206 lines
28 KiB
XML
206 lines
28 KiB
XML
<document id="9484D36EE906E1726EE9E9365EB41457" ID-DOI="10.2478/if-2019-0013" ID-ISSN="2533-4069" ID-Zenodo-Dep="5386203" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630303095841" checkinUser="diego" docAuthor="Friis, Else Marie, Crane, Peter R. & Pedersen, Kaj Raunsgaard" docDate="2019" docId="0396DC10BF23C22DCEFEB46BE5971A5D" docLanguage="en" docName="2_Friis.pdf" docOrigin="Fossil Imprint 75 (2)" docSource="http://dx.doi.org/10.2478/if-2019-0013" docStyle="DocumentStyle:0EC54BF35E227CA4627F894F6BA30634.4:FossilImprint.2016-.journal_article" docStyleId="0EC54BF35E227CA4627F894F6BA30634" docStyleName="FossilImprint.2016-.journal_article" docStyleVersion="4" docTitle="Goczania punctata E. M. FRIIS, P. R. CRANE et K. R. PEDERSEN 2019, sp. nov." docType="treatment" docVersion="5" lastPageNumber="199" masterDocId="FFAFA468BF0FC203CD58B635E7781C51" masterDocTitle="The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community" masterLastPageNumber="257" masterPageNumber="153" pageNumber="197" updateTime="1699221227565" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="CA3BA355915214E033EC0E411AFE99BF">The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community</mods:title>
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<mods:namePart id="FCCAE724878E19300453EDCADAFAFD06">Friis, Else Marie</mods:namePart>
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<mods:roleTerm id="59563CD6CF8F77CB486220EBEE33141E">Author</mods:roleTerm>
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<mods:namePart id="0A1C8345013B43E1649C842594E09AB5">Crane, Peter R.</mods:namePart>
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<mods:namePart id="0FD1F22DC981429583C9231B62355444">Pedersen, Kaj Raunsgaard</mods:namePart>
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<mods:typeOfResource id="D878FF8EC058AD53E74B1BD90E7109D9">text</mods:typeOfResource>
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<mods:title id="6DF6138EB4DE18E22BD1839E129D8F2B">Fossil Imprint</mods:title>
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<mods:date id="3BD5015DB34BCAAC68DE5FC92E6899D8">2019</mods:date>
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<mods:number id="D6815F44BE2D799B0EB9F5DFCF88A047">2019-11-25</mods:number>
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<paragraph id="8B806D06BF23C22FCEFEB46BE26D1EC6" blockId="44.[934,1390,606,663]" pageId="44" pageNumber="197">
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<heading id="D0C8DA6ABF23C22FCEFEB46BE26D1EC6" centered="true" fontSize="8" level="2" pageId="44" pageNumber="197" reason="2">
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<emphasis id="B94BB114BF23C22FCEFEB46BE26D1EC6" bold="true" pageId="44" pageNumber="197">
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<taxonomicName id="4C3F1685BF23C22FCEFEB46BE3C11EC4" authority="E.M.FRIIS, P.R.CRANE et K.R.PEDERSEN" authorityName="E. M. FRIIS, P. R. CRANE et K. R. PEDERSEN" authorityYear="2019" class="Magnoliopsida" family="Chloranthaceae" genus="Goczania" kingdom="Plantae" order="Chloranthales" pageId="44" pageNumber="197" phylum="Tracheophyta" rank="species" species="punctata" status="sp. nov.">
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<emphasis id="B94BB114BF23C22FCEFEB46BE30C1E27" bold="true" box="[934,1140,606,630]" italics="true" pageId="44" pageNumber="197">Goczania punctata</emphasis>
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E.M.FRIIS, P.R.CRANE et K.R.PEDERSEN
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</taxonomicName>
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<taxonomicNameLabel id="A2780C6FBF23C22FC998B44AE26D1EC6" box="[1216,1301,639,663]" pageId="44" pageNumber="197" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B806D06BF23C22FC962B4AAE3A31EE6" blockId="44.[1082,1243,671,695]" box="[1082,1243,671,695]" pageId="44" pageNumber="197">
