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<document ID-DOI="http://doi.org/10.5281/zenodo.4758513" ID-GBIF-Dataset="f0f8f265-10f1-448f-8b15-6c56fc140166" ID-GBIF-Taxon="182067791" ID-ISSN="1854-0392" ID-Zenodo-Dep="4758513" ID-ZooBank="8BFBE3F7-E0EF-4EDD-BA82-9EA087F74688" approvalRequired="2" approvalRequired_for_textStreams="2" checkinTime="1620834297829" checkinUser="felipe" docAuthor="Grubbs, Scott A." docDate="2015" docId="03DD8784FF8AA70AFEC45EC765D5FE83" docLanguage="en" docName="Illiesia.11.12.147-166.pdf" docOrigin="Illiesia 11 (12)" docStyle="DocumentStyle:3ED4ECBC8873166DC296BE619562C54F.3:Illiesia.2009-.journal_article" docStyleId="3ED4ECBC8873166DC296BE619562C54F" docStyleName="Illiesia.2009-.journal_article" docStyleVersion="3" docTitle="Leuctra variabilis Hanson 1941" docType="treatment" docVersion="6" lastPageNumber="165" masterDocId="FFE4FFFCFF98A718FFF65E23604AFFEB" masterDocTitle="Leuctra Schusteri, A New Stonefly Species (Plecoptera: Leuctridae) Of The L. Tenuis (Pictet) Group From The Southeastern Usa" masterLastPageNumber="166" masterPageNumber="147" pageId="18" pageNumber="165" updateTime="1632012907386" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Leuctra Schusteri, A New Stonefly Species (Plecoptera: Leuctridae) Of The L. Tenuis (Pictet) Group From The Southeastern Usa</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Grubbs, Scott A.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Illiesia</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2015</mods:date>
<mods:detail type="volume">
<mods:number>11</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>12</mods:number>
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<mods:extent unit="page">
<mods:start>147</mods:start>
<mods:end>166</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.4752768</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4758513" ID-GBIF-Taxon="182067791" ID-Zenodo-Dep="4758513" LSID="urn:lsid:plazi:treatment:03DD8784FF8AA70AFEC45EC765D5FE83" httpUri="http://treatment.plazi.org/id/03DD8784FF8AA70AFEC45EC765D5FE83" lastPageNumber="165" pageId="18" pageNumber="165">
<subSubSection box="[306,681,227,252]" pageId="18" pageNumber="165" type="nomenclature">
<paragraph blockId="18.[306,681,227,252]" box="[306,681,227,252]" pageId="18" pageNumber="165">
<heading bold="true" box="[306,681,227,252]" centered="true" fontSize="10" level="2" pageId="18" pageNumber="165" reason="2">
<taxonomicName authority="Hanson 1941" authorityName="Hanson" authorityYear="1941" box="[306,681,227,252]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="18" pageNumber="165" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis bold="true" box="[306,681,227,252]" pageId="18" pageNumber="165">
<emphasis bold="true" box="[306,520,227,251]" italics="true" pageId="18" pageNumber="165">Leuctra variabilis</emphasis>
<bibRefCitation author="Hanson, J. F." box="[526,681,227,252]" pageId="18" pageNumber="165" pagination="57 - 66" refId="ref9332" refString="Hanson, J. F. 1941. Studies on the Plecoptera of North America, II. Bulletin of the Brooklyn Entomological Society, 36: 57 - 66." type="journal article" year="1941">Hanson 1941</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[382,604,264,288]" pageId="18" pageNumber="165" type="vernacular_names">
<paragraph blockId="18.[382,604,264,288]" box="[382,604,264,288]" pageId="18" pageNumber="165">Variable Needlefly</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="165" type="multiple">
<paragraph blockId="18.[189,790,332,428]" pageId="18" pageNumber="165">
http://lsid.speciesfile.org/urn:lsid:
