1031 lines
120 KiB
XML
1031 lines
120 KiB
XML
<document id="CED89D5D796958105CB82BD3A01D0103" ID-DOI="10.5281/zenodo.196655" ID-GBIF-Dataset="3e264ade-2499-4077-99d5-b199c1e2f06a" ID-ISSN="1175-5326" ID-Zenodo-Dep="196655" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460248488294" checkinUser="plazi" docAuthor="Piálek, Lubomír, An, Old Ř Ich Ř Í Č, Casciotta, Jorge & Almirón, Adriana" docDate="2010" docId="5E46A70EAC00FFF353DDDB5CFD18F854" docLanguage="en" docName="zt02537p046.pdf" docOrigin="Zootaxa 2537" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Crenicichla Piálek, An, Casciotta & Almirón, 2010, new species" docType="treatment" docVersion="9" lastPageNumber="44" masterDocId="A27FDF76AC05FFF8534ADF27FF89FF83" masterDocTitle="Crenicichla hu, a new species of cichlid fish (Teleostei: Cichlidae) from the Paraná basin in Misiones, Argentina" masterLastPageNumber="46" masterPageNumber="33" pageNumber="38" updateTime="1698242308532" updateUser="plazi">
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<mods:title id="2326DF144C476BD4DE0489F10BFFC6ED">Crenicichla hu, a new species of cichlid fish (Teleostei: Cichlidae) from the Paraná basin in Misiones, Argentina</mods:title>
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<mods:namePart id="57C8875E4E49421D393FE18B9168ED15">Piálek, Lubomír</mods:namePart>
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<mods:namePart id="1AAB6446182CE5F6911ABF0016F7AFBA">An, Old Ř Ich Ř Í Č</mods:namePart>
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<mods:namePart id="EDB84BFD5CEA0BA0BCBA13C62C1A5E59">Casciotta, Jorge</mods:namePart>
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<mods:namePart id="6E24647984E2802B6DD295AEEA43FC8F">Almirón, Adriana</mods:namePart>
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<mods:title id="848CFA8E6C8113D60857B51BB4323097">Zootaxa</mods:title>
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<mods:part id="9140A55CBA2C0054ED9979D1E4B30DFC">
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<mods:date id="5390D40FD5D537DDE4AD4485BF968046">2010</mods:date>
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<mods:number id="AA0D49FCBFEF5D75DF3D943062429ACE">2537</mods:number>
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<mods:identifier id="92863E80CE2DF05CEC224A87C317F118" type="DOI">10.5281/zenodo.196655</mods:identifier>
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<treatment id="5E46A70EAC00FFF353DDDB5CFD18F854" ID-DOI="http://doi.org/10.5281/zenodo.5670194" ID-GBIF-Taxon="119391080" ID-Zenodo-Dep="5670194" LSID="urn:lsid:plazi:treatment:5E46A70EAC00FFF353DDDB5CFD18F854" httpUri="http://treatment.plazi.org/id/5E46A70EAC00FFF353DDDB5CFD18F854" lastPageId="11" lastPageNumber="44" pageId="5" pageNumber="38">
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<subSubSection id="9EF54593AC00FFFD53DDDB5CFE91FB38" pageId="5" pageNumber="38" type="nomenclature">
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<paragraph id="D6501618AC00FFFD53DDDB5CFE62FB16" blockId="5.[151,491,1147,1211]" box="[151,491,1147,1173]" pageId="5" pageNumber="38">
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<heading id="8D18A174AC00FFFD53DDDB5CFE62FB16" bold="true" box="[151,491,1147,1173]" fontSize="11" level="1" pageId="5" pageNumber="38" reason="1">
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<emphasis id="E49BCA0AAC00FFFD53DDDB5CFE62FB16" bold="true" box="[151,491,1147,1173]" pageId="5" pageNumber="38">
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<emphasis id="E49BCA0AAC00FFFD53DDDB5CFEDDFB16" bold="true" box="[151,340,1147,1173]" italics="true" pageId="5" pageNumber="38">
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<taxonomicName id="11EF6D9BAC00FFFD53DDDB5CFEACFB16" ID-CoL="3VK6" ID-ENA="50736" box="[151,293,1147,1173]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="38" phylum="Chordata" rank="genus">Crenicichla</taxonomicName>
|
||
hu,
|
||
</emphasis>
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<taxonomicNameLabel id="FFA87771AC00FFFD5211DB5CFE62FB16" box="[347,491,1147,1173]" pageId="5" pageNumber="38" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="D6501618AC00FFFD53DDDB86FE91FB38" blockId="5.[151,491,1147,1211]" box="[151,280,1185,1211]" pageId="5" pageNumber="38">
|
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(
|
||
<figureCitation id="4ED40A9DAC00FFFD53D5DB86FE86FB38" box="[159,271,1185,1211]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStartId-0="6.[151,255,913,937]" captionStartId-1="6.[151,255,1713,1737]" captionStartId-2="8.[151,255,951,975]" captionStartId-3="8.[151,255,1481,1505]" captionTargetBox-0="[194,1391,521,856]" captionTargetBox-1="[151,1435,988,1691]" captionTargetBox-2="[151,1435,226,929]" captionTargetBox-3="[509,1082,1035,1451]" captionTargetId-0="figure@6.[180,1406,493,892]" captionTargetId-1="figure@6.[151,1436,988,1691]" captionTargetId-2="figure@8.[151,1436,226,929]" captionTargetId-3="figure@8.[501,1085,1025,1460]" captionTargetPageId-0="6" captionTargetPageId-1="6" captionTargetPageId-2="8" captionTargetPageId-3="8" captionText-0="FIGURE 2. Crenicichla hu, female, holotype, MACN-ict 9429, 118.0 mm, arroyo Piray-Miní, 26 ° 20 ' 00 " S 53 ° 52 ' 30 " W." captionText-1="FIGURE 3. Crenicichla hu male, live specimen, paratype, MACN-ict 9430, 153.0 mm." captionText-2="FIGURE 4. Crenicichla hu female, live specimen, paratype, MACN-ict 9430, 122.0 mm." captionText-3="FIGURE 5. Crenicichla hu, lower pharyngeal tooth plate in occlusal view, AI 262, 93.9 mm SL. Scale bar: 1 mm." httpUri-0="https://zenodo.org/record/196657/files/figure.png" httpUri-1="https://zenodo.org/record/196658/files/figure.png" httpUri-2="https://zenodo.org/record/196659/files/figure.png" httpUri-3="https://zenodo.org/record/196660/files/figure.png" pageId="5" pageNumber="38">Figs. 2–5</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9EF54593AC00FFFD53DDDBCCFB57FAF6" pageId="5" pageNumber="38" type="materials_examined">
|
||
<paragraph id="D6501618AC00FFFD53DDDBCCFCF6FAA8" blockId="5.[151,1437,1259,2032]" pageId="5" pageNumber="38">
|
||
<emphasis id="E49BCA0AAC00FFFD53DDDBCCFE94FA86" bold="true" box="[151,285,1259,1285]" pageId="5" pageNumber="38">
|
||
<typeStatus id="0954A8BAAC00FFFD53DDDBCCFE9EFA86" box="[151,279,1259,1285]" pageId="5" pageNumber="38" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
MACN-ict 9429, 118.0 mm,
|
||
<collectingCountry id="AEF85688AC00FFFD51D5DBCCFCADFA86" box="[671,804,1259,1285]" name="Argentina" pageId="5" pageNumber="38">Argentina</collectingCountry>
|
||
, Misiones, río Paraná basin, arroyo Piray–Miní,
|
||
<geoCoordinate id="B3DB70DFAC00FFFD53DDDA36FEBBFAA8" box="[151,306,1297,1323]" direction="south" orientation="latitude" pageId="5" pageNumber="38" precision="1" value="-26.333418">26°20'00.3"S</geoCoordinate>
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||
<geoCoordinate id="B3DB70DFAC00FFFD5273DA36FE56FAA8" box="[313,479,1297,1323]" direction="west" orientation="longitude" pageId="5" pageNumber="38" precision="1" value="-53.875">53°52'30.0"W</geoCoordinate>
|
||
,
|
||
<date id="A25130D8AC00FFFD52A1DA36FDEBFAA8" box="[491,610,1297,1323]" pageId="5" pageNumber="38" value="2007-11">Nov 2007</date>
|
||
,
|
||
<collectingCountry id="AEF85688AC00FFFD5125DA36FD0EFAA8" box="[623,647,1297,1323]" name="Iceland" pageId="5" pageNumber="38">O</collectingCountry>
|
||
. Ř íčan
|
||
<emphasis id="E49BCA0AAC00FFFD5194DA36FC9BFAA9" box="[734,786,1297,1322]" italics="true" pageId="5" pageNumber="38">et al</emphasis>
|
||
. (
|
||
<figureCitation id="4ED40A9DAC00FFFD5063DA36FCF8FAA8" box="[809,881,1297,1323]" captionStart="FIGURE 2" captionStartId="6.[151,255,913,937]" captionTargetBox="[194,1391,521,856]" captionTargetId="figure@6.[180,1406,493,892]" captionTargetPageId="6" captionText="FIGURE 2. Crenicichla hu, female, holotype, MACN-ict 9429, 118.0 mm, arroyo Piray-Miní, 26 ° 20 ' 00 " S 53 ° 52 ' 30 " W." httpUri="https://zenodo.org/record/196657/files/figure.png" pageId="5" pageNumber="38">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
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||
<paragraph id="D6501618AC00FFFD538FDA12FB57FAF6" blockId="5.[151,1437,1259,2032]" pageId="5" pageNumber="38">
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||
<emphasis id="E49BCA0AAC00FFFD538FDA12FEC3FACC" bold="true" box="[197,330,1333,1359]" pageId="5" pageNumber="38">
|
||
<typeStatus id="0954A8BAAC00FFFD538FDA12FECCFACC" box="[197,325,1333,1359]" pageId="5" pageNumber="38" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
All from
|
||
<collectingCountry id="AEF85688AC00FFFD528BDA11FDB0FAD3" box="[449,569,1334,1360]" name="Argentina" pageId="5" pageNumber="38">Argentina</collectingCountry>
|
||
, same data as the
|
||
<typeStatus id="0954A8BAAC00FFFD505BDA11FCF5FAD3" box="[785,892,1334,1360]" pageId="5" pageNumber="38" type="holotype">holotype</typeStatus>
|
||
. MACN-ict 9430, 17 ex., 76.9–153.0 mm. AI 261, 4 ex., 96.3–110.0 mm. AI 262, 1 ex. (C&S)
|
||
<quantity id="1117BBFDAC00FFFD519CDA7CFCCBFAF6" box="[726,834,1371,1397]" metricMagnitude="-2" metricUnit="m" metricValue="9.39" pageId="5" pageNumber="38" unit="mm" value="93.9">93.9 mm</quantity>
|
||
, same data as
|
||
<typeStatus id="0954A8BAAC00FFFD50A2DA7CFBC6FAF6" box="[1000,1103,1371,1397]" pageId="5" pageNumber="38" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="4ED40A9DAC00FFFD5715DA7CFB58FAF6" box="[1119,1233,1371,1397]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="6.[151,255,1713,1737]" captionStartId-1="8.[151,255,951,975]" captionTargetBox-0="[151,1435,988,1691]" captionTargetBox-1="[151,1435,226,929]" captionTargetId-0="figure@6.[151,1436,988,1691]" captionTargetId-1="figure@8.[151,1436,226,929]" captionTargetPageId-0="6" captionTargetPageId-1="8" captionText-0="FIGURE 3. Crenicichla hu male, live specimen, paratype, MACN-ict 9430, 153.0 mm." captionText-1="FIGURE 4. Crenicichla hu female, live specimen, paratype, MACN-ict 9430, 122.0 mm." httpUri-0="https://zenodo.org/record/196658/files/figure.png" httpUri-1="https://zenodo.org/record/196659/files/figure.png" pageId="5" pageNumber="38">Figs. 3–4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9EF54593AC00FFFE538FDAA7FBC5FE34" lastPageId="6" lastPageNumber="39" pageId="5" pageNumber="38" type="diagnosis">
|
||
<paragraph id="D6501618AC00FFFD538FDAA7FB4FF988" blockId="5.[151,1437,1259,2032]" pageId="5" pageNumber="38">
|
||
<emphasis id="E49BCA0AAC00FFFD538FDAA7FEC0FA19" bold="true" box="[197,329,1408,1434]" pageId="5" pageNumber="38">Diagnosis.</emphasis>
|
||
<emphasis id="E49BCA0AAC00FFFD5218DAA6FD8FFA19" box="[338,518,1409,1434]" italics="true" pageId="5" pageNumber="38">
|
||
<taxonomicName id="11EF6D9BAC00FFFD5218DAA6FE69FA19" box="[338,480,1409,1434]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="38" phylum="Chordata" rank="genus">Crenicichla</taxonomicName>
|
||
hu
|
||
</emphasis>
|
||
is distinguished from all known species of the La Plata basin and adjacent coastal rivers by the following combination of characters: 1. dark grey or dark brown to black color of body and fins, 2. 7 to 9 black irregular blotches on the flank, 3. 47–54 scales in row E1, 4. the dorsal fin of adult females with a color pattern formed of black and white longitudinal stripes and/or blotches.
|
||
</paragraph>
|
||
<paragraph id="D6501618AC00FFFD538FD931FC1AF9F8" blockId="5.[151,1437,1259,2032]" pageId="5" pageNumber="38">
|
||
Since the molecular analysis confirmed close relations between
|
||
<emphasis id="E49BCA0AAC00FFFD5086D931FB87F9AC" box="[972,1038,1558,1583]" italics="true" pageId="5" pageNumber="38">C. hu</emphasis>
|
||
and its biogeographic congeners from the Paraná basin (
|
||
<figureCitation id="4ED40A9DAC00FFFD52F0D91CFD8AF9D6" box="[442,515,1595,1621]" captionStart="FIGURE 1" captionStartId="5.[151,255,965,989]" captionTargetBox="[158,1418,179,941]" captionTargetId="figure@5.[151,1436,167,949]" captionTargetPageId="5" captionText="FIGURE 1. Maximum parsimony tree topology based on ND 2 sequences; the dotted line displays the different topology of the alternative Bayesian inference. Numbers for each recovered node represent nonparametric bootstrap support (left) and Bayesian posterior probability (right), respectively." httpUri="https://zenodo.org/record/196656/files/figure.png" pageId="5" pageNumber="38">Fig. 1</figureCitation>
|
||
), a detailed comparative analysis was performed on all 13 known species inhabiting the Paraná drainage basin, either exclusively or partly:
|
||
</paragraph>
|
||
<paragraph id="D6501618AC00FFFE538FD9A1FBC5FE34" blockId="5.[151,1437,1259,2032]" lastBlockId="6.[151,1436,152,439]" lastPageId="6" lastPageNumber="39" pageId="5" pageNumber="38">
|
||
<emphasis id="E49BCA0AAC00FFFD538FD9A1FEFBF91C" box="[197,370,1670,1695]" italics="true" pageId="5" pageNumber="38">
|
||
<taxonomicName id="11EF6D9BAC00FFFD538FD9A1FEC7F91C" box="[197,334,1670,1695]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="38" phylum="Chordata" rank="genus">Crenicichla</taxonomicName>
|
||
hu
|
||
</emphasis>
|
||
is distinguished from
|
||
<taxonomicName id="11EF6D9BAC00FFFD513DD9A1FD65F91C" box="[631,748,1670,1695]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="38" phylum="Chordata" rank="species" species="britskii">
|
||
<emphasis id="E49BCA0AAC00FFFD513DD9A1FD65F91C" box="[631,748,1670,1695]" italics="true" pageId="5" pageNumber="38">C. britskii</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="11EF6D9BAC00FFFD506ED9A1FC2FF91C" box="[804,934,1670,1695]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="38" phylum="Chordata" rank="species" species="lepidota">
|
||
<emphasis id="E49BCA0AAC00FFFD506ED9A1FC2FF91C" box="[804,934,1670,1695]" italics="true" pageId="5" pageNumber="38">C. lepidota</emphasis>
|
||
</taxonomicName>
|
||
(both
|
||
<taxonomicName id="11EF6D9BAC00FFFD50BBD9A1FBFCF91C" box="[1009,1141,1670,1695]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="38" phylum="Chordata" rank="species" species="saxatilis">
|
||
<emphasis id="E49BCA0AAC00FFFD50BBD9A1FBFCF91C" box="[1009,1141,1670,1695]" italics="true" pageId="5" pageNumber="38">C. saxatilis</emphasis>
|
||
</taxonomicName>
|
||
group) by the absence of the distinctive humeral spot vs. a humeral spot present (synapomorphy of the group).
