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<document id="919CEE3A4962E57F924A1AB903E72AD7" ID="10.11646/zootaxa.4236.1.5SLASH10255" ID-CLB-Dataset="36476" ID-DOI="10.11646/zootaxa.4236.1.5" ID-GBIF-Dataset="9b07325e-60db-405c-97a2-831f68682f6e" ID-ISSN="1175-5326" ID-Zenodo-Dep="322026" ID-ZooBank="04D3145C-1C8A-4AE7-968F-7F307E1E732A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1487754882402" checkinUser="plazi" docAuthor="Mašán, Peter" docDate="2017" docId="03DD87E12E0EC467A69EFB83FB9A673A" docLanguage="en" docName="zootaxa.4236.1.5SLASH10255.pdf" docOrigin="Zootaxa 4236 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pachylaelaps (Longipachylaelaps) marcovallei Mašán, 2017, sp. nov." docType="treatment" docVersion="9" lastPageNumber="106" masterDocId="FFE4FF992E06C46CA609FFD4FFAA6059" masterDocTitle="Four new species of the subgenus Pachylaelaps (Longipachylaelaps) from Italy, with a new identification key to the known species (Acari: Pachylaelapidae)" masterLastPageNumber="117" masterPageNumber="95" pageNumber="103" updateTime="1698437172724" updateUser="ExternalLinkService">
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<mods:title id="7B9425CCBEF0D997DD4D1B3132F0BDC8">Four new species of the subgenus Pachylaelaps (Longipachylaelaps) from Italy, with a new identification key to the known species (Acari: Pachylaelapidae)</mods:title>
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<mods:namePart id="0D70938112B1BDC03A9C2B4B1B2741C0">Mašán, Peter</mods:namePart>
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<mods:date id="0655D66803D887D5E7B5B3C5789574E1">2017</mods:date>
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<treatment id="03DD87E12E0EC467A69EFB83FB9A673A" ID-DOI="http://doi.org/10.5281/zenodo.5694786" ID-GBIF-Taxon="127696754" ID-Zenodo-Dep="5694786" LSID="urn:lsid:plazi:treatment:03DD87E12E0EC467A69EFB83FB9A673A" httpUri="http://treatment.plazi.org/id/03DD87E12E0EC467A69EFB83FB9A673A" lastPageId="11" lastPageNumber="106" pageId="8" pageNumber="103">
<subSubSection id="C36E657C2E0EC464A69EFB83FE7A64CA" pageId="8" pageNumber="103" type="nomenclature">
<paragraph id="8BCB36F72E0EC464A69EFB83FC856428" blockId="8.[151,815,1111,1171]" box="[151,815,1111,1137]" pageId="8" pageNumber="103">
<heading id="D083819B2E0EC464A69EFB83FC856428" bold="true" box="[151,815,1111,1137]" fontSize="11" level="1" pageId="8" pageNumber="103" reason="1">
<emphasis id="B900EAE52E0EC464A69EFB83FC856428" bold="true" box="[151,815,1111,1137]" pageId="8" pageNumber="103">
<taxonomicName id="4C744D742E0EC464A69EFB83FD636428" ID-CoL="8Y7KM" box="[151,713,1111,1137]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="103" phylum="Arthropoda" rank="species" species="marcovallei" status="sp. nov." subGenus="Longipachylaelaps">
<emphasis id="B900EAE52E0EC464A69EFB83FE9D6428" bold="true" box="[151,311,1111,1137]" italics="true" pageId="8" pageNumber="103">Pachylaelaps</emphasis>
(
<emphasis id="B900EAE52E0EC464A74EFB83FD816428" bold="true" box="[327,555,1111,1137]" italics="true" pageId="8" pageNumber="103">Longipachylaelaps</emphasis>
)
<emphasis id="B900EAE52E0EC464A435FB83FD636428" bold="true" box="[572,713,1111,1137]" italics="true" pageId="8" pageNumber="103">marcovallei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A233579E2E0EC464A4D8FB83FC856428" box="[721,815,1111,1137]" pageId="8" pageNumber="103" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCB36F72E0EC464A69EFBAEFE7A64CA" blockId="8.[151,815,1111,1171]" box="[151,464,1146,1171]" pageId="8" pageNumber="103">
(
<figureCitation id="134F2A722E0EC464A696FBAEFF5D64CA" box="[159,247,1146,1171]" captionStart="FIGURES 27 28" captionStartId="9.[151,263,1844,1867]" captionTargetBox="[154,1434,737,1819]" captionTargetId="figure@9.[151,1436,733,1823]" captionTargetPageId="9" captionText="FIGURES 27 28. Pachylaelaps (Longipachylaelaps) marcovallei sp. nov., female. 14. Dorsal idiosoma; 15. Ventral idiosoma. Not scaled." httpUri="https://zenodo.org/record/322033/files/figure.png" pageId="8" pageNumber="103">Figs 27</figureCitation>
¯35, 38, 49, 50, 56)
</paragraph>
</subSubSection>
<subSubSection id="C36E657C2E0EC464A69EFB16FB4D6533" pageId="8" pageNumber="103" type="description">
<paragraph id="8BCB36F72E0EC464A69EFB16FB4D6533" blockId="8.[151,1437,1218,2035]" pageId="8" pageNumber="103">
<materialsCitation id="3B1C3CAA2E0EC464A69EFB16FDEE657B" ID-GBIF-Occurrence="1434656170" collectingDate="2015-05-13" collectorName="Oltre Il Colle &amp; P. Masan" country="Italy" county="Oltre Il Colle Village" elevation="1100" location="Specimens" municipality="Prealps" pageId="8" pageNumber="103" specimenCount="1" typeStatus="holotype">
<emphasis id="B900EAE52E0EC464A69EFB16FE326482" bold="true" box="[151,408,1218,1243]" pageId="8" pageNumber="103">
<location id="8EAB602C2E0EC464A69EFB16FEBF6482" LSID="urn:lsid:plazi:treatment:03DD87E12E0EC467A69EFB83FB9A673A:8EAB602C2E0EC464A69EFB16FEBF6482" box="[151,277,1218,1243]" country="Italy" county="Oltre Il Colle Village" municipality="Prealps" name="Specimens" pageId="8" pageNumber="103">Specimens</location>
examined.
