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<document id="6913F03F79F3B31F46D3690B0E0F57F8" ID-CLB-Dataset="26916" ID-DOI="10.5281/zenodo.2669927" ID-GBIF-Dataset="f32a6f5a-0c9f-43c4-a273-4bc325ece96e" ID-ISSN="1175-5326" ID-Zenodo-Dep="2669927" ID-ZooBank="F066A962-743F-4899-AFAE-485C5A51EF2F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1557202401381" checkinUser="plazi" docAuthor="Williams, Kevin A., Lelej, Arkady S., Okayasu, Juriya, Borkent, Christopher J., Malee, Rufeah, Thoawan, Kodeeyah &amp; Thaochan, Narit" docDate="2019" docId="03DCF504FFE89862FF392DD64D33C25B" docLanguage="en" docName="zootaxa.4602.1.1.pdf" docOrigin="Zootaxa 4602 (1)" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Cockerellidia Lelej &amp; Krombein 1999" docType="treatment" docVersion="20" lastPageNumber="30" masterDocId="FFE58D7CFFF5987FFFAE2F624960C60C" masterDocTitle="The female velvet ants (aka modkhong) of southern Thailand (Hymenoptera: Mutillidae), with a key to the genera of southeast Asia" masterLastPageNumber="69" masterPageNumber="1" pageNumber="30" updateTime="1698726454309" updateUser="ExternalLinkService">
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<mods:title id="0F066AD72E2B4912B4E3B6F58D887959">The female velvet ants (aka modkhong) of southern Thailand (Hymenoptera: Mutillidae), with a key to the genera of southeast Asia</mods:title>
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<mods:namePart id="140C33AEE68BFA75B55810BA875E44EF">Williams, Kevin A.</mods:namePart>
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<mods:namePart id="327EE33E5482B655746619771562D6AA">Lelej, Arkady S.</mods:namePart>
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<mods:namePart id="D3D9E77C780B5FD88AF2CF7BAC3CD878">Okayasu, Juriya</mods:namePart>
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<mods:namePart id="293698F6BA9974542BF6DBEE257829C6">Borkent, Christopher J.</mods:namePart>
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<mods:namePart id="B2B31EF39D95F1457885571B9BA4F296">Malee, Rufeah</mods:namePart>
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<mods:namePart id="C3DB0C942DDB1B39A0DAD5AD5B63A78F">Thoawan, Kodeeyah</mods:namePart>
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<mods:namePart id="851336F6BC2EA5C8525F3EF069907F0E">Thaochan, Narit</mods:namePart>
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<treatment id="03DCF504FFE89862FF392DD64D33C25B" ID-DOI="http://doi.org/10.5281/zenodo.5923617" ID-GBIF-Taxon="157062791" ID-Zenodo-Dep="5923617" LSID="urn:lsid:plazi:treatment:03DCF504FFE89862FF392DD64D33C25B" httpUri="http://treatment.plazi.org/id/03DCF504FFE89862FF392DD64D33C25B" lastPageNumber="30" pageId="29" pageNumber="30">
<subSubSection id="C36F1799FFE89862FF392DD64B93C4C2" box="[151,755,692,718]" pageId="29" pageNumber="30" type="nomenclature">
<paragraph id="8BCA4412FFE89862FF392DD64B93C4C2" blockId="29.[151,755,692,718]" box="[151,755,692,718]" pageId="29" pageNumber="30">
<heading id="D082F37EFFE89862FF392DD64B93C4C2" bold="true" box="[151,755,692,718]" fontSize="11" level="1" pageId="29" pageNumber="30" reason="1">
<emphasis id="B9019800FFE89862FF392DD64B93C4C2" bold="true" box="[151,755,692,718]" pageId="29" pageNumber="30">
Genus 16.
