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<document id="CD9A68DA71045DBE40B6E308F38BF270" ID-DOI="10.12651/JSR.2017.6" ID-ISSN="2713-8615" ID-Zenodo-Dep="13161827" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722552417796" checkinUser="felipe" docAuthor="Lee, Sue Yeon, Yoo, Jung Sun &amp; Kim, Seung Tae" docDate="2017" docId="03AFE01C083B7D36AC6BB77E6CF4FE02" docLanguage="en" docName="JSpeciesRes.6.Special.185-214.pdf" docOrigin="Journal of Species Research 6" docSource="http://dx.doi.org/10.5281/zenodo.13161684" docStyle="DocumentStyle:4D398BFF45FF8CA9E08F546BB06AE372.2:JSpeciesRes.2012-.journal_article" docStyleId="4D398BFF45FF8CA9E08F546BB06AE372" docStyleName="JSpeciesRes.2012-.journal_article" docStyleVersion="2" docTitle="Phyllopodopsyllus busanensis Lee, Yoo &amp; Kim, 2017, sp. nov." docType="treatment" docVersion="2" lastPageNumber="208" masterDocId="FF969864082B7D21AF2CB273680FFFDD" masterDocTitle="Two new Phyllopodopsyllus (Copepoda, Harpacticoida) from Korean marine interstitial" masterLastPageNumber="214" masterPageNumber="185" pageNumber="201" updateTime="1722570211887" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="6E42B22217837DF214C972FC9854D355">Two new Phyllopodopsyllus (Copepoda, Harpacticoida) from Korean marine interstitial</mods:title>
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<mods:namePart id="86D682B5BEBEF420E4AAF091A67509A8">Lee, Sue Yeon</mods:namePart>
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<mods:namePart id="932D7DE19B6CD76F2728E522332C9449">Yoo, Jung Sun</mods:namePart>
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<mods:namePart id="812223D7893F1C5D346D2D759FD81ACF">Kim, Seung Tae</mods:namePart>
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<mods:title id="EA5522E7197E4FD927D2DB9E860AA81C">Journal of Species Research</mods:title>
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<mods:date id="1BF9DAF03CA79EE30ABF9C7A5D4CB236">2017</mods:date>
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<treatment id="03AFE01C083B7D36AC6BB77E6CF4FE02" LSID="urn:lsid:plazi:treatment:03AFE01C083B7D36AC6BB77E6CF4FE02" httpUri="http://treatment.plazi.org/id/03AFE01C083B7D36AC6BB77E6CF4FE02" lastPageId="23" lastPageNumber="208" pageId="16" pageNumber="201">
<subSubSection id="C31C0281083B7D31AC6BB77E6CE4FAFB" box="[839,1259,1293,1318]" pageId="16" pageNumber="201" type="nomenclature">
<paragraph id="8BB9510A083B7D31AC6BB77E6CE4FAFB" blockId="16.[839,1259,1293,1350]" box="[839,1259,1293,1318]" pageId="16" pageNumber="201">
<heading id="D0F1E666083B7D31AC6BB77E6CE4FAFB" bold="true" box="[839,1259,1293,1318]" fontSize="10" level="2" pageId="16" pageNumber="201" reason="4">
<emphasis id="B9728D18083B7D31AC6BB77E6CE4FAFB" bold="true" box="[839,1259,1293,1318]" pageId="16" pageNumber="201">
<taxonomicName id="4C062A89083B7D31AC6BB77E6C9FFAF8" ID-CoL="B6CPX" authorityName="Lee &amp; Yoo &amp; Kim" authorityYear="2017" box="[839,1168,1293,1317]" class="Copepoda" family="Tetragonicipitidae" genus="Phyllopodopsyllus" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="201" phylum="Arthropoda" rank="species" species="busanensis" status="sp. nov.">
<emphasis id="B9728D18083B7D31AC6BB77E6C9FFAF8" bold="true" box="[839,1168,1293,1317]" italics="true" pageId="16" pageNumber="201">Phyllopodopsyllus busanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2413063083B7D31ABBAB77D6CE4FAFB" box="[1174,1259,1294,1318]" pageId="16" pageNumber="201" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31C0281083B7D31AC67B75D6C3FFA9B" box="[843,1072,1326,1350]" pageId="16" pageNumber="201" type="description">
<paragraph id="8BB9510A083B7D31AC67B75D6C3FFA9B" blockId="16.[839,1259,1293,1350]" box="[843,1072,1326,1350]" pageId="16" pageNumber="201">
<emphasis id="B9728D18083B7D31AC67B75D6C3FFA9B" bold="true" box="[843,1072,1326,1350]" pageId="16" pageNumber="201">
(
<figureCitation id="133D4D8F083B7D31AC7FB75D6BBDFA9B" box="[851,946,1326,1350]" captionStart="Fig" captionStartId="3.[144,177,1234,1256]" captionTargetBox="[177,1380,257,1202]" captionTargetId="figure-358@3.[177,1380,257,1202]" captionTargetPageId="3" captionText="Fig. 1. Light photograph of two new Korean Phyllopodopsyllus Scot T., 1906 species, on the same slide in glycerol:A, P. kitazimai sp. nov., holotype female; B, P. kitazimai sp. nov., allotype male; C, P. busanensis sp. nov., holotype female." pageId="16" pageNumber="201">Figs. 1C</figureCitation>
and
<figureCitation id="133D4D8F083B7D31ACC6B75D6C27FA9B" box="[1002,1064,1326,1350]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStartId-0="17.[144,177,1843,1865]" captionStartId-1="18.[174,207,1843,1865]" captionStartId-2="19.[144,177,1424,1446]" captionStartId-3="20.[174,207,1910,1932]" captionStartId-4="21.[144,177,1727,1749]" captionTargetBox-0="[223,1334,257,1816]" captionTargetBox-1="[253,1364,258,1816]" captionTargetBox-2="[178,1372,263,1388]" captionTargetBox-4="[178,1348,262,1697]" captionTargetId-0="figure-10@17.[223,1334,257,1816]" captionTargetId-1="figure-11@18.[253,1364,258,1816]" captionTargetId-2="figure-199@19.[189,1429,262,1396]" captionTargetId-4="figure-97@21.[167,1404,261,1697]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionTargetPageId-2="19" captionTargetPageId-4="21" captionText-0="Fig. 11. Phyllopodopsyllus busanensis sp. nov., paratype female, SEM photographs: A, habitus, lateral; B, cephalothorax (Ct), lateral; C, anterior part of cephalothorax (Ct) with rostrum(Ro) and first segment of antennula (A1), lateral; D, ventral part of cephalothorax (Ct), with parts of antennula (A1), antenna (A2), and mandibula, lateral; E, dorso-posterior part of cephalothorax (Ct), lateral; F, ventro-posterior corner of cephalothorax (Ct), lateral; G, tergites of three free prosomites (Fp1-Fp3), lateral; H, detail of ornamentation of first (Fp1) and second (Fp2) prosomites, lateral. Arabic numerals for cuticular sensilla and Roman numerals for pores assigned consecutively from anterior to posterior end of each somite and caudal ramus, and from dorsal to ventral side." captionText-1="Fig. 12. Phyllopodopsyllus busanensis sp. nov., paratype female, SEM photographs: A, first five urosomites (U1-U5) with fifth leg (P5), lateral; B, tergite of first urosomite (U1), lateral; C, parts of fused second (U2) and third (U3) urosomites, with lateral part of fifth leg (P5), lateral; D, part of fourth urosomite (U4) and distal part of fifth leg (P5); E, fourth (U4) and fifth (U5) urosomites, lateral; F, sixth urosomite and caudal rami(Cr), lateral; G, posterior part of caudal rami(Cr), lateral; H, parts of maxilliped (Mxp) and first swimming leg (P1), lateral. Arabic numerals for cuticular sensilla and Roman numerals for pores assigned consecutively from anterior to posterior end of each somite and caudal ramus, and from dorsal to ventral side." captionText-2="Fig. 13. Phyllopodopsyllus busanensis sp. nov., holotype female, line drawings: A, second to sixth urosomites (U2-U6) and caudal rami, dorsal; B, second to sixth urosomites (U2-U6), with caudal rami (Cr), and attached spermatophore (Sp), lateral; C, second to sixth urosomites (U2-U6), with caudal rami (Cr), sixth legs (P6), attached spermatophore (Sp), and internal reseptacula seminis (Rs), ventral. Arabic numerals for cuticular sensilla and Roman numerals for pores assigned consecutively from anterior to posterior end of each somite and caudal ramus, and from dorsal to ventral side. Cuticular pits and minute surface spinules omitted for clarity." captionText-3="Fig. 14. Phyllopodopsyllus busanensis sp. nov., holotype female, line drawings: A, antennula, dorsal. B, antenna, posterior; C, mandibula, anterior; D, praecoxal arthrite of maxillula, anterior; E, coxa of maxillula, anterior; F, endopod of maxillula, anterior; G, maxilla, posterior; H, maxilliped, anterior. Cuticular pits on antennula and antenna omitted for clarity." captionText-4="Fig. 15. Phyllopodopsyllus busanensis sp. nov., holotype female, line drawings: A, first swimming leg, anterior; B, second swimming leg, anterior; C, third swimming leg, anterior; D, fourth swimming leg, anterior; E, fifth leg, compressed and slightly deformed, anterior." pageId="16" pageNumber="201">11-15</figureCitation>
)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C31C0281083B7D31AC6BB71C6BB1F955" pageId="16" pageNumber="201" type="materials_examined">
<paragraph id="8BB9510A083B7D31AC6BB71C6BEFFA3A" blockId="16.[839,1444,1390,1995]" pageId="16" pageNumber="201">
<materialsCitation id="3B6E5B57083B7D31AC6BB71C6BD3FA3A" country="South Korea" county="East Coast" latitude="35.179016" location="Songjeong Beach" longLatPrecision="1" longitude="129.20528" municipality="Gyeongnam-do province" pageId="16" pageNumber="201" specimenCount="1" stateProvince="Busan" typeStatus="holotype">
<emphasis id="B9728D18083B7D31AC6BB71C6BEFFA5B" bold="true" box="[839,992,1390,1415]" pageId="16" pageNumber="201">
<typeStatus id="54BDEFA8083B7D31AC6BB71C6B70FA5A" box="[839,895,1391,1415]" pageId="16" pageNumber="201">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F311119A083B7D31ACC4B71C6C23FA5B" box="[1000,1068,1391,1414]" name="South Korea" pageId="16" pageNumber="201">Korea</collectingCountry>
,
<collectingCounty id="62D82986083B7D31AB15B71C6CA1FA5B" box="[1081,1198,1390,1414]" pageId="16" pageNumber="201">East Coast</collectingCounty>
,
<collectingMunicipality id="6BDDCB70083B7D31AB97B71D6B79FA7A" pageId="16" pageNumber="201">Gyeongnam-do province</collectingMunicipality>
,
<collectingRegion id="49C29FE8083B7D31ACAFB7FC6BC6FA7B" box="[899,969,1423,1446]" country="South Korea" name="Busan" pageId="16" pageNumber="201">Busan</collectingRegion>
,
<location id="8ED907D1083B7D31ACFAB7FC6C97FA7A" LSID="urn:lsid:plazi:treatment:03AFE01C083B7D36AC6BB77E6CF4FE02:8ED907D1083B7D31ACFAB7FC6C97FA7A" box="[982,1176,1423,1447]" country="South Korea" county="East Coast" latitude="35.179016" longLatPrecision="1" longitude="129.20528" municipality="Gyeongnam-do province" name="Songjeong Beach" pageId="16" pageNumber="201" stateProvince="Busan">Songjeong Beach</location>