|
||
<emphasis id="E49BCA0AAC00FFFD578CD98CFAF6F947" box="[1222,1407,1707,1732]" italics="true" pageId="5" pageNumber="38">
|
||
<taxonomicName id="11EF6D9BAC00FFFD578CD98CFADEF947" box="[1222,1367,1707,1732]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="38" phylum="Chordata" rank="genus">Crenicichla</taxonomicName>
|
||
hu
|
||
</emphasis>
|
||
is distinguished from
|
||
<taxonomicName id="11EF6D9BAC00FFFD52CBD9F6FD83F969" box="[385,522,1745,1770]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="38" phylum="Chordata" rank="species" species="haroldoi">
|
||
<emphasis id="E49BCA0AAC00FFFD52CBD9F6FD83F969" box="[385,522,1745,1770]" italics="true" pageId="5" pageNumber="38">C. haroldoi</emphasis>
|
||
</taxonomicName>
|
||
by the absence of dots on lateral line scales vs. brown dots present on each lateral line scale. It differs from
|
||
<taxonomicName id="11EF6D9BAC00FFFD516DD9D1FD5FF88C" box="[551,726,1782,1807]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="38" phylum="Chordata" rank="species" species="iguassensis">
|
||
<emphasis id="E49BCA0AAC00FFFD516DD9D1FD5FF88C" box="[551,726,1782,1807]" italics="true" pageId="5" pageNumber="38">C. iguassensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="11EF6D9BAC00FFFD5059D9D1FCFFF88C" box="[787,886,1782,1807]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="38" phylum="Chordata" rank="species" species="tesay">
|
||
<emphasis id="E49BCA0AAC00FFFD5059D9D1FCFFF88C" box="[787,886,1782,1807]" italics="true" pageId="5" pageNumber="38">C. tesay</emphasis>
|
||
</taxonomicName>
|
||
in the absence of small dots on the flank vs. numerous scattered small dots present.
|
||
<emphasis id="E49BCA0AAC00FFFD513BD83CFCADF8B7" box="[625,804,1819,1844]" italics="true" pageId="5" pageNumber="38">
|
||
<taxonomicName id="11EF6D9BAC00FFFD513BD83CFD74F8B7" box="[625,765,1819,1844]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="38" phylum="Chordata" rank="genus">Crenicichla</taxonomicName>
|
||
hu
|
||
</emphasis>
|
||
is distinguished from
|
||
<taxonomicName id="11EF6D9BAC00FFFD5779D83CFB69F8B7" box="[1075,1248,1819,1844]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="38" phylum="Chordata" rank="species" species="jaguarensis">
|
||
<emphasis id="E49BCA0AAC00FFFD5779D83CFB69F8B7" box="[1075,1248,1819,1844]" italics="true" pageId="5" pageNumber="38">C. jaguarensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="11EF6D9BAC00FFFD57BBD83CFAE9F8B7" box="[1265,1376,1819,1844]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="38" phylum="Chordata" rank="species" species="vittata">
|
||
<emphasis id="E49BCA0AAC00FFFD57BBD83CFAE9F8B7" box="[1265,1376,1819,1844]" italics="true" pageId="5" pageNumber="38">C. vittata</emphasis>
|
||
</taxonomicName>
|
||
, and adults of
|
||
<taxonomicName id="11EF6D9BAC00FFFD5246D866FE6CF8D9" box="[268,485,1857,1882]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="38" phylum="Chordata" rank="species" species="mandelburgeri">
|
||
<emphasis id="E49BCA0AAC00FFFD5246D866FE6CF8D9" box="[268,485,1857,1882]" italics="true" pageId="5" pageNumber="38">C. mandelburgeri</emphasis>
|
||
</taxonomicName>
|
||
by the absence of a lateral band vs. a lateral band present. It differs from
|
||
<taxonomicName id="11EF6D9BAC00FFFD56CAD866FEA4F8FC" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="38" phylum="Chordata" rank="species" species="jaguarensis">
|
||
<emphasis id="E49BCA0AAC00FFFD56CAD866FEA4F8FC" italics="true" pageId="5" pageNumber="38">C. jaguarensis</emphasis>
|
||
</taxonomicName>
|
||
in the absence vs. presence of the caudal spot. Further,
|
||
<emphasis id="E49BCA0AAC00FFFD5745D841FB59F8FC" box="[1039,1232,1894,1919]" italics="true" pageId="5" pageNumber="38">
|
||
<taxonomicName id="11EF6D9BAC00FFFD5745D841FB2EF8FC" box="[1039,1191,1894,1919]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="38" phylum="Chordata" rank="genus">Crenicichla</taxonomicName>
|
||
hu
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="11EF6D9BAC00FFFD56CAD841FECDF827" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="38" phylum="Chordata" rank="species" species="mandelburgeri">
|
||
<emphasis id="E49BCA0AAC00FFFD56CAD841FECDF827" italics="true" pageId="5" pageNumber="38">C. mandelburgeri</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="11EF6D9BAC00FFFD5237D8ACFDAFF827" box="[381,550,1931,1956]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="38" phylum="Chordata" rank="species" species="niederleinii">
|
||
<emphasis id="E49BCA0AAC00FFFD5237D8ACFDAFF827" box="[381,550,1931,1956]" italics="true" pageId="5" pageNumber="38">C. niederleinii</emphasis>
|
||
</taxonomicName>
|
||
in the absence vs. presence of the narrow vertical double-bars on the flank. It is also distinguished by a low number of scales in a lateral row, 47–54 vs.
|
||
<quantity id="1117BBFDAC00FFFD5757D896FB0CF848" box="[1053,1157,1969,1995]" metricMagnitude="0" metricUnit="m" metricValue="1.5367" metricValueMax="1.6509999999999998" metricValueMin="1.4224" pageId="5" pageNumber="38" unit="in" value="60.5" valueMax="65.0" valueMin="56.0">56–65 in</quantity>
|
||
<taxonomicName id="11EF6D9BAC00FFFD57C6D896FABFF849" box="[1164,1334,1969,1994]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="38" phylum="Chordata" rank="species" species="niederleinii">
|
||
<emphasis id="E49BCA0AAC00FFFD57C6D896FABFF849" box="[1164,1334,1969,1994]" italics="true" pageId="5" pageNumber="38">C. niederleinii</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<quantity id="1117BBFDAC00FFFD563AD896FF5DF873" metricMagnitude="0" metricUnit="m" metricValue="2.0701" metricValueMax="2.159" metricValueMin="1.9812" pageId="5" pageNumber="38" unit="in" value="81.5" valueMax="85.0" valueMin="78.0">78– 85 in</quantity>
|
||
<taxonomicName id="11EF6D9BAC00FFFD5397D8F1FEC7F86C" box="[221,334,2006,2031]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="38" phylum="Chordata" rank="species" species="vittata">
|
||
<emphasis id="E49BCA0AAC00FFFD5397D8F1FEC7F86C" box="[221,334,2006,2031]" italics="true" pageId="5" pageNumber="38">C. vittata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="E49BCA0AAC00FFFD5217D8F1FD99F86C" box="[349,528,2006,2031]" italics="true" pageId="5" pageNumber="38">
|
||
<taxonomicName id="11EF6D9BAC00FFFD5217D8F1FE60F86C" box="[349,489,2006,2031]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="38" phylum="Chordata" rank="genus">Crenicichla</taxonomicName>
|
||
hu
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="11EF6D9BAC00FFFD51F8D8F1FCC6F86C" box="[690,847,2006,2031]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="38" phylum="Chordata" rank="species" species="jupiaensis">
|
||
<emphasis id="E49BCA0AAC00FFFD51F8D8F1FCC6F86C" box="[690,847,2006,2031]" italics="true" pageId="5" pageNumber="38">C. jupiaensis</emphasis>
|
||
</taxonomicName>
|
||
in the absence vs. presence of numerous narrow vertical bars on the flank, a well developed (but composed of spots) suborbital stripe vs. reduced to a few spots posteriorly to the orbit, a cheek bearing 4 to 6 scale rows vs. a naked cheek, and the absence vs. presence of a thin black line on the posterior margin of the preopercle.
|
||
<emphasis id="E49BCA0AAC03FFFE50CFDFC5FBB1FF78" box="[901,1080,226,251]" italics="true" pageId="6" pageNumber="39">
|
||
<taxonomicName id="11EF6D9BAC03FFFE50CFDFC5FB9BFF78" box="[901,1042,226,251]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="39" phylum="Chordata" rank="genus">Crenicichla</taxonomicName>
|
||
hu
|
||
</emphasis>
|
||
lacks several regular parallel rows of small dark spots vs. present in
|
||
<taxonomicName id="11EF6D9BAC03FFFE5121DE2FFD5EFEA2" box="[619,727,264,289]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="39" phylum="Chordata" rank="species" species="scottii">
|
||
<emphasis id="E49BCA0AAC03FFFE5121DE2FFD5EFEA2" box="[619,727,264,289]" italics="true" pageId="6" pageNumber="39">C. scottii</emphasis>
|
||
</taxonomicName>
|
||
. The new species is distinguished from
|
||
<taxonomicName id="11EF6D9BAC03FFFE57FDDE2FFAE6FEA2" box="[1207,1391,264,289]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="39" phylum="Chordata" rank="species" species="semifasciata">
|
||
<emphasis id="E49BCA0AAC03FFFE57FDDE2FFAE6FEA2" box="[1207,1391,264,289]" italics="true" pageId="6" pageNumber="39">C. semifasciata</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="11EF6D9BAC03FFFE56CBDE2FFE81FEC5" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="39" phylum="Chordata" rank="species" species="reticulata">
|
||
<emphasis id="E49BCA0AAC03FFFE56CBDE2FFE81FEC5" italics="true" pageId="6" pageNumber="39">C. reticulata</emphasis>
|
||
</taxonomicName>
|
||
group) by having about half of the caudal fin scaled vs. this fin scaled over most of its surface.
|
||
<emphasis id="E49BCA0AAC03FFFE5617DE0AFA12FEC5" box="[1373,1435,301,326]" italics="true" pageId="6" pageNumber="39">C. hu</emphasis>
|
||
further has the ascending arm of the premaxilla longer than the dentigerous one vs. shorter in
|
||
<taxonomicName id="11EF6D9BAC03FFFE57AADE75FA1DFEE8" box="[1248,1428,338,363]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="39" phylum="Chordata" rank="species" species="semifasciata">
|
||
<emphasis id="E49BCA0AAC03FFFE57AADE75FA1DFEE8" box="[1248,1428,338,363]" italics="true" pageId="6" pageNumber="39">C. semifasciata</emphasis>
|
||
</taxonomicName>
|
||
. Finally,
|
||
<emphasis id="E49BCA0AAC03FFFE53BDDE5FFEB1FE12" box="[247,312,376,401]" italics="true" pageId="6" pageNumber="39">C. hu</emphasis>
|
||
is distinguished from
|
||
<taxonomicName id="11EF6D9BAC03FFFE510BDE5FFD14FE12" box="[577,669,376,401]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="39" phylum="Chordata" rank="species" species="yaha">
|
||
<emphasis id="E49BCA0AAC03FFFE510BDE5FFD14FE12" box="[577,669,376,401]" italics="true" pageId="6" pageNumber="39">C. yaha</emphasis>
|
||
</taxonomicName>
|
||
by the head depth 17.9–20.8% vs. 15.1–18.1% of SL, and lower jaw slightly prognathous vs. jaws isognathous or upper jaw slightly prognathous.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="82904690AC03FFFE53DDDCB6FA1EFC2A" box="[151,1431,913,937]" httpUri="https://zenodo.org/record/196657/files/figure.png" pageId="6" pageNumber="39" targetBox="[194,1391,521,856]" targetPageId="6">
|
||
<paragraph id="D6501618AC03FFFE53DDDCB6FA1EFC2A" blockId="6.[151,1431,913,937]" box="[151,1431,913,937]" pageId="6" pageNumber="39">
|
||
<emphasis id="E49BCA0AAC03FFFE53DDDCB6FE90FC2A" bold="true" box="[151,281,913,937]" pageId="6" pageNumber="39">FIGURE 2.</emphasis>
|
||
<emphasis id="E49BCA0AAC03FFFE526ADCB5FE37FC2A" box="[288,446,914,937]" italics="true" pageId="6" pageNumber="39">
|
||
<taxonomicName id="11EF6D9BAC03FFFE526ADCB5FE14FC2A" box="[288,413,914,937]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="39" phylum="Chordata" rank="genus">Crenicichla</taxonomicName>
|
||
hu
|
||
</emphasis>
|
||
, female, holotype, MACN-ict 9429, 118.0 mm, arroyo Piray-Miní, 26°20'00"S 53°52'30"W.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="82904690AC03FFFE53DDD996FBB6F94B" box="[151,1087,1713,1737]" httpUri="https://zenodo.org/record/196658/files/figure.png" pageId="6" pageNumber="39" targetBox="[151,1435,988,1691]" targetPageId="6">
|
||
<paragraph id="D6501618AC03FFFE53DDD996FBB6F94B" blockId="6.[151,1087,1713,1737]" box="[151,1087,1713,1737]" pageId="6" pageNumber="39">
|
||
<emphasis id="E49BCA0AAC03FFFE53DDD996FE90F94A" bold="true" box="[151,281,1713,1737]" pageId="6" pageNumber="39">FIGURE 3.</emphasis>
|
||
<emphasis id="E49BCA0AAC03FFFE526AD996FE37F94B" box="[288,446,1713,1736]" italics="true" pageId="6" pageNumber="39">
|
||
<taxonomicName id="11EF6D9BAC03FFFE526AD996FE14F94B" box="[288,413,1713,1736]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="39" phylum="Chordata" rank="genus">Crenicichla</taxonomicName>
|
||
hu
|
||
</emphasis>
|
||
male, live specimen, paratype, MACN-ict 9430, 153.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="9EF54593AC03FFFF538FD9D1FD8FF8DF" lastPageId="7" lastPageNumber="40" pageId="6" pageNumber="39" type="description">
|
||
<paragraph id="D6501618AC03FFFE538FD9D1FA14F873" blockId="6.[151,1437,1782,2032]" pageId="6" pageNumber="39">
|
||
<emphasis id="E49BCA0AAC03FFFE538FD9D1FED4F893" bold="true" box="[197,349,1782,1808]" pageId="6" pageNumber="39">Description.</emphasis>
|
||
Morphometric data of the
|
||
<typeStatus id="0954A8BAAC03FFFE51D6D9D1FC8AF893" box="[668,771,1782,1808]" pageId="6" pageNumber="39" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="0954A8BAAC03FFFE5077D9D1FC27F893" box="[829,942,1782,1808]" pageId="6" pageNumber="39" type="paratype">paratypes</typeStatus>
|
||
is given in
|
||
<tableCitation id="9B6D23A3AC03FFFE5770D9D1FB1AF893" box="[1082,1171,1782,1808]" captionStart="TABLE 3" captionStartId="7.[151,242,855,879]" captionTargetBox="[151,1436,938,1388]" captionTargetPageId="7" captionText="TABLE 3. Proportional measurements in percents of Standard length of the holotype and 21 paratypes of the new species Crenicichla hu. SD = standard deviation." httpUri="http://table.plazi.org/id/82904690AC02FFFF53DDDC70FD1AFC0C" pageId="6" pageNumber="39" tableUuid="82904690AC02FFFF53DDDC70FD1AFC0C">Table 3</tableCitation>
|
||
. Body elongate, depth 21.5 to 25.6% of SL (
|
||
<figureCitation id="4ED40A9DAC03FFFE52DCD83BFE56F8B5" box="[406,479,1820,1846]" captionStart="FIGURE 2" captionStartId="6.[151,255,913,937]" captionTargetBox="[194,1391,521,856]" captionTargetId="figure@6.[180,1406,493,892]" captionTargetPageId="6" captionText="FIGURE 2. Crenicichla hu, female, holotype, MACN-ict 9429, 118.0 mm, arroyo Piray-Miní, 26 ° 20 ' 00 " S 53 ° 52 ' 30 " W." httpUri="https://zenodo.org/record/196657/files/figure.png" pageId="6" pageNumber="39">Fig. 2</figureCitation>
|
||
). Head slightly deeper than wide. Snout short, bluntly pointed in lateral view, 2.5 to 3.0 times in HL. Lower jaw slightly prognathous. Tip of maxilla not reaching anterior margin of orbit in most specimens (reaching in four specimens, MACN-ict 9429). Lower lip widely interrupted medially. Nostrils dorsolateral, nearer anterior margin of orbit than snout tip. Posterior margin of preopercle weakly serrated (21 ex.*) or smooth (3 ex., MACN-ict 9430). Scales on flank strongly ctenoid. Head scales cycloid. Predorsal scales small, superficially embedded in skin. Prepelvic scales smaller than predorsal ones.
