treatments-xml/data/03/C6/3E/03C63E676805FFAAFF0001D8FBFC07E9.xml

<document id="A5CF6311E33A322023B9499031F59813" ID-DOI="10.5281/zenodo.5328899" ID-ISSN="0253-116X" ID-Zenodo-Dep="5328899" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630160147779" checkinUser="carolina" docAuthor="Casale, A., D. W &amp; P, Antalya" docDate="2012" docId="03C63E676805FFAAFF0001D8FBFC07E9" docLanguage="en" docName="LinzerbiolBeitr.2012.44.2.1111-1127.pdf" docOrigin="Linzer biologische Beiträge 44 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Laemostenus (Pristonychus) bellicosus Casale &amp; D.W &amp; P 2012, nov.sp." docType="treatment" docVersion="3" lastPageNumber="1121" masterDocId="FFFF461F680CFFA0FFB2033BFFE2035B" masterDocTitle="New or little known Laemostenus species from the Near and Middle East (Coleoptera, Carabidae: Sphodrini)" masterLastPageNumber="1127" masterPageNumber="1111" pageNumber="1120" updateTime="1699169291185" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="BC836A0CE78C5605C21F409E8ADC7E0A">New or little known Laemostenus species from the Near and Middle East (Coleoptera, Carabidae: Sphodrini)</mods:title>
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<mods:namePart id="A570EC917541673EDC135A8F2901AE0F">Casale, A.</mods:namePart>
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<mods:namePart id="88B13CD7AB1D6E850B4F2B703DD0EDF0">D. W</mods:namePart>
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<mods:namePart id="58428FC65E95FBEC04FC1687CD3930BF">P, Antalya</mods:namePart>
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<mods:title id="2CE43C348BBD2A091A972F60E6DEC8CD">Linzer biologische Beiträge</mods:title>
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<paragraph id="8BD08F716805FFA9FF0001D8FD7901A3" blockId="9.[178,667,738,760]" box="[178,667,738,760]" pageId="9" pageNumber="1120">
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<taxonomicName id="4C6FF4F26805FFA9FF0001D8FDAB01A3" ID-CoL="8M5S3" authority="Casale &amp; D.W &amp; P, 2012" authorityName="Casale &amp; D.W &amp; P" authorityYear="2012" box="[178,585,738,760]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1120" phylum="Arthropoda" rank="species" species="bellicosus" status="sp. nov." subGenus="Pristonychus">
<emphasis id="B91B53636805FFA9FF0001D8FEDE01A3" bold="true" box="[178,316,739,760]" italics="true" pageId="9" pageNumber="1120">Laemostenus</emphasis>
(
<emphasis id="B91B53636805FFA9FEF801D9FE3001A3" bold="true" box="[330,466,738,760]" italics="true" pageId="9" pageNumber="1120">Pristonychus</emphasis>
) 
<emphasis id="B91B53636805FFA9FE5001D9FDAB01A3" bold="true" box="[482,585,738,760]" italics="true" pageId="9" pageNumber="1120">bellicosus</emphasis>
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<paragraph id="8BD08F716805FFA9FF000029FD940065" blockId="9.[178,1085,786,830]" pageId="9" pageNumber="1120">
<materialsCitation id="3B07852C6805FFA9FF000029FD940065" collectingDate="1998-06-27" collectionCode="T" collectorName="P. Kabatek" country="Syria" elevation="610" latitude="34.75" location="Crac des Chevaliers" longLatPrecision="1197" longitude="36.283333" pageId="9" pageNumber="1120" specimenCount="1" typeStatus="holotype">
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y p e m a t e r i a l: 
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, labelled: &quot; 
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occ., 
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<geoCoordinate id="EE5BE9B66805FFA9FC2C0029FC14007D" box="[926,1014,786,806]" degrees="34" direction="north" minutes="45" orientation="latitude" pageId="9" pageNumber="1120" precision="925" value="34.75">34° 45’ N</geoCoordinate>
<geoCoordinate id="EE5BE9B66805FFA9FC4E0029FF0C0065" degrees="36" direction="east" minutes="17" orientation="longitude" pageId="9" pageNumber="1120" precision="925" value="36.283333">36° 17’ E</geoCoordinate>
, 
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<collectingDate id="EF9550596805FFA9FF450011FEB90065" box="[247,347,810,830]" pageId="9" pageNumber="1120" value="1998-06-27">27.VI.1998</collectingDate>
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leg. 
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, 
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&quot; (cWR).
