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<document id="DF94F7EAF82604652CA0EC30184F863A" ID-DOI="10.11646/zootaxa.3609.5.7" ID-GBIF-Dataset="569c5ee1-2e43-46f7-9f9e-98b2d78da23f" ID-ISSN="1175-5326" ID-Zenodo-Dep="220607" ID-ZooBank="C959B588-0833-47C5-BD1F-D510E884FB3D" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460573025187" checkinUser="plazi" docAuthor="Liu, Xingyue, Hayashi, Fumio &amp; Yang, Ding" docDate="2013" docId="03AFEF04DB51FFB6928B9FACFEEAB6E0" docLanguage="en" docName="zt03609p519.pdf" docOrigin="Zootaxa 3609 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Protohermes sonus Liu, Hayashi &amp; Yang, 2013, sp. nov." docType="treatment" docVersion="8" lastPageNumber="516" masterDocId="FF96977CDB50FFB2921C9902FFBAB634" masterDocTitle="The Protohermes latus species group (Megaloptera: Corydalidae), with description of two new species from India and Myanmar" masterLastPageNumber="519" masterPageNumber="513" pageNumber="514" updateTime="1698322571802" updateUser="plazi">
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<mods:title id="B68C904758A96DCB0C581B40475EAC81">The Protohermes latus species group (Megaloptera: Corydalidae), with description of two new species from India and Myanmar</mods:title>
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<mods:name id="BE7FCD5E5F8DE4818CE6C1C5DAB01558" type="personal">
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<mods:namePart id="DBC7FD26F4C8E9CFFA7F8D5234A1143F">Liu, Xingyue</mods:namePart>
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<mods:namePart id="645BA3B93AC411722C79166D101C6382">Hayashi, Fumio</mods:namePart>
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<mods:namePart id="CADDE99ED40C0508781AF581E939C4AF">Yang, Ding</mods:namePart>
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<mods:title id="910518C740A658B40009A016FE853E7B">Zootaxa</mods:title>
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<mods:date id="68A6173AAF4F7607FD48397295C86229">2013</mods:date>
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<treatment id="03AFEF04DB51FFB6928B9FACFEEAB6E0" ID-DOI="http://doi.org/10.5281/zenodo.6157629" ID-GBIF-Taxon="119559197" ID-Zenodo-Dep="6157629" LSID="urn:lsid:plazi:treatment:03AFEF04DB51FFB6928B9FACFEEAB6E0" httpUri="http://treatment.plazi.org/id/03AFEF04DB51FFB6928B9FACFEEAB6E0" lastPageId="4" lastPageNumber="516" pageId="1" pageNumber="514">
<subSubSection id="C31C0D99DB51FFB3928B9FACFEF0B0DD" pageId="1" pageNumber="514" type="nomenclature">
<paragraph id="8BB95E12DB51FFB3928B9FACFE5EB0FC" blockId="1.[151,484,1710,1769]" box="[151,484,1710,1736]" pageId="1" pageNumber="514">
<heading id="D0F1E97EDB51FFB3928B9FACFE5EB0FC" bold="true" box="[151,484,1710,1736]" fontSize="11" level="1" pageId="1" pageNumber="514" reason="1">
<emphasis id="B9728200DB51FFB3928B9FACFE5EB0FC" bold="true" box="[151,484,1710,1736]" pageId="1" pageNumber="514">
<taxonomicName id="4C062591DB51FFB3928B9FACFEC5B0FC" ID-CoL="4N5JL" box="[151,383,1710,1736]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="1" pageNumber="514" phylum="Arthropoda" rank="species" species="sonus" status="sp. nov.">
