treatments-xml/data/03/DC/6C/03DC6C76CC2A4C38F7DFFF78FDF0FCA0.xml
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<document id="515708C081424AE58E14299AD49743F7" ID-DOI="10.11646/zootaxa.5267.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7840783" ID-ZooBank="9CD59054-8D7D-413F-B9FD-29EAFE7E511D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1681816955386" checkinUser="plazi" docAuthor="Lehmann, Ingo, Zahiri, Reza &amp; Husemann, Martin" docDate="2023" docId="03DC6C76CC2A4C38F7DFFF78FDF0FCA0" docLanguage="en" docName="zootaxa.5267.1.1.pdf" docOrigin="Zootaxa 5267 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5267.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Lukeniana michaelgrzimeki Lehmann, Zahiri &amp; Husemann 2023, sp. nov." docType="treatment" docUuid="40073D25-07E7-472C-8774-89022C1D8722" docUuidSource="ZooBank" docVersion="3" lastPageNumber="29" masterDocId="FFE5140ECC314C24F748FFEFFF9EFFB5" masterDocTitle="Revision of the Metarbelodes Strand, 1909 genus-group (Lepidoptera: Cossoidea: Metarbelidae) with descriptions of two new genera and 33 new species from high elevations of eastern and southern Africa" masterLastPageNumber="106" masterPageNumber="1" pageNumber="28" updateTime="1699487052491" updateUser="plazi">
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<mods:title id="615D59531FE6BFDD6CDCF8A9189AE885">Revision of the Metarbelodes Strand, 1909 genus-group (Lepidoptera: Cossoidea: Metarbelidae) with descriptions of two new genera and 33 new species from high elevations of eastern and southern Africa</mods:title>
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<mods:namePart id="0AB8223D50867486A09D38E749F86B8F">Lehmann, Ingo</mods:namePart>
<mods:nameIdentifier id="0C06F97E28CAD8C20626E339EE425D03" type="email">martin.husemann@uni-hamburg.de</mods:nameIdentifier>
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<mods:namePart id="274A4361E6F0D6A8F4F9C86F1D88C5E1">Zahiri, Reza</mods:namePart>
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<mods:namePart id="6E383831FB6096794114D0A99B238FAA">Husemann, Martin</mods:namePart>
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<subSubSection id="C36F8EEBCC2A4C3FF7DFFF78FC43FF04" box="[151,989,151,178]" pageId="27" pageNumber="28" type="nomenclature">
<paragraph id="8BCADD60CC2A4C3FF7DFFF78FC43FF04" blockId="27.[151,989,151,250]" box="[151,989,151,178]" pageId="27" pageNumber="28">
<heading id="D0826A0CCC2A4C3FF7DFFF78FC43FF04" bold="true" box="[151,989,151,178]" fontSize="11" level="2" pageId="27" pageNumber="28" reason="4">
<emphasis id="B9010172CC2A4C3FF7DFFF78FC43FF04" bold="true" box="[151,989,151,178]" pageId="27" pageNumber="28">
<taxonomicName id="4C75A6E3CC2A4C3FF7DFFF78FCE6FF04" ID-CoL="B8R8T" authority="Lehmann, Zahiri &amp; Husemann" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[151,888,151,178]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="michaelgrzimeki" status="sp. nov.">
<emphasis id="B9010172CC2A4C3FF7DFFF78FE75FF07" bold="true" box="[151,491,151,178]" italics="true" pageId="27" pageNumber="28">Lukeniana michaelgrzimeki</emphasis>
Lehmann, Zahiri &amp; Husemann
</taxonomicName>
<taxonomicNameLabel id="A232BC09CC2A4C3FF437FF78FC43FF04" box="[895,989,151,177]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C36F8EEBCC2A4C3FF7DFFF54FBEEFEF4" pageId="27" pageNumber="28" type="description">
<paragraph id="8BCADD60CC2A4C3FF7DFFF54FE1CFF60" blockId="27.[151,989,151,250]" box="[151,386,187,214]" pageId="27" pageNumber="28">
<figureCitation id="134EC1E5CC2A4C3FF7DFFF54FE8BFF63" box="[151,277,187,214]" captionStart="FIGURE 5" captionStartId="26.[151,250,1484,1508]" captionTargetBox="[151,1436,181,1460]" captionTargetId="figure-28@26.[151,1436,181,1460]" captionTargetPageId="26" captionText="FIGURE 5. a. Lukeniana chapmani, holotype, male, Malawi, Southern Region, Mount Mulanje, Likabula; b. L. michaelgrzimeki, holotype, male, Tanzania, Arusha Region, Mount Meru, Momella; c. L. michaelgrzimeki, paratype, female, Tanzania, Arusha Region, Mount Meru, Momella; d. L. timdavenporti, holotype, male, Kenya, Rift Valley Province, Mount Elgon National Park, Chepnyalil Cave; e. L. mzuzuensis, holotype, male, Malawi, Northern Region, Mzuzu, Nkhorongo; f. L. mzuzuensis, paratype, female, Malawi, Northern Region, Mzuzu, Nkhorongo." figureDoi="http://doi.org/10.5281/zenodo.7840798" httpUri="https://zenodo.org/record/7840798/files/figure.png" pageId="27" pageNumber="28">Figs 5b, 5c</figureCitation>
,