<taxonomicName id="4C753F91FFE89862FEB42DD64B93C4C2" authority="Lelej &amp; Krombein, 1999" authorityName="Lelej &amp; Krombein" authorityYear="1999" box="[282,755,692,718]" class="Insecta" family="Mutillidae" genus="Cockerellidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B9019800FFE89862FEB42DD648DBC4C2" bold="true" box="[282,443,692,718]" italics="true" pageId="29" pageNumber="30">Cockerellidia</emphasis>
<bibRefCitation id="EFE439E3FFE89862FE6C2DD64B93C4C2" author="Lelej, A. S. &amp; Krombein, K. V." box="[450,755,692,718]" pageId="29" pageNumber="30" pagination="1 - 8" refId="ref59248" refString="Lelej, A. S. &amp; Krombein, K. V. (1999) Two remarkable new genera of mutillid wasps (Hymenoptera: Mutillidae, Sphaeropthalminae, Pseudomethocini) from Thailand. Far Eastern entomologist, 79, 1 - 8." type="journal article" year="1999">Lelej &amp; Krombein, 1999</bibRefCitation>
</taxonomicName>
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</paragraph>
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<paragraph id="8BCA4412FFE89862FF392D98480AC557" blockId="29.[151,1437,762,1111]" pageId="29" pageNumber="30">
<emphasis id="B9019800FFE89862FF392D984871C51F" bold="true" box="[151,273,762,787]" pageId="29" pageNumber="30">Diagnosis.</emphasis>
FEMALE. Immediately recognizable by the large red-brown head, small eyes, laterally swollen mesonotum, and dense elongate setae covering the body. See
<bibRefCitation id="EFE439E3FFE89862FCC22C7C4DCBC53B" author="Lelej, A. S. &amp; Krombein, K. V." box="[876,1195,798,823]" pageId="29" pageNumber="30" pagination="1 - 8" refId="ref59248" refString="Lelej, A. S. &amp; Krombein, K. V. (1999) Two remarkable new genera of mutillid wasps (Hymenoptera: Mutillidae, Sphaeropthalminae, Pseudomethocini) from Thailand. Far Eastern entomologist, 79, 1 - 8." type="journal article" year="1999">Lelej and Krombein (1999)</bibRefCitation>
for additional traits. MALE. Unknown.
</paragraph>
<paragraph id="8BCA4412FFE89862FF692C044ACBC573" blockId="29.[151,1437,762,1111]" box="[199,939,870,895]" pageId="29" pageNumber="30">
<emphasis id="B9019800FFE89862FF692C044B68C573" bold="true" box="[199,520,870,895]" pageId="29" pageNumber="30">Diversity and Distribution.</emphasis>
One species from northern
<collectingCountry id="F3620482FFE89862FCEE2C044AC7C573" box="[832,935,870,895]" name="Thailand" pageId="29" pageNumber="30">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C36F1799FFE89862FF692CE84D33C25B" pageId="29" pageNumber="30" type="discussion">
<paragraph id="8BCA4412FFE89862FF692CE84D33C25B" blockId="29.[151,1437,762,1111]" pageId="29" pageNumber="30">
<emphasis id="B9019800FFE89862FF692CE8485BC5AF" bold="true" box="[199,315,906,931]" pageId="29" pageNumber="30">Remarks.</emphasis>
<taxonomicName id="4C753F91FFE89862FEEC2CE84B90C5AF" authority="(Cockerell, 1928)" baseAuthorityName="Cockerell" baseAuthorityYear="1928" box="[322,752,906,931]" class="Insecta" family="Mutillidae" genus="Cockerellidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="sohmi">
<emphasis id="B9019800FFE89862FEEC2CE84B43C5AF" box="[322,547,906,931]" italics="true" pageId="29" pageNumber="30">Cockerellidia sohmi</emphasis>
(
<bibRefCitation id="EFE439E3FFE89862FD9C2CE84B88C5AF" author="Cockerell, T. D. A." box="[562,744,906,931]" pageId="29" pageNumber="30" pagination="596 - 600" refId="ref58143" refString="Cockerell, T. D. A. (1928) Mutillidae from Siam. The Annals and Magazine of Natural History, Series 10, 2, 596 - 600. https: // doi. org / 10.1080 / 00222932808672926" type="journal article" year="1928">Cockerell, 1928</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="134E5897FFE89862FD512CE84A31C5AF" box="[767,849,906,931]" captionStart="FIGURES 128" captionStartId="51.[151,263,1875,1898]" captionTargetBox="[169,1417,193,1814]" captionTargetId="figure@51.[169,1417,193,1854]" captionTargetPageId="51" captionText="FIGURES 128. Mutillidae, dorsal habitus of females. Scale bars represent actual specimen length." httpUri="https://zenodo.org/record/2669929/files/figure.png" pageId="29" pageNumber="30">Fig. 15</figureCitation>
) is known only from the unique
<typeStatus id="54CEFAB0FFE89862FB6F2CE94D91C5AF" box="[1217,1265,907,931]" pageId="29" pageNumber="30">type</typeStatus>
collected in an uncertain locality of northern
<collectingCountry id="F3620482FFE89862FE4B2CCC4B2DC5CB" box="[485,589,942,967]" name="Thailand" pageId="29" pageNumber="30">Thailand</collectingCountry>
: “