<emphasis id="B9728200DB51FFB3928B9FACFEC5B0FC" bold="true" box="[151,383,1710,1736]" italics="true" pageId="1" pageNumber="514">Protohermes sonus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2413F7BDB51FFB3939A9FACFE5EB0FC" box="[390,484,1710,1736]" pageId="1" pageNumber="514" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB95E12DB51FFB3928B9FD3FEF0B0DD" blockId="1.[151,484,1710,1769]" box="[151,330,1745,1769]" pageId="1" pageNumber="514">
(
<figureCitation id="133D4297DB51FFB392829FD3FEB3B0DD" box="[158,265,1745,1769]" captionStart="FIGURE 1 2" captionStartId="2.[151,250,1628,1651]" captionTargetBox="[151,1429,214,1589]" captionTargetId="figure@2.[151,1436,195,1604]" captionTargetPageId="2" captionText="FIGURE 1 2. Habitus of Protohermes spp. 1. Protohermes sonus sp. nov., male holotype; 2. Protohermes sublunatus sp. nov. Scale bar = 5.0 mm." httpUri="https://zenodo.org/record/220608/files/figure.png" pageId="1" pageNumber="514">Figures 1</figureCitation>
,
<figureCitation id="133D4297DB51FFB3930B9FD3FEFBB0DD" box="[279,321,1745,1769]" captionStart="FIGURES 3 5" captionStartId="3.[151,264,810,833]" captionTargetBox="[167,1412,206,765]" captionTargetId="figure@3.[151,1436,193,789]" captionTargetPageId="3" captionText="FIGURES 3 5. Protohermes sonus sp. nov., male holotype. 3. genitalia, dorsal view; 4. genitalia, ventral view; 5. tenth gonocoxite, ventral view. c: cercus, e: ectoproct, gs 9: gonostylus 9, S 9: ninth sternum, T 9: ninth tergum. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/220609/files/figure.png" pageId="1" pageNumber="514">35</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31C0D99DB51FFB3928B9E1BFE37B1A9" pageId="1" pageNumber="514" type="diagnosis">
<paragraph id="8BB95E12DB51FFB3928B9E1BFE37B1A9" blockId="1.[151,1436,1817,1986]" pageId="1" pageNumber="514">
<emphasis id="B9728200DB51FFB3928B9E1BFEABB106" bold="true" box="[151,273,1817,1842]" pageId="1" pageNumber="514">Diagnosis.</emphasis>
Head pale brown with anterior margin of clypeus yellow. Pronotum reddish brown, with anterior ridge black, strongly broadened in male. Wings immaculate, slightly grayish, with costal area slightly darker. Male ectoproct bilobed, dorsal lobe transversely broadened, much shorter than ninth tergum, ventral lobe band-like, visible in dorsal view.
</paragraph>
</subSubSection>
<subSubSection id="C31C0D99DB51FFB192D59EABFE25B05B" lastPageId="3" lastPageNumber="516" pageId="1" pageNumber="514" type="description">
<paragraph id="8BB95E12DB51FFB392D59EABFB7AB1F5" blockId="1.[151,1436,1817,1986]" box="[201,1216,1961,1986]" pageId="1" pageNumber="514">
<emphasis id="B9728200DB51FFB392D59EABFEE0B1F6" bold="true" box="[201,346,1961,1986]" pageId="1" pageNumber="514">Description.</emphasis>
Male. Body length
<quantity id="4CFEF3F7DB51FFB390259EABFD31B1F5" box="[569,651,1961,1985]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="514" unit="mm" value="50.0">50 mm</quantity>
; forewing length
<quantity id="4CFEF3F7DB51FFB391539EABFC18B1F5" box="[847,930,1961,1985]" metricMagnitude="-2" metricUnit="m" metricValue="5.9" pageId="1" pageNumber="514" unit="mm" value="59.0">59 mm</quantity>
, hindwing length
<quantity id="4CFEF3F7DB51FFB396769EABFB06B1F5" box="[1130,1212,1961,1985]" metricMagnitude="-2" metricUnit="m" metricValue="4.9" pageId="1" pageNumber="514" unit="mm" value="49.0">49 mm</quantity>
.
