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<document id="43F83EEE7001B2D968997E76348B80D6" ID-DOI="10.11646/zootaxa.4763.2.5" ID-GBIF-Dataset="85439034-f26e-40e2-ba2a-f5654eb352f1" ID-ISSN="1175-5334" ID-Zenodo-Dep="3758250" ID-ZooBank="urn:lsid:zoobank.org:pub:D179A5F1-C594-4B42-938E-3F8A2734BA84" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587398341573" checkinUser="carolina" docAuthor="Tan, Ming Kai, Baroga-Barbecho, Jessica B., Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul &amp; Yap, Sheryl A." docDate="2020" docId="039F4D5AB965FFB74BDEF8D55A0F0D9C" docLanguage="en" docName="zootaxa.4763.2.5.pdf.imf" docOrigin="Zootaxa 4763 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhicnogryllus bipunctatus Ingrisch 1987" docType="treatment" docVersion="4" lastPageNumber="224" masterDocId="FFA63522B960FFB04B49FFD25F330803" masterDocTitle="A taxonomic study on the Bornean and Philippines Sword-tailed Crickets in the genus Rhicnogryllus Chopard, 1925 (Orthoptera: Trgonidiidae; Trigonidiinae)" masterLastPageNumber="230" masterPageNumber="217" pageNumber="222" updateTime="1698739035565" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="AB3EE4EB424C6F76A209B25572B6DBF2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="769CE893C554EC2E41FB8F8B3750FD19">
<mods:title id="CA58A08F7C4C07D2315B0163F694F593">A taxonomic study on the Bornean and Philippines Sword-tailed Crickets in the genus Rhicnogryllus Chopard, 1925 (Orthoptera: Trgonidiidae; Trigonidiinae)</mods:title>
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<mods:namePart id="723996B071096226A3FC35C81D219282">Tan, Ming Kai</mods:namePart>
<mods:affiliation id="6A01B968B0FFDB544025F4C22370823A">Institut de Systématique, Evolution et Biodiversité (ISYEB), Muséum national d’Histoire naturelle</mods:affiliation>
<mods:nameIdentifier id="50BE1F5264638B01E69BA7754E944ED4" type="email">orthoptera.mingkai@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="FA8BB161F597053E63CF3B6EC182767A" type="personal">
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</mods:role>
<mods:namePart id="4B9D538E9BD3C0C6DC88B1804454696C">Baroga-Barbecho, Jessica B.</mods:namePart>
<mods:affiliation id="77EDCE2CCFA87FF7D889A0BF30872FF8">Office of the Vice Chancellor for Research and Extension, University of the Philippines Los Baños, College</mods:affiliation>
</mods:name>
<mods:name id="6B4EDFD01C70F529FF92C0EE571E11A0" type="personal">
<mods:role id="DE84634C4A24EF6045CA2CC611CAAF68">
<mods:roleTerm id="10D874214AECC4DC04747AEB12225A76">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6159ECFEFF29B5F4A34A175E9EF20E6A">Japir, Razy</mods:namePart>
<mods:affiliation id="3A488C865E5DB1D1895F7D49CECFA1F7">Forest Research Centre (Sepilok), Sabah Forestry Department</mods:affiliation>
</mods:name>
<mods:name id="0AC733872FA7636D7950A34AAB4ED850" type="personal">
<mods:role id="B4273FD767B5177379B940B3875E0C13">
<mods:roleTerm id="61725F865558BDE9D0292E75DA8DC2D3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="39075821012004B9620C0AA5A24C9667">Chung, Arthur Y. C.</mods:namePart>
<mods:affiliation id="B961261E87B54A8C044771CBA1728791">Forest Research Centre (Sepilok), Sabah Forestry Department</mods:affiliation>
</mods:name>
<mods:name id="DB6B88CFC1C9F38FDF85956FD58BFBB6" type="personal">
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<mods:namePart id="C99137D5205DC7DD16DF9580CF6164AD">Wahab, Rodzay Bin Haji Abdul</mods:namePart>
<mods:affiliation id="263EAD11848176E5680F9101CB1ABE23">Institute for Biodiversity and Environmental Research, Universiti Brunei Darussalam, Jalan Universiti</mods:affiliation>
</mods:name>
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<mods:namePart id="6A97C01B161E62C400D5C1473B33AC76">Yap, Sheryl A.</mods:namePart>
<mods:affiliation id="1C5FF272C3E3743E85A38F3DF9F23CA8">Institute of Weed Science, Entomology, and Plant Pathology, College of Agriculture and Food Science, University of the Philippines Los Baños, College. Museum of Natural History, University of the Philippines Los Baños, College.</mods:affiliation>
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<mods:typeOfResource id="50D48C9A828850959F84DB9BF1FBBBBF">text</mods:typeOfResource>
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<mods:title id="29C4DCA580702A255C32F18B1131C069">Zootaxa</mods:title>
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<mods:part id="C81AD229C6E680C28875A933D93C4290">
<mods:date id="6EAB726BC4037D4C6D6D38864B2A6F49">2020</mods:date>
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<mods:number id="9D25509BC32D2EF35F35A66E50A39B75">2020-04-09</mods:number>