<collectingCountry id="F3620482FFE89862FDCF2CCC4BFDC5CB" box="[609,669,942,967]" name="Thailand" pageId="29" pageNumber="30">Siam</collectingCountry>
near Mecatin”. It is similar to the following genus,
<taxonomicName id="4C753F91FFE89862FB4E2CCC48AEC5E7" authority="Lelej in Lelej &amp; Krombein, 1999" authorityName="Lelej in Lelej &amp; Krombein" authorityYear="1999" class="Insecta" family="Mutillidae" genus="Karlidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B9019800FFE89862FB4E2CCC4C5FC5CB" box="[1248,1343,942,967]" italics="true" pageId="29" pageNumber="30">Karlidia</emphasis>
Lelej in
<bibRefCitation id="EFE439E3FFE89862FF392CB048AEC5E7" author="Lelej, A. S. &amp; Krombein, K. V." box="[151,462,978,1003]" pageId="29" pageNumber="30" pagination="1 - 8" refId="ref59248" refString="Lelej, A. S. &amp; Krombein, K. V. (1999) Two remarkable new genera of mutillid wasps (Hymenoptera: Mutillidae, Sphaeropthalminae, Pseudomethocini) from Thailand. Far Eastern entomologist, 79, 1 - 8." type="journal article" year="1999">Lelej &amp; Krombein, 1999</bibRefCitation>
</taxonomicName>
, which was described in the same publication. The most recent phylogenetic reconstruction (
<bibRefCitation id="EFE439E3FFE89862FEEB2C944B25C203" author="Brothers, D. J. &amp; Lelej, A. S." box="[325,581,1014,1039]" pageId="29" pageNumber="30" pagination="1 - 97" refId="ref57992" refString="Brothers, D. J. &amp; Lelej, A. S. (2017) Phylogeny and higher classification of Mutillidae (Hymenoptera) based on morphological reanalyses. Journal of Hymenoptera Research, 60, 1 - 97. https: // doi. org / 10.3897 / jhr. 60.20091" type="journal article" year="2017">Brothers &amp; Lelej 2017</bibRefCitation>
) indicates that
<taxonomicName id="4C753F91FFE89862FD432C944AE4C203" authorityName="Lelej &amp; Krombein" authorityYear="1999" box="[749,900,1014,1039]" class="Insecta" family="Mutillidae" genus="Cockerellidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B9019800FFE89862FD432C944AE4C203" box="[749,900,1014,1039]" italics="true" pageId="29" pageNumber="30">Cockerellidia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C753F91FFE89862FC142C944D79C203" authorityName="Lelej in Lelej &amp; Krombein" authorityYear="1999" box="[954,1049,1014,1039]" class="Insecta" family="Mutillidae" genus="Karlidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B9019800FFE89862FC142C944D79C203" box="[954,1049,1014,1039]" italics="true" pageId="29" pageNumber="30">Karlidia</emphasis>
</taxonomicName>
belong to the subtribe Ephutina of the tribe
<taxonomicName id="4C753F91FFE89862FF542B78483EC23F" authorityName="Latreille" authorityYear="1802" box="[250,350,1050,1075]" class="Insecta" family="Mutillidae" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="tribe" tribe="Mutillini">Mutillini</taxonomicName>
in subfamily
<taxonomicName id="4C753F91FFE89862FE562B784B10C23F" authorityName="Latreille" authorityYear="1802" box="[504,624,1050,1075]" class="Insecta" family="Mutillidae" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="subFamily" subFamily="Mutillinae">Mutillinae</taxonomicName>
, rather than
<taxonomicName id="4C753F91FFE89862FD542B784AB6C23F" box="[762,982,1050,1075]" class="Insecta" family="Mutillidae" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="subFamily" subFamily="Sphaeropthalminae">Sphaeropthalminae</taxonomicName>
. The unknown males of these genera are likely to be confused with
<taxonomicName id="4C753F91FFE89862FE6F2B5C4A7BC25B" authority="Ashmead, 1899" authorityName="Ashmead" authorityYear="1899" box="[449,795,1086,1111]" class="Insecta" family="Mutillidae" genus="Odontomutilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B9019800FFE89862FE6F2B5C4B05C25B" box="[449,613,1086,1111]" italics="true" pageId="29" pageNumber="30">Odontomutilla</emphasis>
<bibRefCitation id="EFE439E3FFE89862FDC52B5C4A7BC25B" author="Ashmead, W. H." box="[619,795,1086,1111]" pageId="29" pageNumber="30" pagination="45 - 60" refId="ref57634" refString="Ashmead, W. H. (1899) Superfamilies in the Hymenoptera and generic synopses of the families Thynnidae, Myrmosidae, and Mutillidae. Journal of the New York Entomological Society, 7, 45 - 60." type="journal article" year="1899">Ashmead, 1899</bibRefCitation>
</taxonomicName>
or
<emphasis id="B9019800FFE89862FCEE2B5C4AA5C25B" box="[832,965,1086,1111]" italics="true" pageId="29" pageNumber="30">Yamanetilla</emphasis>
Lelej, 1996.
</paragraph>
</subSubSection>
</treatment>
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