<figureCitation id="134EC1E5CC2A4C3FF669FF54FED2FF60" box="[289,332,187,213]" captionStart="FIGURE 10" captionStartId="60.[151,250,1754,1778]" captionTargetBox="[151,1436,677,1731]" captionTargetId="figure-255@60.[151,1436,669,1731]" captionTargetPageId="60" captionText="FIGURE 10. An intact CuP vein on the forewing is a plesiomorphy (cf. Fig. 9) and occurs only in basal Metarbelidae in northeastern parts of the Republic of South Africa, mainly on the Southern African Plateau, and on the plateau area of Mount Mulanje, only 90 km to the North of the Zambezi River (Lehmann 2019b). Figured are examples of apomorphies in regard to the CuP on forewing (red arrows): a. CuP absent, CuP fold is weak but not continuous in Lukeniana michaelgrzimeki, female, paratype; b. The CuP fold is almost entirely absent in L. rajaeii, male, paratype; c. Zambezia darrelplowesi, female, holotype and d. Z. madambae, male, paratype, both have a strong CuP fold, continuous and partly sclerotized. Images were taken by Dr. Hossein Rajaei (SMNS) in November 2015." figureDoi="http://doi.org/10.5281/zenodo.7840810" httpUri="https://zenodo.org/record/7840810/files/figure.png" pageId="27" pageNumber="28">10a</figureCitation>
,
<figureCitation id="134EC1E5CC2A4C3FF61FFF54FE1CFF60" box="[343,386,187,213]" captionStart="FIGURE 16" captionStartId="67.[151,250,1673,1697]" captionTargetBox="[159,1428,391,1636]" captionTargetId="figure-109@67.[151,1436,381,1649]" captionTargetPageId="67" captionText="FIGURE 16. a. Lukeniana madrandelei, paratype, male, DRC, Katanga Province, Lubumbashi; b. L. friederikebauerae, paratype, male, Zambia, Lusaka Province, Lusaka; c. L. chapmani, holotype, male, Malawi, Southern Region, Mount Mulanje, Likabula; d. L. michaelgrzimeki, holotype, male, Tanzania, Arusha Region, Mount Meru, Momella; e. L. kammeri, holotype, male, Kenya, County Kiambu, Thika; f. L. timdavenporti, holotype, male, Kenya, Rift Valley Province, Mount Elgon National Park, Chepnyalil Cave." figureDoi="http://doi.org/10.5281/zenodo.7840829" httpUri="https://zenodo.org/record/7840829/files/figure.png" pageId="27" pageNumber="28">16d</figureCitation>
</paragraph>
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<uri id="FFE4D162CC2A4C3FF7DFFF30FC30FF4F" box="[151,942,223,250]" pageId="27" pageNumber="28">
urn:lsid:zoobank.org:act:
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<materialsCitation id="3B1DD73DCC2A4C3FF7DFFEC7FBF2FEF4" box="[151,1132,295,322]" collectionCode="ZSM" country="Tanzania" location="Zoological State Collection" municipality="Munich" pageId="27" pageNumber="28" specimenCount="1" typeStatus="holotype">
<typeStatus id="54CE63C2CC2A4C3FF7DFFEC7FF51FEF7" box="[151,207,296,322]" pageId="27" pageNumber="28">Type</typeStatus>
locality and repository:
<collectingCountry id="F3629DF0CC2A4C3FF6A8FEC8FDD6FEF4" box="[480,584,295,321]" name="Tanzania" pageId="27" pageNumber="28">Tanzania</collectingCountry>
, the
<location id="8EAA8BBBCC2A4C3FF534FEC8FC2DFEF4" LSID="urn:lsid:plazi:treatment:03DC6C76CC2A4C38F7DFFF78FDF0FCA0:8EAA8BBBCC2A4C3FF534FEC8FC2DFEF4" box="[636,947,295,321]" country="Tanzania" municipality="Munich" name="Zoological State Collection" pageId="27" pageNumber="28">Zoological State Collection</location>
,
<collectingMunicipality id="6BAE471ACC2A4C3FF4F6FEC8FB8BFEF4" box="[958,1045,295,321]" pageId="27" pageNumber="28">Munich</collectingMunicipality>
(
<collectionCode id="ED6445A5CC2A4C3FF36CFEC8FBFDFEF4" box="[1060,1123,295,321]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="27" pageNumber="28" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
.
</paragraph>
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<subSubSection id="C36F8EEBCC2A4C3FF7DFFE80FACCFE40" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="8BCADD60CC2A4C3FF7DFFE80FACCFE40" blockId="27.[151,1437,367,2014]" pageId="27" pageNumber="28">
<emphasis id="B9010172CC2A4C3FF7DFFE80FE1CFE3C" bold="true" box="[151,386,367,393]" pageId="27" pageNumber="28">Material examined.</emphasis>
<materialsCitation id="3B1DD73DCC2A4C3FF6C2FE80FA87FE18" collectingDate="1964-01-20" collectingDateMax="1964-01-31" collectingDateMin="1964-01-20" collectionCode="ZSM" collectorName="W. Forster &amp; I. Lehmann" country="Tanzania" county="Arumeru District" elevation="2" location="Momella" municipality="Mount Meru" pageId="27" pageNumber="28" specimenCount="1" stateProvince="Arusha" typeStatus="holotype">
<typeStatus id="54CE63C2CC2A4C3FF6C2FE80FE6CFE3C" box="[394,498,367,393]" pageId="27" pageNumber="28" type="holotype">Holotype</typeStatus>
male,
<collectingCountry id="F3629DF0CC2A4C3FF508FE80FD36FE3C" box="[576,680,367,393]" name="Tanzania" pageId="27" pageNumber="28">Tanzania</collectingCountry>
,
<collectingRegion id="49B11382CC2A4C3FF5FBFE80FCFEFE3C" box="[691,864,367,393]" country="Tanzania" name="Arusha" pageId="27" pageNumber="28">Arusha Region</collectingRegion>
,