</emphasis>
<typeStatus id="54CF88552E0EC464A7A9FB16FDA26482" box="[416,520,1218,1243]" pageId="8" pageNumber="103" type="holotype">Holotype</typeStatus>
female: North
<collectingCountry id="F36376672E0EC464A4BFFB16FD406482" box="[694,746,1218,1243]" name="Italy" pageId="8" pageNumber="103">Italy</collectingCountry>
,
<location id="8EAB602C2E0EC464A4FEFB17FC676482" LSID="urn:lsid:plazi:treatment:03DD87E12E0EC467A69EFB83FB9A673A:8EAB602C2E0EC464A4FEFB17FC676482" box="[759,973,1218,1243]" country="Italy" county="Oltre Il Colle Village" municipality="Prealps" name="Bergamo Province" pageId="8" pageNumber="103">Bergamo Province</location>
,
<location id="8EAB602C2E0EC464A5D0FB17FB046482" LSID="urn:lsid:plazi:treatment:03DD87E12E0EC467A69EFB83FB9A673A:8EAB602C2E0EC464A5D0FB17FB046482" box="[985,1198,1218,1243]" country="Italy" county="Oltre Il Colle Village" municipality="Prealps" name="Bergamasque Alps" pageId="8" pageNumber="103">Bergamasque Alps</location>
and
<collectingMunicipality id="6BAFAC8D2E0EC464A2EEFB16FA966482" box="[1255,1340,1218,1243]" pageId="8" pageNumber="103">Prealps</collectingMunicipality>
,
<collectingCounty id="62AA4E7B2E0EC464A340FB16FE8464A6" pageId="8" pageNumber="103">
<collectorName id="268153212E0EC464A340FB16FF7E64A6" pageId="8" pageNumber="103">Oltre Il Colle</collectorName>
Village
</collectingCounty>
, beech forest (
<taxonomicName id="4C744D742E0EC464A7C6FB33FDD564A7" box="[463,639,1254,1278]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="103" phylum="Tracheophyta" rank="species" species="sylvatica">
<emphasis id="B900EAE52E0EC464A7C6FB33FDD564A7" box="[463,639,1254,1278]" italics="true" pageId="8" pageNumber="103">Fagus sylvatica</emphasis>
</taxonomicName>
) with admixed spruce (
<taxonomicName id="4C744D742E0EC464A58FFB33FBAD64A7" box="[902,1031,1254,1278]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="8" pageNumber="103" phylum="Tracheophyta" rank="species" species="abies">
<emphasis id="B900EAE52E0EC464A58FFB33FBAD64A7" box="[902,1031,1254,1278]" italics="true" pageId="8" pageNumber="103">Picea abies</emphasis>
</taxonomicName>
), leaf litter and soil detritus, altitude
<quantity id="4C8C9B122E0EC464A69EFADEFF5F657B" box="[151,245,1290,1314]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="103" unit="m" value="1100.0">
<elevation id="0059D1C42E0EC464A69EFADEFF5F657B" box="[151,245,1290,1314]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="103" unit="m" value="1100.0">1,100 m</elevation>
</quantity>
,
<date id="FFCA10372E0EC464A709FADEFE33657A" box="[256,409,1290,1315]" pageId="8" pageNumber="103" value="2015-05-13">
<collectingDate id="EF8EE9DF2E0EC464A709FADEFE33657A" box="[256,409,1290,1315]" pageId="8" pageNumber="103" value="2015-05-13">13 May 2015</collectingDate>
</date>
, coll.
<collectorName id="268153212E0EC464A7D5FADFFDEE657B" box="[476,580,1290,1314]" pageId="8" pageNumber="103">P. Mašán</collectorName>
</materialsCitation>
.
<materialsCitation id="3B1C3CAA2E0EC464A446FADFFA95657A" ID-GBIF-Occurrence="1434656316" box="[591,1343,1290,1315]" collectingDate="2015-05-13" collectorName="Oltre Il Colle &amp; P. Masan" country="Italy" county="Oltre Il Colle Village" elevation="1100" location="Specimens" municipality="Prealps" pageId="8" pageNumber="103" specimenCount="5" specimenCount-female="1" specimenCount-male="4" typeStatus="paratype">
<typeStatus id="54CF88552E0EC464A446FADFFD6A657A" box="[591,704,1291,1315]" pageId="8" pageNumber="103" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D72FD7E2E0EC464A4C3FADEFC81657A" box="[714,811,1290,1315]" pageId="8" pageNumber="103" type="female">1 female</specimenCount>
and
<specimenCount id="9D72FD7E2E0EC464A56BFADEFC11657A" box="[866,955,1290,1315]" pageId="8" pageNumber="103" type="male">4 males</specimenCount>
, with the same data as in
<typeStatus id="54CF88552E0EC464A2D0FADEFA95657A" box="[1241,1343,1290,1315]" pageId="8" pageNumber="103" type="holotype">holotype</typeStatus>
</materialsCitation>
;
<materialsCitation id="3B1C3CAA2E0EC464A343FADEFB4E6533" ID-GBIF-Occurrence="1434656284" collectingDate="2015-05-13" collectorName="P. Masan" country="Italy" elevation="1170" location="Zambla Alta Village" municipality="Zambla Pass" pageId="8" pageNumber="103" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D72FD7E2E0EC464A343FADEFA32657A" box="[1354,1432,1290,1315]" pageId="8" pageNumber="103" type="male">1 male</specimenCount>
,
<location id="8EAB602C2E0EC464A69EFAFAFE2A651E" LSID="urn:lsid:plazi:treatment:03DD87E12E0EC467A69EFB83FB9A673A:8EAB602C2E0EC464A69EFAFAFE2A651E" box="[151,384,1326,1351]" country="Italy" municipality="Zambla Pass" name="Zambla Alta Village" pageId="8" pageNumber="103">Zambla Alta Village</location>
, near to
<collectingMunicipality id="6BAFAC8D2E0EC464A7E8FAFAFDD8651F" box="[481,626,1326,1351]" pageId="8" pageNumber="103">Zambla Pass</collectingMunicipality>
, spruce forest (
<taxonomicName id="4C744D742E0EC464A529FAFBFCD2651F" box="[800,888,1326,1350]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="8" pageNumber="103" phylum="Tracheophyta" rank="species" species="abies">
<emphasis id="B900EAE52E0EC464A529FAFBFC84651F" box="[800,814,1327,1350]" italics="true" pageId="8" pageNumber="103">P</emphasis>
.
<emphasis id="B900EAE52E0EC464A534FAFAFCD2651F" box="[829,888,1326,1350]" italics="true" pageId="8" pageNumber="103">abies</emphasis>
</taxonomicName>
) with admixed beech (
<taxonomicName id="4C744D742E0EC464A274FAFBFB55651F" box="[1149,1279,1326,1350]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="103" phylum="Tracheophyta" rank="species" species="sylvatica">
<emphasis id="B900EAE52E0EC464A274FAFBFB26651F" box="[1149,1164,1327,1350]" italics="true" pageId="8" pageNumber="103">F</emphasis>
.
<emphasis id="B900EAE52E0EC464A292FAFAFB55651F" box="[1179,1279,1326,1350]" italics="true" pageId="8" pageNumber="103">sylvatica</emphasis>
</taxonomicName>
), needle litter and soil detritus, with decomposed wood substrate, altitude
<quantity id="4C8C9B122E0EC464A53CFA86FC396533" box="[821,915,1362,1386]" metricMagnitude="3" metricUnit="m" metricValue="1.17" pageId="8" pageNumber="103" unit="m" value="1170.0">
<elevation id="0059D1C42E0EC464A53CFA86FC396533" box="[821,915,1362,1386]" metricMagnitude="3" metricUnit="m" metricValue="1.17" pageId="8" pageNumber="103" unit="m" value="1170.0">1,170 m</elevation>
</quantity>
,
<date id="FFCA10372E0EC464A596FA86FB9D6532" box="[927,1079,1362,1387]" pageId="8" pageNumber="103" value="2015-05-13">
<collectingDate id="EF8EE9DF2E0EC464A596FA86FB9D6532" box="[927,1079,1362,1387]" pageId="8" pageNumber="103" value="2015-05-13">13 May 2015</collectingDate>
</date>
, coll.