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<figureCitation id="13047183BF23C22FC962B4AAE3A31EE6" box="[1082,1243,671,695]" captionStart="Text-fig" captionStartId="47.[124,205,1990,2011]" captionTargetBox="[125,1439,694,1970]" captionTargetId="figure-303@47.[125,1439,694,1972]" captionTargetPageId="47" captionText="Text-fig. 29. Scanning electron microscope (SEM) images of monocolpate pollen of Goczania punctata sp. nov.; Torres Vedras locality, Portugal. a–c) Holotype; anther fragment (a) with group of pollen grains showing the distal face (b, c) with clearly delimited colpus, the almost psilate tectum with occasional small perforations, and poorly differentiated microechinae on the proximal face (b). Specimen, TV44-S148024 (holotype). Scale bars 300 Μm (a), 6 Μm (b, c)." figureDoi="http://doi.org/10.5281/zenodo.5386299" httpUri="https://zenodo.org/record/5386299/files/figure.png" pageId="44" pageNumber="197">Text-fig. 29a–c</figureCitation>
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</paragraph>
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<subSubSection id="C3253E8DBF23C22DCE29B4FAE5971A5D" lastPageId="46" lastPageNumber="199" pageId="44" pageNumber="197" type="description">
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<paragraph id="8B806D06BF23C22FCE29B4FAE3B11F56" blockId="44.[843,1482,719,775]" pageId="44" pageNumber="197">
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H o l o t y p e. Designated here. S148024 (Torres Vedras sample 44; figured
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<figureCitation id="13047183BF23C22FC940B4DAE3C51F56" box="[1048,1213,751,775]" captionStart="Text-fig" captionStartId="47.[124,205,1990,2011]" captionTargetBox="[125,1439,694,1970]" captionTargetId="figure-303@47.[125,1439,694,1972]" captionTargetPageId="47" captionText="Text-fig. 29. Scanning electron microscope (SEM) images of monocolpate pollen of Goczania punctata sp. nov.; Torres Vedras locality, Portugal. a–c) Holotype; anther fragment (a) with group of pollen grains showing the distal face (b, c) with clearly delimited colpus, the almost psilate tectum with occasional small perforations, and poorly differentiated microechinae on the proximal face (b). Specimen, TV44-S148024 (holotype). Scale bars 300 Μm (a), 6 Μm (b, c)." figureDoi="http://doi.org/10.5281/zenodo.5386299" httpUri="https://zenodo.org/record/5386299/files/figure.png" pageId="44" pageNumber="197">Text-fig. 29a–c</figureCitation>
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).
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</paragraph>
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<paragraph id="8B806D06BF23C22FCE29B52AE2BF1F67" blockId="44.[843,1479,799,855]" box="[881,1479,799,823]" pageId="44" pageNumber="197">P l a n t F o s s i l N a m e s R e g i s t r y N u m b e r.</paragraph>
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<paragraph id="8B806D06BF23C22FCE13B50AE3E91F06" blockId="44.[843,1479,799,855]" box="[843,1169,831,855]" pageId="44" pageNumber="197">PFN000464 (for new species).</paragraph>
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<paragraph id="8B806D06BF23C22FCE29B545E3501F99" blockId="44.[843,1482,879,968]" pageId="44" pageNumber="197">
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<materialsCitation id="3B57675BBF23C22FCE29B545E35C1F99" collectionCode="R" country="Sweden" county="Swedish Museum of Natural History" location="Palaeobotanical Collections" municipality="Department of Palaeobiology" pageId="44" pageNumber="197" specimenCount="1" stateProvince="Stockholm">
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<collectionCode id="ED2EF5C3BF23C22FCE29B545E4FB1FD6" box="[881,899,880,903]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="44" pageNumber="197">R</collectionCode>
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e p o s i t o r y.