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<mods:number id="85812EA6135AD384D50B6B52281A1DF5">4763</mods:number>
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<mods:number id="A458279A44F6FE0BC8DA52CF3E5FD182">2</mods:number>
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<mods:start id="7798337245D75184B1F9F420CBBECFB1">217</mods:start>
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<mods:classification id="7066DEB35E8AC40FC25140AE0F549A4C">journal article</mods:classification>
<mods:identifier id="8B50B5026CD71950A6479F55A8CF9A35" type="DOI">10.11646/zootaxa.4763.2.5</mods:identifier>
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<treatment id="039F4D5AB965FFB74BDEF8D55A0F0D9C" ID-GBIF-Taxon="163652845" LSID="urn:lsid:plazi:treatment:039F4D5AB965FFB74BDEF8D55A0F0D9C" httpUri="http://treatment.plazi.org/id/039F4D5AB965FFB74BDEF8D55A0F0D9C" lastPageId="7" lastPageNumber="224" pageId="5" pageNumber="222">
<subSubSection id="C32CAFC7B965FFB54BDEF8D55DA20F21" box="[151,657,1799,1826]" pageId="5" pageNumber="222" type="nomenclature">
<paragraph id="8B89FC4CB965FFB54BDEF8D55DA20F21" blockId="5.[151,657,1799,1862]" box="[151,657,1799,1826]" pageId="5" pageNumber="222">
<heading id="D0C14B20B965FFB54BDEF8D55DA20F21" bold="true" box="[151,657,1799,1826]" fontSize="10" level="2" pageId="5" pageNumber="222" reason="2">
<taxonomicName id="4C3687CFB965FFB54BDEF8D55DA20F21" authority="Ingrisch, 1987" authorityName="Ingrisch" authorityYear="1987" box="[151,657,1799,1826]" class="Insecta" family="Haglotettigoniidae" genus="Rhicnogryllus" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="222" phylum="Arthropoda" rank="species" species="bipunctatus">
<emphasis id="B942205EB965FFB54BDEF8D55DA20F21" bold="true" box="[151,657,1799,1826]" pageId="5" pageNumber="222">
<emphasis id="B942205EB965FFB54BDEF8D55EEB0F21" bold="true" box="[151,472,1799,1826]" italics="true" pageId="5" pageNumber="222">Rhicnogryllus bipunctatus</emphasis>
<bibRefCitation id="EFA781BDB965FFB54A96F8DA5DA20F21" author="Ingrisch" box="[479,657,1800,1826]" firstAuthor="Ingrisch" pageId="5" pageNumber="222" pagination="163 - 185" refId="ref6342" refString="Ingrisch, S. (1987) Neue Grillen von Borneo und aus Thailand (Insecta: Saltatoria: Grylloidea). Senckenbergiana biologica, 68 (1 - 3), 163 - 185." type="journal article" year="1987">Ingrisch, 1987</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32CAFC7B965FFB54BDEF8FE5FCC0F46" box="[151,255,1835,1862]" pageId="5" pageNumber="222" type="description">
<paragraph id="8B89FC4CB965FFB54BDEF8FE5FCC0F46" blockId="5.[151,657,1799,1862]" box="[151,255,1835,1862]" pageId="5" pageNumber="222">
<figureCitation id="130DE0C9B965FFB54BDEF8FE5FD70F45" box="[151,228,1836,1862]" captionStart="FIGURE 3" captionStartId="6.[151,250,1980,2005]" captionTargetBox="[206,1380,381,1955]" captionTargetId="figure@6.[206,1380,381,1955]" captionTargetPageId="6" captionText="FIGURE 3. Live habitus of Rhicnogryllus bipunctatus female from Sandakan, Sabah (locality of R. eximius sensu Chopard) (A) and male from Brunei Darussalam (B)." figureDoi="http://doi.org/10.5281/zenodo.3758256" httpUri="https://zenodo.org/record/3758256/files/figure.png" pageId="5" pageNumber="222">Figs. 3</figureCitation>
,
<figureCitation id="130DE0C9B965FFB54BB8F8F95FCC0F46" box="[241,255,1835,1861]" captionStart="FIGURE 6" captionStartId="10.[151,250,1827,1852]" captionTargetBox="[215,1372,181,1801]" captionTargetId="figure@10.[215,1372,181,1801]" captionTargetPageId="10" captionText="FIGURE 6. Male genitalia of Rhicnogryllus bipunctatus (A–C) and Rhicnogryllus paetensis n. sp. (D–F) in dorsal (A, D), ventral (B, E) and lateral (C, F) views." figureDoi="http://doi.org/10.5281/zenodo.3758262" httpUri="https://zenodo.org/record/3758262/files/figure.png" pageId="5" pageNumber="222">6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32CAFC7B965FFB54BDEF8A25D070FA4" pageId="5" pageNumber="222" type="reference_group">
<paragraph id="8B89FC4CB965FFB54BDEF8A25D070FA4" blockId="5.[151,1323,1903,1959]" pageId="5" pageNumber="222">
<treatmentCitationGroup id="AB26DB62B965FFB54BDEF8A25D070FA4" pageId="5" pageNumber="222">
<taxonomicName id="4C3687CFB965FFB54BDEF8A25E910F8B" authorityName="Ingrisch" authorityYear="1987" box="[151,418,1904,1928]" class="Insecta" family="Haglotettigoniidae" genus="Rhicnogryllus" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="222" phylum="Arthropoda" rank="species" species="bipunctatus">