<collectorName id="268153212E0EC464A272FA87FB4E6533" box="[1147,1252,1362,1386]" pageId="8" pageNumber="103">P. Mašán</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36E657C2E0EC464A6CEFAA2FDA5661B" pageId="8" pageNumber="103" type="diagnosis">
<paragraph id="8BCB36F72E0EC464A6CEFAA2FDA5661B" blockId="8.[151,1437,1218,2035]" pageId="8" pageNumber="103">
<emphasis id="B900EAE52E0EC464A6CEFAA2FEEB65D6" bold="true" box="[199,321,1398,1423]" pageId="8" pageNumber="103">Diagnosis.</emphasis>
In female, tubiform structures of sperm induction system elongated, medially curved, with subdistal portions closely adjacent and directed posteriorly (
<figureCitation id="134F2A722E0EC464A53AFA4EFC2365EB" box="[819,905,1434,1459]" captionStart="FIGURES 29 35" captionStartId="10.[151,263,1834,1857]" captionTargetBox="[155,1415,198,1807]" captionTargetId="figure@10.[151,1436,193,1814]" captionTargetPageId="10" captionText="FIGURES 29 35. Pachylaelaps (Longipachylaelaps) marcovallei sp. nov. 29. Sperm induction structures, female; 30. Epistome, female; 31. Chelicera, female; 32. Tarsus II, female, mediodistal part; 33. Chelicera, male; 34. Ventral spurs on leg II (femur, genu and tibia), male; 35. Palptibial projections, male. Not scaled." httpUri="https://zenodo.org/record/322034/files/figure.png" pageId="8" pageNumber="103">Figs 29</figureCitation>
,
<figureCitation id="134F2A722E0EC464A59CFA4EFC7465EA" box="[917,990,1434,1459]" captionStart="FIGURES 49 52" captionStartId="15.[151,263,982,1005]" captionTargetBox="[151,1436,193,961]" captionTargetId="figure@15.[151,1436,193,961]" captionTargetPageId="15" captionText="FIGURES 49 52. Sperm induction structures, females, variant forms. 49 50. Pachylaelaps (Longipachylaelaps) marcovallei sp. nov. (49. Normal form; 50. Aberrant form in strongly pressed specimen); 51 52. Pachylaelaps (Longipachylaelaps) pantinii sp. nov. (51. Normal form; 52. Aberrant form in strongly pressed specimen). Not scaled." httpUri="https://zenodo.org/record/322038/files/figure.png" pageId="8" pageNumber="103">49, 50</figureCitation>
). Male spermatodactyl elongate, at least 2.5 times as long as movable digit, expanded and foliate proximally, moderately narrowed distally, with anterior submedial margin finely denticulated (
<figureCitation id="134F2A722E0EC464A459FA36FD0365A3" box="[592,681,1506,1531]" captionStart="FIGURES 29 35" captionStartId="10.[151,263,1834,1857]" captionTargetBox="[155,1415,198,1807]" captionTargetId="figure@10.[151,1436,193,1814]" captionTargetPageId="10" captionText="FIGURES 29 35. Pachylaelaps (Longipachylaelaps) marcovallei sp. nov. 29. Sperm induction structures, female; 30. Epistome, female; 31. Chelicera, female; 32. Tarsus II, female, mediodistal part; 33. Chelicera, male; 34. Ventral spurs on leg II (femur, genu and tibia), male; 35. Palptibial projections, male. Not scaled." httpUri="https://zenodo.org/record/322034/files/figure.png" pageId="8" pageNumber="103">Figs 33</figureCitation>
,
<figureCitation id="134F2A722E0EC464A4BEFA36FD7D65A2" box="[695,727,1506,1531]" captionStart="FIGURES 53 56" captionStartId="19.[151,263,1900,1923]" captionTargetBox="[151,1436,195,1879]" captionTargetId="figure@19.[151,1436,195,1879]" captionTargetPageId="19" captionText="FIGURES 53 56. Chelicerae, males, with variant forms of spermatodactyls. 53. Pachylaelaps (Longipachylaelaps) abludens sp. nov.; 54. Pachylaelaps (Longipachylaelaps) bergomensis sp. nov.; 55. Pachylaelaps (Longipachylaelaps) pantinii sp. nov.; 56. Pachylaelaps (Longipachylaelaps) marcovallei sp. nov. Not scaled." httpUri="https://zenodo.org/record/322039/files/figure.png" pageId="8" pageNumber="103">56</figureCitation>
). Palptibia with two divergent petal-like projections: antiaxial projection longer, expanded medially and rounded distally; paraxial projection shorer, subtriangular, and tapered toward obtuse apex (
<figureCitation id="134F2A722E0EC464A776F9FEFE7C661A" box="[383,470,1578,1603]" captionStart="FIGURES 29 35" captionStartId="10.[151,263,1834,1857]" captionTargetBox="[155,1415,198,1807]" captionTargetId="figure@10.[151,1436,193,1814]" captionTargetPageId="10" captionText="FIGURES 29 35. Pachylaelaps (Longipachylaelaps) marcovallei sp. nov. 29. Sperm induction structures, female; 30. Epistome, female; 31. Chelicera, female; 32. Tarsus II, female, mediodistal part; 33. Chelicera, male; 34. Ventral spurs on leg II (femur, genu and tibia), male; 35. Palptibial projections, male. Not scaled." httpUri="https://zenodo.org/record/322034/files/figure.png" pageId="8" pageNumber="103">Figs 35</figureCitation>
,
<figureCitation id="134F2A722E0EC464A7EBF9FEFDA8661B" box="[482,514,1578,1602]" captionStart="FIGURES 36 39" captionStartId="11.[151,263,874,897]" captionTargetBox="[151,1436,193,853]" captionTargetId="figure@11.[151,1436,193,853]" captionTargetPageId="11" captionText="FIGURES 36 39. Palptibial projections, males. 36. Pachylaelaps (Longipachylaelaps) abludens sp. nov.; 37. Pachylaelaps (Longipachylaelaps) bergomensis sp. nov.; 38. Pachylaelaps (Longipachylaelaps) marcovallei sp. nov.; 39. Pachylaelaps (Longipachylaelaps) pantinii sp. nov. Not scaled." httpUri="https://zenodo.org/record/322035/files/figure.png" pageId="8" pageNumber="103">38</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36E657C2E0EC467A6CEF99AFF516631" lastPageId="11" lastPageNumber="106" pageId="8" pageNumber="103" type="description">
<paragraph id="8BCB36F72E0EC464A6CEF99AFE71663E" blockId="8.[151,1437,1218,2035]" box="[199,475,1614,1639]" pageId="8" pageNumber="103">
<emphasis id="B900EAE52E0EC464A6CEF99AFE71663E" bold="true" box="[199,475,1614,1639]" pageId="8" pageNumber="103">
Description.