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<location id="8EE03BDDBF23C22FC97AB55AE23D1FD6" LSID="urn:lsid:plazi:treatment:0396DC10BF23C22DCEFEB46BE5971A5D:8EE03BDDBF23C22FC97AB55AE23D1FD6" box="[1058,1349,879,903]" country="Sweden" county="Swedish Museum of Natural History" municipality="Department of Palaeobiology" name="Palaeobotanical Collections" pageId="44" pageNumber="197" stateProvince="Stockholm">Palaeobotanical Collections</location>
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,
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<collectingMunicipality id="6BE4F77CBF23C22FC817B545E37B1FF6" pageId="44" pageNumber="197">Department of Palaeobiology</collectingMunicipality>
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, the
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<collectingCounty id="62E1158ABF23C22FC96FB5BAE2BE1FF6" box="[1079,1478,911,935]" pageId="44" pageNumber="197">Swedish Museum of Natural History</collectingCounty>
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,
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<collectingRegion id="49FBA3E4BF23C22FCE13B585E4B91F99" box="[843,961,944,968]" country="Sweden" name="Stockholm" pageId="44" pageNumber="197">Stockholm</collectingRegion>
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,
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<collectingCountry id="F3282D96BF23C22FCE94B585E35C1F99" box="[972,1060,944,968]" name="Sweden" pageId="44" pageNumber="197">Sweden</collectingCountry>
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</materialsCitation>
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.
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</paragraph>
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<caption id="DF403D8EBF23C22FCDCBB19DE2DB1462" ID-DOI="http://doi.org/10.5281/zenodo.5386293" ID-Zenodo-Dep="5386293" httpUri="https://zenodo.org/record/5386293/files/figure.png" pageId="44" pageNumber="197" startId="44.[147,228,1960,1981]" targetBox="[151,1463,1039,1939]" targetPageId="44">
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<paragraph id="8B806D06BF23C22FCDCBB19DE2DB1462" blockId="44.[147,1463,1960,2099]" pageId="44" pageNumber="197">
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Text-fig. 27. Scanning electron microscope (SEM) images of monocolpate pollen grains of
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||
<taxonomicName id="4C3F1685BF23C22FC974B19CE3AB1BEF" authority="Friis & Crane & Pedersen, 2019" authorityName="Friis & Crane & Pedersen" authorityYear="2019" box="[1068,1235,1961,1982]" class="Magnoliopsida" family="Chloranthaceae" genus="Goczania" kingdom="Plantae" order="Chloranthales" pageId="44" pageNumber="197" phylum="Tracheophyta" rank="species" species="rugosa" status="gen. et sp. nov.">
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<emphasis id="B94BB114BF23C22FC974B19CE3AB1BEF" bold="true" box="[1068,1235,1961,1982]" italics="true" pageId="44" pageNumber="197">Goczania rugosa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2780C6FBF23C22FC982B19DE2081BEC" box="[1242,1392,1960,1981]" pageId="44" pageNumber="197" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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in situ in dithecate, tetrasporangiate stamens; Torres Vedras locality, Portugal. a, b) Stamens in apical view (holotype a) showing the paired pollen sacs; c) Pollen grain from stamen fragment in (b) showing short aperture and rugulate-microechinate surface; d–f) Pollen grains from stamen fragment in (a) showing a folded distal aperture and rugulate-microechinate surface; note the orbicules of different sizes. Specimens, TV44-S136688 (holotype; a, c–f), TV44-S136664 (b). Scale bars 300 Μm (a,b), 6 Μm (c–e), 3 Μm (f).