<emphasis id="B942205EB965FFB54BDEF8A25E910F8B" box="[151,418,1904,1928]" italics="true" pageId="5" pageNumber="222">Rhicnogryllus bipunctatus</emphasis>
</taxonomicName>
—
<treatmentCitation id="0A97DA5DB965FFB54AF5F8BD5DB20F8B" author="Ingrisch, S." box="[444,641,1903,1928]" page="176" pageId="5" pageNumber="222" year="1987">
<bibRefCitation id="EFA781BDB965FFB54AF5F8BD5DB20F8B" author="Ingrisch" box="[444,641,1903,1928]" firstAuthor="Ingrisch" pageId="5" pageNumber="222" pagination="163 - 185" refId="ref6342" refString="Ingrisch, S. (1987) Neue Grillen von Borneo und aus Thailand (Insecta: Saltatoria: Grylloidea). Senckenbergiana biologica, 68 (1 - 3), 163 - 185." type="journal article" year="1987">Ingrisch, 1987: 176</bibRefCitation>
</treatmentCitation>
&gt; original description;
<treatmentCitation id="0A97DA5DB965FFB5483CF8A25B630F8B" author="Tan, M. K. &amp; Japir, R. &amp; Chung, A. Y. C." box="[885,1104,1903,1928]" httpUri="http://treatment.plazi.org/id/039F8783FFE9AD2510EF6409FC7F6C49" page="306" pageId="5" pageNumber="222" year="2019">
<bibRefCitation id="EFA781BDB965FFB5483CF8A25B630F8B" author="Tan" box="[885,1104,1903,1928]" etAl="et al." firstAuthor="Tan" pageId="5" pageNumber="222" pagination="301 - 349" refId="ref7151" refString="Tan, M. K., Japir, R. &amp; Chung, A. Y. C. (2019 c) Uncovering the Grylloidea and Tettigonioidea (Orthoptera: Ensifera) in the Forest Research Centre (Sepilok) Entomological Collection. Zootaxa, 4701 (4), 301 - 349. https: // doi. org / 10.11646 / zootaxa. 4701.4.1" type="journal article" year="2019" yearSuffix="c">Tan et al., 2019c: 306</bibRefCitation>
</treatmentCitation>
(record in Sandakan)
<taxonomicName id="4C3687CFB965FFB54BDEF85D5E4A0FA5" box="[151,377,1935,1959]" class="Insecta" family="Haglotettigoniidae" genus="Rhicnogryllus" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="222" phylum="Arthropoda" rank="species" species="eximius">
<emphasis id="B942205EB965FFB54BDEF85D5E4A0FA5" box="[151,377,1935,1959]" italics="true" pageId="5" pageNumber="222">Rhicnogryllus eximius</emphasis>
</taxonomicName>
—Chopard, in litt.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C32CAFC7B965FFB74BDEF8065C8D0940" lastPageId="7" lastPageNumber="224" pageId="5" pageNumber="222" type="materials_examined">
<paragraph id="8B89FC4CB965FFB54BDEF8065AAF0FEC" blockId="5.[151,1436,2004,2032]" box="[151,1436,2004,2032]" pageId="5" pageNumber="222">
<emphasis id="B942205EB965FFB54BDEF8065EB70FEC" bold="true" box="[151,388,2004,2031]" pageId="5" pageNumber="222">Material examined.</emphasis>
<materialsCitation id="3B5EF611B965FFB54AC6F8075AAF0FEC" ID-GBIF-Occurrence="2599922924" box="[399,1436,2004,2032]" collectionCode="SMFD" collectorName="S. Ingrisch" country="Malaysia" location="August" municipality="Batu Niah" pageId="5" pageNumber="222" specimenCount="1" specimenCount-male="1" stateProvince="Sarawak" typeStatus="holotype">
<specimenCount id="9D3037C5B965FFB54AC6F8075EFA0FEC" box="[399,457,2005,2031]" pageId="5" pageNumber="222" type="male">Male</specimenCount>
<typeStatus id="548D42EEB965FFB54A9AF8075D060FEC" box="[467,565,2005,2031]" pageId="5" pageNumber="222" type="holotype">holotype</typeStatus>
(only images examined),
<emphasis id="B942205EB965FFB5482EF8075B200FEC" bold="true" box="[871,1043,2004,2031]" pageId="5" pageNumber="222">
East
<collectingCountry id="F321BCDCB965FFB548EFF8065B200FEC" box="[934,1043,2004,2031]" name="Malaysia" pageId="5" pageNumber="222">Malaysia</collectingCountry>
</emphasis>
,
<collectingRegion id="49F232AEB965FFB54F6CF8075BBB0FEC" box="[1061,1160,2005,2031]" country="Malaysia" name="Sarawak" pageId="5" pageNumber="222">Sarawak</collectingRegion>
,
<collectingMunicipality id="6BED6636B965FFB54FDEF8075A3C0FEC" box="[1175,1295,2005,2031]" pageId="5" pageNumber="222">Batu Niah</collectingMunicipality>
, 3/4
<location id="8EE9AA97B965FFB54E02F8075AAF0FEC" LSID="urn:lsid:plazi:treatment:039F4D5AB965FFB74BDEF8D55A0F0D9C:8EE9AA97B965FFB54E02F8075AAF0FEC" box="[1355,1436,2005,2031]" country="Malaysia" municipality="Batu Niah" name="August" pageId="5" pageNumber="222" stateProvince="Sarawak">August</location>
</materialsCitation>
</paragraph>
<paragraph id="8B89FC4CB966FFB64BDEFF4B5B4708D4" blockId="6.[151,1437,152,359]" pageId="6" pageNumber="223">
1984, coll.
<collectorName id="26C3999AB966FFB64A5FFF4B5EA308B0" box="[278,400,153,179]" pageId="6" pageNumber="223">S. Ingrisch</collectorName>
(
<collectionCode id="ED276489B966FFB64AD6FF4B5EC208B0" box="[415,497,153,179]" collectionName="Germany, Frankfurt-am-Main, Forschungsinstitut und Naturmuseum Senckenberg" pageId="6" pageNumber="223">SMFD</collectionCode>
).