<collectingCounty id="62ABA5ECCC2A4C3FF423FE80FBB1FE3C" box="[875,1071,367,393]" pageId="27" pageNumber="28">Arumeru District</collectingCounty>
,
<collectingMunicipality id="6BAE471ACC2A4C3FF374FE9FFB51FE3F" box="[1084,1231,368,394]" pageId="27" pageNumber="28">Mount Meru</collectingMunicipality>
,
<location id="8EAA8BBBCC2A4C3FF393FE80FADDFE3C" LSID="urn:lsid:plazi:treatment:03DC6C76CC2A4C38F7DFFF78FDF0FCA0:8EAA8BBBCC2A4C3FF393FE80FADDFE3C" box="[1243,1347,367,393]" country="Tanzania" county="Arumeru District" municipality="Mount Meru" name="Momella" pageId="27" pageNumber="28" stateProvince="Arusha">Momella</location>
,
<quantity id="4C8D7085CC2A4C3FF207FE80FF69FE1B" metricMagnitude="0" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="27" pageNumber="28" unit="m" value="1.7" valueMax="1.8" valueMin="1.6">
<elevation id="00583A53CC2A4C3FF207FE80FF69FE1B" metricMagnitude="0" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="27" pageNumber="28" unit="m" value="1.7" valueMax="1.8" valueMin="1.6">1.600 1.800 m</elevation>
</quantity>
,
<date id="FFCBFBA0CC2A4C3FF64DFE7CFE6EFE18" box="[261,496,403,430]" pageId="27" pageNumber="28" value="1964-01-20" valueMax="1964-01-31" valueMin="1964-01-20">
<collectingDate id="EF8F0248CC2A4C3FF64DFE7CFE6EFE18" box="[261,496,403,430]" pageId="27" pageNumber="28" value="1964-01-20" valueMax="1964-01-31" valueMin="1964-01-20">2031 January 1964</collectingDate>
</date>
,
<collectorName id="2680B8B6CC2A4C3FF6B4FE7BFDEDFE1B" box="[508,627,404,430]" pageId="27" pageNumber="28">W. Forster</collectorName>
leg., genitalia slide number 24/082012
<collectorName id="2680B8B6CC2A4C3FF375FE7BFB5FFE18" box="[1085,1217,403,430]" pageId="27" pageNumber="28">I. Lehmann</collectorName>
(
<collectionCode id="ED6445A5CC2A4C3FF39AFE7CFA8FFE18" box="[1234,1297,403,429]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="27" pageNumber="28" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B1DD73DCC2A4C3FF26EFE7BFEC9FE40" collectingDate="1964-02-01" collectingDateMax="1964-02-10" collectingDateMin="1964-02-01" collectionCode="ZSM" collectorName="W. Forster &amp; I. Lehmann" country="Tanzania" county="Arumeru District" elevation="2" location="Momella" municipality="Mount Meru" pageId="27" pageNumber="28" specimenCount="1" specimenCount-female="1" stateProvince="Arusha" typeStatus="paratype">
<typeStatus id="54CE63C2CC2A4C3FF26EFE7BFA0DFE1B" box="[1318,1427,404,430]" pageId="27" pageNumber="28" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D7316E9CC2A4C3FF7DFFE57FE8BFE64" box="[151,277,439,466]" pageId="27" pageNumber="28" type="female">one female</specimenCount>
, same locality and elevation,
<date id="FFCBFBA0CC2A4C3FF516FE58FCCCFE64" box="[606,850,439,465]" pageId="27" pageNumber="28" value="1964-02-01" valueMax="1964-02-10" valueMin="1964-02-01">
<collectingDate id="EF8F0248CC2A4C3FF516FE58FCCCFE64" box="[606,850,439,465]" pageId="27" pageNumber="28" value="1964-02-01" valueMax="1964-02-10" valueMin="1964-02-01">0110 February 1964</collectingDate>
</date>
,
<collectorName id="2680B8B6CC2A4C3FF414FE57FC4FFE67" box="[860,977,440,466]" pageId="27" pageNumber="28">W. Forster</collectorName>
leg., genitalia slide number 10/082014
<collectorName id="2680B8B6CC2A4C3FF2C4FE57FE9FFE40" pageId="27" pageNumber="28">I. Lehmann</collectorName>
(
<collectionCode id="ED6445A5CC2A4C3FF658FE34FED1FE40" box="[272,335,475,501]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="27" pageNumber="28" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1DD73DCC2A4C3FF62BFE33FAD0FE40" box="[355,1358,475,502]" collectingDate="1964-02-01" collectingDateMax="1964-02-10" collectingDateMin="1964-02-01" collectionCode="ZSM" collectorName="W. Forster &amp; I. Lehmann" country="Tanzania" county="Arumeru District" elevation="2" location="Momella" municipality="Mount Meru" pageId="27" pageNumber="28" specimenCount="1" specimenCount-male="1" stateProvince="Arusha" typeStatus="paratype">
<specimenCount id="9D7316E9CC2A4C3FF62BFE33FE55FE40" box="[355,459,475,502]" pageId="27" pageNumber="28" type="male">one male</specimenCount>
, same locality, elevation, date and collector, no genitalia dissection done (
<collectionCode id="ED6445A5CC2A4C3FF24FFE34FAD8FE40" box="[1287,1350,475,501]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="27" pageNumber="28" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC2A4C3FF78FFE10FDABF820" pageId="27" pageNumber="28" type="description">
<paragraph id="8BCADD60CC2A4C3FF78FFE10FAE3FDD4" blockId="27.[151,1437,367,2014]" pageId="27" pageNumber="28">
<emphasis id="B9010172CC2A4C3FF78FFE10FEC7FDAC" bold="true" box="[199,345,511,537]" pageId="27" pageNumber="28">Description.</emphasis>
Male.