<emphasis id="B900EAE52E0EC464A756F99AFE7E663F" bold="true" box="[351,468,1614,1638]" italics="true" pageId="8" pageNumber="103">FEMALE</emphasis>
.
</emphasis>
</paragraph>
<paragraph id="8BCB36F72E0EC464A6CEF9A6FB016743" blockId="8.[151,1437,1218,2035]" pageId="8" pageNumber="103">
<emphasis id="B900EAE52E0EC464A6CEF9A6FE2C66D3" box="[199,390,1650,1674]" italics="true" pageId="8" pageNumber="103">Dorsal idiosoma</emphasis>
(
<figureCitation id="134F2A722E0EC464A790F9A6FE5A66D3" box="[409,496,1650,1675]" captionStart="FIGURES 27 28" captionStartId="9.[151,263,1844,1867]" captionTargetBox="[154,1434,737,1819]" captionTargetId="figure@9.[151,1436,733,1823]" captionTargetPageId="9" captionText="FIGURES 27 28. Pachylaelaps (Longipachylaelaps) marcovallei sp. nov., female. 14. Dorsal idiosoma; 15. Ventral idiosoma. Not scaled." httpUri="https://zenodo.org/record/322033/files/figure.png" pageId="8" pageNumber="103">Fig. 27</figureCitation>
). Dorsal shield 830865 µm long and 480505 µm wide, elongated and suboval (length/width 1.651.8), delicately and evenly reticulated on surface, bearing 30 pairs of dorsal setae. Dorsal setae uniform, smooth and needle-shaped, mostly subequal in length, relatively shorter and mostly with tips not reaching beyond insertions of following setae; setae z1 shortest. For length and spacing of some selected dorsal shield setae see
<tableCitation id="C6F6034C2E0EC464A6CBF8D6FEBF6743" box="[194,277,1794,1819]" captionStart="TABLE 1" captionStartId="12.[151,239,152,175]" captionTargetBox="[159,1374,202,1650]" captionTargetPageId="12" captionText="TABLE 1. Measurements (in µm) of four species of Pachylaelaps (Longipachylaelaps)." httpUri="http://table.plazi.org/id/DF0B667F2E0AC460A69EFF4CFBEF60F6" pageId="8" pageNumber="103" tableUuid="DF0B667F2E0AC460A69EFF4CFBEF60F6">Table 1</tableCitation>
. Dorsolateral soft integument with seven pairs of marginal setae (r6, R1, R3R7).
</paragraph>
<paragraph id="8BCB36F72E0EC464A6CEF8F2FA3767AA" blockId="8.[151,1437,1218,2035]" pageId="8" pageNumber="103">
<emphasis id="B900EAE52E0EC464A6CEF8F2FE2F6767" box="[199,389,1830,1854]" italics="true" pageId="8" pageNumber="103">Ventral idiosoma</emphasis>
(
<figureCitation id="134F2A722E0EC464A79DF8F2FE4D6766" box="[404,487,1830,1855]" captionStart="FIGURES 27 28" captionStartId="9.[151,263,1844,1867]" captionTargetBox="[154,1434,737,1819]" captionTargetId="figure@9.[151,1436,733,1823]" captionTargetPageId="9" captionText="FIGURES 27 28. Pachylaelaps (Longipachylaelaps) marcovallei sp. nov., female. 14. Dorsal idiosoma; 15. Ventral idiosoma. Not scaled." httpUri="https://zenodo.org/record/322033/files/figure.png" pageId="8" pageNumber="103">Fig. 28</figureCitation>
). Sternal shield 281296 µm long, slightly shorter than genitiventral shield (length of sternal shield/length of genitiventral shield 0.920.99). Genitiventral shield slightly as long as wide (length 296 306 µm, width 265278 µm, length/width 1.071.12). Anal shield subtriangular, 122134 µm long and 173181 µm wide (length/width 0.680.77); anus with three circum-anal setae situated close to posterior margin of shield. Peritremes normally developed, with anterior tips reaching dorsal surface between insertions of setae z1 and z2. Ventral shields with delicate reticulation on surface. Metapodal platelets minuscule, free on soft integument.
</paragraph>
<paragraph id="8BCB36F72E0FC465A69EFF43FEFB608C" blockId="9.[151,1437,151,680]" pageId="9" pageNumber="104">Ventral soft integument with nine pairs of ventral setae (JV2JV5, ZV1ZV5). Ventral setae similar to those on dorsal idiosoma.</paragraph>
<paragraph id="8BCB36F72E0FC465A6CEFF34FDEF613C" blockId="9.[151,1437,151,680]" pageId="9" pageNumber="104">
<emphasis id="B900EAE52E0FC465A6CEFF34FE5660A1" box="[199,508,224,248]" italics="true" pageId="9" pageNumber="104">Sperm induction structures</emphasis>
(
<figureCitation id="134F2A722E0FC465A404FF0BFDCD60A1" box="[525,615,223,248]" captionStart="FIGURES 29 35" captionStartId="10.[151,263,1834,1857]" captionTargetBox="[155,1415,198,1807]" captionTargetId="figure@10.[151,1436,193,1814]" captionTargetPageId="10" captionText="FIGURES 29 35. Pachylaelaps (Longipachylaelaps) marcovallei sp. nov. 29. Sperm induction structures, female; 30. Epistome, female; 31. Chelicera, female; 32. Tarsus II, female, mediodistal part; 33. Chelicera, male; 34. Ventral spurs on leg II (femur, genu and tibia), male; 35. Palptibial projections, male. Not scaled." httpUri="https://zenodo.org/record/322034/files/figure.png" pageId="9" pageNumber="104">Figs 29</figureCitation>
,
<figureCitation id="134F2A722E0FC465A47FFF34FD6E60A1" box="[630,708,223,248]" captionStart="FIGURES 49 52" captionStartId="15.[151,263,982,1005]" captionTargetBox="[151,1436,193,961]" captionTargetId="figure@15.[151,1436,193,961]" captionTargetPageId="15" captionText="FIGURES 49 52. Sperm induction structures, females, variant forms. 49 50. Pachylaelaps (Longipachylaelaps) marcovallei sp. nov. (49. Normal form; 50. Aberrant form in strongly pressed specimen); 51 52. Pachylaelaps (Longipachylaelaps) pantinii sp. nov. (51. Normal form; 52. Aberrant form in strongly pressed specimen). Not scaled." httpUri="https://zenodo.org/record/322038/files/figure.png" pageId="9" pageNumber="104">49, 50</figureCitation>
). Tubiform structures of sperm induction system contiguous to anterior inner margin of coxae IV, well developed and sclerotised (especially in their distal portions), relatively large, moderately elongated, widened basally, curved medially, regularly rounded apically, and with adjacent subdistal portions directed posteriorly.