<typeStatus id="548D42EEB966FFB6494CFF4B5D5408B0" box="[517,615,153,179]" pageId="6" pageNumber="223" type="paratype">Paratype</typeStatus>
of
<taxonomicName id="4C3687CFB966FFB649C4FF4B5CB408B0" box="[653,903,153,179]" class="Insecta" family="Haglotettigoniidae" genus="Rhicnogryllus" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="223" phylum="Arthropoda" rank="species" species="eximius">
<emphasis id="B942205EB966FFB649C4FF4B5CB408B0" box="[653,903,153,179]" italics="true" pageId="6" pageNumber="223">Rhicnogryllus eximius</emphasis>
</taxonomicName>
(
<specimenCount id="9D3037C5B966FFB648D0FF4B5CD508B0" box="[921,998,153,179]" pageId="6" pageNumber="223" type="male">1 male</specimenCount>
, MNHN-EO-ENSIF6583, only images examined),
<emphasis id="B942205EB966FFB64A52FF6F5EF708D4" bold="true" box="[283,452,188,215]" pageId="6" pageNumber="223">
East
<collectingCountry id="F321BCDCB966FFB64A1EFF6E5EF708D4" box="[343,452,188,215]" name="Malaysia" pageId="6" pageNumber="223">Malaysia</collectingCountry>
</emphasis>
,
<collectingRegion id="49F232AEB966FFB64A9BFF6F5D2508D4" box="[466,534,189,215]" country="Malaysia" name="Sabah" pageId="6" pageNumber="223">Sabah</collectingRegion>
(“North Borneo”), Sandakan,
<date id="FF88DA8CB966FFB64824FF6F5CCE08D4" box="[877,1021,189,215]" pageId="6" pageNumber="223" value="1927-07-24">24 July 1927</date>
(
<collectionCode id="ED276489B966FFB64F45FF6F5B5B08D4" box="[1036,1128,189,215]" country="France" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="223">MNHN</collectionCode>
).
</paragraph>
<paragraph id="8B89FC4CB966FFB64B8EFF335D060964" blockId="6.[151,1437,152,359]" pageId="6" pageNumber="223">
<specimenCount id="9D3037C5B966FFB64B8EFF335E1408F8" box="[199,295,225,251]" pageId="6" pageNumber="223" type="female">1 female</specimenCount>
(BRU.19.61),
<emphasis id="B942205EB966FFB64A85FF325D8708F8" bold="true" box="[460,692,224,251]" pageId="6" pageNumber="223">
<collectingCountry id="F321BCDCB966FFB64A85FF325D8708F8" box="[460,692,224,251]" name="Brunei" pageId="6" pageNumber="223">Brunei Darussalam</collectingCountry>
</emphasis>
,
<collectingRegion id="49F232AEB966FFB64988FF335C5208F8" box="[705,865,225,251]" country="Brunei" name="Belait" pageId="6" pageNumber="223">Belait District</collectingRegion>
, Teraja,
<geoCoordinate id="EE029A8BB966FFB648F4FF335B1C08F8" box="[957,1071,225,251]" degrees="4.28487" direction="north" orientation="latitude" pageId="6" pageNumber="223" precision="1" value="4.28487">N4.28487</geoCoordinate>
,
<geoCoordinate id="EE029A8BB966FFB64F70FF335BF008F8" box="[1081,1219,225,251]" degrees="114.41817" direction="east" orientation="longitude" pageId="6" pageNumber="223" precision="1" value="114.41817">E114.41817</geoCoordinate>
,
<quantity id="4CCE51A9B966FFB64F84FF335A5208F8" box="[1229,1377,225,251]" metricMagnitude="1" metricUnit="m" metricValue="5.18" metricValueMax="7.49" metricValueMin="2.87" pageId="6" pageNumber="223" unit="m" value="51.8" valueMax="74.9" valueMin="28.7">28.7±74.9 m</quantity>
.a.s.l., 1906 hours,
<date id="FF88DA8CB966FFB64A69FED75E8D091C" box="[288,446,261,287]" pageId="6" pageNumber="223" value="2019-03-02">2 March 2019</date>
, coll. M. K. Tan &amp; H. Yeo;
<specimenCount id="9D3037C5B966FFB649A0FED75C00091C" box="[745,819,261,287]" pageId="6" pageNumber="223" type="male">1 male</specimenCount>
(BRU.19.68) and
<specimenCount id="9D3037C5B966FFB648B6FED75B6C091C" box="[1023,1119,261,287]" pageId="6" pageNumber="223" type="female">1 female</specimenCount>
(BRU.19.69), same locality, Wasai Wong Kadir Recreational Park,
<geoCoordinate id="EE029A8BB966FFB64907FEFB5D810940" box="[590,690,297,323]" degrees="4.3421" direction="north" orientation="latitude" pageId="6" pageNumber="223" precision="5" value="4.3421">N4.3421</geoCoordinate>
,
<geoCoordinate id="EE029A8BB966FFB649F7FEFA5C7A0940" box="[702,841,296,323]" degrees="114.44655" direction="east" orientation="longitude" pageId="6" pageNumber="223" precision="1" value="114.44655">E114.44655</geoCoordinate>
, 23.0±
<quantity id="4CCE51A9B966FFB648DCFEFB5CEE0940" box="[917,989,297,323]" metricMagnitude="0" metricUnit="m" metricValue="6.6" pageId="6" pageNumber="223" unit="m" value="6.6">6.6 m</quantity>
.a.s.l., 1639 hours,
<date id="FF88DA8CB966FFB64FE7FEFB5A040940" box="[1198,1335,297,323]" pageId="6" pageNumber="223" value="2019-07-07">7 July 2019</date>
, coll. M. K. Tan &amp; H. Yeo (
<collectionCode id="ED276489B966FFB64A2BFE9F5EA90964" box="[354,410,333,359]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="6" pageNumber="223">ZRC</collectionCode>
and
<collectionCode id="ED276489B966FFB64A98FE9F5D190964" box="[465,554,333,359]" pageId="6" pageNumber="223">UBDM</collectionCode>
).