<emphasis id="B9010172CC2A4C3FF6E7FE10FE72FDAC" box="[431,492,511,537]" italics="true" pageId="27" pageNumber="28">Head</emphasis>
: Yellow-ochre; eyes brown with black patches; antenna 0.48 length of forewing, bipectinate, with branches 6 width of shaft, covered with ivory-yellow scales laterally, shaft covered with ivory-yellow scales dorsally; antennal tips not spatulate with two longer scales, bending towards apex; labial palpi ochre.
</paragraph>
<paragraph id="8BCADD60CC2A4C3FF78FFD84FE60FC58" blockId="27.[151,1437,367,2014]" pageId="27" pageNumber="28">
<emphasis id="B9010172CC2A4C3FF78FFD84FE86FD30" box="[199,280,619,645]" italics="true" pageId="27" pageNumber="28">Thorax</emphasis>
: Patagia and tegulae with long hair-like scales of cream mixed with yellow ochre, glossy. A small crest of ivory-yellow on metathorax. Hindlegs ochre with fine hair-like scales, shiny; one pair of narrow, long tibial spurs present, outer spur
<emphasis id="B9010172CC2A4C3FF625FD5BFE11FD78" box="[365,399,692,717]" italics="true" pageId="27" pageNumber="28">ca.</emphasis>
<quantity id="4C8D7085CC2A4C3FF6DEFD5CFE70FD7B" box="[406,494,691,718]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="27" pageNumber="28" unit="mm" value="1.2">1.2 mm</quantity>
, inner spur
<emphasis id="B9010172CC2A4C3FF539FD5BFD0CFD78" box="[625,658,692,717]" italics="true" pageId="27" pageNumber="28">ca.</emphasis>
1.0 mm. Forewing length of
<typeStatus id="54CE63C2CC2A4C3FF492FD5CFBA2FD78" box="[986,1084,691,717]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
<quantity id="4C8D7085CC2A4C3FF30BFD5CFB38FD7B" box="[1091,1190,691,718]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="27" pageNumber="28" unit="mm" value="12.5">12.5 mm</quantity>
(wingspan
<quantity id="4C8D7085CC2A4C3FF260FD5CFA0EFD78" box="[1320,1424,691,718]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" pageId="27" pageNumber="28" unit="mm" value="27.5">27.5 mm</quantity>
). Forewing upperside ochre with few sepia striae, costal margin sepia with few darker striae; termen with lunules of sepia; a slightly bent subterminal line of sepia from near apex to end of CuA
<subScript id="17F1DF25CC2A4C3FF4A1FCE1FC6FFCA9" attach="left" box="[1001,1009,782,796]" fontSize="6" pageId="27" pageNumber="28">2</subScript>
, with a square design between M
<subScript id="17F1DF25CC2A4C3FF22DFCE1FAF3FCA9" attach="left" box="[1381,1389,782,796]" fontSize="6" pageId="27" pageNumber="28">1</subScript>
and M
<subScript id="17F1DF25CC2A4C3FF7F8FCDDFF26FCF5" attach="left" box="[176,184,818,832]" fontSize="6" pageId="27" pageNumber="28">2</subScript>
(possibly a variable character); at upper end of CuA
<subScript id="17F1DF25CC2A4C3FF449FCDDFC97FCF5" attach="left" box="[769,777,818,832]" fontSize="6" pageId="27" pageNumber="28">2</subScript>
a narrow and almost straight line of sepia to costal margin; CuA
<subScript id="17F1DF25CC2A4C3FF784FCB9FF4AFCD1" attach="left" box="[204,212,854,868]" fontSize="6" pageId="27" pageNumber="28">2</subScript>
broad, ivory-yellow with pure white, edged by a narrow band of sepia above; all remaining veins not distinctly coloured; upper base of forewing dark ochre in both sexes; cilia very long, 2.0 mm, yellow-ochre, shiny. Underside of forewing roughly scaled, primarily ivory-yellow, ochre along costa, glossy, subterminal line not visible. Hindwing upperside ivory-yellow mixed with pure white, glossy, cilia colour as in forewing, very long, 2.0 mm in both sexes; underside as in forewing.