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</paragraph>
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||
</caption>
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||
<caption id="DF403D8EBF22C22ECD25B14BE2D81477" ID-DOI="http://doi.org/10.5281/zenodo.5386295" ID-Zenodo-Dep="5386295" httpUri="https://zenodo.org/record/5386295/files/figure.png" pageId="45" pageNumber="198" startId="45.[125,206,1918,1939]" targetBox="[128,1440,151,1898]" targetPageId="45">
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<paragraph id="8B806D06BF22C22ECD25B14BE2D81477" blockId="45.[125,1440,1918,2086]" pageId="45" pageNumber="198">
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Text-fig. 28. Scanning electron microscope (SEM) images of monocolpate pollen of
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<taxonomicName id="4C3F1685BF22C22ECE85B14BE3D11BC2" authority="Friis & Crane & Pedersen, 2019" authorityName="Friis & Crane & Pedersen" authorityYear="2019" box="[989,1193,1918,1939]" class="Magnoliopsida" family="Chloranthaceae" genus="Goczania" kingdom="Plantae" order="Chloranthales" pageId="45" pageNumber="198" phylum="Tracheophyta" rank="species" species="inaequalis" status="sp. nov.">
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<emphasis id="B94BB114BF22C22ECE85B14BE3D11BC2" bold="true" box="[989,1193,1918,1939]" italics="true" pageId="45" pageNumber="198">Goczania inaequalis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2780C6FBF22C22EC9E8B14BE27A1BC2" box="[1200,1282,1918,1939]" pageId="45" pageNumber="198" rank="species">sp. nov.</taxonomicNameLabel>
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; Torres Vedras locality, Portugal. a) Holotype; stamen fragment that yielded the pollen in (b–f, k); b) Group of pollen grains showing distal and proximal surfaces and inner surface of anther wall with densely spaced orbicules; c) Inner surface of anther wall showing small, spherical orbicules with microechinate sculpturing; d, e, j–l) Pollen grains in distal and equatorial view (k) showing the long colpus with an irregular margin flanked by narrow bands of poorly differentiated microechinate; note that colpus is aligned perpendicular to the longest axes in elliptical grains and that the pollen wall is almost psilate around the equator and in the distal
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</paragraph>
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</caption>
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<paragraph id="8B806D06BF21C22DCDFEB6ADE6181C81" blockId="46.[129,767,151,208]" pageId="46" pageNumber="199">E t y m o l o g y. From Latin: punctatus referring to punctate pollen wall.</paragraph>
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<paragraph id="8B806D06BF21C22DCDFEB6DDE5871D51" blockId="46.[129,767,232,288]" box="[166,767,232,256]" pageId="46" pageNumber="199">T y p e l o c a l i t y. Torres Vedras (NE of Forte de</paragraph>
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<paragraph id="8B806D06BF21C22DCDD9B73CE6941D71" blockId="46.[129,767,232,288]" box="[129,492,263,288]" pageId="46" pageNumber="199">
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Forca;
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<geoCoordinate id="EE0B0BC1BF21C22DCD93B732E62C1D71" box="[203,340,263,288]" degrees="39" direction="north" minutes="06" orientation="latitude" pageId="46" pageNumber="199" precision="15" seconds="13" value="39.10361">39°06′13″ N</geoCoordinate>
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,
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<geoCoordinate id="EE0B0BC1BF21C22DCC06B732E6981D71" box="[350,480,263,288]" degrees="9" direction="west" minutes="14" orientation="longitude" pageId="46" pageNumber="199" precision="15" seconds="47" value="-9.246389">9°14′47″ W</geoCoordinate>
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).
