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<document ID-DOI="10.3853/j.2201-4349.69.2017.1653" ID-ISSN="2201-4349" ID-Zenodo-Dep="5237800" ID-ZooBank="68F315FF-3FEB-410E-96EC-5F494510F440" checkinTime="1629535407606" checkinUser="felipe" docAuthor="Parnaby, Harry E., Ingleby, Sandy &amp; Divljan, Anja" docDate="2017" docId="03DD87C8FFD6735A1936F9A2FC43937E" docLanguage="en" docName="RecAustMus.69.5.277-420.pdf" docOrigin="Records of the Australian Museum 69 (5)" docSource="http://dx.doi.org/10.3853/j.2201-4349.69.2017.1653" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Animalia" docType="treatment" docVersion="2" lastPageNumber="388" masterDocId="FFE4FFB0FFB973351832FFEBFFD5973C" masterDocTitle="Type Specimens of Non-fossil Mammals in the Australian Museum, Sydney" masterLastPageNumber="420" masterPageNumber="277" pageNumber="388" updateTime="1674492396722" updateUser="juliana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Type Specimens of Non-fossil Mammals in the Australian Museum, Sydney</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Parnaby, Harry E.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ingleby, Sandy</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Divljan, Anja</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Records of the Australian Museum</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="pubDate">
<mods:number>2017-10-06</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>69</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>5</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>277</mods:start>
<mods:end>420</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3853/j.2201-4349.69.2017.1653</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3853/j.2201-4349.69.2017.1653</mods:identifier>
<mods:identifier type="ISSN">2201-4349</mods:identifier>
<mods:identifier type="Zenodo-Dep">5237800</mods:identifier>
<mods:identifier type="ZooBank">68F315FF-3FEB-410E-96EC-5F494510F440</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:03DD87C8FFD6735A1936F9A2FC43937E" httpUri="http://treatment.plazi.org/id/03DD87C8FFD6735A1936F9A2FC43937E" lastPageNumber="388" pageId="111" pageNumber="388">
<subSubSection pageId="111" pageNumber="388" type="nomenclature">
<paragraph blockId="111.[260,610,1608,1672]" pageId="111" pageNumber="388">
<taxonomicName ID-CoL="72RG" ID-ENA="187267" authority="Flannery &amp; White, 1991" kingdom="Animalia" pageId="111" pageNumber="388" rank="subSpecies">
<heading box="[260,610,1608,1635]" centered="true" fontSize="11" level="2" pageId="111" pageNumber="388" reason="2">
<emphasis bold="true" box="[260,610,1608,1635]" italics="true" pageId="111" pageNumber="388">Pteropus temmincki ennisae</emphasis>
</heading>
<emphasis bold="true" box="[280,589,1645,1672]" pageId="111" pageNumber="388">Flannery &amp; White, 1991</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="111" pageNumber="388" type="reference_group">
<paragraph blockId="111.[151,718,1692,1741]" pageId="111" pageNumber="388">
<emphasis box="[151,616,1692,1713]" italics="true" pageId="111" pageNumber="388">National Geographic Research and Exploration</emphasis>
7(1): 100, figs 1, 4, tables 23. (1991: May or earlier).
</paragraph>
</subSubSection>
<subSubSection box="[121,539,1762,1786]" pageId="111" pageNumber="388" type="vernacular_names">
<paragraph blockId="111.[121,539,1762,1786]" box="[121,539,1762,1786]" pageId="111" pageNumber="388">
<emphasis bold="true" box="[121,295,1762,1786]" pageId="111" pageNumber="388">Common name</emphasis>
. Bismarck Flying-fox.
</paragraph>
</subSubSection>
<subSubSection pageId="111" pageNumber="388" type="description">
<paragraph blockId="111.[120,748,1807,1861]" pageId="111" pageNumber="388">
<emphasis bold="true" box="[121,282,1807,1832]" pageId="111" pageNumber="388">Current name</emphasis>
.