</paragraph>
<paragraph id="8BCB36F72E0FC465A6CEFEA4FDDE6239" blockId="9.[151,1437,151,680]" pageId="9" pageNumber="104">
<emphasis id="B900EAE52E0FC465A6CEFEA4FE7C61D1" box="[199,470,368,392]" italics="true" pageId="9" pageNumber="104">Gnathosomal structures</emphasis>
(
<figureCitation id="134F2A722E0FC465A7EEFEBBFDC761D1" box="[487,621,367,392]" captionStart="FIGURES 29 35" captionStartId="10.[151,263,1834,1857]" captionTargetBox="[155,1415,198,1807]" captionTargetId="figure@10.[151,1436,193,1814]" captionTargetPageId="10" captionText="FIGURES 29 35. Pachylaelaps (Longipachylaelaps) marcovallei sp. nov. 29. Sperm induction structures, female; 30. Epistome, female; 31. Chelicera, female; 32. Tarsus II, female, mediodistal part; 33. Chelicera, male; 34. Ventral spurs on leg II (femur, genu and tibia), male; 35. Palptibial projections, male. Not scaled." httpUri="https://zenodo.org/record/322034/files/figure.png" pageId="9" pageNumber="104">Figs 30, 31</figureCitation>
). Corniculi elongated and horn-like; laciniae densely pilose, as long as corniculi; deutosternum with six rows of denticles; subcapitular setae smooth and needle-shaped. Epistome with subtriangular and regularly narrowed base, narrow central neck and widened apical part densely crenulate anteriorly; basal part with irregularly denticulate lateral margins (
<figureCitation id="134F2A722E0FC465A593FE08FC5961AD" box="[922,1011,476,501]" captionStart="FIGURES 29 35" captionStartId="10.[151,263,1834,1857]" captionTargetBox="[155,1415,198,1807]" captionTargetId="figure@10.[151,1436,193,1814]" captionTargetPageId="10" captionText="FIGURES 29 35. Pachylaelaps (Longipachylaelaps) marcovallei sp. nov. 29. Sperm induction structures, female; 30. Epistome, female; 31. Chelicera, female; 32. Tarsus II, female, mediodistal part; 33. Chelicera, male; 34. Ventral spurs on leg II (femur, genu and tibia), male; 35. Palptibial projections, male. Not scaled." httpUri="https://zenodo.org/record/322034/files/figure.png" pageId="9" pageNumber="104">Fig. 30</figureCitation>
). Fixed digit of chelicera with bifid terminal hook, one large distal tooth associated with pilus dentilis, and one additional distal denticle on lateral ridge; pilus dentilis well developed, curved and directed backward; movable digit of chelicera unidentate, with simple hook and one distal tooth (
<figureCitation id="134F2A722E0FC465A41AFD93FDCC6239" box="[531,614,583,608]" captionStart="FIGURES 29 35" captionStartId="10.[151,263,1834,1857]" captionTargetBox="[155,1415,198,1807]" captionTargetId="figure@10.[151,1436,193,1814]" captionTargetPageId="10" captionText="FIGURES 29 35. Pachylaelaps (Longipachylaelaps) marcovallei sp. nov. 29. Sperm induction structures, female; 30. Epistome, female; 31. Chelicera, female; 32. Tarsus II, female, mediodistal part; 33. Chelicera, male; 34. Ventral spurs on leg II (femur, genu and tibia), male; 35. Palptibial projections, male. Not scaled." httpUri="https://zenodo.org/record/322034/files/figure.png" pageId="9" pageNumber="104">Fig. 31</figureCitation>
).
</paragraph>
<paragraph id="8BCB36F72E0FC465A6CEFDB9FF6962F1" blockId="9.[151,1437,151,680]" pageId="9" pageNumber="104">
<emphasis id="B900EAE52E0FC465A6CEFDB9FF5662DD" box="[199,252,621,644]" italics="true" pageId="9" pageNumber="104">Legs</emphasis>
. Leg setation normal for genus (Mašán, 2007). Tarsus II with two spur-like distal setae, pl1 and pl2 (
<figureCitation id="134F2A722E0FC465A366FDB8FF1D62F1" captionStart="FIGURES 29 35" captionStartId="10.[151,263,1834,1857]" captionTargetBox="[155,1415,198,1807]" captionTargetId="figure@10.[151,1436,193,1814]" captionTargetPageId="10" captionText="FIGURES 29 35. Pachylaelaps (Longipachylaelaps) marcovallei sp. nov. 29. Sperm induction structures, female; 30. Epistome, female; 31. Chelicera, female; 32. Tarsus II, female, mediodistal part; 33. Chelicera, male; 34. Ventral spurs on leg II (femur, genu and tibia), male; 35. Palptibial projections, male. Not scaled." httpUri="https://zenodo.org/record/322034/files/figure.png" pageId="9" pageNumber="104">Fig. 32</figureCitation>
).
</paragraph>
<caption id="DF0B667F2E0FC465A69EF8E0FEA06730" httpUri="https://zenodo.org/record/322033/files/figure.png" pageId="9" pageNumber="104" targetBox="[154,1434,737,1819]" targetPageId="9">
<paragraph id="8BCB36F72E0FC465A69EF8E0FEA06730" blockId="9.[151,1437,1844,1897]" pageId="9" pageNumber="104">
<emphasis id="B900EAE52E0FC465A69EF8E0FEFE6713" bold="true" box="[151,340,1844,1867]" pageId="9" pageNumber="104">FIGURES 2728.</emphasis>
<taxonomicName id="4C744D742E0FC465A753F8E0FC926713" box="[346,824,1844,1866]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="104" phylum="Arthropoda" rank="species" species="marcovallei" status="sp. nov." subGenus="Longipachylaelaps">
<emphasis id="B900EAE52E0FC465A753F8E0FE496713" box="[346,483,1844,1866]" italics="true" pageId="9" pageNumber="104">Pachylaelaps</emphasis>
(
<emphasis id="B900EAE52E0FC465A7F8F8E0FD186713" box="[497,690,1844,1866]" italics="true" pageId="9" pageNumber="104">Longipachylaelaps</emphasis>
)
<emphasis id="B900EAE52E0FC465A4C9F8E0FC926713" box="[704,824,1844,1866]" italics="true" pageId="9" pageNumber="104">marcovallei</emphasis>
</taxonomicName>
<emphasis id="B900EAE52E0FC465A537F8E1FC276713" bold="true" box="[830,909,1845,1866]" pageId="9" pageNumber="104">
<taxonomicNameLabel id="A233579E2E0FC465A537F8E1FC276713" box="[830,909,1845,1866]" pageId="9" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female. 14. Dorsal idiosoma; 15. Ventral idiosoma. Not scaled.