, intertidal sand, interstitial water from a
<location id="8ED907D1083B7D31AB2BB7DC6CD9FA1A" LSID="urn:lsid:plazi:treatment:03AFE01C083B7D36AC6BB77E6CF4FE02:8ED907D1083B7D31AB2BB7DC6CD9FA1A" box="[1031,1238,1455,1479]" country="South Korea" county="East Coast" latitude="35.179016" longLatPrecision="1" longitude="129.20528" municipality="Gyeongnam-do province" name="Karaman-Chappuis" pageId="16" pageNumber="201" stateProvince="Busan">Karaman-Chappuis</location>
hole,
<geoCoordinate id="EE3237CD083B7D31AA30B7DC6DABFA1B" box="[1308,1444,1453,1479]" degrees="35" direction="north" minutes="10.741" orientation="latitude" pageId="16" pageNumber="201" precision="1" value="35.179016">
35°10.741
<emphasis id="B9728D18083B7D31AAA7B7DE6D9EFA1A" bold="true" box="[1419,1425,1453,1479]" italics="true" pageId="16" pageNumber="201">ʹ</emphasis>
N
</geoCoordinate>
<geoCoordinate id="EE3237CD083B7D31AC6BB7BC6BD3FA3A" box="[839,988,1486,1512]" degrees="129" direction="east" minutes="12.317" orientation="longitude" pageId="16" pageNumber="201" precision="1" value="129.20528">
129°12.317
<emphasis id="B9728D18083B7D31ACEFB7BD6BC6FA35" bold="true" box="[963,969,1486,1512]" italics="true" pageId="16" pageNumber="201">ʹ</emphasis>
E
</geoCoordinate>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB9510A083B7D31AC6BB79C6BB1F955" blockId="16.[839,1444,1390,1995]" pageId="16" pageNumber="201">
<emphasis id="B9728D18083B7D31AC6BB79C6C5BF9DA" bold="true" box="[839,1108,1519,1543]" pageId="16" pageNumber="201">Specimens examined.</emphasis>
<materialsCitation id="3B6E5B57083B7D31AB73B79C6BB1F955" collectingDate="2016-05-06" collectionCode="NIBR, T" pageId="16" pageNumber="201" specimenCode="IV 0000287280, IV 0000287281" specimenCount="2" typeStatus="holotype">
<typeStatus id="54BDEFA8083B7D31AB73B79C6CC4F9DA" box="[1119,1227,1519,1543]" pageId="16" pageNumber="201" type="holotype">Holotype</typeStatus>
female (
<collectionCode id="ED17C9CF083B7D31AA19B7836D74F9D5" box="[1333,1403,1520,1544]" country="South Korea" httpUri="http://grbio.org/cool/r4iq-k5jx" name="National Institute of Biological Resources" pageId="16" pageNumber="201">NIBR</collectionCode>
<specimenCode id="DBA0F971083B7D31AAABB7836BDCF9F5" collectionCode="NIBR" country="South Korea" httpUri="http://grbio.org/cool/r4iq-k5jx" name="National Institute of Biological Resources" pageId="16" pageNumber="201">IV 0000287280</specimenCode>
) dissected on one slide;
<typeStatus id="54BDEFA8083B7D31ABDCB4636D40F9F5" box="[1264,1359,1552,1576]" pageId="16" pageNumber="201" type="paratype">paratype</typeStatus>
female (
<collectionCode id="ED17C9CF083B7D31AC62B4436B80F995" box="[846,911,1584,1608]" country="South Korea" httpUri="http://grbio.org/cool/r4iq-k5jx" name="National Institute of Biological Resources" pageId="16" pageNumber="201">NIBR</collectionCode>
<specimenCode id="DBA0F971083B7D31ACB5B4436C4BF995" box="[921,1092,1584,1608]" collectionCode="NIBR" country="South Korea" httpUri="http://grbio.org/cool/r4iq-k5jx" name="National Institute of Biological Resources" pageId="16" pageNumber="201">IV 0000287281</specimenCode>
)
<emphasis id="B9728D18083B7D31AB79B4426C93F995" box="[1109,1180,1585,1608]" italics="true" pageId="16" pageNumber="201">in toto</emphasis>
on one SEM stub; both collected from the type locality,
<date id="FFB877CA083B7D31ABECB4236D42F9B5" box="[1216,1357,1616,1641]" pageId="16" pageNumber="201" value="2016-05-06">
<collectingDate id="EFFC8E22083B7D31ABECB4236D42F9B5" box="[1216,1357,1616,1641]" pageId="16" pageNumber="201" value="2016-05-06">6 May 2016</collectingDate>
</date>
, leg.
<collectionCode id="ED17C9CF083B7D31AAA2B4226DAFF9B5" box="[1422,1440,1617,1640]" name="Tavera, Department of Geology and Geophysics" pageId="16" pageNumber="201">T</collectionCode>
. Karanovic.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31C0281083B7D31AC6BB4E26D51F934" pageId="16" pageNumber="201" type="etymology">
<paragraph id="8BB9510A083B7D31AC6BB4E26D51F934" blockId="16.[839,1444,1390,1995]" pageId="16" pageNumber="201">
<emphasis id="B9728D18083B7D31AC6BB4E26BC7F974" bold="true" box="[839,968,1681,1705]" pageId="16" pageNumber="201">Etymology.</emphasis>
The species is named after the
<typeStatus id="54BDEFA8083B7D31AA36B4E26D47F974" box="[1306,1352,1681,1705]" pageId="16" pageNumber="201">type</typeStatus>
locality, the city of
<collectingRegion id="49C29FE8083B7D31AC90B4C16BF0F914" box="[956,1023,1714,1737]" country="South Korea" name="Busan" pageId="16" pageNumber="201">Busan</collectingRegion>
in
<collectingCountry id="F311119A083B7D31AB0EB4C26CA2F914" box="[1058,1197,1713,1737]" name="South Korea" pageId="16" pageNumber="201">South Korea</collectingCountry>
. The name is an adjective for place, made with the Latin suffix “-ensis”.
</paragraph>
</subSubSection>
<subSubSection id="C31C0281083B7D36AC6BB4826CF4FE02" lastPageId="23" lastPageNumber="208" pageId="16" pageNumber="201" type="description">
<paragraph id="8BB9510A083B7D31AC6BB4826DABF816" blockId="16.[839,1444,1390,1995]" pageId="16" pageNumber="201">
<emphasis id="B9728D18083B7D31AC6BB4826BDBF8D4" bold="true" box="[839,980,1777,1801]" pageId="16" pageNumber="201">Description.</emphasis>
Female (based on
<typeStatus id="54BDEFA8083B7D31AB89B4826D0CF8D4" box="[1189,1283,1777,1801]" pageId="16" pageNumber="201" type="holotype">holotype</typeStatus>
and
<specimenCount id="9D009A83083B7D31AA10B4816B75F8F7" count="1" pageId="16" pageNumber="201" type="generic" typeStatus="paratype">one paratype</specimenCount>
). Total body length, measured from tip of rostrum to posterior margin of caudal rami (excluding caudal setae and appendages) from 405 (
<typeStatus id="54BDEFA8083B7D31AB94B5216D16F8B7" box="[1208,1305,1874,1898]" pageId="16" pageNumber="201" type="holotype">holotype</typeStatus>
) to 450 μm. Colour of preserved specimen light brown (
<figureCitation id="133D4D8F083B7D31AA10B5006D9AF856" box="[1340,1429,1907,1931]" captionStart="Fig" captionStartId="3.[144,177,1234,1256]" captionTargetBox="[177,1380,257,1202]" captionTargetId="figure-358@3.[177,1380,257,1202]" captionTargetPageId="3" captionText="Fig. 1. Light photograph of two new Korean Phyllopodopsyllus Scot T., 1906 species, on the same slide in glycerol:A, P. kitazimai sp. nov., holotype female; B, P. kitazimai sp. nov., allotype male; C, P. busanensis sp. nov., holotype female." pageId="16" pageNumber="201">Fig. 1C</figureCitation>
). Nauplius eye not visible. Segmentation as in previous species. Habitus (
<figureCitation id="133D4D8F083B7D31AB26B5C06C62F816" box="[1034,1133,1971,1995]" captionStart="Fig" captionStartId="3.[144,177,1234,1256]" captionTargetBox="[177,1380,257,1202]" captionTargetId="figure-358@3.[177,1380,257,1202]" captionTargetPageId="3" captionText="Fig. 1. Light photograph of two new Korean Phyllopodopsyllus Scot T., 1906 species, on the same slide in glycerol:A, P. kitazimai sp. nov., holotype female; B, P. kitazimai sp. nov., allotype male; C, P. busanensis sp. nov., holotype female." pageId="16" pageNumber="201">Figs. 1C</figureCitation>
,
<figureCitation id="133D4D8F083B7D31AB57B5C06CA5F816" box="[1147,1194,1971,1995]" captionStart="Fig" captionStartId="17.[144,177,1843,1865]" captionTargetBox="[223,1334,257,1816]" captionTargetId="figure-10@17.[223,1334,257,1816]" captionTargetPageId="17" captionText="Fig. 11. Phyllopodopsyllus busanensis sp. nov., paratype female, SEM photographs: A, habitus, lateral; B, cephalothorax (Ct), lateral; C, anterior part of cephalothorax (Ct) with rostrum(Ro) and first segment of antennula (A1), lateral; D, ventral part of cephalothorax (Ct), with parts of antennula (A1), antenna (A2), and mandibula, lateral; E, dorso-posterior part of cephalothorax (Ct), lateral; F, ventro-posterior corner of cephalothorax (Ct), lateral; G, tergites of three free prosomites (Fp1-Fp3), lateral; H, detail of ornamentation of first (Fp1) and second (Fp2) prosomites, lateral. Arabic numerals for cuticular sensilla and Roman numerals for pores assigned consecutively from anterior to posterior end of each somite and caudal ramus, and from dorsal to ventral side." pageId="16" pageNumber="201">11A</figureCitation>
) cylindrical but much
</paragraph>
<caption id="DF790182083A7D30AFBCB5406B32F812" pageId="17" pageNumber="202" startId="17.[144,177,1843,1865]" targetBox="[223,1334,257,1816]" targetPageId="17" targetType="figure">
<paragraph id="8BB9510A083A7D30AFBCB5406B32F812" blockId="17.[144,1414,1843,1999]" pageId="17" pageNumber="202">
<emphasis id="B9728D18083A7D30AFBCB54068D8F895" bold="true" box="[144,215,1843,1865]" pageId="17" pageNumber="202">Fig. 11.</emphasis>
<taxonomicName id="4C062A89083A7D30AFF3B54069FCF895" authorityName="Lee &amp; Yoo &amp; Kim" authorityYear="2017" box="[223,499,1843,1864]" class="Copepoda" family="Tetragonicipitidae" genus="Phyllopodopsyllus" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="202" phylum="Arthropoda" rank="species" species="busanensis" status="sp. nov.">
<emphasis id="B9728D18083A7D30AFF3B54069FCF895" box="[223,499,1843,1864]" italics="true" pageId="17" pageNumber="202">Phyllopodopsyllus busanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2413063083A7D30AED6B5476A4AF894" box="[506,581,1844,1865]" pageId="17" pageNumber="202" rank="species">sp. nov.</taxonomicNameLabel>
, paratype female, SEM photographs: A, habitus, lateral; B, cephalothorax (Ct), lateral; C, anterior part of cephalothorax (Ct) with rostrum (Ro) and first segment of antennula (A1), lateral; D, ventral part of cephalothorax (Ct), with parts of antennula (A1), antenna (A2), and mandibula, lateral; E, dorso-posterior part of cephalothorax (Ct), lateral; F, ventro-posterior corner of cephalothorax (Ct), lateral; G, tergites of three free prosomites (Fp1-Fp3), lateral; H, detail of ornamentation of first (Fp1) and second (Fp2) prosomites, lateral. Arabic numerals for cuticular sensilla and Roman numerals for pores assigned consecutively from anterior to posterior end of each somite and caudal ramus, and from dorsal to ventral side.