<taxonomicName authority="Flannery &amp; White, 1991" authorityName="Flannery &amp; White" authorityYear="1991" box="[294,744,1807,1832]" class="Dothideomycetes" family="Pteropodidae" genus="Pteropus" kingdom="Fungi" order="Chiroptera" pageId="111" pageNumber="388" phylum="Ascomycota" rank="species" species="ennisae">
<emphasis box="[294,477,1808,1831]" italics="true" pageId="111" pageNumber="388">Pteropus ennisae</emphasis>
Flannery &amp; White, 1991
</taxonomicName>
; following
<bibRefCitation author="Almeida, F &amp; Giannini, N" box="[229,463,1837,1861]" pageId="111" pageNumber="388" pagination="83 - 95" refId="ref124454" refString="Almeida, F. C., N. P. Giannini, N. B. Simmons, and K. M. Helgen. 2014. Each flying fox on its own branch: A phylogenetic tree for Pteropus and related genera (Chiroptera: Pteropodidae). Molecular Phylogenetics and Evolution 77: 83 - 95. https: // doi. org / 10.1016 / j. ympev. 2014.03.009" type="journal article" year="2014">
Almeida
<emphasis box="[327,381,1837,1861]" italics="true" pageId="111" pageNumber="388">et al.</emphasis>
(2014)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="111" pageNumber="388" type="materials_examined">
<paragraph blockId="111.[121,750,1882,1994]" lastBlockId="111.[792,1423,151,264]" pageId="111" pageNumber="388">
<emphasis bold="true" box="[121,225,1882,1906]" pageId="111" pageNumber="388">
<typeStatus box="[121,225,1882,1906]" pageId="111" pageNumber="388">Holotype</typeStatus>
</emphasis>
. M.20438 by original designation. Female adult, [Field no. FE419], skull, study skin, skinned body in alc., frozen tissue; collected on
<date box="[411,554,1941,1965]" pageId="111" pageNumber="388" value="1988-06-20">20 June 1988</date>
by T. F. Flannery, L. Seri, T. Heinsohn and T. Ennis; registered on
<date box="[618,745,1970,1994]" pageId="111" pageNumber="388" value="1989-06-06">6 June 1989</date>
.
<emphasis bold="true" box="[794,907,151,175]" pageId="111" pageNumber="388">Condition</emphasis>
. Cranium complete, soft palate
<emphasis box="[1236,1299,152,175]" italics="true" pageId="111" pageNumber="388">in situ</emphasis>
, hole in the right maxilla, and left temporal bone; broken left dentary, missing entire section posterior to last molar. Study skin has large bald patch on rump, otherwise in good condition.
</paragraph>
</subSubSection>
<subSubSection pageId="111" pageNumber="388" type="description">
<paragraph blockId="111.[792,1422,285,426]" pageId="111" pageNumber="388">
<emphasis bold="true" box="[794,1121,285,309]" pageId="111" pageNumber="388">Cranial measurements (mm)</emphasis>
. M.20438: GL, 55.5; ConL, 53.25; NasL, 14.70; NasB, 5.83; UC1C1 (alv.), 10.56; PAL, 31.09; UPM (alv.), 3.69; UMR (alv.), 6.72; ZB, 30.69; POC, 8.64; BUL, 4.19 (length of annula); MB, 18.30; DL (condyl.), 42.41 (right dentary); LPM (alv.), 3.76; LMR (alv.), 8.90.