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</paragraph>
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<paragraph id="8BD08F716805FFA9FF000076FE7C00C6" blockId="9.[178,1085,844,925]" pageId="9" pageNumber="1120">
D i a g n o s i s: A medium-sized, completely piceous-black 
<taxonomicName id="4C6FF4F26805FFA9FC0B0076FEA40024" authority="" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1120" phylum="Arthropoda" rank="subGenus" subGenus="Pristonychus">
<emphasis id="B91B53636805FFA9FC0B0076FBDF0039" box="[953,1085,845,866]" italics="true" pageId="9" pageNumber="1120">Laemostenus</emphasis>
(
<emphasis id="B91B53636805FFA9FF080051FEDC00DB" box="[186,318,874,896]" italics="true" pageId="9" pageNumber="1120">Pristonychus</emphasis>
)
</taxonomicName>
species of the 
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<emphasis id="B91B53636805FFA9FE540051FDB800DB" box="[486,602,874,896]" italics="true" pageId="9" pageNumber="1120">L. terricola</emphasis>
</taxonomicName>
species group (in the widest sense of 
<bibRefCitation id="EFFEF2806805FFA9FC5E0051FF0A00C6" author="CASALE A" pageId="9" pageNumber="1120" refId="ref8179" refString="CASALE A. (1988): Revisione degli Sphodrina (Coleoptera, Carabidae, Sphodrini). - Museo regionale di Scienze naturali, Torino, Monografie 5: 1024 pp." type="book" year="1988">CASALE 1988</bibRefCitation>
). Habitus 
<figureCitation id="135493F46805FFA9FEFD00BCFE7C00C6" box="[335,414,903,925]" captionStart="Figs 22-25" captionStartId="16.[178,218,1506,1526]" captionTargetBox="[266,976,232,1475]" captionTargetId="figure-64@16.[178,1085,212,1495]" captionTargetPageId="16" captionText="Figs 22-25: Laemostenus, habitus. (22) L. croyi nov.sp., paratype. (23) L. phoenicius nov.sp., paratype (Maїfouq). (24) L. eggeri nov.sp., paratype. (25) L. bellicosus nov.sp., holotype (various magnifications)." figureDoi="http://doi.org/10.5281/zenodo.5328905" httpUri="https://zenodo.org/record/5328905/files/figure.png" pageId="9" pageNumber="1120">Fig. 25.</figureCitation>
</paragraph>
<paragraph id="8BD08F716805FFA9FF000096FEE9070E" blockId="9.[178,1085,940,1109]" pageId="9" pageNumber="1120">
E t y m o l o g y: The Latin specific epithet (
<emphasis id="B91B53636805FFA9FD3F0097FD100099" box="[653,754,940,962]" italics="true" pageId="9" pageNumber="1120">bellicosus</emphasis>
: bellicose, belligerent) indicates the strategic rule of the 
<typeStatus id="54D431D36805FFA9FE1400F1FE300084" box="[422,466,970,991]" pageId="9" pageNumber="1120">type</typeStatus>
locality of this species, the famous medieval crusader castle Crac (incorrect: Krak) des Chevaliers in 
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(in Arabic language: Qal'at al-Hosn), that T.E. Lawrence (better known as &quot;Lawrence of Arabia&quot;) described as being &quot;the most impressive fortress in the world&quot;. The site is now included in the UNESCO world heritage.
</paragraph>
<paragraph id="8BD08F716805FFA9FF00075FFBD20721" blockId="9.[178,1072,1124,1146]" box="[178,1072,1124,1146]" pageId="9" pageNumber="1120">D e s c r i p t i o n: Body medium-sized 18.0 mm (TL) and 19.0 mm (L), respectively.</paragraph>
<paragraph id="8BD08F716805FFA9FF0007B2FE2207AC" blockId="9.[178,1086,1161,1271]" pageId="9" pageNumber="1120">Colour: Piceous black; labrum, mandibles, palpomeres, and sutural interval of elytra dark reddish brown; elytra without bluish reflection. Surface opaque, elytra not sericeous. Microsculpure: head and pronotum with transverse, evident microlines; elytra with distinct, isodiametric meshes.</paragraph>
<paragraph id="8BD08F716805FFA9FF00063DFC470662" blockId="9.[178,1085,1286,1337]" pageId="9" pageNumber="1120">Head narrow, elongate, with distinct transverse wrinkles; eyes small, shorter than tempora, not prominent laterally; frontal impressions wide, shallow, wrinkled.</paragraph>