</paragraph>
</caption>
<caption id="DF79018208397D33AF82B5406A7BF813" pageId="18" pageNumber="203" startId="18.[174,207,1843,1865]" targetBox="[253,1364,258,1816]" targetPageId="18" targetType="figure">
<paragraph id="8BB9510A08397D33AF82B5406A7BF813" blockId="18.[174,1445,1843,1998]" pageId="18" pageNumber="203">
<emphasis id="B9728D1808397D33AF82B54068F8F895" bold="true" box="[174,247,1843,1865]" pageId="18" pageNumber="203">Fig. 12.</emphasis>
<taxonomicName id="4C062A8908397D33AFD3B5406A1CF895" authorityName="Lee &amp; Yoo &amp; Kim" authorityYear="2017" box="[255,531,1843,1864]" class="Copepoda" family="Tetragonicipitidae" genus="Phyllopodopsyllus" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="203" phylum="Arthropoda" rank="species" species="busanensis" status="sp. nov.">
<emphasis id="B9728D1808397D33AFD3B5406A1CF895" box="[255,531,1843,1864]" italics="true" pageId="18" pageNumber="203">Phyllopodopsyllus busanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241306308397D33AD37B5476A69F894" box="[539,614,1844,1865]" pageId="18" pageNumber="203" rank="species">sp. nov.</taxonomicNameLabel>
, paratype female, SEM photographs: A, first five urosomites (U1-U5) with fifth leg (P5), lateral; B, tergite of first urosomite (U1), lateral; C, parts of fused second (U2) and third (U3) urosomites, with lateral part of fifth leg (P5), lateral; D, part of fourth urosomite (U4) and distal part of fifth leg (P5); E, fourth (U4) and fifth (U5) urosomites, lateral; F, sixth urosomite and caudal rami (Cr), lateral; G, posterior part of caudal rami (Cr), lateral; H, parts of maxilliped (Mxp) and first swimming leg (P1), lateral. Arabic numerals for cuticular sensilla and Roman numerals for pores assigned consecutively from anterior to posterior end of each somite and caudal ramus, and from dorsal to ventral side.
</paragraph>
</caption>
<paragraph id="8BB9510A08387D32AD71B7176AABFAA5" blockId="19.[605,676,1380,1400]" box="[605,676,1380,1400]" pageId="19" pageNumber="204">
<emphasis id="B9728D1808387D32AD71B7176AABFAA5" bold="true" box="[605,676,1380,1400]" pageId="19" pageNumber="204">100 µm</emphasis>
</paragraph>
<caption id="DF79018208387D32AFBCB7E36C4DF9CC" pageId="19" pageNumber="204" startId="19.[144,177,1424,1446]" targetBox="[178,1372,263,1388]" targetPageId="19" targetType="figure">
<paragraph id="8BB9510A08387D32AFBCB7E36C4DF9CC" blockId="19.[144,1414,1424,1553]" pageId="19" pageNumber="204">
<emphasis id="B9728D1808387D32AFBCB7E368D7FA78" bold="true" box="[144,216,1424,1446]" pageId="19" pageNumber="204">Fig. 13.</emphasis>
<taxonomicName id="4C062A8908387D32AFCCB7E369FBFA78" authorityName="Lee &amp; Yoo &amp; Kim" authorityYear="2017" box="[224,500,1424,1445]" class="Copepoda" family="Tetragonicipitidae" genus="Phyllopodopsyllus" kingdom="Animalia" order="Harpacticoida" pageId="19" pageNumber="204" phylum="Arthropoda" rank="species" species="busanensis" status="sp. nov.">
<emphasis id="B9728D1808387D32AFCCB7E369FBFA78" box="[224,500,1424,1445]" italics="true" pageId="19" pageNumber="204">Phyllopodopsyllus busanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241306308387D32AED7B7E26A49FA7B" box="[507,582,1425,1446]" pageId="19" pageNumber="204" rank="species">sp. nov.</taxonomicNameLabel>
, holotype female, line drawings: A, second to sixth urosomites (U2-U6) and caudal rami, dorsal; B, second to sixth urosomites (U2-U6), with caudal rami (Cr), and attached spermatophore (Sp), lateral; C, second to sixth urosomites (U2-U6), with caudal rami (Cr), sixth legs (P6), attached spermatophore (Sp), and internal reseptacula seminis (Rs), ventral. Arabic numerals for cuticular sensilla and Roman numerals for pores assigned consecutively from anterior to posterior end of each somite and caudal ramus, and from dorsal to ventral side. Cuticular pits and minute surface spinules omitted for clarity.
</paragraph>
</caption>
<paragraph id="8BB9510A08387D32AFBCB4246D89F86C" blockId="19.[144,749,1623,1970]" lastBlockId="19.[809,1414,1623,1970]" pageId="19" pageNumber="204">
less slender than in previous species, without distinct demarcation between prosome and urosome in dorsal view, but with sharp bend in lateral view; prosome/urosome ratio about 0.9; greatest width at posterior end of cephalothorax; cephalothorax only 1.2 times as wide as genital double-somite in dorsal view. Body length/ width ratio about 4.2. Free pedigerous somites without pronounced lateral or dorsal expansions. Integument of all somites relatively well sclerotized, generally smooth, covered with numerous shallow cuticular pits (see
<figureCitation id="133D4D8F08387D32AD94B50A68B0F86F" captionStart="Fig" captionStartId="17.[144,177,1843,1865]" captionTargetBox="[223,1334,257,1816]" captionTargetId="figure-10@17.[223,1334,257,1816]" captionTargetPageId="17" captionText="Fig. 11. Phyllopodopsyllus busanensis sp. nov., paratype female, SEM photographs: A, habitus, lateral; B, cephalothorax (Ct), lateral; C, anterior part of cephalothorax (Ct) with rostrum(Ro) and first segment of antennula (A1), lateral; D, ventral part of cephalothorax (Ct), with parts of antennula (A1), antenna (A2), and mandibula, lateral; E, dorso-posterior part of cephalothorax (Ct), lateral; F, ventro-posterior corner of cephalothorax (Ct), lateral; G, tergites of three free prosomites (Fp1-Fp3), lateral; H, detail of ornamentation of first (Fp1) and second (Fp2) prosomites, lateral. Arabic numerals for cuticular sensilla and Roman numerals for pores assigned consecutively from anterior to posterior end of each somite and caudal ramus, and from dorsal to ventral side." pageId="19" pageNumber="204">Figs. 11C</figureCitation>
,
<figureCitation id="133D4D8F08387D32AFE2B5E9690DF86F" box="[206,258,1946,1970]" captionStart="Fig" captionStartId="18.[174,207,1843,1865]" captionTargetBox="[253,1364,258,1816]" captionTargetId="figure-11@18.[253,1364,258,1816]" captionTargetPageId="18" captionText="Fig. 12. Phyllopodopsyllus busanensis sp. nov., paratype female, SEM photographs: A, first five urosomites (U1-U5) with fifth leg (P5), lateral; B, tergite of first urosomite (U1), lateral; C, parts of fused second (U2) and third (U3) urosomites, with lateral part of fifth leg (P5), lateral; D, part of fourth urosomite (U4) and distal part of fifth leg (P5); E, fourth (U4) and fifth (U5) urosomites, lateral; F, sixth urosomite and caudal rami(Cr), lateral; G, posterior part of caudal rami(Cr), lateral; H, parts of maxilliped (Mxp) and first swimming leg (P1), lateral. Arabic numerals for cuticular sensilla and Roman numerals for pores assigned consecutively from anterior to posterior end of each somite and caudal ramus, and from dorsal to ventral side." pageId="19" pageNumber="204">12G</figureCitation>
); free prosomites and all urosomites additionally covered by parallel rows of minute and slender spinules; cephalothorax devoid of spinules. Hyaline fringe of all somites narrow and coarsely serrated. Surface of somites and caudal rami with total maximum of 185 cuticular organs (29 pairs of cuticular pores, 58 pairs of sensilla, one unpaired dorsal sensillum, and ten unpaired dorsal pores). All sensilla and pores homologous to those in previous species, with only small differences in their relative position; only real difference in cuticular organs visible on fifth urosomite (pore U5-II absent; see
<figureCitation id="133D4D8F08387D32AC05B5E96B85F86F" box="[809,906,1946,1970]" captionStart="Fig" captionStartId="19.[144,177,1424,1446]" captionTargetBox="[178,1372,263,1388]" captionTargetId="figure-199@19.[189,1429,262,1396]" captionTargetPageId="19" captionText="Fig. 13. Phyllopodopsyllus busanensis sp. nov., holotype female, line drawings: A, second to sixth urosomites (U2-U6) and caudal rami, dorsal; B, second to sixth urosomites (U2-U6), with caudal rami (Cr), and attached spermatophore (Sp), lateral; C, second to sixth urosomites (U2-U6), with caudal rami (Cr), sixth legs (P6), attached spermatophore (Sp), and internal reseptacula seminis (Rs), ventral. Arabic numerals for cuticular sensilla and Roman numerals for pores assigned consecutively from anterior to posterior end of each somite and caudal ramus, and from dorsal to ventral side. Cuticular pits and minute surface spinules omitted for clarity." pageId="19" pageNumber="204">Fig. 13C</figureCitation>
) and caudal rami (additional pore Cr-III pres-
</paragraph>
<paragraph id="8BB9510A083F7D35AD7DB3226CCEF8CC" pageId="20" pageNumber="205">A B C D G H E F</paragraph>
<paragraph id="8BB9510A083F7D35ACCFB5486C25F892" blockId="20.[995,1066,1851,1871]" box="[995,1066,1851,1871]" pageId="20" pageNumber="205">
<emphasis id="B9728D18083F7D35ACCFB5486C25F892" bold="true" box="[995,1066,1851,1871]" pageId="20" pageNumber="205">100 µm</emphasis>