|
||
</paragraph>
|
||
<paragraph id="D6501618AC02FFFF53DDDFBFFC45FCAF" blockId="7.[151,1437,152,812]" pageId="7" pageNumber="40">
|
||
Interopercle naked. Cheek scaled, 4 to 6 scales below eye embedded in skin. Scales in E1 row 47(1), 51(2), 52(4), 53(5), 54(9*). Scales in transverse row 10/14(1), 11/13(1), 11/14(7), 11/15(2), 11/16(3*), 11/17(3), 12/ 13(1), 12/14(2), 12/15(1). Three scale rows between lateral lines. Upper lateral line scales 18(1), 19(1), 21(3), 22(8), 23(4*), 24(2), 25(2). Lower lateral line scales 10(1), 11(7), 12(5), 13(1), 14(5), 15(2*). Dorsal, anal, pectoral and pelvic fins naked. Dorsal fin XVIII,10(1); XX,12(2); XX,13(1); XXI,10(3*); XXI,11(8); XXI, 12(4); XXII,11(1). Anal fin II,10(1); III,8(2); III,9(14*); III,10(3). Pectoral fin 15(10*), 16(11). Caudal-fin squamation extending almost to middle of fin in larger specimens, no more than the basal third of caudal fin in smaller ones. Soft-dorsal fin rounded or pointed tip, surpassing caudal-fin base. Tip of anal fin reaching caudal-fin base (not reaching in three specimens, AI 261 and MACN-ict 9429). Caudal fin rounded. Pectoral fin rounded, almost reaching the tip of pelvic fin. Microbranchiospines present on second through fourth gill arches. Gill rakers externally on first gill arch: 1 on epibranchial, 1 on angle, and 8 on ceratobranchial. Three to five patches of unicuspid teeth on fourth ceratobranchial. Lower pharyngeal tooth plate with unicuspid recurved and curved crenulated bicuspid teeth, those of posterior and medial row larger than remaining ones (
|
||
<figureCitation id="4ED40A9DAC02FFFF53D5DD5AFF6EFD14" box="[159,231,637,663]" captionStart="FIGURE 5" captionStartId="8.[151,255,1481,1505]" captionTargetBox="[509,1082,1035,1451]" captionTargetId="figure@8.[501,1085,1025,1460]" captionTargetPageId="8" captionText="FIGURE 5. Crenicichla hu, lower pharyngeal tooth plate in occlusal view, AI 262, 93.9 mm SL. Scale bar: 1 mm." httpUri="https://zenodo.org/record/196660/files/figure.png" pageId="7" pageNumber="40">Fig. 5</figureCitation>
|
||
). Upper pharyngeal tooth plate with unicuspid and bicuspid teeth. Frashed zone bearing one concavity with small unicuspid teeth. Premaxillary ascending process longer than dentigerous one. Premaxilla with 24(1) unicuspid teeth on outer row, larger than inner ones. Five teeth rows near symphysis. Dentary with 25(1) unicuspid teeth on outer row, 4 rows near symphysis. Total vertebrae 35 (1 C&S ex.). Premaxillary and dentary outer row teeth slightly movable, inner ones fully depressible.
|
||
</paragraph>
|
||
<caption id="82904690AC02FFFF53DDDC70FD1AFC0C" ID-Table-UUID="82904690AC02FFFF53DDDC70FD1AFC0C" httpUri="http://table.plazi.org/id/82904690AC02FFFF53DDDC70FD1AFC0C" pageId="7" pageNumber="40" targetBox="[151,1436,938,1388]" targetIsTable="true" targetPageId="7">
|
||
<paragraph id="D6501618AC02FFFF53DDDC70FD1AFC0C" blockId="7.[151,1436,855,911]" pageId="7" pageNumber="40">
|
||
<emphasis id="E49BCA0AAC02FFFF53DDDC70FE86FCEC" bold="true" box="[151,271,855,879]" pageId="7" pageNumber="40">TABLE 3.</emphasis>
|
||
Proportional measurements in percents of Standard length of the holotype and 21 paratypes of the new species
|
||
<emphasis id="E49BCA0AAC02FFFF53A0DC5FFE01FC0C" box="[234,392,888,911]" italics="true" pageId="7" pageNumber="40">
|
||
<taxonomicName id="11EF6D9BAC02FFFF53A0DC5FFEEEFC0C" box="[234,359,888,911]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="40" phylum="Chordata" rank="genus">Crenicichla</taxonomicName>
|
||
hu
|
||
</emphasis>
|
||
. SD=standard deviation.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D6501618AC02FFFF513DDC8DFAA3FAEF" pageId="7" pageNumber="40">
|
||
<table id="A4EFE4B8AC02000753DDDC8DFA15FAEF" box="[151,1436,938,1388]" gridcols="3" gridrows="8" pageId="7" pageNumber="40">
|
||
<tr id="68DF145AAC02000753DDDC8DFA15FC42" box="[151,1436,938,961]" gridrow="0" pageId="7" pageNumber="40" rowspan-0="1">
|
||
<th id="2B0E7D26AC020007513DDC8DFC12FC42" box="[631,923,938,961]" gridcol="1" gridrow="0" pageId="7" pageNumber="40">Holotype Range</th>
|
||
<th id="2B0E7D26AC0200075705DC8DFA15FC42" box="[1103,1436,938,961]" gridcol="2" gridrow="0" pageId="7" pageNumber="40">Mean SD</th>
|
||
</tr>
|
||
<tr id="68DF145AAC02000753DDDCF2FA15FC6F" box="[151,1436,981,1004]" gridrow="1" pageId="7" pageNumber="40" rowspan-2="1">
|
||
<th id="2B0E7D26AC02000753DDDCF2FE12FC6F" box="[151,411,981,1004]" gridcol="0" gridrow="1" pageId="7" pageNumber="40">Standard length (mm)</th>
|
||
<td id="2B0E7D26AC020007513DDCF2FC12FC6F" box="[631,923,981,1004]" gridcol="1" gridrow="1" pageId="7" pageNumber="40">118.0 76.9–153.0</td>
|
||
</tr>
|
||
<tr id="68DF145AAC02000753DDDB27FA15FBC2" box="[151,1436,1024,1089]" gridrow="2" pageId="7" pageNumber="40">
|
||
<th id="2B0E7D26AC02000753DDDB27FE12FBC2" box="[151,411,1024,1089]" gridcol="0" gridrow="2" pageId="7" pageNumber="40">Head length Snout length</th>
|
||
<td id="2B0E7D26AC020007513DDB27FC12FBC2" box="[631,923,1024,1089]" gridcol="1" gridrow="2" pageId="7" pageNumber="40">33.7 31.0–35.4 12.5 10.7–13.4</td>
|
||
<td id="2B0E7D26AC0200075705DB27FA15FBC2" box="[1103,1436,1024,1089]" gridcol="2" gridrow="2" pageId="7" pageNumber="40">33.4 1.18 12.0 0.75</td>
|
||
</tr>
|
||
<tr id="68DF145AAC02000753DDDB72FA15FBEF" box="[151,1436,1109,1132]" gridrow="3" pageId="7" pageNumber="40">
|
||
<th id="2B0E7D26AC02000753DDDB72FE12FBEF" box="[151,411,1109,1132]" gridcol="0" gridrow="3" pageId="7" pageNumber="40">Head depth</th>
|
||
<td id="2B0E7D26AC020007513DDB72FC12FBEF" box="[631,923,1109,1132]" gridcol="1" gridrow="3" pageId="7" pageNumber="40">16.9 15.1–18.1</td>
|
||
<td id="2B0E7D26AC0200075705DB72FA15FBEF" box="[1103,1436,1109,1132]" gridcol="2" gridrow="3" pageId="7" pageNumber="40">16.2 0.74</td>
|
||
</tr>
|
||
<tr id="68DF145AAC02000753DDDBA7FA15FB42" box="[151,1436,1152,1217]" gridrow="4" pageId="7" pageNumber="40">
|
||
<th id="2B0E7D26AC02000753DDDBA7FE12FB42" box="[151,411,1152,1217]" gridcol="0" gridrow="4" pageId="7" pageNumber="40">Body depth Orbital diameter</th>
|
||
<td id="2B0E7D26AC020007513DDBA7FC12FB42" box="[631,923,1152,1217]" gridcol="1" gridrow="4" pageId="7" pageNumber="40">25.3 21.5–25.6 5.8 5.0–7.3</td>
|
||
<td id="2B0E7D26AC0200075705DBA7FA15FB42" box="[1103,1436,1152,1217]" gridcol="2" gridrow="4" pageId="7" pageNumber="40">23.2 1.24 6.2 0.54</td>
|
||
</tr>
|
||
<tr id="68DF145AAC02000753DDDBF2FA15FB6F" box="[151,1436,1237,1260]" gridrow="5" pageId="7" pageNumber="40">
|
||
<th id="2B0E7D26AC02000753DDDBF2FE12FB6F" box="[151,411,1237,1260]" gridcol="0" gridrow="5" pageId="7" pageNumber="40">Interorbital width</th>
|
||
<td id="2B0E7D26AC020007513DDBF2FC12FB6F" box="[631,923,1237,1260]" gridcol="1" gridrow="5" pageId="7" pageNumber="40">8.6 6.7–9.0</td>
|
||
<td id="2B0E7D26AC0200075705DBF2FA15FB6F" box="[1103,1436,1237,1260]" gridcol="2" gridrow="5" pageId="7" pageNumber="40">7.9 0.69</td>
|
||
</tr>
|
||
<tr id="68DF145AAC02000753DDDA27FA15FAC2" box="[151,1436,1280,1345]" gridrow="6" pageId="7" pageNumber="40">
|
||
<th id="2B0E7D26AC02000753DDDA27FE12FAC2" box="[151,411,1280,1345]" gridcol="0" gridrow="6" pageId="7" pageNumber="40">Pectoral fin length Caudal peduncle depth</th>
|
||
<td id="2B0E7D26AC020007513DDA27FC12FAC2" box="[631,923,1280,1345]" gridcol="1" gridrow="6" pageId="7" pageNumber="40">19.1 19.1–23.1 14.5 11.7–14.5</td>
|
||
<td id="2B0E7D26AC0200075705DA27FA15FAC2" box="[1103,1436,1280,1345]" gridcol="2" gridrow="6" pageId="7" pageNumber="40">20.5 0.92 12.9 0.60</td>
|
||
</tr>
|
||
<tr id="68DF145AAC02000753DDDA72FA15FAEF" box="[151,1436,1365,1388]" gridrow="7" pageId="7" pageNumber="40">
|
||
<th id="2B0E7D26AC02000753DDDA72FE12FAEF" box="[151,411,1365,1388]" gridcol="0" gridrow="7" pageId="7" pageNumber="40">Caudal peduncle length</th>
|
||
<td id="2B0E7D26AC020007513DDA72FC12FAEF" box="[631,923,1365,1388]" gridcol="1" gridrow="7" pageId="7" pageNumber="40">16.8 13.8–16.8</td>
|
||
<td id="2B0E7D26AC0200075705DA72FA15FAEF" box="[1103,1436,1365,1388]" gridcol="2" gridrow="7" pageId="7" pageNumber="40">15.1 0.99</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="D6501618AC02FFFF538FDA80FD8FF8DF" blockId="7.[151,1437,1447,2034]" pageId="7" pageNumber="40">
|
||
<emphasis id="E49BCA0AAC02FFFF538FDA80FE51FA42" bold="true" box="[197,472,1447,1473]" pageId="7" pageNumber="40">Coloration in alcohol.</emphasis>
|
||
Background of body deeply dark, almost black in large specimens; smaller ones (
|
||
<quantity id="1117BBFDAC02FFFF53D5DAEAFEA9FA64" box="[159,288,1485,1511]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" metricValueMax="9.5" metricValueMin="7.5" pageId="7" pageNumber="40" unit="mm" value="85.0" valueMax="95.0" valueMin="75.0">75–95 mm</quantity>
|
||
) dark brown. Deep grey preorbital stripe between anterior margin of orbit to snout tip, only visible in smaller specimens. Postorbital stripe between posterior margin of orbit to preopercle distal margin, deep grey; only visible in smaller specimens. Suborbital stripe black almost reaching ventral margin of cheek; wide (up to six dots) and fragmented. Flank with 7 to 9 black irregular blotches just below upper lateral line and reaching faintly dorsal-fin base. Posteriormost blotch extending or not onto caudal peduncle. Dorsal, anal, and caudal fins dark grey or black, dorsal and anal fins with numerous dark scattered dots on their surface, also present in caudal fin in smaller specimens. Dorsal fin (females) with an irregular color pattern formed by black and white longitudinal stripes and blotches (3 ex., AI 261 and MACN-ict 9430;
|
||
<figureCitation id="4ED40A9DAC02FFFF578CD9F5FA86F96F" box="[1222,1295,1746,1772]" captionStart="FIGURE 4" captionStartId="8.[151,255,951,975]" captionTargetBox="[151,1435,226,929]" captionTargetId="figure@8.[151,1436,226,929]" captionTargetPageId="8" captionText="FIGURE 4. Crenicichla hu female, live specimen, paratype, MACN-ict 9430, 122.0 mm." httpUri="https://zenodo.org/record/196659/files/figure.png" pageId="7" pageNumber="40">Fig. 4</figureCitation>
|
||
) or a black longitudinal stripe (sometimes reduced to a single blotch) with white margin (2 ex.*, MACN-ict 9429;
|
||
<figureCitation id="4ED40A9DAC02FFFF560DD9DFFA04F891" box="[1351,1421,1784,1810]" captionStart="FIGURE 2" captionStartId="6.[151,255,913,937]" captionTargetBox="[194,1391,521,856]" captionTargetId="figure@6.[180,1406,493,892]" captionTargetPageId="6" captionText="FIGURE 2. Crenicichla hu, female, holotype, MACN-ict 9429, 118.0 mm, arroyo Piray-Miní, 26 ° 20 ' 00 " S 53 ° 52 ' 30 " W." httpUri="https://zenodo.org/record/196657/files/figure.png" pageId="7" pageNumber="40">Fig. 2</figureCitation>
|
||
). Caudal fin with a black subcircular spot well separated from base of fin, just above midline of caudal fin. Pectoral and pelvic fins smoky.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9EF54593AC02FFF0538FD840FC7AF9ED" lastPageId="8" lastPageNumber="41" pageId="7" pageNumber="40" type="materials_examined">
|
||
<paragraph id="D6501618AC02FFFF538FD840FE7EF871" blockId="7.[151,1437,1447,2034]" pageId="7" pageNumber="40">
|
||
<emphasis id="E49BCA0AAC02FFFF538FD840FDC8F802" bold="true" box="[197,577,1895,1921]" pageId="7" pageNumber="40">Coloration in live specimens.</emphasis>
|
||
Same as color in alcohol (
|
||
<figureCitation id="4ED40A9DAC02FFFF50C4D84FFB8EF801" box="[910,1031,1896,1922]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="6.[151,255,1713,1737]" captionStartId-1="8.[151,255,951,975]" captionTargetBox-0="[151,1435,988,1691]" captionTargetBox-1="[151,1435,226,929]" captionTargetId-0="figure@6.[151,1436,988,1691]" captionTargetId-1="figure@8.[151,1436,226,929]" captionTargetPageId-0="6" captionTargetPageId-1="8" captionText-0="FIGURE 3. Crenicichla hu male, live specimen, paratype, MACN-ict 9430, 153.0 mm." captionText-1="FIGURE 4. Crenicichla hu female, live specimen, paratype, MACN-ict 9430, 122.0 mm." httpUri-0="https://zenodo.org/record/196658/files/figure.png" httpUri-1="https://zenodo.org/record/196659/files/figure.png" pageId="7" pageNumber="40">Figs. 3–4</figureCitation>