</paragraph>
<caption id="DF790E9ADB52FFB0928B9F5EFEDEB0A5" httpUri="https://zenodo.org/record/220608/files/figure.png" pageId="2" pageNumber="515" targetBox="[151,1429,214,1589]" targetPageId="2">
<paragraph id="8BB95E12DB52FFB0928B9F5EFEDEB0A5" blockId="2.[151,1435,1628,1681]" pageId="2" pageNumber="515">
<emphasis id="B9728200DB52FFB0928B9F5EFEB6B045" bold="true" box="[151,268,1628,1651]" pageId="2" pageNumber="515">FIGURE 1</emphasis>
<emphasis id="B9728200DB52FFB093059F5EFE9CB045" bold="true" box="[281,294,1628,1649]" pageId="2" pageNumber="515">2</emphasis>
. Habitus of
<taxonomicName id="4C062591DB52FFB093BE9F5EFD9EB046" box="[418,548,1628,1650]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="2" pageNumber="515" phylum="Arthropoda" rank="genus">
<emphasis id="B9728200DB52FFB093BE9F5EFD9EB046" box="[418,548,1628,1650]" italics="true" pageId="2" pageNumber="515">Protohermes</emphasis>
</taxonomicName>
spp. 1.
<taxonomicName id="4C062591DB52FFB0906F9F5EFC8FB046" box="[627,821,1628,1650]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="2" pageNumber="515" phylum="Arthropoda" rank="species" species="sonus" status="sp. nov.">
<emphasis id="B9728200DB52FFB0906F9F5EFC8FB046" box="[627,821,1628,1650]" italics="true" pageId="2" pageNumber="515">Protohermes sonus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2413F7BDB52FFB091279F5FFC31B046" box="[827,907,1629,1650]" pageId="2" pageNumber="515" rank="species">
<emphasis id="B9728200DB52FFB091279F5FFC31B046" bold="true" box="[827,907,1629,1650]" pageId="2" pageNumber="515">sp. nov.</emphasis>
</taxonomicNameLabel>
, male holotype; 2.
<taxonomicName id="4C062591DB52FFB096539F5EFAFFB046" box="[1103,1349,1628,1650]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="2" pageNumber="515" phylum="Arthropoda" rank="species" species="sublunatus" status="sp. nov.">
<emphasis id="B9728200DB52FFB096539F5EFAFFB046" box="[1103,1349,1628,1650]" italics="true" pageId="2" pageNumber="515">Protohermes sublunatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2413F7BDB52FFB097579F5FFA21B046" box="[1355,1435,1629,1650]" pageId="2" pageNumber="515" rank="species">
<emphasis id="B9728200DB52FFB097579F5FFA21B046" bold="true" box="[1355,1435,1629,1650]" pageId="2" pageNumber="515">sp. nov.</emphasis>
</taxonomicNameLabel>
Scale bar = 5.0 mm.
</paragraph>
</caption>
<caption id="DF790E9ADB53FFB1928B9A28FA30B56B" httpUri="https://zenodo.org/record/220609/files/figure.png" pageId="3" pageNumber="516" targetBox="[167,1412,206,765]" targetPageId="3">
<paragraph id="8BB95E12DB53FFB1928B9A28FA30B56B" blockId="3.[151,1436,810,863]" pageId="3" pageNumber="516">
<emphasis id="B9728200DB53FFB1928B9A28FEA5B574" bold="true" box="[151,287,810,833]" pageId="3" pageNumber="516">FIGURES 3</emphasis>
<emphasis id="B9728200DB53FFB193379A29FE82B574" bold="true" box="[299,312,811,832]" pageId="3" pageNumber="516">5</emphasis>
.