</paragraph>
<caption id="DF49ACC4B966FFB64BDEF86E5D100FFA" ID-DOI="http://doi.org/10.5281/zenodo.3758256" ID-Zenodo-Dep="3758256" httpUri="https://zenodo.org/record/3758256/files/figure.png" pageId="6" pageNumber="223" startId="6.[151,250,1980,2005]" targetBox="[206,1380,381,1955]" targetPageId="6">
<paragraph id="8B89FC4CB966FFB64BDEF86E5D100FFA" blockId="6.[151,1436,1980,2041]" pageId="6" pageNumber="223">
<emphasis id="B942205EB966FFB64BDEF86E5E210FD6" bold="true" box="[151,274,1980,2005]" pageId="6" pageNumber="223">FIGURE 3.</emphasis>
Live habitus of
<taxonomicName id="4C3687CFB966FFB64AFDF86F5D8E0FD6" authorityName="Ingrisch" authorityYear="1987" box="[436,701,1981,2005]" class="Insecta" family="Haglotettigoniidae" genus="Rhicnogryllus" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="223" phylum="Arthropoda" rank="species" species="bipunctatus">
<emphasis id="B942205EB966FFB64AFDF86F5D8E0FD6" box="[436,701,1981,2005]" italics="true" pageId="6" pageNumber="223">Rhicnogryllus bipunctatus</emphasis>
</taxonomicName>
female from Sandakan, Sabah (locality of
<taxonomicName id="4C3687CFB966FFB64F20F86F5A5E0FD6" authority="sensu Chopard" authorityName="Chopard" box="[1129,1389,1980,2005]" class="Insecta" family="Haglotettigoniidae" genus="Rhicnogryllus" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="223" phylum="Arthropoda" rank="species" species="eximius">
<emphasis id="B942205EB966FFB64F20F86F5BE20FD7" box="[1129,1233,1981,2004]" italics="true" pageId="6" pageNumber="223">R. eximius</emphasis>
sensu Chopard
</taxonomicName>
) (A) and male from Brunei Darussalam (B).
</paragraph>
</caption>
<paragraph id="8B89FC4CB967FFB74B8EFF4B5BF6091C" blockId="7.[151,1437,152,1439]" pageId="7" pageNumber="224">
<specimenCount id="9D3037C5B967FFB74B8EFF4B5E1208B0" box="[199,289,153,179]" pageId="7" pageNumber="224" type="male">4 males</specimenCount>
(SDK.19.42–45),
<emphasis id="B942205EB967FFB74AB0FF4B5D9508B0" bold="true" box="[505,678,152,179]" pageId="7" pageNumber="224">
East
<collectingCountry id="F321BCDCB967FFB74970FF4A5D9508B0" box="[569,678,152,179]" name="Malaysia" pageId="7" pageNumber="224">Malaysia</collectingCountry>
</emphasis>
,
<collectingRegion id="49F232AEB967FFB749F1FF4B5DCC08B0" box="[696,767,153,179]" country="Malaysia" name="Sabah" pageId="7" pageNumber="224">Sabah</collectingRegion>
, Sandakan, Ulu Dusun,
<geoCoordinate id="EE029A8BB967FFB74F54FF4A5BBE08B0" box="[1053,1165,152,179]" degrees="5.78364" direction="north" orientation="latitude" pageId="7" pageNumber="224" precision="1" value="5.78364">N5.78364</geoCoordinate>
<geoCoordinate id="EE029A8BB967FFB74FC7FF4A5BCB08B0" box="[1166,1272,152,179]" degrees="5.78379" direction="west" orientation="longitude" pageId="7" pageNumber="224" precision="1" value="-5.78379">–5.78379</geoCoordinate>
,
<geoCoordinate id="EE029A8BB967FFB74E4EFF4A5ABE08B0" box="[1287,1421,152,179]" degrees="117.76051" direction="east" orientation="longitude" pageId="7" pageNumber="224" precision="1" value="117.76051">E117.76051</geoCoordinate>
– 117.76094, 38.3–41.6±
<quantity id="4CCE51A9B967FFB74AD5FF6E5D2608D4" box="[412,533,188,215]" metricMagnitude="0" metricUnit="m" metricValue="5.9" metricValueMax="6.7" metricValueMin="5.1" pageId="7" pageNumber="224" unit="m" value="5.9" valueMax="6.7" valueMin="5.1">5.1–6.7 m</quantity>
.a.s.l.,
<date id="FF88DA8CB967FFB74910FF6F5DE508D4" box="[601,726,189,215]" pageId="7" pageNumber="224" value="1008" valueMax="1038">1008–1038</date>
hours,
<date id="FF88DA8CB967FFB74864FF6F5CDF08D4" box="[813,1004,189,215]" pageId="7" pageNumber="224" value="2019-01-10">10 January 2019</date>
, coll. M. K. Tan, R. Japir, M. Binti &amp; J. L. Yukang;
<specimenCount id="9D3037C5B967FFB74A7DFF335EA608F8" box="[308,405,225,251]" pageId="7" pageNumber="224" type="female">1 female</specimenCount>
(SDK.19.67), same locality, Sepilok, Rainforest Discovery Centre,
<geoCoordinate id="EE029A8BB967FFB74FD9FF325A3108F8" box="[1168,1282,224,251]" degrees="5.87575" direction="north" orientation="latitude" pageId="7" pageNumber="224" precision="1" value="5.87575">N5.87575</geoCoordinate>
,
<geoCoordinate id="EE029A8BB967FFB74E47FF325AAB08F8" box="[1294,1432,224,251]" degrees="117.94059" direction="east" orientation="longitude" pageId="7" pageNumber="224" precision="1" value="117.94059">E117.94059</geoCoordinate>
, 54.5±