|
||
). Live specimens lack almost all carotenoid or physical reflective colors, the overall color is dark grey or dark brown to black. Some female specimens show a faint orange area behind the pectoral fin. Outline of the black areas in the dorsal fin of females milk-colored (
|
||
<figureCitation id="4ED40A9DAC02FFFF52EBD8FFFE60F871" box="[417,489,2008,2034]" captionStart="FIGURE 4" captionStartId="8.[151,255,951,975]" captionTargetBox="[151,1435,226,929]" captionTargetId="figure@8.[151,1436,226,929]" captionTargetPageId="8" captionText="FIGURE 4. Crenicichla hu female, live specimen, paratype, MACN-ict 9430, 122.0 mm." httpUri="https://zenodo.org/record/196659/files/figure.png" pageId="7" pageNumber="40">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="82904690AC0DFFF053DDDC90FBDDFC4D" box="[151,1108,951,975]" httpUri="https://zenodo.org/record/196659/files/figure.png" pageId="8" pageNumber="41" targetBox="[151,1435,226,929]" targetPageId="8">
|
||
<paragraph id="D6501618AC0DFFF053DDDC90FBDDFC4D" blockId="8.[151,1108,951,975]" box="[151,1108,951,975]" pageId="8" pageNumber="41">
|
||
<emphasis id="E49BCA0AAC0DFFF053DDDC90FE90FC4C" bold="true" box="[151,281,951,975]" pageId="8" pageNumber="41">FIGURE 4.</emphasis>
|
||
<emphasis id="E49BCA0AAC0DFFF0526ADC90FE37FC4D" box="[288,446,951,974]" italics="true" pageId="8" pageNumber="41">
|
||
<taxonomicName id="11EF6D9BAC0DFFF0526ADC90FE14FC4D" box="[288,413,951,974]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="41" phylum="Chordata" rank="genus">Crenicichla</taxonomicName>
|
||
hu
|
||
</emphasis>
|
||
female, live specimen, paratype, MACN-ict 9430, 122.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="82904690AC0DFFF053DDDAEEFAD5FA62" box="[151,1372,1481,1505]" httpUri="https://zenodo.org/record/196660/files/figure.png" pageId="8" pageNumber="41" targetBox="[509,1082,1035,1451]" targetPageId="8">
|
||
<paragraph id="D6501618AC0DFFF053DDDAEEFAD5FA62" blockId="8.[151,1372,1481,1505]" box="[151,1372,1481,1505]" pageId="8" pageNumber="41">
|
||
<emphasis id="E49BCA0AAC0DFFF053DDDAEEFE93FA62" bold="true" box="[151,282,1481,1505]" pageId="8" pageNumber="41">FIGURE 5.</emphasis>
|
||
<emphasis id="E49BCA0AAC0DFFF0526ADAEDFE37FA62" box="[288,446,1482,1505]" italics="true" pageId="8" pageNumber="41">
|
||
<taxonomicName id="11EF6D9BAC0DFFF0526ADAEDFE14FA62" box="[288,413,1482,1505]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="41" phylum="Chordata" rank="genus">Crenicichla</taxonomicName>
|
||
hu
|
||
</emphasis>
|
||
, lower pharyngeal tooth plate in occlusal view, AI 262, 93.9 mm SL. Scale bar: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D6501618AC0DFFF0538FD92EFC7AF9ED" blockId="8.[151,1436,1545,2020]" pageId="8" pageNumber="41">
|
||
<emphasis id="E49BCA0AAC0DFFF0538FD92EFE1DF9A0" bold="true" box="[197,404,1545,1571]" pageId="8" pageNumber="41">Ecological notes.</emphasis>
|
||
The arroyo Piray–Miní (the
|
||
<typeStatus id="0954A8BAAC0DFFF051A8D92DFC9CF9A7" box="[738,789,1546,1572]" pageId="8" pageNumber="41">type</typeStatus>
|
||
and only-known locality) has clear and rapidly flowing water. The depth of the arroyo Piray–Miní is variable,
|
||
<quantity id="1117BBFDAC0DFFF05050D908FC40F9CA" box="[794,969,1583,1609]" metricMagnitude="-1" metricUnit="m" metricValue="7.999999999999999" metricValueMax="14.0" metricValueMin="2.0" pageId="8" pageNumber="41" unit="m" value="0.7999999999999999" valueMax="1.4" valueMin="0.2">0.20 to 1.40 m</quantity>
|
||
. The bottom consists of mud, sand, and mostly stones. Some areas have scarce submerged vegetation (
|
||
<figureCitation id="4ED40A9DAC0DFFF0503ED973FC6FF9ED" box="[884,998,1620,1646]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="9.[151,255,912,936]" captionStartId-1="10.[151,255,1899,1923]" captionTargetBox-1="[199,1385,341,1863]" captionTargetId-1="figure@10.[187,1400,329,1878]" captionTargetPageId-1="10" captionText-0="FIGURE 6. Arroyo Piray – Miní, the type locality of Crenicichla hu." captionText-1="FIGURE 7. Hydrological map of the Province of Misiones. The type locality is marked with a solid circle." httpUri-1="https://zenodo.org/record/196661/files/figure.png" pageId="8" pageNumber="41">Figs. 6–7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9EF54593AC0DFFF0538FD95EFE00F93A" pageId="8" pageNumber="41" type="etymology">
|
||
<paragraph id="D6501618AC0DFFF0538FD95EFE00F93A" blockId="8.[151,1436,1545,2020]" pageId="8" pageNumber="41">
|
||
<emphasis id="E49BCA0AAC0DFFF0538FD95EFEDBF910" bold="true" box="[197,338,1657,1683]" pageId="8" pageNumber="41">Etymology.</emphasis>
|
||
The specific epithet hu is a
|
||
<taxonomicName id="11EF6D9BAC0DFFF051D7D95DFD72F917" box="[669,763,1658,1684]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="44" phylum="Chordata" rank="species" species="guarani">Guaraní</taxonomicName>
|
||
word
|
||
<emphasis id="E49BCA0AAC0DFFF0500DD95DFCEDF910" box="[839,868,1658,1683]" italics="true" pageId="8" pageNumber="41">hû</emphasis>
|
||
that means black in allusion to the ground color of the body and fins.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9EF54593AC0DFFF3538FD9E3FDF1FBD4" lastPageId="11" lastPageNumber="44" pageId="8" pageNumber="41" type="discussion">
|
||
<paragraph id="D6501618AC0DFFF0538FD9E3FA15F867" blockId="8.[151,1436,1545,2020]" pageId="8" pageNumber="41">
|
||
<emphasis id="E49BCA0AAC0DFFF0538FD9E3FEDDF95D" bold="true" box="[197,340,1732,1758]" pageId="8" pageNumber="41">Discussion.</emphasis>
|
||
Molecular phylogeny divides the Misioneran crenicichlas into several clades (
|
||
<figureCitation id="4ED40A9DAC0DFFF0565ED9E3FADFF95D" box="[1300,1366,1732,1758]" captionStart="FIGURE 1" captionStartId="5.[151,255,965,989]" captionTargetBox="[158,1418,179,941]" captionTargetId="figure@5.[151,1436,167,949]" captionTargetPageId="5" captionText="FIGURE 1. Maximum parsimony tree topology based on ND 2 sequences; the dotted line displays the different topology of the alternative Bayesian inference. Numbers for each recovered node represent nonparametric bootstrap support (left) and Bayesian posterior probability (right), respectively." httpUri="https://zenodo.org/record/196656/files/figure.png" pageId="8" pageNumber="41">Fig.1</figureCitation>
|
||
). The basal-most species is
|
||
<taxonomicName id="11EF6D9BAC0DFFF052D9D9CDFD9CF880" box="[403,533,1770,1795]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="41" phylum="Chordata" rank="species" species="lepidota">
|
||
<emphasis id="E49BCA0AAC0DFFF052D9D9CDFD9CF880" box="[403,533,1770,1795]" italics="true" pageId="8" pageNumber="41">C. lepidota</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="11EF6D9BAC0DFFF0516FD9CDFD21F880" box="[549,680,1770,1795]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="41" phylum="Chordata" rank="species" species="saxatilis">
|
||
<emphasis id="E49BCA0AAC0DFFF0516FD9CDFD21F880" box="[549,680,1770,1795]" italics="true" pageId="8" pageNumber="41">C. saxatilis</emphasis>
|
||
</taxonomicName>
|
||
group;
|
||
<figureCitation id="4ED40A9DAC0DFFF05049D9CDFCCEF887" box="[771,839,1770,1796]" captionStart="FIGURE 1" captionStartId="5.[151,255,965,989]" captionTargetBox="[158,1418,179,941]" captionTargetId="figure@5.[151,1436,167,949]" captionTargetPageId="5" captionText="FIGURE 1. Maximum parsimony tree topology based on ND 2 sequences; the dotted line displays the different topology of the alternative Bayesian inference. Numbers for each recovered node represent nonparametric bootstrap support (left) and Bayesian posterior probability (right), respectively." httpUri="https://zenodo.org/record/196656/files/figure.png" pageId="8" pageNumber="41">Fig. 1</figureCitation>
|
||
; tree additionaly rooted with
|
||
<taxonomicName id="11EF6D9BAC0DFFF057EAD9CDFA15F880" box="[1184,1436,1770,1795]" class="Actinopterygii" family="Cichlidae" genus="Satanoperca" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="41" phylum="Chordata" rank="species" species="jurupari">
|
||
<emphasis id="E49BCA0AAC0DFFF057EAD9CDFA15F880" box="[1184,1436,1770,1795]" italics="true" pageId="8" pageNumber="41">Satanoperca jurupari</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="11EF6D9BAC0DFFF05386D828FE41F8AB" box="[204,456,1807,1832]" class="Actinopterygii" family="Cichlidae" genus="Astronotus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="41" phylum="Chordata" rank="species" species="ocellatus">
|
||
<emphasis id="E49BCA0AAC0DFFF05386D828FE41F8AB" box="[204,456,1807,1832]" italics="true" pageId="8" pageNumber="41">Astronotus ocellatus</emphasis>
|
||
</taxonomicName>
|
||
; see Methods). The two philosophically distinct computing methods (MP, BA) inferred a robust phylogenetic hypothesis of nearly identical topologies that supports the biogeographic foundation of the recognized species groups. All species from the Misioneran part of the Paraná drainage basin (
|
||
<emphasis id="E49BCA0AAC0DFFF053ACD858FEAEF81B" box="[230,295,1919,1944]" italics="true" pageId="8" pageNumber="41">C. hu</emphasis>
|
||
,
|
||
<taxonomicName id="11EF6D9BAC0DFFF0527CD858FE1AF81B" box="[310,403,1919,1944]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="41" phylum="Chordata" rank="species" species="tesay">
|
||
<emphasis id="E49BCA0AAC0DFFF0527CD858FE1AF81B" box="[310,403,1919,1944]" italics="true" pageId="8" pageNumber="41">C. tesay</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="11EF6D9BAC0DFFF052E9D858FE76F81B" box="[419,511,1919,1944]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="41" phylum="Chordata" rank="species" species="yaha">
|
||
<emphasis id="E49BCA0AAC0DFFF052E9D858FE76F81B" box="[419,511,1919,1944]" italics="true" pageId="8" pageNumber="41">C. yaha</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="11EF6D9BAC0DFFF0510BD858FC98F81B" box="[577,785,1919,1944]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="41" phylum="Chordata" rank="species" species="mandelburgeri">
|
||
<emphasis id="E49BCA0AAC0DFFF0510BD858FC98F81B" box="[577,785,1919,1944]" italics="true" pageId="8" pageNumber="41">C. mandelburgeri</emphasis>
|
||
</taxonomicName>
|
||
) were grouped together in one clade with conclusively high support (
|
||
<figureCitation id="4ED40A9DAC0DFFF05270D883FE08F83D" box="[314,385,1956,1982]" captionStart="FIGURE 1" captionStartId="5.[151,255,965,989]" captionTargetBox="[158,1418,179,941]" captionTargetId="figure@5.[151,1436,167,949]" captionTargetPageId="5" captionText="FIGURE 1. Maximum parsimony tree topology based on ND 2 sequences; the dotted line displays the different topology of the alternative Bayesian inference. Numbers for each recovered node represent nonparametric bootstrap support (left) and Bayesian posterior probability (right), respectively." httpUri="https://zenodo.org/record/196656/files/figure.png" pageId="8" pageNumber="41">Fig. 1</figureCitation>
|
||
; Paraná endemic species; bootstrap, 96; PP, 1.00). The newly described
|
||
<emphasis id="E49BCA0AAC0DFFF05799D882FA09F83D" box="[1235,1408,1957,1982]" italics="true" pageId="8" pageNumber="41">
|
||
<taxonomicName id="11EF6D9BAC0DFFF05799D882FAD5F83D" box="[1235,1372,1957,1982]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="41" phylum="Chordata" rank="genus">Crenicichla</taxonomicName>
|
||
hu
|
||
</emphasis>
|
||
is recovered in a basal position of this clade.