<taxonomicName id="4C062591DB53FFB193549A29FDB4B575" box="[328,526,811,833]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="3" pageNumber="516" phylum="Arthropoda" rank="species" species="sonus" status="sp. nov.">
<emphasis id="B9728200DB53FFB193549A29FDB4B575" box="[328,526,811,833]" italics="true" pageId="3" pageNumber="516">Protohermes sonus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2413F7BDB53FFB190059A29FDD7B574" box="[537,621,811,832]" pageId="3" pageNumber="516" rank="species">
<emphasis id="B9728200DB53FFB190059A29FDD7B574" bold="true" box="[537,621,811,832]" pageId="3" pageNumber="516">sp. nov.</emphasis>
</taxonomicNameLabel>
, male holotype. 3. genitalia, dorsal view; 4. genitalia, ventral view; 5. tenth gonocoxite, ventral view. c: cercus, e: ectoproct, gs9: gonostylus 9, S9: ninth sternum, T9: ninth tergum. Scale bar = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BB95E12DB53FFB192D59A85FC1DB23F" blockId="3.[151,1437,903,2008]" pageId="3" pageNumber="516">
Head (
<figureCitation id="133D4297DB53FFB1930A9A85FEE0B5AB" box="[278,346,903,927]" captionStart="FIGURE 1 2" captionStartId="2.[151,250,1628,1651]" captionTargetBox="[151,1429,214,1589]" captionTargetId="figure@2.[151,1436,195,1604]" captionTargetPageId="2" captionText="FIGURE 1 2. Habitus of Protohermes spp. 1. Protohermes sonus sp. nov., male holotype; 2. Protohermes sublunatus sp. nov. Scale bar = 5.0 mm." httpUri="https://zenodo.org/record/220608/files/figure.png" pageId="3" pageNumber="516">Fig. 1</figureCitation>
) pale brown, immaculate, with anterior margin of clypeus yellow; postocular spine short and bluntly produced. Compound eyes pale brown; ocelli pale yellow, medially margined black, lateral ocelli nearly touching median ocellus. Antenna subserrate, black, with scape and pedicel pale brown. Mouthparts yellow; mandibles reddish brown with apex and inner margin blackish brown.
</paragraph>
<paragraph id="8BB95E12DB53FFB192D59D15FCB6B31F" blockId="3.[151,1437,903,2008]" pageId="3" pageNumber="516">
Prothorax (
<figureCitation id="133D4297DB53FFB193559D15FE31B21B" box="[329,395,1047,1071]" captionStart="FIGURE 1 2" captionStartId="2.[151,250,1628,1651]" captionTargetBox="[151,1429,214,1589]" captionTargetId="figure@2.[151,1436,195,1604]" captionTargetPageId="2" captionText="FIGURE 1 2. Habitus of Protohermes spp. 1. Protohermes sonus sp. nov., male holotype; 2. Protohermes sublunatus sp. nov. Scale bar = 5.0 mm." httpUri="https://zenodo.org/record/220608/files/figure.png" pageId="3" pageNumber="516">Fig. 1</figureCitation>
) reddish brown, pronotum strongly broadened on anterolateral and posterolateral portions, and anteriorly produced into a black ridge. Meso- and metathorax yellow. Legs reddish brown with dense, yellowish, short setae; tibiae and tarsi blackish brown, except for distal three tarsomeres yellow; tarsal claws reddish brown. Wings hyaline, slightly grayish, immaculate, except for costal cellules with indistinct grayish stripes; veins yellow, forewing with bases of costal vein and bases of first and second anal veins black. Rs 10- branched; MA distally bifurcate; anterior branch of MP 5-branched in forewings but 4-branched in hindwings, posterior branch of MP 2-branched; CuA 4 or 5-branched; humeral area well developed, with a feebly developed humeral crossvein; 1114 crossveins between R and Rs.
</paragraph>
<paragraph id="8BB95E12DB53FFB192D59C35FB46B07F" blockId="3.[151,1437,903,2008]" pageId="3" pageNumber="516">
Abdomen blackish brown. Ninth tergum (