<paragraph id="8BD08F716805FFA9FF000672FBFA068F" blockId="9.[178,1085,1353,1492]" pageId="9" pageNumber="1120">Pronotum (Fig. 15) narrow (PL/PW: 0.91), cordate, its lateral sides moderately reflexed and markedly sinuate in front of the basolateral angles, which are acute and prominent outside; anterolateral angles small, but markedly prominent; base fully beaded. Disc flattened, with deep transverse wrinkles; basal impressions elongate, deep, each with some large, deep punctures; anterolateral and basolateral setiferous punctures present.</paragraph>
<paragraph id="8BD08F716805FFA9FF0006DFFE3D06A2" blockId="9.[178,479,1507,1529]" box="[178,479,1507,1529]" pageId="9" pageNumber="1120">Mesosternum not denticulate.</paragraph>
<paragraph id="8BD08F716805FFA9FF000533FBDF0567" blockId="9.[178,1085,1544,1596]" pageId="9" pageNumber="1120">Elytra elongate-ovate (EL/EW: 1.71), subconvex. Base narrow; basal ridge weakly concave; humeral tooth and shoulders absent. Striae deep, smooth; intervals slightly</paragraph>
<paragraph id="8BD08F716806FFAAFF0003E0FDE30256" blockId="10.[178,1085,219,270]" pageId="10" pageNumber="1121">convex. Chaetotaxy: Basal pore present; umbilicate series with 24-26 setiferous punctures; 2-3 setae at apex of stria 7.</paragraph>
<paragraph id="8BD08F716806FFAAFF000225FE9802B8" blockId="10.[178,1086,285,483]" pageId="10" pageNumber="1121">
Legs long and slender; profemora on ventral side longitudinally shallowly concave for entire lengtht, with both external and internal margins smooth, their outer side with 3 short setae, the apical one reaching the outer margin of femur; mesotibiae almost straight (female 
<typeStatus id="54D431D36806FFAAFEB4024EFE8102D0" box="[262,355,373,395]" pageId="10" pageNumber="1121" type="holotype">holotype</typeStatus>
); metatibiae each with apical brush of rather long, yellow reddish setae; tarsomeres with dorsal pubescence dense, short; metatarsomers short, thickened. Tarsal claws with a series of very small teeth, reduced in number (4-5), along the basal half of the internal margin.
</paragraph>
<paragraph id="8BD08F716806FFAAFF0002C9FE5E0153" blockId="10.[178,444,498,520]" box="[178,444,498,520]" pageId="10" pageNumber="1121">Male genitalia: Unknown.</paragraph>
<paragraph id="8BD08F716806FFAAFF00012CFEC80110" blockId="10.[178,1085,535,587]" pageId="10" pageNumber="1121">Female genitalia: Internal tract, spermatheca, and gonocoxites 1 and 2 of ovipositor as in Figs 17, 18.</paragraph>
<paragraph id="8BD08F716806FFAAFF000161FDE00001" blockId="10.[178,1086,602,858]" pageId="10" pageNumber="1121">
C o m p a r i s o n s: The new species is related to the Anatolian species 
<taxonomicName id="4C6FF4F26806FFAAFC460160FE3A01D6" authority="(G. MULLER 1931)" baseAuthorityName="G. MULLER" baseAuthorityYear="1931" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1121" phylum="Arthropoda" rank="species" species="pisidicus" subGenus="Pristonychus">
<emphasis id="B91B53636806FFAAFC460160FBEB012B" box="[1012,1033,603,624]" italics="true" pageId="10" pageNumber="1121">L.</emphasis>
(
<emphasis id="B91B53636806FFAAFBAC0160FBD7012B" box="[1054,1077,603,624]" italics="true" pageId="10" pageNumber="1121">P.</emphasis>
) 
<emphasis id="B91B53636806FFAAFF00014CFEEE01D6" box="[178,268,631,653]" italics="true" pageId="10" pageNumber="1121">pisidicus</emphasis>
(G. MÜLLER 1931)
</taxonomicName>
, from which it differs by the more elongate head, the more reduced, not prominent eyes, the narrower, markedly cordiform pronotum, with anterolateral angles prominent in front and basolateral angles prominent outside, the more elongate elytra, with smooth striae and shoulders and humeral tooth completely absent. It is not related to other 
<taxonomicName id="4C6FF4F26806FFAAFE1C01D6FD650059" box="[430,647,748,770]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1121" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B91B53636806FFAAFE1C01D6FDD00059" box="[430,562,749,770]" italics="true" pageId="10" pageNumber="1121">Laemostenus</emphasis>
species
</taxonomicName>
reported from this area (from Crac des Chevaliers, 
<taxonomicName id="4C6FF4F26806FFAAFE9E0031FDE5007B" baseAuthorityName="REICHE" baseAuthorityYear="1855" box="[300,519,778,800]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1121" phylum="Arthropoda" rank="species" species="parallelocollis" subGenus="Laemostenus">