</paragraph>
<paragraph id="8BCADD60CC2A4C3FF78FFC18FF59FA50" blockId="27.[151,1437,367,2014]" pageId="27" pageNumber="28">
<emphasis id="B9010172CC2A4C3FF78FFC18FEAFFBA4" box="[199,305,1015,1041]" italics="true" pageId="27" pageNumber="28">Abdomen</emphasis>
: Ochre mixed with ivory-yellow, shiny; abdominal tuft one-fifth of abdominal length. Genitalia (
<figureCitation id="134EC1E5CC2A4C3FF227FC18FF57FB80" captionStart="FIGURE 16" captionStartId="67.[151,250,1673,1697]" captionTargetBox="[159,1428,391,1636]" captionTargetId="figure-109@67.[151,1436,381,1649]" captionTargetPageId="67" captionText="FIGURE 16. a. Lukeniana madrandelei, paratype, male, DRC, Katanga Province, Lubumbashi; b. L. friederikebauerae, paratype, male, Zambia, Lusaka Province, Lusaka; c. L. chapmani, holotype, male, Malawi, Southern Region, Mount Mulanje, Likabula; d. L. michaelgrzimeki, holotype, male, Tanzania, Arusha Region, Mount Meru, Momella; e. L. kammeri, holotype, male, Kenya, County Kiambu, Thika; f. L. timdavenporti, holotype, male, Kenya, Rift Valley Province, Mount Elgon National Park, Chepnyalil Cave." figureDoi="http://doi.org/10.5281/zenodo.7840829" httpUri="https://zenodo.org/record/7840829/files/figure.png" pageId="27" pageNumber="28">Fig. 16 d</figureCitation>
) with uncus represents a pair of broad rounded lobes, outer edge slightly C-shaped, basal edge pointed as well as strongly bent at middle towards emargination of uncus, emargination small, width slightly broader than tip of uncus lobe; lobes densely covered with long setae ventrally; gnathos arms long (almost 1.2 basal width of valva), but not touching coastal margin of valva, slightly bent towards uncus; valva ovoid, broader at base, costa slightly bent, with few long setae; sacculus with long setae; weakly-sclerotized projection with a prickly appearance due to many long setae, tip rounded, its end is equal in length to thickly sclerotized, narrow and short thorn-like process; latter slightly bent, hollow, with a rounded tip and many dots with tiny setae; from base of thorn-like process to beginning of sacculus edge S-shaped ventrally; median sector of valva with scattered long setae on inner side; long, broad, ovoid emargination extending between weakly-sclerotized projection and thorn-like process,
<emphasis id="B9010172CC2A4C3FF3BCFAD3FA88FAE0" box="[1268,1302,1340,1365]" italics="true" pageId="27" pageNumber="28">ca.</emphasis>
40% length of valva; ventral side of valva not bent at middle. Saccus as long as juxta, digitate, narrow, rounded distally. Juxta broad with two acuminate tips, each bearing a thorn-like process, emargination between tips deep, 90% length of juxta. Phallus long, 1.2 length of valva, almost trumpet-like, strongly bent near narrower end, bilobed, cleft on one end.
</paragraph>
<paragraph id="8BCADD60CC2A4C3FF78FFA00FAA0F9E4" blockId="27.[151,1437,367,2014]" pageId="27" pageNumber="28">
Female.
<emphasis id="B9010172CC2A4C3FF660FA00FEFBF9BC" box="[296,357,1519,1545]" italics="true" pageId="27" pageNumber="28">Head</emphasis>
and
<emphasis id="B9010172CC2A4C3FF6D5FA00FE70F9BC" box="[413,494,1519,1545]" italics="true" pageId="27" pageNumber="28">Thorax</emphasis>
: Essentially as described for male, except antenna unipectinate, slightly bipectinate towards tip; branches of antenna 1 width of shaft; antenna 0.41 length of forewing; scales of patagia and tegulae not shiny; lunules of termen faded compared to those of male; forewing length
<quantity id="4C8D7085CC2A4C3FF4ACF9D8FBD6F9E7" box="[996,1096,1591,1618]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" pageId="27" pageNumber="28" unit="mm" value="21.5">21.5 mm</quantity>
(wingspan
<quantity id="4C8D7085CC2A4C3FF382F9D8FAACF9E4" box="[1226,1330,1591,1618]" metricMagnitude="-2" metricUnit="m" metricValue="4.65" pageId="27" pageNumber="28" unit="mm" value="46.5">46.5 mm</quantity>
).
</paragraph>
<paragraph id="8BCADD60CC2A4C3FF78FF9B4FDABF820" blockId="27.[151,1437,367,2014]" pageId="27" pageNumber="28">
<emphasis id="B9010172CC2A4C3FF78FF9B4FEAFF9C0" box="[199,305,1627,1653]" italics="true" pageId="27" pageNumber="28">Abdomen</emphasis>
: Genitalia with papillae anales broad, dorsal part 8-shaped in posterior view, densely covered with long and short setae. Segment 8 with scattered long setae along posterior margin and on entire surface; two broad latero-ventral sclerotized bands, completely fused along posterior margin and separated along anterior margin by a rectangular end towards base of anterior apophysis; bands covering more than 100% of ventral side of segment 8 (in ventral view) and are thickly sclerotized. Dorso-anterior margin of abdominal plate without an emargination. Posterior apophysis straight with slightly broader base 1.5 as broad as width of base of anterior apophysis. Anterior apophyses straight, rounded at tip. Posterior apophyses and anterior apophyses of almost equal length. Ductus and corpus bursae broad, thinly membranous and without any processes or distinct characters; corpus bursae elliptical,
<emphasis id="B9010172CC2A4C3FF7DFF893FF27F820" box="[151,185,1916,1941]" italics="true" pageId="27" pageNumber="28">ca.</emphasis>
3.0 as long as anterior apophysis.
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC2A4C38F78FF870FB94FE18" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" type="diagnosis">
<paragraph id="8BCADD60CC2A4C38F78FF870FB94FE18" blockId="27.[151,1437,367,2014]" lastBlockId="28.[151,1437,151,789]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
<emphasis id="B9010172CC2A4C3FF78FF870FEDCF80C" bold="true" box="[199,322,1951,1977]" pageId="27" pageNumber="28">Diagnosis.</emphasis>
One character in male genitalia shared by
<taxonomicName id="4C75A6E3CC2A4C3FF455F84FFC6AF80C" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[797,1012,1951,1977]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="michaelgrzimeki">
<emphasis id="B9010172CC2A4C3FF455F84FFC6AF80C" box="[797,1012,1951,1977]" italics="true" pageId="27" pageNumber="28">L. michaelgrzimeki</emphasis>
</taxonomicName>
and
<taxonomicName id="4C75A6E3CC2A4C3FF361F84FFB59F80C" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[1065,1223,1951,1977]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="lutztoepferi">
<emphasis id="B9010172CC2A4C3FF361F84FFB59F80C" box="[1065,1223,1951,1977]" italics="true" pageId="27" pageNumber="28">L. lutztoepferi</emphasis>
</taxonomicName>
is the dense covering of the ventral side of the uncus with long setae. However, the species most similar to
<taxonomicName id="4C75A6E3CC2A4C3FF3C3F82BFAFDF868" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[1163,1379,1987,2013]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="michaelgrzimeki">
<emphasis id="B9010172CC2A4C3FF3C3F82BFAFDF868" box="[1163,1379,1987,2013]" italics="true" pageId="27" pageNumber="28">L. michaelgrzimeki</emphasis>
</taxonomicName>
with respect to the male genitalia is
<taxonomicName id="4C75A6E3CC2D4C38F540FF78FD15FF04" authorityName="Lehmann &amp; Zahiri &amp; Husemann" authorityYear="2023" box="[520,651,151,177]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="kammeri">
<emphasis id="B9010172CC2D4C38F540FF78FD15FF04" box="[520,651,151,177]" italics="true" pageId="28" pageNumber="29">L. kammeri</emphasis>
</taxonomicName>
(diagnosis below).