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</paragraph>
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<paragraph id="8B806D06BF21C22DCDFEB70DE66C1DC0" blockId="46.[129,768,312,401]" pageId="46" pageNumber="199">T y p e s t r a t u m a n d a g e. Lower member of the Almargem Formation; Early Cretaceous (late Barremianearly Aptian).</paragraph>
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<paragraph id="8B806D06BF21C22DCDFEB79CE7931E13" blockId="46.[129,768,424,578]" pageId="46" pageNumber="199">D i a g n o s i s. As for the genus with the following additions: Equatorial outline of pollen circular to transversely elliptical. Microechinae on proximal surface and bordering the colpus; distal face, except along the colpus, punctatefoveolate.</paragraph>
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<paragraph id="8B806D06BF21C22DCDFEB46FE5DD1F64" blockId="46.[128,768,602,821]" pageId="46" pageNumber="199">
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D i s t i n g u i s h i n g f e a t u r e s. Pollen of
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<taxonomicName id="4C3F1685BF21C22DCFC1B46FE7A51EC3" authorityName="E. M. FRIIS, P. R. CRANE et K. R. PEDERSEN" authorityYear="2019" class="Magnoliopsida" family="Chloranthaceae" genus="Goczania" kingdom="Plantae" order="Chloranthales" pageId="46" pageNumber="199" phylum="Tracheophyta" rank="species" species="punctata">
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<emphasis id="B94BB114BF21C22DCFC1B46FE7A51EC3" italics="true" pageId="46" pageNumber="199">Goczania punctata</emphasis>
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</taxonomicName>
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is closely similar to that of
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<taxonomicName id="4C3F1685BF21C22DCF6BB44EE5B11EC3" authorityName="Friis & Crane & Pedersen" authorityYear="2019" box="[563,713,634,658]" class="Magnoliopsida" family="Chloranthaceae" genus="Goczania" kingdom="Plantae" order="Chloranthales" pageId="46" pageNumber="199" phylum="Tracheophyta" rank="species" species="inaequalis">
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<emphasis id="B94BB114BF21C22DCF6BB44EE5B11EC3" box="[563,713,634,658]" italics="true" pageId="46" pageNumber="199">G. inaequalis</emphasis>
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</taxonomicName>
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, but differs in the slightly larger grains that have a distinctly perforate tectum on their distal face. Pollen of
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<taxonomicName id="4C3F1685BF21C22DCFC1B489E7A61EA2" authorityName="E. M. FRIIS, P. R. CRANE et K. R. PEDERSEN" authorityYear="2019" class="Magnoliopsida" family="Chloranthaceae" genus="Goczania" kingdom="Plantae" order="Chloranthales" pageId="46" pageNumber="199" phylum="Tracheophyta" rank="species" species="punctata">
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<emphasis id="B94BB114BF21C22DCFC1B489E7A61EA2" italics="true" pageId="46" pageNumber="199">Goczania punctata</emphasis>
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</taxonomicName>
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is also distinguished from that of
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<taxonomicName id="4C3F1685BF21C22DCF37B4E9E5A61EA2" authorityName="Friis & Crane & Pedersen" authorityYear="2019" box="[623,734,732,755]" class="Magnoliopsida" family="Chloranthaceae" genus="Goczania" kingdom="Plantae" order="Chloranthales" pageId="46" pageNumber="199" phylum="Tracheophyta" rank="species" species="rugosa">
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||
<emphasis id="B94BB114BF21C22DCF37B4E9E5A61EA2" box="[623,734,732,755]" italics="true" pageId="46" pageNumber="199">G. rugosa</emphasis>
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</taxonomicName>
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in having a smoother surface and in having the microechinae restricted mainly to the proximal face of the grains.
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||
</paragraph>
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<paragraph id="8B806D06BF21C22DCDFEB578E5841F34" blockId="46.[166,764,844,869]" box="[166,764,844,869]" pageId="46" pageNumber="199">D i m e n s i o n s. Diameter of pollen grains: 13–14 µm.</paragraph>
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<paragraph id="8B806D06BF21C22DCDFEB549E5371847" blockId="46.[129,768,892,1338]" pageId="46" pageNumber="199">
|
||
D e s c r i p t i o n a n d r e m a r k s.
