption pageId="20" pageNumber="221" startId="20.[136,166,1889,1909]" targetBox="[239,1342,411,1716]" targetPageId="20">
<paragraph blockId="20.[136,1418,1888,1960]" pageId="20" pageNumber="221">
<emphasis bold="true" box="[136,215,1888,1909]" pageId="20" pageNumber="221">Fig. 10. </emphasis>
<taxonomicName authority="G. E. Schatz &amp; Lowry. A. Branch" box="[220,698,1889,1909]" class="Magnoliopsida" family="Ebenaceae" genus="Diospyros" kingdom="Plantae" order="Ericales" pageId="20" pageNumber="221" phylum="Tracheophyta" rank="species" species="grandiflora">
<emphasis box="[220,415,1889,1909]" italics="true" pageId="20" pageNumber="221">Diospyros grandiflora</emphasis>
G.E. Schatz &amp; Lowry.
<emphasis bold="true" box="[608,628,1889,1909]" pageId="20" pageNumber="221">A.</emphasis>
Branch
</taxonomicName>
in fruit;
<emphasis bold="true" box="[774,793,1889,1909]" pageId="20" pageNumber="221">B.</emphasis>
Branch with female flowers;
<emphasis bold="true" box="[1060,1080,1889,1909]" pageId="20" pageNumber="221">C.</emphasis>
Schematic section of female flower;
<emphasis bold="true" box="[136,156,1915,1935]" pageId="20" pageNumber="221">D.</emphasis>
Corolla of female flower;
<emphasis bold="true" box="[391,409,1915,1935]" pageId="20" pageNumber="221">E.</emphasis>
Fruit.
</paragraph>
</caption>
<paragraph blockId="21.[113,764,279,399]" pageId="21" pageNumber="222">
plausible threat of exploitation of trees for firewood and house construction material,
<taxonomicName authorityName="G. E. Schatz. A. Branch" authorityYear="2021" box="[352,450,311,335]" class="Magnoliopsida" family="Ebenaceae" genus="Diospyros" kingdom="Plantae" order="Ericales" pageId="21" pageNumber="222" phylum="Tracheophyta" rank="species" species="lowryi">
<emphasis box="[352,450,311,335]" italics="true" pageId="21" pageNumber="222">D. lowryi</emphasis>
</taxonomicName>
exists at two locations, and it was recently assessed for its risk of extinction as “Endangered” [EN B2ab(iii,v)] (
<bibRefCitation author="IUCN" box="[305,444,375,399]" pageId="21" pageNumber="222" refId="ref25081" refString="IUCN (2021). The IUCN Red List of Threatened Species. [https: // www. iucnredlist. org]" type="url" year="2021">IUCN, 2021</bibRefCitation>
).
</paragraph>
<paragraph blockId="21.[113,764,439,847]" pageId="21" pageNumber="222">
<emphasis box="[151,404,439,463]" italics="true" pageId="21" pageNumber="222">
Notes.
<taxonomicName authorityName="G. E. Schatz. A. Branch" authorityYear="2021" box="[237,404,439,463]" class="Magnoliopsida" family="Ebenaceae" genus="Diospyros" kingdom="Plantae" order="Ericales" pageId="21" pageNumber="222" phylum="Tracheophyta" rank="species" species="lowryi">Diospyros lowryi</taxonomicName>
</emphasis>
belongs to a loosely defined group of species in
<collectingCountry box="[254,384,471,495]" name="Madagascar" pageId="21" pageNumber="222">Madagascar</collectingCountry>
characterized by 3-merous flowers and a small calyx that is generally cupuliform and entire or shallowly lobed, which includes several species previously placed by
<bibRefCitation author="PERRIER DE LA BATHIE, H." box="[223,553,561,591]" pageId="21" pageNumber="222" pagination="165" refId="ref25291" refString="PERRIER DE LA BATHIE, H. (1952). Ebenacees. In: HUMBERT, H. (ed.), Fl. Madagascar Comores 165." type="book chapter" year="1952">PERRIER DE LA BÂTHIE (1952)</bibRefCitation>
in the genus
<taxonomicName box="[702,764,567,591]" class="Magnoliopsida" family="Ebenaceae" genus="Maba" higherTaxonomySource="GBIF" kingdom="Plantae" order="Ericales" pageId="21" pageNumber="222" phylum="Tracheophyta" rank="genus">
<emphasis box="[702,764,567,591]" italics="true" pageId="21" pageNumber="222">Maba</emphasis>
</taxonomicName>
(see above under
<taxonomicName box="[293,469,599,623]" class="Magnoliopsida" family="Ebenaceae" genus="Diospyros" kingdom="Plantae" order="Ericales" pageId="21" pageNumber="222" phylum="Tracheophyta" rank="species" species="andohahelensis">
<emphasis box="[293,469,599,623]" italics="true" pageId="21" pageNumber="222">D. andohahelensis</emphasis>
</taxonomicName>
), as well as perhaps 25 additional new species that remain to be described. Within this group,
<taxonomicName authorityName="G. E. Schatz. A. Branch" authorityYear="2021" box="[188,289,663,687]" class="Magnoliopsida" family="Ebenaceae" genus="Diospyros" kingdom="Plantae" order="Ericales" pageId="21" pageNumber="222" phylum="Tracheophyta" rank="species" species="lowryi">
<emphasis box="[188,289,663,687]" italics="true" pageId="21" pageNumber="222">D. lowryi</emphasis>
</taxonomicName>
can be distinguished by its ovate to elliptic, glabrous lamina with an abruptly acuminate apex and its axillary, paired infructescences, each with 36 fruits, the ovoid to elliptic fruits densely covered with appressed, beige trichomes
<quantity box="[113,243,790,815]" metricMagnitude="-4" metricUnit="m" metricValue="3" metricValueMax="4.0" metricValueMin="2.0" pageId="21" pageNumber="222" unit="mm" value="0.3" valueMax="0.4" valueMin="0.2">0.20.4 mm</quantity>
long, rendering the fruit beige in vivo, light brown in sicco.