<quantity id="4CCE51A9B967FFB74B91FED65E2C091C" box="[216,287,260,287]" metricMagnitude="0" metricUnit="m" metricValue="6.3" pageId="7" pageNumber="224" unit="m" value="6.3">6.3 m</quantity>
.a.s.l., 0946 hours,
<date id="FF88DA8CB967FFB74AA5FED75DAE091C" box="[492,669,261,287]" pageId="7" pageNumber="224" value="2019-10-01">1 October 2019</date>
, coll. M. K. Tan &amp; J. L. Yukang (
<collectionCode id="ED276489B967FFB74F58FED75B7A091C" box="[1041,1097,261,287]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="224">ZRC</collectionCode>
and
<collectionCode id="ED276489B967FFB74F36FED75B8A091C" box="[1151,1209,261,287]" country="USA" httpUri="http://grbio.org/cool/z9c1-g2b0" name="Fusarium Research Center" pageId="7" pageNumber="224">FRC</collectionCode>
).
</paragraph>
<paragraph id="8B89FC4CB967FFB74B8EFEFB5C8D0940" blockId="7.[151,1437,152,1439]" box="[199,958,296,323]" pageId="7" pageNumber="224">
<emphasis id="B942205EB967FFB74B8EFEFB5E6D0940" bold="true" box="[199,350,296,323]" pageId="7" pageNumber="224">Type details.</emphasis>
Male
<typeStatus id="548D42EEB967FFB74AECFEFB5D390940" box="[421,522,297,323]" pageId="7" pageNumber="224" type="holotype">holotype</typeStatus>
, Borneo:
<collectingCountry id="F321BCDCB967FFB7493FFEFB5DE80940" box="[630,731,297,323]" name="Malaysia" pageId="7" pageNumber="224">Sarawak</collectingCountry>
: Batu Niah (
<collectionCode id="ED276489B967FFB7482EFEFB5C850940" box="[871,950,297,323]" pageId="7" pageNumber="224">SMFD</collectionCode>
)
</paragraph>
</subSubSection>
<subSubSection id="C32CAFC7B967FFB74B8EFE9F5AA40A3C" pageId="7" pageNumber="224" type="discussion">
<paragraph id="8B89FC4CB967FFB74B8EFE9F5E730A18" blockId="7.[151,1437,152,1439]" pageId="7" pageNumber="224">
<emphasis id="B942205EB967FFB74B8EFE9F5E080964" bold="true" box="[199,315,333,359]" pageId="7" pageNumber="224">Remarks.</emphasis>
A male “
<typeStatus id="548D42EEB967FFB74AE9FE9F5D350964" box="[416,518,333,359]" pageId="7" pageNumber="224" type="paratype">paratype</typeStatus>
” of
<taxonomicName id="4C3687CFB967FFB7497AFE9F5CA20964" authority="Chopard" authorityName="Chopard" box="[563,913,333,359]" class="Insecta" family="Haglotettigoniidae" genus="Rhicnogryllus" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="eximius">
<emphasis id="B942205EB967FFB7497AFE9F5C1F0964" box="[563,812,333,359]" italics="true" pageId="7" pageNumber="224">Rhicnogryllus eximius</emphasis>
Chopard
</taxonomicName>
collected from Sandakan was found in MNHN and it resembles
<taxonomicName id="4C3687CFB967FFB74A14FEA05D310988" authorityName="Ingrisch" authorityYear="1987" box="[349,514,369,395]" class="Insecta" family="Haglotettigoniidae" genus="Rhicnogryllus" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="bipunctatus">
<emphasis id="B942205EB967FFB74A14FEA05D310988" box="[349,514,369,395]" italics="true" pageId="7" pageNumber="224">R. bipunctatus</emphasis>
</taxonomicName>
as described by
<bibRefCitation id="EFA781BDB967FFB7498EFEA35C4B0988" author="Ingrisch" box="[711,888,369,395]" firstAuthor="Ingrisch" pageId="7" pageNumber="224" pagination="163 - 185" refId="ref6342" refString="Ingrisch, S. (1987) Neue Grillen von Borneo und aus Thailand (Insecta: Saltatoria: Grylloidea). Senckenbergiana biologica, 68 (1 - 3), 163 - 185." type="journal article" year="1987">Ingrisch (1987)</bibRefCitation>
as well as the recent specimens collected from
<collectingCountry id="F321BCDCB967FFB74BDEFE475FD009AC" box="[151,227,405,431]" name="Brunei" pageId="7" pageNumber="224">Brunei</collectingCountry>
and Sandakan. However, we could not find any publication of the description associated with
<taxonomicName id="4C3687CFB967FFB74FB3FE445A5D09AC" box="[1274,1390,406,431]" class="Insecta" family="Haglotettigoniidae" genus="Rhicnogryllus" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="eximius">
<emphasis id="B942205EB967FFB74FB3FE445A5D09AC" box="[1274,1390,406,431]" italics="true" pageId="7" pageNumber="224">R. eximius</emphasis>
</taxonomicName>
and concluded that Chopard may not have described this species despite specifying the type specimens until
<bibRefCitation id="EFA781BDB967FFB74E09FE6B5FD109F4" author="Ingrisch" firstAuthor="Ingrisch" pageId="7" pageNumber="224" pagination="163 - 185" refId="ref6342" refString="Ingrisch, S. (1987) Neue Grillen von Borneo und aus Thailand (Insecta: Saltatoria: Grylloidea). Senckenbergiana biologica, 68 (1 - 3), 163 - 185." type="journal article" year="1987">Ingrisch (1987)</bibRefCitation>