|
||
<taxonomicName id="4C3F1685BF21C22DCF6DB549E5871FC5" authorityName="E. M. FRIIS, P. R. CRANE et K. R. PEDERSEN" authorityYear="2019" box="[565,767,892,916]" class="Magnoliopsida" family="Chloranthaceae" genus="Goczania" kingdom="Plantae" order="Chloranthales" pageId="46" pageNumber="199" phylum="Tracheophyta" rank="species" species="punctata">
|
||
<emphasis id="B94BB114BF21C22DCF6DB549E5871FC5" box="[565,767,892,916]" italics="true" pageId="46" pageNumber="199">Goczania punctata</emphasis>
|
||
</taxonomicName>
|
||
is based on a single anther fragment (
|
||
<figureCitation id="13047183BF21C22DCF76B5A9E5C51FE5" box="[558,701,924,948]" captionStart="Text-fig" captionStartId="47.[124,205,1990,2011]" captionTargetBox="[125,1439,694,1970]" captionTargetId="figure-303@47.[125,1439,694,1972]" captionTargetPageId="47" captionText="Text-fig. 29. Scanning electron microscope (SEM) images of monocolpate pollen of Goczania punctata sp. nov.; Torres Vedras locality, Portugal. a–c) Holotype; anther fragment (a) with group of pollen grains showing the distal face (b, c) with clearly delimited colpus, the almost psilate tectum with occasional small perforations, and poorly differentiated microechinae on the proximal face (b). Specimen, TV44-S148024 (holotype). Scale bars 300 Μm (a), 6 Μm (b, c)." figureDoi="http://doi.org/10.5281/zenodo.5386299" httpUri="https://zenodo.org/record/5386299/files/figure.png" pageId="46" pageNumber="199">Text-fig. 29a</figureCitation>
|
||
) with pollen in situ. Orbicules on the inner surface of the anther and among the pollen grains are rare. They are small, spherical and with a microechinate surface.
|
||
</paragraph>
|
||
<paragraph id="8B806D06BF21C22DCDFEB22BE571196B" blockId="46.[129,768,892,1338]" pageId="46" pageNumber="199">
|
||
Pollen grains are small, circular to slightly transversely elliptical in equatorial outline, about 13–14 µm in equatorial diameter (
|
||
<figureCitation id="13047183BF21C22DCDB5B26AE6ED1826" box="[237,405,1119,1143]" captionStart="Text-fig" captionStartId="47.[124,205,1990,2011]" captionTargetBox="[125,1439,694,1970]" captionTargetId="figure-303@47.[125,1439,694,1972]" captionTargetPageId="47" captionText="Text-fig. 29. Scanning electron microscope (SEM) images of monocolpate pollen of Goczania punctata sp. nov.; Torres Vedras locality, Portugal. a–c) Holotype; anther fragment (a) with group of pollen grains showing the distal face (b, c) with clearly delimited colpus, the almost psilate tectum with occasional small perforations, and poorly differentiated microechinae on the proximal face (b). Specimen, TV44-S148024 (holotype). Scale bars 300 Μm (a), 6 Μm (b, c)." figureDoi="http://doi.org/10.5281/zenodo.5386299" httpUri="https://zenodo.org/record/5386299/files/figure.png" pageId="46" pageNumber="199">Text-fig. 29b, c</figureCitation>
|
||
) and monocolpate. The colpus is long, reaching almost to the equator (
|
||