<taxonomicName id="4C75A6E3CC2D4C38F43BFF78FB33FF04" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[883,1197,151,177]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="michaelgrzimeki">
<emphasis id="B9010172CC2D4C38F43BFF78FB33FF04" box="[883,1197,151,177]" italics="true" pageId="28" pageNumber="29">Lukeniana michaelgrzimeki</emphasis>
</taxonomicName>
has three characters unique among all
<taxonomicName id="4C75A6E3CC2D4C38F62CFF54FDA9FF60" box="[356,567,187,213]" pageId="28" pageNumber="29">
<emphasis id="B9010172CC2D4C38F62CFF54FE44FF60" box="[356,474,187,213]" italics="true" pageId="28" pageNumber="29">Lukeniana</emphasis>
species
</taxonomicName>
: (i) the thorn-like process of the valva has many dots with tiny setae; (ii) the weakly-sclerotized projection has a prickly appearance due to many long setae; and (iii) the costa and the sacculus have scattered long setae. In the female postabdominal structure cover the ventral bands more than 100% of the ventral side of segment 8 (in ventral view). They are similar in shape and structure to those of
<taxonomicName id="4C75A6E3CC2D4C38F3F1FEC8FAFAFEF4" authorityName="Lehmann &amp; Zahiri &amp; Husemann" authorityYear="2023" box="[1209,1380,295,321]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="mikerobertsi">
<emphasis id="B9010172CC2D4C38F3F1FEC8FAFAFEF4" box="[1209,1380,295,321]" italics="true" pageId="28" pageNumber="29">L. mikerobertsi</emphasis>
</taxonomicName>
. The corpus bursae is elliptical in
<taxonomicName id="4C75A6E3CC2D4C38F697FEA4FD29FED0" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[479,695,331,357]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="michaelgrzimeki">
<emphasis id="B9010172CC2D4C38F697FEA4FD29FED0" box="[479,695,331,357]" italics="true" pageId="28" pageNumber="29">L. michaelgrzimeki</emphasis>
</taxonomicName>
but round in
<taxonomicName id="4C75A6E3CC2D4C38F41CFEA4FC61FED0" authorityName="Lehmann &amp; Zahiri &amp; Husemann" authorityYear="2023" box="[852,1023,331,357]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="mikerobertsi">
<emphasis id="B9010172CC2D4C38F41CFEA4FC61FED0" box="[852,1023,331,357]" italics="true" pageId="28" pageNumber="29">L. mikerobertsi</emphasis>
</taxonomicName>
. Additionally, the base of the posterior apophysis is only 1.5 broader than the width of the base of the anterior apophyses in
<taxonomicName id="4C75A6E3CC2D4C38F336FE80FACBFE3C" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[1150,1365,367,393]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="michaelgrzimeki">
<emphasis id="B9010172CC2D4C38F336FE80FACBFE3C" box="[1150,1365,367,393]" italics="true" pageId="28" pageNumber="29">L. michaelgrzimeki</emphasis>
</taxonomicName>
, but 4 broader than the width of the base of the anterior apophyses in
<taxonomicName id="4C75A6E3CC2D4C38F411FE7CFB9DFE18" authorityName="Lehmann &amp; Zahiri &amp; Husemann" authorityYear="2023" box="[857,1027,403,429]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="mikerobertsi">
<emphasis id="B9010172CC2D4C38F411FE7CFB9DFE18" box="[857,1027,403,429]" italics="true" pageId="28" pageNumber="29">L. mikerobertsi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC2D4C38F78FFE59FE77FDD4" pageId="28" pageNumber="29" type="distribution">
<paragraph id="8BCADD60CC2D4C38F78FFE59FE10FD88" blockId="28.[151,1437,151,789]" pageId="28" pageNumber="29">
<emphasis id="B9010172CC2D4C38F78FFE59FEFFFE65" bold="true" box="[199,353,438,464]" pageId="28" pageNumber="29">Distribution.</emphasis>
<taxonomicName id="4C75A6E3CC2D4C38F620FE58FD00FE64" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[360,670,439,465]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="michaelgrzimeki">
<emphasis id="B9010172CC2D4C38F620FE58FD00FE64" box="[360,670,439,465]" italics="true" pageId="28" pageNumber="29">Lukeniana michaelgrzimeki</emphasis>
</taxonomicName>
is known only from Momella (also “Momela”, elevation
<emphasis id="B9010172CC2D4C38F26EFE58FAD6FE65" box="[1318,1352,439,464]" italics="true" pageId="28" pageNumber="29">ca.</emphasis>
<quantity id="4C8D7085CC2D4C38F207FE58FF68FE40" metricMagnitude="3" metricUnit="m" metricValue="1.521" metricValueMax="1.61" metricValueMin="1.432" pageId="28" pageNumber="29" unit="m" value="1521.0" valueMax="1610.0" valueMin="1432.0">
<geoCoordinate id="EE41BBA7CC2D4C38F207FE58FA13FE64" box="[1359,1421,439,465]" degrees="1,432" direction="north" orientation="latitude" pageId="28" pageNumber="29" precision="55" value="1.432">1,432</geoCoordinate>
<geoCoordinate id="EE41BBA7CC2D4C38F2C5FE58FF48FE40" degrees="1,610" direction="west" orientation="longitude" pageId="28" pageNumber="29" precision="55" value="-1.61"> 1,610</geoCoordinate>
m
</quantity>
), which is mostly within the