</paragraph>
<paragraph blockId="21.[113,764,885,960]" pageId="21" pageNumber="222">
<emphasis box="[151,399,885,904]" italics="true" pageId="21" pageNumber="222">Additional specimen examined.</emphasis>
<materialsCitation collectingDate="1955-03-04" collectingDateMax="1955-11-25" collectingDateMin="1955-03-04" collectionCode="MO, P, TEF, G" country="Madagascar" location="Service Forestier" municipality="Farankaraina" pageId="21" pageNumber="222" specimenCount="1" specimenCount-male="1" stateProvince="Toamasina">
<emphasis bold="true" pageId="21" pageNumber="222">
<collectingCountry box="[418,542,885,904]" name="Madagascar" pageId="21" pageNumber="222">MADAGASCAR</collectingCountry>
.Reg. Analanjirofo [Prov.
<collectingRegion box="[113,219,912,931]" country="Madagascar" name="Toamasina" pageId="21" pageNumber="222">Toamasina</collectingRegion>
]:
</emphasis>
<collectingMunicipality box="[235,352,912,931]" pageId="21" pageNumber="222">Farankaraina</collectingMunicipality>
,
<date box="[362,449,912,931]" pageId="21" pageNumber="222" value="1955-03-04">
<collectingDate box="[362,449,912,931]" pageId="21" pageNumber="222" value="1955-03-04">4.III.1955</collectingDate>
</date>
, fr.,
<emphasis box="[488,685,912,931]" italics="true" pageId="21" pageNumber="222">
<location LSID="urn:lsid:plazi:treatment:03F5091DFFBC5F11FF94FABC9B9C01F9:8E83EED0FFBA5F11FE0DFC3B9B0101E5" box="[488,628,912,931]" country="Madagascar" municipality="Farankaraina" name="Service Forestier" pageId="21" pageNumber="222" stateProvince="Toamasina">Service Forestier</location>
12978
</emphasis>
(
<collectionCode box="[698,738,912,931]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="21" pageNumber="222" type="Herbarium">MO</collectionCode>
,
<collectionCode box="[749,760,912,931]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="21" pageNumber="222" type="Herbarium">P</collectionCode>
,
<collectionCode box="[113,157,939,958]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12774" lsid="urn:lsid:biocol.org:col:12774" name="Centre National de la Recherche Appliquée au Developement Rural" pageId="21" pageNumber="222" type="Herbarium">TEF</collectionCode>
);
<emphasis box="[173,243,938,957]" italics="true" pageId="21" pageNumber="222">ibid. loco</emphasis>
,
<date box="[252,342,938,957]" pageId="21" pageNumber="222" value="1955-11-25">
<collectingDate box="[252,342,938,957]" pageId="21" pageNumber="222" value="1955-11-25">25.XI.1955</collectingDate>
</date>
,
<specimenCount box="[351,367,939,960]" pageId="21" pageNumber="222" type="male"></specimenCount>
fl.,
<emphasis box="[398,584,938,958]" italics="true" pageId="21" pageNumber="222">Service Forestier 15365</emphasis>
(
<collectionCode box="[596,614,939,958]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="21" pageNumber="222" type="Herbarium">G</collectionCode>
,
<collectionCode box="[623,662,939,958]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="21" pageNumber="222" type="Herbarium">MO</collectionCode>
,
<collectionCode box="[670,681,939,958]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="21" pageNumber="222" type="Herbarium">P</collectionCode>
,
<collectionCode box="[689,734,939,958]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12774" lsid="urn:lsid:biocol.org:col:12774" name="Centre National de la Recherche Appliquée au Developement Rural" pageId="21" pageNumber="222" type="Herbarium">TEF</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>