treatments-xml/data/03/DD/87/03DD8791F018897DFF0EFD81FE2BFF75.xml

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<mods:title id="782404933B08562DAA2EB36BF2A0308F">A new species of the genus Misgurnus (Cypriniformes, Cobitidae) from Ryukyu Islands, Japan</mods:title>
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<mods:namePart id="130AC906155BC944C0893D956F682EC7">Nakajima, Jun</mods:namePart>
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<mods:affiliation id="B554EF903E418A9FA401B37621BCAB15">Fukuoka Institute of Health and Environmental Sciences, Mukaizano 39, Dazaifu, Fukuoka, 811 - 0135 Japan. oikawamaru @ gmail. com; https: // orcid. org / 0000 - 0002 - 3542 - 9523</mods:affiliation>
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<mods:affiliation id="7B98A190B74653BE1DFACFCA4123C429">Osaka Medical and Pharmaceutical University, Daigakumachi 2 - 7, Takatsuki, Osaka, 569 - 0801 Japan. yasuyuki. hashiguchi @ ompu. ac. jp</mods:affiliation>
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<heading id="D08381EBF0188974FF0EFD81FDB9FD9D" bold="true" box="[151,533,547,573]" fontSize="11" level="1" pageId="4" pageNumber="529" reason="1">
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<taxonomicName id="4C744D04F0188974FF0EFD81FE1DFD9D" authority="Nakajima &amp; Hashiguchi, 2022" authorityName="Nakajima &amp; Hashiguchi" authorityYear="2022" box="[151,433,547,573]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="529" phylum="Chordata" rank="species" species="amamianus" status="sp. nov.">
<emphasis id="B900EA95F0188974FF0EFD81FE1DFD9D" bold="true" box="[151,433,547,573]" italics="true" pageId="4" pageNumber="529">Misgurnus amamianus</emphasis>
</taxonomicName>
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<paragraph id="8BCB3687F0188974FF0EFDE5FEDEFDC1" blockId="4.[151,533,547,609]" box="[151,370,582,609]" pageId="4" pageNumber="529">
(
<figureCitation id="134F2A02F0188974FF06FDE5FF42FDC1" box="[159,238,583,609]" captionStart="FIGURE 3" captionStartId="4.[151,250,1596,1620]" captionTargetBox="[151,1436,930,1572]" captionTargetId="figure-239@4.[151,1436,930,1572]" captionTargetPageId="4" captionText="FIGURE 3. Misgurnus amamianus sp. nov. a, holotype (male, KPM-NI 68404); b, paratype (female, TKPM-P 26178)." figureDoi="http://doi.org/10.5281/zenodo.6816838" httpUri="https://zenodo.org/record/6816838/files/figure.png" pageId="4" pageNumber="529">Figs. 3</figureCitation>
, 
<figureCitation id="134F2A02F0188974FF62FDE5FEA7FDC1" box="[251,267,583,609]" captionStart="FIGURE 4" captionStartId="5.[151,250,797,821]" captionTargetBox="[163,1436,364,773]" captionTargetId="figure-99@5.[151,1436,345,773]" captionTargetPageId="5" captionText="FIGURE 4. Misgurnus amamianus sp. nov., KPM-NI 68404, holotype. a, mouth; b, lamina circularis (allow indicating the neckline at the base of lamina circularis), bar 1 mm; c, dorsal fin." figureDoi="http://doi.org/10.5281/zenodo.6816840" httpUri="https://zenodo.org/record/6816840/files/figure.png" pageId="4" pageNumber="529">4</figureCitation>
, 
<figureCitation id="134F2A02F0188974FE8EFDE4FE8BFDC1" box="[279,295,582,609]" captionStart="FIGURE 5" captionStartId="5.[151,250,1865,1889]" captionTargetBox="[151,1436,881,1841]" captionTargetId="figure-136@5.[151,1436,881,1841]" captionTargetPageId="5" captionText="FIGURE 5. Misgurnus amamianus sp. nov., KPM-NI 68404, holotype. a, radiograph; b, radiograph of dorsal fin; c, radiograph of anal fin; d, radiograph of caudal fin; e, living body." figureDoi="http://doi.org/10.5281/zenodo.6816842" httpUri="https://zenodo.org/record/6816842/files/figure.png" pageId="4" pageNumber="529">5</figureCitation>
, 
<figureCitation id="134F2A02F0188974FEAAFDE5FEEFFDC1" box="[307,323,583,609]" captionStart="FIGURE 6" captionStartId="6.[151,250,1663,1687]" captionTargetBox="[151,1435,181,1637]" captionTargetId="figure-25@6.[151,1436,181,1639]" captionTargetPageId="6" captionText="FIGURE 6. Maximum likelihood tree of the partial sequences of mitochondrial DNA control region (CR) in Misgurnus amamianus sp. nov. and Japanese and Northeast Asian species of Misgurnus. CR sequences of Barbatula barbatula and Leptobotia elongata are used as outgroups. Numbers on each branch indicate bootstrap values with 1,000 replications." figureDoi="http://doi.org/10.5281/zenodo.6816844" httpUri="https://zenodo.org/record/6816844/files/figure.png" pageId="4" pageNumber="529">6</figureCitation>
, 
<figureCitation id="134F2A02F0188974FED6FDE5FEC6FDC1" box="[335,362,583,609]" captionStart="FIGURE 7" captionStartId="7.[151,250,972,996]" captionTargetBox="[151,1436,309,948]" captionTargetId="figure-95@7.[151,1436,309,948]" captionTargetPageId="7" captionText="FIGURE 7. Lamina circularis of Northeast Asian species of Misgurnus. a, M. amamianus sp. nov. (KPM-NI 68404, holotype, Tokuno-shima Is., Japan, 83.8 mm SL); b, M. anguillicaudatus (Japan clade) (KYUM-PI 4520, Fukuoka, Japan, 93.4 mm SL); c, M. anguillicaudatus (China clade) (KMNH VR 100186, Zheijiang, China, 128.9 mm SL); d, M. dabryanus (JNC130, Fukuoka, Japan, 115.2 mm SL); e, M. mohoity (JNC563, Ibaraki, Japan, 71.7 mm SL); f, M. nikolskyi (JNC301, Tokyo, Japan, 121.3 mm SL); g, M. sp. (Clade A) (JNC345, Aomori, Japan, 110.5 mm SL); e, M. sp. OK (KMNH VR100189, Okinawa, Japan, 77.1 mm SL). Bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.6816846" httpUri="https://zenodo.org/record/6816846/files/figure.png" pageId="4" pageNumber="529">7a</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36E650CF0188974FF0EFD2EFACAFCBE" pageId="4" pageNumber="529" type="reference_group">
<paragraph id="8BCB3687F0188974FF0EFD2EFEABFD62" blockId="4.[151,1437,651,798]" pageId="4" pageNumber="529">
<treatmentCitationGroup id="AB6411A9F0188974FF0EFD2EFEABFD62" pageId="4" pageNumber="529">
<taxonomicName id="4C744D04F0188974FF0EFD2EFDF3FD03" ID-CoL="43NK4" authority="(Cantor, 1842)" baseAuthorityName="Cantor" baseAuthorityYear="1842" box="[151,607,651,675]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="529" phylum="Chordata" rank="species" species="anguillicaudatus">
<emphasis id="B900EA95F0188974FF0EFD2EFE14FD03" box="[151,440,651,675]" italics="true" pageId="4" pageNumber="529">Misgurnus anguillicaudatus</emphasis>
(Cantor, 1842)
</taxonomicName>
: 
<treatmentCitation id="0AD51096F0188974FDF5FD29FCB1FD03" author="Ogawa, K." box="[620,797,651,675]" page="18" pageId="4" pageNumber="529" year="1937">
<bibRefCitation id="EFE54B76F0188974FDF5FD29FCB1FD03" author="Ogawa, K." box="[620,797,651,675]" pageId="4" pageNumber="529" pagination="1 - 27" refId="ref8429" refString="Ogawa, K. (1937) Regional distribution of freshwater fishes of Kagoshima Prefecture. Natural Science Association Magazine, Hiroshima University, 5, 1 - 27." type="journal article" year="1937">Ogawa 1937: 18</bibRefCitation>
</treatmentCitation>
, Fig. 15 (
<collectingCountry id="F3637617F0188974FC10FD29FC64FD03" box="[905,968,651,675]" name="Japan" pageId="4" pageNumber="529">Japan</collectingCountry>
: Tokuno-shima Is., Amami Isls., 
<collectingRegion id="49B0F865F0188974FAB0FD29FF51FD62" country="Japan" name="Kagoshima" pageId="4" pageNumber="529">Kagoshima Pref.</collectingRegion>
).
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</paragraph>
<paragraph id="8BCB3687F0188974FF0EFD6BFACAFCBE" blockId="4.[151,1437,651,798]" pageId="4" pageNumber="529">
<taxonomicName id="4C744D04F0188974FF0EFD6BFE8BFD40" box="[151,295,713,736]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="529" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B900EA95F0188974FF0EFD6BFEAFFD40" box="[151,259,713,736]" italics="true" pageId="4" pageNumber="529">Misgurnus</emphasis>
sp.
</taxonomicName>
IR (
<emphasis id="B900EA95F0188974FECAFD6BFE27FD40" box="[339,395,713,736]" italics="true" pageId="4" pageNumber="529">sensu</emphasis>
Shmizu et al. 2011): 
<bibRefCitation id="EFE54B76F0188974FDF8FD6AFCF4FD41" author="Shimizu, T. &amp; Suzuki, T. &amp; Takagi, M. &amp; Oseko, N." box="[609,856,712,737]" pageId="4" pageNumber="529" pagination="141 - 153" refId="ref8808" refString="Shimizu, T., Suzuki, T., Takagi, M. &amp; Oseko, N. (2011) Morphological and genetic characteristics of misgurnid loach from Okinawa and Iriomote Islands. Bulletin of the Biogeographical Society of Japan, 66, 141 - 153. [in Japanese with English abstract]" type="journal article" year="2011">Shimizu et al. 2011: 144</bibRefCitation>
, 
<figureCitation id="134F2A02F0188974FCFBFD6AFC30FD41" box="[866,924,712,737]" captionStart="FIGURE 1" captionStartId="1.[151,250,1699,1723]" captionTargetBox="[151,1434,708,1673]" captionTargetId="figure-286@1.[151,1436,702,1673]" captionTargetPageId="1" captionText="FIGURE 1. Map showing the collection sites of the type series of Misgurnus amamianus sp. nov. A double circle indicates the locality of holotype. Double and closed circles indicate native populations, and open circle indicate an introduced population." figureDoi="http://doi.org/10.5281/zenodo.6816834" httpUri="https://zenodo.org/record/6816834/files/figure.png" pageId="4" pageNumber="529">Fig. 1</figureCitation>
(OMNH-P20968, 
<collectingCountry id="F3637617F0188974FBC0FD6BFB3BFD40" box="[1113,1175,713,736]" name="Japan" pageId="4" pageNumber="529">Japan</collectingCountry>
: Iriomote-jima Is., Yaeyama Isls., 
<collectingRegion id="49B0F865F0188974FEB8FD45FE14FD5F" box="[289,440,743,767]" country="Japan" name="Okinawa" pageId="4" pageNumber="529">Okinawa Pref.</collectingRegion>
); 
<bibRefCitation id="EFE54B76F0188974FE51FD45FCBFFD5F" author="Nakajima, J. &amp; Uchiyama, R." box="[456,787,743,767]" pageId="4" pageNumber="529" refId="ref8396" refString="Nakajima, J. &amp; Uchiyama, R. (2017) Loaches of Japan - Natural History and Culture. Yama-Kei Publishers, Tokyo, 223 pp. [in Japanese]" type="book" year="2017">Nakajima &amp; Uchiyama 2017: 72</bibRefCitation>
, 74 (
<collectingCountry id="F3637617F0188974FCDAFD45FC2EFD5F" box="[835,898,743,767]" name="Japan" pageId="4" pageNumber="529">Japan</collectingCountry>
: Amami Isls., 
<collectingRegion id="49B0F865F0188974FB88FD45FB13FD5F" box="[1041,1215,743,767]" country="Japan" name="Kagoshima" pageId="4" pageNumber="529">Kagoshima Pref.</collectingRegion>
), 75 (
<collectingCountry id="F3637617F0188974FB6CFD45FA98FD5F" box="[1269,1332,743,767]" name="Japan" pageId="4" pageNumber="529">Japan</collectingCountry>
: Iriomotejima Is., Yaeyama Isls., 
<collectingRegion id="49B0F865F0188974FE20FCA4FDFCFCBE" box="[441,592,774,798]" country="Japan" name="Okinawa" pageId="4" pageNumber="529">Okinawa Pref.</collectingRegion>
); 
<bibRefCitation id="EFE54B76F0188974FDF8FCA4FC82FCBE" author="Nakajima, J." box="[609,814,774,798]" pageId="4" pageNumber="529" pagination="110 - 115" refId="ref8280" refString="Nakajima, J. (2018) Cobitidae. In: Nakabo, T. (Ed.), The Natural History of the Fishes of Japan. Shogakukan, Tokyo, pp. 110 - 115. [in Japanese]" type="book chapter" year="2018">Nakajima 2018: 111</bibRefCitation>
(no data); 
<bibRefCitation id="EFE54B76F0188974FC05FCA4FBFAFCBE" author="Hosoya, K." box="[924,1110,774,798]" pageId="4" pageNumber="529" refId="ref7344" refString="Hosoya, K. (2019) Zouho-kaitei Nihon-No-Tansuigyo [Freshwater fishes of Japan Revised Edition]. Yama-Kei Publishers, Tokyo, 559 pp. [in Japanese]" type="book" year="2019">Hosoya 2019: 180</bibRefCitation>
(
<collectingCountry id="F3637617F0188974FBFAFCA4FB0EFCBE" box="[1123,1186,774,798]" name="Japan" pageId="4" pageNumber="529">Japan</collectingCountry>
: 
<collectingRegion id="49B0F865F0188974FB35FCA4FAF7FCBE" box="[1196,1371,774,798]" country="Japan" name="Kagoshima" pageId="4" pageNumber="529">Kagoshima Pref.</collectingRegion>
).
</paragraph>
</subSubSection>
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<paragraph id="8BCB3687F0188974FF0EFCEEFE1EFC2A" blockId="4.[151,1436,844,907]" pageId="4" pageNumber="529">
<materialsCitation id="3B1C3CDAF0188974FF0EFCEEFE1EFC2A" ID-GBIF-Occurrence="3861475315" collectingDate="2015-12-12" collectorName="Tokuno-shima Is. &amp; T. Kitano" country="Japan" location="Isen" pageId="4" pageNumber="529" specimenCount="1" stateProvince="Kagoshima" typeStatus="holotype">
<emphasis id="B900EA95F0188974FF0EFCEEFEA7FCC6" bold="true" box="[151,267,844,870]" pageId="4" pageNumber="529">
<typeStatus id="54CF8825F0188974FF0EFCEEFEAAFCC6" box="[151,262,844,870]" pageId="4" pageNumber="529" type="holotype">Holotype</typeStatus>
.
</emphasis>
KPM-NI 68404, male, 
<quantity id="4C8C9B62F0188974FD80FCEEFDD2FCC7" box="[537,638,844,871]" metricMagnitude="-2" metricUnit="m" metricValue="8.379999999999999" pageId="4" pageNumber="529" unit="mm" value="83.8">83.8 mm</quantity>
SL, 
<collectingCountry id="F3637617F0188974FD2CFCEFFD55FCC7" box="[693,761,845,871]" name="Japan" pageId="4" pageNumber="529">Japan</collectingCountry>
: 
<location id="8EAB605CF0188974FC9CFCEFFC9AFCC7" LSID="urn:lsid:plazi:treatment:03DD8791F018897DFF0EFD81FE2BFF75:8EAB605CF0188974FC9CFCEFFC9AFCC7" box="[773,822,845,871]" country="Japan" name="Isen" pageId="4" pageNumber="529" stateProvince="Kagoshima">Isen</location>
, 
<collectorName id="26815351F0188974FCDBFCEEFBA4FCC7" box="[834,1032,844,871]" pageId="4" pageNumber="529">Tokuno-shima Is.</collectorName>
, Amami Isls., 
<collectingRegion id="49B0F865F0188974FB34FCEEFAC2FCC6" box="[1197,1390,844,870]" country="Japan" name="Kagoshima" pageId="4" pageNumber="529">Kagoshima Pref.</collectingRegion>
, 
<date id="FFCA1047F0188974FAE0FCEEFEA6FC2B" pageId="4" pageNumber="529" value="2015-12-12">
<collectingDate id="EF8EE9AFF0188974FAE0FCEEFEA6FC2B" pageId="4" pageNumber="529" value="2015-12-12">12. XII. 2015</collectingDate>
</date>
., 
<collectorName id="26815351F0188974FE80FCD3FE2EFC2A" box="[281,386,880,907]" pageId="4" pageNumber="529">T. Kitano</collectorName>
leg.
</materialsCitation>
</paragraph>
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<paragraph id="8BCB3687F0188974FF0EF99EFAFCF9F4" blockId="4.[151,1360,1596,1621]" box="[151,1360,1596,1621]" pageId="4" pageNumber="529">
<emphasis id="B900EA95F0188974FF0EF99EFEBFF9F4" bold="true" box="[151,275,1596,1620]" pageId="4" pageNumber="529">FIGURE 3.</emphasis>
<taxonomicName id="4C744D04F0188974FE80F99FFDADF9F4" authority="Nakajima &amp; Hashiguchi, 2022" authorityName="Nakajima &amp; Hashiguchi" authorityYear="2022" box="[281,513,1597,1620]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="529" phylum="Chordata" rank="species" species="amamianus" status="sp. nov.">
<emphasis id="B900EA95F0188974FE80F99FFDADF9F4" box="[281,513,1597,1620]" italics="true" pageId="4" pageNumber="529">Misgurnus amamianus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23357EEF0188974FD9EF99FFDFAF9F4" box="[519,598,1597,1620]" pageId="4" pageNumber="529" rank="species">sp. nov.</taxonomicNameLabel>
a, holotype (male, KPM-NI 68404); b, paratype (female, TKPM-P 26178).
</paragraph>
</caption>
<paragraph id="8BCB3687F0188975FF54F921FBA9FEE1" blockId="4.[151,1437,1666,2017]" lastBlockId="5.[151,1436,151,321]" lastPageId="5" lastPageNumber="530" pageId="4" pageNumber="529">
<materialsCitation id="3B1C3CDAF0188974FF54F921FCCAF961" ID-GBIF-Occurrence="3861475304" collectingDate="2015-12-12" collectionCode="JNC" collectorName="Tokuno-shima Is. &amp; T. Kitano" country="Japan" location="Ryukyu Islands" pageId="4" pageNumber="529" specimenCode="JNC353, 354" specimenCount="26" specimenCount-female="10" specimenCount-male="16" stateProvince="Kagoshima" typeStatus="paratype">
<emphasis id="B900EA95F0188974FF54F921FEE0F93D" bold="true" box="[205,332,1667,1693]" pageId="4" pageNumber="529">
<typeStatus id="54CF8825F0188974FF54F921FEEBF93D" box="[205,327,1667,1693]" pageId="4" pageNumber="529" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="9D72FD0EF0188974FECBF920FE1AF93C" box="[338,438,1666,1692]" pageId="4" pageNumber="529" type="male">16 males</specimenCount>
and 
<specimenCount id="9D72FD0EF0188974FE72F921FDCBF93C" box="[491,615,1666,1693]" pageId="4" pageNumber="529" type="female">10 females</specimenCount>
, 
<quantity id="4C8C9B62F0188974FDE8F920FC8EF93D" box="[625,802,1666,1693]" metricMagnitude="-2" metricUnit="m" metricValue="9.07" metricValueMax="12.629999999999999" metricValueMin="5.51" pageId="4" pageNumber="529" unit="mm" value="90.7" valueMax="126.3" valueMin="55.1">55.1–126.3 mm</quantity>
SL, all from the 
<location id="8EAB605CF0188974FC45F920FB20F93C" LSID="urn:lsid:plazi:treatment:03DD8791F018897DFF0EFD81FE2BFF75:8EAB605CF0188974FC45F920FB20F93C" box="[988,1164,1666,1692]" country="Japan" name="Ryukyu Islands" pageId="4" pageNumber="529">Ryukyu Islands</location>
, 
<collectingCountry id="F3637617F0188974FB0EF921FB76F93D" box="[1175,1242,1667,1693]" name="Japan" pageId="4" pageNumber="529">Japan</collectingCountry>
. TKPM-P 26178, 26179, male &amp; female, same data as the holotype; 
<specimenCode id="DBD29EFCF0188974FD56F904FC82F961" box="[719,814,1702,1729]" collectionCode="JNC" pageId="4" pageNumber="529">JNC353</specimenCode>
, 
<specimenCode id="DBD29EFCF0188974FCA0F904FCCAF961" box="[825,870,1702,1729]" collectionCode="JNC" pageId="4" pageNumber="529">354</specimenCode>
</materialsCitation>
, 
<materialsCitation id="3B1C3CDAF0188974FCE8F904FD48F944" ID-GBIF-Occurrence="3861475301" accessionNumber="VR 100463" collectingDate="2015-12-12" collectionCode="KMNH" collectorName="J. Nakajima" country="Japan" location="Isen" pageId="4" pageNumber="529" specimenCount="1" stateProvince="Kagoshima" typeStatus="paratype">
male &amp; female, same locality as the holotype, 29. VII. 29. 2016., 
<collectorName id="26815351F0188974FED8F969FE6BF944" box="[321,455,1738,1765]" pageId="4" pageNumber="529">J. Nakajima</collectorName>
leg.; 
<collectionCode id="ED65AE42F0188974FD9DF969FDF6F945" box="[516,602,1739,1765]" pageId="4" pageNumber="529">KMNH</collectionCode>
VR 100463
</materialsCitation>
, 
<materialsCitation id="3B1C3CDAF0188974FD76F968FD2DF8A9" ID-GBIF-Occurrence="3861475314" collectingDate="2015-08-20" collectionCode="JNC" collectorName="Tokuno-shima Is. &amp; Y. Kano" country="Japan" county="Noda" location="Amami Isls." municipality="Isen" pageId="4" pageNumber="529" specimenCode="JNC554" specimenCount="1" stateProvince="Kagoshima" typeStatus="paratype">
male, 
<collectingCounty id="62AA4E0BF0188974FCABF968FCDDF944" box="[818,881,1738,1764]" pageId="4" pageNumber="529">Noda</collectingCounty>
, 
<collectingMunicipality id="6BAFACFDF0188974FCE5F969FC01F945" box="[892,941,1739,1765]" pageId="4" pageNumber="529">Isen</collectingMunicipality>
, 
<collectorName id="26815351F0188974FC21F968FBD0F945" box="[952,1148,1738,1765]" pageId="4" pageNumber="529">Tokuno-shima Is.</collectorName>
, 
<location id="8EAB605CF0188974FB1EF968FABEF944" LSID="urn:lsid:plazi:treatment:03DD8791F018897DFF0EFD81FE2BFF75:8EAB605CF0188974FB1EF968FABEF944" box="[1159,1298,1738,1764]" country="Japan" county="Noda" municipality="Isen" name="Amami Isls." pageId="4" pageNumber="529" stateProvince="Kagoshima">Amami Isls.</location>
, 
<collectingRegion id="49B0F865F0188974FA84F968FF7CF8A8" country="Japan" name="Kagoshima" pageId="4" pageNumber="529">Kagoshima Pref.</collectingRegion>
, 
<date id="FFCA1047F0188974FF43F94DFED2F8A9" box="[218,382,1774,1801]" pageId="4" pageNumber="529" value="2015-08-20">
<collectingDate id="EF8EE9AFF0188974FF43F94DFED2F8A9" box="[218,382,1774,1801]" pageId="4" pageNumber="529" value="2015-08-20">20. VIII. 2015</collectingDate>
</date>
., 
<collectorName id="26815351F0188974FE12F94DFE4AF8A9" box="[395,486,1775,1801]" pageId="4" pageNumber="529">Y. Kano</collectorName>
leg.; 
<specimenCode id="DBD29EFCF0188974FDBAF94CFD2DF8A9" box="[547,641,1774,1801]" collectionCode="JNC" pageId="4" pageNumber="529">JNC554</specimenCode>
</materialsCitation>
, 
<materialsCitation id="3B1C3CDAF0188974FD15F94CFE66F8F0" ID-GBIF-Occurrence="3861475305" accessionNumber="VR 100188" collectingDate="1958-08-03" collectingDateMax="2019-11-10" collectingDateMin="1958-08-03" collectionCode="KMNH" collectorName="S. Ueno &amp; Y. Morimoto" country="China" county="Okinoerabu-jima Is." location="Amami Isls." municipality="Okinoerabu-jima Is." pageId="4" pageNumber="529" specimenCount="1" stateProvince="Kagoshima" typeStatus="paratype">
female, same locality, 
<date id="FFCA1047F0188974FC1DF94DFBBAF8A9" box="[900,1046,1775,1801]" pageId="4" pageNumber="529" value="2019-11-10">
<collectingDate id="EF8EE9AFF0188974FC1DF94DFBBAF8A9" box="[900,1046,1775,1801]" pageId="4" pageNumber="529" value="2019-11-10">10. XI. 2019</collectingDate>
</date>
., 
<collectorName id="26815351F0188974FBBDF94DFB05F8A8" box="[1060,1193,1774,1801]" pageId="4" pageNumber="529">J. Nakajima</collectorName>
leg.; NSMT-P 34733, male, 
<collectorName id="26815351F0188974FF45F8B0FE90F88C" box="[220,316,1810,1836]" pageId="4" pageNumber="529">Otsukan</collectorName>
, 
<collectingCountry id="F3637617F0188974FEDEF8B0FE21F88C" box="[327,397,1810,1836]" name="China" pageId="4" pageNumber="529">China</collectingCountry>
, 
<collectorName id="26815351F0188974FE01F8B0FDD7F88D" box="[408,635,1810,1837]" pageId="4" pageNumber="529">Okinoerabu-jima Is.</collectorName>
, 
<collectorName id="26815351F0188974FD1FF8B0FCBEF88C" box="[646,786,1810,1836]" pageId="4" pageNumber="529">Amami Isls.</collectorName>
, 
<collectorName id="26815351F0188974FC84F8B0FC71F88C" box="[797,989,1810,1836]" pageId="4" pageNumber="529">Kagoshima Pref.</collectorName>
, 
<date id="FFCA1047F0188974FC71F8B1FB2CF88D" box="[1000,1152,1810,1837]" pageId="4" pageNumber="529" value="1958-08-03">
<collectingDate id="EF8EE9AFF0188974FC71F8B1FB2CF88D" box="[1000,1152,1810,1837]" pageId="4" pageNumber="529" value="1958-08-03">3. VIII. 1958</collectingDate>
</date>
., 
<collectorName id="26815351F0188974FB16F8B1FB45F88D" box="[1167,1257,1811,1837]" pageId="4" pageNumber="529">S. Ueno</collectorName>
&amp; 
<collectorName id="26815351F0188974FA95F8B1FA31F88C" box="[1292,1437,1810,1837]" pageId="4" pageNumber="529">Y. Morimoto</collectorName>