described the exact same species from
<collectingCountry id="F321BCDCB967FFB749EDFE0F5C3B09F4" box="[676,776,477,503]" name="Malaysia" pageId="7" pageNumber="224">Sarawak</collectingCountry>
. Thus, we consider Chopard’s
<taxonomicName id="4C3687CFB967FFB74F2DFE0C5BE809F4" box="[1124,1243,478,503]" class="Insecta" family="Haglotettigoniidae" genus="Rhicnogryllus" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="eximius">
<emphasis id="B942205EB967FFB74F2DFE0C5BE809F4" box="[1124,1243,478,503]" italics="true" pageId="7" pageNumber="224">R. eximius</emphasis>
</taxonomicName>
as a synonym of
<taxonomicName id="4C3687CFB967FFB74BDEFDD05E0A0A18" authorityName="Ingrisch" authorityYear="1987" box="[151,313,513,539]" class="Insecta" family="Haglotettigoniidae" genus="Rhicnogryllus" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="bipunctatus">
<emphasis id="B942205EB967FFB74BDEFDD05E0A0A18" box="[151,313,513,539]" italics="true" pageId="7" pageNumber="224">R. bipunctatus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B89FC4CB967FFB74B8EFDF75AA40A3C" blockId="7.[151,1437,152,1439]" box="[199,1431,549,575]" pageId="7" pageNumber="224">
<bibRefCitation id="EFA781BDB967FFB74B8EFDF75E470A3C" author="Ingrisch" box="[199,372,549,575]" firstAuthor="Ingrisch" pageId="7" pageNumber="224" pagination="163 - 185" refId="ref6342" refString="Ingrisch, S. (1987) Neue Grillen von Borneo und aus Thailand (Insecta: Saltatoria: Grylloidea). Senckenbergiana biologica, 68 (1 - 3), 163 - 185." type="journal article" year="1987">Ingrisch (1987)</bibRefCitation>
provided a comprehensive description of the male, but the female was not known at that time.
</paragraph>
</subSubSection>
<subSubSection id="C32CAFC7B967FFB74B8EFD9A5A300AF0" pageId="7" pageNumber="224" type="diagnosis">
<paragraph id="8B89FC4CB967FFB74B8EFD9A5A300AF0" blockId="7.[151,1437,152,1439]" pageId="7" pageNumber="224">
<emphasis id="B942205EB967FFB74B8EFD9A5E710A60" bold="true" box="[199,322,584,611]" pageId="7" pageNumber="224">Diagnosis.</emphasis>
This species differs from congeners by the combination of these characters: generally black colouration, with dorsum of head with a broad white line connecting the eyes; antennal scapus red brown; posterior femora white with two black bands, posterior tibia red brown; tarsal segments whitish; tegmen with a big, round, white spot at the base; supra-anal plate and base of cerci white; and male genitalia (
<figureCitation id="130DE0C9B967FFB748F0FD675CCE0ACC" box="[953,1021,693,719]" captionStart="FIGURE 3" captionStartId="6.[151,250,1980,2005]" captionTargetBox="[206,1380,381,1955]" captionTargetId="figure@6.[206,1380,381,1955]" captionTargetPageId="6" captionText="FIGURE 3. Live habitus of Rhicnogryllus bipunctatus female from Sandakan, Sabah (locality of R. eximius sensu Chopard) (A) and male from Brunei Darussalam (B)." figureDoi="http://doi.org/10.5281/zenodo.3758256" httpUri="https://zenodo.org/record/3758256/files/figure.png" pageId="7" pageNumber="224">Fig. 3</figureCitation>
). The male genitalia is also similar to that of
<taxonomicName id="4C3687CFB967FFB74BAFFD0B5E430AF0" authorityName="Rambur" authorityYear="1838" box="[230,368,729,755]" class="Insecta" family="Haglotettigoniidae" genus="Trigonidium" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="B942205EB967FFB74BAFFD0B5E430AF0" box="[230,368,729,755]" italics="true" pageId="7" pageNumber="224">Trigonidium</emphasis>
</taxonomicName>
but differs by virga as long as rachis (usually shorter than rachis in
<taxonomicName id="4C3687CFB967FFB74F21FD0B5BC10AF0" authorityName="Rambur" authorityYear="1838" box="[1128,1266,729,755]" class="Insecta" family="Haglotettigoniidae" genus="Trigonidium" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="B942205EB967FFB74F21FD0B5BC10AF0" box="[1128,1266,729,755]" italics="true" pageId="7" pageNumber="224">Trigonidium</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C32CAFC7B967FFB74B8EFD2E5B460B14" box="[199,1141,764,792]" pageId="7" pageNumber="224" type="distribution">
<paragraph id="8B89FC4CB967FFB74B8EFD2E5B460B14" blockId="7.[151,1437,152,1439]" box="[199,1141,764,792]" pageId="7" pageNumber="224">
<emphasis id="B942205EB967FFB74B8EFD2E5E520B14" bold="true" box="[199,353,764,791]" pageId="7" pageNumber="224">Distribution.</emphasis>