<emphasis id="B91B53636806FFAAFE9E0031FEA30044" box="[300,321,778,799]" italics="true" pageId="10" pageNumber="1121">L.</emphasis>
(
<emphasis id="B91B53636806FFAAFEE20031FE870044" box="[336,357,778,799]" italics="true" pageId="10" pageNumber="1121">L.</emphasis>
) 
<emphasis id="B91B53636806FFAAFEC60031FDE5007B" box="[372,519,778,800]" italics="true" pageId="10" pageNumber="1121">parallelocollis</emphasis>
</taxonomicName>
[REICHE 1855] was cited by 
<bibRefCitation id="EFFEF2806806FFAAFC810031FC20007B" author="CASALE A" box="[819,962,778,800]" pageId="10" pageNumber="1121" refId="ref8179" refString="CASALE A. (1988): Revisione degli Sphodrina (Coleoptera, Carabidae, Sphodrini). - Museo regionale di Scienze naturali, Torino, Monografie 5: 1024 pp." type="book" year="1988">CASALE 1988</bibRefCitation>
), but apparently closer to taxa distributed in Southern 
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and in other areas of 
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(see 
<taxonomicName id="4C6FF4F26806FFAAFB9A0013FF100001" authority="Casale &amp; D.W &amp; P, 2012" authorityName="Casale &amp; D.W &amp; P" authorityYear="2012" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1121" phylum="Arthropoda" rank="species" species="eggeri" status="sp. nov.">
<emphasis id="B91B53636806FFAAFB9A0013FF100001" italics="true" pageId="10" pageNumber="1121">L. eggeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A228EE186806FFAAFF4B007EFEA70001" box="[249,325,837,858]" pageId="10" pageNumber="1121" rank="species">nov.sp.</taxonomicNameLabel>
, described above).
</paragraph>
<paragraph id="8BD08F716806FFAAFF000051FC4E0749" blockId="10.[178,1085,873,1042]" pageId="10" pageNumber="1121">R e m a r k s: This species is a further, unexpected new taxon in a well known site of the Eastern Mediterranean area, sampled in a locality famous and visited by both tourists and entomologists for many years. Its apparent rarity could be attributed to its putative subterranean way of life, indicated by some morphological features, as the slender habitus, the small-sized eyes, the long appendages, the reduced denticulation of the internal side of tarsal claws, and the absence of bluish, metallic reflection on elytra.</paragraph>
<paragraph id="8BD08F716806FFAAFF000719FE1D0729" blockId="10.[178,1085,1057,1138]" pageId="10" pageNumber="1121">
D i s t r i b u t i o n a n d h a b i t a t: This markedly characterized species is so far known from only 
<specimenCount id="9D6944F86806FFAAFE3E0704FE19070F" box="[396,507,1086,1108]" pageId="10" pageNumber="1121" type="female">one female</specimenCount>
sampled in the 
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locality, at 
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altitude. We have no information about the habitat.
</paragraph>
<paragraph id="8BD08F716806FFAAFF0007A7FBFC07E9" blockId="10.[178,1054,1180,1229]" box="[178,1054,1180,1202]" pageId="10" pageNumber="1121">
<emphasis id="B91B53636806FFAAFF0007A7FBFC07E9" bold="true" box="[178,1054,1180,1202]" pageId="10" pageNumber="1121">
Additional note on a 
<taxonomicName id="4C6FF4F26806FFAAFE2707A6FD5F07E9" authority="" box="[405,701,1180,1202]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1121" phylum="Arthropoda" rank="subGenus" subGenus="Pristonychus">
<emphasis id="B91B53636806FFAAFE2707A6FDFD07E9" bold="true" box="[405,543,1181,1202]" italics="true" pageId="10" pageNumber="1121">Laemostenus</emphasis>
(
<emphasis id="B91B53636806FFAAFD9F07A7FD5707E9" bold="true" box="[557,693,1180,1202]" italics="true" pageId="10" pageNumber="1121">Pristonychus</emphasis>
)
</taxonomicName>
species of the 
<taxonomicName id="4C6FF4F26806FFAAFCEB07A6FC2F07E9" baseAuthorityName="Herbst" baseAuthorityYear="1784" box="[857,973,1180,1202]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1121" phylum="Arthropoda" rank="species" species="terricola">
<emphasis id="B91B53636806FFAAFCEB07A6FC2F07E9" bold="true" box="[857,973,1180,1202]" italics="true" pageId="10" pageNumber="1121">L. terricola</emphasis>
</taxonomicName>
species
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>