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<document id="0604B52999E7933EA2C9AFA90CF86C82" ID-ISSN="1175­5334" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629574900641" checkinUser="felipe" docAuthor="Tormos, José, Polidori, Carlo, Asís, Josep Daniel &amp; Gayubo, S. F." docDate="2008" docId="03AC122CFFA1FFF2AAE1FFE3FD2EFE4F" docLanguage="en" docName="zt01946p054.pdf" docOrigin="Zootaxa 1946" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Symmorphus murarius" docType="treatment" docVersion="3" lastPageNumber="50" masterDocId="FF956A54FFA7FFFAAA76F96FFFAEFF85" masterDocTitle="Description of mature larvae of Allodynerus rossii (Lepeletier), Ancistrocerus auctus (Fabricius), Euodynerus dantici (Rossi) and Symmorphus murarius (Linnaeus) (Hymenoptera, Vespidae)" masterLastPageNumber="54" masterPageNumber="42" pageNumber="48" updateTime="1699078586606" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="AEC971E967517D46775605118C31D376">Description of mature larvae of Allodynerus rossii (Lepeletier), Ancistrocerus auctus (Fabricius), Euodynerus dantici (Rossi) and Symmorphus murarius (Linnaeus) (Hymenoptera, Vespidae)</mods:title>
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<mods:namePart id="17CE9DE46D6FCCDFD02A06CE8C171C48">Tormos, José</mods:namePart>
<mods:affiliation id="678C04E7B6487E514CBC85736B04EFBB">Área de Zoología, Facultad de Biología, Universidad de Salamanca, 37071 - Salamanca, Spain. E-mail: tormos @ usal. es Dipartimento di Biologia, Sezione di Zoologia e Citologia, Università degli Studi di Milano, Via Celoria, 26, 20133, Milan, Italy</mods:affiliation>
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<mods:namePart id="5700BBF76CBFA1968AF568F056FF0DC1">Polidori, Carlo</mods:namePart>
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<mods:namePart id="5D006F7662A2C06CF57B3A566DECE5E6">Asís, Josep Daniel</mods:namePart>
<mods:affiliation id="7E4E94CC1C75948B1F22FFF0F4CD8BBA">Área de Zoología, Facultad de Biología, Universidad de Salamanca, 37071 - Salamanca, Spain. E-mail: tormos @ usal. es Dipartimento di Biologia, Sezione di Zoologia e Citologia, Università degli Studi di Milano, Via Celoria, 26, 20133, Milan, Italy</mods:affiliation>
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<mods:namePart id="B63805022FDF9DE1D691565C04C64100">Gayubo, S. F.</mods:namePart>
<mods:affiliation id="7E78408DDE46D94FA7392818AEC47A4B">Área de Zoología, Facultad de Biología, Universidad de Salamanca, 37071 - Salamanca, Spain. E-mail: tormos @ usal. es Dipartimento di Biologia, Sezione di Zoologia e Citologia, Università degli Studi di Milano, Via Celoria, 26, 20133, Milan, Italy</mods:affiliation>
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<treatment id="03AC122CFFA1FFF2AAE1FFE3FD2EFE4F" ID-DOI="http://doi.org/10.5281/zenodo.5231659" ID-Zenodo-Dep="5231659" LSID="urn:lsid:plazi:treatment:03AC122CFFA1FFF2AAE1FFE3FD2EFE4F" httpUri="http://treatment.plazi.org/id/03AC122CFFA1FFF2AAE1FFE3FD2EFE4F" lastPageId="8" lastPageNumber="50" pageId="6" pageNumber="48">
<subSubSection id="C31FF0B1FFA1FFFCAAE1FFE3FD92F923" box="[151,572,1676,1702]" pageId="6" pageNumber="48" type="nomenclature">
<paragraph id="8BBAA33AFFA1FFFCAAE1FFE3FD92F923" blockId="6.[151,572,1676,1737]" box="[151,572,1676,1702]" pageId="6" pageNumber="48">
<heading id="D0F21456FFA1FFFCAAE1FFE3FD92F923" bold="true" box="[151,572,1676,1702]" fontSize="11" level="1" pageId="6" pageNumber="48" reason="1">