</paragraph>
</caption>
<caption id="DF0B667F2E0CC466A69EF8FEFC2F6727" httpUri="https://zenodo.org/record/322034/files/figure.png" pageId="10" pageNumber="105" targetBox="[155,1415,198,1807]" targetPageId="10">
<paragraph id="8BCB36F72E0CC466A69EF8FEFC2F6727" blockId="10.[151,1436,1834,1918]" pageId="10" pageNumber="105">
<emphasis id="B900EAE52E0CC466A69EF8FEFEF16719" bold="true" box="[151,347,1834,1857]" pageId="10" pageNumber="105">FIGURES 2935.</emphasis>
<taxonomicName id="4C744D742E0CC466A761F8FFFCFE6718" box="[360,852,1835,1857]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="105" phylum="Arthropoda" rank="species" species="marcovallei" status="sp. nov." subGenus="Longipachylaelaps">
<emphasis id="B900EAE52E0CC466A761F8FFFE5B6718" box="[360,497,1835,1857]" italics="true" pageId="10" pageNumber="105">Pachylaelaps</emphasis>
(
<emphasis id="B900EAE52E0CC466A40FF8FFFD6D6718" box="[518,711,1835,1857]" italics="true" pageId="10" pageNumber="105">Longipachylaelaps</emphasis>
)
<emphasis id="B900EAE52E0CC466A4D5F8FFFCFE6718" box="[732,852,1835,1857]" italics="true" pageId="10" pageNumber="105">marcovallei</emphasis>
</taxonomicName>
<emphasis id="B900EAE52E0CC466A568F8FFFC126719" bold="true" box="[865,952,1835,1856]" pageId="10" pageNumber="105">
<taxonomicNameLabel id="A233579E2E0CC466A568F8FFFC126719" box="[865,952,1835,1856]" pageId="10" pageNumber="105" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
29. Sperm induction structures, female; 30. Epistome, female; 31. Chelicera, female; 32. Tarsus II, female, mediodistal part; 33. Chelicera, male; 34. Ventral spurs on leg II (femur, genu and tibia), male; 35. Palptibial projections, male. Not scaled.
</paragraph>
</caption>
<caption id="DF0B667F2E0DC467A69EFCBEFD2463E7" httpUri="https://zenodo.org/record/322035/files/figure.png" pageId="11" pageNumber="106" targetBox="[151,1436,193,853]" targetPageId="11">
<paragraph id="8BCB36F72E0DC467A69EFCBEFD2463E7" blockId="11.[151,1436,874,958]" pageId="11" pageNumber="106">
<emphasis id="B900EAE52E0DC467A69EFCBEFEFD63D9" bold="true" box="[151,343,874,897]" pageId="11" pageNumber="106">FIGURES 3639.</emphasis>
Palptibial projections, males. 36.
<taxonomicName id="4C744D742E0DC467A4B3FCBFFB2B63D8" box="[698,1153,875,897]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="106" phylum="Arthropoda" rank="species" species="abludens" status="sp. nov." subGenus="Longipachylaelaps">
<emphasis id="B900EAE52E0DC467A4B3FCBFFCE963D8" box="[698,835,875,897]" italics="true" pageId="11" pageNumber="106">Pachylaelaps</emphasis>
(
<emphasis id="B900EAE52E0DC467A55AFCBFFBBE63D8" box="[851,1044,875,897]" italics="true" pageId="11" pageNumber="106">Longipachylaelaps</emphasis>
)
<emphasis id="B900EAE52E0DC467A22CFCBFFB2B63D8" box="[1061,1153,875,897]" italics="true" pageId="11" pageNumber="106">abludens</emphasis>
</taxonomicName>
<emphasis id="B900EAE52E0DC467A280FCBFFB7163D9" bold="true" box="[1161,1243,875,896]" pageId="11" pageNumber="106">
<taxonomicNameLabel id="A233579E2E0DC467A280FCBFFB7163D9" box="[1161,1243,875,896]" pageId="11" pageNumber="106" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 37.
<taxonomicName id="4C744D742E0DC467A31DFCBFFE5863C6" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="106" phylum="Arthropoda" rank="species" species="bergomensis" status="sp. nov." subGenus="Longipachylaelaps">
<emphasis id="B900EAE52E0DC467A31DFCBFFA3663D8" box="[1300,1436,875,897]" italics="true" pageId="11" pageNumber="106">Pachylaelaps</emphasis>
(
<emphasis id="B900EAE52E0DC467A696FC5DFECA63C6" box="[159,352,905,927]" italics="true" pageId="11" pageNumber="106">Longipachylaelaps</emphasis>
)
<emphasis id="B900EAE52E0DC467A77AFC5DFE5863C6" box="[371,498,905,927]" italics="true" pageId="11" pageNumber="106">bergomensis</emphasis>
</taxonomicName>
<emphasis id="B900EAE52E0DC467A7F4FC5EFDF863C6" bold="true" box="[509,594,906,927]" pageId="11" pageNumber="106">
<taxonomicNameLabel id="A233579E2E0DC467A7F4FC5EFDF863C6" box="[509,594,906,927]" pageId="11" pageNumber="106" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 38.
<taxonomicName id="4C744D742E0DC467A486FC5DFBDD63C6" box="[655,1143,905,927]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="106" phylum="Arthropoda" rank="species" species="marcovallei" status="sp. nov." subGenus="Longipachylaelaps">
<emphasis id="B900EAE52E0DC467A486FC5DFCBD63C6" box="[655,791,905,927]" italics="true" pageId="11" pageNumber="106">Pachylaelaps</emphasis>
(
<emphasis id="B900EAE52E0DC467A522FC5DFC4663C6" box="[811,1004,905,927]" italics="true" pageId="11" pageNumber="106">Longipachylaelaps</emphasis>
)
<emphasis id="B900EAE52E0DC467A5F6FC5DFBDD63C6" box="[1023,1143,905,927]" italics="true" pageId="11" pageNumber="106">marcovallei</emphasis>
</taxonomicName>
<emphasis id="B900EAE52E0DC467A28BFC5EFB7D63C6" bold="true" box="[1154,1239,906,927]" pageId="11" pageNumber="106">
<taxonomicNameLabel id="A233579E2E0DC467A28BFC5EFB7D63C6" box="[1154,1239,906,927]" pageId="11" pageNumber="106" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 39.
<taxonomicName id="4C744D742E0DC467A31DFC5DFE1463E7" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="106" phylum="Arthropoda" rank="species" species="pantinii" status="sp. nov." subGenus="Longipachylaelaps">
<emphasis id="B900EAE52E0DC467A31DFC5DFA3663C6" box="[1300,1436,905,927]" italics="true" pageId="11" pageNumber="106">Pachylaelaps</emphasis>
(
<emphasis id="B900EAE52E0DC467A696FC7CFECA63E7" box="[159,352,936,958]" italics="true" pageId="11" pageNumber="106">Longipachylaelaps</emphasis>
)
<emphasis id="B900EAE52E0DC467A766FC7DFE1463E7" box="[367,446,937,958]" italics="true" pageId="11" pageNumber="106">pantinii</emphasis>
</taxonomicName>
<emphasis id="B900EAE52E0DC467A7CDFC7CFDBE63E4" bold="true" box="[452,532,936,957]" pageId="11" pageNumber="106">
<taxonomicNameLabel id="A233579E2E0DC467A7CDFC7CFDBE63E4" box="[452,532,936,957]" pageId="11" pageNumber="106" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Not scaled.