</paragraph>
<paragraph id="8BB9510A083F7D35AF82B2DA6DACFF62" blockId="20.[174,1443,169,191]" box="[174,1443,169,191]" pageId="20" pageNumber="205">
<materialsCitation id="3B6E5B57083F7D35AF82B2DA6C9EFF63" box="[174,1169,169,190]" collectingDate="2017-12" country="South Korea" location="KOREA" pageId="20" pageNumber="205" specimenCount="1">
<emphasis id="B9728D18083F7D35AF82B2DA6931FF63" box="[174,318,169,190]" italics="true" pageId="20" pageNumber="205">
<date id="FFB877CA083F7D35AF82B2DA6931FF63" box="[174,318,169,190]" pageId="20" pageNumber="205" value="2017-12">
<collectingDate id="EFFC8E22083F7D35AF82B2DA6931FF63" box="[174,318,169,190]" pageId="20" pageNumber="205" value="2017-12">December 2017</collectingDate>
</date>
</emphasis>
KARANOVIC-NEW SPECIES OF
<taxonomicName id="4C062A89083F7D35AC26B2DA6C31FF63" authority="FROM" authorityName="FROM" box="[778,1086,169,190]" class="Copepoda" family="Tetragonicipitidae" genus="Phyllopodopsyllus" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="205" phylum="Arthropoda" rank="genus">
<emphasis id="B9728D18083F7D35AC26B2DA6BF7FF63" box="[778,1016,169,190]" italics="true" pageId="20" pageNumber="205">PHYLLOPODOPSYLLUS</emphasis>
FROM
</taxonomicName>
<collectingCountry id="F311119A083F7D35AB68B2DA6C9EFF63" box="[1092,1169,169,190]" name="South Korea" pageId="20" pageNumber="205">KOREA</collectingCountry>
</materialsCitation>
205
</paragraph>
<caption id="DF790182083F7D35AF82B5056B8DF81C" pageId="20" pageNumber="205" startId="20.[174,207,1910,1932]" targetType="figure">
<paragraph id="8BB9510A083F7D35AF82B5056B8DF81C" blockId="20.[174,1445,1910,1985]" pageId="20" pageNumber="205">
<emphasis id="B9728D18083F7D35AF82B50568F9F856" bold="true" box="[174,246,1910,1932]" pageId="20" pageNumber="205">Fig. 14.</emphasis>
<taxonomicName id="4C062A89083F7D35AFD1B5056A01F856" authorityName="Lee &amp; Yoo &amp; Kim" authorityYear="2017" box="[253,526,1910,1931]" class="Copepoda" family="Tetragonicipitidae" genus="Phyllopodopsyllus" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="205" phylum="Arthropoda" rank="species" species="busanensis" status="sp. nov.">
<emphasis id="B9728D18083F7D35AFD1B5056A01F856" box="[253,526,1910,1931]" italics="true" pageId="20" pageNumber="205">Phyllopodopsyllus busanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2413063083F7D35AD3AB5046A6FF851" box="[534,608,1911,1932]" pageId="20" pageNumber="205" rank="species">sp. nov.</taxonomicNameLabel>
, holotype female, line drawings: A, antennula, dorsal. B, antenna, posterior; C, mandibula, anterior; D, praecoxal arthrite of maxillula, anterior; E, coxa of maxillula, anterior; F, endopod of maxillula, anterior; G, maxilla, posterior; H, maxilliped, anterior. Cuticular pits on antennula and antenna omitted for clarity.
</paragraph>
</caption>
<caption id="DF790182083E7D34AFBCB4CC6D32F932" pageId="21" pageNumber="206" startId="21.[144,177,1727,1749]" targetBox="[178,1348,262,1697]" targetPageId="21" targetType="figure">
<paragraph id="8BB9510A083E7D34AFBCB4CC6D32F932" blockId="21.[144,1414,1727,1776]" pageId="21" pageNumber="206">
<emphasis id="B9728D18083E7D34AFBCB4CC68D8F909" bold="true" box="[144,215,1727,1749]" pageId="21" pageNumber="206">Fig. 15.</emphasis>
<taxonomicName id="4C062A89083E7D34AFF3B4CC69FFF909" authorityName="Lee &amp; Yoo &amp; Kim" authorityYear="2017" box="[223,496,1727,1748]" class="Copepoda" family="Tetragonicipitidae" genus="Phyllopodopsyllus" kingdom="Animalia" order="Harpacticoida" pageId="21" pageNumber="206" phylum="Arthropoda" rank="species" species="busanensis" status="sp. nov.">
<emphasis id="B9728D18083E7D34AFF3B4CC69FFF909" box="[223,496,1727,1748]" italics="true" pageId="21" pageNumber="206">Phyllopodopsyllus busanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2413063083E7D34AEDBB4CC6A4EF909" box="[503,577,1727,1748]" pageId="21" pageNumber="206" rank="species">sp. nov.</taxonomicNameLabel>
, holotype female, line drawings: A, first swimming leg, anterior; B, second swimming leg, anterior; C, third swimming leg, anterior; D, fourth swimming leg, anterior; E, fifth leg, compressed and slightly deformed, anterior.
</paragraph>
</caption>
<paragraph id="8BB9510A083E7D34AFBCB5456A0CF852" blockId="21.[144,749,1846,1999]" pageId="21" pageNumber="206">
ent; see
<figureCitation id="133D4D8F083E7D34AFC6B5456958F893" box="[234,343,1846,1870]" captionStart="Fig" captionStartId="18.[174,207,1843,1865]" captionTargetBox="[253,1364,258,1816]" captionTargetId="figure-11@18.[253,1364,258,1816]" captionTargetPageId="18" captionText="Fig. 12. Phyllopodopsyllus busanensis sp. nov., paratype female, SEM photographs: A, first five urosomites (U1-U5) with fifth leg (P5), lateral; B, tergite of first urosomite (U1), lateral; C, parts of fused second (U2) and third (U3) urosomites, with lateral part of fifth leg (P5), lateral; D, part of fourth urosomite (U4) and distal part of fifth leg (P5); E, fourth (U4) and fifth (U5) urosomites, lateral; F, sixth urosomite and caudal rami(Cr), lateral; G, posterior part of caudal rami(Cr), lateral; H, parts of maxilliped (Mxp) and first swimming leg (P1), lateral. Arabic numerals for cuticular sensilla and Roman numerals for pores assigned consecutively from anterior to posterior end of each somite and caudal ramus, and from dorsal to ventral side." pageId="21" pageNumber="206">Figs. 12G</figureCitation>
,
<figureCitation id="133D4D8F083E7D34AE48B545699BF893" box="[356,404,1846,1870]" captionStart="Fig" captionStartId="19.[144,177,1424,1446]" captionTargetBox="[178,1372,263,1388]" captionTargetId="figure-199@19.[189,1429,262,1396]" captionTargetPageId="19" captionText="Fig. 13. Phyllopodopsyllus busanensis sp. nov., holotype female, line drawings: A, second to sixth urosomites (U2-U6) and caudal rami, dorsal; B, second to sixth urosomites (U2-U6), with caudal rami (Cr), and attached spermatophore (Sp), lateral; C, second to sixth urosomites (U2-U6), with caudal rami (Cr), sixth legs (P6), attached spermatophore (Sp), and internal reseptacula seminis (Rs), ventral. Arabic numerals for cuticular sensilla and Roman numerals for pores assigned consecutively from anterior to posterior end of each somite and caudal ramus, and from dorsal to ventral side. Cuticular pits and minute surface spinules omitted for clarity." pageId="21" pageNumber="206">13B</figureCitation>
); pore Ct-XI larger than in previous species (see
<figureCitation id="133D4D8F083E7D34AE7AB52569BDF8B3" box="[342,434,1878,1902]" captionStart="Fig" captionStartId="17.[144,177,1843,1865]" captionTargetBox="[223,1334,257,1816]" captionTargetId="figure-10@17.[223,1334,257,1816]" captionTargetPageId="17" captionText="Fig. 11. Phyllopodopsyllus busanensis sp. nov., paratype female, SEM photographs: A, habitus, lateral; B, cephalothorax (Ct), lateral; C, anterior part of cephalothorax (Ct) with rostrum(Ro) and first segment of antennula (A1), lateral; D, ventral part of cephalothorax (Ct), with parts of antennula (A1), antenna (A2), and mandibula, lateral; E, dorso-posterior part of cephalothorax (Ct), lateral; F, ventro-posterior corner of cephalothorax (Ct), lateral; G, tergites of three free prosomites (Fp1-Fp3), lateral; H, detail of ornamentation of first (Fp1) and second (Fp2) prosomites, lateral. Arabic numerals for cuticular sensilla and Roman numerals for pores assigned consecutively from anterior to posterior end of each somite and caudal ramus, and from dorsal to ventral side." pageId="21" pageNumber="206">Fig. 11F</figureCitation>
) and sensilla U1-3 and U1-4 closer to each other (see
<figureCitation id="133D4D8F083E7D34AEBAB50569F9F852" box="[406,502,1910,1935]" captionStart="Fig" captionStartId="18.[174,207,1843,1865]" captionTargetBox="[253,1364,258,1816]" captionTargetId="figure-11@18.[253,1364,258,1816]" captionTargetPageId="18" captionText="Fig. 12. Phyllopodopsyllus busanensis sp. nov., paratype female, SEM photographs: A, first five urosomites (U1-U5) with fifth leg (P5), lateral; B, tergite of first urosomite (U1), lateral; C, parts of fused second (U2) and third (U3) urosomites, with lateral part of fifth leg (P5), lateral; D, part of fourth urosomite (U4) and distal part of fifth leg (P5); E, fourth (U4) and fifth (U5) urosomites, lateral; F, sixth urosomite and caudal rami(Cr), lateral; G, posterior part of caudal rami(Cr), lateral; H, parts of maxilliped (Mxp) and first swimming leg (P1), lateral. Arabic numerals for cuticular sensilla and Roman numerals for pores assigned consecutively from anterior to posterior end of each somite and caudal ramus, and from dorsal to ventral side." pageId="21" pageNumber="206">Fig. 12B</figureCitation>
).