leg.; OMNH-P 1145, 1146,
</materialsCitation>
<materialsCitation id="3B1C3CDAF0188974FE4AF894FF6FF839" ID-GBIF-Occurrence="3861475303" accessionNumber="VR 100188" collectingDate="1958-08-03" collectingDateMax="2019-11-10" collectingDateMin="1958-08-03" collectionCode="KMNH" collectorName="Y. Shibata" country="China" county="Okinoerabu-jima Is." location="Amami Isls." municipality="Okinoerabu-jima Is." pageId="4" pageNumber="529" specimenCode="JNC140, 141" specimenCount="1" stateProvince="Kagoshima" typeStatus="paratype">
male &amp; female, 
<collectorName id="26815351F0188974FD15F894FC98F8F0" box="[652,820,1846,1872]" pageId="4" pageNumber="529">Kamihirakawa</collectorName>
, 
<collectingCountry id="F3637617F0188974FCD9F894FC2AF8F0" box="[832,902,1846,1872]" name="China" pageId="4" pageNumber="529">China</collectingCountry>
, 
<collectingMunicipality id="6BAFACFDF0188974FC0BF894FBDBF8F1" box="[914,1143,1846,1873]" pageId="4" pageNumber="529">
<collectingCounty id="62AA4E0BF0188974FC0BF894FBDBF8F1" box="[914,1143,1846,1873]" pageId="4" pageNumber="529">Okinoerabu-jima Is.</collectingCounty>
</collectingMunicipality>
, 
<location id="8EAB605CF0188974FB1AF894FABCF8F0" LSID="urn:lsid:plazi:treatment:03DD8791F018897DFF0EFD81FE2BFF75:8EAB605CF0188974FB1AF894FABCF8F0" box="[1155,1296,1846,1872]" country="China" county="Okinoerabu-jima Is." municipality="Okinoerabu-jima Is." name="Amami Isls." pageId="4" pageNumber="529">Amami Isls.</location>
, 
<location id="8EAB605CF0188974FA84F894FF7CF8D4" LSID="urn:lsid:plazi:treatment:03DD8791F018897DFF0EFD81FE2BFF75:8EAB605CF0188974FA84F894FF7CF8D4" country="China" county="Okinoerabu-jima Is." municipality="Okinoerabu-jima Is." name="Kagoshima Pref." pageId="4" pageNumber="529">Kagoshima Pref.</location>
, 
<date id="FFCA1047F0188974FF40F8F9FED0F8D4" box="[217,380,1882,1909]" pageId="4" pageNumber="529" value="1964-08-29">
<collectingDate id="EF8EE9AFF0188974FF40F8F9FED0F8D4" box="[217,380,1882,1909]" pageId="4" pageNumber="529" value="1964-08-29">29. VIII. 1964</collectingDate>
</date>
., 
<collectorName id="26815351F0188974FE11F8F9FE55F8D4" box="[392,505,1882,1909]" pageId="4" pageNumber="529">Y. Shibata</collectorName>
leg.; KPM-NI 68405, 68406, 
<collectionCode id="ED65AE42F0188974FCDAF8F9FC35F8D5" box="[835,921,1883,1909]" pageId="4" pageNumber="529">KMNH</collectionCode>
VR 100188, TKPM-P 26180, 26181, 
<specimenCode id="DBD29EFCF0188974FAA0F8F9FA34F8D5" box="[1337,1432,1883,1909]" collectionCode="JNC" pageId="4" pageNumber="529">JNC140</specimenCode>
, 
<specimenCode id="DBD29EFCF0188974FF0EF8DDFF6FF839" box="[151,195,1919,1945]" collectionCode="JNC" pageId="4" pageNumber="529">141</specimenCode>
</materialsCitation>
, 
<materialsCitation id="3B1C3CDAF0188974FF48F8DCFD29F81D" ID-GBIF-Occurrence="3861475311" collectingDate="1970-08-07" collectingDateMax="2013-11-17" collectingDateMin="1970-08-07" collectorName="J. Nakajima" country="China" county="Kamihirakawa" location="Uken" municipality="Amuro" pageId="4" pageNumber="529" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D72FD0EF0188974FF48F8DCFEE5F838" box="[209,329,1918,1944]" pageId="4" pageNumber="529" type="male">four males</specimenCount>
&amp; 
<specimenCount id="9D72FD0EF0188974FEE8F8DCFDA0F838" box="[369,524,1918,1944]" pageId="4" pageNumber="529" type="female">three females</specimenCount>
, 
<collectingCounty id="62AA4E0BF0188974FD81F8DCFD6CF838" box="[536,704,1918,1944]" pageId="4" pageNumber="529">Kamihirakawa</collectingCounty>
, 
<collectingCountry id="F3637617F0188974FD54F8DCFCBFF838" box="[717,787,1918,1944]" name="China" pageId="4" pageNumber="529">China</collectingCountry>
, 
<collectorName id="26815351F0188974FCB9F8DCFBA8F839" box="[800,1028,1918,1945]" pageId="4" pageNumber="529">Okinoerabu-jima Is.</collectorName>
, 
<collectorName id="26815351F0188974FB88F8DCFB32F838" box="[1041,1182,1918,1944]" pageId="4" pageNumber="529">Amami Isls.</collectorName>
, 
<collectorName id="26815351F0188974FB35F8DCFAC1F838" box="[1196,1389,1918,1944]" pageId="4" pageNumber="529">Kagoshima Pref.</collectorName>
, 
<date id="FFCA1047F0188974FAE0F8DDFEAEF81D" pageId="4" pageNumber="529" value="2013-11-17">
<collectingDate id="EF8EE9AFF0188974FAE0F8DDFEAEF81D" pageId="4" pageNumber="529" value="2013-11-17">17. XI. 2013</collectingDate>
</date>
., 
<collectorName id="26815351F0188974FE8BF801FE34F81C" box="[274,408,1954,1981]" pageId="4" pageNumber="529">J. Nakajima</collectorName>
leg.; YCM-P 31002,
</materialsCitation>
<materialsCitation id="3B1C3CDAF0188974FD14F800FCAFF840" ID-GBIF-Occurrence="3861475312" collectingDate="1970-08-07" collectingDateMax="2013-11-17" collectingDateMin="1970-08-07" collectorName="H. Ikeda" country="China" county="Kamihirakawa" location="Uken" municipality="Amuro" pageId="4" pageNumber="529" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
male, Suitengu-no-ike, 
<collectorName id="26815351F0188974FC0CF800FB9FF81D" box="[917,1075,1954,1981]" pageId="4" pageNumber="529">Kikai-jima Is.</collectorName>
, 
<collectorName id="26815351F0188974FBA6F800FB67F81C" box="[1087,1227,1954,1980]" pageId="4" pageNumber="529">Amami Isls.</collectorName>
, 
<collectorName id="26815351F0188974FB41F800FA35F81C" box="[1240,1433,1954,1980]" pageId="4" pageNumber="529">Kagoshima Pref.</collectorName>
, data unknown (1930s)., 
<collectorName id="26815351F0188974FE30F865FDA7F840" box="[425,523,1990,2017]" pageId="4" pageNumber="529">H. Ikeda</collectorName>
leg.; NSMT-P 14268,
</materialsCitation>
<materialsCitation id="3B1C3CDAF0188975FC95F864FC41FF11" ID-GBIF-Occurrence="3861475302" collectingDate="1970-08-07" collectingDateMax="2013-11-17" collectingDateMin="1970-08-07" country="China" county="Kamihirakawa" lastPageId="5" lastPageNumber="530" location="Uken" municipality="Amuro" pageId="4" pageNumber="529" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
male &amp; female, 
<collectingMunicipality id="6BAFACFDF0188974FC58F865FBBFF841" box="[961,1043,1991,2017]" pageId="4" pageNumber="529">Amuro</collectingMunicipality>
, 
<location id="8EAB605CF0188974FB86F864FBF3F840" LSID="urn:lsid:plazi:treatment:03DD8791F018897DFF0EFD81FE2BFF75:8EAB605CF0188974FB86F864FBF3F840" box="[1055,1119,1990,2016]" country="China" county="Kamihirakawa" municipality="Amuro" name="Uken" pageId="4" pageNumber="529">Uken</location>
, 
<location id="8EAB605CF0188974FBF0F864FA90F841" LSID="urn:lsid:plazi:treatment:03DD8791F018897DFF0EFD81FE2BFF75:8EAB605CF0188974FBF0F864FA90F841" box="[1129,1340,1990,2017]" country="China" county="Kamihirakawa" municipality="Amuro" name="Amami-oshima Is." pageId="4" pageNumber="529">Amami-oshima Is.</location>
, 
<location id="8EAB605CF0188975FAD1F864FF6BFF11" LSID="urn:lsid:plazi:treatment:03DD8791F018897DFF0EFD81FE2BFF75:8EAB605CF0188975FAD1F864FF6BFF11" country="China" county="Kamihirakawa" lastPageId="5" lastPageNumber="530" municipality="Amuro" name="Amami Isls." pageId="4" pageNumber="529">Amami Isls.</location>
, 
<location id="8EAB605CF0198975FF4FFF35FE3BFF11" LSID="urn:lsid:plazi:treatment:03DD8791F018897DFF0EFD81FE2BFF75:8EAB605CF0198975FF4FFF35FE3BFF11" box="[214,407,151,177]" country="China" county="Kamihirakawa" municipality="Amuro" name="Kagoshima Pref." pageId="5" pageNumber="530">Kagoshima Pref.</location>
, 
<date id="FFCA1047F0198975FE3CFF35FDEDFF11" box="[421,577,151,177]" pageId="5" pageNumber="530" value="1970-08-07">
<collectingDate id="EF8EE9AFF0198975FE3CFF35FDEDFF11" box="[421,577,151,177]" pageId="5" pageNumber="530" value="1970-08-07">7. VIII. 1970</collectingDate>
</date>
., collector unknown; NSMT-P 28933
</materialsCitation>
, 
<materialsCitation id="3B1C3CDAF0198975FC63FF35FA90FF75" ID-GBIF-Occurrence="3861475307" collectingDate="1975-04-04" collectionCode="JNC" collectorName="H. Sakai &amp; M. Sato" country="China" county="Nishi-nakama" location="Kagoshima Pref." municipality="Amami Isls." pageId="5" pageNumber="530" specimenCode="JNC128" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D72FD0EF0198975FC63FF35FBC3FF11" box="[1018,1135,151,177]" pageId="5" pageNumber="530" type="male">two males</specimenCount>
&amp; 
<specimenCount id="9D72FD0EF0198975FB01FF35FA89FF11" box="[1176,1317,151,177]" pageId="5" pageNumber="530" type="female">two females</specimenCount>
, 
<collectingCounty id="62AA4E0BF0198975FAABFF35FF7BFF75" pageId="5" pageNumber="530">Nishi-nakama</collectingCounty>
, 
<collectorName id="26815351F0198975FF78FF19FE18FF75" box="[225,436,187,213]" pageId="5" pageNumber="530">Amami-oshima Is.</collectorName>
, 
<collectingMunicipality id="6BAFACFDF0198975FE58FF19FDE1FF75" box="[449,589,187,213]" pageId="5" pageNumber="530">Amami Isls.</collectingMunicipality>
, 
<location id="8EAB605CF0198975FDC3FF19FCB7FF75" LSID="urn:lsid:plazi:treatment:03DD8791F018897DFF0EFD81FE2BFF75:8EAB605CF0198975FDC3FF19FCB7FF75" box="[602,795,187,213]" country="China" county="Nishi-nakama" municipality="Amami Isls." name="Kagoshima Pref." pageId="5" pageNumber="530">Kagoshima Pref.</location>
, 
<date id="FFCA1047F0198975FCBFFF19FC07FF75" box="[806,939,186,213]" pageId="5" pageNumber="530" value="1975-04-04">
<collectingDate id="EF8EE9AFF0198975FCBFFF19FC07FF75" box="[806,939,186,213]" pageId="5" pageNumber="530" value="1975-04-04">4. IV. 1975</collectingDate>
</date>
., 
<collectorName id="26815351F0198975FC22FF19FBB0FF75" box="[955,1052,187,213]" pageId="5" pageNumber="530">H. Sakai</collectorName>
&amp; 
<collectorName id="26815351F0198975FBDAFF19FB31FF75" box="[1091,1181,187,213]" pageId="5" pageNumber="530">M. Sato</collectorName>
leg.; 
<specimenCode id="DBD29EFCF0198975FB47FF19FA90FF75" box="[1246,1340,187,213]" collectionCode="JNC" pageId="5" pageNumber="530">JNC128</specimenCode>
</materialsCitation>
, 
<materialsCitation id="3B1C3CDAF0198975FAD0FF19FAFAFF59" ID-GBIF-Occurrence="3861475309" collectingDate="2007-10-26" collectingDateMax="2011-09-08" collectingDateMin="2007-10-26" collectionCode="JNC" collectorName="H. Mizutani" country="China" county="Nishi-nakama" location="Okinawa Pref." municipality="Yaeyama Isls." pageId="5" pageNumber="530" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D72FD0EF0198975FAD0FF19FA34FF75" box="[1353,1432,187,213]" pageId="5" pageNumber="530" type="male">1 male</specimenCount>
, 
<collectorName id="26815351F0198975FF0EFF7DFF58FF59" box="[151,244,223,249]" pageId="5" pageNumber="530">Urauchi</collectorName>
, 
<collectorName id="26815351F0198975FE98FF7DFE6DFF59" box="[257,449,223,249]" pageId="5" pageNumber="530">Iriomote-jima Is.</collectorName>
, 
<collectorName id="26815351F0198975FE57FF7DFDC1FF59" box="[462,621,223,249]" pageId="5" pageNumber="530">Yaeyama Isls.</collectorName>
, 
<collectorName id="26815351F0198975FDE3FF7DFC8DFF59" box="[634,801,223,249]" pageId="5" pageNumber="530">Okinawa Pref.</collectorName>
, 
<date id="FFCA1047F0198975FCB4FF7DFC15FF59" box="[813,953,223,249]" pageId="5" pageNumber="530" value="2007-10-26">
<collectingDate id="EF8EE9AFF0198975FCB4FF7DFC15FF59" box="[813,953,223,249]" pageId="5" pageNumber="530" value="2007-10-26">26. X. 2007</collectingDate>
</date>
., 
<collectorName id="26815351F0198975FC53FF7DFBFEFF59" box="[970,1106,223,249]" pageId="5" pageNumber="530">H. Mizutani</collectorName>
leg.; OMNH-P 37946,
</materialsCitation>
<materialsCitation id="3B1C3CDAF0198975FAC6FF7DFBD0FEBD" ID-GBIF-Occurrence="3861475306" collectingDate="2007-10-26" collectingDateMax="2011-09-08" collectingDateMin="2007-10-26" collectionCode="JNC" collectorName="Urauchi &amp; Iriomote-jima Is. &amp; Yaeyama Isls. &amp; Okinawa Pref. &amp; H. Mizutani &amp; T. Suzuki &amp; M. Hosokawa &amp; M. Baba &amp; Kumotta &amp; J. Nakajima" country="China" county="Nishi-nakama" location="Okinawa Pref." municipality="Yaeyama Isls." pageId="5" pageNumber="530" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
male, same locality, 
<date id="FFCA1047F0198975FEA3FEA1FE6BFEBD" box="[314,455,259,285]" pageId="5" pageNumber="530" value="2008-10-24">
<collectingDate id="EF8EE9AFF0198975FEA3FEA1FE6BFEBD" box="[314,455,259,285]" pageId="5" pageNumber="530" value="2008-10-24">24. X. 2008</collectingDate>
</date>
., 
<collectorName id="26815351F0198975FE4FFEA1FDEAFEBD" box="[470,582,259,285]" pageId="5" pageNumber="530">T. Suzuki</collectorName>
, 
<collectorName id="26815351F0198975FDC8FEA1FD43FEBD" box="[593,751,259,285]" pageId="5" pageNumber="530">M. Hosokawa</collectorName>
&amp; 
<collectorName id="26815351F0198975FC8FFEA1FCD4FEBD" box="[790,888,259,285]" pageId="5" pageNumber="530">M. Baba</collectorName>
leg.; WMNH-PI 0311,
</materialsCitation>
<materialsCitation id="3B1C3CDAF0198975FB1DFEA1FBA9FEE1" ID-GBIF-Occurrence="3861475310" collectingDate="2007-10-26" collectingDateMax="2011-09-08" collectingDateMin="2007-10-26" collectionCode="JNC" collectorName="J. Nakajima" country="China" county="Nishi-nakama" location="Okinawa Pref." municipality="Yaeyama Isls." pageId="5" pageNumber="530" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
male, 
<collectorName id="26815351F0198975FB53FEA1FA83FEBD" box="[1226,1327,259,285]" pageId="5" pageNumber="530">Kumotta</collectorName>
, 
<collectorName id="26815351F0198975FAA2FEA1FA34FEBD" box="[1339,1432,259,285]" pageId="5" pageNumber="530">Urauchi</collectorName>
, 
<collectorName id="26815351F0198975FF0EFE85FEFAFEE1" box="[151,342,295,321]" pageId="5" pageNumber="530">Iriomote-jima Is.</collectorName>
, 
<collectingMunicipality id="6BAFACFDF0198975FEFBFE85FDACFEE1" box="[354,512,295,321]" pageId="5" pageNumber="530">Yaeyama Isls.</collectingMunicipality>
, 
<location id="8EAB605CF0198975FD95FE85FD1DFEE1" LSID="urn:lsid:plazi:treatment:03DD8791F018897DFF0EFD81FE2BFF75:8EAB605CF0198975FD95FE85FD1DFEE1" box="[524,689,295,321]" municipality="Yaeyama Isls." name="Okinawa Pref." pageId="5" pageNumber="530">Okinawa Pref.</location>
, 
<date id="FFCA1047F0198975FD22FE85FCECFEE1" box="[699,832,295,321]" pageId="5" pageNumber="530" value="2011-09-08">
<collectingDate id="EF8EE9AFF0198975FD22FE85FCECFEE1" box="[699,832,295,321]" pageId="5" pageNumber="530" value="2011-09-08">8. IX. 2011</collectingDate>
</date>
., 
<collectorName id="26815351F0198975FCD6FE85FC79FEE1" box="[847,981,295,321]" pageId="5" pageNumber="530">J. Nakajima</collectorName>
leg.
</materialsCitation>
</paragraph>
<caption id="DF0B660FF0198975FF0EFCBFFC81FCF9" ID-DOI="http://doi.org/10.5281/zenodo.6816840" ID-Zenodo-Dep="6816840" httpUri="https://zenodo.org/record/6816840/files/figure.png" pageId="5" pageNumber="530" startId="5.[151,250,797,821]" targetBox="[163,1436,364,773]" targetPageId="5">
<paragraph id="8BCB3687F0198975FF0EFCBFFC81FCF9" blockId="5.[151,1436,797,857]" pageId="5" pageNumber="530">
<emphasis id="B900EA95F0198975FF0EFCBFFEB9FC95" bold="true" box="[151,277,797,821]" pageId="5" pageNumber="530">FIGURE 4.</emphasis>
<taxonomicName id="4C744D04F0198975FE87FCBCFDA5FC95" authority="Nakajima &amp; Hashiguchi, 2022" authorityName="Nakajima &amp; Hashiguchi" authorityYear="2022" box="[286,521,798,821]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="530" phylum="Chordata" rank="species" species="amamianus" status="sp. nov.">
<emphasis id="B900EA95F0198975FE87FCBCFDA5FC95" box="[286,521,798,821]" italics="true" pageId="5" pageNumber="530">Misgurnus amamianus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23357EEF0198975FD88FCBFFDC9FC94" box="[529,613,797,820]" pageId="5" pageNumber="530" rank="species">sp. nov.</taxonomicNameLabel>
, KPM-NI 68404, holotype. a, mouth; b, lamina circularis (allow indicating the neckline at the base of lamina circularis), bar 1 mm; c, dorsal fin.
</paragraph>
</caption>
<caption id="DF0B660FF0198975FF0EF8EBFD15F825" ID-DOI="http://doi.org/10.5281/zenodo.6816842" ID-Zenodo-Dep="6816842" httpUri="https://zenodo.org/record/6816842/files/figure.png" pageId="5" pageNumber="530" startId="5.[151,250,1865,1889]" targetBox="[151,1436,881,1841]" targetPageId="5">
<paragraph id="8BCB3687F0198975FF0EF8EBFD15F825" blockId="5.[151,1437,1865,1925]" pageId="5" pageNumber="530">
<emphasis id="B900EA95F0198975FF0EF8EBFEBEF8C1" bold="true" box="[151,274,1865,1889]" pageId="5" pageNumber="530">FIGURE 5.</emphasis>
<taxonomicName id="4C744D04F0198975FE80F8E8FDACF8C1" authority="Nakajima &amp; Hashiguchi, 2022" authorityName="Nakajima &amp; Hashiguchi" authorityYear="2022" box="[281,512,1866,1889]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="530" phylum="Chordata" rank="species" species="amamianus" status="sp. nov.">
<emphasis id="B900EA95F0198975FE80F8E8FDACF8C1" box="[281,512,1866,1889]" italics="true" pageId="5" pageNumber="530">Misgurnus amamianus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23357EEF0198975FD9FF8EBFDFAF8C0" box="[518,598,1865,1888]" pageId="5" pageNumber="530" rank="species">sp. nov.</taxonomicNameLabel>
, KPM-NI 68404, holotype. a, radiograph; b, radiograph of dorsal fin; c, radiograph of anal fin; d, radiograph of caudal fin; e, living body.
</paragraph>
</caption>
<paragraph id="8BCB3687F0198975FF5EF811FB02F851" blockId="5.[151,1436,1971,2034]" pageId="5" pageNumber="530">
<emphasis id="B900EA95F0198975FF5EF811FE13F86D" bold="true" box="[199,447,1971,1997]" pageId="5" pageNumber="530">Non-type specimens.</emphasis>
<materialsCitation id="3B1C3CDAF0198975FE51F811FB02F851" ID-GBIF-Occurrence="3861475313" collectingDate="2008-10-24" collectorName="Iriomotejima Is. &amp; T. Suzuki &amp; M. Hosokawa &amp; M. Baba" country="Japan" location="Urauchi" pageId="5" pageNumber="530" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Okinawa">
OMNH-P 37941-37945, 37947, 
<specimenCount id="9D72FD0EF0198975FCA2F816FC03F86D" box="[827,943,1971,1998]" pageId="5" pageNumber="530" type="male">two males</specimenCount>
&amp; 
<specimenCount id="9D72FD0EF0198975FC41F811FBC6F86D" box="[984,1130,1971,1997]" pageId="5" pageNumber="530" type="female">four females</specimenCount>
, 
<collectingCountry id="F3637617F0198975FBEEF816FB17F86E" box="[1143,1211,1972,1998]" name="Japan" pageId="5" pageNumber="530">Japan</collectingCountry>
: 
<location id="8EAB605CF0198975FB51F811FA89F86D" LSID="urn:lsid:plazi:treatment:03DD8791F018897DFF0EFD81FE2BFF75:8EAB605CF0198975FB51F811FA89F86D" box="[1224,1317,1971,1997]" country="Japan" name="Urauchi" pageId="5" pageNumber="530" stateProvince="Okinawa">Urauchi</location>
, 
<collectorName id="26815351F0198975FAABF811FF41F852" pageId="5" pageNumber="530">Iriomotejima Is.</collectorName>
, Yaeyama Isls., 
<collectingRegion id="49B0F865F0198975FE3BF875FDE8F851" box="[418,580,2007,2033]" country="Japan" name="Okinawa" pageId="5" pageNumber="530">Okinawa Pref.</collectingRegion>
<date id="FFCA1047F0198975FDD2F875FD79F851" box="[587,725,2007,2034]" pageId="5" pageNumber="530" value="2008-10-24">
<collectingDate id="EF8EE9AFF0198975FDD2F875FD79F851" box="[587,725,2007,2034]" pageId="5" pageNumber="530" value="2008-10-24">24. X. 2008</collectingDate>
</date>
., 
<collectorName id="26815351F0198975FD7AF87AFCFDF851" box="[739,849,2007,2034]" pageId="5" pageNumber="530">T. Suzuki</collectorName>
, 
<collectorName id="26815351F0198975FCC5F87AFC55F851" box="[860,1017,2007,2034]" pageId="5" pageNumber="530">M. Hosokawa</collectorName>
&amp; 
<collectorName id="26815351F0198975FB84F87AFBD2F851" box="[1053,1150,2007,2034]" pageId="5" pageNumber="530">M. Baba</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF0B660FF01A8976FF0EF9DDFB7FF97F" ID-DOI="http://doi.org/10.5281/zenodo.6816844" ID-Zenodo-Dep="6816844" httpUri="https://zenodo.org/record/6816844/files/figure.png" pageId="6" pageNumber="531" startId="6.[151,250,1663,1687]" targetBox="[151,1435,181,1637]" targetPageId="6">
<paragraph id="8BCB3687F01A8976FF0EF9DDFB7FF97F" blockId="6.[151,1436,1663,1760]" pageId="6" pageNumber="531">
<emphasis id="B900EA95F01A8976FF0EF9DDFEBFF937" bold="true" box="[151,275,1663,1687]" pageId="6" pageNumber="531">FIGURE 6.</emphasis>
Maximum likelihood tree of the partial sequences of mitochondrial DNA control region (CR) in 
<taxonomicName id="4C744D04F01A8976FB6CF922FF4CF91B" authority="Nakajima &amp; Hashiguchi, 2022" authorityName="Nakajima &amp; Hashiguchi" authorityYear="2022" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="531" phylum="Chordata" rank="species" species="amamianus" status="sp. nov.">
<emphasis id="B900EA95F01A8976FB6CF922FF4CF91B" italics="true" pageId="6" pageNumber="531">Misgurnus amamianus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23357EEF01A8976FF7EF906FE99F91B" box="[231,309,1700,1723]" pageId="6" pageNumber="531" rank="species">sp. nov.</taxonomicNameLabel>
and Japanese and Northeast Asian species of 
<taxonomicName id="4C744D04F01A8976FC9DF906FCDCF91B" authorityName="La Cepede" authorityYear="1803" box="[772,880,1700,1723]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="531" phylum="Chordata" rank="genus">
<emphasis id="B900EA95F01A8976FC9DF906FCDCF91B" box="[772,880,1700,1723]" italics="true" pageId="6" pageNumber="531">Misgurnus</emphasis>
</taxonomicName>
. CR sequences of 
<taxonomicName id="4C744D04F01A8976FBB4F901FB52F91B" baseAuthorityName="C.Linnaeus" baseAuthorityYear="1758" box="[1069,1278,1699,1723]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="531" phylum="Chordata" rank="species" species="barbatula">
<emphasis id="B900EA95F01A8976FBB4F901FB52F91B" box="[1069,1278,1699,1723]" italics="true" pageId="6" pageNumber="531">Barbatula barbatula</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C744D04F01A8976FAB7F901FF5CF97F" baseAuthorityName="Bleeker" baseAuthorityYear="1870" class="Actinopterygii" family="Cobitidae" genus="Leptobotia" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="531" phylum="Chordata" rank="species" species="elongata">
<emphasis id="B900EA95F01A8976FAB7F901FF5CF97F" italics="true" pageId="6" pageNumber="531">Leptobotia elongata</emphasis>
</taxonomicName>
are used as outgroups. Numbers on each branch indicate bootstrap values with 1,000 replications.