</paragraph>
</caption>
<paragraph id="8BCB36F72E0DC467A6CEFC3FFE336471" blockId="11.[151,1437,1003,1892]" pageId="11" pageNumber="106">
<emphasis id="B900EAE52E0DC467A6CEFC3FFEA6645B" bold="true" box="[199,268,1003,1026]" italics="true" pageId="11" pageNumber="106">MALE</emphasis>
<emphasis id="B900EAE52E0DC467A702FC3FFEB8645D" bold="true" box="[267,274,1003,1028]" pageId="11" pageNumber="106">.</emphasis>
<emphasis id="B900EAE52E0DC467A717FC3FFE75645A" box="[286,479,1003,1027]" italics="true" pageId="11" pageNumber="106">Dorsal idiosoma</emphasis>
. Dorsal shield 710750 µm long and 430500 µm wide, elongated and suboval (length/width 1.51.7).
</paragraph>
<paragraph id="8BCB36F72E0DC467A6CEFBE7FAF664E1" blockId="11.[151,1437,1003,1892]" pageId="11" pageNumber="106">
<emphasis id="B900EAE52E0DC467A6CEFBE7FE216412" box="[199,395,1075,1099]" italics="true" pageId="11" pageNumber="106">Ventral idiosoma</emphasis>
. Sternal, genitiventral, peritrematal, metapodal and anal plates fused to form an entire holoventral shield bearing nine pairs of setae (st1st5, JV1JV3, ZV2), not including three circum-anal setae close to anus; shield with reticulate sculpture on surface. Dorsolateral and ventral soft integument with 13 pairs of setae (r6, R1, R3R7, JV4, JV5, Zv1, ZV3ZV5). Dorsal and ventral chaetotaxy and other characters as in female.
</paragraph>
<paragraph id="8BCB36F72E0DC467A6CEFB17FE696581" blockId="11.[151,1437,1003,1892]" pageId="11" pageNumber="106">
<emphasis id="B900EAE52E0DC467A6CEFB17FE7C6482" box="[199,470,1219,1243]" italics="true" pageId="11" pageNumber="106">Gnathosomal structures</emphasis>
(
<figureCitation id="134F2A722E0DC467A7EEFB17FDC66485" box="[487,620,1219,1244]" captionStart="FIGURES 29 35" captionStartId="10.[151,263,1834,1857]" captionTargetBox="[155,1415,198,1807]" captionTargetId="figure@10.[151,1436,193,1814]" captionTargetPageId="10" captionText="FIGURES 29 35. Pachylaelaps (Longipachylaelaps) marcovallei sp. nov. 29. Sperm induction structures, female; 30. Epistome, female; 31. Chelicera, female; 32. Tarsus II, female, mediodistal part; 33. Chelicera, male; 34. Ventral spurs on leg II (femur, genu and tibia), male; 35. Palptibial projections, male. Not scaled." httpUri="https://zenodo.org/record/322034/files/figure.png" pageId="11" pageNumber="106">Figs 33, 35</figureCitation>
,
<figureCitation id="134F2A722E0DC467A472FB17FD336482" box="[635,665,1219,1243]" captionStart="FIGURES 36 39" captionStartId="11.[151,263,874,897]" captionTargetBox="[151,1436,193,853]" captionTargetId="figure@11.[151,1436,193,853]" captionTargetPageId="11" captionText="FIGURES 36 39. Palptibial projections, males. 36. Pachylaelaps (Longipachylaelaps) abludens sp. nov.; 37. Pachylaelaps (Longipachylaelaps) bergomensis sp. nov.; 38. Pachylaelaps (Longipachylaelaps) marcovallei sp. nov.; 39. Pachylaelaps (Longipachylaelaps) pantinii sp. nov. Not scaled." httpUri="https://zenodo.org/record/322035/files/figure.png" pageId="11" pageNumber="106">38</figureCitation>
,
<figureCitation id="134F2A722E0DC467A4AEFB17FD6D6485" box="[679,711,1219,1244]" captionStart="FIGURES 53 56" captionStartId="19.[151,263,1900,1923]" captionTargetBox="[151,1436,195,1879]" captionTargetId="figure@19.[151,1436,195,1879]" captionTargetPageId="19" captionText="FIGURES 53 56. Chelicerae, males, with variant forms of spermatodactyls. 53. Pachylaelaps (Longipachylaelaps) abludens sp. nov.; 54. Pachylaelaps (Longipachylaelaps) bergomensis sp. nov.; 55. Pachylaelaps (Longipachylaelaps) pantinii sp. nov.; 56. Pachylaelaps (Longipachylaelaps) marcovallei sp. nov. Not scaled." httpUri="https://zenodo.org/record/322039/files/figure.png" pageId="11" pageNumber="106">56</figureCitation>
). Cheliceral spermatodactyl well developed, elongate, 16 5185 µm long, 2.52.85 times as long as movable digit, foliately expanded proximally, moderately narrowed in distalmost portion, with anterior submedial margin finely denticulated (
<figureCitation id="134F2A722E0DC467A5C8FADFFBB3657A" box="[961,1049,1291,1316]" captionStart="FIGURES 29 35" captionStartId="10.[151,263,1834,1857]" captionTargetBox="[155,1415,198,1807]" captionTargetId="figure@10.[151,1436,193,1814]" captionTargetPageId="10" captionText="FIGURES 29 35. Pachylaelaps (Longipachylaelaps) marcovallei sp. nov. 29. Sperm induction structures, female; 30. Epistome, female; 31. Chelicera, female; 32. Tarsus II, female, mediodistal part; 33. Chelicera, male; 34. Ventral spurs on leg II (femur, genu and tibia), male; 35. Palptibial projections, male. Not scaled." httpUri="https://zenodo.org/record/322034/files/figure.png" pageId="11" pageNumber="106">Figs 33</figureCitation>
,
<figureCitation id="134F2A722E0DC467A22FFADFFBEC657D" box="[1062,1094,1291,1316]" captionStart="FIGURES 53 56" captionStartId="19.[151,263,1900,1923]" captionTargetBox="[151,1436,195,1879]" captionTargetId="figure@19.[151,1436,195,1879]" captionTargetPageId="19" captionText="FIGURES 53 56. Chelicerae, males, with variant forms of spermatodactyls. 53. Pachylaelaps (Longipachylaelaps) abludens sp. nov.; 54. Pachylaelaps (Longipachylaelaps) bergomensis sp. nov.; 55. Pachylaelaps (Longipachylaelaps) pantinii sp. nov.; 56. Pachylaelaps (Longipachylaelaps) marcovallei sp. nov. Not scaled." httpUri="https://zenodo.org/record/322039/files/figure.png" pageId="11" pageNumber="106">56</figureCitation>
). Palptibiae slightly thickened basally, each bearing a pair of petal-like projections on proximal ventral surface, as in
<figureCitation id="134F2A722E0DC467A261FAFBFB166511" box="[1128,1212,1327,1352]" captionStart="FIGURES 29 35" captionStartId="10.[151,263,1834,1857]" captionTargetBox="[155,1415,198,1807]" captionTargetId="figure@10.[151,1436,193,1814]" captionTargetPageId="10" captionText="FIGURES 29 35. Pachylaelaps (Longipachylaelaps) marcovallei sp. nov. 29. Sperm induction structures, female; 30. Epistome, female; 31. Chelicera, female; 32. Tarsus II, female, mediodistal part; 33. Chelicera, male; 34. Ventral spurs on leg II (femur, genu and tibia), male; 35. Palptibial projections, male. Not scaled." httpUri="https://zenodo.org/record/322034/files/figure.png" pageId="11" pageNumber="106">Figs 35</figureCitation>
and
<figureCitation id="134F2A722E0DC467A2FDFAFBFAB9651E" box="[1268,1299,1327,1351]" captionStart="FIGURES 36 39" captionStartId="11.[151,263,874,897]" captionTargetBox="[151,1436,193,853]" captionTargetId="figure@11.[151,1436,193,853]" captionTargetPageId="11" captionText="FIGURES 36 39. Palptibial projections, males. 36. Pachylaelaps (Longipachylaelaps) abludens sp. nov.; 37. Pachylaelaps (Longipachylaelaps) bergomensis sp. nov.; 38. Pachylaelaps (Longipachylaelaps) marcovallei sp. nov.; 39. Pachylaelaps (Longipachylaelaps) pantinii sp. nov. Not scaled." httpUri="https://zenodo.org/record/322035/files/figure.png" pageId="11" pageNumber="106">38</figureCitation>
; projections relatively large, with divergent adjacent margins: antiaxial projection longer, with narrow base, medial expansion and rounded distal margin, lacking small lamellar structure in its inner basal margin; paraxial projection shorer, subtriangular, widened basally and tapered toward obtuse apex. Palpfemur with distinct papular tubercle on anterolateral basal surface.