<taxonomicName box="[579,710,332,356]" class="Insecta" kingdom="Animalia" order="Plecoptera" pageId="18" pageNumber="165" phylum="Arthropoda" rank="order">Plecoptera</taxonomicName>
.specie sfile.org:TaxonName:460700
</paragraph>
</subSubSection>
<subSubSection box="[420,567,404,428]" pageId="18" pageNumber="165" type="description">
<paragraph blockId="18.[189,790,332,428]" box="[420,567,404,428]" pageId="18" pageNumber="165">
(
<figureCitation box="[428,558,404,428]" captionStart="Figs" captionStartId="16.[189,241,1664,1688]" captionTargetBox="[249,1383,290,1581]" captionTargetId="figure-100@16.[249,1383,290,1581]" captionTargetPageId="16" captionText="Figs. 4852. Leuctra variabilis, Tennessee, Twin Springs. 48, 7th abdominal process, dorsal view, 350X; 49, paraprocts, caudal view, 350X; 50, paraprocts, caudal view, 1500X; 51, paraprocts, lateral view, 350X; 52, L. variabilis, paraprocts, lateral view, 1500X. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752794" httpUri="https://zenodo.org/record/4752794/files/figure.png" pageId="18" pageNumber="165">Figs. 4852</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="165" type="reference_group">
<paragraph blockId="18.[227,802,471,558]" pageId="18" pageNumber="165">
<treatmentCitationGroup pageId="18" pageNumber="165">
<treatmentCitation author="Hanson, J. F." box="[227,631,472,494]" page="62" pageId="18" pageNumber="165" year="1941">
<taxonomicName authority="Hanson 1941: 62" authorityName="Hanson" authorityPageNumber="62" authorityYear="1941" box="[227,600,472,494]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="18" pageNumber="165" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis box="[227,416,472,493]" italics="true" pageId="18" pageNumber="165">Leuctra variabilis</emphasis>
<bibRefCitation author="Hanson, J. F." box="[441,631,472,494]" pageId="18" pageNumber="165" pagination="57 - 66" refId="ref9332" refString="Hanson, J. F. 1941. Studies on the Plecoptera of North America, II. Bulletin of the Brooklyn Entomological Society, 36: 57 - 66." type="journal article" year="1941">Hanson 1941:62</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<materialsCitation ID-GBIF-Occurrence="3111876303" collectionCode="USNM" location="Paradise Trail" pageId="18" pageNumber="165" specimenCount="1" specimenCount-male="1" stateProvince="Sunderland" typeStatus="holotype">
<typeStatus box="[659,759,472,494]" pageId="18" pageNumber="165">Holotype</typeStatus>
<specimenCount box="[784,802,471,494]" pageId="18" pageNumber="165" type="male"></specimenCount>
(
<collectionCode box="[234,312,504,526]" pageId="18" pageNumber="165">USNM</collectionCode>
),
<location LSID="urn:lsid:plazi:treatment:03DD8784FF8AA70AFEC45EC765D5FE83:8EAB6049FF8AA70AFEBF5FDB6195FDE5" box="[329,479,504,526]" name="Paradise Trail" pageId="18" pageNumber="165" stateProvince="Sunderland">Paradise Trail</location>
,
<collectingRegion box="[489,615,504,526]" country="United Kingdom" name="Sunderland" pageId="18" pageNumber="165">Sunderland</collectingRegion>
,
<collectingRegion box="[625,778,504,526]" country="United States of America" name="Massachusetts" pageId="18" pageNumber="165">Massachusetts</collectingRegion>
</materialsCitation>
<taxonomicName authority=": Illies 1966: 111" authorityName=": Illies" authorityPageNumber="111" authorityYear="1966" box="[227,397,536,558]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="18" pageNumber="165" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis box="[227,397,536,558]" italics="true" pageId="18" pageNumber="165">Leuctra variabilis</emphasis>
</taxonomicName>
:
<treatmentCitation author="Illies, J." box="[409,555,536,558]" page="111" pageId="18" pageNumber="165" year="1966">
<bibRefCitation author="Illies, J." box="[409,555,536,558]" pageId="18" pageNumber="165" refId="ref9520" refString="Illies, J. 1966. Katalog der rezenten Plecoptera. Das Tierreich 82. Walter de Gruyter and Company, Berlin. 632 pp." type="book" year="1966">Illies 1966:111</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="165" type="description">
<paragraph blockId="18.[189,798,602,1922]" pageId="18" pageNumber="165">
<emphasis bold="true" box="[189,258,602,626]" pageId="18" pageNumber="165">Male.</emphasis>
The 7th abdominal dorsal process shown herein from eastern
<collectingRegion box="[450,571,639,662]" country="United States of America" name="Tennessee" pageId="18" pageNumber="165">Tennessee</collectingRegion>
is symmetricallytrapezoidal with no appearance of lateral lobes (
<figureCitation box="[197,284,710,734]" captionStart="Figs" captionStartId="16.[189,241,1664,1688]" captionTargetBox="[249,1383,290,1581]" captionTargetId="figure-100@16.[249,1383,290,1581]" captionTargetPageId="16" captionText="Figs. 4852. Leuctra variabilis, Tennessee, Twin Springs. 48, 7th abdominal process, dorsal view, 350X; 49, paraprocts, caudal view, 350X; 50, paraprocts, caudal view, 1500X; 51, paraprocts, lateral view, 350X; 52, L. variabilis, paraprocts, lateral view, 1500X. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752794" httpUri="https://zenodo.org/record/4752794/files/figure.png" pageId="18" pageNumber="165">Fig. 48</figureCitation>
). The specimens examined from eastern
<collectingRegion box="[189,303,746,770]" country="United States of America" name="Kentucky" pageId="18" pageNumber="165">Kentucky</collectingRegion>
have small, laterally-directed spurs, yet still lack the appearance of lateral lobes. The specilla are broad basally, tapering slightly and not recurved; specilla mostly smooth in appearance, bearing shallow pits along the distal outer margin and very few raised tubercles (
<figureCitation box="[588,723,926,950]" captionStart="Figs" captionStartId="16.[189,241,1664,1688]" captionTargetBox="[249,1383,290,1581]" captionTargetId="figure-100@16.[249,1383,290,1581]" captionTargetPageId="16" captionText="Figs. 4852. Leuctra variabilis, Tennessee, Twin Springs. 48, 7th abdominal process, dorsal view, 350X; 49, paraprocts, caudal view, 350X; 50, paraprocts, caudal view, 1500X; 51, paraprocts, lateral view, 350X; 52, L. variabilis, paraprocts, lateral view, 1500X. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752794" httpUri="https://zenodo.org/record/4752794/files/figure.png" pageId="18" pageNumber="165">Figs. 4952</figureCitation>
). The subanal lobes are narrow throughout length, gently recurved anteriorly, and ca. 90% the length of the inner lobes (
<figureCitation box="[335,503,1034,1058]" captionStart="Figs" captionStartId="16.[189,241,1664,1688]" captionTargetBox="[249,1383,290,1581]" captionTargetId="figure-100@16.[249,1383,290,1581]" captionTargetPageId="16" captionText="Figs. 4852. Leuctra variabilis, Tennessee, Twin Springs. 48, 7th abdominal process, dorsal view, 350X; 49, paraprocts, caudal view, 350X; 50, paraprocts, caudal view, 1500X; 51, paraprocts, lateral view, 350X; 52, L. variabilis, paraprocts, lateral view, 1500X. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752794" httpUri="https://zenodo.org/record/4752794/files/figure.png" pageId="18" pageNumber="165">Figs. 49, 5152</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="165" type="materials_examined">
<paragraph blockId="18.[189,798,602,1922]" pageId="18" pageNumber="165">
<emphasis bold="true" box="[189,430,1069,1093]" pageId="18" pageNumber="165">Material examined.</emphasis>
<materialsCitation ID-GBIF-Occurrence="3111206338" collectingDate="2009-09-09" collectingDateMax="2009-11-04" collectingDateMin="2009-09-09" collectionCode="DCTC" collectorName="M. A. Floyd" country="United States of America" county="Harlan Co." latitude="36.8619" location="Blanton Forest" longLatPrecision="6" longitude="-83.3753" municipality="Watts Creek" pageId="18" pageNumber="165" specimenCount="4" specimenCount-male="4" stateProvince="Kentucky">
<emphasis bold="true" box="[442,500,1069,1093]" pageId="18" pageNumber="165">
<collectingCountry box="[442,500,1069,1093]" name="United States of America" pageId="18" pageNumber="165">USA</collectingCountry>
</emphasis>
,
<emphasis bold="true" box="[519,637,1070,1094]" pageId="18" pageNumber="165">
<collectingRegion box="[519,637,1070,1094]" country="United States of America" name="Kentucky" pageId="18" pageNumber="165">Kentucky</collectingRegion>
</emphasis>
,
<collectingCounty box="[656,792,1070,1094]" pageId="18" pageNumber="165">Harlan Co.</collectingCounty>
,
<collectingMunicipality box="[189,338,1106,1130]" pageId="18" pageNumber="165">Watts Creek</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03DD8784FF8AA70AFEC45EC765D5FE83:8EAB6049FF8AA70AFEA85A716241FB81" box="[350,523,1106,1130]" country="United States of America" county="Harlan Co." latitude="36.8619" longLatPrecision="6" longitude="-83.3753" municipality="Watts Creek" name="Blanton Forest" pageId="18" pageNumber="165" stateProvince="Kentucky">Blanton Forest</location>
State Nature Preserve,
<geoCoordinate box="[189,276,1142,1166]" degrees="36.8619" direction="north" orientation="latitude" pageId="18" pageNumber="165" precision="5" value="36.8619">36.8619</geoCoordinate>
,
<geoCoordinate box="[295,391,1142,1166]" degrees="83.3753" direction="west" orientation="longitude" pageId="18" pageNumber="165" precision="5" value="-83.3753">-83.3753</geoCoordinate>
,
<date box="[409,793,1142,1166]" pageId="18" pageNumber="165" value="2009-09-09" valueMax="2009-11-04" valueMin="2009-09-09">
<collectingDate box="[409,793,1142,1166]" pageId="18" pageNumber="165" value="2009-09-09" valueMax="2009-11-04" valueMin="2009-09-09">9 September4 November 2009</collectingDate>
</date>
,
<collectorName box="[189,328,1177,1202]" pageId="18" pageNumber="165">M.A. Floyd</collectorName>
,
<specimenCount box="[343,377,1177,1202]" pageId="18" pageNumber="165" type="male">4♂</specimenCount>
(
<collectionCode box="[394,474,1178,1202]" pageId="18" pageNumber="165">DCTC</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3111206393" collectingDate="2014-10-23" collectionCode="WKU" collectorName="A. L. Sheldon" country="United States of America" latitude="36.1018" location="Holder Creek" longLatPrecision="6" longitude="-82.1266" pageId="18" pageNumber="165" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="North Carolina">
<collectingRegion box="[496,683,1177,1202]" country="United States of America" name="North Carolina" pageId="18" pageNumber="165">
<emphasis bold="true" box="[496,568,1178,1202]" pageId="18" pageNumber="165">North</emphasis>
<taxonomicName authority=", Mitchell Co." authorityName="Mitchell Co." box="[578,683,1177,1201]" genus="Carolina" kingdom="Animalia" pageId="18" pageNumber="165" rank="genus">
<emphasis bold="true" box="[578,683,1177,1201]" pageId="18" pageNumber="165">Carolina</emphasis>
</taxonomicName>
</collectingRegion>
, Mitchell Co., tributary to
<location LSID="urn:lsid:plazi:treatment:03DD8784FF8AA70AFEC45EC765D5FE83:8EAB6049FF8AA70AFE665A9D6272FB3D" box="[400,568,1214,1238]" country="United States of America" latitude="36.1018" longLatPrecision="6" longitude="-82.1266" name="Holder Creek" pageId="18" pageNumber="165" stateProvince="North Carolina">Holder Creek</location>
, FS 130,
<geoCoordinate box="[683,772,1214,1238]" degrees="36.1018" direction="north" orientation="latitude" pageId="18" pageNumber="165" precision="5" value="36.1018">36.1018</geoCoordinate>
,
<geoCoordinate degrees="82.1266" direction="west" orientation="longitude" pageId="18" pageNumber="165" precision="5" value="-82.1266">- 82.1266</geoCoordinate>
,
<date box="[298,502,1250,1274]" pageId="18" pageNumber="165" value="2014-10-23">
<collectingDate box="[298,502,1250,1274]" pageId="18" pageNumber="165" value="2014-10-23">23 October 2014</collectingDate>
</date>
,
<collectorName box="[525,689,1250,1274]" pageId="18" pageNumber="165">A.L. Sheldon</collectorName>
,
<specimenCount box="[709,742,1250,1274]" pageId="18" pageNumber="165" type="male">1♂</specimenCount>
,
<specimenCount box="[764,798,1250,1274]" pageId="18" pageNumber="165" type="female">1♀</specimenCount>
(
<collectionCode box="[197,266,1286,1310]" country="USA" lsid="urn:lsid:biocol.org:col:15761" name="Western Kentucky University" pageId="18" pageNumber="165" type="Herbarium">WKU</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3111206301" collectionCode="NC" collectorName="Twin Springs" country="United States of America" county="Carter Co." location="26 km N Spruce Pine" pageId="18" pageNumber="165" specimenCount="1" stateProvince="Tennessee">
<emphasis bold="true" box="[293,420,1286,1310]" pageId="18" pageNumber="165">
<collectingRegion box="[293,420,1286,1310]" country="United States of America" name="Tennessee" pageId="18" pageNumber="165">Tennessee</collectingRegion>
</emphasis>
,
<collectingCounty box="[440,569,1286,1310]" pageId="18" pageNumber="165">Carter Co.</collectingCounty>
,
<collectorName box="[587,753,1286,1310]" pageId="18" pageNumber="165">Twin Springs</collectorName>
,
<location LSID="urn:lsid:plazi:treatment:03DD8784FF8AA70AFEC45EC765D5FE83:8EAB6049FF8AA70AFCF45B2561DFFAA9" country="United States of America" county="Carter Co." name="26 km N Spruce Pine" pageId="18" pageNumber="165" stateProvince="Tennessee">
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.6" pageId="18" pageNumber="165" unit="km" value="26.0">26 km</quantity>
N Spruce Pine
</location>
(
<collectionCode box="[422,466,1322,1346]" pageId="18" pageNumber="165">NC</collectionCode>
)
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="3111206323" collectingDate="2002-11-29" collectionCode="WKU" collectorName="S. A. Grubbs" country="United States of America" latitude="36.1213" location="Cherokee National Forest" longLatPrecision="6" longitude="-82.0865" pageId="18" pageNumber="165" specimenCount="3" specimenCount-male="3" stateProvince="Tennessee">
<location LSID="urn:lsid:plazi:treatment:03DD8784FF8AA70AFEC45EC765D5FE83:8EAB6049FF8AA70AFE105B096352FAAA" box="[486,792,1322,1346]" country="United States of America" latitude="36.1213" longLatPrecision="6" longitude="-82.0865" name="Cherokee National Forest" pageId="18" pageNumber="165" stateProvince="Tennessee">Cherokee National Forest</location>
,
<geoCoordinate box="[189,278,1358,1382]" degrees="36.1213" direction="north" orientation="latitude" pageId="18" pageNumber="165" precision="5" value="36.1213">36.1213</geoCoordinate>
,
<geoCoordinate box="[292,387,1358,1382]" degrees="82.0865" direction="west" orientation="longitude" pageId="18" pageNumber="165" precision="5" value="-82.0865">-82.0865</geoCoordinate>
,
<date box="[404,631,1358,1382]" pageId="18" pageNumber="165" value="2002-11-29">
<collectingDate box="[404,631,1358,1382]" pageId="18" pageNumber="165" value="2002-11-29">29 November 2002</collectingDate>
</date>
,
<collectorName box="[645,792,1358,1382]" pageId="18" pageNumber="165">S.A. Grubbs</collectorName>