</paragraph>
</caption>
<subSubSection id="C36E650CF01A8977FF5EF8AFFB47FEBD" lastPageId="7" lastPageNumber="532" pageId="6" pageNumber="531" type="diagnosis">
<paragraph id="8BCB3687F01A8977FF5EF8AFFB47FEBD" blockId="6.[151,1437,1805,2012]" lastBlockId="7.[151,1437,151,285]" lastPageId="7" lastPageNumber="532" pageId="6" pageNumber="531">
<emphasis id="B900EA95F01A8976FF5EF8AFFEEEF887" bold="true" box="[199,322,1805,1831]" pageId="6" pageNumber="531">Diagnosis.</emphasis>
The new species is distinguished from other species of 
<taxonomicName id="4C744D04F01A8976FC33F8ACFB8EF887" authorityName="La Cepede" authorityYear="1803" box="[938,1058,1806,1831]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="531" phylum="Chordata" rank="genus">
<emphasis id="B900EA95F01A8976FC33F8ACFB8EF887" box="[938,1058,1806,1831]" italics="true" pageId="6" pageNumber="531">Misgurnus</emphasis>
</taxonomicName>
by the combination of the following characteristics (
<tableCitation id="C6F6033CF01A8976FEE9F893FE6AF8EC" box="[368,454,1841,1868]" captionStart="TABLE 3" captionStartId="9.[151,239,618,644]" captionTargetPageId="9" captionText="TABLE 3. Comparison of morphological features between Misgurnus" httpUri="http://table.plazi.org/id/DF0B660FF0158979FF0EFDC8FC29FD24" pageId="6" pageNumber="531" tableUuid="DF0B660FF0158979FF0EFDC8FC29FD24">Table 3</tableCitation>
): lamina circularis at base of pectoral fin in adult male poleaxe-shape (vs. spatula-shape in 
<taxonomicName id="4C744D04F01A8976FF2DF8F4FEF4F8CF" authorityName="Kim &amp; Pak" authorityYear="1995" box="[180,344,1877,1903]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="531" phylum="Chordata" rank="species" species="buphoensis">
<emphasis id="B900EA95F01A8976FF2DF8F4FEF4F8CF" box="[180,344,1877,1903]" italics="true" pageId="6" pageNumber="531">M. buphoensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C744D04F01A8976FE16F8F4FDB4F8CF" authorityName="Vasil'eva" authorityYear="2001" box="[399,536,1877,1903]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="531" phylum="Chordata" rank="species" species="nikolskyi">
<emphasis id="B900EA95F01A8976FE16F8F4FDB4F8CF" box="[399,536,1877,1903]" italics="true" pageId="6" pageNumber="531">M. nikolskyi</emphasis>
</taxonomicName>
; elongated horizontally in 
<taxonomicName id="4C744D04F01A8976FCDEF8F4FC4EF8CF" baseAuthorityName="Dabry de Thiersant" baseAuthorityYear="1872" box="[839,994,1877,1903]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="531" phylum="Chordata" rank="species" species="dabryanus">
<emphasis id="B900EA95F01A8976FCDEF8F4FC4EF8CF" box="[839,994,1877,1903]" italics="true" pageId="6" pageNumber="531">M. dabryanus</emphasis>
</taxonomicName>
; absent in 
<taxonomicName id="4C744D04F01A8976FBC4F8F4FB7DF8CF" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1117,1233,1877,1903]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="531" phylum="Chordata" rank="species" species="fossilis">
<emphasis id="B900EA95F01A8976FBC4F8F4FB7DF8CF" box="[1117,1233,1877,1903]" italics="true" pageId="6" pageNumber="531">M. fossilis</emphasis>
</taxonomicName>
); neckline at base of lamina circularis shallow (vs. deep in 
<taxonomicName id="4C744D04F01A8976FDE9F8D8FCFFF833" baseAuthorityName="Cantor" baseAuthorityYear="1842" box="[624,851,1913,1939]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="531" phylum="Chordata" rank="species" species="anguillicaudatus">
<emphasis id="B900EA95F01A8976FDE9F8D8FCFFF833" box="[624,851,1913,1939]" italics="true" pageId="6" pageNumber="531">M. anguillicaudatus</emphasis>
</taxonomicName>
); dorsal fin ray iii+6 (vs. commonly iii+ 
<quantity id="4C8C9B62F01A8976FAB5F8DBFAD9F833" box="[1324,1397,1913,1939]" metricMagnitude="-1" metricUnit="m" metricValue="1.905" metricValueMax="2.032" metricValueMin="1.778" pageId="6" pageNumber="531" unit="in" value="7.5" valueMax="8.0" valueMin="7.0">7–8 in</quantity>
<taxonomicName id="4C744D04F01A8976FAE7F8D8FEFFF817" baseAuthorityName="Cantor" baseAuthorityYear="1842" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="531" phylum="Chordata" rank="species" species="anguillicaudatus">
<emphasis id="B900EA95F01A8976FAE7F8D8FEFFF817" italics="true" pageId="6" pageNumber="531">M. anguillicaudatus</emphasis>
</taxonomicName>
); last ray of dorsal fin commonly not branched (vs. commonly branched in 
<taxonomicName id="4C744D04F01A8976FB2AF83CFA39F817" baseAuthorityName="Cantor" baseAuthorityYear="1842" box="[1203,1429,1949,1975]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="531" phylum="Chordata" rank="species" species="anguillicaudatus">
<emphasis id="B900EA95F01A8976FB2AF83CFA39F817" box="[1203,1429,1949,1975]" italics="true" pageId="6" pageNumber="531">M. anguillicaudatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C744D04F01A8976FF0EF860FE95F87B" authorityName="Kim &amp; Park" authorityYear="1995" box="[151,313,1985,2011]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="531" phylum="Chordata" rank="species" species="buohoensis">
<emphasis id="B900EA95F01A8976FF0EF860FE95F87B" box="[151,313,1985,2011]" italics="true" pageId="6" pageNumber="531">M. buohoensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C744D04F01A8976FEDFF860FE4DF87B" baseAuthorityName="Dabry de Thiersant" baseAuthorityYear="1872" box="[326,481,1985,2011]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="531" phylum="Chordata" rank="species" species="dabryanus">
<emphasis id="B900EA95F01A8976FEDFF860FE4DF87B" box="[326,481,1985,2011]" italics="true" pageId="6" pageNumber="531">M. dabryanus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C744D04F01A8976FE77F860FDCCF87B" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[494,608,1985,2011]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="531" phylum="Chordata" rank="species" species="fossilis">
<emphasis id="B900EA95F01A8976FE77F860FDCCF87B" box="[494,608,1985,2011]" italics="true" pageId="6" pageNumber="531">M. fossilis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C744D04F01A8976FDF4F860FD47F87B" baseAuthorityName="Dybowski" baseAuthorityYear="1869" box="[621,747,1985,2011]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="531" phylum="Chordata" rank="species" species="mohoity">
<emphasis id="B900EA95F01A8976FDF4F860FD47F87B" box="[621,747,1985,2011]" italics="true" pageId="6" pageNumber="531">M. mohoity</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C744D04F01A8976FC86F860FC04F87B" authorityName="Vasil'eva" authorityYear="2001" box="[799,936,1985,2011]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="531" phylum="Chordata" rank="species" species="nikolskyi">
<emphasis id="B900EA95F01A8976FC86F860FC04F87B" box="[799,936,1985,2011]" italics="true" pageId="6" pageNumber="531">M. nikolskyi</emphasis>
</taxonomicName>
); L/S less than 0.5 (vs. over 
<quantity id="4C8C9B62F01A8976FB7BF863FA8DF87B" box="[1250,1313,1985,2012]" metricMagnitude="-2" metricUnit="m" metricValue="1.27" pageId="6" pageNumber="531" unit="in" value="0.5">0.5 in</quantity>
<taxonomicName id="4C744D04F01A8977FABEF860FEA0FF11" baseAuthorityName="Cantor" baseAuthorityYear="1842" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" lastPageId="7" lastPageNumber="532" order="Cypriniformes" pageId="6" pageNumber="531" phylum="Chordata" rank="species" species="anguillicaudatus">
<emphasis id="B900EA95F01A8977FABEF860FEA0FF11" italics="true" lastPageId="7" lastPageNumber="532" pageId="6" pageNumber="531">M. anguillicaudatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C744D04F01B8977FE83FF35FE11FF11" authorityName="Kim &amp; Pak" authorityYear="1995" box="[282,445,151,177]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="532" phylum="Chordata" rank="species" species="buphoensis">
<emphasis id="B900EA95F01B8977FE83FF35FE11FF11" box="[282,445,151,177]" italics="true" pageId="7" pageNumber="532">M. buphoensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C744D04F01B8977FE53FF35FDC9FF11" baseAuthorityName="Dabry de Thiersant" baseAuthorityYear="1872" box="[458,613,151,177]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="532" phylum="Chordata" rank="species" species="dabryanus">
<emphasis id="B900EA95F01B8977FE53FF35FDC9FF11" box="[458,613,151,177]" italics="true" pageId="7" pageNumber="532">M. dabryanus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C744D04F01B8977FDEAFF35FD5EFF11" baseAuthorityName="Dybowski" baseAuthorityYear="1869" box="[627,754,151,177]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="532" phylum="Chordata" rank="species" species="mohoity">
<emphasis id="B900EA95F01B8977FDEAFF35FD5EFF11" box="[627,754,151,177]" italics="true" pageId="7" pageNumber="532">M. mohoity</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C744D04F01B8977FCB1FF35FC1DFF11" authorityName="Vasil'eva" authorityYear="2001" box="[808,945,151,177]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="532" phylum="Chordata" rank="species" species="nikolskyi">
<emphasis id="B900EA95F01B8977FCB1FF35FC1DFF11" box="[808,945,151,177]" italics="true" pageId="7" pageNumber="532">M. nikolskyi</emphasis>
</taxonomicName>
); LPC/DPA over 1.1 (vs. less than 1.0 in 
<taxonomicName id="4C744D04F01B8977FAE7FF35FEFFFF75" baseAuthorityName="Cantor" baseAuthorityYear="1842" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="532" phylum="Chordata" rank="species" species="anguillicaudatus">
<emphasis id="B900EA95F01B8977FAE7FF35FEFFFF75" italics="true" pageId="7" pageNumber="532">M. anguillicaudatus</emphasis>
</taxonomicName>
(
<collectingCountry id="F3637617F01B8977FEFBFF19FE0BFF75" box="[354,423,187,213]" name="China" pageId="7" pageNumber="532">China</collectingCountry>
clade), 
<taxonomicName id="4C744D04F01B8977FE66FF19FD36FF75" baseAuthorityName="Dabry de Thiersant" baseAuthorityYear="1872" box="[511,666,187,213]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="532" phylum="Chordata" rank="species" species="dabryanus">
<emphasis id="B900EA95F01B8977FE66FF19FD36FF75" box="[511,666,187,213]" italics="true" pageId="7" pageNumber="532">M. dabryanus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C744D04F01B8977FD49FF19FCDFFF75" authorityName="Rendahl" authorityYear="1937" box="[720,883,187,213]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="532" phylum="Chordata" rank="species" species="tonkinensis">
<emphasis id="B900EA95F01B8977FD49FF19FCDFFF75" box="[720,883,187,213]" italics="true" pageId="7" pageNumber="532">M. tonkinensis</emphasis>
</taxonomicName>
); barbel length general (vs. long in 
<taxonomicName id="4C744D04F01B8977FA98FF19FA30FF75" baseAuthorityName="Dabry de Thiersant" baseAuthorityYear="1872" box="[1281,1436,187,213]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="532" phylum="Chordata" rank="species" species="dabryanus">
<emphasis id="B900EA95F01B8977FA98FF19FA30FF75" box="[1281,1436,187,213]" italics="true" pageId="7" pageNumber="532">M. dabryanus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C744D04F01B8977FF5CFF7DFEC4FF59" authorityName="Rendahl" authorityYear="1937" box="[197,360,223,249]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="532" phylum="Chordata" rank="species" species="tonkinensis">
<emphasis id="B900EA95F01B8977FF5CFF7DFEC4FF59" box="[197,360,223,249]" italics="true" pageId="7" pageNumber="532">M. tonkinensis</emphasis>
</taxonomicName>
); caudal-peduncle depth shallow (vs. deep in 
<emphasis id="B900EA95F01B8977FCFBFF7DFB77FF59" box="[866,1243,223,249]" italics="true" pageId="7" pageNumber="532">
<taxonomicName id="4C744D04F01B8977FCFBFF7DFC53FF59" baseAuthorityName="Dabry de Thiersant" baseAuthorityYear="1872" box="[866,1023,223,249]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="532" phylum="Chordata" rank="species" species="dabryanus">M. dabryanus</taxonomicName>
, 
<taxonomicName id="4C744D04F01B8977FB90FF7DFB77FF59" authorityName="Rendahl" authorityYear="1944" box="[1033,1243,223,249]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="532" phylum="Chordata" rank="species" species="multimaculatus">M. multimaculatus</taxonomicName>
</emphasis>
and 
<emphasis id="B900EA95F01B8977FA97FF7DFF12FEBD" italics="true" pageId="7" pageNumber="532">
<taxonomicName id="4C744D04F01B8977FA97FF7DFF19FEBD" authorityName="Rendahl" authorityYear="1937" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="532" phylum="Chordata" rank="species" species="tonkinensis">M. tonkinensis</taxonomicName>
)
</emphasis>
; and total vertebrae number 44–48 (vs. over 
<quantity id="4C8C9B62F01B8977FD2EFEA1FD5CFEBD" box="[695,752,259,285]" metricMagnitude="0" metricUnit="m" metricValue="1.2446" pageId="7" pageNumber="532" unit="in" value="49.0">49 in</quantity>
<taxonomicName id="4C744D04F01B8977FD6FFEA1FC36FEBD" authorityName="Kim &amp; Pak" authorityYear="1995" box="[758,922,259,285]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="532" phylum="Chordata" rank="species" species="buphoensis">
<emphasis id="B900EA95F01B8977FD6FFEA1FC36FEBD" box="[758,922,259,285]" italics="true" pageId="7" pageNumber="532">M. buphoensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C744D04F01B8977FC31FEA1FBB7FEBD" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[936,1051,259,285]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="532" phylum="Chordata" rank="species" species="fossilis">
<emphasis id="B900EA95F01B8977FC31FEA1FBB7FEBD" box="[936,1051,259,285]" italics="true" pageId="7" pageNumber="532">M. fossilis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C744D04F01B8977FBC8FEA1FB77FEBD" authorityName="Vasil'eva" authorityYear="2001" box="[1105,1243,259,285]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="532" phylum="Chordata" rank="species" species="nikolskyi">
<emphasis id="B900EA95F01B8977FBC8FEA1FB77FEBD" box="[1105,1243,259,285]" italics="true" pageId="7" pageNumber="532">M. nikolskyi</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<caption id="DF0B660FF01B8977FF0EFC6EFE9CFB38" ID-DOI="http://doi.org/10.5281/zenodo.6816846" ID-Zenodo-Dep="6816846" httpUri="https://zenodo.org/record/6816846/files/figure.png" pageId="7" pageNumber="532" startId="7.[151,250,972,996]" targetBox="[151,1436,309,948]" targetPageId="7">
<paragraph id="8BCB3687F01B8977FF0EFC6EFE9CFB38" blockId="7.[151,1437,972,1176]" pageId="7" pageNumber="532">
<emphasis id="B900EA95F01B8977FF0EFC6EFEBFFC44" bold="true" box="[151,275,972,996]" pageId="7" pageNumber="532">FIGURE 7.</emphasis>
Lamina circularis of Northeast Asian species of 
<taxonomicName id="4C744D04F01B8977FC9DFC6FFCDCFC44" authorityName="La Cepede" authorityYear="1803" box="[772,880,973,996]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="532" phylum="Chordata" rank="genus">
<emphasis id="B900EA95F01B8977FC9DFC6FFCDCFC44" box="[772,880,973,996]" italics="true" pageId="7" pageNumber="532">Misgurnus</emphasis>
</taxonomicName>
. a, 
<taxonomicName id="4C744D04F01B8977FC0CFC6FFB80FC44" authority="Nakajima &amp; Hashiguchi, 2022" authorityName="Nakajima &amp; Hashiguchi" authorityYear="2022" box="[917,1068,973,996]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="532" phylum="Chordata" rank="species" species="amamianus" status="sp. nov.">
<emphasis id="B900EA95F01B8977FC0CFC6FFB80FC44" box="[917,1068,973,996]" italics="true" pageId="7" pageNumber="532">M. amamianus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23357EEF01B8977FBAAFC6EFB2EFC43" box="[1075,1154,972,995]" pageId="7" pageNumber="532" rank="species">sp. nov.</taxonomicNameLabel>
(KPM-NI 68404, holotype, Tokuno-shima Is., Japan, 83.8 mm SL); b, 
<taxonomicName id="4C744D04F01B8977FDA6FC53FCA6FBA8" baseAuthorityName="Cantor" baseAuthorityYear="1842" box="[575,778,1008,1032]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="532" phylum="Chordata" rank="species" species="anguillicaudatus">
<emphasis id="B900EA95F01B8977FDA6FC53FCA6FBA8" box="[575,778,1008,1032]" italics="true" pageId="7" pageNumber="532">M. anguillicaudatus</emphasis>
</taxonomicName>
(Japan clade) (KYUM-PI 4520, Fukuoka, Japan, 93.4 mm SL); c, 
<taxonomicName id="4C744D04F01B8977FF0EFBB7FECFFB8C" baseAuthorityName="Cantor" baseAuthorityYear="1842" box="[151,355,1044,1068]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="532" phylum="Chordata" rank="species" species="anguillicaudatus">
<emphasis id="B900EA95F01B8977FF0EFBB7FECFFB8C" box="[151,355,1044,1068]" italics="true" pageId="7" pageNumber="532">M. anguillicaudatus</emphasis>
</taxonomicName>
(China clade) (KMNH VR 100186, Zheijiang, China, 128.9 mm SL); d, 
<taxonomicName id="4C744D04F01B8977FBDAFBB7FB63FB8C" baseAuthorityName="Dabry de Thiersant" baseAuthorityYear="1872" box="[1091,1231,1044,1068]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="532" phylum="Chordata" rank="species" species="dabryanus">
<emphasis id="B900EA95F01B8977FBDAFBB7FB63FB8C" box="[1091,1231,1044,1068]" italics="true" pageId="7" pageNumber="532">M. dabryanus</emphasis>
</taxonomicName>
(JNC130, Fukuoka, Japan, 115.2 mm SL); e, 
<taxonomicName id="4C744D04F01B8977FE0DFB9BFDA4FBF0" baseAuthorityName="Dybowski" baseAuthorityYear="1869" box="[404,520,1080,1104]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="532" phylum="Chordata" rank="species" species="mohoity">
<emphasis id="B900EA95F01B8977FE0DFB9BFDA4FBF0" box="[404,520,1080,1104]" italics="true" pageId="7" pageNumber="532">M. mohoity</emphasis>
</taxonomicName>
(JNC563, Ibaraki, Japan, 71.7 mm SL); f, 
<taxonomicName id="4C744D04F01B8977FC20FB9BFB99FBF0" authorityName="Vasil'eva" authorityYear="2001" box="[953,1077,1080,1104]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="532" phylum="Chordata" rank="species" species="nikolskyi">
<emphasis id="B900EA95F01B8977FC20FB9BFB99FBF0" box="[953,1077,1080,1104]" italics="true" pageId="7" pageNumber="532">M. nikolskyi</emphasis>
</taxonomicName>
(JNC301, Tokyo, Japan, 121.3 mm SL); g, 
<emphasis id="B900EA95F01B8977FF7AFBFFFEACFBD4" box="[227,256,1117,1140]" italics="true" pageId="7" pageNumber="532">M.</emphasis>
sp. (Clade A) (JNC345, Aomori, Japan, 110.5 mm SL); e, 
<emphasis id="B900EA95F01B8977FCD6FBFFFCC7FBD4" box="[847,875,1117,1140]" italics="true" pageId="7" pageNumber="532">M.</emphasis>
sp. OK (KMNH VR100189, Okinawa, Japan, 77.1 mm SL). Bar 1mm.
</paragraph>
</caption>
<subSubSection id="C36E650CF01B8978FF5EFB64FB5CFA95" lastPageId="8" lastPageNumber="533" pageId="7" pageNumber="532" type="description">
<paragraph id="8BCB3687F01B8977FF5EFB64FF53FA88" blockId="7.[151,1437,1222,2040]" pageId="7" pageNumber="532">
<emphasis id="B900EA95F01B8977FF5EFB64FEF5FB40" bold="true" box="[199,345,1222,1248]" pageId="7" pageNumber="532">Description.</emphasis>
The morphometric and meristic data for the type specimens (
<specimenCount id="9D72FD0EF01B8977FB88FB64FBDAFB40" box="[1041,1142,1222,1248]" pageId="7" pageNumber="532" type="male">17 males</specimenCount>
and 
<specimenCount id="9D72FD0EF01B8977FB37FB64FA87FB40" box="[1198,1323,1222,1248]" pageId="7" pageNumber="532" type="female">10 females</specimenCount>
) are summarised in 
<tableCitation id="C6F6033CF01B8977FE8DFB48FEC7FAA4" box="[276,363,1258,1284]" captionStart="TABLE 2" captionStartId="8.[151,244,150,176]" captionTargetPageId="8" captionText="TABLE 2. Counts and morphometric measurements of Misgurnus amamianus sp. nov." httpUri="http://table.plazi.org/id/DF0B660FF0148978FF0EFF34FBCAFF11" pageId="7" pageNumber="532" tableUuid="DF0B660FF0148978FF0EFF34FBCAFF11">Table 2</tableCitation>
. Data for the 
<typeStatus id="54CF8825F01B8977FD9CFB48FDCBFAA4" box="[517,615,1258,1284]" pageId="7" pageNumber="532" type="holotype">holotype</typeStatus>
are presented first, followed by those for the 
<typeStatus id="54CF8825F01B8977FBF3FB49FB79FAA5" box="[1130,1237,1259,1285]" pageId="7" pageNumber="532" type="paratype">paratypes</typeStatus>
in parentheses, if different.
</paragraph>
<paragraph id="8BCB3687F01B8977FF5EFA90FBF4F95D" blockId="7.[151,1437,1222,2040]" pageId="7" pageNumber="532">
Body elongated, slightly compressed. Head elongated, snout rather short. Interorbital space convex. Eye moderately small, positioned upper and middle on head. Caudal peduncle compressed; ridges on the caudal peduncle at upper and lower edge weakly developed. Mouth small, inferior arched with fleshy lips; lower lip with 4 long, barbellike mental lobes; upper lip with transverse wrinkles on surface (
<figureCitation id="134F2A02F01B8977FCEBFA3CFC69FA18" box="[882,965,1438,1464]" captionStart="FIGURE 4" captionStartId="5.[151,250,797,821]" captionTargetBox="[163,1436,364,773]" captionTargetId="figure-99@5.[151,1436,345,773]" captionTargetPageId="5" captionText="FIGURE 4. Misgurnus amamianus sp. nov., KPM-NI 68404, holotype. a, mouth; b, lamina circularis (allow indicating the neckline at the base of lamina circularis), bar 1 mm; c, dorsal fin." figureDoi="http://doi.org/10.5281/zenodo.6816840" httpUri="https://zenodo.org/record/6816840/files/figure.png" pageId="7" pageNumber="532">Fig. 4a</figureCitation>
). Barbels 3 pairs, first on rostrum, second on maxilla, and third on mandible; each barbels well developed, approximately 30% of head length. No lateral line. Ridges on caudal peduncle at upper and lower edge weakly developed. Small cycloid scales on trunk; head without scale. Lamina circularis at base of pectoral fin in adult males, poleaxe shape, backend round; neckline at base of lamina circularis shallow (
<figureCitation id="134F2A02F01B8977FE27F98CFDB7F9E8" box="[446,539,1582,1608]" captionStart="FIGURE 4" captionStartId="5.[151,250,797,821]" captionTargetBox="[163,1436,364,773]" captionTargetId="figure-99@5.[151,1436,345,773]" captionTargetPageId="5" captionText="FIGURE 4. Misgurnus amamianus sp. nov., KPM-NI 68404, holotype. a, mouth; b, lamina circularis (allow indicating the neckline at the base of lamina circularis), bar 1 mm; c, dorsal fin." figureDoi="http://doi.org/10.5281/zenodo.6816840" httpUri="https://zenodo.org/record/6816840/files/figure.png" pageId="7" pageNumber="532">Figs. 4b</figureCitation>
, 
<figureCitation id="134F2A02F01B8977FDBFF98CFDE8F9E8" box="[550,580,1582,1608]" captionStart="FIGURE 7" captionStartId="7.[151,250,972,996]" captionTargetBox="[151,1436,309,948]" captionTargetId="figure-95@7.[151,1436,309,948]" captionTargetPageId="7" captionText="FIGURE 7. Lamina circularis of Northeast Asian species of Misgurnus. a, M. amamianus sp. nov. (KPM-NI 68404, holotype, Tokuno-shima Is., Japan, 83.8 mm SL); b, M. anguillicaudatus (Japan clade) (KYUM-PI 4520, Fukuoka, Japan, 93.4 mm SL); c, M. anguillicaudatus (China clade) (KMNH VR 100186, Zheijiang, China, 128.9 mm SL); d, M. dabryanus (JNC130, Fukuoka, Japan, 115.2 mm SL); e, M. mohoity (JNC563, Ibaraki, Japan, 71.7 mm SL); f, M. nikolskyi (JNC301, Tokyo, Japan, 121.3 mm SL); g, M. sp. (Clade A) (JNC345, Aomori, Japan, 110.5 mm SL); e, M. sp. OK (KMNH VR100189, Okinawa, Japan, 77.1 mm SL). Bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.6816846" httpUri="https://zenodo.org/record/6816846/files/figure.png" pageId="7" pageNumber="532">7a</figureCitation>
). First branched soft ray of pectoral fin longer than others, broad and slightly extended; pectoral fin in males longer than that in females. Dorsal-fin ray thin and weak; last ray unbranched, its length less than half of second branched soft ray (
<figureCitation id="134F2A02F01B8977FD23F9D4FCB8F930" box="[698,788,1654,1680]" captionStart="FIGURE 4" captionStartId="5.[151,250,797,821]" captionTargetBox="[163,1436,364,773]" captionTargetId="figure-99@5.[151,1436,345,773]" captionTargetPageId="5" captionText="FIGURE 4. Misgurnus amamianus sp. nov., KPM-NI 68404, holotype. a, mouth; b, lamina circularis (allow indicating the neckline at the base of lamina circularis), bar 1 mm; c, dorsal fin." figureDoi="http://doi.org/10.5281/zenodo.6816840" httpUri="https://zenodo.org/record/6816840/files/figure.png" pageId="7" pageNumber="532">Figs. 4c</figureCitation>
, 
<figureCitation id="134F2A02F01B8977FC86F9D4FC93F931" box="[799,831,1654,1681]" captionStart="FIGURE 5" captionStartId="5.[151,250,1865,1889]" captionTargetBox="[151,1436,881,1841]" captionTargetId="figure-136@5.[151,1436,881,1841]" captionTargetPageId="5" captionText="FIGURE 5. Misgurnus amamianus sp. nov., KPM-NI 68404, holotype. a, radiograph; b, radiograph of dorsal fin; c, radiograph of anal fin; d, radiograph of caudal fin; e, living body." figureDoi="http://doi.org/10.5281/zenodo.6816842" httpUri="https://zenodo.org/record/6816842/files/figure.png" pageId="7" pageNumber="532">5b</figureCitation>
). Margin of dorsal fin rounded, slightly pointed at center tip. Dorsal-fin base behind middle of body. Pelvic-fin origin below third or fourth branched dorsal-fin ray. Anal fin not reaching caudal-fin base. Margin of caudal fin rounded, slightly pointed at center tip. Abdominal vertebrae 30 (27–30); caudal vertebrae 18 (15–19); total vertebrae 48 (44–48) (
<emphasis id="B900EA95F01B8977FC06F941FC01F95C" box="[927,941,1763,1788]" italics="true" pageId="7" pageNumber="532">n</emphasis>
= 20) (
<figureCitation id="134F2A02F01B8977FC66F940FBE7F95D" box="[1023,1099,1762,1789]" captionStart="FIGURE 5" captionStartId="5.[151,250,1865,1889]" captionTargetBox="[151,1436,881,1841]" captionTargetId="figure-136@5.[151,1436,881,1841]" captionTargetPageId="5" captionText="FIGURE 5. Misgurnus amamianus sp. nov., KPM-NI 68404, holotype. a, radiograph; b, radiograph of dorsal fin; c, radiograph of anal fin; d, radiograph of caudal fin; e, living body." figureDoi="http://doi.org/10.5281/zenodo.6816842" httpUri="https://zenodo.org/record/6816842/files/figure.png" pageId="7" pageNumber="532">Fig 5a</figureCitation>
).
</paragraph>
<paragraph id="8BCB3687F01B8977FF5EF8A4FF13F810" blockId="7.[151,1437,1222,2040]" pageId="7" pageNumber="532">
<emphasis id="B900EA95F01B8977FF5EF8A4FDEAF880" bold="true" box="[199,582,1798,1824]" pageId="7" pageNumber="532">Colouration of fresh specimens.</emphasis>
Head and back blackish brown with irregular and indistinct spots; slightly purplish on cheeks and body; abdomen yellowish brown (
<figureCitation id="134F2A02F01B8977FCBCF888FCD9F8E4" box="[805,885,1834,1860]" captionStart="FIGURE 3" captionStartId="4.[151,250,1596,1620]" captionTargetBox="[151,1436,930,1572]" captionTargetId="figure-239@4.[151,1436,930,1572]" captionTargetPageId="4" captionText="FIGURE 3. Misgurnus amamianus sp. nov. a, holotype (male, KPM-NI 68404); b, paratype (female, TKPM-P 26178)." figureDoi="http://doi.org/10.5281/zenodo.6816838" httpUri="https://zenodo.org/record/6816838/files/figure.png" pageId="7" pageNumber="532">
Figs. 
<geoCoordinate id="EE405040F01B8977FCFFF888FCD9F8E4" box="[870,885,1834,1860]" degrees="3, 5" direction="east" orientation="longitude" pageId="7" pageNumber="532" precision="5555" value="3.5">3</geoCoordinate>
</figureCitation>
, 
<figureCitation id="134F2A02F01B8977FC1BF888FC0CF8E5" box="[898,928,1834,1861]" captionStart="FIGURE 5" captionStartId="5.[151,250,1865,1889]" captionTargetBox="[151,1436,881,1841]" captionTargetId="figure-136@5.[151,1436,881,1841]" captionTargetPageId="5" captionText="FIGURE 5. Misgurnus amamianus sp. nov., KPM-NI 68404, holotype. a, radiograph; b, radiograph of dorsal fin; c, radiograph of anal fin; d, radiograph of caudal fin; e, living body." figureDoi="http://doi.org/10.5281/zenodo.6816842" httpUri="https://zenodo.org/record/6816842/files/figure.png" pageId="7" pageNumber="532">5e</figureCitation>
). Unclear streak running from tip of snout to occiput, crossing to eye. Dorsal and caudal fins with irregular and indistinct spots. Upper spot at caudal-fin base ambiguous, size smaller than eye diameter. Ridges on caudal peduncle at upper and lower edge tinged with yellowish.
</paragraph>
<paragraph id="8BCB3687F01B8977FF5EF818FCD0F858" blockId="7.[151,1437,1222,2040]" pageId="7" pageNumber="532">
<emphasis id="B900EA95F01B8977FF5EF818FE1BF874" bold="true" box="[199,439,1978,2004]" pageId="7" pageNumber="532">Sexual dimorphism.</emphasis>
A lamina circularis of pectoral fin only in males; pectoral fin in males longer than that in females. Generally, body size of females larger than that of males.