<taxonomicName id="4C05D8B9FFA1FFFCAAE1FFE3FD92F923" ID-CoL="53NZL" authority="(Linnaeus)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[151,572,1676,1702]" class="Insecta" family="Eumenidae" genus="Symmorphus" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="48" phylum="Arthropoda" rank="species" species="murarius">
<emphasis id="B9717F28FFA1FFFCAAE1FFE3FD92F923" bold="true" box="[151,572,1676,1702]" pageId="6" pageNumber="48">
<emphasis id="B9717F28FFA1FFFCAAE1FFE3FE00F923" bold="true" box="[151,430,1676,1702]" italics="true" pageId="6" pageNumber="48">Symmorphus murarius</emphasis>
(Linnaeus)
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<subSubSection id="C31FF0B1FFA1FFF2AAE1FFC0FD2EFE4F" lastPageId="8" lastPageNumber="50" pageId="6" pageNumber="48" type="description">
<paragraph id="8BBAA33AFFA1FFFCAAE1FFC0FE9AF94C" blockId="6.[151,572,1676,1737]" box="[151,308,1711,1737]" pageId="6" pageNumber="48">
(
<figureCitation id="133EBFBFFFA1FFFCAAE9FFC0FE82F94C" box="[159,300,1711,1737]" captionStart-0="FIGURES 2830" captionStart-1="FIGURE 31" captionStartId-0="9.[151,269,1548,1572]" captionStartId-1="10.[151,255,1381,1405]" captionTargetBox-0="[190,1393,384,1524]" captionTargetBox-1="[242,1348,321,1346]" captionTargetId-0="figure-294@9.[185,1406,384,1528]" captionTargetId-1="figure-283@10.[239,1349,318,1352]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURES 2830. Mature larva of Symmorphus murarius: (28) body, lateral view (anus (a), transverse crease on abdominal segment (tc), spiracle (s)); (29) integument: setae (se, sed (detail of a seta)), spiracle (s). (30) prothoracic spiracle (arrows point the spinules (s) and the spines (sp))." captionText-1="FIGURE 31. Mature larva of Symmorphus murarius: cranium, lateral view (detail of setae (a), antennal papilla (b), labrum (c) and mouthparts (d): maxilla (m), maxillary palpus (mp), galea (g), labial palpus (lp), salivary orifice (so)." pageId="6" pageNumber="48">Figs. 2831</figureCitation>
)
</paragraph>
<paragraph id="8BBAA33AFFA1FFFCAAE1FF90FD8FF83C" blockId="6.[151,1436,1791,1977]" pageId="6" pageNumber="48">
A nest of this species (length =
<quantity id="4CFD0EDFFFA1FFFCA87FFF90FDF0F89C" box="[521,606,1791,1817]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="48" unit="mm" value="80.0">80 mm</quantity>
; diameter =
<quantity id="4CFD0EDFFFA1FFFCA886FF90FC9BF89C" box="[752,821,1791,1817]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="48" unit="mm" value="6.0">6 mm</quantity>
) was obtained from a trap-nest (number 465) placed in Ticino Regional Park (Riserva Orientata La Fagiana, Milano,
<collectingCountry id="F312E3AAFFA1FFFCA9ECFE48FC7FF8C4" box="[922,977,1831,1857]" name="Italy" pageId="6" pageNumber="48">Italy</collectingCountry>
) in 2003. The nest had 3 cells
<quantity id="4CFD0EDFFFA1FFFCAF31FE48FA35F8C4" box="[1351,1435,1831,1857]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="6" pageNumber="48" unit="mm" value="18.0">18 mm</quantity>
long on average, from one of which emerged a male (
<date id="FFBB85FAFFA1FFFCA969FE20FC49F8EC" box="[799,999,1871,1897]" pageId="6" pageNumber="48" value="2003-09">September 2003</date>
);
<specimenCount id="9D0368B3FFA1FFFCA988FE20FBD5F8EC" box="[1022,1147,1871,1897]" pageId="6" pageNumber="48" type="larva">two larvae</specimenCount>
were obtained from the other two cells. The nest lacked of the typical mud closing plug, revealing that it was not completely provisioned at the moment of removal.