|
||
<taxonomicName id="11EF6D9BAC0DFFF051C0D8EDFC4EF860" box="[650,967,1994,2019]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="41" phylum="Chordata" rank="species" species="mandelburgeri">
|
||
<emphasis id="E49BCA0AAC0DFFF051C0D8EDFC4EF860" box="[650,967,1994,2019]" italics="true" pageId="8" pageNumber="41">Crenicichla mandelburgeri</emphasis>
|
||
</taxonomicName>
|
||
appears non-monophyletic (see below).
|
||
</paragraph>
|
||
<paragraph id="D6501618AC0CFFF153DDDFBFFC34FEC4" blockId="9.[151,1436,152,327]" pageId="9" pageNumber="42">
|
||
The position of
|
||
<taxonomicName id="11EF6D9BAC0CFFF15219DFBFFE48FF32" box="[339,449,152,177]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="vittata">
|
||
<emphasis id="E49BCA0AAC0CFFF15219DFBFFE48FF32" box="[339,449,152,177]" italics="true" pageId="9" pageNumber="42">C. vittata</emphasis>
|
||
</taxonomicName>
|
||
inhabiting both the Paraná and the
|
||
<collectingCountry id="AEF85688AC0CFFF1502FDFBFFC42FF31" box="[869,971,152,178]" name="Uruguay" pageId="9" pageNumber="42">Uruguay</collectingCountry>
|
||
basins differs between the BA and MP hypotheses, in the latter forming a monophyly together with the endemic Paraná species (
|
||
<figureCitation id="4ED40A9DAC0CFFF15787DF9AFA9FFF54" box="[1229,1302,189,215]" captionStart="FIGURE 1" captionStartId="5.[151,255,965,989]" captionTargetBox="[158,1418,179,941]" captionTargetId="figure@5.[151,1436,167,949]" captionTargetPageId="5" captionText="FIGURE 1. Maximum parsimony tree topology based on ND 2 sequences; the dotted line displays the different topology of the alternative Bayesian inference. Numbers for each recovered node represent nonparametric bootstrap support (left) and Bayesian posterior probability (right), respectively." httpUri="https://zenodo.org/record/196656/files/figure.png" pageId="9" pageNumber="42">Fig. 1</figureCitation>
|
||
; bootstrap, 66). The morphologically distinct
|
||
<taxonomicName id="11EF6D9BAC0CFFF1516CDFC5FD26FF78" box="[550,687,226,251]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="genus">
|
||
<emphasis id="E49BCA0AAC0CFFF1516CDFC5FD26FF78" box="[550,687,226,251]" italics="true" pageId="9" pageNumber="42">Crenicichla</emphasis>
|
||
</taxonomicName>
|
||
species from the
|
||
<collectingCountry id="AEF85688AC0CFFF15036DFC5FC6AFF7F" box="[892,995,226,252]" name="Uruguay" pageId="9" pageNumber="42">Uruguay</collectingCountry>
|
||
basin formed two independent clades (
|
||
<figureCitation id="4ED40A9DAC0CFFF153D5DE2FFF6FFEA1" box="[159,230,264,290]" captionStart="FIGURE 1" captionStartId="5.[151,255,965,989]" captionTargetBox="[158,1418,179,941]" captionTargetId="figure@5.[151,1436,167,949]" captionTargetPageId="5" captionText="FIGURE 1. Maximum parsimony tree topology based on ND 2 sequences; the dotted line displays the different topology of the alternative Bayesian inference. Numbers for each recovered node represent nonparametric bootstrap support (left) and Bayesian posterior probability (right), respectively." httpUri="https://zenodo.org/record/196656/files/figure.png" pageId="9" pageNumber="42">Fig. 1</figureCitation>
|
||
;
|
||
<taxonomicName id="11EF6D9BAC0CFFF153BCDE2FFE2AFEA2" box="[246,419,264,289]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="missioneira">
|
||
<emphasis id="E49BCA0AAC0CFFF153BCDE2FFE2AFEA2" box="[246,419,264,289]" italics="true" pageId="9" pageNumber="42">C. missioneira</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="11EF6D9BAC0CFFF15294DE2FFDC0FEA2" box="[478,585,264,289]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="scottii">
|
||
<emphasis id="E49BCA0AAC0CFFF15294DE2FFDC0FEA2" box="[478,585,264,289]" italics="true" pageId="9" pageNumber="42">C. scottii</emphasis>
|
||
</taxonomicName>
|
||
groups). Two species from the coastal rivers (
|
||
<emphasis id="E49BCA0AAC0CFFF1573ADE2FFA1BFEA2" box="[1136,1426,264,289]" italics="true" pageId="9" pageNumber="42">
|
||
<taxonomicName id="11EF6D9BAC0CFFF1573ADE2FFB72FEA2" box="[1136,1275,264,289]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="lacustris">C. lacustris</taxonomicName>
|
||
,
|
||
<taxonomicName id="11EF6D9BAC0CFFF15642DE2FFA1BFEA2" box="[1288,1426,264,289]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="punctata">C. punctata</taxonomicName>
|
||
</emphasis>
|
||
) do not form a monophyletic lineage in either of the two phylogenies.
|
||
</paragraph>
|
||
<caption id="82904690AC0CFFF153DDDCB7FCE6FC24" box="[151,879,912,936]" pageId="9" pageNumber="42">
|
||
<paragraph id="D6501618AC0CFFF153DDDCB7FCE6FC24" blockId="9.[151,879,912,936]" box="[151,879,912,936]" pageId="9" pageNumber="42">
|
||
<emphasis id="E49BCA0AAC0CFFF153DDDCB7FE93FC2B" bold="true" box="[151,282,912,936]" pageId="9" pageNumber="42">FIGURE 6.</emphasis>
|
||
Arroyo Piray–Miní, the type locality of
|
||
<emphasis id="E49BCA0AAC0CFFF15183DCB6FCEEFC2B" box="[713,871,913,936]" italics="true" pageId="9" pageNumber="42">
|
||
<taxonomicName id="11EF6D9BAC0CFFF15183DCB6FCCFFC2B" box="[713,838,913,936]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="genus">Crenicichla</taxonomicName>
|
||
hu
|
||
</emphasis>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D6501618AC0CFFF1538FDCF1FA1FFB28" blockId="9.[151,1437,982,2016]" pageId="9" pageNumber="42">
|
||
The phylogeny of a few species of
|
||
<taxonomicName id="11EF6D9BAC0CFFF15139DCF1FC88FC6C" box="[627,769,982,1007]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="genus">
|
||
<emphasis id="E49BCA0AAC0CFFF15139DCF1FC88FC6C" box="[627,769,982,1007]" italics="true" pageId="9" pageNumber="42">Crenicichla</emphasis>
|
||
</taxonomicName>
|
||
from southern South
|
||
<collectingCountry id="AEF85688AC0CFFF15746DCF1FBFDFC73" box="[1036,1140,982,1008]" name="United States of America" pageId="9" pageNumber="42">America</collectingCountry>
|
||
was recently studied by Kullander
|
||
<emphasis id="E49BCA0AAC0CFFF1525EDCDCFEC7FB97" box="[276,334,1019,1044]" italics="true" pageId="9" pageNumber="42">et al.</emphasis>
|
||
(in press). That study supports virtually the same relationships between the above-mentioned clades. The endemic Paraná species group was represented in their analysis by only one taxon (
|
||
<taxonomicName id="11EF6D9BAC0CFFF156CADB06FEA2FBDC" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="iguassuensis">
|
||
<emphasis id="E49BCA0AAC0CFFF156CADB06FEA2FBDC" italics="true" pageId="9" pageNumber="42">C. iguassuensis</emphasis>
|
||
</taxonomicName>
|
||
) as were the coastal-river drainages (
|
||
<taxonomicName id="11EF6D9BAC0CFFF151A9DB61FCE5FBDC" box="[739,876,1094,1119]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="punctata">
|
||
<emphasis id="E49BCA0AAC0CFFF151A9DB61FCE5FBDC" box="[739,876,1094,1119]" italics="true" pageId="9" pageNumber="42">C. punctata</emphasis>
|
||
</taxonomicName>
|
||
). The
|
||
<collectingCountry id="AEF85688AC0CFFF150F0DB61FBA9FBE3" box="[954,1056,1094,1120]" name="Uruguay" pageId="9" pageNumber="42">Uruguay</collectingCountry>
|
||
basin was represented by the
|
||
<taxonomicName id="11EF6D9BAC0CFFF156CBDB61FF6DFB07" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="scottii">
|
||
<emphasis id="E49BCA0AAC0CFFF156CBDB61FF6DFB07" italics="true" pageId="9" pageNumber="42">C. scottii</emphasis>
|
||
</taxonomicName>
|
||
group (
|
||
<taxonomicName id="11EF6D9BAC0CFFF15202DB4CFE33FB07" box="[328,442,1131,1156]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="scottii">
|
||
<emphasis id="E49BCA0AAC0CFFF15202DB4CFE33FB07" box="[328,442,1131,1156]" italics="true" pageId="9" pageNumber="42">C. scottii</emphasis>
|
||
</taxonomicName>
|
||
) and the
|
||
<taxonomicName id="11EF6D9BAC0CFFF1517BDB4CFD6EFB07" box="[561,743,1131,1156]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="missioneira">
|
||
<emphasis id="E49BCA0AAC0CFFF1517BDB4CFD6EFB07" box="[561,743,1131,1156]" italics="true" pageId="9" pageNumber="42">C. missioneira</emphasis>
|
||
</taxonomicName>
|
||
group (
|
||
<taxonomicName id="11EF6D9BAC0CFFF15001DB4CFB88FB07" box="[843,1025,1131,1156]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="missioneira">
|
||
<emphasis id="E49BCA0AAC0CFFF15001DB4CFB88FB07" box="[843,1025,1131,1156]" italics="true" pageId="9" pageNumber="42">C. missioneira</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="11EF6D9BAC0CFFF15758DB4CFB2BFB07" box="[1042,1186,1131,1156]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="minuano">
|
||
<emphasis id="E49BCA0AAC0CFFF15758DB4CFB2BFB07" box="[1042,1186,1131,1156]" italics="true" pageId="9" pageNumber="42">C. minuano</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="11EF6D9BAC0CFFF157FEDB4CFAE6FB07" box="[1204,1391,1131,1156]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="celidochilus">
|
||
<emphasis id="E49BCA0AAC0CFFF157FEDB4CFAE6FB07" box="[1204,1391,1131,1156]" italics="true" pageId="9" pageNumber="42">C. celidochilus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="11EF6D9BAC0CFFF156CADB4CFE8FFB29" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="empheres">
|
||
<emphasis id="E49BCA0AAC0CFFF156CADB4CFE8FFB29" italics="true" pageId="9" pageNumber="42">C. empheres</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="11EF6D9BAC0CFFF1525EDBB6FE6CFB29" box="[276,485,1169,1194]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="tendybaguassu">
|
||
<emphasis id="E49BCA0AAC0CFFF1525EDBB6FE6CFB29" box="[276,485,1169,1194]" italics="true" pageId="9" pageNumber="42">C. tendybaguassu</emphasis>
|
||
</taxonomicName>
|
||
). There is thus no overlap with our taxon sampling of the endemic Paraná clade.