</paragraph>
<caption id="DF0B660FF0148978FF0EFF34FBCAFF11" ID-Table-UUID="DF0B660FF0148978FF0EFF34FBCAFF11" box="[151,1126,150,177]" httpUri="http://table.plazi.org/id/DF0B660FF0148978FF0EFF34FBCAFF11" pageId="8" pageNumber="533" startId="8.[151,244,150,176]" targetBox="[163,1312,192,1182]" targetIsTable="true" targetPageId="8">
<paragraph id="8BCB3687F0148978FF0EFF34FBCAFF11" blockId="8.[151,1126,150,177]" box="[151,1126,150,177]" pageId="8" pageNumber="533">
<emphasis id="B900EA95F0148978FF0EFF34FEBCFF10" bold="true" box="[151,272,150,176]" pageId="8" pageNumber="533">TABLE 2.</emphasis>
Counts and morphometric measurements of 
<taxonomicName id="4C744D04F0148978FC91FF35FBA4FF10" authority="Nakajima &amp; Hashiguchi, 2022" authorityName="Nakajima &amp; Hashiguchi" authorityYear="2022" box="[776,1032,151,176]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="amamianus" status="sp. nov.">
<emphasis id="B900EA95F0148978FC91FF35FBA4FF10" box="[776,1032,151,176]" italics="true" pageId="8" pageNumber="533">Misgurnus amamianus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23357EEF0148978FB96FF35FBCAFF11" box="[1039,1126,151,177]" pageId="8" pageNumber="533" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BCB3687F0148978FD3CFF76FB41FB3E" pageId="8" pageNumber="533">
<table id="F974C427F014768FFF3AFF62FA8CFB3E" box="[163,1312,192,1182]" gridcols="4" gridrows="24" pageId="8" pageNumber="533">
<tr id="354434C5F014768FFF3AFF62FA8CFEA0" box="[163,1312,192,256]" gridrow="0" pageId="8" pageNumber="533" rowspan-0="1">
<th id="76955DB9F014768FFD3CFF62FCA4FEA0" box="[677,776,192,256]" gridcol="1" gridrow="0" pageId="8" pageNumber="533">
<emphasis id="B900EA95F0148978FD3CFF76FCA4FF4C" bold="true" box="[677,776,212,236]" pageId="8" pageNumber="533">Holotype</emphasis>
</th>
<th id="76955DB9F014768FFCD0FF62FC5EFEA0" box="[841,1010,192,256]" gridcol="2" gridrow="0" pageId="8" pageNumber="533">
<emphasis id="B900EA95F0148978FCD0FF4AFC78FEA0" bold="true" box="[841,980,232,256]" pageId="8" pageNumber="533">males (n=16)</emphasis>
</th>
<th id="76955DB9F014768FFBAAFF62FA8CFEA0" box="[1075,1312,192,256]" gridcol="3" gridrow="0" pageId="8" pageNumber="533">
<emphasis id="B900EA95F0148978FBAAFF62FAA4FEA0" bold="true" box="[1075,1288,192,256]" pageId="8" pageNumber="533">Paratypes females (n=10)</emphasis>
</th>
</tr>
<tr id="354434C5F014768FFF3AFEB3FA8CFE89" box="[163,1312,273,297]" gridrow="1" pageId="8" pageNumber="533">
<th id="76955DB9F014768FFF3AFEB3FDB9FE89" box="[163,533,273,297]" gridcol="0" gridrow="1" pageId="8" pageNumber="533">Standard length (mm); mean (range)</th>
<td id="76955DB9F014768FFD3CFEB3FCA4FE89" box="[677,776,273,297]" gridcol="1" gridrow="1" pageId="8" pageNumber="533">83.8</td>
<td id="76955DB9F014768FFCD0FEB3FC5EFE89" box="[841,1010,273,297]" gridcol="2" gridrow="1" pageId="8" pageNumber="533">75.2 (63.3–89.4)</td>
<td id="76955DB9F014768FFBAAFEB3FA8CFE89" box="[1075,1312,273,297]" gridcol="3" gridrow="1" pageId="8" pageNumber="533">92.1 (55.1–126.3)</td>
</tr>
<tr id="354434C5F014768FFF3AFE9BFA8CFEF1" box="[163,1312,313,337]" gridrow="2" pageId="8" pageNumber="533">
<th id="76955DB9F014768FFF3AFE9BFA8CFEF1" box="[163,1312,313,337]" colspan="4" colspanRight="3" gridcol="0" gridrow="2" pageId="8" pageNumber="533">Counts</th>
</tr>
<tr id="354434C5F014768FFF3AFEC3FA8CFEDA" box="[163,1312,353,378]" gridrow="3" pageId="8" pageNumber="533">
<th id="76955DB9F014768FFF3AFEC3FDB9FEDA" box="[163,533,353,378]" gridcol="0" gridrow="3" pageId="8" pageNumber="533">Dorsal fin rays</th>
<td id="76955DB9F014768FFD3CFEC3FCA4FEDA" box="[677,776,353,378]" gridcol="1" gridrow="3" pageId="8" pageNumber="533">iii+6</td>
<td id="76955DB9F014768FFCD0FEC3FC5EFEDA" box="[841,1010,353,378]" gridcol="2" gridrow="3" pageId="8" pageNumber="533">iii+6</td>
<td id="76955DB9F014768FFBAAFEC3FA8CFEDA" box="[1075,1312,353,378]" gridcol="3" gridrow="3" pageId="8" pageNumber="533">iii+6</td>
</tr>
<tr id="354434C5F014768FFF3AFE2BFA8CFE02" box="[163,1312,393,418]" gridrow="4" pageId="8" pageNumber="533">
<th id="76955DB9F014768FFF3AFE2BFDB9FE02" box="[163,533,393,418]" gridcol="0" gridrow="4" pageId="8" pageNumber="533">Anal fin rays</th>
<td id="76955DB9F014768FFD3CFE2BFCA4FE02" box="[677,776,393,418]" gridcol="1" gridrow="4" pageId="8" pageNumber="533">ii+5</td>
<td id="76955DB9F014768FFCD0FE2BFC5EFE02" box="[841,1010,393,418]" gridcol="2" gridrow="4" pageId="8" pageNumber="533">ii+4–5</td>
<td id="76955DB9F014768FFBAAFE2BFA8CFE02" box="[1075,1312,393,418]" gridcol="3" gridrow="4" pageId="8" pageNumber="533">ii+4–6</td>
</tr>
<tr id="354434C5F014768FFF3AFE10FA8CFE6A" box="[163,1312,434,458]" gridrow="5" pageId="8" pageNumber="533">
<th id="76955DB9F014768FFF3AFE10FDB9FE6A" box="[163,533,434,458]" gridcol="0" gridrow="5" pageId="8" pageNumber="533">Pectoral fin rays</th>
<td id="76955DB9F014768FFD3CFE10FCA4FE6A" box="[677,776,434,458]" gridcol="1" gridrow="5" pageId="8" pageNumber="533">i+9</td>
<td id="76955DB9F014768FFCD0FE10FC5EFE6A" box="[841,1010,434,458]" gridcol="2" gridrow="5" pageId="8" pageNumber="533">i+8-9</td>
<td id="76955DB9F014768FFBAAFE10FA8CFE6A" box="[1075,1312,434,458]" gridcol="3" gridrow="5" pageId="8" pageNumber="533">i+8–9</td>
</tr>
<tr id="354434C5F014768FFF3AFE78FA8CFE52" box="[163,1312,474,498]" gridrow="6" pageId="8" pageNumber="533">
<th id="76955DB9F014768FFF3AFE78FDB9FE52" box="[163,533,474,498]" gridcol="0" gridrow="6" pageId="8" pageNumber="533">Pelvic fin rays</th>
<td id="76955DB9F014768FFD3CFE78FCA4FE52" box="[677,776,474,498]" gridcol="1" gridrow="6" pageId="8" pageNumber="533">i+5</td>
<td id="76955DB9F014768FFCD0FE78FC5EFE52" box="[841,1010,474,498]" gridcol="2" gridrow="6" pageId="8" pageNumber="533">i+4–5</td>
<td id="76955DB9F014768FFBAAFE78FA8CFE52" box="[1075,1312,474,498]" gridcol="3" gridrow="6" pageId="8" pageNumber="533">i+3–5</td>
</tr>
<tr id="354434C5F014768FFF3AFDA0FA8CFDBA" box="[163,1312,514,538]" gridrow="7" pageId="8" pageNumber="533">
<th id="76955DB9F014768FFF3AFDA0FDB9FDBA" box="[163,533,514,538]" gridcol="0" gridrow="7" pageId="8" pageNumber="533">Caudal fin rays*</th>
<td id="76955DB9F014768FFD3CFDA0FCA4FDBA" box="[677,776,514,538]" gridcol="1" gridrow="7" pageId="8" pageNumber="533">5+5</td>
<td id="76955DB9F014768FFCD0FDA0FC5EFDBA" box="[841,1010,514,538]" gridcol="2" gridrow="7" pageId="8" pageNumber="533">6–8+7</td>
<td id="76955DB9F014768FFBAAFDA0FA8CFDBA" box="[1075,1312,514,538]" gridcol="3" gridrow="7" pageId="8" pageNumber="533">7–8+6–7</td>
</tr>
<tr id="354434C5F014768FFF3AFD88FA8CFDE3" box="[163,1312,554,579]" gridrow="8" pageId="8" pageNumber="533" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="76955DB9F014768FFF3AFD88FDB9FDE3" box="[163,533,554,579]" gridcol="0" gridrow="8" pageId="8" pageNumber="533">In % standard length; mean (range)</th>
</tr>
<tr id="354434C5F014768FFF3AFDF0FA8CFDCB" box="[163,1312,594,619]" gridrow="9" pageId="8" pageNumber="533">
<th id="76955DB9F014768FFF3AFDF0FDB9FDCB" box="[163,533,594,619]" gridcol="0" gridrow="9" pageId="8" pageNumber="533">Head length</th>
<td id="76955DB9F014768FFD3CFDF0FCA4FDCB" box="[677,776,594,619]" gridcol="1" gridrow="9" pageId="8" pageNumber="533">16.8</td>
<td id="76955DB9F014768FFCD0FDF0FC5EFDCB" box="[841,1010,594,619]" gridcol="2" gridrow="9" pageId="8" pageNumber="533">17.0 (15.4–18.7)</td>
<td id="76955DB9F014768FFBAAFDF0FA8CFDCB" box="[1075,1312,594,619]" gridcol="3" gridrow="9" pageId="8" pageNumber="533">17.7 (16.1–18.7)</td>
</tr>
<tr id="354434C5F014768FFF3AFDD9FA8CFD33" box="[163,1312,635,659]" gridrow="10" pageId="8" pageNumber="533">
<th id="76955DB9F014768FFF3AFDD9FDB9FD33" box="[163,533,635,659]" gridcol="0" gridrow="10" pageId="8" pageNumber="533">Body depth</th>
<td id="76955DB9F014768FFD3CFDD9FCA4FD33" box="[677,776,635,659]" gridcol="1" gridrow="10" pageId="8" pageNumber="533">10.9</td>
<td id="76955DB9F014768FFCD0FDD9FC5EFD33" box="[841,1010,635,659]" gridcol="2" gridrow="10" pageId="8" pageNumber="533">12.4 (10.1–15.6)</td>
<td id="76955DB9F014768FFBAAFDD9FA8CFD33" box="[1075,1312,635,659]" gridcol="3" gridrow="10" pageId="8" pageNumber="533">12.1 (10.5–15.3)</td>
</tr>
<tr id="354434C5F014768FFF3AFD01FA8CFD1B" box="[163,1312,675,699]" gridrow="11" pageId="8" pageNumber="533">
<th id="76955DB9F014768FFF3AFD01FDB9FD1B" box="[163,533,675,699]" gridcol="0" gridrow="11" pageId="8" pageNumber="533">Depth of caudal peduncle</th>
<td id="76955DB9F014768FFD3CFD01FCA4FD1B" box="[677,776,675,699]" gridcol="1" gridrow="11" pageId="8" pageNumber="533">8.8</td>
<td id="76955DB9F014768FFCD0FD01FC5EFD1B" box="[841,1010,675,699]" gridcol="2" gridrow="11" pageId="8" pageNumber="533">10.0 (8.4–11.4)</td>
<td id="76955DB9F014768FFBAAFD01FA8CFD1B" box="[1075,1312,675,699]" gridcol="3" gridrow="11" pageId="8" pageNumber="533">9.9 (8.0–11.0)</td>
</tr>
<tr id="354434C5F014768FFF3AFD69FA8CFD43" box="[163,1312,715,739]" gridrow="12" pageId="8" pageNumber="533">
<th id="76955DB9F014768FFF3AFD69FDB9FD43" box="[163,533,715,739]" gridcol="0" gridrow="12" pageId="8" pageNumber="533">Predorsal length</th>
<td id="76955DB9F014768FFD3CFD69FCA4FD43" box="[677,776,715,739]" gridcol="1" gridrow="12" pageId="8" pageNumber="533">56.3</td>
<td id="76955DB9F014768FFCD0FD69FC5EFD43" box="[841,1010,715,739]" gridcol="2" gridrow="12" pageId="8" pageNumber="533">56.2 (54.2–58.9)</td>
<td id="76955DB9F014768FFBAAFD69FA8CFD43" box="[1075,1312,715,739]" gridcol="3" gridrow="12" pageId="8" pageNumber="533">57.1 (56.0–59.2)</td>
</tr>
<tr id="354434C5F014768FFF3AFD51FA8CFCAC" box="[163,1312,755,780]" gridrow="13" pageId="8" pageNumber="533">
<th id="76955DB9F014768FFF3AFD51FDB9FCAC" box="[163,533,755,780]" gridcol="0" gridrow="13" pageId="8" pageNumber="533">Prepelvic length</th>
<td id="76955DB9F014768FFD3CFD51FCA4FCAC" box="[677,776,755,780]" gridcol="1" gridrow="13" pageId="8" pageNumber="533">57.5</td>
<td id="76955DB9F014768FFCD0FD51FC5EFCAC" box="[841,1010,755,780]" gridcol="2" gridrow="13" pageId="8" pageNumber="533">58.1 (56.0–60.2)</td>
<td id="76955DB9F014768FFBAAFD51FA8CFCAC" box="[1075,1312,755,780]" gridcol="3" gridrow="13" pageId="8" pageNumber="533">59.2 (57.3–60.6)</td>
</tr>
<tr id="354434C5F014768FFF3AFCBEFA8CFC94" box="[163,1312,796,820]" gridrow="14" pageId="8" pageNumber="533">
<th id="76955DB9F014768FFF3AFCBEFDB9FC94" box="[163,533,796,820]" gridcol="0" gridrow="14" pageId="8" pageNumber="533">Preanal length</th>
<td id="76955DB9F014768FFD3CFCBEFCA4FC94" box="[677,776,796,820]" gridcol="1" gridrow="14" pageId="8" pageNumber="533">69.1</td>
<td id="76955DB9F014768FFCD0FCBEFC5EFC94" box="[841,1010,796,820]" gridcol="2" gridrow="14" pageId="8" pageNumber="533">68.9 (66.8–71.8)</td>
<td id="76955DB9F014768FFBAAFCBEFA8CFC94" box="[1075,1312,796,820]" gridcol="3" gridrow="14" pageId="8" pageNumber="533">69.9 (68.4–72.0)</td>
</tr>
<tr id="354434C5F014768FFF3AFCE6FA8CFCFC" box="[163,1312,836,860]" gridrow="15" pageId="8" pageNumber="533">
<th id="76955DB9F014768FFF3AFCE6FA8CFCFC" box="[163,1312,836,860]" colspan="4" colspanRight="3" gridcol="0" gridrow="15" pageId="8" pageNumber="533">In % head length; mean (range)</th>
</tr>
<tr id="354434C5F014768FFF3AFCCEFA8CFC24" box="[163,1312,876,900]" gridrow="16" pageId="8" pageNumber="533">
<th id="76955DB9F014768FFF3AFCCEFDB9FC24" box="[163,533,876,900]" gridcol="0" gridrow="16" pageId="8" pageNumber="533">Snout length</th>
<td id="76955DB9F014768FFD3CFCCEFCA4FC24" box="[677,776,876,900]" gridcol="1" gridrow="16" pageId="8" pageNumber="533">38.3</td>
<td id="76955DB9F014768FFCD0FCCEFC5EFC24" box="[841,1010,876,900]" gridcol="2" gridrow="16" pageId="8" pageNumber="533">38.0 (31.2–42.5)</td>
<td id="76955DB9F014768FFBAAFCCEFA8CFC24" box="[1075,1312,876,900]" gridcol="3" gridrow="16" pageId="8" pageNumber="533">38.8 (36.9–42.3)</td>
</tr>
<tr id="354434C5F014768FFF3AFC36FA8CFC0D" box="[163,1312,916,941]" gridrow="17" pageId="8" pageNumber="533">
<th id="76955DB9F014768FFF3AFC36FDB9FC0D" box="[163,533,916,941]" gridcol="0" gridrow="17" pageId="8" pageNumber="533">Orbit diameter</th>
<td id="76955DB9F014768FFD3CFC36FCA4FC0D" box="[677,776,916,941]" gridcol="1" gridrow="17" pageId="8" pageNumber="533">12.8</td>
<td id="76955DB9F014768FFCD0FC36FC5EFC0D" box="[841,1010,916,941]" gridcol="2" gridrow="17" pageId="8" pageNumber="533">14.8 (12.3–16.8)</td>
<td id="76955DB9F014768FFBAAFC36FA8CFC0D" box="[1075,1312,916,941]" gridcol="3" gridrow="17" pageId="8" pageNumber="533">12.8 (10.1–14.7)</td>
</tr>
<tr id="354434C5F014768FFF3AFC1FFA8CFC75" box="[163,1312,957,981]" gridrow="18" pageId="8" pageNumber="533">
<th id="76955DB9F014768FFF3AFC1FFDB9FC75" box="[163,533,957,981]" gridcol="0" gridrow="18" pageId="8" pageNumber="533">Interorbital width</th>
<td id="76955DB9F014768FFD3CFC1FFCA4FC75" box="[677,776,957,981]" gridcol="1" gridrow="18" pageId="8" pageNumber="533">23.4</td>
<td id="76955DB9F014768FFCD0FC1FFC5EFC75" box="[841,1010,957,981]" gridcol="2" gridrow="18" pageId="8" pageNumber="533">22.9 (16.0–26.8)</td>
<td id="76955DB9F014768FFBAAFC1FFA8CFC75" box="[1075,1312,957,981]" gridcol="3" gridrow="18" pageId="8" pageNumber="533">23.2 (16.9–28.5)</td>
</tr>
<tr id="354434C5F014768FFF3AFC47FA8CFC5D" box="[163,1312,997,1021]" gridrow="19" pageId="8" pageNumber="533">
<th id="76955DB9F014768FFF3AFC47FDB9FC5D" box="[163,533,997,1021]" gridcol="0" gridrow="19" pageId="8" pageNumber="533">Rostral barbel</th>
<td id="76955DB9F014768FFD3CFC47FCA4FC5D" box="[677,776,997,1021]" gridcol="1" gridrow="19" pageId="8" pageNumber="533">31.9</td>
<td id="76955DB9F014768FFCD0FC47FC5EFC5D" box="[841,1010,997,1021]" gridcol="2" gridrow="19" pageId="8" pageNumber="533">30.1 (21.7–36.2)</td>
<td id="76955DB9F014768FFBAAFC47FA8CFC5D" box="[1075,1312,997,1021]" gridcol="3" gridrow="19" pageId="8" pageNumber="533">25.1 (13.7–31.1)</td>
</tr>
<tr id="354434C5F014768FFF3AFBAFFA8CFB85" box="[163,1312,1037,1061]" gridrow="20" pageId="8" pageNumber="533">
<th id="76955DB9F014768FFF3AFBAFFDB9FB85" box="[163,533,1037,1061]" gridcol="0" gridrow="20" pageId="8" pageNumber="533">Maxillary barbel</th>
<td id="76955DB9F014768FFD3CFBAFFCA4FB85" box="[677,776,1037,1061]" gridcol="1" gridrow="20" pageId="8" pageNumber="533">31.2</td>
<td id="76955DB9F014768FFCD0FBAFFC5EFB85" box="[841,1010,1037,1061]" gridcol="2" gridrow="20" pageId="8" pageNumber="533">32.4 (28.5–37.7)</td>
<td id="76955DB9F014768FFBAAFBAFFA8CFB85" box="[1075,1312,1037,1061]" gridcol="3" gridrow="20" pageId="8" pageNumber="533">27.4 (16.5–32.2)</td>
</tr>
<tr id="354434C5F014768FFF3AFB97FA8CFBEE" box="[163,1312,1077,1102]" gridrow="21" pageId="8" pageNumber="533">
<th id="76955DB9F014768FFF3AFB97FDB9FBEE" box="[163,533,1077,1102]" gridcol="0" gridrow="21" pageId="8" pageNumber="533">Mandibular barbel</th>
<td id="76955DB9F014768FFD3CFB97FCA4FBEE" box="[677,776,1077,1102]" gridcol="1" gridrow="21" pageId="8" pageNumber="533">33.3</td>
<td id="76955DB9F014768FFCD0FB97FC5EFBEE" box="[841,1010,1077,1102]" gridcol="2" gridrow="21" pageId="8" pageNumber="533">36.1 (30.7–45.1)</td>
<td id="76955DB9F014768FFBAAFB97FA8CFBEE" box="[1075,1312,1077,1102]" gridcol="3" gridrow="21" pageId="8" pageNumber="533">32.1 (19.4–40.8)</td>
</tr>
<tr id="354434C5F014768FFF3AFBFCFA8CFBD6" box="[163,1312,1118,1142]" gridrow="22" pageId="8" pageNumber="533">
<th id="76955DB9F014768FFF3AFBFCFDB9FBD6" box="[163,533,1118,1142]" gridcol="0" gridrow="22" pageId="8" pageNumber="533">LPC/DPA</th>
<td id="76955DB9F014768FFD3CFBFCFCA4FBD6" box="[677,776,1118,1142]" gridcol="1" gridrow="22" pageId="8" pageNumber="533">1.5</td>
<td id="76955DB9F014768FFCD0FBFCFC5EFBD6" box="[841,1010,1118,1142]" gridcol="2" gridrow="22" pageId="8" pageNumber="533">1.3 (1.1–1.6)</td>
<td id="76955DB9F014768FFBAAFBFCFA8CFBD6" box="[1075,1312,1118,1142]" gridcol="3" gridrow="22" pageId="8" pageNumber="533">1.2 (1.1–1.4)</td>
</tr>
<tr id="354434C5F014768FFF3AFB24FA8CFB3E" box="[163,1312,1158,1182]" gridrow="23" pageId="8" pageNumber="533">
<th id="76955DB9F014768FFF3AFB24FDB9FB3E" box="[163,533,1158,1182]" gridcol="0" gridrow="23" pageId="8" pageNumber="533">L/S</th>
<td id="76955DB9F014768FFD3CFB24FCA4FB3E" box="[677,776,1158,1182]" gridcol="1" gridrow="23" pageId="8" pageNumber="533">0.4</td>
<td id="76955DB9F014768FFCD0FB24FC5EFB3E" box="[841,1010,1158,1182]" gridcol="2" gridrow="23" pageId="8" pageNumber="533">0.4 (0.4–0.5)</td>
<td id="76955DB9F014768FFBAAFB24FA8CFB3E" box="[1075,1312,1158,1182]" gridcol="3" gridrow="23" pageId="8" pageNumber="533">0.4 (0.3–0.5)</td>
</tr>
</table>
</paragraph>
<paragraph id="8BCB3687F0148978FF3AFB0CFBDCFB44" blockId="8.[163,1418,1198,1252]" pageId="8" pageNumber="533">
<tableNote id="76923709F0148978FF3AFB0CFBDCFB44" pageId="8" pageNumber="533" targetBox="[163,1312,192,1182]" targetPageId="8">DPA, distance between the pelvic and anal fin; LPC, length of caudal peduncle. L, the length of last soft ray of dorsal fin; S, the length of the second soft ray of dorsal fin. See also Figure 2. *, Counted only branched rays.</tableNote>
</paragraph>
<paragraph id="8BCB3687F0148978FF5EFAB8FB5CFA95" blockId="8.[151,1437,1306,2017]" box="[199,1264,1306,1333]" pageId="8" pageNumber="533">
<emphasis id="B900EA95F0148978FF5EFAB8FD83FA95" bold="true" box="[199,559,1306,1333]" pageId="8" pageNumber="533">Mitochondrial DNA sequence.</emphasis>
<materialsCitation id="3B1C3CDAF0148978FDAFFAB9FB40FA95" ID-GBIF-Occurrence="3861475308" accessionNumber="LC672439, LC278291" box="[566,1260,1306,1333]" location="Control region" pageId="8" pageNumber="533" specimenCount="1" typeStatus="holotype">
<location id="8EAB605CF0148978FDAFFAB9FD73FA95" LSID="urn:lsid:plazi:treatment:03DD8791F018897DFF0EFD81FE2BFF75:8EAB605CF0148978FDAFFAB9FD73FA95" box="[566,735,1307,1333]" name="Control region" pageId="8" pageNumber="533">Control region</location>
: 
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(
<typeStatus id="54CF8825F0148978FCE9FAB9FC73FA95" box="[880,991,1307,1333]" pageId="8" pageNumber="533" type="holotype">Holotype</typeStatus>
); 12SrRNA: 
<accessionNumber id="9427AB64F0148978FBE9FAB9FB40FA95" box="[1136,1260,1307,1333]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC278291" pageId="8" pageNumber="533">LC278291</accessionNumber>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36E650CF0148978FF5EFA9CFDFDF9AD" pageId="8" pageNumber="533" type="distribution">
<paragraph id="8BCB3687F0148978FF5EFA9CFF5EFA01" blockId="8.[151,1437,1306,2017]" pageId="8" pageNumber="533">
<emphasis id="B900EA95F0148978FF5EFA9CFECDFAF8" bold="true" box="[199,353,1342,1368]" pageId="8" pageNumber="533">Distribution.</emphasis>
<collectingCountry id="F3637617F0148978FEF4FA9DFE1DFAF9" box="[365,433,1343,1369]" name="Japan" pageId="8" pageNumber="533">Japan</collectingCountry>
: Amami Islands. (Kikai-jima Island, Amami-oshima Isand, Tokuno-shima Island and Okinoerabu-jima Islamd), 
<collectingRegion id="49B0F865F0148978FE5CFAC1FD68FADD" box="[453,708,1379,1405]" country="Japan" name="Kagoshima" pageId="8" pageNumber="533">Kagoshima Prefecture</collectingRegion>
, introduced to Iriomote-jima Island, 
<collectingRegion id="49B0F865F0148978FBF2FAC1FAE3FADD" box="[1131,1359,1379,1405]" country="Japan" name="Okinawa" pageId="8" pageNumber="533">Okinawa Prefecture</collectingRegion>
, 
<collectingCountry id="F3637617F0148978FAC5FAC1FA30FADD" box="[1372,1436,1379,1405]" name="Japan" pageId="8" pageNumber="533">Japan</collectingCountry>
(
<figureCitation id="134F2A02F0148978FF06FA25FF4DFA01" box="[159,225,1415,1441]" captionStart="FIGURE 1" captionStartId="1.[151,250,1699,1723]" captionTargetBox="[151,1434,708,1673]" captionTargetId="figure-286@1.[151,1436,702,1673]" captionTargetPageId="1" captionText="FIGURE 1. Map showing the collection sites of the type series of Misgurnus amamianus sp. nov. A double circle indicates the locality of holotype. Double and closed circles indicate native populations, and open circle indicate an introduced population." figureDoi="http://doi.org/10.5281/zenodo.6816834" httpUri="https://zenodo.org/record/6816834/files/figure.png" pageId="8" pageNumber="533">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8BCB3687F0148978FF5EFA08FDFDF9AD" blockId="8.[151,1437,1306,2017]" pageId="8" pageNumber="533">
<emphasis id="B900EA95F0148978FF5EFA08FE11FA64" bold="true" box="[199,445,1450,1477]" pageId="8" pageNumber="533">Habitat and biology.</emphasis>
<taxonomicName id="4C744D04F0148978FE5CFA09FD6AFA64" authorityName="Nakajima &amp; Hashiguchi" authorityYear="2022" box="[453,710,1451,1476]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="amamianus">
<emphasis id="B900EA95F0148978FE5CFA09FD6AFA64" box="[453,710,1451,1476]" italics="true" pageId="8" pageNumber="533">Misgurnus amamianus</emphasis>
</taxonomicName>
inhabits the muddy bottoms of small streams, paddy fields, and ponds with rich vegetation. The life cycle is unknown, but it breeds from May to August in captivity. The loach has a strong tendency to hide in vegetation.
</paragraph>
</subSubSection>
<subSubSection id="C36E650CF0148978FF5EF9B5FC85F9F5" pageId="8" pageNumber="533" type="etymology">
<paragraph id="8BCB3687F0148978FF5EF9B5FC85F9F5" blockId="8.[151,1437,1306,2017]" pageId="8" pageNumber="533">
<emphasis id="B900EA95F0148978FF5EF9B5FEE2F991" bold="true" box="[199,334,1559,1585]" pageId="8" pageNumber="533">Etymology.</emphasis>
The specific name was derived from the Amami Islands, which is the primary distribution area of this species. Japanese name is suggested as Shinobi-dojyô.
</paragraph>
</subSubSection>
<subSubSection id="C36E650CF014897DFF57F9FDFE2BFF75" lastPageId="13" lastPageNumber="538" pageId="8" pageNumber="533" type="discussion">
<paragraph id="8BCB3687F0148978FF57F9FDFAEFF8F1" blockId="8.[151,1437,1306,2017]" pageId="8" pageNumber="533">
<emphasis id="B900EA95F0148978FF57F9FDFEEEF9D9" bold="true" box="[206,322,1631,1657]" pageId="8" pageNumber="533">Remarks.</emphasis>
There are many synonyms for 
<taxonomicName id="4C744D04F0148978FD39F9FDFC76F9D9" baseAuthorityName="Cantor" baseAuthorityYear="1842" box="[672,986,1631,1657]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="anguillicaudatus">
<emphasis id="B900EA95F0148978FD39F9FDFC76F9D9" box="[672,986,1631,1657]" italics="true" pageId="8" pageNumber="533">Misgurnus anguillicaudatus</emphasis>
</taxonomicName>
, but no species or subspecies have been described from the Ryukyu Islands, including the Amami Islands (
<bibRefCitation id="EFE54B76F0148978FC1AF921FB88F93D" author="Kottelat, M." box="[899,1060,1667,1693]" pageId="8" pageNumber="533" pagination="1 - 199" refId="ref7717" refString="Kottelat, M. (2012) Conspectus cobitidum: an inventory of the loaches of the world (Teleostei: Cypriniformes: Cobitoidei). The Raffles Bulletin of Zoology, Supplement 26, 1 - 199." type="journal article" year="2012">Kottelat 2012</bibRefCitation>
). This species is morphologically and genetically distinguished from the known species and populations distributed in Northeast Asia (
<figureCitation id="134F2A02F0148978FABCF905FADBF960" box="[1317,1399,1702,1729]" captionStart="FIGURE 6" captionStartId="6.[151,250,1663,1687]" captionTargetBox="[151,1435,181,1637]" captionTargetId="figure-25@6.[151,1436,181,1639]" captionTargetPageId="6" captionText="FIGURE 6. Maximum likelihood tree of the partial sequences of mitochondrial DNA control region (CR) in Misgurnus amamianus sp. nov. and Japanese and Northeast Asian species of Misgurnus. CR sequences of Barbatula barbatula and Leptobotia elongata are used as outgroups. Numbers on each branch indicate bootstrap values with 1,000 replications." figureDoi="http://doi.org/10.5281/zenodo.6816844" httpUri="https://zenodo.org/record/6816844/files/figure.png" pageId="8" pageNumber="533">Figs. 6</figureCitation>
, 
<figureCitation id="134F2A02F0148978FA1FF905FA3AF961" box="[1414,1430,1703,1729]" captionStart="FIGURE 7" captionStartId="7.[151,250,972,996]" captionTargetBox="[151,1436,309,948]" captionTargetId="figure-95@7.[151,1436,309,948]" captionTargetPageId="7" captionText="FIGURE 7. Lamina circularis of Northeast Asian species of Misgurnus. a, M. amamianus sp. nov. (KPM-NI 68404, holotype, Tokuno-shima Is., Japan, 83.8 mm SL); b, M. anguillicaudatus (Japan clade) (KYUM-PI 4520, Fukuoka, Japan, 93.4 mm SL); c, M. anguillicaudatus (China clade) (KMNH VR 100186, Zheijiang, China, 128.9 mm SL); d, M. dabryanus (JNC130, Fukuoka, Japan, 115.2 mm SL); e, M. mohoity (JNC563, Ibaraki, Japan, 71.7 mm SL); f, M. nikolskyi (JNC301, Tokyo, Japan, 121.3 mm SL); g, M. sp. (Clade A) (JNC345, Aomori, Japan, 110.5 mm SL); e, M. sp. OK (KMNH VR100189, Okinawa, Japan, 77.1 mm SL). Bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.6816846" httpUri="https://zenodo.org/record/6816846/files/figure.png" pageId="8" pageNumber="533">7</figureCitation>
; 
<tableCitation id="C6F6033CF0148978FF0EF969FF43F945" box="[151,239,1739,1765]" captionStart="TABLE 3" captionStartId="9.[151,239,618,644]" captionTargetPageId="9" captionText="TABLE 3. Comparison of morphological features between Misgurnus" httpUri="http://table.plazi.org/id/DF0B660FF0158979FF0EFDC8FC29FD24" pageId="8" pageNumber="533" tableUuid="DF0B660FF0158979FF0EFDC8FC29FD24">Table 3</tableCitation>
). In this study, we also compared the 
<typeStatus id="54CF8825F0148978FD30F969FD75F945" box="[681,729,1739,1765]" pageId="8" pageNumber="533">type</typeStatus>
specimens of 
<taxonomicName id="4C744D04F0148978FC1BF969FB6CF945" baseAuthorityName="Cantor" baseAuthorityYear="1842" box="[898,1216,1739,1765]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="anguillicaudatus">
<emphasis id="B900EA95F0148978FC1BF969FB6CF945" box="[898,1216,1739,1765]" italics="true" pageId="8" pageNumber="533">Misgurnus anguillicaudatus</emphasis>
</taxonomicName>
as well as the five nominal species described from 
<collectingCountry id="F3637617F0148978FD90F94DFDE5F8A9" box="[521,585,1775,1801]" name="Japan" pageId="8" pageNumber="533">Japan</collectingCountry>
and 
<collectingCountry id="F3637617F0148978FD1AF94DFC8AF8A9" box="[643,806,1775,1801]" name="Taiwan Island" pageId="8" pageNumber="533">Taiwan Island</collectingCountry>
. We confirmed that the combination of the ray number in the dorsal fin, the features of the last ray of the dorsal fin, the LPC/DPA ratio, the L/S ratio, and total vertebrae number morphologically can aid in the distinguishing of this species from 
<taxonomicName id="4C744D04F0148978FC43F895FB2CF8F0" authorityName="Nakajima &amp; Hashiguchi" authorityYear="2022" box="[986,1152,1847,1872]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="amamianus">
<emphasis id="B900EA95F0148978FC43F895FB2CF8F0" box="[986,1152,1847,1872]" italics="true" pageId="8" pageNumber="533">M. amamianus</emphasis>
</taxonomicName>
(
<figureCitation id="134F2A02F0148978FB16F895FB78F8F1" box="[1167,1236,1847,1873]" captionStart="FIGURE 8" captionStartId="12.[151,250,1150,1174]" captionTargetBox="[151,1436,309,1127]" captionTargetId="figure-84@12.[151,1436,309,1127]" captionTargetPageId="12" captionText="FIGURE 8. Type specimens of Misgurnus anguillicaudatus and five Misgurnus loaches described from Japan and Taiwan. a, BMNH 1843.7.21.15-18a (syntype of Cobitis anguillicaudata Cantor, 1842, Chusan Island [Zhoushan Island], China, 103.9 mm SL); b, BMNH 1843.7.21.15-18b (syntype of Cobitis anguillicaudata Cantor, 1842, 114.9 mm SL); c, BMNH 1843.7.21.15- 18c (syntype of Cobitis anguillicaudata Cantor, 1842, 87.0 mm SL); d, BMNH 1843.7.21.15-18d (syntype of Cobitis anguillicaudata Cantor, 1842, 91.5 mm SL); e, RMNH.PISC.2705 (lectotype of Cobitis rubripinnis Temminck &amp; Schlegel, 1846, Nagasaki, Japan, 147.5 mm SL); f, BMNH1866.5.2.26 (lectotype of Cobitichthys enalios Bleeker, 1860, Kaminoseki, Japan, 64.6 mm SL); g, BMNH1866.5.2.96 (lectotype of Cobitichthys dichachrous Bleeker, 1860, Jedo [Tokyo], Japan, 102.8 mm SL); h, BMNH1866.5.2.24 (lectotype of Cobitichthys polynema Bleeker, 1860, Jedo [Tokyo], Japan, 123.6 mm SL); i, NRM 10354 (holotype of Misgurnus anguillicaudatus formosanus Rendahl, 1936, Lake Candidius, Taiwan, 105.1 mm SL). Photos by James Maclaine &amp; Lucie Goodayle (BMNH) (a, b, c, d, f, g, h), Esther Dondorp (RMNH) (e), and Andrea Hennyey (NRM) (i)." figureDoi="http://doi.org/10.5281/zenodo.6816848" httpUri="https://zenodo.org/record/6816848/files/figure.png" pageId="8" pageNumber="533">Fig. 8</figureCitation>
; 
<tableCitation id="C6F6033CF0148978FB79F895FA9AF8F1" box="[1248,1334,1847,1873]" captionStart="TABLE 4" captionStartId="11.[151,239,547,573]" captionTargetPageId="11" captionText="TABLE 4. Comparison of morphological features between type speciemns of Misgurnus described from Japan and Taiwan, and syntypes of Misgurnus anguillicaudatus." httpUri="http://table.plazi.org/id/DF0B660FF017897BFF0EFD81F80BFD9D" pageId="8" pageNumber="533" tableUuid="DF0B660FF017897BFF0EFD81F80BFD9D">Table 4</tableCitation>
).