</paragraph>
<paragraph id="8BB9510A083E7D34AF8AB5E46C3BF872" blockId="21.[144,749,1846,1999]" lastBlockId="21.[809,1414,1846,1999]" pageId="21" pageNumber="206">
Rostrum (
<figureCitation id="133D4D8F083E7D34AE35B5E4698DF872" box="[281,386,1943,1967]" captionStart="Fig" captionStartId="17.[144,177,1843,1865]" captionTargetBox="[223,1334,257,1816]" captionTargetId="figure-10@17.[223,1334,257,1816]" captionTargetPageId="17" captionText="Fig. 11. Phyllopodopsyllus busanensis sp. nov., paratype female, SEM photographs: A, habitus, lateral; B, cephalothorax (Ct), lateral; C, anterior part of cephalothorax (Ct) with rostrum(Ro) and first segment of antennula (A1), lateral; D, ventral part of cephalothorax (Ct), with parts of antennula (A1), antenna (A2), and mandibula, lateral; E, dorso-posterior part of cephalothorax (Ct), lateral; F, ventro-posterior corner of cephalothorax (Ct), lateral; G, tergites of three free prosomites (Fp1-Fp3), lateral; H, detail of ornamentation of first (Fp1) and second (Fp2) prosomites, lateral. Arabic numerals for cuticular sensilla and Roman numerals for pores assigned consecutively from anterior to posterior end of each somite and caudal ramus, and from dorsal to ventral side." pageId="21" pageNumber="206">Fig. 11C</figureCitation>
) as in previous species small, weakly demarcated at base from cephalothorax, linguiform, about as wide as long, its anterior tip hardly reaching beyond anterior margin of lateral wings of cephalothoracic shield, with single dorsal pair of sensilla (Ct-1) at about 2/3 of its length.
</paragraph>
<paragraph id="8BB9510A083E7D37AC13B5C469EDFE43" blockId="21.[809,1414,1846,1999]" lastBlockId="22.[174,779,261,1995]" lastPageId="22" lastPageNumber="207" pageId="21" pageNumber="206">
Cephalothorax (
<figureCitation id="133D4D8F083E7D34ACD9B5C46C49F812" box="[1013,1094,1975,1999]" captionStart="Fig" captionStartId="17.[144,177,1843,1865]" captionTargetBox="[223,1334,257,1816]" captionTargetId="figure-10@17.[223,1334,257,1816]" captionTargetPageId="17" captionText="Fig. 11. Phyllopodopsyllus busanensis sp. nov., paratype female, SEM photographs: A, habitus, lateral; B, cephalothorax (Ct), lateral; C, anterior part of cephalothorax (Ct) with rostrum(Ro) and first segment of antennula (A1), lateral; D, ventral part of cephalothorax (Ct), with parts of antennula (A1), antenna (A2), and mandibula, lateral; E, dorso-posterior part of cephalothorax (Ct), lateral; F, ventro-posterior corner of cephalothorax (Ct), lateral; G, tergites of three free prosomites (Fp1-Fp3), lateral; H, detail of ornamentation of first (Fp1) and second (Fp2) prosomites, lateral. Arabic numerals for cuticular sensilla and Roman numerals for pores assigned consecutively from anterior to posterior end of each somite and caudal ramus, and from dorsal to ventral side." pageId="21" pageNumber="206">Fig. 11</figureCitation>
B-E) smooth, cylindrical in dorsal view, tapering towards anterior end in lateral view, about 1.5 times as long as wide in dorsal view (including rostrum); representing 33% of total body length. Hyaline fringe of cephalothoracic shield narrow, serrat- ed dorsally, smooth laterally.
</paragraph>
<paragraph id="8BB9510A083D7D37AFE9B3D56AE9FE22" blockId="22.[174,779,261,1995]" pageId="22" pageNumber="207">
Pleurons of first free prosomites (
<figureCitation id="133D4D8F083D7D37AD01B3D56ABFFE62" box="[557,688,422,447]" captionStart="Fig" captionStartId="17.[144,177,1843,1865]" captionTargetBox="[223,1334,257,1816]" captionTargetId="figure-10@17.[223,1334,257,1816]" captionTargetPageId="17" captionText="Fig. 11. Phyllopodopsyllus busanensis sp. nov., paratype female, SEM photographs: A, habitus, lateral; B, cephalothorax (Ct), lateral; C, anterior part of cephalothorax (Ct) with rostrum(Ro) and first segment of antennula (A1), lateral; D, ventral part of cephalothorax (Ct), with parts of antennula (A1), antenna (A2), and mandibula, lateral; E, dorso-posterior part of cephalothorax (Ct), lateral; F, ventro-posterior corner of cephalothorax (Ct), lateral; G, tergites of three free prosomites (Fp1-Fp3), lateral; H, detail of ornamentation of first (Fp1) and second (Fp2) prosomites, lateral. Arabic numerals for cuticular sensilla and Roman numerals for pores assigned consecutively from anterior to posterior end of each somite and caudal ramus, and from dorsal to ventral side." pageId="22" pageNumber="207">Fig. 11G, H</figureCitation>
) shorter than in previous species and densely covered by slender spinules, with only small lateral surface free of them.
</paragraph>
<paragraph id="8BB9510A083D7D37AFE9B07468EFFDBD" blockId="22.[174,779,261,1995]" pageId="22" pageNumber="207">
All urosomites (
<figureCitation id="133D4D8F083D7D37AE58B07469CDFDC2" box="[372,450,519,543]" captionStart="Fig" captionStartId="18.[174,207,1843,1865]" captionTargetBox="[253,1364,258,1816]" captionTargetId="figure-11@18.[253,1364,258,1816]" captionTargetPageId="18" captionText="Fig. 12. Phyllopodopsyllus busanensis sp. nov., paratype female, SEM photographs: A, first five urosomites (U1-U5) with fifth leg (P5), lateral; B, tergite of first urosomite (U1), lateral; C, parts of fused second (U2) and third (U3) urosomites, with lateral part of fifth leg (P5), lateral; D, part of fourth urosomite (U4) and distal part of fifth leg (P5); E, fourth (U4) and fifth (U5) urosomites, lateral; F, sixth urosomite and caudal rami(Cr), lateral; G, posterior part of caudal rami(Cr), lateral; H, parts of maxilliped (Mxp) and first swimming leg (P1), lateral. Arabic numerals for cuticular sensilla and Roman numerals for pores assigned consecutively from anterior to posterior end of each somite and caudal ramus, and from dorsal to ventral side." pageId="22" pageNumber="207">Fig. 12</figureCitation>
A-F) also densely covered by slender spinules, and more so on dorsal than on ventral side.
</paragraph>
<paragraph id="8BB9510A083D7D37AFE9B01B69FDFC7F" blockId="22.[174,779,261,1995]" pageId="22" pageNumber="207">
Genital double-somite (
<figureCitation id="133D4D8F083D7D37AEFEB01B6A65FD5D" box="[466,618,616,640]" captionStart="Fig" captionStartId="18.[174,207,1843,1865]" captionTargetBox="[253,1364,258,1816]" captionTargetId="figure-11@18.[253,1364,258,1816]" captionTargetPageId="18" captionText="Fig. 12. Phyllopodopsyllus busanensis sp. nov., paratype female, SEM photographs: A, first five urosomites (U1-U5) with fifth leg (P5), lateral; B, tergite of first urosomite (U1), lateral; C, parts of fused second (U2) and third (U3) urosomites, with lateral part of fifth leg (P5), lateral; D, part of fourth urosomite (U4) and distal part of fifth leg (P5); E, fourth (U4) and fifth (U5) urosomites, lateral; F, sixth urosomite and caudal rami(Cr), lateral; G, posterior part of caudal rami(Cr), lateral; H, parts of maxilliped (Mxp) and first swimming leg (P1), lateral. Arabic numerals for cuticular sensilla and Roman numerals for pores assigned consecutively from anterior to posterior end of each somite and caudal ramus, and from dorsal to ventral side." pageId="22" pageNumber="207">Figs. 12A, C</figureCitation>
,
<figureCitation id="133D4D8F083D7D37AD56B01B6A9CFD5D" box="[634,659,616,640]" captionStart="Fig" captionStartId="19.[144,177,1424,1446]" captionTargetBox="[178,1372,263,1388]" captionTargetId="figure-199@19.[189,1429,262,1396]" captionTargetPageId="19" captionText="Fig. 13. Phyllopodopsyllus busanensis sp. nov., holotype female, line drawings: A, second to sixth urosomites (U2-U6) and caudal rami, dorsal; B, second to sixth urosomites (U2-U6), with caudal rami (Cr), and attached spermatophore (Sp), lateral; C, second to sixth urosomites (U2-U6), with caudal rami (Cr), sixth legs (P6), attached spermatophore (Sp), and internal reseptacula seminis (Rs), ventral. Arabic numerals for cuticular sensilla and Roman numerals for pores assigned consecutively from anterior to posterior end of each somite and caudal ramus, and from dorsal to ventral side. Cuticular pits and minute surface spinules omitted for clarity." pageId="22" pageNumber="207">13</figureCitation>
A-C) only slightly longer than wide in ventral view, with finely serrated dorsal fringe and lateral internal ridges as only evidence of ancestral segmentation. Genital complex (
<figureCitation id="133D4D8F083D7D37AF9AB09A6918FCDC" box="[182,279,745,769]" captionStart="Fig" captionStartId="19.[144,177,1424,1446]" captionTargetBox="[178,1372,263,1388]" captionTargetId="figure-199@19.[189,1429,262,1396]" captionTargetPageId="19" captionText="Fig. 13. Phyllopodopsyllus busanensis sp. nov., holotype female, line drawings: A, second to sixth urosomites (U2-U6) and caudal rami, dorsal; B, second to sixth urosomites (U2-U6), with caudal rami (Cr), and attached spermatophore (Sp), lateral; C, second to sixth urosomites (U2-U6), with caudal rami (Cr), sixth legs (P6), attached spermatophore (Sp), and internal reseptacula seminis (Rs), ventral. Arabic numerals for cuticular sensilla and Roman numerals for pores assigned consecutively from anterior to posterior end of each somite and caudal ramus, and from dorsal to ventral side. Cuticular pits and minute surface spinules omitted for clarity." pageId="22" pageNumber="207">Fig. 13C</figureCitation>
) with single large and round copulatory pore in distal part of second urosomite, short and narrow copulatory duct, two small and ovoid seminal receptacles in central part of second urosomite, and central genital aperture in anterior part of second urosomite; aperture covered by reduced sixth legs.