<figureCitation id="13047183BF21C22DCF4FB2B5E5B918C9" box="[535,705,1152,1176]" captionStart="Text-fig" captionStartId="47.[124,205,1990,2011]" captionTargetBox="[125,1439,694,1970]" captionTargetId="figure-303@47.[125,1439,694,1972]" captionTargetPageId="47" captionText="Text-fig. 29. Scanning electron microscope (SEM) images of monocolpate pollen of Goczania punctata sp. nov.; Torres Vedras locality, Portugal. a–c) Holotype; anther fragment (a) with group of pollen grains showing the distal face (b, c) with clearly delimited colpus, the almost psilate tectum with occasional small perforations, and poorly differentiated microechinae on the proximal face (b). Specimen, TV44-S148024 (holotype). Scale bars 300 Μm (a), 6 Μm (b, c)." figureDoi="http://doi.org/10.5281/zenodo.5386299" httpUri="https://zenodo.org/record/5386299/files/figure.png" pageId="46" pageNumber="199">Text-fig. 29b, c</figureCitation>
|
||
). The colpus margin is distinct (
|
||
<figureCitation id="13047183BF21C22DCCCAB295E55918E9" box="[402,545,1184,1208]" captionStart="Text-fig" captionStartId="47.[124,205,1990,2011]" captionTargetBox="[125,1439,694,1970]" captionTargetId="figure-303@47.[125,1439,694,1972]" captionTargetPageId="47" captionText="Text-fig. 29. Scanning electron microscope (SEM) images of monocolpate pollen of Goczania punctata sp. nov.; Torres Vedras locality, Portugal. a–c) Holotype; anther fragment (a) with group of pollen grains showing the distal face (b, c) with clearly delimited colpus, the almost psilate tectum with occasional small perforations, and poorly differentiated microechinae on the proximal face (b). Specimen, TV44-S148024 (holotype). Scale bars 300 Μm (a), 6 Μm (b, c)." figureDoi="http://doi.org/10.5281/zenodo.5386299" httpUri="https://zenodo.org/record/5386299/files/figure.png" pageId="46" pageNumber="199">Text-fig. 29b</figureCitation>
|
||
). The exine is tectate with a supratectal ornamentation of solitary microechinae on the proximal surface and bordering the colpus. On the distal surface, beyond the colpus margin, the tectum is psilatepunctate-foveolate (
|
||
<figureCitation id="13047183BF21C22DCC0DB317E683196B" box="[341,507,1314,1338]" captionStart="Text-fig" captionStartId="47.[124,205,1990,2011]" captionTargetBox="[125,1439,694,1970]" captionTargetId="figure-303@47.[125,1439,694,1972]" captionTargetPageId="47" captionText="Text-fig. 29. Scanning electron microscope (SEM) images of monocolpate pollen of Goczania punctata sp. nov.; Torres Vedras locality, Portugal. a–c) Holotype; anther fragment (a) with group of pollen grains showing the distal face (b, c) with clearly delimited colpus, the almost psilate tectum with occasional small perforations, and poorly differentiated microechinae on the proximal face (b). Specimen, TV44-S148024 (holotype). Scale bars 300 Μm (a), 6 Μm (b, c)." figureDoi="http://doi.org/10.5281/zenodo.5386299" httpUri="https://zenodo.org/record/5386299/files/figure.png" pageId="46" pageNumber="199">Text-fig. 29b, c</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B806D06BF21C22DCDFEB367E5971A5D" blockId="46.[128,768,1362,1548]" pageId="46" pageNumber="199">
|
||
A f f i n i t y a n d o t h e r o c c u r r e n c e s.