</paragraph>
<paragraph id="8BCB3687F014897CFF5EF8F9FEE9FEBD" blockId="8.[151,1437,1306,2017]" lastBlockId="12.[151,1436,151,285]" lastPageId="12" lastPageNumber="537" pageId="8" pageNumber="533">
There are two different mitochondrial DNA lineages in the genus 
<taxonomicName id="4C744D04F0148978FC23F8F9FB9EF8D4" authorityName="La Cepede" authorityYear="1803" box="[954,1074,1883,1908]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="533" phylum="Chordata" rank="genus">
<emphasis id="B900EA95F0148978FC23F8F9FB9EF8D4" box="[954,1074,1883,1908]" italics="true" pageId="8" pageNumber="533">Misgurnus</emphasis>
</taxonomicName>
: clades A and B (or 
<typeStatus id="54CF8825F0148978FAB8F8F9FAF5F8D5" box="[1313,1369,1883,1909]" pageId="8" pageNumber="533">Type</typeStatus>
I and 
<typeStatus id="54CF8825F0148978FF0EF8DDFF63F839" box="[151,207,1919,1945]" pageId="8" pageNumber="533">Type</typeStatus>
II) (
<bibRefCitation id="EFE54B76F0148978FF67F8DDFE5BF839" author="Morishima, K. &amp; Nakamura-Shiokawa, Y. &amp; Bando, E. &amp; Li, Y. J. &amp; Boron, A. &amp; Khan, M. M. R. &amp; Arai, K." box="[254,503,1919,1945]" pageId="8" pageNumber="533" pagination="159 - 171" refId="ref8188" refString="Morishima, K., Nakamura-Shiokawa, Y., Bando, E., Li, Y. J., Boron, A., Khan, M. M. R. &amp; Arai, K. (2008) Cryptic clonal lineages and genetic diversity in the loach Misgurnus anguillicaudatus (Teleostei: Cobitidae) inferred from nuclear and mitochondrial DNA analyses. Genetica, 132, 159 - 171. https: // doi. org / 10.1007 / s 10709 - 007 - 9158 - 1" type="journal article" year="2008">Morishima et al. 2008</bibRefCitation>
; 
<bibRefCitation id="EFE54B76F0148978FD9BF8DDFD4CF839" author="Koizumi, N. &amp; Takemura, T. &amp; Watabe, K. &amp; Mori, A." box="[514,736,1919,1945]" pageId="8" pageNumber="533" pagination="7 - 16" refId="ref7636" refString="Koizumi, N., Takemura, T., Watabe, K. &amp; Mori, A. (2009) Genetic variation and diversity of Japanese loach inferred from mitochondrial DNA - phylogenetic analysis using the cytochrome b gene sequences. Transactions of the Japanese Society of Irrigation, Drainage and Rural Engineering, 259, 7 - 16. [in Japanese with English abstract] https: // doi. org / 10.11408 / jsidre. 77.7" type="journal article" year="2009">Koizumi et al. 2009</bibRefCitation>
; 
<bibRefCitation id="EFE54B76F0148978FD72F8DDFC7DF839" author="Kitagawa, T. &amp; Fujii, Y. &amp; Koizumi, N." box="[747,977,1919,1945]" pageId="8" pageNumber="533" pagination="340 - 346" refId="ref7512" refString="Kitagawa, T., Fujii, Y. &amp; Koizumi, N. (2011) Origin of the two major distinct mtDNA clades of the Japanese population of the oriental weather loach Misgurnus anguillicaudatus (Teleostei: Cobitidae). Folia Zoologica, 60, 340 - 346. https: // doi. org / 10.25225 / fozo. v 60. i 4. a 12.2011" type="journal article" year="2011">Kitagawa et al. 2011</bibRefCitation>
; 
<bibRefCitation id="EFE54B76F0148978FC45F8DDFB13F839" author="Fujimoto, T. &amp; Yamada, A. &amp; Kodo, Y. &amp; Nakaya, K. &amp; Okubo-Murata, M. &amp; Saito, T. &amp; Ninomiya, K. &amp; Inaba, M. &amp; Kuroda, M. &amp; Arai, K. &amp; Murakami, M." box="[988,1215,1919,1945]" pageId="8" pageNumber="533" pagination="743 - 756" refId="ref7143" refString="Fujimoto, T., Yamada, A., Kodo, Y., Nakaya, K., Okubo-Murata, M., Saito, T., Ninomiya, K., Inaba, M., Kuroda, M., Arai, K. &amp; Murakami, M. (2017) Development of nuclear DNA markers to characterize genetically diverse groups of Misgurnus anguillicaudatus and its closely related species. Fisheries Science, 83, 743 - 756. https: // doi. org / 10.1007 / s 12562 - 017 - 1108 - y" type="journal article" year="2017">Fujimoto et al. 2017</bibRefCitation>
; 
<bibRefCitation id="EFE54B76F0148978FB53F8DDFA3DF839" author="Okada, R. &amp; Inui, T. &amp; Iguchi, Y. &amp; Kitagawa, T. &amp; Takata, K. &amp; Kitagawa, T." box="[1226,1425,1919,1945]" pageId="8" pageNumber="533" pagination="989 - 996" refId="ref8529" refString="Okada, R., Inui, T., Iguchi, Y., Kitagawa, T., Takata, K. &amp; Kitagawa, T. (2017) Molecular and morphological analyses revealed a cryptic species of dojo loach Misgurnus anguillicaudatus (Cypriniformes: Cobitidae) in Japan. Journal of Fish Biology, 91, 989 - 996. https: // doi. org / 10.1111 / jfb. 13393" type="journal article" year="2017">Okada et al. 2017</bibRefCitation>
). These two lineages were confirmed by phylogenetic analysis with high bootstrap values (
<figureCitation id="134F2A02F0148978FB30F801FB5DF81C" box="[1193,1265,1954,1981]" captionStart="FIGURE 6" captionStartId="6.[151,250,1663,1687]" captionTargetBox="[151,1435,181,1637]" captionTargetId="figure-25@6.[151,1436,181,1639]" captionTargetPageId="6" captionText="FIGURE 6. Maximum likelihood tree of the partial sequences of mitochondrial DNA control region (CR) in Misgurnus amamianus sp. nov. and Japanese and Northeast Asian species of Misgurnus. CR sequences of Barbatula barbatula and Leptobotia elongata are used as outgroups. Numbers on each branch indicate bootstrap values with 1,000 replications." figureDoi="http://doi.org/10.5281/zenodo.6816844" httpUri="https://zenodo.org/record/6816844/files/figure.png" pageId="8" pageNumber="533">Fig. 6</figureCitation>
). Phylogenetic analysis revealed that all 
<taxonomicName id="4C744D04F0148978FE21F865FDCCF840" authorityName="Nakajima &amp; Hashiguchi" authorityYear="2022" box="[440,608,1991,2016]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="amamianus">
<emphasis id="B900EA95F0148978FE21F865FDCCF840" box="[440,608,1991,2016]" italics="true" pageId="8" pageNumber="533">M. amamianus</emphasis>
</taxonomicName>
individuals belonged to clade B and clearly diverged from other species within this clade (
<figureCitation id="134F2A02F010897CFEF8FF35FE09FF11" box="[353,421,151,177]" captionStart="FIGURE 6" captionStartId="6.[151,250,1663,1687]" captionTargetBox="[151,1435,181,1637]" captionTargetId="figure-25@6.[151,1436,181,1639]" captionTargetPageId="6" captionText="FIGURE 6. Maximum likelihood tree of the partial sequences of mitochondrial DNA control region (CR) in Misgurnus amamianus sp. nov. and Japanese and Northeast Asian species of Misgurnus. CR sequences of Barbatula barbatula and Leptobotia elongata are used as outgroups. Numbers on each branch indicate bootstrap values with 1,000 replications." figureDoi="http://doi.org/10.5281/zenodo.6816844" httpUri="https://zenodo.org/record/6816844/files/figure.png" pageId="12" pageNumber="537">Fig. 6</figureCitation>
). The monophyly of 
<taxonomicName id="4C744D04F010897CFD0AFF35FC95FF10" authorityName="Nakajima &amp; Hashiguchi" authorityYear="2022" box="[659,825,151,176]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="537" phylum="Chordata" rank="species" species="amamianus">
<emphasis id="B900EA95F010897CFD0AFF35FC95FF10" box="[659,825,151,176]" italics="true" pageId="12" pageNumber="537">M. amamianus</emphasis>
</taxonomicName>
is supported by a 100% bootstrap value (
<figureCitation id="134F2A02F010897CFA95FF35FAFDFF11" box="[1292,1361,151,177]" captionStart="FIGURE 6" captionStartId="6.[151,250,1663,1687]" captionTargetBox="[151,1435,181,1637]" captionTargetId="figure-25@6.[151,1436,181,1639]" captionTargetPageId="6" captionText="FIGURE 6. Maximum likelihood tree of the partial sequences of mitochondrial DNA control region (CR) in Misgurnus amamianus sp. nov. and Japanese and Northeast Asian species of Misgurnus. CR sequences of Barbatula barbatula and Leptobotia elongata are used as outgroups. Numbers on each branch indicate bootstrap values with 1,000 replications." figureDoi="http://doi.org/10.5281/zenodo.6816844" httpUri="https://zenodo.org/record/6816844/files/figure.png" pageId="12" pageNumber="537">Fig. 6</figureCitation>
). Pairwise differences (i.e., p-distances) of the CR nucleotide sequences between 
<taxonomicName id="4C744D04F010897CFC67FF19FB0BFF74" authorityName="Nakajima &amp; Hashiguchi" authorityYear="2022" box="[1022,1191,187,212]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="537" phylum="Chordata" rank="species" species="amamianus">
<emphasis id="B900EA95F010897CFC67FF19FB0BFF74" box="[1022,1191,187,212]" italics="true" pageId="12" pageNumber="537">M. amamianus</emphasis>
</taxonomicName>
and other 
<taxonomicName id="4C744D04F010897CFABDFF19FF44FF59" pageId="12" pageNumber="537">
<emphasis id="B900EA95F010897CFABDFF19FA30FF74" box="[1316,1436,187,212]" italics="true" pageId="12" pageNumber="537">Misgurnus</emphasis>
species
</taxonomicName>
in clade B ranged from 6.3% to 7.8% (mean: 7.2%). These results support the validity of a distinct species, 
<taxonomicName id="4C744D04F010897CFF0EFEA1FE92FEBC" authorityName="Nakajima &amp; Hashiguchi" authorityYear="2022" box="[151,318,259,284]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="537" phylum="Chordata" rank="species" species="amamianus">
<emphasis id="B900EA95F010897CFF0EFEA1FE92FEBC" box="[151,318,259,284]" italics="true" pageId="12" pageNumber="537">M. amamianus</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="DF0B660FF0158979FF0EFDC8FC29FD24" ID-Table-UUID="DF0B660FF0158979FF0EFDC8FC29FD24" box="[151,901,618,644]" httpUri="http://table.plazi.org/id/DF0B660FF0158979FF0EFDC8FC29FD24" pageId="9" pageNumber="534" startId="9.[151,239,618,644]" targetBox="[161,2034,675,1535]" targetIsTable="true" targetPageId="9">
<paragraph id="8BCB3687F0158979FF0EFDC8FC29FD24" blockId="9.[151,901,618,644]" box="[151,901,618,644]" pageId="9" pageNumber="534">
<emphasis id="B900EA95F0158979FF0EFDC8FEA6FD24" bold="true" box="[151,266,618,644]" pageId="9" pageNumber="534">TABLE 3.</emphasis>
Comparison of morphological features between 
<taxonomicName id="4C744D04F0158979FC8AFDC8FC29FD24" authorityName="La Cepede" authorityYear="1803" box="[787,901,618,644]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="534" phylum="Chordata" rank="genus">Misgurnus</taxonomicName>
</paragraph>
</caption>
<paragraph id="8BCB3687F0158979FF38FD6DF8FCFA5F" pageId="9" pageNumber="534">
<table id="F974C427F015768FFF38FD01F85EFA5F" box="[161,2034,675,1535]" colsContinueIn="10.[161,2033,622,1584]" gridcols="13" gridrows="6" pageId="9" pageNumber="534">
<tr id="354434C5F015768FFF38FD01F85EFCB4" box="[161,2034,675,788]" gridrow="0" pageId="9" pageNumber="534">
<th id="76955DB9F015768FFF38FD01FEFAFCB4" box="[161,342,675,788]" gridcol="0" gridrow="0" pageId="9" pageNumber="534">
<emphasis id="B900EA95F0158979FF38FD6DFEEAFD48" bold="true" box="[161,326,719,744]" pageId="9" pageNumber="534">Scientific name</emphasis>
</th>
<th id="76955DB9F015768FFEF3FD01FDB0FCB4" box="[362,540,675,788]" gridcol="1" gridrow="0" pageId="9" pageNumber="534">
<emphasis id="B900EA95F0158979FEF5FD62FE5AFD57" bold="true" pageId="9" pageNumber="534">Shape of lamina circularis</emphasis>
</th>
<th id="76955DB9F015768FFDA8FD01FD3DFCB4" box="[561,657,675,788]" gridcol="2" gridrow="0" pageId="9" pageNumber="534">
<emphasis id="B900EA95F0158979FDD2FD01FDDCFCB4" bold="true" pageId="9" pageNumber="534">Ray number in dorsal fin</emphasis>
</th>
<th id="76955DB9F015768FFD30FD01FCB3FCB4" box="[681,799,675,788]" gridcol="3" gridrow="0" pageId="9" pageNumber="534">
<emphasis id="B900EA95F0158979FD30FD62FCB4FD57" bold="true" pageId="9" pageNumber="534">Last ray of dorsal fin</emphasis>
</th>
<th id="76955DB9F015768FFCA1FD01FCC0FCB4" box="[824,876,675,788]" gridcol="4" gridrow="0" pageId="9" pageNumber="534">
<emphasis id="B900EA95F0158979FCA6FD62FCC0FD57" bold="true" pageId="9" pageNumber="534">L/S ratio</emphasis>
</th>
<th id="76955DB9F015768FFC1BFD01FC40FCB4" box="[898,1004,675,788]" gridcol="5" gridrow="0" pageId="9" pageNumber="534">
<emphasis id="B900EA95F0158979FC1BFD62FC7DFD57" bold="true" pageId="9" pageNumber="534">LPC/DPA ratio</emphasis>
</th>
<th id="76955DB9F015768FFB9BFD01FBE1FCB4" box="[1026,1101,675,788]" gridcol="6" gridrow="0" pageId="9" pageNumber="534">
<emphasis id="B900EA95F0158979FB9BFD62FBE5FD57" bold="true" box="[1026,1099,704,759]" pageId="9" pageNumber="534">Barbel length</emphasis>
</th>
<th id="76955DB9F015768FFBF8FD01FB7FFCB4" box="[1121,1235,675,788]" gridcol="7" gridrow="0" pageId="9" pageNumber="534">
<emphasis id="B900EA95F0158979FBEAFD10FB15FCA5" bold="true" pageId="9" pageNumber="534">Caudal peduncle depth</emphasis>
</th>
<th id="76955DB9F015768FFB74FD01FAF0FCB4" box="[1261,1372,675,788]" gridcol="8" gridrow="0" pageId="9" pageNumber="534">
<emphasis id="B900EA95F0158979FA91FD62FAE1FD57" bold="true" pageId="9" pageNumber="534">Body pattern</emphasis>
</th>
<th id="76955DB9F015768FFAE0FD01FA4DFCB4" box="[1401,1505,675,788]" gridcol="9" gridrow="0" pageId="9" pageNumber="534">
<emphasis id="B900EA95F0158979FA08FD10FA74FCA5" bold="true" pageId="9" pageNumber="534">Total vertebrae number</emphasis>
</th>
<th id="76955DB9F015768FFA63FD01F9DFFCB4" box="[1530,1651,675,788]" gridcol="10" gridrow="0" pageId="9" pageNumber="534">
<emphasis id="B900EA95F0158979FA63FD10F9CEFCA5" bold="true" box="[1530,1651,690,773]" pageId="9" pageNumber="534">Abdominal vertebrae number</emphasis>
</th>
<th id="76955DB9F015768FF914FD01F959FCB4" box="[1677,1781,675,788]" gridcol="11" gridrow="0" pageId="9" pageNumber="534">
<emphasis id="B900EA95F0158979F903FD10F940FCA5" bold="true" pageId="9" pageNumber="534">Caudal vertebrae number</emphasis>
</th>
<th id="76955DB9F015768FF894FD01F85EFCB4" box="[1805,2034,675,788]" gridcol="12" gridrow="0" pageId="9" pageNumber="534">
<emphasis id="B900EA95F0158979F894FD6DF8D5FD48" bold="true" box="[1805,1913,719,744]" pageId="9" pageNumber="534">Reference</emphasis>
</th>
</tr>
<tr id="354434C5F015768FFF38FC90F85EFC03" box="[161,2034,818,931]" gridrow="1" pageId="9" pageNumber="534">
<th id="76955DB9F015768FFF38FC90FEFAFC03" box="[161,342,818,931]" gridcol="0" gridrow="1" pageId="9" pageNumber="534">
<taxonomicName id="4C744D04F0158979FF38FCEDFE94FCC8" authority="Nakajima &amp; Hashiguchi, 2022" authorityName="Nakajima &amp; Hashiguchi" authorityYear="2022" box="[161,312,847,872]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="534" phylum="Chordata" rank="species" species="amamianus" status="sp. nov.">
<emphasis id="B900EA95F0158979FF38FCEDFE94FCC8" box="[161,312,847,872]" italics="true" pageId="9" pageNumber="534">M. amamianus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23357EEF0158979FF38FCCEFF5CFC25" box="[161,240,876,901]" pageId="9" pageNumber="534" rank="species">sp. nov.</taxonomicNameLabel>
</th>
<td id="76955DB9F015768FFEF3FC90FDB0FC03" box="[362,540,818,931]" gridcol="1" gridrow="1" pageId="9" pageNumber="534">poleaxe, neckline at the base of lamina circularis shallow</td>
<td id="76955DB9F015768FFDA8FC90FD3DFC03" box="[561,657,818,931]" gridcol="2" gridrow="1" pageId="9" pageNumber="534">iii+6</td>
<td id="76955DB9F015768FFD30FC90FCB3FC03" box="[681,799,818,931]" gridcol="3" gridrow="1" pageId="9" pageNumber="534">not branched</td>
<td id="76955DB9F015768FFCA1FC90FCC0FC03" box="[824,876,818,931]" gridcol="4" gridrow="1" pageId="9" pageNumber="534">less 0.5</td>
<td id="76955DB9F015768FFC1BFC90FC40FC03" box="[898,1004,818,931]" gridcol="5" gridrow="1" pageId="9" pageNumber="534">over 1.1</td>
<td id="76955DB9F015768FFB9BFC90FBE1FC03" box="[1026,1101,818,931]" gridcol="6" gridrow="1" pageId="9" pageNumber="534">general</td>
<td id="76955DB9F015768FFBF8FC90FB7FFC03" box="[1121,1235,818,931]" gridcol="7" gridrow="1" pageId="9" pageNumber="534">shallow</td>
<td id="76955DB9F015768FFB74FC90FAF0FC03" box="[1261,1372,818,931]" gridcol="8" gridrow="1" pageId="9" pageNumber="534">ambiguous</td>
<td id="76955DB9F015768FFAE0FC90FA4DFC03" box="[1401,1505,818,931]" gridcol="9" gridrow="1" pageId="9" pageNumber="534">44–48</td>
<td id="76955DB9F015768FFA63FC90F9DFFC03" box="[1530,1651,818,931]" gridcol="10" gridrow="1" pageId="9" pageNumber="534">27–30</td>
<td id="76955DB9F015768FF914FC90F959FC03" box="[1677,1781,818,931]" gridcol="11" gridrow="1" pageId="9" pageNumber="534">15–19</td>
<td id="76955DB9F015768FF894FC90F85EFC03" box="[1805,2034,818,931]" gridcol="12" gridrow="1" pageId="9" pageNumber="534">present study</td>
</tr>
<tr id="354434C5F015768FFF38FC0DF85EFB80" box="[161,2034,943,1056]" gridrow="2" pageId="9" pageNumber="534">
<th id="76955DB9F015768FFF38FC0DFEFAFB80" box="[161,342,943,1056]" gridcol="0" gridrow="2" pageId="9" pageNumber="534">
<taxonomicName id="4C744D04F0158979FF38FC1CFEFAFC54" baseAuthorityName="Cantor" baseAuthorityYear="1842" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="534" phylum="Chordata" rank="species" species="anguillicaudatus">
<emphasis id="B900EA95F0158979FF38FC1CFEFAFC54" italics="true" pageId="9" pageNumber="534">M. anguillicaudatus</emphasis>
</taxonomicName>
(Japan clade)
</th>
<td id="76955DB9F015768FFEF3FC0DFDB0FB80" box="[362,540,943,1056]" gridcol="1" gridrow="2" pageId="9" pageNumber="534">poleaxe, neckline at the base of lamina circularis deep</td>
<td id="76955DB9F015768FFDA8FC0DFD3DFB80" box="[561,657,943,1056]" gridcol="2" gridrow="2" pageId="9" pageNumber="534">iii+6–7</td>
<td id="76955DB9F015768FFD30FC0DFCB3FB80" box="[681,799,943,1056]" gridcol="3" gridrow="2" pageId="9" pageNumber="534">branched</td>
<td id="76955DB9F015768FFCA1FC0DFCC0FB80" box="[824,876,943,1056]" gridcol="4" gridrow="2" pageId="9" pageNumber="534">over 0.5</td>
<td id="76955DB9F015768FFC1BFC0DFC40FB80" box="[898,1004,943,1056]" gridcol="5" gridrow="2" pageId="9" pageNumber="534">over 1.1</td>
<td id="76955DB9F015768FFB9BFC0DFBE1FB80" box="[1026,1101,943,1056]" gridcol="6" gridrow="2" pageId="9" pageNumber="534">general</td>
<td id="76955DB9F015768FFBF8FC0DFB7FFB80" box="[1121,1235,943,1056]" gridcol="7" gridrow="2" pageId="9" pageNumber="534">shallow</td>
<td id="76955DB9F015768FFB74FC0DFAF0FB80" box="[1261,1372,943,1056]" gridcol="8" gridrow="2" pageId="9" pageNumber="534">ambiguous</td>
<td id="76955DB9F015768FFAE0FC0DFA4DFB80" box="[1401,1505,943,1056]" gridcol="9" gridrow="2" pageId="9" pageNumber="534">46–47</td>
<td id="76955DB9F015768FFA63FC0DF9DFFB80" box="[1530,1651,943,1056]" gridcol="10" gridrow="2" pageId="9" pageNumber="534">28–30</td>
<td id="76955DB9F015768FF914FC0DF959FB80" box="[1677,1781,943,1056]" gridcol="11" gridrow="2" pageId="9" pageNumber="534">17–18</td>
<td id="76955DB9F015768FF894FC0DF85EFB80" box="[1805,2034,943,1056]" gridcol="12" gridrow="2" pageId="9" pageNumber="534">
<bibRefCitation id="EFE54B76F0158979F894FC6EF8FFFBA3" author="Matsui, S. &amp; Nakajima, J." pageId="9" pageNumber="534" pagination="1 - 15" refId="ref7984" refString="Matsui, S. &amp; Nakajima, J. (2020) Distribution of native and non-native lineages of Misgurnus anguillicaudatus in Osaka Prefecture, Japan, and development of morphological identification for each lineage. Bulletin of the Osaka Museum of Natural History, 74, 1 - 15. [in Japanese with English abstract]" type="journal article" year="2020">Matsui &amp; Nakajima (2020)</bibRefCitation>
; present study
</td>
</tr>
<tr id="354434C5F015768FFF38FB95F85EFB08" box="[161,2034,1079,1192]" gridrow="3" pageId="9" pageNumber="534">
<th id="76955DB9F015768FFF38FB95FEFAFB08" box="[161,342,1079,1192]" gridcol="0" gridrow="3" pageId="9" pageNumber="534">
<taxonomicName id="4C744D04F0158979FF38FBE7FEFAFBDC" baseAuthorityName="Cantor" baseAuthorityYear="1842" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="534" phylum="Chordata" rank="species" species="anguillicaudatus">
<emphasis id="B900EA95F0158979FF38FBE7FEFAFBDC" italics="true" pageId="9" pageNumber="534">M. anguillicaudatus</emphasis>
</taxonomicName>
(China clade)
</th>
<td id="76955DB9F015768FFEF3FB95FDB0FB08" box="[362,540,1079,1192]" gridcol="1" gridrow="3" pageId="9" pageNumber="534">poleaxe, neckline at the base of lamina circularis deep</td>
<td id="76955DB9F015768FFDA8FB95FD3DFB08" box="[561,657,1079,1192]" gridcol="2" gridrow="3" pageId="9" pageNumber="534">iii- iv+7–8</td>
<td id="76955DB9F015768FFD30FB95FCB3FB08" box="[681,799,1079,1192]" gridcol="3" gridrow="3" pageId="9" pageNumber="534">branched</td>
<td id="76955DB9F015768FFCA1FB95FCC0FB08" box="[824,876,1079,1192]" gridcol="4" gridrow="3" pageId="9" pageNumber="534">over 0.5</td>
<td id="76955DB9F015768FFC1BFB95FC40FB08" box="[898,1004,1079,1192]" gridcol="5" gridrow="3" pageId="9" pageNumber="534">less 1.0</td>
<td id="76955DB9F015768FFB9BFB95FBE1FB08" box="[1026,1101,1079,1192]" gridcol="6" gridrow="3" pageId="9" pageNumber="534">long</td>
<td id="76955DB9F015768FFBF8FB95FB7FFB08" box="[1121,1235,1079,1192]" gridcol="7" gridrow="3" pageId="9" pageNumber="534">moderately deep</td>
<td id="76955DB9F015768FFB74FB95FAF0FB08" box="[1261,1372,1079,1192]" gridcol="8" gridrow="3" pageId="9" pageNumber="534">ambiguous</td>
<td id="76955DB9F015768FFAE0FB95FA4DFB08" box="[1401,1505,1079,1192]" gridcol="9" gridrow="3" pageId="9" pageNumber="534">45–46</td>
<td id="76955DB9F015768FFA63FB95F9DFFB08" box="[1530,1651,1079,1192]" gridcol="10" gridrow="3" pageId="9" pageNumber="534">28–29</td>
<td id="76955DB9F015768FF914FB95F959FB08" box="[1677,1781,1079,1192]" gridcol="11" gridrow="3" pageId="9" pageNumber="534">17</td>
<td id="76955DB9F015768FF894FB95F85EFB08" box="[1805,2034,1079,1192]" gridcol="12" gridrow="3" pageId="9" pageNumber="534">
<bibRefCitation id="EFE54B76F0158979F894FBE7F8FFFBDC" author="Matsui, S. &amp; Nakajima, J." pageId="9" pageNumber="534" pagination="1 - 15" refId="ref7984" refString="Matsui, S. &amp; Nakajima, J. (2020) Distribution of native and non-native lineages of Misgurnus anguillicaudatus in Osaka Prefecture, Japan, and development of morphological identification for each lineage. Bulletin of the Osaka Museum of Natural History, 74, 1 - 15. [in Japanese with English abstract]" type="journal article" year="2020">Matsui &amp; Nakajima (2020)</bibRefCitation>
; 
<bibRefCitation id="EFE54B76F0158979F8FEFBC1F8F8FB39" author="Zhang, H. &amp; Wang, Y. X. &amp; Yang, H. L. &amp; Tan, H. M. &amp; Chen, Y. X." pageId="9" pageNumber="534" pagination="414 - 427" refId="ref9254" refString="Zhang, H., Wang, Y. X., Yang, H. L., Tan, H. M. &amp; Chen, Y. X. (2021) Taxonomic revision of Chinese species of the genera Misgurnus and Paramisguenus (Cypriniformes: Cobitidae). Acta Hydrobiologica Sinica, 45, 414 - 427. [in Chinese with English abstract] https: // doi. org / 10.7541 / 2021.2019.166" type="journal article" year="2021">Zhang et al. (2021)</bibRefCitation>
; present study
</td>
</tr>
<tr id="354434C5F015768FFF38FB41F85EFA97" box="[161,2034,1251,1335]" gridrow="4" pageId="9" pageNumber="534">