|
||
</paragraph>
|
||
<paragraph id="D6501618AC0CFFF1538FDB91FD1AFA78" blockId="9.[151,1437,982,2016]" pageId="9" pageNumber="42">
|
||
Evaluating the uncorrected pairwise divergences between the gene sequences of
|
||
<emphasis id="E49BCA0AAC0CFFF157C1DB91FAB4FB4C" box="[1163,1341,1206,1231]" italics="true" pageId="9" pageNumber="42">
|
||
<taxonomicName id="11EF6D9BAC0CFFF157C1DB91FA9FFB4C" box="[1163,1302,1206,1231]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="genus">Crenicichla</taxonomicName>
|
||
hu
|
||
</emphasis>
|
||
and the other species, the supposed higher evolutionary rate in the
|
||
<taxonomicName id="11EF6D9BAC0CFFF15025DBFCFC74FB76" box="[879,1021,1243,1269]" class="Actinopterygii" family="Cichlidae" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="tribe" tribe="Geophagini">Geophagini</taxonomicName>
|
||
tribe of cichlids must be taken in account (
|
||
<bibRefCitation id="B27E6BE9AC0CFFF15246DA26FE7CFA98" author="Farias" box="[268,501,1281,1307]" pageId="9" pageNumber="45" refString="Farias, I. P., Orti, G., Sampaio, I., Schneider, H. & Meyer, A. (1999) Mitochondrial DNA phylogeny of the family Cichlidae: monophyly and fast molecular evolution of the Neotropical assemblage. Journal of Molecular Evolution, 48, 703 - 711." type="journal article" year="1999">
|
||
Farias
|
||
<emphasis id="E49BCA0AAC0CFFF1522CDA26FE21FA99" box="[358,424,1281,1306]" italics="true" pageId="9" pageNumber="42">et al.</emphasis>
|
||
1999
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="B27E6BE9AC0CFFF15148DA26FDCFFA98" author="Farias" box="[514,582,1281,1307]" pageId="9" pageNumber="45" refString="Farias, I. P., Orti, G. & Meyer, A. (2000) Total evidence: molecules, morphology, and the phylogenetics of cichlid fishes. Journal of Experimental Zoology, 288, 76 - 92." type="journal article" year="2000">2000</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="B27E6BE9AC0CFFF1511EDA26FC0AFA98" author="Pereyra" box="[596,899,1281,1307]" pageId="9" pageNumber="46" refString="Pereyra, S. & Garcia, G. (2008) Patterns of genetic differentiation in the Gymnogeophagus gymnogenys species complex, a neotropical cichlid from South American basins. Environmental Biology of Fishes, 83, 245 - 257." type="journal article" year="2008">Pereyra & García 2008</bibRefCitation>
|
||
). Referring to ND2 sequences, the lowest divergence between a haplotype of
|
||
<emphasis id="E49BCA0AAC0CFFF15109DA01FD0DFABC" box="[579,644,1318,1343]" italics="true" pageId="9" pageNumber="42">C. hu</emphasis>
|
||
and a haplotype of the nearest species (
|
||
<taxonomicName id="11EF6D9BAC0CFFF15722DA01FAB5FABC" box="[1128,1340,1318,1343]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="mandelburgeri">
|
||
<emphasis id="E49BCA0AAC0CFFF15722DA01FAB5FABC" box="[1128,1340,1318,1343]" italics="true" pageId="9" pageNumber="42">C. mandelburgeri</emphasis>
|
||
</taxonomicName>
|
||
C44) is 3.3% (Table 2). On the other hand, substantially lower values of divergences between formerly described species can be found (
|
||
<emphasis id="E49BCA0AAC0CFFF152D5DA56FE40FA09" box="[415,457,1393,1418]" italics="true" pageId="9" pageNumber="42">e.g.</emphasis>
|
||
,
|
||
<taxonomicName id="11EF6D9BAC0CFFF15293DA56FD0CFA09" box="[473,645,1393,1418]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="missioneira">
|
||
<emphasis id="E49BCA0AAC0CFFF15293DA56FD0CFA09" box="[473,645,1393,1418]" italics="true" pageId="9" pageNumber="42">C. missioneira</emphasis>
|
||
</taxonomicName>
|
||
C36 vs.
|
||
<taxonomicName id="11EF6D9BAC0CFFF151BADA56FCF3FA09" box="[752,890,1393,1418]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="minuano">
|
||
<emphasis id="E49BCA0AAC0CFFF151BADA56FCF3FA09" box="[752,890,1393,1418]" italics="true" pageId="9" pageNumber="42">C. minuano</emphasis>
|
||
</taxonomicName>
|
||
C80, 0.6%;
|
||
<taxonomicName id="11EF6D9BAC0CFFF1575ADA56FBE6FA09" box="[1040,1135,1393,1418]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="tesay">
|
||
<emphasis id="E49BCA0AAC0CFFF1575ADA56FBE6FA09" box="[1040,1135,1393,1418]" italics="true" pageId="9" pageNumber="42">C. tesay</emphasis>
|
||
</taxonomicName>
|
||
C1 vs.
|
||
<taxonomicName id="11EF6D9BAC0CFFF15781DA56FA15FA09" box="[1227,1436,1393,1418]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="mandelburgeri">
|
||
<emphasis id="E49BCA0AAC0CFFF15781DA56FA15FA09" box="[1227,1436,1393,1418]" italics="true" pageId="9" pageNumber="42">C. mandelburgeri</emphasis>
|
||
</taxonomicName>
|
||
C51, 1.0%;
|
||
<taxonomicName id="11EF6D9BAC0CFFF15264DAB1FE18FA2C" box="[302,401,1430,1455]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="yaha">
|
||
<emphasis id="E49BCA0AAC0CFFF15264DAB1FE18FA2C" box="[302,401,1430,1455]" italics="true" pageId="9" pageNumber="42">C. yaha</emphasis>
|
||
</taxonomicName>
|
||
C5 vs.
|
||
<taxonomicName id="11EF6D9BAC0CFFF152BEDAB1FD5AFA2C" box="[500,723,1430,1455]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="mandelburgeri">
|
||
<emphasis id="E49BCA0AAC0CFFF152BEDAB1FD5AFA2C" box="[500,723,1430,1455]" italics="true" pageId="9" pageNumber="42">C. mandelburgeri</emphasis>
|
||
</taxonomicName>
|
||
C44, 1.0%). Despite the little-known divergence rate in
|
||
<taxonomicName id="11EF6D9BAC0CFFF153DDDA9CFEA8FA56" box="[151,289,1467,1493]" class="Actinopterygii" family="Cichlidae" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="tribe" tribe="Geophagini">Geophagini</taxonomicName>
|
||
(
|
||
<bibRefCitation id="B27E6BE9AC0CFFF1527ADA9CFD05FA56" author="Lopez-Fernandez" box="[304,652,1467,1493]" lastPageId="9" lastPageNumber="46" pageId="9" pageNumber="45" refString="Lopez-Fernandez, H., Honeycutt, R. L. & Winemiller, K. O. (2005) Molecular phylogeny and evidence for an adaptive radiation of geophagine cichlids from South America (Perciformes: Labroidei). Molecular Phylogenetics and Evolution, 34, 227 - 244." type="journal article" year="2005">
|
||
López-Fernández
|
||
<emphasis id="E49BCA0AAC0CFFF15143DA9CFDCCFA57" box="[521,581,1467,1492]" italics="true" pageId="9" pageNumber="42">et al.</emphasis>
|
||
2005
|
||
</bibRefCitation>
|
||
), molecular divergences between the newly described species and its phylogenetic neighbours are substantial.
|
||
</paragraph>
|
||
<paragraph id="D6501618AC0CFFF1538FD921FB30F883" blockId="9.[151,1437,982,2016]" pageId="9" pageNumber="42">
|
||
With the newly described
|
||
<emphasis id="E49BCA0AAC0CFFF1514ED921FDCFF99C" box="[516,582,1542,1567]" italics="true" pageId="9" pageNumber="42">C. hu</emphasis>
|
||
, fourteen species of
|
||
<taxonomicName id="11EF6D9BAC0CFFF1500ED921FC58F99C" box="[836,977,1542,1567]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="genus">
|
||
<emphasis id="E49BCA0AAC0CFFF1500ED921FC58F99C" box="[836,977,1542,1567]" italics="true" pageId="9" pageNumber="42">Crenicichla</emphasis>
|
||
</taxonomicName>
|
||
have now been recorded from the río Paraná basin. Some of them (i.e.
|
||
<emphasis id="E49BCA0AAC0CFFF1515CD90CFC97F9C7" box="[534,798,1579,1604]" italics="true" pageId="9" pageNumber="42">
|
||
<taxonomicName id="11EF6D9BAC0CFFF1515CD90CFD07F9C7" box="[534,654,1579,1604]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="britskii">C. britskii</taxonomicName>
|
||
,
|
||
<taxonomicName id="11EF6D9BAC0CFFF151D3D90CFC97F9C7" box="[665,798,1579,1604]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="haroldoi">C. haroldoi</taxonomicName>
|
||
</emphasis>
|
||
, and
|
||
<taxonomicName id="11EF6D9BAC0CFFF15017D90CFB8FF9C7" box="[861,1030,1579,1604]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="jaguarensis">
|
||
<emphasis id="E49BCA0AAC0CFFF15017D90CFB8FF9C7" box="[861,1030,1579,1604]" italics="true" pageId="9" pageNumber="42">C. jaguarensis</emphasis>
|
||
</taxonomicName>
|
||
) are restricted to the Upper Paraná basin (
|
||
<bibRefCitation id="B27E6BE9AC0CFFF153AED976FE07F9E8" author="Resende" box="[228,398,1617,1643]" pageId="9" pageNumber="46" refString="Resende, E. K. (2003) Migratory Fishes of the Paraguay - Parana Basin, Excluding the Upper Parana Basin. In: Carolsfeld, J., Harvey, B., Ross, C. & Baer, A. (Eds.), Migratory fishes of South America: Biology, Fisheries and Conservation Status. International Development Research Center and The World Bank, pp. 99 - 156." type="book chapter" year="2003">Resende 2003</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="B27E6BE9AC0CFFF152D0D976FDDFF9E8" author="Reis" box="[410,598,1617,1643]" pageId="9" pageNumber="46" refString="Reis, R. E., Kullander, S. O. & Ferraris Jr., C. J. (2003) Check list of the freshwater fishes of South and Central America. Edipucrs, Porto Alegre, Brazil, 729 pp." type="book" year="2003">
|
||
Reis
|
||
<emphasis id="E49BCA0AAC0CFFF1529FD976FD86F9E9" box="[469,527,1617,1642]" italics="true" pageId="9" pageNumber="42">et al.</emphasis>
|
||
2003
|
||
</bibRefCitation>
|
||
).
|
||
<taxonomicName id="11EF6D9BAC0CFFF15123D976FCF9F9E9" box="[617,880,1617,1642]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="jupiaensis">
|
||
<emphasis id="E49BCA0AAC0CFFF15123D976FCF9F9E9" box="[617,880,1617,1642]" italics="true" pageId="9" pageNumber="42">Crenicichla jupiaensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="11EF6D9BAC0CFFF150E2D976FBD8F9E9" box="[936,1105,1617,1642]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="niederleinii">
|
||
<emphasis id="E49BCA0AAC0CFFF150E2D976FBD8F9E9" box="[936,1105,1617,1642]" italics="true" pageId="9" pageNumber="42">C. niederleinii</emphasis>
|
||
</taxonomicName>
|
||
are found both in the Upper and Middle Paraná basin. Several species (i.e.
|
||
<emphasis id="E49BCA0AAC0CFFF15188D951FAE3F90C" box="[706,1386,1654,1679]" italics="true" pageId="9" pageNumber="42">
|
||
<taxonomicName id="11EF6D9BAC0CFFF15188D951FCC0F90C" box="[706,841,1654,1679]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="lepidota">C. lepidota</taxonomicName>
|
||
,
|
||
<taxonomicName id="11EF6D9BAC0CFFF1501CD951FBA3F90C" box="[854,1066,1654,1679]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="mandelburgeri">C. mandelburgeri</taxonomicName>
|
||
,
|
||
<taxonomicName id="11EF6D9BAC0CFFF1577DD951FB2CF90C" box="[1079,1189,1654,1679]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="scottii">C. scottii</taxonomicName>
|
||
,
|
||
<taxonomicName id="11EF6D9BAC0CFFF157F8D951FAE3F90C" box="[1202,1386,1654,1679]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="semifasciata">C. semifasciata</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="11EF6D9BAC0CFFF153DDD9BCFE85F937" box="[151,268,1691,1716]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="vittata">
|
||
<emphasis id="E49BCA0AAC0CFFF153DDD9BCFE85F937" box="[151,268,1691,1716]" italics="true" pageId="9" pageNumber="42">C. vittata</emphasis>
|
||
</taxonomicName>
|
||
) inhabit only the lower–middle part of the river;
|
||
<taxonomicName id="11EF6D9BAC0CFFF15031D9BCFC79F937" box="[891,1008,1691,1716]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="vittata">
|
||
<emphasis id="E49BCA0AAC0CFFF15031D9BCFC79F937" box="[891,1008,1691,1716]" italics="true" pageId="9" pageNumber="42">C. vittata</emphasis>
|
||
</taxonomicName>
|
||
occurs also in the
|
||
<collectingCountry id="AEF85688AC0CFFF15794D9BCFAC3F936" box="[1246,1354,1691,1717]" name="Uruguay" pageId="9" pageNumber="42">Uruguay</collectingCountry>
|
||
basin.
|
||
<taxonomicName id="11EF6D9BAC0CFFF153DDD9E6FE4FF959" box="[151,454,1729,1754]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="iguassuensis">
|
||
<emphasis id="E49BCA0AAC0CFFF153DDD9E6FE4FF959" box="[151,454,1729,1754]" italics="true" pageId="9" pageNumber="42">Crenicichla iguassuensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="11EF6D9BAC0CFFF15148D9E6FDEDF959" box="[514,612,1729,1754]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="tesay">
|
||
<emphasis id="E49BCA0AAC0CFFF15148D9E6FDEDF959" box="[514,612,1729,1754]" italics="true" pageId="9" pageNumber="42">C. tesay</emphasis>
|
||
</taxonomicName>
|
||
are only present in the Iguazú basin.
|
||
<taxonomicName id="11EF6D9BAC0CFFF15765D9E6FA89F959" box="[1071,1280,1729,1754]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="yaha">
|
||
<emphasis id="E49BCA0AAC0CFFF15765D9E6FA89F959" box="[1071,1280,1729,1754]" italics="true" pageId="9" pageNumber="42">Crenicichla yaha</emphasis>
|
||
</taxonomicName>
|
||
is registered from the arroyo Urugua–í (Paraná basin) and Iguazú basin above the Cataratas del Iguazú.
|
||
</paragraph>
|
||
<paragraph id="D6501618AC0CFFF1538FD82CFE93F816" blockId="9.[151,1437,982,2016]" pageId="9" pageNumber="42">
|
||
The high diversity of ichthyofauna in the Argentinean province of Misiones has already been stressed. Three new
|
||
<taxonomicName id="11EF6D9BAC0CFFF15254D816FE20F8C9" box="[286,425,1841,1866]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="genus">
|
||
<emphasis id="E49BCA0AAC0CFFF15254D816FE20F8C9" box="[286,425,1841,1866]" italics="true" pageId="9" pageNumber="42">Crenicichla</emphasis>
|
||
</taxonomicName>
|
||
species have recently been described from the río Paraná tributaries in Misiones (
|
||
<taxonomicName id="11EF6D9BAC0CFFF156CBD816FF59F8EC" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="yaha">
|
||
<emphasis id="E49BCA0AAC0CFFF156CBD816FF59F8EC" italics="true" pageId="9" pageNumber="42">C. yaha</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="11EF6D9BAC0CFFF15394D871FEB0F8EC" box="[222,313,1878,1903]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="tesay">
|
||
<emphasis id="E49BCA0AAC0CFFF15394D871FEB0F8EC" box="[222,313,1878,1903]" italics="true" pageId="9" pageNumber="42">C. tesay</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<emphasis id="E49BCA0AAC0CFFF15232D871FE3EF8EC" box="[376,439,1878,1903]" italics="true" pageId="9" pageNumber="42">C. hu</emphasis>
|
||
), and several additional putatively new species are known to us from these tributaries (pers. obs).