<collectingRegion id="49B11382CC2D4C38F57DFE34FD18FE40" box="[565,646,475,501]" country="Tanzania" name="Arusha" pageId="28" pageNumber="29">Arusha</collectingRegion>
National Park, and located
<emphasis id="B9010172CC2D4C38F4F3FE34FC42FE41" box="[955,988,475,500]" italics="true" pageId="28" pageNumber="29">ca.</emphasis>
<quantity id="4C8D7085CC2D4C38F4ABFE35FBB7FE40" box="[995,1065,474,501]" metricMagnitude="4" metricUnit="m" metricValue="5.5" pageId="28" pageNumber="29" unit="km" value="55.0">55 km</quantity>
southwest of
<collectingRegion id="49B11382CC2D4C38F38CFE34FA02FE40" box="[1220,1436,475,501]" country="Tanzania" name="Kilimanjaro" pageId="28" pageNumber="29">Mount Kilimanjaro</collectingRegion>
on the eastern foothills of Mount Meru (north-central
<collectingCountry id="F3629DF0CC2D4C38F5A0FE10FCD1FDAC" box="[744,847,511,537]" name="Tanzania" pageId="28" pageNumber="29">Tanzania</collectingCountry>
) (
<figureCitation id="134EC1E5CC2D4C38F42CFE10FC28FDAC" box="[868,950,511,537]" captionStart="FIGURE 19" captionStartId="71.[151,250,150,174]" captionText="FIGURE 19 a. Collection sites of species of Lukeniana; 2. Lukeniana rajaeii; 3. L. timdavenporti; 4. L. kakamegaensis; 5. / 6. L. mikerobertsi; 7. / 8. L. enaiposha.; 9 / 11. L. obliqualinea; 10. L. kammeri; 11. L. lutztoepferi; 12. L. georgeadamsoni; 13. / 15. L. bergsteni; 14. L. michaelgrzimeki / L. andreashempi; 16. L. tubiraensis; 17. L. jankiellandi; 18. L. tubiraensis; 19. L. mbalaensis / L. robplowesi; 20. L. hausmanni; 21. L. stueningi; 22. L. stevecollinsi; 23. L. mzuzuensis; 24. / 25. L. madrandelei; 26. L. lenzi; 27. L. carolae; 28. L. chapmani; 29. L. friederikebauerae; 30. / 31. / 32. L. butleri; 34. L. utaheidenreichae; 38. L. kollhorsti; 39. / 40. / 41. L. raymondrevellii (location codes in the EARS sensu McConnell (1972), Eastern Branch and Western Branch, and associated areas sensu Oxburgh (1978), as well as on the Southern African Plateau sensu Nyblade &amp; Sleep (2003); for detailed information on geological/geomorphological ages for all areas herein cf. Lehmann (2019b, 330─341): 2. Main Ethiopian Rift (MER), Ethiopian Plateau (ca. 31 Ma), Ethiopia; 3. west of Kenya Rift (KR), Mount Elgon (ca. 20 Ma), border of Uganda and Kenya; 4. west of KR, Kakamega (ca. 20 Ma), Kenya; 5. east of KR, on Laikipia Plateau (ca. 13.5─11 Ma), Kenya; 6. east of KR, eastern edge of Laikipia Plateau (ca. 13.5─11 Ma), Kenya; 7. within KR, near Lake Nakuru (ca. 2.6 Ma), Kenya; 8. within KR, near Lake Naivasha (ca. 2.6 Ma), Kenya; 9. east of KR, foothills zone of Mount Kenya (ca. 5.1─2.8 Ma), Kenya; 10. / 11. / 12. east of KR, Kenya Dome (ca. 13.5─11 Ma), Kenya; 13. within Northern Tanzanian Divergence Zone (NTDZ), Mount Kilimanjaro (ca. 2.5 Ma), Tanzania; 14. / 15. within NTDZ, Mount Meru and Usa River (ca. 2.0 Ma), Tanzania; 16. / 17. east of Tanganyika Rift (ca. 30─20 Ma), Tanzania; 18. within Rukwa Rift, Southern Highlands (ca. 30─25 Ma), Tan- zania; 19. / 20. northeastern edge of Southern African Plateau (ca. 150 Ma); Zambia; 21. / 22. / 23. west of Malawi Rift, on and south of Nyika Plateau (ca. 30─25 Ma), Zambia, Malawi; 24. / 25. south and west of various rifts of EARS of unknown ages, Democratic Republic of the Congo; 26. west of Luangwa Graben, escarpment of Muchinga Mountains (ca. 150 Ma?), Zambia; 27. west of “Malawi Rift, Mount Ntchisi (ca. 25 Ma), Malawi; 28. Mount Mulanje (ca. 105─138 Ma), Malawi; 29. south of Luangwa Graben (ca. 150 Ma), Zambia; 30. / 31. / 32. south of Mid-Zambezi Rift, Highveld Plateau (ca. 150 Ma), Zimbabwe; 34. east of Bvumba Mountains (younger than ca. 15 Ma), Mozambique; 38. west of Urema Graben (younger than ca. 15 Ma), Zimbabwe; 39. northwest from Waterberg Plateau (ca. 75 Ma), Namibia; 40. on Waterberg Plateau (ca. 75 Ma), Namibia; 41. west-central edge of Southern African Plateau (ca. 150 Ma), Namibia. b. Collection sites of species of Metarbelodes (turquoise circle, white font) and Zambesia (red circles, black font): 1. Zambezia diredaouaensis; 30. / 32. /36. Zambezia jennyhuntae sp. nov.; 35. Zambezia durrellbarnesi; 33. / 36. Metarbelodes umtaliana; 36. Z. darrelplowesi; 37. Z. madambae (location codes: 1. East of MER, Somalian Plateau including Harar Plateau (at least 30 Ma), Ethiopia; 30. / 32. south of Mid-Zambezi Rift, Highveld Plateau (ca. 150 Ma), Zimbabwe; 33. / 35. / 36. areas near and on the Bvumba Mountains, Eastern Highlands of Zimbabwe (ca. 600─500 Ma), Zimbabwe; 37. ca. 2.3 km from Zambezi River (ca. 280 Ma) and on the termination of Southwestern Branch of EARS (ca. 2 Ma), Botswana; for detailed information on geological/geomorphological ages for all areas herein Lehmann (2019b, 330─341))." pageId="28" pageNumber="29">Fig. 19</figureCitation>
). The species is considered an Afromontane near-endemic species.