</paragraph>
</subSubSection>
<subSubSection pageId="111" pageNumber="388" type="distribution">
<paragraph blockId="111.[792,1421,447,501]" pageId="111" pageNumber="388">
<emphasis bold="true" box="[794,934,447,472]" pageId="111" pageNumber="388">
<typeStatus box="[794,849,448,472]" pageId="111" pageNumber="388">Type</typeStatus>
locality
</emphasis>
. Near Medina (
<geoCoordinate box="[1095,1164,447,471]" degrees="2" direction="south" minutes="55" orientation="latitude" pageId="111" pageNumber="388" precision="925" value="-2.9166667">2°55'S</geoCoordinate>
<geoCoordinate box="[1169,1269,447,471]" degrees="151" direction="east" minutes="23" orientation="longitude" pageId="111" pageNumber="388" precision="925" value="151.38333">151°23'E</geoCoordinate>
),
<collectingRegion box="[1286,1417,448,472]" country="Papua New Guinea" name="New Ireland" pageId="111" pageNumber="388">New Ireland</collectingRegion>
,
<collectingRegion box="[792,1029,477,501]" country="Papua New Guinea" name="New Ireland" pageId="111" pageNumber="388">New Ireland Province</collectingRegion>
,
<collectingCountry box="[1039,1249,477,501]" name="Papua New Guinea" pageId="111" pageNumber="388">Papua New Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection pageId="111" pageNumber="388" type="materials_examined">
<paragraph blockId="111.[792,1422,522,722]" pageId="111" pageNumber="388">
<emphasis bold="true" box="[794,898,522,546]" pageId="111" pageNumber="388">
<typeStatus box="[794,898,522,546]" pageId="111" pageNumber="388">Paratype</typeStatus>
</emphasis>
. M.19904 by original designation. Male, study skin, skull, skinned body in alc., collected
<date box="[1272,1421,552,576]" pageId="111" pageNumber="388" value="1988-06-24">24 June 1988</date>
(paper states 23 June), in the Medina area by same collectors as the
<typeStatus box="[861,958,610,634]" pageId="111" pageNumber="388">holotype</typeStatus>
. The remaining
<typeStatus box="[1131,1226,611,635]" pageId="111" pageNumber="388">paratype</typeStatus>
, M.20804, female adult, skull, body in alc., frozen tissue; collected
<date pageId="111" pageNumber="388" value="1988-06-26">26 June 1988</date>
, with the same locality and collectors as
<typeStatus box="[1274,1370,669,693]" pageId="111" pageNumber="388">holotype</typeStatus>
, was sent to
<collectionCode box="[869,949,699,722]" pageId="111" pageNumber="388">BMNH</collectionCode>
in
<date box="[983,1157,698,722]" pageId="111" pageNumber="388" value="1989-12">December 1989</date>
.
</paragraph>
</subSubSection>
<subSubSection pageId="111" pageNumber="388" type="discussion">
<paragraph blockId="111.[792,1422,743,1090]" pageId="111" pageNumber="388">
<emphasis bold="true" box="[794,917,743,767]" pageId="111" pageNumber="388">Comments</emphasis>
. Type series of
<specimenCount box="[1079,1253,744,768]" pageId="111" pageNumber="388" type="generic">three specimens</specimenCount>
. Photographs of the
<typeStatus box="[829,921,773,797]" pageId="111" pageNumber="388">holotype</typeStatus>
skull are given in the original description, along with individual body measurements and summary statistics of two cranial and three dental measurements of the type series.