|
||
</paragraph>
|
||
<paragraph id="D6501618AC0CFFF2538FD886FEC2FEA1" blockId="9.[151,1437,982,2016]" lastBlockId="10.[151,1437,152,290]" lastPageId="10" lastPageNumber="43" pageId="9" pageNumber="42">
|
||
We also confirm the presence of
|
||
<taxonomicName id="11EF6D9BAC0CFFF15109D886FC09F839" box="[579,896,1953,1978]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="mandelburgeri">
|
||
<emphasis id="E49BCA0AAC0CFFF15109D886FC09F839" box="[579,896,1953,1978]" italics="true" pageId="9" pageNumber="42">Crenicichla mandelburgeri</emphasis>
|
||
</taxonomicName>
|
||
in Misiones. Our material was compared with material from the
|
||
<typeStatus id="0954A8BAAC0CFFF15238D8E1FE2FF863" box="[370,422,1990,2016]" pageId="9" pageNumber="42">type</typeStatus>
|
||
localities of
|
||
<taxonomicName id="11EF6D9BAC0CFFF1510FD8E1FC92F85C" box="[581,795,1990,2015]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="mandelburgeri">
|
||
<emphasis id="E49BCA0AAC0CFFF1510FD8E1FC92F85C" box="[581,795,1990,2015]" italics="true" pageId="9" pageNumber="42">C. mandelburgeri</emphasis>
|
||
</taxonomicName>
|
||
(both morphology and the ND2 gene sequences; not shown). This taxon, however, demonstrates geographical variation as well as phylogeographic structure (
|
||
<figureCitation id="4ED40A9DAC0FFFF25627DFBFFF2FFF54" captionStart="FIGURE 1" captionStartId="5.[151,255,965,989]" captionTargetBox="[158,1418,179,941]" captionTargetId="figure@5.[151,1436,167,949]" captionTargetPageId="5" captionText="FIGURE 1. Maximum parsimony tree topology based on ND 2 sequences; the dotted line displays the different topology of the alternative Bayesian inference. Numbers for each recovered node represent nonparametric bootstrap support (left) and Bayesian posterior probability (right), respectively." httpUri="https://zenodo.org/record/196656/files/figure.png" pageId="10" pageNumber="43">Fig. 1</figureCitation>
|
||
), and we thus cannot rule out the presence of a species complex; the specimens from Piray-Guazu (haplotypes C15, C17) that infringe on the monophyly of this species are therefore referred to as
|
||
<taxonomicName id="11EF6D9BAC0FFFF25613DFC5FECDFEA2" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="43" phylum="Chordata" rank="species" species="mandelburgeri">
|
||
<emphasis id="E49BCA0AAC0FFFF25613DFC5FECDFEA2" italics="true" pageId="10" pageNumber="43">C. cf. mandelburgeri</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="82904690AC0FFFF253DDD84CFA84F800" box="[151,1293,1899,1923]" httpUri="https://zenodo.org/record/196661/files/figure.png" pageId="10" pageNumber="43" targetBox="[199,1385,341,1863]" targetPageId="10">
|
||
<paragraph id="D6501618AC0FFFF253DDD84CFA84F800" blockId="10.[151,1293,1899,1923]" box="[151,1293,1899,1923]" pageId="10" pageNumber="43">
|
||
<emphasis id="E49BCA0AAC0FFFF253DDD84CFE90F800" bold="true" box="[151,281,1899,1923]" pageId="10" pageNumber="43">FIGURE 7.</emphasis>
|
||
Hydrological map of the Province of Misiones. The type locality is marked with a solid circle.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D6501618AC0FFFF3538FD896FBC3FEA1" blockId="10.[151,1437,1969,2032]" lastBlockId="11.[151,1437,152,2007]" lastPageId="11" lastPageNumber="44" pageId="10" pageNumber="43">
|
||
Based on our field observations,
|
||
<taxonomicName id="11EF6D9BAC0FFFF25100D896FCEDF849" box="[586,868,1969,1994]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="43" phylum="Chordata" rank="species" species="missioneira">
|
||
<emphasis id="E49BCA0AAC0FFFF25100D896FCEDF849" box="[586,868,1969,1994]" italics="true" pageId="10" pageNumber="43">Crenicichla missioneira</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="11EF6D9BAC0FFFF25039D896FC74F849" box="[883,1021,1969,1994]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="43" phylum="Chordata" rank="species" species="minuano">
|
||
<emphasis id="E49BCA0AAC0FFFF25039D896FC74F849" box="[883,1021,1969,1994]" italics="true" pageId="10" pageNumber="43">C. minuano</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="11EF6D9BAC0FFFF2577CD896FB26F849" box="[1078,1199,1969,1994]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="43" phylum="Chordata" rank="species" species="gaucho">
|
||
<emphasis id="E49BCA0AAC0FFFF2577CD896FB26F849" box="[1078,1199,1969,1994]" italics="true" pageId="10" pageNumber="43">C. gaucho</emphasis>
|
||
</taxonomicName>
|
||
, described from the Middle río
|
||
<collectingCountry id="AEF85688AC0FFFF2526DD8F1FE1DF873" box="[295,404,2006,2032]" name="Uruguay" pageId="10" pageNumber="43">Uruguay</collectingCountry>
|
||
in Rio Grande do Sul,
|
||
<collectingCountry id="AEF85688AC0FFFF251F4D8F1FC82F873" box="[702,779,2006,2032]" name="Brazil" pageId="10" pageNumber="43">Brazil</collectingCountry>
|
||
and also cited by
|
||
<bibRefCitation id="B27E6BE9AC0FFFF250BCD8F1FADEF873" author="Lucena" box="[1014,1367,2006,2032]" pageId="10" pageNumber="46" refString="Lucena, C. A. S. & Kullander, S. O. (1992) The Crenicichla (Teleostei: Cichlidae) species of the Uruguai River drainage in Brazil. Ichthyological Exploration of Freshwaters, 3, 97 - 160." type="journal article" year="1992">Lucena & Kullander (1992)</bibRefCitation>
|
||
from Misiones, are quite common in the río
|
||
<collectingCountry id="AEF85688AC0EFFF3512DDFBFFD46FF31" box="[615,719,152,178]" name="Uruguay" pageId="11" pageNumber="44">Uruguay</collectingCountry>
|
||
tributaries in Misiones.
|
||
<taxonomicName id="11EF6D9BAC0EFFF350B9DFBFFA92FF32" box="[1011,1307,152,177]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="44" phylum="Chordata" rank="species" species="hadrostigma">
|
||
<emphasis id="E49BCA0AAC0EFFF350B9DFBFFA92FF32" box="[1011,1307,152,177]" italics="true" pageId="11" pageNumber="44">Crenicichla hadrostigma</emphasis>
|
||
</taxonomicName>
|
||
, described from the Upper río
|
||
<collectingCountry id="AEF85688AC0EFFF35234DF9AFE6CFF54" box="[382,485,189,215]" name="Uruguay" pageId="11" pageNumber="44">Uruguay</collectingCountry>
|
||
in Santa Catarina,
|
||
<collectingCountry id="AEF85688AC0EFFF35182DF9AFC99FF54" box="[712,784,189,215]" name="Brazil" pageId="11" pageNumber="44">Brazil</collectingCountry>
|
||
is known so far from one locality in Misiones (
|
||
<bibRefCitation id="B27E6BE9AC0EFFF3560EDF9AFF5EFF7F" author="Lucena" pageId="11" pageNumber="46" refString="Lucena, C. A. S. (2007) Two new species of the genus Crenicichla Heckel, 1840 from the upper rio Uruguay drainage (Perciformes: Cichlidae). Neotropical Ichthyology, 5, 449 - 456." type="journal article" year="2007">Lucena 2007</bibRefCitation>
|
||
), confirming the possibility of finding additional Upper
|
||
<collectingCountry id="AEF85688AC0EFFF35030DFC5FC6BFF7F" box="[890,994,226,252]" name="Uruguay" pageId="11" pageNumber="44">Uruguay</collectingCountry>
|
||
<taxonomicName id="11EF6D9BAC0EFFF350A0DFC5FBFCFF78" box="[1002,1141,226,251]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="44" phylum="Chordata" rank="genus">
|
||
<emphasis id="E49BCA0AAC0EFFF350A0DFC5FBFCFF78" box="[1002,1141,226,251]" italics="true" pageId="11" pageNumber="44">Crenicichla</emphasis>
|
||
</taxonomicName>
|
||
species in Misiones. On this note,
|
||
<bibRefCitation id="B27E6BE9AC0EFFF35240DE2FFDC6FEA1" author="Lucena" box="[266,591,264,290]" pageId="11" pageNumber="46" refString="Lucena, C. A. S. & Kullander, S. O. (1992) The Crenicichla (Teleostei: Cichlidae) species of the Uruguai River drainage in Brazil. Ichthyological Exploration of Freshwaters, 3, 97 - 160." type="journal article" year="1992">Lucena & Kullander (1992)</bibRefCitation>
|
||
also cite
|
||
<taxonomicName id="11EF6D9BAC0EFFF351F5DE2FFC19FEA2" box="[703,912,264,289]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="44" phylum="Chordata" rank="species" species="tendybaguassu">
|
||
<emphasis id="E49BCA0AAC0EFFF351F5DE2FFC19FEA2" box="[703,912,264,289]" italics="true" pageId="11" pageNumber="44">C. tendybaguassu</emphasis>
|
||
</taxonomicName>
|
||
from Misiones.
|
||
</paragraph>
|
||
<paragraph id="D6501618AC0EFFF3538FDE0AFBCFFDA4" blockId="11.[151,1437,152,2007]" pageId="11" pageNumber="44">
|
||
So far there are no records in Misiones of four
|
||
<taxonomicName id="11EF6D9BAC0EFFF351BDDE0AFC08FEC5" box="[759,897,301,326]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="44" phylum="Chordata" rank="genus">
|
||
<emphasis id="E49BCA0AAC0EFFF351BDDE0AFC08FEC5" box="[759,897,301,326]" italics="true" pageId="11" pageNumber="44">Crenicichla</emphasis>
|
||
</taxonomicName>
|
||
species which are known from the Upper río
|
||
<collectingCountry id="AEF85688AC0EFFF353DDDE75FF77FEEF" box="[151,254,338,364]" name="Uruguay" pageId="11" pageNumber="44">Uruguay</collectingCountry>
|
||
basin in
|
||
<collectingCountry id="AEF85688AC0EFFF35226DE75FE3DFEEF" box="[364,436,338,364]" name="Brazil" pageId="11" pageNumber="44">Brazil</collectingCountry>
|
||
(
|
||
<taxonomicName id="11EF6D9BAC0EFFF3528CDE75FDB5FEE8" box="[454,572,338,363]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="44" phylum="Chordata" rank="species" species="prenda">
|
||
<emphasis id="E49BCA0AAC0EFFF3528CDE75FDB5FEE8" box="[454,572,338,363]" italics="true" pageId="11" pageNumber="44">C. prenda</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="11EF6D9BAC0EFFF35106DE75FD56FEE8" box="[588,735,338,363]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="44" phylum="Chordata" rank="species" species="empheres">
|
||
<emphasis id="E49BCA0AAC0EFFF35106DE75FD56FEE8" box="[588,735,338,363]" italics="true" pageId="11" pageNumber="44">C. empheres</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="11EF6D9BAC0EFFF351A5DE75FCDBFEE8" box="[751,850,338,363]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="44" phylum="Chordata" rank="species" species="igara">
|
||
<emphasis id="E49BCA0AAC0EFFF351A5DE75FCDBFEE8" box="[751,850,338,363]" italics="true" pageId="11" pageNumber="44">C. igara</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="11EF6D9BAC0EFFF350C7DE75FC71FEE8" box="[909,1016,338,363]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="44" phylum="Chordata" rank="species" species="jurubi">
|
||
<emphasis id="E49BCA0AAC0EFFF350C7DE75FC71FEE8" box="[909,1016,338,363]" italics="true" pageId="11" pageNumber="44">C. jurubi</emphasis>
|
||
</taxonomicName>
|
||
), whose boundary with the Middle río
|
||
<collectingCountry id="AEF85688AC0EFFF353F4DE5FFEACFE11" box="[190,293,376,402]" name="Uruguay" pageId="11" pageNumber="44">Uruguay</collectingCountry>
|
||
is recognised as being at the Salto Moconá (Yucumã), just downstream from the río Pepirí–Guazú which forms the eastern border between Misiones,
|
||
<collectingCountry id="AEF85688AC0EFFF351B4DEBAFCFCFE34" box="[766,885,413,439]" name="Argentina" pageId="11" pageNumber="44">Argentina</collectingCountry>
|
||
and Santa Catarina,
|
||
<collectingCountry id="AEF85688AC0EFFF35725DEBAFB31FE34" box="[1135,1208,413,439]" name="Brazil" pageId="11" pageNumber="44">Brazil</collectingCountry>
|
||
(
|
||
<bibRefCitation id="B27E6BE9AC0EFFF35782DEBAFEBBFE5F" author="Zaniboni" pageId="11" pageNumber="46" refString="Zaniboni Filho, E. & Schulz, U. H. (2003) Migratory fishes of the Uruguay river. In: Carolsfeld, J., Harvey, B., Ross, C. & Baer, A. (Eds.), Migratory fishes of South America: Biology, Fisheries and Conservation Status. International Development Research Center and The World Bank, pp. 157 - 194." type="book chapter" year="2003">Zaniboni Filho & Schulz 2003</bibRefCitation>
|
||
). The ranges of these species could thus primarily be outside of Misiones as they are also not known from the Middle río
|
||
<collectingCountry id="AEF85688AC0EFFF352AADECFFDCFFD81" box="[480,582,488,514]" name="Uruguay" pageId="11" pageNumber="44">Uruguay</collectingCountry>
|
||
in Rio Grande do Sul,
|
||
<collectingCountry id="AEF85688AC0EFFF35010DECFFC2CFD81" box="[858,933,488,514]" name="Brazil" pageId="11" pageNumber="44">Brazil</collectingCountry>
|
||
. However, the presence of these species in Misiones cannot be ruled out (see
|
||
<taxonomicName id="11EF6D9BAC0EFFF35163DD2AFD57FDA5" box="[553,734,525,550]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="44" phylum="Chordata" rank="species" species="hadrostigma">
|
||
<emphasis id="E49BCA0AAC0EFFF35163DD2AFD57FDA5" box="[553,734,525,550]" italics="true" pageId="11" pageNumber="44">C. hadrostigma</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="11EF6D9BAC0EFFF3505DDD2AFC6EFDA5" box="[791,999,525,550]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="44" phylum="Chordata" rank="species" species="tendybaguassu">
|
||
<emphasis id="E49BCA0AAC0EFFF3505DDD2AFC6EFDA5" box="[791,999,525,550]" italics="true" pageId="11" pageNumber="44">C. tendybaguassu</emphasis>
|
||
</taxonomicName>
|
||
above).
|
||
</paragraph>
|
||
<paragraph id="D6501618AC0EFFF3538FDD15FDF1FBD4" blockId="11.[151,1437,152,2007]" pageId="11" pageNumber="44">
|
||
Despite its small size, the province of Misiones shows biogeographic structuring which cannot be explained merely by diversity on a broader scale. Both the río Paraná and río
|
||
<collectingCountry id="AEF85688AC0EFFF3576FDD7FFB05FDF1" box="[1061,1164,600,626]" name="Uruguay" pageId="11" pageNumber="44">Uruguay</collectingCountry>
|
||
tributaries in Misiones are divided from the main rivers by waterfalls close to their mouths, but the northern tributaries of the Paraná in particular and the Iguazú itself in Misiones have a significant number of endemics which are so far not known outside of Misiones (i.e.