,
<specimenCount box="[189,223,1394,1418]" pageId="18" pageNumber="165" type="male">3♂</specimenCount>
(
<collectionCode box="[237,309,1394,1418]" country="USA" lsid="urn:lsid:biocol.org:col:15761" name="Western Kentucky University" pageId="18" pageNumber="165" type="Herbarium">WKU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="165" type="distribution">
<paragraph blockId="18.[189,798,602,1922]" pageId="18" pageNumber="165">
<emphasis bold="true" box="[189,348,1429,1453]" pageId="18" pageNumber="165">Distribution.</emphasis>
<collectingCountry box="[355,414,1430,1454]" name="United States of America" pageId="18" pageNumber="165">USA</collectingCountry>
: MA, MD, ME, NH, NJ, NY, PA, VA, VT (
<bibRefCitation author="DeWalt R. E. &amp; M. D. Maehr &amp; U. Neu-Becker &amp; G. Steuber" box="[305,533,1466,1490]" pageId="18" pageNumber="165" refId="ref9097" refString="DeWalt R. E., M. D. Maehr, U. Neu-Becker, &amp; G. Steuber. 2015. Plecoptera species file online. Version 5.0 / 5.0. Available online: http: // Plecoptera. SpeciesFile. org [retrieved 19 July 2015]" type="url" year="2015">DeWalt et al. 2015</bibRefCitation>
), NC, TN (new state records).
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="165" type="discussion">
<paragraph blockId="18.[189,798,602,1922]" lastBlockId="18.[835,1444,228,361]" pageId="18" pageNumber="165">
<emphasis bold="true" box="[189,304,1538,1562]" pageId="18" pageNumber="165">Remarks.</emphasis>
The abdominal dorsal process of
<taxonomicName authorityName=": Illies" authorityYear="1966" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="18" pageNumber="165" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis italics="true" pageId="18" pageNumber="165">L. variabilis</emphasis>
</taxonomicName>
is similar only to
<taxonomicName authorityName="Grubbs" authorityYear="2010" box="[526,605,1574,1598]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="18" pageNumber="165" phylum="Arthropoda" rank="species" species="usdi">
<emphasis box="[526,605,1574,1598]" italics="true" pageId="18" pageNumber="165">L. usdi</emphasis>
</taxonomicName>
.
<bibRefCitation author="Hanson, J. F." box="[624,792,1574,1598]" pageId="18" pageNumber="165" pagination="57 - 66" refId="ref9332" refString="Hanson, J. F. 1941. Studies on the Plecoptera of North America, II. Bulletin of the Brooklyn Entomological Society, 36: 57 - 66." type="journal article" year="1941">Hanson (1941</bibRefCitation>
, fig. 7) depicted three forms of the dorsal process, namely with the absence or presence of small lateral lobes that appear superficially-similar to
<taxonomicName authorityName="Grubbs" authorityYear="2010" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="18" pageNumber="165" phylum="Arthropoda" rank="species" species="usdi">
<emphasis italics="true" pageId="18" pageNumber="165">L. usdi</emphasis>
</taxonomicName>
(
<figureCitation box="[254,336,1718,1742]" captionStart="Figs" captionStartId="15.[189,241,1594,1618]" captionTargetBox="[249,1383,254,1548]" captionTargetId="figure-163@15.[249,1383,254,1548]" captionTargetPageId="15" captionText="Figs. 4347. Leuctra usdi, Tennessee, spring into Pinewood Branch. 43, 7th abdominal process, dorsal view, 350X; 44, paraprocts, caudal view, 500X; 45, paraprocts, caudal view, 2000X; 46, paraprocts, lateral view, 350X; 47, L. usdi, paraprocts, lateral view, 1500X. sp = specillum, sa = subanal lobe, ll = lateral lobe, ml = medial lobe." figureDoi="http://doi.org/10.5281/zenodo.4752792" httpUri="https://zenodo.org/record/4752792/files/figure.png" pageId="18" pageNumber="165">Fig. 43</figureCitation>