</paragraph>
<caption id="DF7AF3B2FFA0FFFDAAE1FCF4FE6EFA7D" pageId="7" pageNumber="49" startId="7.[151,269,1435,1459]" targetBox="[192,1402,200,1410]" targetPageId="7">
<paragraph id="8BBAA33AFFA0FFFDAAE1FCF4FE6EFA7D" blockId="7.[151,1436,1435,1528]" pageId="7" pageNumber="49">
<emphasis id="B9717F28FFA0FFFDAAE1FCF4FEF1FA36" bold="true" box="[151,351,1435,1459]" pageId="7" pageNumber="49">FIGURES 2023.</emphasis>
Mature larva of
<taxonomicName id="4C05D8B9FFA0FFFDA863FCF4FD49FA37" box="[533,743,1435,1458]" class="Insecta" family="Eumenidae" genus="Euodynerus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="49" phylum="Arthropoda" rank="species" species="dantici">
<emphasis id="B9717F28FFA0FFFDA863FCF4FD49FA37" box="[533,743,1435,1458]" italics="true" pageId="7" pageNumber="49">Euodynerus dantici</emphasis>
</taxonomicName>
: (20) last abdominal segment: anus (a), setae of integument (b); (21) first abdominal spiracle; (22) detail of anterior spiracle showing the asperities (as); (23) cranium (frontal view): parietal bands (pb), antenna (a).
</paragraph>
</caption>
<paragraph id="8BBAA33AFFA0FFFDAAB0FF49FD8BF8B5" blockId="7.[151,1436,1574,2000]" pageId="7" pageNumber="49">
<emphasis id="B9717F28FFA0FFFDAAB0FF49FEACF9BA" box="[198,258,1574,1599]" italics="true" pageId="7" pageNumber="49">Body</emphasis>
(
<figureCitation id="133EBFBFFFA0FFFDAB65FF49FEC5F9C5" box="[275,363,1574,1600]" captionStart="FIGURES 2830" captionStartId="9.[151,269,1548,1572]" captionTargetBox="[190,1393,384,1524]" captionTargetId="figure-294@9.[185,1406,384,1528]" captionTargetPageId="9" captionText="FIGURES 2830. Mature larva of Symmorphus murarius: (28) body, lateral view (anus (a), transverse crease on abdominal segment (tc), spiracle (s)); (29) integument: setae (se, sed (detail of a seta)), spiracle (s). (30) prothoracic spiracle (arrows point the spinules (s) and the spines (sp))." pageId="7" pageNumber="49">Fig. 28</figureCitation>
) (l =
<quantity id="4CFD0EDFFFA0FFFDABD9FF49FDA3F9C5" box="[431,525,1574,1600]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="7" pageNumber="49" unit="mm" value="8.5">8.5 mm</quantity>
, w =
<quantity id="4CFD0EDFFFA0FFFDA824FF49FD36F9C5" box="[594,664,1574,1600]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="49" unit="mm" value="3.0">3 mm</quantity>
) fusiform, robust, with dorsum of thoracic and first 8 abdominal segments divided into two annulets by transverse crease (
<figureCitation id="133EBFBFFFA0FFFDA945FF21FC24F9ED" box="[819,906,1614,1640]" captionStart="FIGURES 2830" captionStartId="9.[151,269,1548,1572]" captionTargetBox="[190,1393,384,1524]" captionTargetId="figure-294@9.[185,1406,384,1528]" captionTargetPageId="9" captionText="FIGURES 2830. Mature larva of Symmorphus murarius: (28) body, lateral view (anus (a), transverse crease on abdominal segment (tc), spiracle (s)); (29) integument: setae (se, sed (detail of a seta)), spiracle (s). (30) prothoracic spiracle (arrows point the spinules (s) and the spines (sp))." pageId="7" pageNumber="49">Fig. 28</figureCitation>
, tc). Anus (
<figureCitation id="133EBFBFFFA0FFFDAE66FF21FBC8F9ED" box="[1040,1126,1614,1640]" captionStart="FIGURES 2830" captionStartId="9.[151,269,1548,1572]" captionTargetBox="[190,1393,384,1524]" captionTargetId="figure-294@9.[185,1406,384,1528]" captionTargetPageId="9" captionText="FIGURES 2830. Mature larva of Symmorphus murarius: (28) body, lateral view (anus (a), transverse crease on abdominal segment (tc), spiracle (s)); (29) integument: setae (se, sed (detail of a seta)), spiracle (s). (30) prothoracic spiracle (arrows point the spinules (s) and the spines (sp))." pageId="7" pageNumber="49">Fig. 28</figureCitation>
, a) as transverse slit. Pleural lobes weakly developed. Integument with setae (
<figureCitation id="133EBFBFFFA0FFFDA972FF19FCF3F915" box="[772,861,1654,1680]" captionStart="FIGURES 2830" captionStartId="9.[151,269,1548,1572]" captionTargetBox="[190,1393,384,1524]" captionTargetId="figure-294@9.[185,1406,384,1528]" captionTargetPageId="9" captionText="FIGURES 2830. Mature larva of Symmorphus murarius: (28) body, lateral view (anus (a), transverse crease on abdominal segment (tc), spiracle (s)); (29) integument: setae (se, sed (detail of a seta)), spiracle (s). (30) prothoracic spiracle (arrows point the spinules (s) and the spines (sp))." pageId="7" pageNumber="49">Fig. 29</figureCitation>
, se, sed) (l =
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), arranged in transverse band behind of the spiracles; 710 abdominal segments with scattered setae and sparse minute spinules on anterior margin of each segment. First two spiracles (
<figureCitation id="133EBFBFFFA0FFFDA96CFFA9FC0FF965" box="[794,929,1734,1760]" captionStart="FIGURES 2830" captionStartId="9.[151,269,1548,1572]" captionTargetBox="[190,1393,384,1524]" captionTargetId="figure-294@9.[185,1406,384,1528]" captionTargetPageId="9" captionText="FIGURES 2830. Mature larva of Symmorphus murarius: (28) body, lateral view (anus (a), transverse crease on abdominal segment (tc), spiracle (s)); (29) integument: setae (se, sed (detail of a seta)), spiracle (s). (30) prothoracic spiracle (arrows point the spinules (s) and the spines (sp))." pageId="7" pageNumber="49">Fig. 28, 30</figureCitation>
) with subatrium (d =
<quantity id="4CFD0EDFFFA0FFFDAEDAFFA9FAA4F965" box="[1196,1290,1734,1760]" metricMagnitude="-1" metricUnit="m" metricValue="1.8" pageId="7" pageNumber="49" unit="mm" value="180.0">180 µm</quantity>
) larger than atrium (d =
<quantity id="4CFD0EDFFFA0FFFDAB57FF81FED0F88D" box="[289,382,1774,1800]" metricMagnitude="-1" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="49" unit="mm" value="130.0">130 µm</quantity>
), and 1.2 times wider as successive ones (d =
<quantity id="4CFD0EDFFFA0FFFDA9E0FF81FC5DF88D" box="[918,1011,1774,1800]" metricMagnitude="-1" metricUnit="m" metricValue="1.1" pageId="7" pageNumber="49" unit="mm" value="110.0">110 µm</quantity>
); walls of atrium spinulose; opening into subatrium armed with spines.
</paragraph>
<paragraph id="8BBAA33AFFA0FFF2AAB3FE51FF77FF5F" blockId="7.[151,1436,1574,2000]" lastBlockId="8.[151,1436,152,458]" lastPageId="8" lastPageNumber="50" pageId="7" pageNumber="49">
<emphasis id="B9717F28FFA0FFFDAAB3FE51FE83F8D2" box="[197,301,1854,1879]" italics="true" pageId="7" pageNumber="49">Cranium</emphasis>
(
<figureCitation id="133EBFBFFFA0FFFDAB49FE51FE3AF8DD" box="[319,404,1854,1880]" captionStart="FIGURE 31" captionStartId="10.[151,255,1381,1405]" captionTargetBox="[242,1348,321,1346]" captionTargetId="figure-283@10.[239,1349,318,1352]" captionTargetPageId="10" captionText="FIGURE 31. Mature larva of Symmorphus murarius: cranium, lateral view (detail of setae (a), antennal papilla (b), labrum (c) and mouthparts (d): maxilla (m), maxillary palpus (mp), galea (g), labial palpus (lp), salivary orifice (so)." pageId="7" pageNumber="49">Fig. 31</figureCitation>
) (w =
<quantity id="4CFD0EDFFFA0FFFDAB93FE51FDEDF8DD" box="[485,579,1854,1880]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="7" pageNumber="49" unit="mm" value="1.1">1.1 mm</quantity>
, h (excluding of labrum) =
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), with setigerous punctures and scattered setae (
<figureCitation id="133EBFBFFFA0FFFDAB52FE09FE29F805" box="[292,391,1894,1920]" captionStart="FIGURE 31" captionStartId="10.[151,255,1381,1405]" captionTargetBox="[242,1348,321,1346]" captionTargetId="figure-283@10.[239,1349,318,1352]" captionTargetPageId="10" captionText="FIGURE 31. Mature larva of Symmorphus murarius: cranium, lateral view (detail of setae (a), antennal papilla (b), labrum (c) and mouthparts (d): maxilla (m), maxillary palpus (mp), galea (g), labial palpus (lp), salivary orifice (so)." pageId="7" pageNumber="49">Fig. 31a</figureCitation>
) (l=
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) more numerous near insertion of mandibles. Coronal suture and parietal bands present. Antennal orbits circular (d =
<quantity id="4CFD0EDFFFA0FFFDA8C2FEE1FCBCF82D" box="[692,786,1934,1960]" metricMagnitude="-1" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="49" unit="mm" value="130.0">130 µm</quantity>
) with papilla (
<figureCitation id="133EBFBFFFA0FFFDA9B1FEE1FB81F82D" box="[967,1071,1934,1960]" captionStart="FIGURE 31" captionStartId="10.[151,255,1381,1405]" captionTargetBox="[242,1348,321,1346]" captionTargetId="figure-283@10.[239,1349,318,1352]" captionTargetPageId="10" captionText="FIGURE 31. Mature larva of Symmorphus murarius: cranium, lateral view (detail of setae (a), antennal papilla (b), labrum (c) and mouthparts (d): maxilla (m), maxillary palpus (mp), galea (g), labial palpus (lp), salivary orifice (so)." pageId="7" pageNumber="49">Fig. 31b</figureCitation>
) (d =
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) with 3 sensilla. Clypeus with scattered setae (l =
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). Labrum (
<figureCitation id="133EBFBFFFA0FFFDA96EFED9FCD4F855" box="[792,890,1974,2000]" captionStart="FIGURE 31" captionStartId="10.[151,255,1381,1405]" captionTargetBox="[242,1348,321,1346]" captionTargetId="figure-283@10.[239,1349,318,1352]" captionTargetPageId="10" captionText="FIGURE 31. Mature larva of Symmorphus murarius: cranium, lateral view (detail of setae (a), antennal papilla (b), labrum (c) and mouthparts (d): maxilla (m), maxillary palpus (mp), galea (g), labial palpus (lp), salivary orifice (so)." pageId="7" pageNumber="49">Fig. 31c</figureCitation>
) (w =
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) strongly bilobed, with 18 short conical sensilla (w =
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) on each side. Epipharynx with 6 protuberant sensilla on distal edges of both sides.