</paragraph>
<paragraph id="8BCB36F72E0DC467A6CEFA30FF516631" blockId="11.[151,1437,1003,1892]" pageId="11" pageNumber="106">
<emphasis id="B900EAE52E0DC467A6CEFA30FF5665A2" box="[199,252,1508,1531]" italics="true" pageId="11" pageNumber="106">Legs</emphasis>
. Medial segments of legs II spurred on their distal ventral surface: femur with one robust spur, genu and tibia each with a peg-like spur, as in
<figureCitation id="134F2A722E0DC467A424F9D3FDD56646" box="[557,639,1543,1568]" captionStart="FIGURES 29 35" captionStartId="10.[151,263,1834,1857]" captionTargetBox="[155,1415,198,1807]" captionTargetId="figure@10.[151,1436,193,1814]" captionTargetPageId="10" captionText="FIGURES 29 35. Pachylaelaps (Longipachylaelaps) marcovallei sp. nov. 29. Sperm induction structures, female; 30. Epistome, female; 31. Chelicera, female; 32. Tarsus II, female, mediodistal part; 33. Chelicera, male; 34. Ventral spurs on leg II (femur, genu and tibia), male; 35. Palptibial projections, male. Not scaled." httpUri="https://zenodo.org/record/322034/files/figure.png" pageId="11" pageNumber="106">Fig. 34</figureCitation>
. Femoral spur broadened in basal part, produced into narrowly rounded apex, with two small tubercles on axillary side (
<figureCitation id="134F2A722E0DC467A4B5F9FFFCB8661A" box="[700,786,1579,1604]" captionStart="FIGURES 29 35" captionStartId="10.[151,263,1834,1857]" captionTargetBox="[155,1415,198,1807]" captionTargetId="figure@10.[151,1436,193,1814]" captionTargetPageId="10" captionText="FIGURES 29 35. Pachylaelaps (Longipachylaelaps) marcovallei sp. nov. 29. Sperm induction structures, female; 30. Epistome, female; 31. Chelicera, female; 32. Tarsus II, female, mediodistal part; 33. Chelicera, male; 34. Ventral spurs on leg II (femur, genu and tibia), male; 35. Palptibial projections, male. Not scaled." httpUri="https://zenodo.org/record/322034/files/figure.png" pageId="11" pageNumber="106">Fig. 34</figureCitation>
). Terminal part of tarsus II with only one spur-like distal seta, pl1.
</paragraph>
</subSubSection>
<subSubSection id="C36E657C2E0DC467A6CEF9A7FB9A673A" pageId="11" pageNumber="106" type="etymology">
<paragraph id="8BCB36F72E0DC467A6CEF9A7FA93668D" blockId="11.[151,1437,1003,1892]" pageId="11" pageNumber="106">
<emphasis id="B900EAE52E0DC467A6CEF9A7FEE466D5" bold="true" box="[199,334,1651,1676]" pageId="11" pageNumber="106">Etymology.</emphasis>
This species is named in honour of Marco Valle, eminent and esteemed zoologist and museologist, director of the Natural History Museum in Bergamo (Museo Civico di Scienze Naturali &quot;Enrico Caffi&quot;), who did many years of patient work especially on the diversity and biology of the Italian caddisflies (Trichoptera).
</paragraph>
<paragraph id="8BCB36F72E0DC467A6CEF90BFB9A673A" blockId="11.[151,1437,1003,1892]" pageId="11" pageNumber="106">
<emphasis id="B900EAE52E0DC467A6CEF90BFE3C66A1" bold="true" box="[199,406,1759,1784]" pageId="11" pageNumber="106">Taxonomic notes.</emphasis>
The main diagnostic character states for
<taxonomicName id="4C744D742E0DC467A56BF90BFAD966AE" box="[866,1395,1759,1783]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="106" phylum="Arthropoda" rank="species" species="marcovallei" subGenus="Longipachylaelaps">
<emphasis id="B900EAE52E0DC467A56BF90BFC5366AE" box="[866,1017,1759,1783]" italics="true" pageId="11" pageNumber="106">Pachylaelaps</emphasis>
(
<emphasis id="B900EAE52E0DC467A200F90BFB7466AE" box="[1033,1246,1759,1783]" italics="true" pageId="11" pageNumber="106">Longipachylaelaps</emphasis>
)
<emphasis id="B900EAE52E0DC467A2E7F90BFAD966AE" box="[1262,1395,1759,1783]" italics="true" pageId="11" pageNumber="106">marcovallei</emphasis>
</taxonomicName>
are the presence of &quot;pendulous&quot; sperm tubes in females and the denticulation of the anterior submedial margin of the spermatodactyl in males. This denticulation has not been reported before in the family
<taxonomicName id="4C744D742E0DC467A2AEF8F3FACB6719" box="[1191,1377,1831,1856]" class="Arachnida" family="Pachylaelapidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="106" phylum="Arthropoda" rank="family">Pachylaelapidae</taxonomicName>
. The characters that separate the new species from its congeners are detailed in
<tableCitation id="C6F6034C2E0DC467A5DEF89FFB87673A" box="[983,1069,1867,1892]" captionStart="TABLE 2" captionStartId="16.[146,221,530,550]" captionText="TABLE 2. Comparative characteristics of selected features in the Pachylaelaps (Longipachylaelaps) species. SL = length of cheliceral spermatodactyl, MDL = length of movable digit of chelicera, ― = unknown adult stage,? = unknown character)." pageId="11" pageNumber="106">Table 2</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>