</paragraph>
<paragraph id="8BB9510A083D7D37AFE9B1D869A1FC3E" blockId="22.[174,779,261,1995]" pageId="22" pageNumber="207">
Fourth urosomite (
<figureCitation id="133D4D8F083D7D37AEA1B1D86A13FC1F" box="[397,540,939,963]" captionStart="Fig" captionStartId="18.[174,207,1843,1865]" captionTargetBox="[253,1364,258,1816]" captionTargetId="figure-11@18.[253,1364,258,1816]" captionTargetPageId="18" captionText="Fig. 12. Phyllopodopsyllus busanensis sp. nov., paratype female, SEM photographs: A, first five urosomites (U1-U5) with fifth leg (P5), lateral; B, tergite of first urosomite (U1), lateral; C, parts of fused second (U2) and third (U3) urosomites, with lateral part of fifth leg (P5), lateral; D, part of fourth urosomite (U4) and distal part of fifth leg (P5); E, fourth (U4) and fifth (U5) urosomites, lateral; F, sixth urosomite and caudal rami(Cr), lateral; G, posterior part of caudal rami(Cr), lateral; H, parts of maxilliped (Mxp) and first swimming leg (P1), lateral. Arabic numerals for cuticular sensilla and Roman numerals for pores assigned consecutively from anterior to posterior end of each somite and caudal ramus, and from dorsal to ventral side." pageId="22" pageNumber="207">Figs. 12D, E</figureCitation>
,
<figureCitation id="133D4D8F083D7D37AD05B1D86A4EFC1E" box="[553,577,939,963]" captionStart="Fig" captionStartId="19.[144,177,1424,1446]" captionTargetBox="[178,1372,263,1388]" captionTargetId="figure-199@19.[189,1429,262,1396]" captionTargetPageId="19" captionText="Fig. 13. Phyllopodopsyllus busanensis sp. nov., holotype female, line drawings: A, second to sixth urosomites (U2-U6) and caudal rami, dorsal; B, second to sixth urosomites (U2-U6), with caudal rami (Cr), and attached spermatophore (Sp), lateral; C, second to sixth urosomites (U2-U6), with caudal rami (Cr), sixth legs (P6), attached spermatophore (Sp), and internal reseptacula seminis (Rs), ventral. Arabic numerals for cuticular sensilla and Roman numerals for pores assigned consecutively from anterior to posterior end of each somite and caudal ramus, and from dorsal to ventral side. Cuticular pits and minute surface spinules omitted for clarity." pageId="22" pageNumber="207">13</figureCitation>
A-C) with hyaline fringe smooth ventrally.
</paragraph>
<paragraph id="8BB9510A083D7D37AFE9B1986A56FB18" blockId="22.[174,779,261,1995]" pageId="22" pageNumber="207">
Sixth urosomite (
<figureCitation id="133D4D8F083D7D37AE53B19869E7FBDE" box="[383,488,1003,1027]" captionStart="Fig" captionStartId="18.[174,207,1843,1865]" captionTargetBox="[253,1364,258,1816]" captionTargetId="figure-11@18.[253,1364,258,1816]" captionTargetPageId="18" captionText="Fig. 12. Phyllopodopsyllus busanensis sp. nov., paratype female, SEM photographs: A, first five urosomites (U1-U5) with fifth leg (P5), lateral; B, tergite of first urosomite (U1), lateral; C, parts of fused second (U2) and third (U3) urosomites, with lateral part of fifth leg (P5), lateral; D, part of fourth urosomite (U4) and distal part of fifth leg (P5); E, fourth (U4) and fifth (U5) urosomites, lateral; F, sixth urosomite and caudal rami(Cr), lateral; G, posterior part of caudal rami(Cr), lateral; H, parts of maxilliped (Mxp) and first swimming leg (P1), lateral. Arabic numerals for cuticular sensilla and Roman numerals for pores assigned consecutively from anterior to posterior end of each somite and caudal ramus, and from dorsal to ventral side." pageId="22" pageNumber="207">Figs. 12F</figureCitation>
,
<figureCitation id="133D4D8F083D7D37AED9B1986A02FBDE" box="[501,525,1003,1027]" captionStart="Fig" captionStartId="19.[144,177,1424,1446]" captionTargetBox="[178,1372,263,1388]" captionTargetId="figure-199@19.[189,1429,262,1396]" captionTargetPageId="19" captionText="Fig. 13. Phyllopodopsyllus busanensis sp. nov., holotype female, line drawings: A, second to sixth urosomites (U2-U6) and caudal rami, dorsal; B, second to sixth urosomites (U2-U6), with caudal rami (Cr), and attached spermatophore (Sp), lateral; C, second to sixth urosomites (U2-U6), with caudal rami (Cr), sixth legs (P6), attached spermatophore (Sp), and internal reseptacula seminis (Rs), ventral. Arabic numerals for cuticular sensilla and Roman numerals for pores assigned consecutively from anterior to posterior end of each somite and caudal ramus, and from dorsal to ventral side. Cuticular pits and minute surface spinules omitted for clarity." pageId="22" pageNumber="207">13</figureCitation>
A-C) slightly narrower and significantly longer than fifth urosomite, cleft medially in posterior part, with posterior row of spinules; anal operculum broad but less so than in previous species, also longer and more convex, with finely serrated posterior margin, representing about 46% of somites width; anal sinus as in previous species.
</paragraph>
<paragraph id="8BB9510A083D7D37AFE9B6BE6ADFF955" blockId="22.[174,779,261,1995]" pageId="22" pageNumber="207">
Caudal rami (
<figureCitation id="133D4D8F083D7D37AE48B6BE69F5FB38" box="[356,506,1229,1253]" captionStart="Fig" captionStartId="18.[174,207,1843,1865]" captionTargetBox="[253,1364,258,1816]" captionTargetId="figure-11@18.[253,1364,258,1816]" captionTargetPageId="18" captionText="Fig. 12. Phyllopodopsyllus busanensis sp. nov., paratype female, SEM photographs: A, first five urosomites (U1-U5) with fifth leg (P5), lateral; B, tergite of first urosomite (U1), lateral; C, parts of fused second (U2) and third (U3) urosomites, with lateral part of fifth leg (P5), lateral; D, part of fourth urosomite (U4) and distal part of fifth leg (P5); E, fourth (U4) and fifth (U5) urosomites, lateral; F, sixth urosomite and caudal rami(Cr), lateral; G, posterior part of caudal rami(Cr), lateral; H, parts of maxilliped (Mxp) and first swimming leg (P1), lateral. Arabic numerals for cuticular sensilla and Roman numerals for pores assigned consecutively from anterior to posterior end of each somite and caudal ramus, and from dorsal to ventral side." pageId="22" pageNumber="207">Figs. 12F, G</figureCitation>
,
<figureCitation id="133D4D8F083D7D37AD26B6BE6A2BFB38" box="[522,548,1229,1253]" captionStart="Fig" captionStartId="19.[144,177,1424,1446]" captionTargetBox="[178,1372,263,1388]" captionTargetId="figure-199@19.[189,1429,262,1396]" captionTargetPageId="19" captionText="Fig. 13. Phyllopodopsyllus busanensis sp. nov., holotype female, line drawings: A, second to sixth urosomites (U2-U6) and caudal rami, dorsal; B, second to sixth urosomites (U2-U6), with caudal rami (Cr), and attached spermatophore (Sp), lateral; C, second to sixth urosomites (U2-U6), with caudal rami (Cr), sixth legs (P6), attached spermatophore (Sp), and internal reseptacula seminis (Rs), ventral. Arabic numerals for cuticular sensilla and Roman numerals for pores assigned consecutively from anterior to posterior end of each somite and caudal ramus, and from dorsal to ventral side. Cuticular pits and minute surface spinules omitted for clarity." pageId="22" pageNumber="207">13</figureCitation>
A-C) strongly sclerotized, cylindrical but with strong dorsal ridge, with slightly inflated posterior ventro-outer corner, with slender spinules along outer-dorsal surface but without spinules along inner margin, about 1.8 times as long as greatest width in ventral view and nearly 1.7 times in lateral view, with narrow space between them; ornamented three lateral pores, two of them homologous to those in previous species (Cr-I and Cr-II); armed as in previous species with seven setae but no setae fused basally, anterior lateral setae inserted slightly more anteriorly, longer inner apical seta, and central apical seta with bulbous base; novel pore (Cr-III) situated between two other pores; doral seta marking end of dorsal ridge.
</paragraph>
<paragraph id="8BB9510A083D7D37AFE9B4E26D04FE63" blockId="22.[174,779,261,1995]" lastBlockId="22.[839,1445,261,1995]" pageId="22" pageNumber="207">
Antennula (
<figureCitation id="133D4D8F083D7D37AE6DB4E269C2F975" box="[321,461,1681,1705]" captionStart="Fig" captionStartId="17.[144,177,1843,1865]" captionTargetBox="[223,1334,257,1816]" captionTargetId="figure-10@17.[223,1334,257,1816]" captionTargetPageId="17" captionText="Fig. 11. Phyllopodopsyllus busanensis sp. nov., paratype female, SEM photographs: A, habitus, lateral; B, cephalothorax (Ct), lateral; C, anterior part of cephalothorax (Ct) with rostrum(Ro) and first segment of antennula (A1), lateral; D, ventral part of cephalothorax (Ct), with parts of antennula (A1), antenna (A2), and mandibula, lateral; E, dorso-posterior part of cephalothorax (Ct), lateral; F, ventro-posterior corner of cephalothorax (Ct), lateral; G, tergites of three free prosomites (Fp1-Fp3), lateral; H, detail of ornamentation of first (Fp1) and second (Fp2) prosomites, lateral. Arabic numerals for cuticular sensilla and Roman numerals for pores assigned consecutively from anterior to posterior end of each somite and caudal ramus, and from dorsal to ventral side." pageId="22" pageNumber="207">Figs. 11C, D</figureCitation>
,
<figureCitation id="133D4D8F083D7D37AEF5B4E26A06F974" box="[473,521,1681,1705]" captionStart="Fig" captionStartId="20.[174,207,1910,1932]" captionText="Fig. 14. Phyllopodopsyllus busanensis sp. nov., holotype female, line drawings: A, antennula, dorsal. B, antenna, posterior; C, mandibula, anterior; D, praecoxal arthrite of maxillula, anterior; E, coxa of maxillula, anterior; F, endopod of maxillula, anterior; G, maxilla, posterior; H, maxilliped, anterior. Cuticular pits on antennula and antenna omitted for clarity." pageId="22" pageNumber="207">14A</figureCitation>
) nine-segmented, about as long as cephalothorax, with prominent integumental beak on second segment, with robust and long aesthetasc on fourth segment fused basally to slightly longer seta, slender and much shorter apical aesthetasc on ninth segment fused basally to two slightly longer setae, and setal formula 1.9.7.4.2.4.2.2.7. All setae slender and smooth; one lateral seta on seventh and eight segments and four lateral setae on sixth segment biarticulate. Only ornamentation short arched row of spinules at base of first segment, on dorso-median surface. Length ratio of antennular segments, from proximal end and along caudal margin, 1: 0.4: 0.2: 0.2: 0.15: 0.15: 0.1: 0.1: 0.4. First segment also strongest, about 2.9 times as long as wide; fourth segment with prominently protruded anteri- or distal corner at base of large easthetasc.