Borneo: Sandakan (
<collectingRegion id="49F232AEB967FFB7490CFD2F5DBC0B1B" box="[581,655,765,792]" country="Malaysia" name="Sabah" pageId="7" pageNumber="224">Sabah</collectingRegion>
); Batu Niah (
<collectingCountry id="F321BCDCB967FFB7486DFD2F5CB80B1B" box="[804,907,765,792]" name="Malaysia" pageId="7" pageNumber="224">Sarawak</collectingCountry>
);
<collectingCountry id="F321BCDCB967FFB748D7FD2F5B460B14" box="[926,1141,765,791]" name="Brunei" pageId="7" pageNumber="224">Brunei Darussalam</collectingCountry>
</paragraph>
</subSubSection>
<subSubSection id="C32CAFC7B967FFB74B8EFCF35A0F0D9C" pageId="7" pageNumber="224" type="description">
<paragraph id="8B89FC4CB967FFB74B8EFCF35D760C34" blockId="7.[151,1437,152,1439]" pageId="7" pageNumber="224">
<emphasis id="B942205EB967FFB74B8EFCF35DFA0B38" bold="true" box="[199,713,800,827]" pageId="7" pageNumber="224">
Male genitalia characteristics (
<figureCitation id="130DE0C9B967FFB7497DFCF25DB00B38" box="[564,643,800,827]" captionStart="FIGURE 6" captionStartId="10.[151,250,1827,1852]" captionTargetBox="[215,1372,181,1801]" captionTargetId="figure@10.[215,1372,181,1801]" captionTargetPageId="10" captionText="FIGURE 6. Male genitalia of Rhicnogryllus bipunctatus (A–C) and Rhicnogryllus paetensis n. sp. (D–F) in dorsal (A, D), ventral (B, E) and lateral (C, F) views." figureDoi="http://doi.org/10.5281/zenodo.3758262" httpUri="https://zenodo.org/record/3758262/files/figure.png" pageId="7" pageNumber="224">Figs. 6</figureCitation>
A–C).
</emphasis>
Pseudepiphallus separated into two lateral parts joined by an anterior sclerotized bridge; bridge narrow and gently curved. Posterior apex of pseudepiphallus forming thin and long, sclerotized lophi; lophus feebly curved inwards, with inner margin with 4–5 tooth-like spines, with apex forming a tooth; posterior edges of pseudepiphallic sclerite with a stout, angular lobule in the middle, lobule with stout teeth along margin. Pseudepiphallic parameres elongated heart-shaped, not exceeding apex of pseudepiphallic lobules. Ectophallic fold large forming an elongated sclerotized virga with acute apex. Ectophallic apodeme weakly sclerotized but long. Ramus straight. Endophallic sclerite elongated along dorsal cavity, with a median crest, with lamella of apodemes appearing flat and broad.
</paragraph>
<paragraph id="8B89FC4CB967FFB74B8EFB935B000D0C" blockId="7.[151,1437,152,1439]" pageId="7" pageNumber="224">
<emphasis id="B942205EB967FFB74B8EFB935E810C58" bold="true" box="[199,434,1088,1115]" pageId="7" pageNumber="224">Female description.</emphasis>
Habitus similar to males, does not exhibit sexual dimorphism. Supra-anal plate with basal half black, with a longitudinal groove and angularly emarginated at the apex; epiproct with broad and stout, with obtuse apex, white. Subgenital plate stout, distinctly broader than wide, with anterior margin straight, apex narrowly excised in the middle; basal part black, thereafter white. Ovipositor not surpassing cerci; basal third with margins straight and smooth; apical third with dorsal margin distinctly denticulated, ventral margin also denticulated but less dense and only towards the apex. Ventral valves slightly longer than dorsal valves.
</paragraph>
<paragraph id="8B89FC4CB967FFB74B8EFACB5A0F0D9C" blockId="7.[151,1437,152,1439]" pageId="7" pageNumber="224">
<emphasis id="B942205EB967FFB74B8EFACB5EB60D30" bold="true" box="[199,389,1304,1331]" pageId="7" pageNumber="224">Natural history.</emphasis>
This species tends to be found among foliage of shrubs or low-lying vegetation in the understory of relatively disturbed forest and/or forest edge (at least in Sandakan and
<collectingCountry id="F321BCDCB967FFB74F58FAEF5B510D5B" box="[1041,1122,1341,1368]" name="Brunei" pageId="7" pageNumber="224">Brunei</collectingCountry>
). Ulu Dusun and Rainforest Discovery Centre (Sandakan) as well as Teraja and Wasai Wong Kadir Recreational Park (
<collectingCountry id="F321BCDCB967FFB74FE4FAB35BCF0D7F" box="[1197,1276,1377,1404]" name="Brunei" pageId="7" pageNumber="224">Brunei</collectingCountry>
) typically are disturbed forest with the presence of human traffic. The species can be found active in both day and night.
</paragraph>
</subSubSection>
</treatment>
</document>
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