<figureCitation id="133D4297DB53FFB190AC9C35FD4DB37B" box="[688,759,1335,1359]" captionStart="FIGURES 3 5" captionStartId="3.[151,264,810,833]" captionTargetBox="[167,1412,206,765]" captionTargetId="figure@3.[151,1436,193,789]" captionTargetPageId="3" captionText="FIGURES 3 5. Protohermes sonus sp. nov., male holotype. 3. genitalia, dorsal view; 4. genitalia, ventral view; 5. tenth gonocoxite, ventral view. c: cercus, e: ectoproct, gs 9: gonostylus 9, S 9: ninth sternum, T 9: ninth tergum. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/220609/files/figure.png" pageId="3" pageNumber="516">Fig. 3</figureCitation>
) subtrapezoidal, anterior margin nearly truncate, posterior margin medially concaved. Ninth sternum (
<figureCitation id="133D4297DB53FFB190609C59FD05B347" box="[636,703,1371,1395]" captionStart="FIGURES 3 5" captionStartId="3.[151,264,810,833]" captionTargetBox="[167,1412,206,765]" captionTargetId="figure@3.[151,1436,193,789]" captionTargetPageId="3" captionText="FIGURES 3 5. Protohermes sonus sp. nov., male holotype. 3. genitalia, dorsal view; 4. genitalia, ventral view; 5. tenth gonocoxite, ventral view. c: cercus, e: ectoproct, gs 9: gonostylus 9, S 9: ninth sternum, T 9: ninth tergum. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/220609/files/figure.png" pageId="3" pageNumber="516">Fig. 4</figureCitation>
) about 1.5 times as long as ninth tergum, but much narrower than ninth tergum, posterior margin deeply incised, forming a pair of broadly subtriangular lobes. Ninth gonostylus (
<figureCitation id="133D4297DB53FFB197739C7DFF1CB38F" captionStart="FIGURES 3 5" captionStartId="3.[151,264,810,833]" captionTargetBox="[167,1412,206,765]" captionTargetId="figure@3.[151,1436,193,789]" captionTargetPageId="3" captionText="FIGURES 3 5. Protohermes sonus sp. nov., male holotype. 3. genitalia, dorsal view; 4. genitalia, ventral view; 5. tenth gonocoxite, ventral view. c: cercus, e: ectoproct, gs 9: gonostylus 9, S 9: ninth sternum, T 9: ninth tergum. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/220609/files/figure.png" pageId="3" pageNumber="516">Fig. 4</figureCitation>
) unguiform and strongly curved inward, with sclerotized claw extremely slender and elongated. Ectoproct (
<figureCitation id="133D4297DB53FFB197789CA1FF78B3EB" captionStart="FIGURES 3 5" captionStartId="3.[151,264,810,833]" captionTargetBox="[167,1412,206,765]" captionTargetId="figure@3.[151,1436,193,789]" captionTargetPageId="3" captionText="FIGURES 3 5. Protohermes sonus sp. nov., male holotype. 3. genitalia, dorsal view; 4. genitalia, ventral view; 5. tenth gonocoxite, ventral view. c: cercus, e: ectoproct, gs 9: gonostylus 9, S 9: ninth sternum, T 9: ninth tergum. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/220609/files/figure.png" pageId="3" pageNumber="516">Figs. 34</figureCitation>
) bilobed dorsoventrally; dorsal lobe transversely wide, much shorter than ninth tergum, with straight outer margin and arcuate inner margin; ventral lobe visible in dorsal view, elongate and band-like, much longer than dorsal lobe as well as ninth tergum, slightly narrowed toward apex. Cercus invisible in dorsal view. Tenth gonocoxite (
<figureCitation id="133D4297DB53FFB1933E9F31FEDFB07F" box="[290,357,1587,1611]" captionStart="FIGURES 3 5" captionStartId="3.[151,264,810,833]" captionTargetBox="[167,1412,206,765]" captionTargetId="figure@3.[151,1436,193,789]" captionTargetPageId="3" captionText="FIGURES 3 5. Protohermes sonus sp. nov., male holotype. 3. genitalia, dorsal view; 4. genitalia, ventral view; 5. tenth gonocoxite, ventral view. c: cercus, e: ectoproct, gs 9: gonostylus 9, S 9: ninth sternum, T 9: ninth tergum. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/220609/files/figure.png" pageId="3" pageNumber="516">Fig. 5</figureCitation>
) extremely small, arched, with lateral lobes short, digitiform and ventrally setose.