<th id="76955DB9F015768FFF38FB41FEFAFA97" box="[161,342,1251,1335]" gridcol="0" gridrow="4" pageId="9" pageNumber="534">
<taxonomicName id="4C744D04F0158979FF38FAA3FE99FABA" authorityName="Kim &amp; Pak" authorityYear="1995" box="[161,309,1281,1306]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="534" phylum="Chordata" rank="species" species="buphoensis">
<emphasis id="B900EA95F0158979FF38FAA3FE99FABA" box="[161,309,1281,1306]" italics="true" pageId="9" pageNumber="534">M. buphoensis</emphasis>
</taxonomicName>
</th>
<td id="76955DB9F015768FFEF3FB41FDB0FA97" box="[362,540,1251,1335]" gridcol="1" gridrow="4" pageId="9" pageNumber="534">spatula</td>
<td id="76955DB9F015768FFDA8FB41FD3DFA97" box="[561,657,1251,1335]" gridcol="2" gridrow="4" pageId="9" pageNumber="534">iii+6–7</td>
<td id="76955DB9F015768FFD30FB41FCB3FA97" box="[681,799,1251,1335]" gridcol="3" gridrow="4" pageId="9" pageNumber="534">branched</td>
<td id="76955DB9F015768FFCA1FB41FCC0FA97" box="[824,876,1251,1335]" gridcol="4" gridrow="4" pageId="9" pageNumber="534">over 0.5</td>
<td id="76955DB9F015768FFC1BFB41FC40FA97" box="[898,1004,1251,1335]" gridcol="5" gridrow="4" pageId="9" pageNumber="534">over 1.1</td>
<td id="76955DB9F015768FFB9BFB41FBE1FA97" box="[1026,1101,1251,1335]" gridcol="6" gridrow="4" pageId="9" pageNumber="534">general</td>
<td id="76955DB9F015768FFBF8FB41FB7FFA97" box="[1121,1235,1251,1335]" gridcol="7" gridrow="4" pageId="9" pageNumber="534">shallow</td>
<td id="76955DB9F015768FFB74FB41FAF0FA97" box="[1261,1372,1251,1335]" gridcol="8" gridrow="4" pageId="9" pageNumber="534">ambiguous</td>
<td id="76955DB9F015768FFAE0FB41FA4DFA97" box="[1401,1505,1251,1335]" gridcol="9" gridrow="4" pageId="9" pageNumber="534">50–53</td>
<td id="76955DB9F015768FFA63FB41F9DFFA97" box="[1530,1651,1251,1335]" gridcol="10" gridrow="4" pageId="9" pageNumber="534">33</td>
<td id="76955DB9F015768FF914FB41F959FA97" box="[1677,1781,1251,1335]" gridcol="11" gridrow="4" pageId="9" pageNumber="534">17–20</td>
<td id="76955DB9F015768FF894FB41F85EFA97" box="[1805,2034,1251,1335]" gridcol="12" gridrow="4" pageId="9" pageNumber="534">
<bibRefCitation id="EFE54B76F0158979F894FB41F803FB5C" author="Uchida, K." box="[1805,1967,1251,1276]" pageId="9" pageNumber="534" pagination="1 - 458" refId="ref9075" refString="Uchida, K. (1939) The fishes of Tyosen (Korea), Part I: Nematognathi, Eventognathi. Bulletin of the Fisheries Experiment Station of the Government-General of Tyosen, 6, 1 - 458. [in Japanese]" type="journal article" year="1939">Uchida (1939)</bibRefCitation>
; 
<bibRefCitation id="EFE54B76F0158979F85CFB41F831FABA" author="Kim, R. T. &amp; Pak, S. Y." pageId="9" pageNumber="534" pagination="54 - 56" refId="ref7582" refString="Kim, R. T. &amp; Pak, S. Y. (1995) A new species of a loach, Misgurnus from D. P. R. of Korea. Bulletin of the Academy of Sciences of the Democratic People's Republic of Korea, 1, 54 - 56." type="journal article" year="1995">Kim &amp; Pak (1995)</bibRefCitation>
; present study
</td>
</tr>
<tr id="354434C5F015768FFF38FAD3F85EFA5F" box="[161,2034,1393,1535]" gridrow="5" pageId="9" pageNumber="534">
<th id="76955DB9F015768FFF38FAD3FEFAFA5F" box="[161,342,1393,1535]" gridcol="0" gridrow="5" pageId="9" pageNumber="534">
<taxonomicName id="4C744D04F0158979FF38FA0EFE82FA65" baseAuthorityName="Dabry de Thiersant" baseAuthorityYear="1872" box="[161,302,1452,1477]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="534" phylum="Chordata" rank="species" species="dabryanus">
<emphasis id="B900EA95F0158979FF38FA0EFE82FA65" box="[161,302,1452,1477]" italics="true" pageId="9" pageNumber="534">M. dabryanus</emphasis>
</taxonomicName>
</th>
<td id="76955DB9F015768FFEF3FAD3FDB0FA5F" box="[362,540,1393,1535]" gridcol="1" gridrow="5" pageId="9" pageNumber="534">elongated horizontally</td>
<td id="76955DB9F015768FFDA8FAD3FD3DFA5F" box="[561,657,1393,1535]" gridcol="2" gridrow="5" pageId="9" pageNumber="534">iii- iv+6–8</td>
<td id="76955DB9F015768FFD30FAD3FCB3FA5F" box="[681,799,1393,1535]" gridcol="3" gridrow="5" pageId="9" pageNumber="534">branched</td>
<td id="76955DB9F015768FFCA1FAD3FCC0FA5F" box="[824,876,1393,1535]" gridcol="4" gridrow="5" pageId="9" pageNumber="534">over 0.5</td>
<td id="76955DB9F015768FFC1BFAD3FC40FA5F" box="[898,1004,1393,1535]" gridcol="5" gridrow="5" pageId="9" pageNumber="534">less 1.0</td>
<td id="76955DB9F015768FFB9BFAD3FBE1FA5F" box="[1026,1101,1393,1535]" gridcol="6" gridrow="5" pageId="9" pageNumber="534">long</td>
<td id="76955DB9F015768FFBF8FAD3FB7FFA5F" box="[1121,1235,1393,1535]" gridcol="7" gridrow="5" pageId="9" pageNumber="534">deep</td>
<td id="76955DB9F015768FFB74FAD3FAF0FA5F" box="[1261,1372,1393,1535]" gridcol="8" gridrow="5" pageId="9" pageNumber="534">ambiguous</td>
<td id="76955DB9F015768FFAE0FAD3FA4DFA5F" box="[1401,1505,1393,1535]" gridcol="9" gridrow="5" pageId="9" pageNumber="534">46–53</td>
<td id="76955DB9F015768FFA63FAD3F9DFFA5F" box="[1530,1651,1393,1535]" gridcol="10" gridrow="5" pageId="9" pageNumber="534">24–27</td>
<td id="76955DB9F015768FF914FAD3F959FA5F" box="[1677,1781,1393,1535]" gridcol="11" gridrow="5" pageId="9" pageNumber="534">17–19</td>
<td id="76955DB9F015768FF894FAD3F85EFA5F" box="[1805,2034,1393,1535]" gridcol="12" gridrow="5" pageId="9" pageNumber="534">
<bibRefCitation id="EFE54B76F0158979F894FAD3F842FA2A" author="Vasil'eva, E. D." box="[1805,2030,1393,1418]" pageId="9" pageNumber="534" pagination="553 - 563" refId="ref9119" refString="Vasil'eva, E. D. (2001) Loaches (genus Misgurnus, Cobitidae) of Russian Asia. I. The species composition in waters of Russia (with a description of a new species) and some nomenclature and taxonomic problems of related forms from adjacent countries. Journal of Ichthyology, 41, 553 - 563." type="journal article" year="2001">Vasil’eva (2001)</bibRefCitation>
; 
<bibRefCitation id="EFE54B76F0158979F894FA2CF8FFFA64" author="Takahashi, H. &amp; Shimizu, T." pageId="9" pageNumber="534" pagination="27 - 33" refId="ref8864" refString="Takahashi, H. &amp; Shimizu, T. (2016) Ehime-ken-de-saishu-sareta-kokugai-inyushu-karadojyo-no-shichigyo-no-keitai [Morphology of larvae and juveniles of invasive loach species, Misgurnus dabryanus, collected in Ehime Prefecture]. Nanyo Seibutsu, 18, 27 - 33. [in Japanese]" type="journal article" year="2016">Takahashi &amp; Shimizu (2016)</bibRefCitation>
; 
<bibRefCitation id="EFE54B76F0158979F83CFA09F8FFFA42" author="Hosoya, K." pageId="9" pageNumber="534" refId="ref7344" refString="Hosoya, K. (2019) Zouho-kaitei Nihon-No-Tansuigyo [Freshwater fishes of Japan Revised Edition]. Yama-Kei Publishers, Tokyo, 559 pp. [in Japanese]" type="book" year="2019">Hosoya (2019)</bibRefCitation>
; 
<bibRefCitation id="EFE54B76F0158979F8FEFA6BF8FCFA5F" author="Zhang, H. &amp; Wang, Y. X. &amp; Yang, H. L. &amp; Tan, H. M. &amp; Chen, Y. X." pageId="9" pageNumber="534" pagination="414 - 427" refId="ref9254" refString="Zhang, H., Wang, Y. X., Yang, H. L., Tan, H. M. &amp; Chen, Y. X. (2021) Taxonomic revision of Chinese species of the genera Misgurnus and Paramisguenus (Cypriniformes: Cobitidae). Acta Hydrobiologica Sinica, 45, 414 - 427. [in Chinese with English abstract] https: // doi. org / 10.7541 / 2021.2019.166" type="journal article" year="2021">Zhang et al. (2021)</bibRefCitation>
</td>
</tr>
</table>
</paragraph>
<paragraph id="8BCB3687F0158979FF38F9B6FE71F98D" blockId="9.[161,477,1556,1581]" box="[161,477,1556,1581]" pageId="9" pageNumber="534">
<tableNote id="76923709F0158979FF38F9B6FE71F98D" box="[161,477,1556,1581]" pageId="9" pageNumber="534" targetBox="[161,2034,675,1535]" targetPageId="9">......Continued on the next page</tableNote>
</paragraph>
<paragraph id="8BCB3687F016897AFF0EFD94FE20FDF0" blockId="10.[151,396,566,592]" box="[151,396,566,592]" pageId="10" pageNumber="535">
<tableNote id="76923709F016897AFF0EFD94FE20FDF0" box="[151,396,566,592]" pageId="10" pageNumber="535" targetBox="[161,2033,622,1584]" targetPageId="10">
<emphasis id="B900EA95F016897AFF0EFD94FEA6FDF0" bold="true" box="[151,266,566,592]" pageId="10" pageNumber="535">TABLE 3.</emphasis>
(continued)
</tableNote>
</paragraph>
<paragraph id="8BCB3687F016897AFF38FD38F872F9B3" pageId="10" pageNumber="535">
<table id="F974C427F016768FFF38FDCCF85DF990" box="[161,2033,622,1584]" colsContinueFrom="9.[161,2034,675,1535]" gridcols="13" gridrows="7" pageId="10" pageNumber="535">
<tr id="354434C5F016768FFF38FDCCF85DFD7F" box="[161,2033,622,735]" gridrow="0" pageId="10" pageNumber="535">
<th id="76955DB9F016768FFF38FDCCFEFAFD7F" box="[161,342,622,735]" gridcol="0" gridrow="0" pageId="10" pageNumber="535">
<emphasis id="B900EA95F016897AFF38FD38FEEAFD13" bold="true" box="[161,326,666,691]" pageId="10" pageNumber="535">Scientific name</emphasis>
</th>
<th id="76955DB9F016768FFEF3FDCCFDB0FD7F" box="[362,540,622,735]" gridcol="1" gridrow="0" pageId="10" pageNumber="535">
<emphasis id="B900EA95F016897AFEF5FD2EFE5AFD62" bold="true" pageId="10" pageNumber="535">Shape of lamina circularis</emphasis>
</th>
<th id="76955DB9F016768FFDAAFDCCFD3EFD7F" box="[563,658,622,735]" gridcol="2" gridrow="0" pageId="10" pageNumber="535">
<emphasis id="B900EA95F016897AFDD4FDCCFDDEFD7F" bold="true" pageId="10" pageNumber="535">Ray number in dorsal fin</emphasis>
</th>
<th id="76955DB9F016768FFD28FDCCFCBFFD7F" box="[689,787,622,735]" gridcol="3" gridrow="0" pageId="10" pageNumber="535">
<emphasis id="B900EA95F016897AFD2DFDDFFD5CFD71" bold="true" pageId="10" pageNumber="535">Last ray of dorsal fin</emphasis>
</th>
<th id="76955DB9F016768FFCA9FDCCFCC8FD7F" box="[816,868,622,735]" gridcol="4" gridrow="0" pageId="10" pageNumber="535">
<emphasis id="B900EA95F016897AFCAEFD2EFCC8FD62" bold="true" pageId="10" pageNumber="535">L/S ratio</emphasis>
</th>
<th id="76955DB9F016768FFCE5FDCCFC4BFD7F" box="[892,999,622,735]" gridcol="5" gridrow="0" pageId="10" pageNumber="535">
<emphasis id="B900EA95F016897AFCE5FD2EFC60FD62" bold="true" pageId="10" pageNumber="535">LPC/DPA ratio</emphasis>
</th>
<th id="76955DB9F016768FFC67FDCCFBE5FD7F" box="[1022,1097,622,735]" gridcol="6" gridrow="0" pageId="10" pageNumber="535">
<emphasis id="B900EA95F016897AFC66FD2EFBEAFD62" bold="true" box="[1023,1096,652,706]" pageId="10" pageNumber="535">Barbel length</emphasis>
</th>
<th id="76955DB9F016768FFBF9FDCCFB7EFD7F" box="[1120,1234,622,735]" gridcol="7" gridrow="0" pageId="10" pageNumber="535">
<emphasis id="B900EA95F016897AFBE8FDDFFB14FD71" bold="true" pageId="10" pageNumber="535">Caudal peduncle depth</emphasis>
</th>
<th id="76955DB9F016768FFB75FDCCFAF7FD7F" box="[1260,1371,622,735]" gridcol="8" gridrow="0" pageId="10" pageNumber="535">
<emphasis id="B900EA95F016897AFA9EFD2EFAE7FD62" bold="true" pageId="10" pageNumber="535">Body pattern</emphasis>
</th>
<th id="76955DB9F016768FFAE0FDCCFA4DFD7F" box="[1401,1505,622,735]" gridcol="9" gridrow="0" pageId="10" pageNumber="535">
<emphasis id="B900EA95F016897AFA08FDDFFA74FD71" bold="true" pageId="10" pageNumber="535">Total vertebrae number</emphasis>
</th>
<th id="76955DB9F016768FFA65FDCCF9D9FD7F" box="[1532,1653,622,735]" gridcol="10" gridrow="0" pageId="10" pageNumber="535">
<emphasis id="B900EA95F016897AFA65FDDFF9CFFD71" bold="true" box="[1532,1653,637,721]" pageId="10" pageNumber="535">Abdominal vertebrae number</emphasis>
</th>
<th id="76955DB9F016768FF913FDCCF95EFD7F" box="[1674,1778,622,735]" gridcol="11" gridrow="0" pageId="10" pageNumber="535">
<emphasis id="B900EA95F016897AF90EFDDFF946FD71" bold="true" pageId="10" pageNumber="535">Caudal vertebrae number</emphasis>
</th>
<th id="76955DB9F016768FF89FFDCCF85DFD7F" box="[1798,2033,622,735]" gridcol="12" gridrow="0" pageId="10" pageNumber="535">
<emphasis id="B900EA95F016897AF89FFD38F8DEFD13" bold="true" box="[1798,1906,666,691]" pageId="10" pageNumber="535">Reference</emphasis>
</th>
</tr>
<tr id="354434C5F016768FFF38FCA8F85DFCDB" box="[161,2033,778,891]" gridrow="1" pageId="10" pageNumber="535">
<th id="76955DB9F016768FFF38FCA8FEFAFCDB" box="[161,342,778,891]" gridcol="0" gridrow="1" pageId="10" pageNumber="535">
<taxonomicName id="4C744D04F016897AFF38FC94FEB9FCEF" baseAuthorityName="Dybowski" baseAuthorityYear="1869" box="[161,277,822,847]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="535" phylum="Chordata" rank="species" species="mohoity">
<emphasis id="B900EA95F016897AFF38FC94FEB9FCEF" box="[161,277,822,847]" italics="true" pageId="10" pageNumber="535">M. mohoity</emphasis>
</taxonomicName>
*1
</th>
<td id="76955DB9F016768FFEF3FCA8FDB0FCDB" box="[362,540,778,891]" gridcol="1" gridrow="1" pageId="10" pageNumber="535">poleaxe, neckline at the base of lamina circularis shallow</td>
<td id="76955DB9F016768FFDAAFCA8FD3EFCDB" box="[563,658,778,891]" gridcol="2" gridrow="1" pageId="10" pageNumber="535">iii-iv+6</td>
<td id="76955DB9F016768FFD28FCA8FCBFFCDB" box="[689,787,778,891]" gridcol="3" gridrow="1" pageId="10" pageNumber="535">branched</td>
<td id="76955DB9F016768FFCA9FCA8FCC8FCDB" box="[816,868,778,891]" gridcol="4" gridrow="1" pageId="10" pageNumber="535">over 0.5</td>
<td id="76955DB9F016768FFCE5FCA8FC4BFCDB" box="[892,999,778,891]" gridcol="5" gridrow="1" pageId="10" pageNumber="535">over 1.1</td>
<td id="76955DB9F016768FFC67FCA8FBE5FCDB" box="[1022,1097,778,891]" gridcol="6" gridrow="1" pageId="10" pageNumber="535">general</td>
<td id="76955DB9F016768FFBF9FCA8FB7EFCDB" box="[1120,1234,778,891]" gridcol="7" gridrow="1" pageId="10" pageNumber="535">shallow</td>
<td id="76955DB9F016768FFB75FCA8FAF7FCDB" box="[1260,1371,778,891]" gridcol="8" gridrow="1" pageId="10" pageNumber="535">ambiguous</td>
<td id="76955DB9F016768FFAE0FCA8FA4DFCDB" box="[1401,1505,778,891]" gridcol="9" gridrow="1" pageId="10" pageNumber="535">51</td>
<td id="76955DB9F016768FFA65FCA8F9D9FCDB" box="[1532,1653,778,891]" gridcol="10" gridrow="1" pageId="10" pageNumber="535">31</td>
<td id="76955DB9F016768FF913FCA8F95EFCDB" box="[1674,1778,778,891]" gridcol="11" gridrow="1" pageId="10" pageNumber="535">20</td>
<td id="76955DB9F016768FF89FFCA8F85DFCDB" box="[1798,2033,778,891]" gridcol="12" gridrow="1" pageId="10" pageNumber="535">
<bibRefCitation id="EFE54B76F016897AF89FFCBAF841FC91" author="Vasil'eva, E. D." box="[1798,2029,792,817]" pageId="10" pageNumber="535" pagination="553 - 563" refId="ref9119" refString="Vasil'eva, E. D. (2001) Loaches (genus Misgurnus, Cobitidae) of Russian Asia. I. The species composition in waters of Russia (with a description of a new species) and some nomenclature and taxonomic problems of related forms from adjacent countries. Journal of Ichthyology, 41, 553 - 563." type="journal article" year="2001">Vasil’eva (2001)</bibRefCitation>
; 
<bibRefCitation id="EFE54B76F016897AF89FFC94F844FCEF" author="Zhang, H. &amp; Wang, Y. X. &amp; Yang, H. L. &amp; Tan, H. M. &amp; Chen, Y. X." box="[1798,2024,822,847]" pageId="10" pageNumber="535" pagination="414 - 427" refId="ref9254" refString="Zhang, H., Wang, Y. X., Yang, H. L., Tan, H. M. &amp; Chen, Y. X. (2021) Taxonomic revision of Chinese species of the genera Misgurnus and Paramisguenus (Cypriniformes: Cobitidae). Acta Hydrobiologica Sinica, 45, 414 - 427. [in Chinese with English abstract] https: // doi. org / 10.7541 / 2021.2019.166" type="journal article" year="2021">Zhang et al. (2021)</bibRefCitation>
; present study
</td>
</tr>
<tr id="354434C5F016768FFF38FC1FF85DFC53" box="[161,2033,957,1011]" gridrow="2" pageId="10" pageNumber="535">
<th id="76955DB9F016768FFF38FC1FFEFAFC53" box="[161,342,957,1011]" gridcol="0" gridrow="2" pageId="10" pageNumber="535">
<taxonomicName id="4C744D04F016897AFF38FC1FFE91FC53" authorityName="Rendahl" authorityYear="1944" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="535" phylum="Chordata" rank="species" species="multimaculatus">
<emphasis id="B900EA95F016897AFF38FC1FFE91FC53" italics="true" pageId="10" pageNumber="535">M. multimaculatus</emphasis>
</taxonomicName>
</th>
<td id="76955DB9F016768FFEF3FC1FFDB0FC53" box="[362,540,957,1011]" gridcol="1" gridrow="2" pageId="10" pageNumber="535">-</td>
<td id="76955DB9F016768FFDAAFC1FFD3EFC53" box="[563,658,957,1011]" gridcol="2" gridrow="2" pageId="10" pageNumber="535">iii+6</td>
<td id="76955DB9F016768FFD28FC1FFCBFFC53" box="[689,787,957,1011]" gridcol="3" gridrow="2" pageId="10" pageNumber="535">-</td>
<td id="76955DB9F016768FFCA9FC1FFCC8FC53" box="[816,868,957,1011]" gridcol="4" gridrow="2" pageId="10" pageNumber="535">-</td>
<td id="76955DB9F016768FFCE5FC1FFC4BFC53" box="[892,999,957,1011]" gridcol="5" gridrow="2" pageId="10" pageNumber="535">over 1.1</td>
<td id="76955DB9F016768FFC67FC1FFBE5FC53" box="[1022,1097,957,1011]" gridcol="6" gridrow="2" pageId="10" pageNumber="535">general</td>
<td id="76955DB9F016768FFBF9FC1FFB7EFC53" box="[1120,1234,957,1011]" gridcol="7" gridrow="2" pageId="10" pageNumber="535">deep</td>
<td id="76955DB9F016768FFB75FC1FFAF7FC53" box="[1260,1371,957,1011]" gridcol="8" gridrow="2" pageId="10" pageNumber="535">spotted</td>
<td id="76955DB9F016768FFAE0FC1FFA4DFC53" box="[1401,1505,957,1011]" gridcol="9" gridrow="2" pageId="10" pageNumber="535">-</td>
<td id="76955DB9F016768FFA65FC1FF9D9FC53" box="[1532,1653,957,1011]" gridcol="10" gridrow="2" pageId="10" pageNumber="535">-</td>
<td id="76955DB9F016768FF913FC1FF95EFC53" box="[1674,1778,957,1011]" gridcol="11" gridrow="2" pageId="10" pageNumber="535">-</td>
<td id="76955DB9F016768FF89FFC1FF85DFC53" box="[1798,2033,957,1011]" gridcol="12" gridrow="2" pageId="10" pageNumber="535">
<bibRefCitation id="EFE54B76F016897AF89FFC6EF808FC45" author="Rendahl, H." box="[1798,1956,972,997]" pageId="10" pageNumber="535" pagination="1 - 54" refId="ref8632" refString="Rendahl, H. (1944) Einige Cobitiden von Annam und Tonkin. Goteborgs Kungliga Vetenskaps och Vitterhets Samhallas Handlingar, Series B, Matematiska och Naturvetenskapliga Skrifter, 3 (3), 1 - 54." type="journal article" year="1944">Rendahl (1944)</bibRefCitation>
</td>
</tr>
<tr id="354434C5F016768FFF38FBE5F85DFB3B" box="[161,2033,1095,1179]" gridrow="3" pageId="10" pageNumber="535">
<th id="76955DB9F016768FFF38FBE5FEFAFB3B" box="[161,342,1095,1179]" gridcol="0" gridrow="3" pageId="10" pageNumber="535">
<taxonomicName id="4C744D04F016897AFF38FBC7FEB1FBDE" authorityName="Vasil'eva" authorityYear="2001" box="[161,285,1125,1150]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="535" phylum="Chordata" rank="species" species="nikolskyi">
<emphasis id="B900EA95F016897AFF38FBC7FEB1FBDE" box="[161,285,1125,1150]" italics="true" pageId="10" pageNumber="535">M. nikolskyi</emphasis>
</taxonomicName>
*2
</th>
<td id="76955DB9F016768FFEF3FBE5FDB0FB3B" box="[362,540,1095,1179]" gridcol="1" gridrow="3" pageId="10" pageNumber="535">spatula or elongated poleaxe</td>
<td id="76955DB9F016768FFDAAFBE5FD3EFB3B" box="[563,658,1095,1179]" gridcol="2" gridrow="3" pageId="10" pageNumber="535">iii-iv+6</td>
<td id="76955DB9F016768FFD28FBE5FCBFFB3B" box="[689,787,1095,1179]" gridcol="3" gridrow="3" pageId="10" pageNumber="535">branched</td>
<td id="76955DB9F016768FFCA9FBE5FCC8FB3B" box="[816,868,1095,1179]" gridcol="4" gridrow="3" pageId="10" pageNumber="535">over 0.5</td>
<td id="76955DB9F016768FFCE5FBE5FC4BFB3B" box="[892,999,1095,1179]" gridcol="5" gridrow="3" pageId="10" pageNumber="535">over 1.1</td>
<td id="76955DB9F016768FFC67FBE5FBE5FB3B" box="[1022,1097,1095,1179]" gridcol="6" gridrow="3" pageId="10" pageNumber="535">general</td>
<td id="76955DB9F016768FFBF9FBE5FB7EFB3B" box="[1120,1234,1095,1179]" gridcol="7" gridrow="3" pageId="10" pageNumber="535">shallow</td>
<td id="76955DB9F016768FFB75FBE5FAF7FB3B" box="[1260,1371,1095,1179]" gridcol="8" gridrow="3" pageId="10" pageNumber="535">ambiguous</td>
<td id="76955DB9F016768FFAE0FBE5FA4DFB3B" box="[1401,1505,1095,1179]" gridcol="9" gridrow="3" pageId="10" pageNumber="535">49–53</td>
<td id="76955DB9F016768FFA65FBE5F9D9FB3B" box="[1532,1653,1095,1179]" gridcol="10" gridrow="3" pageId="10" pageNumber="535">33–34</td>
<td id="76955DB9F016768FF913FBE5F95EFB3B" box="[1674,1778,1095,1179]" gridcol="11" gridrow="3" pageId="10" pageNumber="535">16–18</td>
<td id="76955DB9F016768FF89FFBE5F85DFB3B" box="[1798,2033,1095,1179]" gridcol="12" gridrow="3" pageId="10" pageNumber="535">
<bibRefCitation id="EFE54B76F016897AF89FFBE5F844FBC0" author="Vasil'eva, E. D." box="[1798,2024,1095,1120]" pageId="10" pageNumber="535" pagination="553 - 563" refId="ref9119" refString="Vasil'eva, E. D. (2001) Loaches (genus Misgurnus, Cobitidae) of Russian Asia. I. The species composition in waters of Russia (with a description of a new species) and some nomenclature and taxonomic problems of related forms from adjacent countries. Journal of Ichthyology, 41, 553 - 563." type="journal article" year="2001">Vasil’eva (2001)</bibRefCitation>
; 
<bibRefCitation id="EFE54B76F016897AF89FFBC7F844FBDE" author="Zhang, H. &amp; Wang, Y. X. &amp; Yang, H. L. &amp; Tan, H. M. &amp; Chen, Y. X." box="[1798,2024,1125,1150]" pageId="10" pageNumber="535" pagination="414 - 427" refId="ref9254" refString="Zhang, H., Wang, Y. X., Yang, H. L., Tan, H. M. &amp; Chen, Y. X. (2021) Taxonomic revision of Chinese species of the genera Misgurnus and Paramisguenus (Cypriniformes: Cobitidae). Acta Hydrobiologica Sinica, 45, 414 - 427. [in Chinese with English abstract] https: // doi. org / 10.7541 / 2021.2019.166" type="journal article" year="2021">Zhang et al. (2021)</bibRefCitation>
; present study
</td>
</tr>
<tr id="354434C5F016768FFF38FAA3F85DFABA" box="[161,2033,1281,1306]" gridrow="4" pageId="10" pageNumber="535">
<th id="76955DB9F016768FFF38FAA3FEFAFABA" box="[161,342,1281,1306]" gridcol="0" gridrow="4" pageId="10" pageNumber="535">
<taxonomicName id="4C744D04F016897AFF38FAA3FE99FABA" authorityName="Rendahl" authorityYear="1937" box="[161,309,1281,1306]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="535" phylum="Chordata" rank="species" species="tonkinensis">
<emphasis id="B900EA95F016897AFF38FAA3FE99FABA" box="[161,309,1281,1306]" italics="true" pageId="10" pageNumber="535">M. tonkinensis</emphasis>
</taxonomicName>
</th>
<td id="76955DB9F016768FFEF3FAA3FDB0FABA" box="[362,540,1281,1306]" gridcol="1" gridrow="4" pageId="10" pageNumber="535">-</td>
<td id="76955DB9F016768FFDAAFAA3FD3EFABA" box="[563,658,1281,1306]" gridcol="2" gridrow="4" pageId="10" pageNumber="535">-</td>
<td id="76955DB9F016768FFD28FAA3FCBFFABA" box="[689,787,1281,1306]" gridcol="3" gridrow="4" pageId="10" pageNumber="535">-</td>
<td id="76955DB9F016768FFCA9FAA3FCC8FABA" box="[816,868,1281,1306]" gridcol="4" gridrow="4" pageId="10" pageNumber="535">-</td>