</paragraph>
<paragraph id="8BB9510A083D7D37AC72B3B46D4BFBDE" blockId="22.[839,1445,261,1995]" pageId="22" pageNumber="207">
Antenna (
<figureCitation id="133D4D8F083D7D37ACEBB3B46C38FE02" box="[967,1079,455,479]" captionStart="Fig" captionStartId="17.[144,177,1843,1865]" captionTargetBox="[223,1334,257,1816]" captionTargetId="figure-10@17.[223,1334,257,1816]" captionTargetPageId="17" captionText="Fig. 11. Phyllopodopsyllus busanensis sp. nov., paratype female, SEM photographs: A, habitus, lateral; B, cephalothorax (Ct), lateral; C, anterior part of cephalothorax (Ct) with rostrum(Ro) and first segment of antennula (A1), lateral; D, ventral part of cephalothorax (Ct), with parts of antennula (A1), antenna (A2), and mandibula, lateral; E, dorso-posterior part of cephalothorax (Ct), lateral; F, ventro-posterior corner of cephalothorax (Ct), lateral; G, tergites of three free prosomites (Fp1-Fp3), lateral; H, detail of ornamentation of first (Fp1) and second (Fp2) prosomites, lateral. Arabic numerals for cuticular sensilla and Roman numerals for pores assigned consecutively from anterior to posterior end of each somite and caudal ramus, and from dorsal to ventral side." pageId="22" pageNumber="207">Figs. 11D</figureCitation>
,
<figureCitation id="133D4D8F083D7D37AB69B3B46C7DFE02" box="[1093,1138,455,479]" captionStart="Fig" captionStartId="20.[174,207,1910,1932]" captionText="Fig. 14. Phyllopodopsyllus busanensis sp. nov., holotype female, line drawings: A, antennula, dorsal. B, antenna, posterior; C, mandibula, anterior; D, praecoxal arthrite of maxillula, anterior; E, coxa of maxillula, anterior; F, endopod of maxillula, anterior; G, maxilla, posterior; H, maxilliped, anterior. Cuticular pits on antennula and antenna omitted for clarity." pageId="22" pageNumber="207">14B</figureCitation>
) slenderer than in previous species, but with same segmentation and armature. Coxa 0.8 times as long as wide, unornamented. Allobasis 2.5 times as long as wide and about three times as long as coxa, ornamented with longitudinal row of minute spinules along inner margin. First endopodal segment 2.4 times as long as wide and about as long as allobasis, ornamented with single transverse arched row of minute spinules in proximal part. Second endopodal segment most robust, slightly slenderer proximally but also generally cylindrical, 3.7 times as long as wide, 1.4 times as long as first endopodal segment, with two surface frills distally and row of small spinules along inner margin. Exopod slender, cylindrical but slightly curved, about 3.6 times as long as wide and half as long as allobasis, unornamented; outer apical seta strongest and spiniform, fused basally to exopod, about 1.3 times as long as inner apical seta, and 0.8 times as long as lateral seta.
</paragraph>
<paragraph id="8BB9510A083D7D37AC72B6786BC3FB99" blockId="22.[839,1445,261,1995]" pageId="22" pageNumber="207">Labrum as in previous species, not mounted properly for drawing.</paragraph>
<paragraph id="8BB9510A083D7D37AC72B63F6D0FFABB" blockId="22.[839,1445,261,1995]" pageId="22" pageNumber="207">
Mandibula (
<figureCitation id="133D4D8F083D7D37ACCAB63F6C55FBB9" box="[998,1114,1100,1124]" captionStart="Fig" captionStartId="17.[144,177,1843,1865]" captionTargetBox="[223,1334,257,1816]" captionTargetId="figure-10@17.[223,1334,257,1816]" captionTargetPageId="17" captionText="Fig. 11. Phyllopodopsyllus busanensis sp. nov., paratype female, SEM photographs: A, habitus, lateral; B, cephalothorax (Ct), lateral; C, anterior part of cephalothorax (Ct) with rostrum(Ro) and first segment of antennula (A1), lateral; D, ventral part of cephalothorax (Ct), with parts of antennula (A1), antenna (A2), and mandibula, lateral; E, dorso-posterior part of cephalothorax (Ct), lateral; F, ventro-posterior corner of cephalothorax (Ct), lateral; G, tergites of three free prosomites (Fp1-Fp3), lateral; H, detail of ornamentation of first (Fp1) and second (Fp2) prosomites, lateral. Arabic numerals for cuticular sensilla and Roman numerals for pores assigned consecutively from anterior to posterior end of each somite and caudal ramus, and from dorsal to ventral side." pageId="22" pageNumber="207">Figs. 11D</figureCitation>
,
<figureCitation id="133D4D8F083D7D37AB44B63F6C98FBB9" box="[1128,1175,1100,1124]" captionStart="Fig" captionStartId="20.[174,207,1910,1932]" captionText="Fig. 14. Phyllopodopsyllus busanensis sp. nov., holotype female, line drawings: A, antennula, dorsal. B, antenna, posterior; C, mandibula, anterior; D, praecoxal arthrite of maxillula, anterior; E, coxa of maxillula, anterior; F, endopod of maxillula, anterior; G, maxilla, posterior; H, maxilliped, anterior. Cuticular pits on antennula and antenna omitted for clarity." pageId="22" pageNumber="207">14C</figureCitation>
) segmentation, general shape, and most armature and ornamentation as in previous species, but endopod proportionately smaller and exopod much larger and armed with three apical setae. Basis twice as long as wide. Endopod 3.9 times as long as wide, 0.7 times as long as basis, and 1.4 times as long as exopod. Exopod cylindrical, very slender, 4.8 times as long as wide, unornamented, armed with one lateral and four apical slender and smooth setae.
</paragraph>
<paragraph id="8BB9510A083D7D37AC72B71D6D6AF9F5" blockId="22.[839,1445,261,1995]" pageId="22" pageNumber="207">
Maxillula (
<figureCitation id="133D4D8F083D7D37ACFAB71D6C2BFA5B" box="[982,1060,1390,1414]" captionStart="Fig" captionStartId="20.[174,207,1910,1932]" captionText="Fig. 14. Phyllopodopsyllus busanensis sp. nov., holotype female, line drawings: A, antennula, dorsal. B, antenna, posterior; C, mandibula, anterior; D, praecoxal arthrite of maxillula, anterior; E, coxa of maxillula, anterior; F, endopod of maxillula, anterior; G, maxilla, posterior; H, maxilliped, anterior. Cuticular pits on antennula and antenna omitted for clarity." pageId="22" pageNumber="207">Fig. 14</figureCitation>
D-F) segmentation, general shape, and most armature and ornamentation as in previous species, but praexocal arthrite armed apically only seven elements, and coxa without outer seta but with five inner elements (one strong and geniculate, four slender and smooth); endopod about 2.7 times as long as wide.
</paragraph>
<paragraph id="8BB9510A083D7D37AC72B4436D34F9B5" blockId="22.[839,1445,261,1995]" pageId="22" pageNumber="207">
Maxilla (
<figureCitation id="133D4D8F083D7D37AC91B4436C11F995" box="[957,1054,1584,1608]" captionStart="Fig" captionStartId="20.[174,207,1910,1932]" captionText="Fig. 14. Phyllopodopsyllus busanensis sp. nov., holotype female, line drawings: A, antennula, dorsal. B, antenna, posterior; C, mandibula, anterior; D, praecoxal arthrite of maxillula, anterior; E, coxa of maxillula, anterior; F, endopod of maxillula, anterior; G, maxilla, posterior; H, maxilliped, anterior. Cuticular pits on antennula and antenna omitted for clarity." pageId="22" pageNumber="207">Fig. 14G</figureCitation>
) segmentation, general shape, armature, and ornamentation as in previous species.
</paragraph>
<paragraph id="8BB9510A083D7D37AC72B4036D2AF816" blockId="22.[839,1445,261,1995]" pageId="22" pageNumber="207">
Maxilliped (
<figureCitation id="133D4D8F083D7D37ACCBB4036C54F954" box="[999,1115,1648,1673]" captionStart="Fig" captionStartId="18.[174,207,1843,1865]" captionTargetBox="[253,1364,258,1816]" captionTargetId="figure-11@18.[253,1364,258,1816]" captionTargetPageId="18" captionText="Fig. 12. Phyllopodopsyllus busanensis sp. nov., paratype female, SEM photographs: A, first five urosomites (U1-U5) with fifth leg (P5), lateral; B, tergite of first urosomite (U1), lateral; C, parts of fused second (U2) and third (U3) urosomites, with lateral part of fifth leg (P5), lateral; D, part of fourth urosomite (U4) and distal part of fifth leg (P5); E, fourth (U4) and fifth (U5) urosomites, lateral; F, sixth urosomite and caudal rami(Cr), lateral; G, posterior part of caudal rami(Cr), lateral; H, parts of maxilliped (Mxp) and first swimming leg (P1), lateral. Arabic numerals for cuticular sensilla and Roman numerals for pores assigned consecutively from anterior to posterior end of each somite and caudal ramus, and from dorsal to ventral side." pageId="22" pageNumber="207">Figs. 12H</figureCitation>
,
<figureCitation id="133D4D8F083D7D37AB45B4026C95F954" box="[1129,1178,1649,1673]" captionStart="Fig" captionStartId="20.[174,207,1910,1932]" captionText="Fig. 14. Phyllopodopsyllus busanensis sp. nov., holotype female, line drawings: A, antennula, dorsal. B, antenna, posterior; C, mandibula, anterior; D, praecoxal arthrite of maxillula, anterior; E, coxa of maxillula, anterior; F, endopod of maxillula, anterior; G, maxilla, posterior; H, maxilliped, anterior. Cuticular pits on antennula and antenna omitted for clarity." pageId="22" pageNumber="207">14H</figureCitation>
) segmentation, general shape, and most armature and ornamentation as in previous species, but all spinules shorter and endopod armed with additional smooth and slender seta. Syncoxa 2.5 times as long as wide, cylindrical, and only slightly restricted in central part. Basis 3.4 times as long as wide and 1.1 times as long as syncoxa, cylindrical but wider in central part. Endopod nearly three times as long as wide and 0.3 times as long as basis, cylindrical but narrower in proximal half; apical spine 2.2 times as long as basis and 1.6 times as long as subapical seta.
</paragraph>
<paragraph id="8BB9510A083C7D36AF8AB3766AE7FE80" blockId="23.[144,749,261,1995]" pageId="23" pageNumber="208">
Swimming legs (
<figureCitation id="133D4D8F083C7D36AE70B37669A8FEC0" box="[348,423,261,285]" captionStart="Fig" captionStartId="21.[144,177,1727,1749]" captionTargetBox="[178,1348,262,1697]" captionTargetId="figure-97@21.[167,1404,261,1697]" captionTargetPageId="21" captionText="Fig. 15. Phyllopodopsyllus busanensis sp. nov., holotype female, line drawings: A, first swimming leg, anterior; B, second swimming leg, anterior; C, third swimming leg, anterior; D, fourth swimming leg, anterior; E, fifth leg, compressed and slightly deformed, anterior." pageId="23" pageNumber="208">Fig. 15</figureCitation>
A-D) slightly shorter and less slender than in previous species, but with same segmentation, general shape, most ornamentation and armature.