</paragraph>
<paragraph id="8BB95E12DB53FFB192D59F55FE25B05B" blockId="3.[151,1437,903,2008]" box="[201,415,1623,1647]" pageId="3" pageNumber="516">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C31C0D99DB53FFB192D59F79FAE4B0A7" box="[201,1374,1658,1683]" pageId="3" pageNumber="516" type="materials_examined">
<paragraph id="8BB95E12DB53FFB192D59F79FAE4B0A7" blockId="3.[151,1437,903,2008]" box="[201,1374,1658,1683]" pageId="3" pageNumber="516">
<emphasis id="B9728200DB53FFB192D59F79FECDB0A7" bold="true" box="[201,375,1658,1683]" pageId="3" pageNumber="516">
<typeStatus id="54BDE0B0DB53FFB192D59F79FEBEB0A7" box="[201,260,1659,1683]" pageId="3" pageNumber="516">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54BDE0B0DB53FFB193619F79FE5FB0A7" box="[381,485,1659,1683]" pageId="3" pageNumber="516" type="holotype">Holotype</typeStatus>
3,
<collectingCountry id="F3111E82DB53FFB1900C9F79FD1DB0A7" box="[528,679,1659,1683]" name="Myanmar" pageId="3" pageNumber="516">MYANMAR</collectingCountry>
:
<collectingCountry id="F3111E82DB53FFB190AD9F79FD5DB0A7" box="[689,743,1659,1683]" name="China" pageId="3" pageNumber="516">Chin</collectingCountry>
State, near Tiddim [23°17ʹN, 93°41ʹE],
<date id="FFB878D2DB53FFB196B69F79FABDB0A7" box="[1194,1287,1659,1683]" pageId="3" pageNumber="516" value="2009-06">VI.2009</date>
(HFC).
</paragraph>
</subSubSection>
<subSubSection id="C31C0D99DB53FFB192D59F9CFD94B083" box="[201,558,1694,1720]" pageId="3" pageNumber="516" type="distribution">
<paragraph id="8BB95E12DB53FFB192D59F9CFD94B083" blockId="3.[151,1437,903,2008]" box="[201,558,1694,1720]" pageId="3" pageNumber="516">
<emphasis id="B9728200DB53FFB192D59F9CFED8B083" bold="true" box="[201,354,1694,1719]" pageId="3" pageNumber="516">Distribution.</emphasis>
<collectingCountry id="F3111E82DB53FFB193759FA2FE6CB08C" box="[361,470,1696,1720]" name="Myanmar" pageId="3" pageNumber="516">Myanmar</collectingCountry>
(
<collectingCountry id="F3111E82DB53FFB193F99F9DFD9BB083" box="[485,545,1695,1719]" name="China" pageId="3" pageNumber="516">Chin</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C31C0D99DB53FFB192D59FC1FBDCB0CB" pageId="3" pageNumber="516" type="etymology">
<paragraph id="8BB95E12DB53FFB192D59FC1FBDCB0CB" blockId="3.[151,1437,903,2008]" pageId="3" pageNumber="516">
<emphasis id="B9728200DB53FFB192D59FC1FEEAB0EF" bold="true" box="[201,336,1731,1755]" pageId="3" pageNumber="516">Etymology.</emphasis>
The specific epithet
<taxonomicName id="4C062591DB53FFB190539FC6FD35B0EF" box="[591,655,1732,1755]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="3" pageNumber="516" phylum="Arthropoda" rank="species" species="sonus">
<emphasis id="B9728200DB53FFB190539FC6FD35B0EF" box="[591,655,1732,1755]" italics="true" pageId="3" pageNumber="516">sonus</emphasis>
</taxonomicName>
means sound and refers to the male call during courtship possibly made by the beating between broadened pronotum and humeral areas of the forewings.