</paragraph>
<paragraph id="8BCADD60CC2D4C38F78FFDA9FE77FDD4" blockId="28.[151,1437,151,789]" box="[199,489,582,609]" pageId="28" pageNumber="29">
<emphasis id="B9010172CC2D4C38F78FFDA9FEB4FDD5" bold="true" box="[199,298,582,608]" pageId="28" pageNumber="29">Habitat.</emphasis>
See Appendix 1.
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC2D4C38F78FFD84FDF0FCA0" pageId="28" pageNumber="29" type="etymology">
<paragraph id="8BCADD60CC2D4C38F78FFD84FDF0FCA0" blockId="28.[151,1437,151,789]" pageId="28" pageNumber="29">
<emphasis id="B9010172CC2D4C38F78FFD84FED0FD30" bold="true" box="[199,334,619,645]" pageId="28" pageNumber="29">Etymology.</emphasis>
<taxonomicName id="4C75A6E3CC2D4C38F61EFD84FD10FD30" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[342,654,619,645]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="michaelgrzimeki">
<emphasis id="B9010172CC2D4C38F61EFD84FD10FD30" box="[342,654,619,645]" italics="true" pageId="28" pageNumber="29">Lukeniana michaelgrzimeki</emphasis>
</taxonomicName>
is named for the German cinematographer Michael Grzimek (1934 1959). Grzimek died near the Sanjan River Gorge during the filming of “Serengeti Shall Not Die” when he collided with his legendary Dornier 27 (rebuilt in the year 2000 on the Schacksdorf-Finsterwalde Airport) with a Rüppells Griffon Vulture,
<taxonomicName id="4C75A6E3CC2D4C38F61AFD38FD79FD44" authority="(A. E. Brehm, 1852)" baseAuthorityName="A. E. Brehm" baseAuthorityYear="1852" box="[338,743,726,753]" class="Aves" family="Accipitridae" genus="Gyps" kingdom="Animalia" order="Accipitriformes" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="rueppellii">
<emphasis id="B9010172CC2D4C38F61AFD38FD9EFD44" box="[338,512,727,753]" italics="true" pageId="28" pageNumber="29">Gyps rueppellii</emphasis>
(A.E.
<bibRefCitation id="EFE4A091CC2D4C38F502FD38FD7EFD44" author="Brehm, A. E." box="[586,736,726,753]" pageId="28" pageNumber="29" pagination="38 - 51" refId="ref75730" refString="Brehm, A. E. (1852) Beitrage zur Ornithologie Nord-Ost-Afrika's, mit besonderer Rucksicht auf die in Europa vorkommenden Arten der Vogel. In: Baldamus, E. (Ed.), Naumannia - Archiv fur die Ornithologie. Band 2. Heft 3. s. n., Stuttgart, pp. 38 - 51." type="book chapter" year="1852">Brehm, 1852</bibRefCitation>
)
</taxonomicName>
, once a commonly occurring species (
<bibRefCitation id="EFE4A091CC2D4C38F3D9FD38FAC3FD44" author="Brown, L. H. &amp; Urban, E. K. &amp; Newman, K." box="[1169,1373,727,753]" pageId="28" pageNumber="29" refId="ref75793" refString="Brown, L. H., Urban, E. K. &amp; Newman, K. (1982) The Birds of Africa. Vol. 1. Academic Press, London and New York, xiii + 521 pp." type="book" year="1982">
Brown
<emphasis id="B9010172CC2D4C38F3AEFD38FA80FD44" box="[1254,1310,727,753]" italics="true" pageId="28" pageNumber="29">et al.</emphasis>
1982
</bibRefCitation>
) now at risk of extinction (
<bibRefCitation id="EFE4A091CC2D4C38F6C8FD14FDFCFCA0" author="Nature Kenya" box="[384,610,763,789]" pageId="28" pageNumber="29" pagination="1 - 2" refId="ref81425" refString="Nature Kenya (2016) Africa's vultures are sliding towards extinction warns BirdLife. Nature net (Nature Kenya Newsletter), Dec 2015 - Jan 2016, 1 - 2." type="journal article" year="2016">
Nature
<collectingCountry id="F3629DF0CC2D4C38F69DFD14FD80FCA0" box="[469,542,763,789]" name="Kenya" pageId="28" pageNumber="29">Kenya</collectingCountry>
2016
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>