</paragraph>
<paragraph id="8BB9510A083C7D36AF8AB3156A6BFC7F" blockId="23.[144,749,261,1995]" pageId="23" pageNumber="208">
First swimming leg (
<figureCitation id="133D4D8F083C7D36AEA6B31569E2FEA3" box="[394,493,358,382]" captionStart="Fig" captionStartId="21.[144,177,1727,1749]" captionTargetBox="[178,1348,262,1697]" captionTargetId="figure-97@21.[167,1404,261,1697]" captionTargetPageId="21" captionText="Fig. 15. Phyllopodopsyllus busanensis sp. nov., holotype female, line drawings: A, first swimming leg, anterior; B, second swimming leg, anterior; C, third swimming leg, anterior; D, fourth swimming leg, anterior; E, fifth leg, compressed and slightly deformed, anterior." pageId="23" pageNumber="208">Fig. 15A</figureCitation>
) with prehensile endopod as in previous species, but coxa and basis wider, fist endopodal segment shorter and additionally armed with inner seta, and first exopodal segment with inner bulge. Coxa 0.8 times as long as wide, narrower in distal part than proximal, with short outer spiniform process, ornamented with three anterior and one posterior row of spinules. Basis about as long as wide, inner spine slightly stronger and about 1.4 times as long as outer. Exopod with first segment longest and third shortest; inner geniculate seta on third segment 1.3 times as long as entire exopod and about 1.6 times as long as outer geniculate seta. Endopod longer than exopod; first endopodal segment 1.15 times as long as entire exopod, more than six times as long as wide, and 3.1 times as long as second endopodal segment; endopodal apical seta as long as exopodal outer apical seta but slightly stronger, and about 1.4 times as long as apical endopodal spine.
</paragraph>
<paragraph id="8BB9510A083C7D36AF8AB1D86A48FA98" blockId="23.[144,749,261,1995]" pageId="23" pageNumber="208">
Second swimming leg (
<figureCitation id="133D4D8F083C7D36AE83B1D86A1BFC1F" box="[431,532,938,963]" captionStart="Fig" captionStartId="21.[144,177,1727,1749]" captionTargetBox="[178,1348,262,1697]" captionTargetId="figure-97@21.[167,1404,261,1697]" captionTargetPageId="21" captionText="Fig. 15. Phyllopodopsyllus busanensis sp. nov., holotype female, line drawings: A, first swimming leg, anterior; B, second swimming leg, anterior; C, third swimming leg, anterior; D, fourth swimming leg, anterior; E, fifth leg, compressed and slightly deformed, anterior." pageId="23" pageNumber="208">Fig. 15B</figureCitation>
) with all segments shorter than in previous species, with two large spiniform processes on basis, and additionally armed with inner spiniform seta on first endopodal segment, as well as with inner slender seta on third exopodal segment. Coxa 0.7 times as long as wide, ornamented with six rows of small spinules on anterior surface and one row of spinules on posterior surface. All exopodal segments of similar length. Endopod about 0.6 times as long as exopod, first segment 0.8 times as long as second but much wider; apical edopodal spine about 0.7 times as long as endopod, 0.6 times as long as central slender seta, and less than half as long as inner apical seta.
</paragraph>
<paragraph id="8BB9510A083C7D36AF8AB73D6958FA3A" blockId="23.[144,749,261,1995]" pageId="23" pageNumber="208">
Third swimming leg (
<figureCitation id="133D4D8F083C7D36AEB6B73D69F0FABB" box="[410,511,1358,1382]" captionStart="Fig" captionStartId="21.[144,177,1727,1749]" captionTargetBox="[178,1348,262,1697]" captionTargetId="figure-97@21.[167,1404,261,1697]" captionTargetPageId="21" captionText="Fig. 15. Phyllopodopsyllus busanensis sp. nov., holotype female, line drawings: A, first swimming leg, anterior; B, second swimming leg, anterior; C, third swimming leg, anterior; D, fourth swimming leg, anterior; E, fifth leg, compressed and slightly deformed, anterior." pageId="23" pageNumber="208">Fig. 15C</figureCitation>
) very similar to second, except slightly slenderer and with longer inner seta on first endopodal segment; endopod 0.4 times as long as exopod; first endopodal segment 0.7 times as long as second endopodal.
</paragraph>
<paragraph id="8BB9510A083C7D36AF8AB79C6ACEF9F5" blockId="23.[144,749,261,1995]" pageId="23" pageNumber="208">
Fourth swimming leg (
<figureCitation id="133D4D8F083C7D36AEB5B79C69F5F9DA" box="[409,506,1519,1543]" captionStart="Fig" captionStartId="21.[144,177,1727,1749]" captionTargetBox="[178,1348,262,1697]" captionTargetId="figure-97@21.[167,1404,261,1697]" captionTargetPageId="21" captionText="Fig. 15. Phyllopodopsyllus busanensis sp. nov., holotype female, line drawings: A, first swimming leg, anterior; B, second swimming leg, anterior; C, third swimming leg, anterior; D, fourth swimming leg, anterior; E, fifth leg, compressed and slightly deformed, anterior." pageId="23" pageNumber="208">Fig. 15D</figureCitation>
) very similar to previous species, except for spiniform processes on basis.
</paragraph>
<paragraph id="8BB9510A083C7D36AF8AB4436A6EF8D7" blockId="23.[144,749,261,1995]" pageId="23" pageNumber="208">
Fifth leg (
<figureCitation id="133D4D8F083C7D36AE23B4436961F995" box="[271,366,1584,1608]" captionStart="Fig" captionStartId="3.[144,177,1234,1256]" captionTargetBox="[177,1380,257,1202]" captionTargetId="figure-358@3.[177,1380,257,1202]" captionTargetPageId="3" captionText="Fig. 1. Light photograph of two new Korean Phyllopodopsyllus Scot T., 1906 species, on the same slide in glycerol:A, P. kitazimai sp. nov., holotype female; B, P. kitazimai sp. nov., allotype male; C, P. busanensis sp. nov., holotype female." pageId="23" pageNumber="208">Figs. 1A</figureCitation>
,
<figureCitation id="133D4D8F083C7D36AE57B44369E3F99A" box="[379,492,1584,1608]" captionStart="Fig" captionStartId="18.[174,207,1843,1865]" captionTargetBox="[253,1364,258,1816]" captionTargetId="figure-11@18.[253,1364,258,1816]" captionTargetPageId="18" captionText="Fig. 12. Phyllopodopsyllus busanensis sp. nov., paratype female, SEM photographs: A, first five urosomites (U1-U5) with fifth leg (P5), lateral; B, tergite of first urosomite (U1), lateral; C, parts of fused second (U2) and third (U3) urosomites, with lateral part of fifth leg (P5), lateral; D, part of fourth urosomite (U4) and distal part of fifth leg (P5); E, fourth (U4) and fifth (U5) urosomites, lateral; F, sixth urosomite and caudal rami(Cr), lateral; G, posterior part of caudal rami(Cr), lateral; H, parts of maxilliped (Mxp) and first swimming leg (P1), lateral. Arabic numerals for cuticular sensilla and Roman numerals for pores assigned consecutively from anterior to posterior end of each somite and caudal ramus, and from dorsal to ventral side." pageId="23" pageNumber="208">12A, C, D</figureCitation>
,
<figureCitation id="133D4D8F083C7D36AED4B4436A2BF995" box="[504,548,1584,1608]" captionStart="Fig" captionStartId="21.[144,177,1727,1749]" captionTargetBox="[178,1348,262,1697]" captionTargetId="figure-97@21.[167,1404,261,1697]" captionTargetPageId="21" captionText="Fig. 15. Phyllopodopsyllus busanensis sp. nov., holotype female, line drawings: A, first swimming leg, anterior; B, second swimming leg, anterior; C, third swimming leg, anterior; D, fourth swimming leg, anterior; E, fifth leg, compressed and slightly deformed, anterior." pageId="23" pageNumber="208">15E</figureCitation>
) shape, segmentation, and armature as in previous species, but covered by dense spinules in outer part, with serrate outer margin and smooth distal margin; short exopodal setae smooth or sparsely pinnate; forming brooding chamber with genital double-somite and anterior part of fourth urosomite normally containing six to
<specimenCount id="9D009A83083C7D36AEC9B4816A52F8D7" box="[485,605,1778,1802]" count="7" pageId="23" pageNumber="208" type="egg">seven eggs</specimenCount>
.
</paragraph>
<paragraph id="8BB9510A083C7D36AF8AB5616988F816" blockId="23.[144,749,261,1995]" pageId="23" pageNumber="208">
Sixth leg (
<figureCitation id="133D4D8F083C7D36AE37B561698FF8F7" box="[283,384,1810,1834]" captionStart="Fig" captionStartId="19.[144,177,1424,1446]" captionTargetBox="[178,1372,263,1388]" captionTargetId="figure-199@19.[189,1429,262,1396]" captionTargetPageId="19" captionText="Fig. 13. Phyllopodopsyllus busanensis sp. nov., holotype female, line drawings: A, second to sixth urosomites (U2-U6) and caudal rami, dorsal; B, second to sixth urosomites (U2-U6), with caudal rami (Cr), and attached spermatophore (Sp), lateral; C, second to sixth urosomites (U2-U6), with caudal rami (Cr), sixth legs (P6), attached spermatophore (Sp), and internal reseptacula seminis (Rs), ventral. Arabic numerals for cuticular sensilla and Roman numerals for pores assigned consecutively from anterior to posterior end of each somite and caudal ramus, and from dorsal to ventral side. Cuticular pits and minute surface spinules omitted for clarity." pageId="23" pageNumber="208">Fig. 13C</figureCitation>
) shape and ornamentation as in previous species; also armed with three setae, but outermost seta extremely slender and shortest, while central seta longest, pinnate, not reaching posterior margin of genital double-somite, and more than twice as long as innermost pinnate seta.
</paragraph>
<paragraph id="8BB9510A083C7D36AC13B3766BE7FEC0" blockId="23.[809,1414,261,479]" box="[831,1000,261,285]" pageId="23" pageNumber="208">Male unknown.</paragraph>
<paragraph id="8BB9510A083C7D36AC05B3566CF4FE02" blockId="23.[809,1414,261,479]" pageId="23" pageNumber="208">
<emphasis id="B9728D18083C7D36AC05B3566BA4FEE0" bold="true" box="[809,939,293,317]" pageId="23" pageNumber="208">Variability.</emphasis>
No obviously variable structures were observed between the two collected females, but the comparison was hindered by the fact that they were examined by different methods: the
<typeStatus id="54BDEFA8083C7D36ABA9B3F56CEBFE43" box="[1157,1252,390,414]" pageId="23" pageNumber="208" type="holotype">holotype</typeStatus>
was dissected and studied using light microscopy, while the
<typeStatus id="54BDEFA8083C7D36AA05B3D46D89FE62" box="[1321,1414,423,447]" pageId="23" pageNumber="208" type="paratype">paratype</typeStatus>
was permanently mounted on an SEM stub.
</paragraph>
</subSubSection>
</treatment>
</document>