</paragraph>
</subSubSection>
<subSubSection id="C31C0D99DB53FFB692D59E09FEEAB6E0" lastPageId="4" lastPageNumber="517" pageId="3" pageNumber="516" type="discussion">
<paragraph id="8BB95E12DB53FFB692D59E09FEEAB6E0" blockId="3.[151,1437,903,2008]" lastBlockId="4.[151,1436,151,212]" lastPageId="4" lastPageNumber="517" pageId="3" pageNumber="516">
<emphasis id="B9728200DB53FFB192D59E09FE87B117" bold="true" box="[201,317,1803,1827]" pageId="3" pageNumber="516">Remarks.</emphasis>
The new species distinctly differs from
<taxonomicName id="4C062591DB53FFB191299E0EFC34B117" box="[821,910,1803,1827]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="3" pageNumber="516" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="B9728200DB53FFB191299E0EFC34B117" box="[821,910,1803,1827]" italics="true" pageId="3" pageNumber="516">P. latus</emphasis>
</taxonomicName>
by the male ectoproct, which has a short, transversely wide dorsal lobe and an elongate, band-like ventral lobe (
<figureCitation id="133D4297DB53FFB191D99E2DFBB6B173" box="[965,1036,1839,1863]" captionStart="FIGURES 3 5" captionStartId="3.[151,264,810,833]" captionTargetBox="[167,1412,206,765]" captionTargetId="figure@3.[151,1436,193,789]" captionTargetPageId="3" captionText="FIGURES 3 5. Protohermes sonus sp. nov., male holotype. 3. genitalia, dorsal view; 4. genitalia, ventral view; 5. tenth gonocoxite, ventral view. c: cercus, e: ectoproct, gs 9: gonostylus 9, S 9: ninth sternum, T 9: ninth tergum. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/220609/files/figure.png" pageId="3" pageNumber="516">Fig. 3</figureCitation>
), and by the strongly curved male ninth gonostylus (
<figureCitation id="133D4297DB53FFB1937E9E51FE13B15F" box="[354,425,1875,1899]" captionStart="FIGURES 3 5" captionStartId="3.[151,264,810,833]" captionTargetBox="[167,1412,206,765]" captionTargetId="figure@3.[151,1436,193,789]" captionTargetPageId="3" captionText="FIGURES 3 5. Protohermes sonus sp. nov., male holotype. 3. genitalia, dorsal view; 4. genitalia, ventral view; 5. tenth gonocoxite, ventral view. c: cercus, e: ectoproct, gs 9: gonostylus 9, S 9: ninth sternum, T 9: ninth tergum. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/220609/files/figure.png" pageId="3" pageNumber="516">Fig. 4</figureCitation>
). In
<taxonomicName id="4C062591DB53FFB193C29E56FD8BB15F" box="[478,561,1875,1899]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="3" pageNumber="516" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="B9728200DB53FFB193C29E56FD8BB15F" box="[478,561,1875,1899]" italics="true" pageId="3" pageNumber="516">P. latus</emphasis>
</taxonomicName>
the male ectoproct has a rather broad and rounded dorsal lobe, which covers the digitiform ventral lobe (see Liu &amp; Yang 2006:
<figureCitation id="133D4297DB53FFB190D49E75FCEAB1BB" box="[712,848,1911,1935]" captionStart="FIGURES 9 14" captionStartId="5.[151,264,1894,1917]" captionTargetBox="[361,1214,826,1854]" captionTargetId="figure@5.[347,1230,807,1869]" captionTargetPageId="5" captionText="FIGURES 9 14. Pronotum and base of forewing costal area of Protohermes spp. 9. P. latus Liu &amp; Yang, male; 10. same, female; 11. P. sublunatus sp. nov., male; 12. same, female; 13. P. s o n u s sp. nov., male; 14. P. davidi van der Weele, male. Arrows show well developed humeral area." httpUri="https://zenodo.org/record/220611/files/figure.png" pageId="3" pageNumber="516">Figs. 1314</figureCitation>
). Compared with
<taxonomicName id="4C062591DB53FFB1960B9E7AFB11B1BB" box="[1047,1195,1911,1935]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="3" pageNumber="516" phylum="Arthropoda" rank="species" species="sublunatus" status="sp. nov.">
<emphasis id="B9728200DB53FFB1960B9E7AFB11B1BB" box="[1047,1195,1911,1935]" italics="true" pageId="3" pageNumber="516">P. sublunatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2413F7BDB53FFB196AE9E75FAB1B1BB" box="[1202,1291,1911,1935]" pageId="3" pageNumber="516" rank="species">
<emphasis id="B9728200DB53FFB196AE9E75FAB1B1BB" bold="true" box="[1202,1291,1911,1935]" pageId="3" pageNumber="516">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<taxonomicName id="4C062591DB53FFB197059E7AFACEB1BB" box="[1305,1396,1912,1935]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="3" pageNumber="516" phylum="Arthropoda" rank="species" species="sonus" status="sp. nov.">
<emphasis id="B9728200DB53FFB197059E7AFACEB1BB" box="[1305,1396,1912,1935]" italics="true" pageId="3" pageNumber="516">P. sonus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2413F7BDB53FFB197669E75FF72B187" pageId="3" pageNumber="516" rank="species">
<emphasis id="B9728200DB53FFB197669E75FF72B187" bold="true" pageId="3" pageNumber="516">sp. nov.</emphasis>
</taxonomicNameLabel>
has a subtrapezoidal posterior incision on male ninth tergum (
<figureCitation id="133D4297DB53FFB191969E99FC77B187" box="[906,973,1947,1971]" captionStart="FIGURES 3 5" captionStartId="3.[151,264,810,833]" captionTargetBox="[167,1412,206,765]" captionTargetId="figure@3.[151,1436,193,789]" captionTargetPageId="3" captionText="FIGURES 3 5. Protohermes sonus sp. nov., male holotype. 3. genitalia, dorsal view; 4. genitalia, ventral view; 5. tenth gonocoxite, ventral view. c: cercus, e: ectoproct, gs 9: gonostylus 9, S 9: ninth sternum, T 9: ninth tergum. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/220609/files/figure.png" pageId="3" pageNumber="516">Fig. 3</figureCitation>
) and an ectoproct with nearly suboblong dorsal lobe and band-like ventral lobe (
<figureCitation id="133D4297DB53FFB190539EBDFD06B1E3" box="[591,700,1983,2007]" captionStart="FIGURES 3 5" captionStartId="3.[151,264,810,833]" captionTargetBox="[167,1412,206,765]" captionTargetId="figure@3.[151,1436,193,789]" captionTargetPageId="3" captionText="FIGURES 3 5. Protohermes sonus sp. nov., male holotype. 3. genitalia, dorsal view; 4. genitalia, ventral view; 5. tenth gonocoxite, ventral view. c: cercus, e: ectoproct, gs 9: gonostylus 9, S 9: ninth sternum, T 9: ninth tergum. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/220609/files/figure.png" pageId="3" pageNumber="516">Figs. 34</figureCitation>
). In
<taxonomicName id="4C062591DB53FFB190F29EC2FC39B1E3" box="[750,899,1983,2007]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="3" pageNumber="516" phylum="Arthropoda" rank="species" species="sublunatus" status="sp. nov.">
<emphasis id="B9728200DB53FFB190F29EC2FC39B1E3" box="[750,899,1983,2007]" italics="true" pageId="3" pageNumber="516">P. sublunatus</emphasis>
</taxonomicName>
<emphasis id="B9728200DB53FFB191969EBDFC5EB1E3" bold="true" box="[906,996,1983,2007]" pageId="3" pageNumber="516">
<taxonomicNameLabel id="A2413F7BDB53FFB191969EBDFC5EB1E3" box="[906,996,1983,2007]" pageId="3" pageNumber="516" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, the male ninth tergum has a deeply Vshaped posterior incision (
<figureCitation id="133D4297DB54FFB693DC9995FDBFB69B" box="[448,517,151,175]" captionStart="FIGURES 6 8" captionStartId="4.[151,264,1759,1782]" captionTargetBox="[170,1420,1081,1721]" captionTargetId="figure@4.[151,1436,1055,1738]" captionTargetPageId="4" captionText="FIGURES 6 8. Protohermes sublunatus sp. nov., male holotype. 6. genitalia, dorsal view; 7. genitalia, ventral view; 8. tenth gonocoxite, ventral view. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/220610/files/figure.png" pageId="4" pageNumber="517">Fig. 6</figureCitation>
), and the ectoproct has nearly semilune dorsal lobe and thickly digitiform ventral lobe (
<figureCitation id="133D4297DB54FFB692CA99BEFEF9B6E0" box="[214,323,188,212]" captionStart="FIGURES 6 8" captionStartId="4.[151,264,1759,1782]" captionTargetBox="[170,1420,1081,1721]" captionTargetId="figure@4.[151,1436,1055,1738]" captionTargetPageId="4" captionText="FIGURES 6 8. Protohermes sublunatus sp. nov., male holotype. 6. genitalia, dorsal view; 7. genitalia, ventral view; 8. tenth gonocoxite, ventral view. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/220610/files/figure.png" pageId="4" pageNumber="517">Figs. 67</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>