). These two species can more readily be separated by characteristics of the specilla. The distal outer margin of the specilla of
<taxonomicName authorityName="Grubbs" authorityYear="2010" box="[647,725,1790,1814]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="18" pageNumber="165" phylum="Arthropoda" rank="species" species="usdi">
<emphasis box="[647,725,1790,1814]" italics="true" pageId="18" pageNumber="165">L. usdi</emphasis>
</taxonomicName>
bears numerous raised tubercles (
<figureCitation box="[517,648,1825,1850]" captionStart="Figs" captionStartId="15.[189,241,1594,1618]" captionTargetBox="[249,1383,254,1548]" captionTargetId="figure-163@15.[249,1383,254,1548]" captionTargetPageId="15" captionText="Figs. 4347. Leuctra usdi, Tennessee, spring into Pinewood Branch. 43, 7th abdominal process, dorsal view, 350X; 44, paraprocts, caudal view, 500X; 45, paraprocts, caudal view, 2000X; 46, paraprocts, lateral view, 350X; 47, L. usdi, paraprocts, lateral view, 1500X. sp = specillum, sa = subanal lobe, ll = lateral lobe, ml = medial lobe." figureDoi="http://doi.org/10.5281/zenodo.4752792" httpUri="https://zenodo.org/record/4752792/files/figure.png" pageId="18" pageNumber="165">Figs. 45, 47</figureCitation>
) whereas on
<taxonomicName authorityName=": Illies" authorityYear="1966" box="[189,327,1861,1885]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="18" pageNumber="165" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis box="[189,327,1861,1885]" italics="true" pageId="18" pageNumber="165">L. variabilis</emphasis>
</taxonomicName>
the tubercles are lacking along the distal outer margin and at the tips (
<figureCitation box="[646,786,1897,1922]" captionStart="Figs" captionStartId="16.[189,241,1664,1688]" captionTargetBox="[249,1383,290,1581]" captionTargetId="figure-100@16.[249,1383,290,1581]" captionTargetPageId="16" captionText="Figs. 4852. Leuctra variabilis, Tennessee, Twin Springs. 48, 7th abdominal process, dorsal view, 350X; 49, paraprocts, caudal view, 350X; 50, paraprocts, caudal view, 1500X; 51, paraprocts, lateral view, 350X; 52, L. variabilis, paraprocts, lateral view, 1500X. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752794" httpUri="https://zenodo.org/record/4752794/files/figure.png" pageId="18" pageNumber="165">Figs. 4952</figureCitation>
). The shallow pits present along the outer margin on
<taxonomicName authorityName=": Illies" authorityYear="1966" box="[835,966,264,288]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="18" pageNumber="165" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis box="[835,966,264,288]" italics="true" pageId="18" pageNumber="165">L. variabilis</emphasis>
</taxonomicName>
are lacking on
<taxonomicName authorityName="Grubbs" authorityYear="2010" box="[1159,1237,264,288]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="18" pageNumber="165" phylum="Arthropoda" rank="species" species="usdi">
<emphasis box="[1159,1237,264,288]" italics="true" pageId="18" pageNumber="165">L. usdi</emphasis>
</taxonomicName>
. Additionally,
<taxonomicName authorityName="Grubbs" authorityYear="2010" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="18" pageNumber="165" phylum="Arthropoda" rank="species" species="usdi">
<emphasis italics="true" pageId="18" pageNumber="165">L. usdi</emphasis>
</taxonomicName>
is a spring-emergent species whereas
<taxonomicName authorityName=": Illies" authorityYear="1966" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="18" pageNumber="165" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis italics="true" pageId="18" pageNumber="165">L. variabilis</emphasis>
</taxonomicName>
emerges during late summer and autumn.
</paragraph>
</subSubSection>
</treatment>
</document>