<td id="76955DB9F016768FFCE5FAA3FC4BFABA" box="[892,999,1281,1306]" gridcol="5" gridrow="4" pageId="10" pageNumber="535">less 1.0</td>
<td id="76955DB9F016768FFC67FAA3FBE5FABA" box="[1022,1097,1281,1306]" gridcol="6" gridrow="4" pageId="10" pageNumber="535">long</td>
<td id="76955DB9F016768FFBF9FAA3FB7EFABA" box="[1120,1234,1281,1306]" gridcol="7" gridrow="4" pageId="10" pageNumber="535">deep</td>
<td id="76955DB9F016768FFB75FAA3FAF7FABA" box="[1260,1371,1281,1306]" gridcol="8" gridrow="4" pageId="10" pageNumber="535">spotted</td>
<td id="76955DB9F016768FFAE0FAA3FA4DFABA" box="[1401,1505,1281,1306]" gridcol="9" gridrow="4" pageId="10" pageNumber="535">-</td>
<td id="76955DB9F016768FFA65FAA3F9D9FABA" box="[1532,1653,1281,1306]" gridcol="10" gridrow="4" pageId="10" pageNumber="535">-</td>
<td id="76955DB9F016768FF913FAA3F95EFABA" box="[1674,1778,1281,1306]" gridcol="11" gridrow="4" pageId="10" pageNumber="535">-</td>
<td id="76955DB9F016768FF89FFAA3F85DFABA" box="[1798,2033,1281,1306]" gridcol="12" gridrow="4" pageId="10" pageNumber="535">
<bibRefCitation id="EFE54B76F016897AF89FFAA3F808FABA" author="Rendahl, H." box="[1798,1956,1281,1306]" pageId="10" pageNumber="535" pagination="1 - 4" refId="ref8605" refString="Rendahl, H. (1937) Uber einen Misgurnus aus Tonkin. Arkiv for Zoologi, 29 A (12), 1 - 4." type="journal article" year="1937">Rendahl (1937)</bibRefCitation>
</td>
</tr>
<tr id="354434C5F016768FFF38FACDF85DFA05" box="[161,2033,1391,1445]" gridrow="5" pageId="10" pageNumber="535">
<th id="76955DB9F016768FFF38FACDFEFAFA05" box="[161,342,1391,1445]" gridcol="0" gridrow="5" pageId="10" pageNumber="535">
<emphasis id="B900EA95F016897AFF38FACDFF11FA28" box="[161,189,1391,1416]" italics="true" pageId="10" pageNumber="535">M.</emphasis>
sp. (Clade A)*3
</th>
<td id="76955DB9F016768FFEF3FACDFDB0FA05" box="[362,540,1391,1445]" gridcol="1" gridrow="5" pageId="10" pageNumber="535">spatula</td>
<td id="76955DB9F016768FFDAAFACDFD3EFA05" box="[563,658,1391,1445]" gridcol="2" gridrow="5" pageId="10" pageNumber="535">iii+6–7</td>
<td id="76955DB9F016768FFD28FACDFCBFFA05" box="[689,787,1391,1445]" gridcol="3" gridrow="5" pageId="10" pageNumber="535">branched</td>
<td id="76955DB9F016768FFCA9FACDFCC8FA05" box="[816,868,1391,1445]" gridcol="4" gridrow="5" pageId="10" pageNumber="535">over 0.5</td>
<td id="76955DB9F016768FFCE5FACDFC4BFA05" box="[892,999,1391,1445]" gridcol="5" gridrow="5" pageId="10" pageNumber="535">over 1.1</td>
<td id="76955DB9F016768FFC67FACDFBE5FA05" box="[1022,1097,1391,1445]" gridcol="6" gridrow="5" pageId="10" pageNumber="535">general</td>
<td id="76955DB9F016768FFBF9FACDFB7EFA05" box="[1120,1234,1391,1445]" gridcol="7" gridrow="5" pageId="10" pageNumber="535">shallow</td>
<td id="76955DB9F016768FFB75FACDFAF7FA05" box="[1260,1371,1391,1445]" gridcol="8" gridrow="5" pageId="10" pageNumber="535">ambiguous</td>
<td id="76955DB9F016768FFAE0FACDFA4DFA05" box="[1401,1505,1391,1445]" gridcol="9" gridrow="5" pageId="10" pageNumber="535">48–49</td>
<td id="76955DB9F016768FFA65FACDF9D9FA05" box="[1532,1653,1391,1445]" gridcol="10" gridrow="5" pageId="10" pageNumber="535">29–30</td>
<td id="76955DB9F016768FF913FACDF95EFA05" box="[1674,1778,1391,1445]" gridcol="11" gridrow="5" pageId="10" pageNumber="535">17–19</td>
<td id="76955DB9F016768FF89FFACDF85DFA05" box="[1798,2033,1391,1445]" gridcol="12" gridrow="5" pageId="10" pageNumber="535">
<bibRefCitation id="EFE54B76F016897AF89FFACDF8E0FA05" author="Nakajima, J. &amp; Uchiyama, R." pageId="10" pageNumber="535" refId="ref8396" refString="Nakajima, J. &amp; Uchiyama, R. (2017) Loaches of Japan - Natural History and Culture. Yama-Kei Publishers, Tokyo, 223 pp. [in Japanese]" type="book" year="2017">Nakajima &amp;Uchiyama (2017)</bibRefCitation>
; present study
</td>
</tr>
<tr id="354434C5F016768FFF38FA1DF85DF990" box="[161,2033,1471,1584]" gridrow="6" pageId="10" pageNumber="535">
<th id="76955DB9F016768FFF38FA1DFEFAF990" box="[161,342,1471,1584]" gridcol="0" gridrow="6" pageId="10" pageNumber="535">
<emphasis id="B900EA95F016897AFF38FA49FF1AF9A4" box="[161,182,1515,1540]" italics="true" pageId="10" pageNumber="535">M</emphasis>
. sp. OK*3
</th>
<td id="76955DB9F016768FFEF3FA1DFDB0F990" box="[362,540,1471,1584]" gridcol="1" gridrow="6" pageId="10" pageNumber="535">poleaxe, neckline at the base of lamina circularis deep</td>
<td id="76955DB9F016768FFDAAFA1DFD3EF990" box="[563,658,1471,1584]" gridcol="2" gridrow="6" pageId="10" pageNumber="535">iii+7–8</td>
<td id="76955DB9F016768FFD28FA1DFCBFF990" box="[689,787,1471,1584]" gridcol="3" gridrow="6" pageId="10" pageNumber="535">branched</td>
<td id="76955DB9F016768FFCA9FA1DFCC8F990" box="[816,868,1471,1584]" gridcol="4" gridrow="6" pageId="10" pageNumber="535">over 0.5</td>
<td id="76955DB9F016768FFCE5FA1DFC4BF990" box="[892,999,1471,1584]" gridcol="5" gridrow="6" pageId="10" pageNumber="535">less 1.1</td>
<td id="76955DB9F016768FFC67FA1DFBE5F990" box="[1022,1097,1471,1584]" gridcol="6" gridrow="6" pageId="10" pageNumber="535">long</td>
<td id="76955DB9F016768FFBF9FA1DFB7EF990" box="[1120,1234,1471,1584]" gridcol="7" gridrow="6" pageId="10" pageNumber="535">moderately deep</td>
<td id="76955DB9F016768FFB75FA1DFAF7F990" box="[1260,1371,1471,1584]" gridcol="8" gridrow="6" pageId="10" pageNumber="535">spotted</td>
<td id="76955DB9F016768FFAE0FA1DFA4DF990" box="[1401,1505,1471,1584]" gridcol="9" gridrow="6" pageId="10" pageNumber="535">43–45</td>
<td id="76955DB9F016768FFA65FA1DF9D9F990" box="[1532,1653,1471,1584]" gridcol="10" gridrow="6" pageId="10" pageNumber="535">27–29</td>
<td id="76955DB9F016768FF913FA1DF95EF990" box="[1674,1778,1471,1584]" gridcol="11" gridrow="6" pageId="10" pageNumber="535">16–17</td>
<td id="76955DB9F016768FF89FFA1DF85DF990" box="[1798,2033,1471,1584]" gridcol="12" gridrow="6" pageId="10" pageNumber="535">
<bibRefCitation id="EFE54B76F016897AF89FFA7EF8E0F9B3" author="Nakajima, J. &amp; Uchiyama, R." pageId="10" pageNumber="535" refId="ref8396" refString="Nakajima, J. &amp; Uchiyama, R. (2017) Loaches of Japan - Natural History and Culture. Yama-Kei Publishers, Tokyo, 223 pp. [in Japanese]" type="book" year="2017">Nakajima &amp;Uchiyama (2017)</bibRefCitation>
; present study
</td>
</tr>
</table>
</paragraph>
<paragraph id="8BCB3687F016897AFF38F9E0F8B1F9FB" blockId="10.[161,1821,1602,1627]" box="[161,1821,1602,1627]" pageId="10" pageNumber="535">
<tableNote id="76923709F016897AFF38F9E0F8B1F9FB" box="[161,1821,1602,1627]" pageId="10" pageNumber="535" targetBox="[161,2033,622,1584]" targetPageId="10">
*1, Treated as 
<taxonomicName id="4C744D04F016897AFEA2F9E0FE6CF9FB" baseAuthorityName="Sauvage &amp; Dabry de Thiersant" baseAuthorityYear="1874" box="[315,448,1602,1627]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="535" phylum="Chordata" rank="species" species="bipartitus">
<emphasis id="B900EA95F016897AFEA2F9E0FE6CF9FB" box="[315,448,1602,1627]" italics="true" pageId="10" pageNumber="535">M. bipartitus</emphasis>
</taxonomicName>
in 
<bibRefCitation id="EFE54B76F016897AFE79F9E0FD09F9FB" author="Zhang, H. &amp; Wang, Y. X. &amp; Yang, H. L. &amp; Tan, H. M. &amp; Chen, Y. X." box="[480,677,1602,1627]" pageId="10" pageNumber="535" pagination="414 - 427" refId="ref9254" refString="Zhang, H., Wang, Y. X., Yang, H. L., Tan, H. M. &amp; Chen, Y. X. (2021) Taxonomic revision of Chinese species of the genera Misgurnus and Paramisguenus (Cypriniformes: Cobitidae). Acta Hydrobiologica Sinica, 45, 414 - 427. [in Chinese with English abstract] https: // doi. org / 10.7541 / 2021.2019.166" type="journal article" year="2021">Zhang et al. (2021)</bibRefCitation>
; *2, treated as 
<taxonomicName id="4C744D04F016897AFCA5F9E0FC1CF9FB" baseAuthorityName="Dybowski" baseAuthorityYear="1869" box="[828,944,1602,1627]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="535" phylum="Chordata" rank="species" species="mohoity">
<emphasis id="B900EA95F016897AFCA5F9E0FC1CF9FB" box="[828,944,1602,1627]" italics="true" pageId="10" pageNumber="535">M. mohoity</emphasis>
</taxonomicName>
in 
<bibRefCitation id="EFE54B76F016897AFC49F9E0FB39F9FB" author="Zhang, H. &amp; Wang, Y. X. &amp; Yang, H. L. &amp; Tan, H. M. &amp; Chen, Y. X." box="[976,1173,1602,1627]" pageId="10" pageNumber="535" pagination="414 - 427" refId="ref9254" refString="Zhang, H., Wang, Y. X., Yang, H. L., Tan, H. M. &amp; Chen, Y. X. (2021) Taxonomic revision of Chinese species of the genera Misgurnus and Paramisguenus (Cypriniformes: Cobitidae). Acta Hydrobiologica Sinica, 45, 414 - 427. [in Chinese with English abstract] https: // doi. org / 10.7541 / 2021.2019.166" type="journal article" year="2021">Zhang et al. (2021)</bibRefCitation>
; *3, definitions are according to 
<bibRefCitation id="EFE54B76F016897AFA78F9E0F8B5F9FB" author="Nakajima, J. &amp; Uchiyama, R." box="[1505,1817,1602,1627]" pageId="10" pageNumber="535" refId="ref8396" refString="Nakajima, J. &amp; Uchiyama, R. (2017) Loaches of Japan - Natural History and Culture. Yama-Kei Publishers, Tokyo, 223 pp. [in Japanese]" type="book" year="2017">Nakajima &amp; Uchiyama (2017)</bibRefCitation>
.
</tableNote>
</paragraph>
<caption id="DF0B660FF017897BFF0EFD81F80BFD9D" ID-Table-UUID="DF0B660FF017897BFF0EFD81F80BFD9D" box="[151,1959,547,573]" httpUri="http://table.plazi.org/id/DF0B660FF017897BFF0EFD81F80BFD9D" pageId="11" pageNumber="536" startId="11.[151,239,547,573]" targetBox="[161,2026,586,1650]" targetIsTable="true" targetPageId="11">
<paragraph id="8BCB3687F017897BFF0EFD81F80BFD9D" blockId="11.[151,1959,547,573]" box="[151,1959,547,573]" pageId="11" pageNumber="536">
<emphasis id="B900EA95F017897BFF0EFD81FEA6FD9D" bold="true" box="[151,266,547,573]" pageId="11" pageNumber="536">TABLE 4.</emphasis>
Comparison of morphological features between type speciemns of 
<taxonomicName id="4C744D04F017897BFC42FD81FBE1FD9D" authorityName="La Cepede" authorityYear="1803" box="[987,1101,547,573]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="536" phylum="Chordata" rank="genus">
<emphasis id="B900EA95F017897BFC42FD81FBE1FD9D" box="[987,1101,547,573]" italics="true" pageId="11" pageNumber="536">Misgurnus</emphasis>
</taxonomicName>
described from Japan and Taiwan, and syntypes of 
<taxonomicName id="4C744D04F017897BF9EDFD81F80CFD9D" baseAuthorityName="Cantor" baseAuthorityYear="1842" box="[1652,1952,547,573]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="536" phylum="Chordata" rank="species" species="anguillicaudatus">
<emphasis id="B900EA95F017897BF9EDFD81F80CFD9D" box="[1652,1952,547,573]" italics="true" pageId="11" pageNumber="536">Misgurnus anguillicaudatus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BCB3687F017897BFF38FDE8F824F997" pageId="11" pageNumber="536">
<table id="F974C427F017768FFF38FDE8F846F9D2" box="[161,2026,586,1650]" gridcols="9" gridrows="12" pageId="11" pageNumber="536">
<tr id="354434C5F017768FFF38FDE8F846FDC3" box="[161,2026,586,611]" gridrow="0" pageId="11" pageNumber="536">
<th id="76955DB9F017768FFF38FDE8FD28FDC3" box="[161,644,586,611]" gridcol="0" gridrow="0" pageId="11" pageNumber="536">
<emphasis id="B900EA95F017897BFF38FDE8FEEAFDC3" bold="true" box="[161,326,586,611]" pageId="11" pageNumber="536">Scientific name</emphasis>
</th>
<th id="76955DB9F017768FFD3CFDE8FCA6FDC3" box="[677,778,586,611]" gridcol="1" gridrow="0" pageId="11" pageNumber="536">
<emphasis id="B900EA95F017897BFD3CFDE8FD76FDC3" bold="true" box="[677,730,586,611]" pageId="11" pageNumber="536">Type</emphasis>
</th>
<th id="76955DB9F017768FFCB2FDE8FC59FDC3" box="[811,1013,586,611]" gridcol="2" gridrow="0" pageId="11" pageNumber="536">
<emphasis id="B900EA95F017897BFCB2FDE8FC1AFDC3" bold="true" box="[811,950,586,611]" pageId="11" pageNumber="536">Type locality</emphasis>
</th>
<th id="76955DB9F017768FFB91FDE8FAA3FDC3" box="[1032,1295,586,611]" gridcol="3" gridrow="0" pageId="11" pageNumber="536">
<emphasis id="B900EA95F017897BFB91FDE8FB37FDC3" bold="true" box="[1032,1179,586,611]" pageId="11" pageNumber="536">Deposited no.</emphasis>
</th>
<th id="76955DB9F017768FFABFFDE8FA11FDC3" box="[1318,1469,586,611]" gridcol="4" gridrow="0" pageId="11" pageNumber="536">
<emphasis id="B900EA95F017897BFABFFDE8FA11FDC3" bold="true" box="[1318,1469,586,611]" pageId="11" pageNumber="536">Ray number</emphasis>
</th>
<th id="76955DB9F017768FFA48FDE8F9CFFDC3" box="[1489,1635,586,611]" gridcol="5" gridrow="0" pageId="11" pageNumber="536">
<emphasis id="B900EA95F017897BFA48FDE8F9CFFDC3" bold="true" box="[1489,1635,586,611]" pageId="11" pageNumber="536">Last ray of</emphasis>
</th>
<th id="76955DB9F017768FF9EFFDE8F951FDC3" box="[1654,1789,586,611]" gridcol="6" gridrow="0" pageId="11" pageNumber="536">
<emphasis id="B900EA95F017897BF9EEFDE8F951FDC3" bold="true" box="[1655,1789,586,611]" pageId="11" pageNumber="536">L P C / D PA</emphasis>
</th>
<th id="76955DB9F017768FF888FDE8F8F7FDC3" box="[1809,1883,586,611]" gridcol="7" gridrow="0" pageId="11" pageNumber="536">
<emphasis id="B900EA95F017897BF888FDE8F8F7FDC3" bold="true" box="[1809,1883,586,611]" pageId="11" pageNumber="536">L / S</emphasis>
</th>
<th id="76955DB9F017768FF8F6FDE8F846FDC3" box="[1903,2026,586,611]" gridcol="8" gridrow="0" pageId="11" pageNumber="536">
<emphasis id="B900EA95F017897BF8F6FDE8F846FDC3" bold="true" box="[1903,2026,586,611]" pageId="11" pageNumber="536">T o t a l</emphasis>
</th>
</tr>
<tr id="354434C5F017768FFF38FDCAF846FD3E" box="[161,2026,616,670]" gridrow="1" pageId="11" pageNumber="536" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<td id="76955DB9F017768FFABFFDCAFA11FD3E" box="[1318,1469,616,670]" gridcol="4" gridrow="1" pageId="11" pageNumber="536">
<emphasis id="B900EA95F017897BFABFFDCAFA06FD21" bold="true" box="[1318,1450,616,641]" pageId="11" pageNumber="536">in dorsal fin</emphasis>
</td>
<td id="76955DB9F017768FFA48FDCAF9CFFD3E" box="[1489,1635,616,670]" gridcol="5" gridrow="1" pageId="11" pageNumber="536">
<emphasis id="B900EA95F017897BFA48FDCAF995FD21" bold="true" box="[1489,1593,616,641]" pageId="11" pageNumber="536">dorsal fin</emphasis>
</td>
<td id="76955DB9F017768FF9EFFDCAF951FD3E" box="[1654,1789,616,670]" gridcol="6" gridrow="1" pageId="11" pageNumber="536">
<emphasis id="B900EA95F017897BF9EFFDCAF906FD21" bold="true" box="[1654,1706,616,641]" pageId="11" pageNumber="536">ratio</emphasis>
</td>
<td id="76955DB9F017768FF888FDCAF8F7FD3E" box="[1809,1883,616,670]" gridcol="7" gridrow="1" pageId="11" pageNumber="536">
<emphasis id="B900EA95F017897BF888FDCAF8E9FD21" bold="true" box="[1809,1861,616,641]" pageId="11" pageNumber="536">ratio</emphasis>
</td>
<td id="76955DB9F017768FF8F6FDCAF846FD3E" box="[1903,2026,616,670]" gridcol="8" gridrow="1" pageId="11" pageNumber="536">
<emphasis id="B900EA95F017897BF8F6FDCAF869FD3E" bold="true" pageId="11" pageNumber="536">v e r t e b r a e number</emphasis>
</td>
</tr>
<tr id="354434C5F017768FFF38FD0DF846FC80" box="[161,2026,687,800]" gridrow="2" pageId="11" pageNumber="536">
<th id="76955DB9F017768FFF38FD0DFD28FC80" box="[161,644,687,800]" gridcol="0" gridrow="2" pageId="11" pageNumber="536">
<taxonomicName id="4C744D04F017897BFF38FD0DFE94FD68" authority="Nakajima &amp; Hashiguchi, 2022" authorityName="Nakajima &amp; Hashiguchi" authorityYear="2022" box="[161,312,687,712]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="536" phylum="Chordata" rank="species" species="amamianus" status="sp. nov.">
<emphasis id="B900EA95F017897BFF38FD0DFE94FD68" box="[161,312,687,712]" italics="true" pageId="11" pageNumber="536">M. amamianus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23357EEF017897BFEA7FD0DFE21FD68" box="[318,397,687,712]" pageId="11" pageNumber="536" rank="species">sp. nov.</taxonomicNameLabel>
</th>
<td id="76955DB9F017768FFD3CFD0DFCA6FC80" box="[677,778,687,800]" gridcol="1" gridrow="2" pageId="11" pageNumber="536">Holotype</td>
<td id="76955DB9F017768FFCB2FD0DFC59FC80" box="[811,1013,687,800]" gridcol="2" gridrow="2" pageId="11" pageNumber="536">Tokuno-shima Island, Amami Islands, Kagoshima Prefecuture, Japan</td>
<td id="76955DB9F017768FFB91FD0DFAA3FC80" box="[1032,1295,687,800]" gridcol="3" gridrow="2" pageId="11" pageNumber="536">KPM-NI 68404</td>
<td id="76955DB9F017768FFABFFD0DFA11FC80" box="[1318,1469,687,800]" gridcol="4" gridrow="2" pageId="11" pageNumber="536">iii+6</td>
<td id="76955DB9F017768FFA48FD0DF9CFFC80" box="[1489,1635,687,800]" gridcol="5" gridrow="2" pageId="11" pageNumber="536">not branched</td>
<td id="76955DB9F017768FF9EFFD0DF951FC80" box="[1654,1789,687,800]" gridcol="6" gridrow="2" pageId="11" pageNumber="536">1.5</td>
<td id="76955DB9F017768FF888FD0DF8F7FC80" box="[1809,1883,687,800]" gridcol="7" gridrow="2" pageId="11" pageNumber="536">0.4</td>
<td id="76955DB9F017768FF8F6FD0DF846FC80" box="[1903,2026,687,800]" gridcol="8" gridrow="2" pageId="11" pageNumber="536">48</td>
</tr>
<tr id="354434C5F017768FFF38FCFBF846FCD2" box="[161,2026,857,882]" gridrow="3" pageId="11" pageNumber="536">
<th id="76955DB9F017768FFF38FCFBFD28FCD2" box="[161,644,857,882]" gridcol="0" gridrow="3" pageId="11" pageNumber="536">
<taxonomicName id="4C744D04F017897BFF38FCFBFDABFCD2" authority="(Cantor, 1842)" baseAuthorityName="Cantor" baseAuthorityYear="1842" box="[161,519,857,882]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="536" phylum="Chordata" rank="species" species="anguillicaudatus">
<emphasis id="B900EA95F017897BFF38FCFBFEC1FCD2" box="[161,365,857,882]" italics="true" pageId="11" pageNumber="536">M. anguillicaudatus</emphasis>
(Cantor, 1842)
</taxonomicName>
</th>
<td id="76955DB9F017768FFD3CFCFBFCA6FCD2" box="[677,778,857,882]" gridcol="1" gridrow="3" pageId="11" pageNumber="536">Syntypes</td>
<td id="76955DB9F017768FFCB2FCFBFC59FCD2" box="[811,1013,857,882]" gridcol="2" gridrow="3" pageId="11" pageNumber="536">Chusan Island</td>
<td id="76955DB9F017768FFB91FCFBFAA3FCD2" box="[1032,1295,857,882]" gridcol="3" gridrow="3" pageId="11" pageNumber="536">BMNH 1843.7.21.15-18a</td>
<td id="76955DB9F017768FFABFFCFBFA11FCD2" box="[1318,1469,857,882]" gridcol="4" gridrow="3" pageId="11" pageNumber="536">iii+7</td>
<td id="76955DB9F017768FFA48FCFBF9CFFCD2" box="[1489,1635,857,882]" gridcol="5" gridrow="3" pageId="11" pageNumber="536">branched</td>
<td id="76955DB9F017768FF9EFFCFBF951FCD2" box="[1654,1789,857,882]" gridcol="6" gridrow="3" pageId="11" pageNumber="536">0.6</td>
<td id="76955DB9F017768FF888FCFBF8F7FCD2" box="[1809,1883,857,882]" gridcol="7" gridrow="3" pageId="11" pageNumber="536">0.5</td>
<td id="76955DB9F017768FF8F6FCFBF846FCD2" box="[1903,2026,857,882]" gridcol="8" gridrow="3" pageId="11" pageNumber="536">46</td>
</tr>
<tr id="354434C5F017768FFF38FCD4F846FC0D" box="[161,2026,886,941]" gridrow="4" pageId="11" pageNumber="536" rowspan-0="1" rowspan-1="1">
<td id="76955DB9F017768FFCB2FCD4FC59FC0D" box="[811,1013,886,941]" gridcol="2" gridrow="4" pageId="11" pageNumber="536">[Zhoushan Island, Zhejiang Province,</td>
<td id="76955DB9F017768FFB91FCD4FAA3FC0D" box="[1032,1295,886,941]" gridcol="3" gridrow="4" pageId="11" pageNumber="536">BMNH 1843.7.21.15-18b</td>
<td id="76955DB9F017768FFABFFCD4FA11FC0D" box="[1318,1469,886,941]" gridcol="4" gridrow="4" pageId="11" pageNumber="536">iii+7</td>
<td id="76955DB9F017768FFA48FCD4F9CFFC0D" box="[1489,1635,886,941]" gridcol="5" gridrow="4" pageId="11" pageNumber="536">branched</td>
<td id="76955DB9F017768FF9EFFCD4F951FC0D" box="[1654,1789,886,941]" gridcol="6" gridrow="4" pageId="11" pageNumber="536">0.9</td>
<td id="76955DB9F017768FF888FCD4F8F7FC0D" box="[1809,1883,886,941]" gridcol="7" gridrow="4" pageId="11" pageNumber="536">0.5</td>
<td id="76955DB9F017768FF8F6FCD4F846FC0D" box="[1903,2026,886,941]" gridcol="8" gridrow="4" pageId="11" pageNumber="536">48</td>
</tr>
<tr id="354434C5F017768FFF38FC13F846FC7D" box="[161,2026,945,989]" gridrow="5" pageId="11" pageNumber="536" rowspan-0="1" rowspan-1="1">
<td id="76955DB9F017768FFCB2FC13FC59FC7D" box="[811,1013,945,989]" gridcol="2" gridrow="5" pageId="11" pageNumber="536">China]</td>
<td id="76955DB9F017768FFB91FC13FAA3FC7D" box="[1032,1295,945,989]" gridcol="3" gridrow="5" pageId="11" pageNumber="536">BMNH 1843.7.21.15-18c</td>
<td id="76955DB9F017768FFABFFC13FA11FC7D" box="[1318,1469,945,989]" gridcol="4" gridrow="5" pageId="11" pageNumber="536">iii+7</td>
<td id="76955DB9F017768FFA48FC13F9CFFC7D" box="[1489,1635,945,989]" gridcol="5" gridrow="5" pageId="11" pageNumber="536">branched</td>
<td id="76955DB9F017768FF9EFFC13F951FC7D" box="[1654,1789,945,989]" gridcol="6" gridrow="5" pageId="11" pageNumber="536">0.6</td>
<td id="76955DB9F017768FF888FC13F8F7FC7D" box="[1809,1883,945,989]" gridcol="7" gridrow="5" pageId="11" pageNumber="536">0.7</td>
<td id="76955DB9F017768FF8F6FC13F846FC7D" box="[1903,2026,945,989]" gridcol="8" gridrow="5" pageId="11" pageNumber="536">46</td>
</tr>
<tr id="354434C5F017768FFF38FC58F846FBB3" box="[161,2026,1018,1043]" gridrow="6" pageId="11" pageNumber="536" rowspan-0="1" rowspan-1="1" rowspan-2="1">
<td id="76955DB9F017768FFB91FC58FAA3FBB3" box="[1032,1295,1018,1043]" gridcol="3" gridrow="6" pageId="11" pageNumber="536">BMNH 1843.7.21.15-18d</td>
<td id="76955DB9F017768FFABFFC58FA11FBB3" box="[1318,1469,1018,1043]" gridcol="4" gridrow="6" pageId="11" pageNumber="536">iii+7</td>
<td id="76955DB9F017768FFA48FC58F9CFFBB3" box="[1489,1635,1018,1043]" gridcol="5" gridrow="6" pageId="11" pageNumber="536">branched</td>
<td id="76955DB9F017768FF9EFFC58F951FBB3" box="[1654,1789,1018,1043]" gridcol="6" gridrow="6" pageId="11" pageNumber="536">0.6</td>
<td id="76955DB9F017768FF888FC58F8F7FBB3" box="[1809,1883,1018,1043]" gridcol="7" gridrow="6" pageId="11" pageNumber="536">0.7</td>
<td id="76955DB9F017768FF8F6FC58F846FBB3" box="[1903,2026,1018,1043]" gridcol="8" gridrow="6" pageId="11" pageNumber="536">45</td>
</tr>
<tr id="354434C5F017768FFF38FB8DF846FB1E" box="[161,2026,1071,1214]" gridrow="7" pageId="11" pageNumber="536">
<th id="76955DB9F017768FFF38FB8DFD28FB1E" box="[161,644,1071,1214]" gridcol="0" gridrow="7" pageId="11" pageNumber="536">
<taxonomicName id="4C744D04F017897BFF38FB8DFDC3FBE8" authority="(Temminck &amp; Schlegel, 1846)" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1846" box="[161,623,1071,1096]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="536" phylum="Chordata" rank="species" species="rubripinnis">
<emphasis id="B900EA95F017897BFF38FB8DFE99FBE8" box="[161,309,1071,1096]" italics="true" pageId="11" pageNumber="536">M. rubripinnis</emphasis>
(Temminck &amp; Schlegel, 1846)
</taxonomicName>
</th>