|
||
<taxonomicName id="4C3F1685BF21C22DCFC3B367E7A319DB" authorityName="E. M. FRIIS, P. R. CRANE et K. R. PEDERSEN" authorityYear="2019" class="Magnoliopsida" family="Chloranthaceae" genus="Goczania" kingdom="Plantae" order="Chloranthales" pageId="46" pageNumber="199" phylum="Tracheophyta" rank="species" species="punctata">
|
||
<emphasis id="B94BB114BF21C22DCFC3B367E7A319DB" italics="true" pageId="46" pageNumber="199">Goczania punctata</emphasis>
|
||
</taxonomicName>
|
||
is currently known only from the Torres Vedras mesofossil flora. Like
|
||
<taxonomicName id="4C3F1685BF21C22DCC2EB3A6E52319FB" authorityName="Friis & Crane & Pedersen" authorityYear="2019" box="[374,603,1427,1450]" class="Magnoliopsida" genus="Appomattoxia" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="46" pageNumber="199" phylum="Tracheophyta" rank="species" species="minuta">
|
||
<emphasis id="B94BB114BF21C22DCC2EB3A6E52319FB" box="[374,603,1427,1450]" italics="true" pageId="46" pageNumber="199">Appomattoxia minuta</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3F1685BF21C22DCF35B3A6E66C199A" class="Magnoliopsida" genus="Appomattoxia" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="46" pageNumber="199" phylum="Tracheophyta" rank="species" species="ancistrophora">
|
||
<emphasis id="B94BB114BF21C22DCF35B3A6E66C199A" italics="true" pageId="46" pageNumber="199">Appomattoxia ancistrophora</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3F1685BF21C22DCC78B386E69C199A" authorityName="Friis & Crane & Pedersen" authorityYear="2019" box="[288,484,1459,1483]" class="Magnoliopsida" family="Chloranthaceae" genus="Appofructus" kingdom="Plantae" order="Chloranthales" pageId="46" pageNumber="199" phylum="Tracheophyta" rank="species" species="nudus">
|
||
<emphasis id="B94BB114BF21C22DCC78B386E69C199A" box="[288,484,1459,1483]" italics="true" pageId="46" pageNumber="199">Appofructus nudus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3F1685BF21C22DCF4DB386E5C0199A" box="[533,696,1459,1483]" class="Magnoliopsida" family="Chloranthaceae" genus="Appofructus" kingdom="Plantae" order="Chloranthales" pageId="46" pageNumber="199" phylum="Tracheophyta" rank="species" species="undetermined">
|
||
<emphasis id="B94BB114BF21C22DCF4DB386E5ED199A" box="[533,661,1459,1483]" italics="true" pageId="46" pageNumber="199">Appofructus</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
, along with
|
||
<taxonomicName id="4C3F1685BF21C22DCDE2B3E1E60A19BA" authorityName="Friis & Crane & Pedersen" authorityYear="2019" box="[186,370,1492,1515]" class="Magnoliopsida" family="Chloranthaceae" genus="Goczania" kingdom="Plantae" order="Chloranthales" pageId="46" pageNumber="199" phylum="Tracheophyta" rank="species" species="rugosa">
|
||
<emphasis id="B94BB114BF21C22DCDE2B3E1E60A19BA" box="[186,370,1492,1515]" italics="true" pageId="46" pageNumber="199">Goczania rugosa</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3F1685BF21C22DCCF6B3E1E5F119BD" authorityName="Friis & Crane & Pedersen" authorityYear="2019" box="[430,649,1492,1516]" class="Magnoliopsida" family="Chloranthaceae" genus="Goczania" kingdom="Plantae" order="Chloranthales" pageId="46" pageNumber="199" phylum="Tracheophyta" rank="species" species="inaequalis">
|
||
<emphasis id="B94BB114BF21C22DCCF6B3E1E5F119BD" box="[430,649,1492,1516]" italics="true" pageId="46" pageNumber="199">Goczania inaequalis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3F1685BF21C22DCFC3B3E1E7A41A5D" authorityName="E. M. FRIIS, P. R. CRANE et K. R. PEDERSEN" authorityYear="2019" class="Magnoliopsida" family="Chloranthaceae" genus="Goczania" kingdom="Plantae" order="Chloranthales" pageId="46" pageNumber="199" phylum="Tracheophyta" rank="species" species="punctata">
|
||
<emphasis id="B94BB114BF21C22DCFC3B3E1E7A41A5D" italics="true" pageId="46" pageNumber="199">Goczania punctata</emphasis>
|
||
</taxonomicName>
|
||
is probably related to extant
|
||
<taxonomicName id="4C3F1685BF21C22DCF53B3C1E5131A5D" box="[523,619,1524,1548]" class="Magnoliopsida" kingdom="Plantae" order="Piperales" pageId="46" pageNumber="199" phylum="Tracheophyta" rank="order">Piperales</taxonomicName>
|
||
(see above).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |