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<document id="F7C884E012402AB15BDA28FCCEF18146" ID-DOI="10.5281/zenodo.158742" ID-GBIF-Dataset="21e05249-5427-4500-b740-c128c3442a99" ID-ISSN="1175­5326" ID-Zenodo-Dep="158742" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461026603859" checkinUser="plazi" docAuthor="Freire, Pablo R. &amp; Serejo, Cristiana S." docDate="2004" docId="03A012206C0E807BD033FC449613FD52" docLanguage="en" docName="zt00645.pdf" docOrigin="Zootaxa 645" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Trischizostoma costai Freire &amp; Serejo, 2004, sp. nov." docType="treatment" docVersion="8" lastPageNumber="14" masterDocId="FF996A586C058076D13BFFA89301FF94" masterDocTitle="The genus Trischizostoma (Crustacea: Amphipoda: Trischizostomidae) from the Southwest Atlantic, collected by the REVIZEE Program" masterLastPageNumber="15" masterPageNumber="1" pageNumber="12" updateTime="1698572329815" updateUser="plazi">
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<mods:title id="5EDC31A3CC50DFC314AA84AC8C5750BA">The genus Trischizostoma (Crustacea: Amphipoda: Trischizostomidae) from the Southwest Atlantic, collected by the REVIZEE Program</mods:title>
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<mods:namePart id="37CE105F921AEC3AB6F7E5A1A93D8EBF">Freire, Pablo R.</mods:namePart>
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<mods:namePart id="65EDAFA49D74D56460095F678CC4C8E7">Serejo, Cristiana S.</mods:namePart>
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<mods:date id="0E785D88ACD5D3FDED47EB83F6BD409D">2004</mods:date>
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<mods:identifier id="1EEFC70920F0B7717BECF1525CAB7AA5" type="DOI">10.5281/zenodo.158742</mods:identifier>
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<treatment id="03A012206C0E807BD033FC449613FD52" ID-DOI="http://doi.org/10.5281/zenodo.6271772" ID-GBIF-Taxon="119606488" ID-Zenodo-Dep="6271772" LSID="urn:lsid:plazi:treatment:03A012206C0E807BD033FC449613FD52" httpUri="http://treatment.plazi.org/id/03A012206C0E807BD033FC449613FD52" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12">
<subSubSection id="C313F0BD6C0E807DD033FC4491D3FB92" box="[264,722,1004,1030]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="8BB6A3366C0E807DD033FC4491D3FB92" blockId="11.[264,722,1004,1030]" box="[264,722,1004,1030]" pageId="11" pageNumber="12">
<heading id="D0FE145A6C0E807DD033FC4491D3FB92" bold="true" box="[264,722,1004,1030]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<emphasis id="B97D7F246C0E807DD033FC4491D3FB92" bold="true" box="[264,722,1004,1030]" pageId="11" pageNumber="12">
<taxonomicName id="4C09D8B56C0E807DD033FC44910AFB92" box="[264,523,1004,1030]" class="Malacostraca" family="Lysianassidae" genus="Trischizostoma" kingdom="Animalia" order="Amphipoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="costai" status="sp. nov.">
<emphasis id="B97D7F246C0E807DD033FC44910AFB92" bold="true" box="[264,523,1004,1030]" italics="true" pageId="11" pageNumber="12">Trischizostoma costai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24EC25F6C0E807DD328FC449170FB92" box="[531,625,1004,1030]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="1332BFB36C0E807DD3BAFC4491C8FB92" box="[641,713,1004,1030]" captionStart="FIGURE 6" captionStartId="12.[264,367,1806,1830]" captionTargetBox="[302,1219,730,1805]" captionTargetId="figure@12.[302,1285,706,1807]" captionTargetPageId="12" captionText="FIGURE 6. Trischizostoma costai sp. nov., female, 34 mm, MNRJ 15144. A, habitus; B, gnathopod 1; C, gnathopod 2; D, pereopod 3; E, telson. Scales bars: A, 5 mm; B, 1 mm; C, 2 mm; D E, 1 mm." httpUri="https://zenodo.org/record/158747/files/figure.png" pageId="11" pageNumber="12">Fig. 6</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C313F0BD6C0E807DD033FB9E9163FB35" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="8BB6A3366C0E807DD033FB9E9163FB35" blockId="11.[264,1324,1078,1905]" pageId="11" pageNumber="12">
<emphasis id="B97D7F246C0E807DD033FB9E92FCFBC4" bold="true" box="[264,509,1078,1104]" pageId="11" pageNumber="12">Material examined.</emphasis>
REVIZEE Program, N.
<collectingCountry id="F31EE3A66C0E807DD220FB9F9032FBC5" box="[795,819,1079,1105]" name="Iceland" pageId="11" pageNumber="12">O</collectingCountry>
.
<taxonomicName id="4C09D8B56C0E807DD204FB9F90A8FBC4" box="[831,937,1079,1104]" class="Insecta" family="Coccinellidae" genus="Thalassa" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B97D7F246C0E807DD204FB9F90A8FBC4" box="[831,937,1079,1104]" italics="true" pageId="11" pageNumber="12">Thalassa</emphasis>
</taxonomicName>
col., stn. E­0540,
<geoCoordinate id="EE3DC5F16C0E807DD5BDFB9F9624FBC5" box="[1158,1317,1079,1105]" direction="south" orientation="latitude" pageId="11" pageNumber="12" precision="1" value="-21.204884">21º 12.293' S</geoCoordinate>
,
<geoCoordinate id="EE3DC5F16C0E807DD033FBF792BFFBED" box="[264,446,1119,1145]" direction="west" orientation="longitude" pageId="11" pageNumber="12" precision="1" value="-40.014732">040º 00.884' W</geoCoordinate>
to
<geoCoordinate id="EE3DC5F16C0E807DD0DFFBF79180FBED" box="[484,641,1119,1145]" direction="south" orientation="latitude" pageId="11" pageNumber="12" precision="1" value="-21.1596">21º 09.576' S</geoCoordinate>
,
<geoCoordinate id="EE3DC5F16C0E807DD3B5FBF79044FBED" box="[654,837,1119,1145]" direction="west" orientation="longitude" pageId="11" pageNumber="12" precision="1" value="-40.007683">040º 00.461' W</geoCoordinate>
,
<quantity id="4CF10ED36C0E807DD26AFBF790A5FBED" box="[849,932,1119,1145]" metricMagnitude="3" metricUnit="m" metricValue="1.364" pageId="11" pageNumber="12" unit="m" value="1364.0">1364m</quantity>
,
<date id="FFB785F66C0E807DD28AFBF7973BFBED" box="[945,1082,1119,1145]" pageId="11" pageNumber="12" value="2000-07-04">04/07/2000</date>
;
<specimenCount id="9D0F68BF6C0E807DD57DFBF797ADFBED" box="[1094,1196,1119,1145]" pageId="11" pageNumber="12" type="female">1 female</specimenCount>
with oostegites,
<quantity id="4CF10ED36C0E807DD060FB2F92B1FB35" box="[347,432,1159,1185]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" pageId="11" pageNumber="12" unit="mm" value="34.0">34 mm</quantity>
,
<collectionCode id="ED183BF36C0E807DD087FB2F910BFB35" box="[444,522,1159,1185]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="11" pageNumber="12">MNRJ</collectionCode>
15144.
</paragraph>
</subSubSection>
<subSubSection id="C313F0BD6C0E807DD003FB0690ECFA8D" pageId="11" pageNumber="12" type="etymology">
<paragraph id="8BB6A3366C0E807DD003FB0690ECFA8D" blockId="11.[264,1324,1078,1905]" pageId="11" pageNumber="12">
<emphasis id="B97D7F246C0E807DD003FB0692C5FB5C" bold="true" box="[312,452,1198,1224]" pageId="11" pageNumber="12">Etymology.</emphasis>
The name of this species is dedicated in honor to Dr. Paulo Costa, head of the Brazilian Fisheries REVIZEE program — Central Campaign, who always encourages the research on the material obtained in this scientific program.
</paragraph>
</subSubSection>
<subSubSection id="C313F0BD6C0E807DD003FA8E90B1FA2D" pageId="11" pageNumber="12" type="diagnosis">
<paragraph id="8BB6A3366C0E807DD003FA8E90B1FA2D" blockId="11.[264,1324,1078,1905]" pageId="11" pageNumber="12">
<emphasis id="B97D7F246C0E807DD003FA8E92B6FAD4" bold="true" box="[312,439,1318,1344]" pageId="11" pageNumber="12">Diagnosis.</emphasis>
Maxilliped palp longer than outer plate. Propodus of gnathopod 1 oval and with 11 short and long stout setae on its extension and 3 pairs of long stout setae distally. Coxa 2 with anterior margin rounded. Carpus of gnathopod 2 longer than ischium. Basis of pereopod 3
<date id="FFB785F66C0E807DD0A9FA3792BCFA2D" box="[402,445,1439,1465]" pageId="11" pageNumber="12">3 X</date>
propodus length. Telson entire, triangular.
</paragraph>
</subSubSection>
<subSubSection id="C313F0BD6C0E807BD003FA6E91F0FE3A" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12" type="description">
<paragraph id="8BB6A3366C0E807DD003FA6E92C7F945" blockId="11.[264,1324,1078,1905]" pageId="11" pageNumber="12">
<emphasis id="B97D7F246C0E807DD003FA6E92D1FA74" bold="true" box="[312,464,1478,1504]" pageId="11" pageNumber="12">Description.</emphasis>
Female,
<quantity id="4CF10ED36C0E807DD37CFA6F919FFA75" box="[583,670,1479,1505]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" pageId="11" pageNumber="12" unit="mm" value="34.0">34 mm</quantity>
(
<figureCitation id="1332BFB36C0E807DD38FFA6F91FEFA75" box="[692,767,1479,1505]" captionStart="FIGURE 6" captionStartId="12.[264,367,1806,1830]" captionTargetBox="[302,1219,730,1805]" captionTargetId="figure@12.[302,1285,706,1807]" captionTargetPageId="12" captionText="FIGURE 6. Trischizostoma costai sp. nov., female, 34 mm, MNRJ 15144. A, habitus; B, gnathopod 1; C, gnathopod 2; D, pereopod 3; E, telson. Scales bars: A, 5 mm; B, 1 mm; C, 2 mm; D E, 1 mm." httpUri="https://zenodo.org/record/158747/files/figure.png" pageId="11" pageNumber="12">Fig. 6</figureCitation>
A). Rostrum reaching the end of article 1 of peduncle of antenna 1. Eyes well developed. Antenna 1, article 1 of peduncle longer than articles 23; callynophore present. Antenna 2, article 5 of peduncle as long as article 4. Mandibles with distal incisors; palp 3­articulate with lateral setae. Maxilla 1, outer plate with 8 distal setae. Maxilla 2, inner and outer plates subequal in length; outer plate with 3 lateral setae. Maxilliped, outer plate longer than inner plate; palp with 4 articles, longer than outer plate.
</paragraph>
<paragraph id="8BB6A3366C0E807AD003F9779142FD0A" blockId="11.[264,1324,1078,1905]" lastBlockId="12.[264,1323,284,670]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
Gnathopod 1 (
<figureCitation id="1332BFB36C0E807DD0D2F9779132F96D" box="[489,563,1759,1785]" captionStart="FIGURE 6" captionStartId="12.[264,367,1806,1830]" captionTargetBox="[302,1219,730,1805]" captionTargetId="figure@12.[302,1285,706,1807]" captionTargetPageId="12" captionText="FIGURE 6. Trischizostoma costai sp. nov., female, 34 mm, MNRJ 15144. A, habitus; B, gnathopod 1; C, gnathopod 2; D, pereopod 3; E, telson. Scales bars: A, 5 mm; B, 1 mm; C, 2 mm; D E, 1 mm." httpUri="https://zenodo.org/record/158747/files/figure.png" pageId="11" pageNumber="12">Fig. 6</figureCitation>
B) strongly subchelate; coxa reduced and partially covered by coxa 2; propodus inverted, robust, oval; palm with 11 short and long stout setae in its extension and 3 pairs of long stout setae distally. Gnathopod 2 (
<figureCitation id="1332BFB36C0E807DD2C4F8879747F8DD" box="[1023,1094,1839,1865]" captionStart="FIGURE 6" captionStartId="12.[264,367,1806,1830]" captionTargetBox="[302,1219,730,1805]" captionTargetId="figure@12.[302,1285,706,1807]" captionTargetPageId="12" captionText="FIGURE 6. Trischizostoma costai sp. nov., female, 34 mm, MNRJ 15144. A, habitus; B, gnathopod 1; C, gnathopod 2; D, pereopod 3; E, telson. Scales bars: A, 5 mm; B, 1 mm; C, 2 mm; D E, 1 mm." httpUri="https://zenodo.org/record/158747/files/figure.png" pageId="11" pageNumber="12">Fig. 6</figureCitation>
C), coxa with anterior margin rounded; carpus slightly longer than ischium; merus shorter than ischium; propodus shorter than half of carpus; dactylus reduced. Pereopod 3 (
<figureCitation id="1332BFB36C09807AD51FFEB4976CFEA2" box="[1060,1133,284,310]" captionStart="FIGURE 6" captionStartId="12.[264,367,1806,1830]" captionTargetBox="[302,1219,730,1805]" captionTargetId="figure@12.[302,1285,706,1807]" captionTargetPageId="12" captionText="FIGURE 6. Trischizostoma costai sp. nov., female, 34 mm, MNRJ 15144. A, habitus; B, gnathopod 1; C, gnathopod 2; D, pereopod 3; E, telson. Scales bars: A, 5 mm; B, 1 mm; C, 2 mm; D E, 1 mm." httpUri="https://zenodo.org/record/158747/files/figure.png" pageId="12" pageNumber="13">Fig. 6</figureCitation>
D), coxa longer than broad; basis
<date id="FFB785F66C09807AD0DBFEEC910FFECA" box="[480,526,324,350]" pageId="12" pageNumber="13">3 X</date>
length of propodus; carpus as long as merus, with stout setae on anterior and posterior margins; propodus more than twice length of dactylus, with stout setae on posterior margin; propodus and carpus covered with minute setae on posterior margin. Pereopod 4, coxa posteriorly excavated; basis more than
<date id="FFB785F66C09807AD53FFE14972EFE42" box="[1028,1071,444,470]" pageId="12" pageNumber="13">2 X</date>
length of merus; carpus as long as merus and longer than propodus; dactylus with half length of propodus. Pereopod 5, coxa broader than long; basis with posterior lobe; merus as long as carpus and shorter than propodus; dactylus 1/3 of propodus. Pereopods 6 and 7, coxa broader than long; basis with posterior lobe; merus as long as carpus and longer than propodus; dactylus
<date id="FFB785F66C09807AD033FD2C9232FD0A" box="[264,307,644,670]" pageId="12" pageNumber="13">3 X</date>
shorter than propodus.
</paragraph>
<caption id="DF76F3BE6C09807AD033F8A69238F8FF" httpUri="https://zenodo.org/record/158747/files/figure.png" pageId="12" pageNumber="13" targetBox="[302,1219,730,1805]" targetPageId="12">
<paragraph id="8BB6A3366C09807AD033F8A69238F8FF" blockId="12.[264,1323,1806,1899]" pageId="12" pageNumber="13">
<emphasis id="B97D7F246C09807AD033F8A6928AF8B2" bold="true" box="[264,395,1806,1830]" pageId="12" pageNumber="13">FIGURE 6.</emphasis>
<taxonomicName id="4C09D8B56C09807AD0AFF8A7917FF8B2" box="[404,638,1807,1830]" class="Malacostraca" family="Lysianassidae" genus="Trischizostoma" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="costai" status="sp. nov.">
<emphasis id="B97D7F246C09807AD0AFF8A7917FF8B2" box="[404,638,1807,1830]" italics="true" pageId="12" pageNumber="13">Trischizostoma costai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24EC25F6C09807AD3BEF8A691DDF8B1" box="[645,732,1806,1829]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
, female, 34 mm, MNRJ 15144. A, habitus; B, gnathopod 1; C, gnathopod 2; D, pereopod 3; E, telson. Scales bars: A, 5 mm; B, 1mm; C, 2 mm; DE, 1 mm.
</paragraph>
</caption>
<paragraph id="8BB6A3366C08807BD003FEB491F0FE3A" blockId="13.[264,1323,284,710]" pageId="13" pageNumber="14">
Uropods 1 and 2, peduncle subequal to outer ramus; inner ramus slightly longer than outer ramus. Uropod 3, peduncle slightly less than half of outer ramus; inner and outer rami subequal in length; outer ramus 2­articulate, with basal article
<date id="FFB785F66C08807BD506FEC4976AFE12" box="[1085,1131,364,390]" pageId="13" pageNumber="14">5 X</date>
length of distal article. Telson entire, triangular (
<figureCitation id="1332BFB36C08807BD3B3FE3C91CEFE3A" box="[648,719,404,430]" captionStart="FIGURE 6" captionStartId="12.[264,367,1806,1830]" captionTargetBox="[302,1219,730,1805]" captionTargetId="figure@12.[302,1285,706,1807]" captionTargetPageId="12" captionText="FIGURE 6. Trischizostoma costai sp. nov., female, 34 mm, MNRJ 15144. A, habitus; B, gnathopod 1; C, gnathopod 2; D, pereopod 3; E, telson. Scales bars: A, 5 mm; B, 1 mm; C, 2 mm; D E, 1 mm." httpUri="https://zenodo.org/record/158747/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
E).
</paragraph>
</subSubSection>
<subSubSection id="C313F0BD6C08807BD003FE139613FD52" pageId="13" pageNumber="14" type="discussion">
<paragraph id="8BB6A3366C08807BD003FE139613FD52" blockId="13.[264,1323,284,710]" pageId="13" pageNumber="14">
<emphasis id="B97D7F246C08807BD003FE1392B0FE41" bold="true" box="[312,433,443,469]" pageId="13" pageNumber="14">Remarks.</emphasis>
<taxonomicName id="4C09D8B56C08807BD08CFE1491B7FE41" box="[439,694,444,469]" class="Malacostraca" family="Lysianassidae" genus="Trischizostoma" kingdom="Animalia" order="Amphipoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="costai" status="sp. nov.">
<emphasis id="B97D7F246C08807BD08CFE1491B7FE41" box="[439,694,444,469]" italics="true" pageId="13" pageNumber="14">Trischizostoma costai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24EC25F6C08807BD387FE149017FE42" box="[700,790,444,470]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
is similar to
<taxonomicName id="4C09D8B56C08807BD297FE149719FE41" box="[940,1048,444,469]" class="Malacostraca" family="Lysianassidae" genus="Trischizostoma" kingdom="Animalia" order="Amphipoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="richeri">
<emphasis id="B97D7F246C08807BD297FE149719FE41" box="[940,1048,444,469]" italics="true" pageId="13" pageNumber="14">T. richeri</emphasis>
</taxonomicName>
in the shape of the propodus of gnathopod 1, and shape of coxa 2. However, there is a conspicuous distinction in the size of the eyes, much larger in
<taxonomicName id="4C09D8B56C08807BD39CFDA49015FDB1" box="[679,788,524,549]" class="Malacostraca" family="Lysianassidae" genus="Trischizostoma" kingdom="Animalia" order="Amphipoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="richeri">
<emphasis id="B97D7F246C08807BD39CFDA49015FDB1" box="[679,788,524,549]" italics="true" pageId="13" pageNumber="14">T. richeri</emphasis>
</taxonomicName>
, and in the number of stout setae on the palm of gnathopod 1, with 11 setae on
<taxonomicName id="4C09D8B56C08807BD3B0FD9C91ECFDD9" box="[651,749,564,589]" class="Malacostraca" family="Lysianassidae" genus="Trischizostoma" kingdom="Animalia" order="Amphipoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="costai" status="sp. nov.">
<emphasis id="B97D7F246C08807BD3B0FD9C91ECFDD9" box="[651,749,564,589]" italics="true" pageId="13" pageNumber="14">T. costai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24EC25F6C08807BD3CFFD9C9050FDDA" box="[756,849,564,590]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
, and 19 setae on
<taxonomicName id="4C09D8B56C08807BD52CFD9C9782FDD9" box="[1047,1155,564,589]" class="Malacostraca" family="Lysianassidae" genus="Trischizostoma" kingdom="Animalia" order="Amphipoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="richeri">
<emphasis id="B97D7F246C08807BD52CFD9C9782FDD9" box="[1047,1155,564,589]" italics="true" pageId="13" pageNumber="14">T. richeri</emphasis>
</taxonomicName>
. According to observations on juveniles of
<taxonomicName id="4C09D8B56C08807BD362FDF491BFFDE1" box="[601,702,604,629]" class="Malacostraca" family="Lysianassidae" genus="Trischizostoma" kingdom="Animalia" order="Amphipoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="raschi">
<emphasis id="B97D7F246C08807BD362FDF491BFFDE1" box="[601,702,604,629]" italics="true" pageId="13" pageNumber="14">T. raschi</emphasis>
</taxonomicName>
, it seems that all the setae on the palm appears at the same time and not gradually by each molt. Also,
<taxonomicName id="4C09D8B56C08807BD252FD2C90CCFD09" box="[873,973,644,669]" class="Malacostraca" family="Lysianassidae" genus="Trischizostoma" kingdom="Animalia" order="Amphipoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="costai" status="sp. nov.">
<emphasis id="B97D7F246C08807BD252FD2C90CCFD09" box="[873,973,644,669]" italics="true" pageId="13" pageNumber="14">T. costai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24EC25F6C08807BD2E1FD2C9738FD0A" box="[986,1081,644,670]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
presents oostegites, indicating that this specimen is a mature female and not a possible juvenile of
<taxonomicName id="4C09D8B56C08807BD5A5FD04960AFD51" box="[1182,1291,684,709]" class="Malacostraca" family="Lysianassidae" genus="Trischizostoma" kingdom="Animalia" order="Amphipoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="richeri">
<emphasis id="B97D7F246C08807BD5A5FD04960AFD51" box="[1182,1291,684,709]" italics="true" pageId="13" pageNumber="14">T. richeri</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>