<bibRefCitation author="Flannery, T" box="[866,1049,861,885]" pageId="111" pageNumber="388" refId="ref129017" refString="Flannery, T. 1995 c. Mammals of the South-West Pacific and Moluccan Islands. Chatswood, New South Wales: Australian Museum / Reed Books. 464 pp." type="book" year="1995">Flannery (1995c)</bibRefCitation>
and subsequent authors considered
<taxonomicName authority="Peters, 1867 b" authorityName="Peters" authorityYear="1867" box="[792,1080,890,914]" class="Dothideomycetes" family="Pteropodidae" genus="Pteropus" kingdom="Fungi" order="Chiroptera" pageId="111" pageNumber="388" phylum="Ascomycota" rank="species" species="temminckii">
<emphasis box="[792,934,890,914]" italics="true" pageId="111" pageNumber="388">P. temminckii</emphasis>
<bibRefCitation author="Peters, W" box="[938,1080,890,914]" pageId="111" pageNumber="388" pagination="319 - 333" refId="ref141312" refString="Peters, W. 1867 b. Uber die Flederhunde, Pteropi, und insbesondere uber die Arten der Gattung Pteropus s. s. Monatsberichte der Koniglich Preussischen Akademie der Wissenschaften zu Berlin 1867: 319 - 333." type="journal article" year="1867">Peters, 1867b</bibRefCitation>
</taxonomicName>
and
<taxonomicName authority="Peters, 1876 b" authorityName="Peters" authorityYear="1876" box="[1129,1421,890,914]" class="Dothideomycetes" family="Pteropodidae" genus="Pteropus" kingdom="Fungi" order="Chiroptera" pageId="111" pageNumber="388" phylum="Ascomycota" rank="species" species="capistratus">
<emphasis box="[1129,1271,891,914]" italics="true" pageId="111" pageNumber="388">P. capistratus</emphasis>
<bibRefCitation author="Peters, W" box="[1278,1421,890,914]" pageId="111" pageNumber="388" pagination="315 - 319" refId="ref141474" refString="Peters, W. 1876 b. Mittheilung uber die Pelzrobbe von den Inseln St. Paul und Amsterdam und uber die von S. M. S. Gazelle mitgebrachten Flederthiere. Monatsberichte der Koniglich Preussischen Akademie der Wissenschaften zu Berlin 1876: 315 - 319, + pl." type="journal article" year="1876">Peters, 1876b</bibRefCitation>
</taxonomicName>
to be separate species, with
<taxonomicName authorityName="Flannery &amp; White" authorityYear="1991" box="[1099,1180,920,943]" class="Dothideomycetes" family="Pteropodidae" genus="Pteropus" kingdom="Fungi" order="Chiroptera" pageId="111" pageNumber="388" phylum="Ascomycota" rank="species" species="ennisae">
<emphasis box="[1099,1180,920,943]" italics="true" pageId="111" pageNumber="388">ennisae</emphasis>
</taxonomicName>
as a subspecies of the latter.
<bibRefCitation author="Almeida, F &amp; Giannini, N" box="[866,1113,949,973]" pageId="111" pageNumber="388" pagination="83 - 95" refId="ref124454" refString="Almeida, F. C., N. P. Giannini, N. B. Simmons, and K. M. Helgen. 2014. Each flying fox on its own branch: A phylogenetic tree for Pteropus and related genera (Chiroptera: Pteropodidae). Molecular Phylogenetics and Evolution 77: 83 - 95. https: // doi. org / 10.1016 / j. ympev. 2014.03.009" type="journal article" year="2014">
Almeida
<emphasis box="[970,1029,949,973]" italics="true" pageId="111" pageNumber="388">et al.</emphasis>
(2014)
</bibRefCitation>
suggest that
<taxonomicName authorityName="Peters" authorityYear="1876" box="[1268,1421,950,973]" class="Dothideomycetes" family="Pteropodidae" genus="Pteropus" kingdom="Fungi" order="Chiroptera" pageId="111" pageNumber="388" phylum="Ascomycota" rank="species" species="capistratus">
<emphasis box="[1268,1421,950,973]" italics="true" pageId="111" pageNumber="388">P. capistratus</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Flannery &amp; White" authorityYear="1991" box="[842,952,979,1002]" class="Dothideomycetes" family="Pteropodidae" genus="Pteropus" kingdom="Fungi" order="Chiroptera" pageId="111" pageNumber="388" phylum="Ascomycota" rank="species" species="ennisae">
<emphasis box="[842,952,979,1002]" italics="true" pageId="111" pageNumber="388">P. ennisae</emphasis>
</taxonomicName>
are best treated as sister species based on marked morphological differences combined with a level of gene sequence divergence more typical of interspecific differences.
</paragraph>
</subSubSection>
</treatment>
</document>