<td id="76955DB9F017768FFD3CFB8DFCA6FB1E" box="[677,778,1071,1214]" gridcol="1" gridrow="7" pageId="11" pageNumber="536">Lectotype</td>
<td id="76955DB9F017768FFCB2FB8DFC59FB1E" box="[811,1013,1071,1214]" gridcol="2" gridrow="7" pageId="11" pageNumber="536">Surroundings of Nagasaki [Nagasaki Prefecture, Kyushu, Japan]</td>
<td id="76955DB9F017768FFB91FB8DFAA3FB1E" box="[1032,1295,1071,1214]" gridcol="3" gridrow="7" pageId="11" pageNumber="536">RMNH.PISC.2705</td>
<td id="76955DB9F017768FFABFFB8DFA11FB1E" box="[1318,1469,1071,1214]" gridcol="4" gridrow="7" pageId="11" pageNumber="536">iii+6</td>
<td id="76955DB9F017768FFA48FB8DF9CFFB1E" box="[1489,1635,1071,1214]" gridcol="5" gridrow="7" pageId="11" pageNumber="536">branched</td>
<td id="76955DB9F017768FF9EFFB8DF951FB1E" box="[1654,1789,1071,1214]" gridcol="6" gridrow="7" pageId="11" pageNumber="536">1.1</td>
<td id="76955DB9F017768FF888FB8DF8F7FB1E" box="[1809,1883,1071,1214]" gridcol="7" gridrow="7" pageId="11" pageNumber="536">0.6</td>
<td id="76955DB9F017768FF8F6FB8DF846FB1E" box="[1903,2026,1071,1214]" gridcol="8" gridrow="7" pageId="11" pageNumber="536">47</td>
</tr>
<tr id="354434C5F017768FFF38FB68F846FA9B" box="[161,2026,1226,1339]" gridrow="8" pageId="11" pageNumber="536">
<th id="76955DB9F017768FFF38FB68FD28FA9B" box="[161,644,1226,1339]" gridcol="0" gridrow="8" pageId="11" pageNumber="536">
<taxonomicName id="4C744D04F017897BFF38FB68FE1CFB43" authority="(Bleeker, 1860)" baseAuthorityName="Bleeker" baseAuthorityYear="1860" box="[161,432,1226,1251]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="536" phylum="Chordata" rank="species" species="enalios">
<emphasis id="B900EA95F017897BFF38FB68FEA0FB43" box="[161,268,1226,1251]" italics="true" pageId="11" pageNumber="536">M. enalios</emphasis>
(Bleeker, 1860)
</taxonomicName>
</th>
<td id="76955DB9F017768FFD3CFB68FCA6FA9B" box="[677,778,1226,1339]" gridcol="1" gridrow="8" pageId="11" pageNumber="536">Lectotype</td>
<td id="76955DB9F017768FFCB2FB68FC59FA9B" box="[811,1013,1226,1339]" gridcol="2" gridrow="8" pageId="11" pageNumber="536">Kaminoseki [Yamaguchi Prefecture., Honshu, Japan]</td>
<td id="76955DB9F017768FFB91FB68FAA3FA9B" box="[1032,1295,1226,1339]" gridcol="3" gridrow="8" pageId="11" pageNumber="536">BMNH 1866.5.2.26</td>
<td id="76955DB9F017768FFABFFB68FA11FA9B" box="[1318,1469,1226,1339]" gridcol="4" gridrow="8" pageId="11" pageNumber="536">iii+6</td>
<td id="76955DB9F017768FFA48FB68F9CFFA9B" box="[1489,1635,1226,1339]" gridcol="5" gridrow="8" pageId="11" pageNumber="536">branched</td>
<td id="76955DB9F017768FF9EFFB68F951FA9B" box="[1654,1789,1226,1339]" gridcol="6" gridrow="8" pageId="11" pageNumber="536">0.4</td>
<td id="76955DB9F017768FF888FB68F8F7FA9B" box="[1809,1883,1226,1339]" gridcol="7" gridrow="8" pageId="11" pageNumber="536">0.5</td>
<td id="76955DB9F017768FF8F6FB68F846FA9B" box="[1903,2026,1226,1339]" gridcol="8" gridrow="8" pageId="11" pageNumber="536">37</td>
</tr>
<tr id="354434C5F017768FFF38FAECF846FA02" box="[161,2026,1358,1442]" gridrow="9" pageId="11" pageNumber="536">
<th id="76955DB9F017768FFF38FAECFD28FA02" box="[161,644,1358,1442]" gridcol="0" gridrow="9" pageId="11" pageNumber="536">
<taxonomicName id="4C744D04F017897BFF38FAECFE4FFAC7" authority="(Bleeker, 1860)" baseAuthorityName="Bleeker" baseAuthorityYear="1860" box="[161,483,1358,1383]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="536" phylum="Chordata" rank="species" species="dichachrous">
<emphasis id="B900EA95F017897BFF38FAECFE93FAC7" box="[161,319,1358,1383]" italics="true" pageId="11" pageNumber="536">M. dichachrous</emphasis>
(Bleeker, 1860)
</taxonomicName>
</th>
<td id="76955DB9F017768FFD3CFAECFCA6FA02" box="[677,778,1358,1442]" gridcol="1" gridrow="9" pageId="11" pageNumber="536">Lectotype</td>
<td id="76955DB9F017768FFCB2FAECFC59FA02" box="[811,1013,1358,1442]" gridcol="2" gridrow="9" pageId="11" pageNumber="536">Jedo [Tokyo Metropolis, Honshu, Japan]</td>
<td id="76955DB9F017768FFB91FAECFAA3FA02" box="[1032,1295,1358,1442]" gridcol="3" gridrow="9" pageId="11" pageNumber="536">BMNH 1866.5.2.96</td>
<td id="76955DB9F017768FFABFFAECFA11FA02" box="[1318,1469,1358,1442]" gridcol="4" gridrow="9" pageId="11" pageNumber="536">iii+6</td>
<td id="76955DB9F017768FFA48FAECF9CFFA02" box="[1489,1635,1358,1442]" gridcol="5" gridrow="9" pageId="11" pageNumber="536">branched</td>
<td id="76955DB9F017768FF9EFFAECF951FA02" box="[1654,1789,1358,1442]" gridcol="6" gridrow="9" pageId="11" pageNumber="536">1.1</td>
<td id="76955DB9F017768FF888FAECF8F7FA02" box="[1809,1883,1358,1442]" gridcol="7" gridrow="9" pageId="11" pageNumber="536">0.6</td>
<td id="76955DB9F017768FF8F6FAECF846FA02" box="[1903,2026,1358,1442]" gridcol="8" gridrow="9" pageId="11" pageNumber="536">48</td>
</tr>
<tr id="354434C5F017768FFF38FA1AF846F9AC" box="[161,2026,1464,1548]" gridrow="10" pageId="11" pageNumber="536">
<th id="76955DB9F017768FFF38FA1AFD28F9AC" box="[161,644,1464,1548]" gridcol="0" gridrow="10" pageId="11" pageNumber="536">
<taxonomicName id="4C744D04F017897BFF38FA1AFE65FA71" authority="(Bleeker, 1860)" baseAuthorityName="Bleeker" baseAuthorityYear="1860" box="[161,457,1464,1489]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="536" phylum="Chordata" rank="species" species="polynema">
<emphasis id="B900EA95F017897BFF38FA1AFE89FA71" box="[161,293,1464,1489]" italics="true" pageId="11" pageNumber="536">M. polynema</emphasis>
(Bleeker, 1860)
</taxonomicName>
</th>
<td id="76955DB9F017768FFD3CFA1AFCA6F9AC" box="[677,778,1464,1548]" gridcol="1" gridrow="10" pageId="11" pageNumber="536">Lectotype</td>
<td id="76955DB9F017768FFCB2FA1AFC59F9AC" box="[811,1013,1464,1548]" gridcol="2" gridrow="10" pageId="11" pageNumber="536">Jedo [Tokyo Metropolis, Honshu, Japan]</td>
<td id="76955DB9F017768FFB91FA1AFAA3F9AC" box="[1032,1295,1464,1548]" gridcol="3" gridrow="10" pageId="11" pageNumber="536">BMNH 1866.5.2.24</td>
<td id="76955DB9F017768FFABFFA1AFA11F9AC" box="[1318,1469,1464,1548]" gridcol="4" gridrow="10" pageId="11" pageNumber="536">iii+6</td>
<td id="76955DB9F017768FFA48FA1AF9CFF9AC" box="[1489,1635,1464,1548]" gridcol="5" gridrow="10" pageId="11" pageNumber="536">branched</td>
<td id="76955DB9F017768FF9EFFA1AF951F9AC" box="[1654,1789,1464,1548]" gridcol="6" gridrow="10" pageId="11" pageNumber="536">1.2</td>
<td id="76955DB9F017768FF888FA1AF8F7F9AC" box="[1809,1883,1464,1548]" gridcol="7" gridrow="10" pageId="11" pageNumber="536">-</td>
<td id="76955DB9F017768FF8F6FA1AF846F9AC" box="[1903,2026,1464,1548]" gridcol="8" gridrow="10" pageId="11" pageNumber="536">48</td>
</tr>
<tr id="354434C5F017768FFF38F9BCF846F9D2" box="[161,2026,1566,1650]" gridrow="11" pageId="11" pageNumber="536">
<th id="76955DB9F017768FFF38F9BCFD28F9D2" box="[161,644,1566,1650]" gridcol="0" gridrow="11" pageId="11" pageNumber="536">
<taxonomicName id="4C744D04F017897BFF38F9BCFD28F997" authority="Rendahl, 1936" authorityName="Rendahl" authorityYear="1936" box="[161,644,1566,1591]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="536" phylum="Chordata" rank="subSpecies" species="anguillicaudatus" subSpecies="formosanus">
<emphasis id="B900EA95F017897BFF38F9BCFE45F997" box="[161,489,1566,1591]" italics="true" pageId="11" pageNumber="536">M. anguillicaudatus formosanus</emphasis>
Rendahl, 1936
</taxonomicName>
</th>
<td id="76955DB9F017768FFD3CF9BCFCA6F9D2" box="[677,778,1566,1650]" gridcol="1" gridrow="11" pageId="11" pageNumber="536">Holotype</td>
<td id="76955DB9F017768FFCB2F9BCFC59F9D2" box="[811,1013,1566,1650]" gridcol="2" gridrow="11" pageId="11" pageNumber="536">Lake Candidius [Nantou County, Taiwan]</td>
<td id="76955DB9F017768FFB91F9BCFAA3F9D2" box="[1032,1295,1566,1650]" gridcol="3" gridrow="11" pageId="11" pageNumber="536">NRM 10354</td>
<td id="76955DB9F017768FFABFF9BCFA11F9D2" box="[1318,1469,1566,1650]" gridcol="4" gridrow="11" pageId="11" pageNumber="536">iii+7</td>
<td id="76955DB9F017768FFA48F9BCF9CFF9D2" box="[1489,1635,1566,1650]" gridcol="5" gridrow="11" pageId="11" pageNumber="536">branched</td>
<td id="76955DB9F017768FF9EFF9BCF951F9D2" box="[1654,1789,1566,1650]" gridcol="6" gridrow="11" pageId="11" pageNumber="536">0.9</td>
<td id="76955DB9F017768FF888F9BCF8F7F9D2" box="[1809,1883,1566,1650]" gridcol="7" gridrow="11" pageId="11" pageNumber="536">0.5</td>
<td id="76955DB9F017768FF8F6F9BCF846F9D2" box="[1903,2026,1566,1650]" gridcol="8" gridrow="11" pageId="11" pageNumber="536">47</td>
</tr>
</table>
</paragraph>
<caption id="DF0B660FF010897CFF0EFBDCFAF0FA7A" ID-DOI="http://doi.org/10.5281/zenodo.6816848" ID-Zenodo-Dep="6816848" httpUri="https://zenodo.org/record/6816848/files/figure.png" pageId="12" pageNumber="537" startId="12.[151,250,1150,1174]" targetBox="[151,1436,309,1127]" targetPageId="12">
<paragraph id="8BCB3687F010897CFF0EFBDCFAF0FA7A" blockId="12.[151,1437,1150,1499]" pageId="12" pageNumber="537">
<emphasis id="B900EA95F010897CFF0EFBDCFEB8FB36" bold="true" box="[151,276,1150,1174]" pageId="12" pageNumber="537">FIGURE 8.</emphasis>
Type specimens of 
<taxonomicName id="4C744D04F010897CFE78FBDDFD53FB36" baseAuthorityName="Cantor" baseAuthorityYear="1842" box="[481,767,1150,1174]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="537" phylum="Chordata" rank="species" species="anguillicaudatus">
<emphasis id="B900EA95F010897CFE78FBDDFD53FB36" box="[481,767,1150,1174]" italics="true" pageId="12" pageNumber="537">Misgurnus anguillicaudatus</emphasis>
</taxonomicName>
and five 
<taxonomicName id="4C744D04F010897CFCF9FBDDFC60FB36" authorityName="La Cepede" authorityYear="1803" box="[864,972,1151,1174]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="537" phylum="Chordata" rank="genus">
<emphasis id="B900EA95F010897CFCF9FBDDFC60FB36" box="[864,972,1151,1174]" italics="true" pageId="12" pageNumber="537">Misgurnus</emphasis>
</taxonomicName>
loaches described from Japan and Taiwan. a, BMNH 1843.7.21.15-18a (syntype of 
<taxonomicName id="4C744D04F010897CFD8FFB00FC22FB1B" authority="Cantor, 1842" authorityName="Cantor" authorityYear="1842" box="[534,910,1186,1211]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="537" phylum="Chordata" rank="species" species="anguillicaudata">
<emphasis id="B900EA95F010897CFD8FFB00FCA8FB1A" box="[534,772,1186,1210]" italics="true" pageId="12" pageNumber="537">Cobitis anguillicaudata</emphasis>
Cantor, 1842
</taxonomicName>
, Chusan Island [Zhoushan Island], China, 103.9 mm SL); b, BMNH 1843.7.21.15-18b (syntype of 
<taxonomicName id="4C744D04F010897CFDEBFB64FC5EFB7F" authority="Cantor, 1842" authorityName="Cantor" authorityYear="1842" box="[626,1010,1222,1247]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="537" phylum="Chordata" rank="species" species="anguillicaudata">
<emphasis id="B900EA95F010897CFDEBFB64FCCFFB7E" box="[626,867,1222,1246]" italics="true" pageId="12" pageNumber="537">Cobitis anguillicaudata</emphasis>
Cantor, 1842
</taxonomicName>
, 114.9 mm SL); c, BMNH 1843.7.21.15- 18c (syntype of 
<taxonomicName id="4C744D04F010897CFEA7FB48FD11FAA3" authority="Cantor, 1842" authorityName="Cantor" authorityYear="1842" box="[318,701,1258,1283]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="537" phylum="Chordata" rank="species" species="anguillicaudata">
<emphasis id="B900EA95F010897CFEA7FB48FD83FAA2" box="[318,559,1258,1282]" italics="true" pageId="12" pageNumber="537">Cobitis anguillicaudata</emphasis>
Cantor, 1842
</taxonomicName>
, 87.0 mm SL); d, BMNH 1843.7.21.15-18d (syntype of 
<taxonomicName id="4C744D04F010897CFA9AFB48FE24FA87" authority="Cantor, 1842" authorityName="Cantor" authorityYear="1842" class="Actinopterygii" family="Cobitidae" genus="Cobitis" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="537" phylum="Chordata" rank="species" species="anguillicaudata">
<emphasis id="B900EA95F010897CFA9AFB48FF5BFA86" italics="true" pageId="12" pageNumber="537">Cobitis anguillicaudata</emphasis>
Cantor, 1842
</taxonomicName>
, 91.5 mm SL); e, RMNH.PISC.2705 (lectotype of 
<taxonomicName id="4C744D04F010897CFC06FAACFA34FA86" authority="Temminck &amp; Schlegel, 1846" authorityName="Temminck &amp; Schlegel" authorityYear="1846" box="[927,1432,1294,1319]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="537" phylum="Chordata" rank="species" species="rubripinnis">
<emphasis id="B900EA95F010897CFC06FAACFBCFFA86" box="[927,1123,1294,1318]" italics="true" pageId="12" pageNumber="537">Cobitis rubripinnis</emphasis>
Temminck &amp; Schlegel, 1846
</taxonomicName>
, Nagasaki, Japan, 147.5 mm SL); f, BMNH1866.5.2.26 (lectotype of 
<taxonomicName id="4C744D04F010897CFCC7FA90FB64FAEA" authority="Bleeker, 1860" authorityName="Bleeker" authorityYear="1860" box="[862,1224,1330,1354]" class="Actinopterygii" family="Cobitidae" genus="Cobitichthys" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="537" phylum="Chordata" rank="species" species="enalios">
<emphasis id="B900EA95F010897CFCC7FA90FB83FAEA" box="[862,1071,1330,1354]" italics="true" pageId="12" pageNumber="537">Cobitichthys enalios</emphasis>
Bleeker, 1860
</taxonomicName>
, Kaminoseki, Japan, 64.6 mm SL); g, BMNH1866.5.2.96 (lectotype of 
<taxonomicName id="4C744D04F010897CFD16FAF4FB89FACE" authority="Bleeker, 1860" authorityName="Bleeker" authorityYear="1860" box="[655,1061,1366,1390]" class="Actinopterygii" family="Cobitidae" genus="Cobitichthys" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="537" phylum="Chordata" rank="species" species="dichachrous">
<emphasis id="B900EA95F010897CFD16FAF4FC3DFACE" box="[655,913,1366,1390]" italics="true" pageId="12" pageNumber="537">Cobitichthys dichachrous</emphasis>
Bleeker, 1860
</taxonomicName>
, Jedo [Tokyo], Japan, 102.8 mm SL); h, BMNH1866.5.2.24 (lectotype of 
<taxonomicName id="4C744D04F010897CFD9AFAD8FC2FFA32" authority="Bleeker, 1860" authorityName="Bleeker" authorityYear="1860" box="[515,899,1402,1426]" class="Actinopterygii" family="Cobitidae" genus="Cobitichthys" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="537" phylum="Chordata" rank="species" species="polynema">
<emphasis id="B900EA95F010897CFD9AFAD8FD40FA32" box="[515,748,1402,1426]" italics="true" pageId="12" pageNumber="537">Cobitichthys polynema</emphasis>
Bleeker, 1860
</taxonomicName>
, Jedo [Tokyo], Japan, 123.6 mm SL); i, NRM 10354 (holotype of 
<taxonomicName id="4C744D04F010897CFE80FA3DFCE0FA16" authority="Rendahl, 1936" authorityName="Rendahl" authorityYear="1936" box="[281,844,1438,1462]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="537" phylum="Chordata" rank="subSpecies" species="anguillicaudatus" subSpecies="formosanus">
<emphasis id="B900EA95F010897CFE80FA3DFD1DFA16" box="[281,689,1438,1462]" italics="true" pageId="12" pageNumber="537">Misgurnus anguillicaudatus formosanus</emphasis>
Rendahl, 1936
</taxonomicName>
, Lake Candidius, Taiwan, 105.1 mm SL). Photos by James Maclaine &amp; Lucie Goodayle (BMNH) (a, b, c, d, f, g, h), Esther Dondorp (RMNH) (e), and Andrea Hennyey (NRM) (i).
</paragraph>
</caption>
<paragraph id="8BCB3687F010897CFF56F9ABFD8BF977" blockId="12.[151,1437,1545,2039]" pageId="12" pageNumber="537">
Many amphibian and reptile species are known to have evolved in this way in the central part of the Ryukyu Islands, including the Amami Islands (e.g., 
<bibRefCitation id="EFE54B76F010897CFDE4F98FFC88F9E7" author="Lin, S. M. &amp; Chen, C. A. &amp; Lue, K. Y." box="[637,804,1581,1607]" pageId="12" pageNumber="537" pagination="276 - 288" refId="ref7847" refString="Lin, S. M., Chen, C. A. &amp; Lue, K. Y. (2002) Molecular phylogeny and biogeography of the grass lizards genus Takydromus (Reptilia: Lacertidae) of East Asia. Molecular Phylogenetics and Evolution, 22, 276 - 288. https: // doi. org / 10.1006 / mpev. 2001.1059" type="journal article" year="2002">Lin et al. 2002</bibRefCitation>
; 
<bibRefCitation id="EFE54B76F010897CFCB6F98FFC50F9E7" author="Matsui, M. &amp; Shimada, T. &amp; Ota, H. &amp; Tanaka-Ueno, T." box="[815,1020,1580,1607]" pageId="12" pageNumber="537" pagination="733 - 742" refId="ref7912" refString="Matsui, M., Shimada, T., Ota, H. &amp; Tanaka-Ueno, T. (2005) Multiple invasions of the Ryukyu Archipelago by Oriental frogs of the subgenus Odorrana with phylogenetic reassessment of the related subgenera of the genus Rana. Molecular Phylogenetics and Evolution, 37, 733 - 742. https: // doi. org / 10.1016 / j. ympev. 2005.04.030" type="journal article" year="2005">Matsui et al. 2005</bibRefCitation>
; 
<bibRefCitation id="EFE54B76F010897CFB9EF98FFB5BF9E7" author="Tominaga, A. &amp; Matsui, M. &amp; Yoshikawa, N. &amp; Nishikawa, K. &amp; Hayashi, T. &amp; Misawa, Y. &amp; Tanabe, S. &amp; Ota, H." box="[1031,1271,1581,1607]" pageId="12" pageNumber="537" pagination="654 - 667" refId="ref8985" refString="Tominaga, A., Matsui, M., Yoshikawa, N., Nishikawa, K., Hayashi, T., Misawa, Y., Tanabe, S. &amp; Ota, H. (2013) Phylogeny and historical demography of Cynops pyrrhogaster (Amphibia: Urodela): taxonomic relationships and distributional changes associated with climatic oscillations. Molecular Phylogenetics and Evolution, 66, 654 - 667. https: // doi. org / 10.1016 / j. ympev. 2012.10.015" type="journal article" year="2013">Tominaga et al. 2013</bibRefCitation>
), and the same has been reported for freshwater fishes such as the Asian swamp eel, 
<taxonomicName id="4C744D04F010897CFC0BF9F3FA82F9CB" authority="(Matsumoto et al. 2010)" baseAuthorityName="Matsumoto" baseAuthorityYear="2010" box="[914,1326,1617,1643]" class="Actinopterygii" family="Synbranchidae" genus="Monopterus" kingdom="Animalia" order="Synbranchiformes" pageId="12" pageNumber="537" phylum="Chordata" rank="genus">
<emphasis id="B900EA95F010897CFC0BF9F3FBB5F9CA" box="[914,1049,1617,1642]" italics="true" pageId="12" pageNumber="537">Monopterus</emphasis>
(
<bibRefCitation id="EFE54B76F010897CFBBFF9F3FA8AF9CB" author="Matsumoto, S. &amp; Kon, T. &amp; Yamaguchi, M. &amp; Takeshima, H. &amp; Yamazaki, Y. &amp; Mukai, T. &amp; Kuriiwa, K. &amp; Kohda, M. &amp; Nishida, M." box="[1062,1318,1617,1643]" pageId="12" pageNumber="537" pagination="71 - 77" refId="ref8042" refString="Matsumoto, S., Kon, T., Yamaguchi, M., Takeshima, H., Yamazaki, Y., Mukai, T., Kuriiwa, K., Kohda, M. &amp; Nishida, M. (2010) Cryptic diversification of the swamp eel Monopterus albus in East and Southeast Asia, with special reference to the Ryukyuan populations. Ichthyological Research, 57, 71 - 77. https: // doi. org / 10.1007 / s 10228 - 009 - 0125 - y" type="journal article" year="2010">Matsumoto et al. 2010</bibRefCitation>
)
</taxonomicName>
. Our phylogenetic analysis suggests that 
<taxonomicName id="4C744D04F010897CFE63F9D7FD0CF92E" authorityName="Nakajima &amp; Hashiguchi" authorityYear="2022" box="[506,672,1653,1678]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="537" phylum="Chordata" rank="species" species="amamianus">
<emphasis id="B900EA95F010897CFE63F9D7FD0CF92E" box="[506,672,1653,1678]" italics="true" pageId="12" pageNumber="537">M. amamianus</emphasis>
</taxonomicName>
is also an endemic species that was isolated when the Amani Islands were separated from the Asian continent. Further detailed molecular phylogenetic analyses will help to clarify the evolutionary origins of this species.
</paragraph>
<paragraph id="8BCB3687F010897DFF49F943FE2BFF75" blockId="12.[151,1437,1545,2039]" lastBlockId="13.[151,1436,151,213]" lastPageId="13" lastPageNumber="538" pageId="12" pageNumber="537">
<taxonomicName id="4C744D04F010897CFF49F943FE7DF95A" authorityName="Nakajima &amp; Hashiguchi" authorityYear="2022" box="[208,465,1761,1786]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="537" phylum="Chordata" rank="species" species="amamianus">
<emphasis id="B900EA95F010897CFF49F943FE7DF95A" box="[208,465,1761,1786]" italics="true" pageId="12" pageNumber="537">Misgurnus amamianus</emphasis>
</taxonomicName>
is listed as Data Deficient (DD) in the Japanese Red List under the name 
<taxonomicName id="4C744D04F010897CFABDF943FF1BF8BF" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="537" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B900EA95F010897CFABDF943FA30F95A" box="[1316,1436,1761,1786]" italics="true" pageId="12" pageNumber="537">Misgurnus</emphasis>
sp.
</taxonomicName>
IR (
<bibRefCitation id="EFE54B76F010897CFF70F8A7FDDDF8BF" author="Ministry of the Environment" box="[233,625,1797,1823]" pageId="12" pageNumber="537" refId="ref8137" refString="Ministry of the Environment (2020) Red List 2020. Ministry of the Environment, Government of Japan, Tokyo. Available from: https: // www. env. go. jp / press / files / jp / 114457. pdf (accessed 12 January 2022)" type="book" year="2020">Ministry of the Environment 2020</bibRefCitation>
). Our survey showed that the last collection on Kikai-jima Island was in the 1930s, and the last collection on Amami-oshima Island was in the 1970s; the species has not been rediscovered since then. In addition, a pond that was the only habitat in Okinoerabu-jima Island (the collection site of the specimen used as the 
<typeStatus id="54CF8825F010897CFECEF8D3FE6EF82B" box="[343,450,1905,1931]" pageId="12" pageNumber="537" type="paratype">paratypes</typeStatus>
in this study) underwent renovation in the spring of 2021; the species may already be extinct. Therefore, there are only three definite habitats for this species: two ponds on Tokuno-shima Island and one paddy field on Iriomote-jima Island. In the Amami Islands, paddy fields which are the main habitat of this species, have been largely lost because of the rapid conversion from the once-common paddy rice cultivation to sugarcane cultivation (
<bibRefCitation id="EFE54B76F011897DFE87FF35FE66FF11" author="Hagihara, S." box="[286,458,151,177]" pageId="13" pageNumber="538" pagination="35 - 45" refId="ref7245" refString="Hagihara, S. (1992) Amami-nougyou-no-henbou [The transformation of agriculture in Amami]. Japanese Journal of Farm Management, 30, 35 - 45. [in Japanese]" type="journal article" year="1992">Hagihara 1992</bibRefCitation>
). We believe that 
<taxonomicName id="4C744D04F011897DFD0DFF35FC97FF10" authorityName="Nakajima &amp; Hashiguchi" authorityYear="2022" box="[660,827,151,176]" class="Actinopterygii" family="Cobitidae" genus="Misgurnus" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="538" phylum="Chordata" rank="species" species="amamianus">
<emphasis id="B900EA95F011897DFD0DFF35FC97FF10" box="[660,827,151,176]" italics="true" pageId="13" pageNumber="538">M. amamianus</emphasis>
</taxonomicName>
is on the verge of extinction, and urgent conservation measures are needed.
</paragraph>
</subSubSection>
</treatment>
</document>