</paragraph>
<paragraph id="8BBAA33AFFAFFFF2AAB3F987FD2EFE4F" blockId="8.[151,1436,152,458]" pageId="8" pageNumber="50">
<emphasis id="B9717F28FFAFFFF2AAB3F987FEE3FE84" box="[197,333,232,257]" italics="true" pageId="8" pageNumber="50">Mouthparts</emphasis>
. Mandibles (l = 500, w =
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) tridentate, brown; tooth truncate apically. Maxillae (
<figureCitation id="133EBFBFFFAFFFF2AF1BF987FF47FEAF" captionStart="FIGURE 31" captionStartId="10.[151,255,1381,1405]" captionTargetBox="[242,1348,321,1346]" captionTargetId="figure-283@10.[239,1349,318,1352]" captionTargetPageId="10" captionText="FIGURE 31. Mature larva of Symmorphus murarius: cranium, lateral view (detail of setae (a), antennal papilla (b), labrum (c) and mouthparts (d): maxilla (m), maxillary palpus (mp), galea (g), labial palpus (lp), salivary orifice (so)." pageId="8" pageNumber="50">Fig. 31d, m</figureCitation>
) truncate at apex, external surface with setae (l =
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), upper surface of apical portion spinulose; maxillary palpi (
<figureCitation id="133EBFBFFFAFFFF2AB2EF857FE13FED7" box="[344,445,312,338]" captionStart="FIGURE 31" captionStartId="10.[151,255,1381,1405]" captionTargetBox="[242,1348,321,1346]" captionTargetId="figure-283@10.[239,1349,318,1352]" captionTargetPageId="10" captionText="FIGURE 31. Mature larva of Symmorphus murarius: cranium, lateral view (detail of setae (a), antennal papilla (b), labrum (c) and mouthparts (d): maxilla (m), maxillary palpus (mp), galea (g), labial palpus (lp), salivary orifice (so)." pageId="8" pageNumber="50">Fig. 31d</figureCitation>
, mp) (50 x
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) with 2 apical sensilla (d = 8, 10 µm); galeae (
<figureCitation id="133EBFBFFFAFFFF2AEDDF857FA85FED7" box="[1195,1323,312,338]" captionStart="FIGURE 31" captionStartId="10.[151,255,1381,1405]" captionTargetBox="[242,1348,321,1346]" captionTargetId="figure-283@10.[239,1349,318,1352]" captionTargetPageId="10" captionText="FIGURE 31. Mature larva of Symmorphus murarius: cranium, lateral view (detail of setae (a), antennal papilla (b), labrum (c) and mouthparts (d): maxilla (m), maxillary palpus (mp), galea (g), labial palpus (lp), salivary orifice (so)." pageId="8" pageNumber="50">Fig. 31d, g</figureCitation>
) (20 x
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). Labium (w =
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) with short palpi (
<figureCitation id="133EBFBFFFAFFFF2A8DEF80FFCA2FEFF" box="[680,780,352,378]" captionStart="FIGURE 31" captionStartId="10.[151,255,1381,1405]" captionTargetBox="[242,1348,321,1346]" captionTargetId="figure-283@10.[239,1349,318,1352]" captionTargetPageId="10" captionText="FIGURE 31. Mature larva of Symmorphus murarius: cranium, lateral view (detail of setae (a), antennal papilla (b), labrum (c) and mouthparts (d): maxilla (m), maxillary palpus (mp), galea (g), labial palpus (lp), salivary orifice (so)." pageId="8" pageNumber="50">Fig. 31d</figureCitation>
, lp) (
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x
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), which have 4 apical sensilla; prementum with four setae (l =
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) in area ventral to each palpus; salivar orifice (
<figureCitation id="133EBFBFFFAFFFF2AE13F8E7FB65FE27" box="[1125,1227,392,418]" captionStart="FIGURE 31" captionStartId="10.[151,255,1381,1405]" captionTargetBox="[242,1348,321,1346]" captionTargetId="figure-283@10.[239,1349,318,1352]" captionTargetPageId="10" captionText="FIGURE 31. Mature larva of Symmorphus murarius: cranium, lateral view (detail of setae (a), antennal papilla (b), labrum (c) and mouthparts (d): maxilla (m), maxillary palpus (mp), galea (g), labial palpus (lp), salivary orifice (so)." pageId="8" pageNumber="50">Fig. 31d</figureCitation>
, so) as transverse slit (l =
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) with strongly raised lips.
</paragraph>
</subSubSection>
</treatment>
</document>