|
||
<collectingCountry id="AEF85688AC0EFFF35153DDEFFD0EFD61" box="[537,647,712,738]" name="Paraguay" pageId="11" pageNumber="44">Paraguay</collectingCountry>
|
||
or
|
||
<collectingCountry id="AEF85688AC0EFFF351FBDDEFFD76FD61" box="[689,767,712,738]" name="Brazil" pageId="11" pageNumber="44">Brazil</collectingCountry>
|
||
). Among
|
||
<taxonomicName id="11EF6D9BAC0EFFF3503EDDEFFC76FD62" box="[884,1023,712,737]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="44" phylum="Chordata" rank="genus">
|
||
<emphasis id="E49BCA0AAC0EFFF3503EDDEFFC76FD62" box="[884,1023,712,737]" italics="true" pageId="11" pageNumber="44">Crenicichla</emphasis>
|
||
</taxonomicName>
|
||
these are
|
||
<taxonomicName id="11EF6D9BAC0EFFF35732DDEFFB50FD62" box="[1144,1241,712,737]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="44" phylum="Chordata" rank="species" species="tesay">
|
||
<emphasis id="E49BCA0AAC0EFFF35732DDEFFB50FD62" box="[1144,1241,712,737]" italics="true" pageId="11" pageNumber="44">C. tesay</emphasis>
|
||
</taxonomicName>
|
||
from río Iguazú and
|
||
<taxonomicName id="11EF6D9BAC0EFFF35380DDCAFEAEFC85" box="[202,295,749,774]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="44" phylum="Chordata" rank="species" species="yaha">
|
||
<emphasis id="E49BCA0AAC0EFFF35380DDCAFEAEFC85" box="[202,295,749,774]" italics="true" pageId="11" pageNumber="44">C. yaha</emphasis>
|
||
</taxonomicName>
|
||
from río Iguazú and arroyo Urugua–í and
|
||
<emphasis id="E49BCA0AAC0EFFF3506CDDCAFCEEFC85" box="[806,871,749,774]" italics="true" pageId="11" pageNumber="44">C. hu</emphasis>
|
||
from arroyo Piray–Miní (
|
||
<figureCitation id="4ED40A9DAC0EFFF357D5DDCAFB60FC84" box="[1183,1257,749,775]" captionStart="FIGURE 7" captionStartId="10.[151,255,1899,1923]" captionTargetBox="[199,1385,341,1863]" captionTargetId="figure@10.[187,1400,329,1878]" captionTargetPageId="10" captionText="FIGURE 7. Hydrological map of the Province of Misiones. The type locality is marked with a solid circle." httpUri="https://zenodo.org/record/196661/files/figure.png" pageId="11" pageNumber="44">Fig. 7</figureCitation>
|
||
). Two putative new species are further known from the arroyo Urugua–í (pers. obs.), which is located between the río Iguazú and arroyo Piray–Miní. These three drainages together with arroyo Aguaray–Guazú form the northern part of Misiones. From the southeastern-most point of this part of Misiones starts the watershed between the río Paraná and río
|
||
<collectingCountry id="AEF85688AC0EFFF35201DCA5FE3DFC1F" box="[331,436,898,924]" name="Uruguay" pageId="11" pageNumber="44">Uruguay</collectingCountry>
|
||
. Tributaries of the río Paraná from here to the southeast (starting with arroyo Piray– Guazú,
|
||
<figureCitation id="4ED40A9DAC0EFFF353B9DC8FFEB1FC41" box="[243,312,936,962]" captionStart="FIGURE 7" captionStartId="10.[151,255,1899,1923]" captionTargetBox="[199,1385,341,1863]" captionTargetId="figure@10.[187,1400,329,1878]" captionTargetPageId="10" captionText="FIGURE 7. Hydrological map of the Province of Misiones. The type locality is marked with a solid circle." httpUri="https://zenodo.org/record/196661/files/figure.png" pageId="11" pageNumber="44">Fig. 7</figureCitation>
|
||
) have a diferent fauna of
|
||
<taxonomicName id="11EF6D9BAC0EFFF35121DC8FFD7DFC42" box="[619,756,936,961]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="44" phylum="Chordata" rank="genus">
|
||
<emphasis id="E49BCA0AAC0EFFF35121DC8FFD7DFC42" box="[619,756,936,961]" italics="true" pageId="11" pageNumber="44">Crenicichla</emphasis>
|
||
</taxonomicName>
|
||
(dominated by
|
||
<taxonomicName id="11EF6D9BAC0EFFF350E5DC8FFBF6FC42" box="[943,1151,936,962]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="44" phylum="Chordata" rank="species" species="mandelburgeri">
|
||
<emphasis id="E49BCA0AAC0EFFF350E5DC8FFC4BFC42" box="[943,962,936,961]" italics="true" pageId="11" pageNumber="44">C</emphasis>
|
||
.
|
||
<emphasis id="E49BCA0AAC0EFFF35098DC8FFBF6FC42" box="[978,1151,936,961]" italics="true" pageId="11" pageNumber="44">mandelburgeri</emphasis>
|
||
</taxonomicName>
|
||
). A very similar pattern is also observed among
|
||
<taxonomicName id="11EF6D9BAC0EFFF352F5DCEAFDE2FC65" box="[447,619,973,998]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="44" phylum="Chordata" rank="genus">
|
||
<emphasis id="E49BCA0AAC0EFFF352F5DCEAFDE2FC65" box="[447,619,973,998]" italics="true" pageId="11" pageNumber="44">Australoheros</emphasis>
|
||
</taxonomicName>
|
||
Ř íčan & Kullander, but with the exception that south from arroyo Piray–Miní there are so far no known species of
|
||
<taxonomicName id="11EF6D9BAC0EFFF351A9DCD5FC02FB88" box="[739,907,1010,1035]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="44" phylum="Chordata" rank="genus">
|
||
<emphasis id="E49BCA0AAC0EFFF351A9DCD5FC02FB88" box="[739,907,1010,1035]" italics="true" pageId="11" pageNumber="44">Australoheros</emphasis>
|
||
</taxonomicName>
|
||
in the río Paraná tributaries in Misiones (
|
||
<taxonomicName id="11EF6D9BAC0EFFF356C8DCD5FE33FBB1" authority="Casciotta et al." authorityName="Casciotta et al." class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="44" phylum="Chordata" rank="species" species="kaaygua">
|
||
<emphasis id="E49BCA0AAC0EFFF356C8DCD5FF74FBB2" italics="true" pageId="11" pageNumber="44">A. kaaygua</emphasis>
|
||
Casciotta
|
||
<emphasis id="E49BCA0AAC0EFFF35237DB3FFE3DFBB2" box="[381,436,1048,1073]" italics="true" pageId="11" pageNumber="44">et al</emphasis>
|
||
.
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="11EF6D9BAC0EFFF35281DB3FFC8EFBB1" authority="Casciotta et al." authorityName="Casciotta et al." box="[459,775,1048,1074]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="44" phylum="Chordata" rank="species" species="tembe">
|
||
<emphasis id="E49BCA0AAC0EFFF35281DB3FFDBCFBB2" box="[459,565,1048,1073]" italics="true" pageId="11" pageNumber="44">A. tembe</emphasis>
|
||
(Casciotta
|
||
<emphasis id="E49BCA0AAC0EFFF3518ADB3FFD7CFBB2" box="[704,757,1048,1073]" italics="true" pageId="11" pageNumber="44">et al</emphasis>
|
||
.)
|
||
</taxonomicName>
|
||
[and likely
|
||
<taxonomicName id="11EF6D9BAC0EFFF350D1DB3FFB95FBB2" box="[923,1052,1048,1073]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="44" phylum="Chordata" rank="species" species="guarani">
|
||
<emphasis id="E49BCA0AAC0EFFF350D1DB3FFB95FBB2" box="[923,1052,1048,1073]" italics="true" pageId="11" pageNumber="44">A. guaraní</emphasis>
|
||
</taxonomicName>
|
||
Ř íčan & Kullander] are known again only from the northern tributaries).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9EF54593AC0EFFF3538FDB45FD18F854" pageId="11" pageNumber="44" type="materials_examined">
|
||
<paragraph id="D6501618AC0EFFF3538FDB45FD18F854" blockId="11.[151,1437,152,2007]" pageId="11" pageNumber="44">
|
||
<emphasis id="E49BCA0AAC0EFFF3538FDB45FE6BFBFF" bold="true" box="[197,482,1122,1148]" pageId="11" pageNumber="44">Comparative material.</emphasis>
|
||
A list of comparative material of
|
||
<taxonomicName id="11EF6D9BAC0EFFF35024DB44FC50FBFF" box="[878,985,1122,1148]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="44" phylum="Chordata" rank="species" species="scottii">
|
||
<emphasis id="E49BCA0AAC0EFFF35024DB44FC08FBFF" box="[878,897,1123,1148]" italics="true" pageId="11" pageNumber="44">C</emphasis>
|
||
.
|
||
<emphasis id="E49BCA0AAC0EFFF350DBDB44FC50FBFF" box="[913,985,1123,1148]" italics="true" pageId="11" pageNumber="44">scottii</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="11EF6D9BAC0EFFF3575BDB44FBF7FBFF" box="[1041,1150,1122,1148]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="44" phylum="Chordata" rank="species" species="vittata">
|
||
<emphasis id="E49BCA0AAC0EFFF3575BDB44FBADFBFF" box="[1041,1060,1123,1148]" italics="true" pageId="11" pageNumber="44">C</emphasis>
|
||
.
|
||
<emphasis id="E49BCA0AAC0EFFF35779DB44FBF7FBFF" box="[1075,1150,1123,1148]" italics="true" pageId="11" pageNumber="44">vittata</emphasis>
|
||
</taxonomicName>
|
||
is available in
|
||
<bibRefCitation id="B27E6BE9AC0EFFF35665DB45FF63FB21" author="Casciotta" pageId="11" pageNumber="45" refString="Casciotta, J. R. (1987) Crenicichla celidochilus n. sp. from Uruguay and a multivariate analysis of the lacustris group (Perciformes, Cichlidae). Copeia, 1987, 883 - 891." type="journal article" year="1987">Casciotta (1987)</bibRefCitation>
|
||
. In addition, the following material was studied:
|
||
<taxonomicName id="11EF6D9BAC0EFFF3507DDBAFFBD6FB22" box="[823,1119,1160,1185]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="44" phylum="Chordata" rank="species" species="hadrostigma">
|
||
<emphasis id="E49BCA0AAC0EFFF3507DDBAFFBD6FB22" box="[823,1119,1160,1185]" italics="true" pageId="11" pageNumber="44">Crenicichla hadrostigma</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<collectingCountry id="AEF85688AC0EFFF3573ADBAFFB62FB21" box="[1136,1259,1160,1186]" name="Argentina" pageId="11" pageNumber="44">Argentina</collectingCountry>
|
||
. AI 220, 1 ex.,
|
||
<quantity id="1117BBFDAC0EFFF353DDDB8AFE8BFB44" box="[151,258,1197,1223]" metricMagnitude="-2" metricUnit="m" metricValue="7.279999999999999" pageId="11" pageNumber="44" unit="mm" value="72.8">72.8 mm</quantity>
|
||
, Misiones, Itacaruare, río
|
||
<collectingCountry id="AEF85688AC0EFFF3517ADB8AFD1EFB44" box="[560,663,1197,1223]" name="Uruguay" pageId="11" pageNumber="44">Uruguay</collectingCountry>
|
||
basin.
|
||
<taxonomicName id="11EF6D9BAC0EFFF351A3DB8AFB84FB45" box="[745,1037,1197,1222]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="44" phylum="Chordata" rank="species" species="iguassuensis">
|
||
<emphasis id="E49BCA0AAC0EFFF351A3DB8AFB84FB45" box="[745,1037,1197,1222]" italics="true" pageId="11" pageNumber="44">Crenicichla iguassuensis</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<collectingCountry id="AEF85688AC0EFFF35756DB8AFBEFFB44" box="[1052,1126,1197,1223]" name="Brazil" pageId="11" pageNumber="44">Brasil</collectingCountry>
|
||
.
|
||
<collectionCode id="B0FE8EDDAC0EFFF35725DB8AFB4DFB44" LSID="urn:lsid:biocol.org:col:34795" box="[1135,1220,1197,1223]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="11" pageNumber="44">FMNH</collectionCode>
|
||
54159 (
|
||
<typeStatus id="0954A8BAAC0EFFF35669DB8AFA06FB44" box="[1315,1423,1197,1223]" pageId="11" pageNumber="44" type="holotype">holotype</typeStatus>
|
||
),
|
||
<quantity id="1117BBFDAC0EFFF353DDDBF5FF74FB6F" box="[151,253,1234,1260]" metricMagnitude="-1" metricUnit="m" metricValue="1.3699999999999999" pageId="11" pageNumber="44" unit="mm" value="137.0">137 mm</quantity>
|
||
, Porto Uniao da Victoria, Rio Iguassu.
|
||
<taxonomicName id="11EF6D9BAC0EFFF3519EDBF4FC68FB6F" box="[724,993,1235,1260]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="44" phylum="Chordata" rank="species" species="jupiaensis">
|
||
<emphasis id="E49BCA0AAC0EFFF3519EDBF4FC68FB6F" box="[724,993,1235,1260]" italics="true" pageId="11" pageNumber="44">Crenicichla jupiaensis</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<collectingCountry id="AEF85688AC0EFFF350BBDBF5FBE2FB6F" box="[1009,1131,1234,1260]" name="Argentina" pageId="11" pageNumber="44">Argentina</collectingCountry>
|
||
. AI 226, 2 ex., 87.7–93.0 mm, Corrientes, río Paraná at Yahapé. AI 227, 1 ex.,
|
||
<quantity id="1117BBFDAC0EFFF35047DBDFFCF0FA91" box="[781,889,1272,1298]" metricMagnitude="-2" metricUnit="m" metricValue="6.07" pageId="11" pageNumber="44" unit="mm" value="60.7">60.7 mm</quantity>
|
||
, Corrientes, río Paraná at Yahapé.
|
||
<taxonomicName id="11EF6D9BAC0EFFF35658DBDFFF7EFAB5" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="44" phylum="Chordata" rank="species" species="lepidota">
|
||
<emphasis id="E49BCA0AAC0EFFF35658DBDFFF7EFAB5" italics="true" pageId="11" pageNumber="44">Crenicichla lepidota</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<collectingCountry id="AEF85688AC0EFFF3524CDA3AFEF7FAB4" box="[262,382,1309,1335]" name="Argentina" pageId="11" pageNumber="44">Argentina</collectingCountry>
|
||
. MACN-ict 5067, 4 ex.,
|
||
<quantity id="1117BBFDAC0EFFF351EBDA3AFCD5FAB4" box="[673,860,1309,1335]" metricMagnitude="-2" metricUnit="m" metricValue="9.055" metricValueMax="11.34" metricValueMin="6.77" pageId="11" pageNumber="44" unit="mm" value="90.55" valueMax="113.4" valueMin="67.7">67.7–113.4 mm</quantity>
|
||
, Misiones, Represa Estación Experimental Cerro Azul.
|
||
<collectionCode id="B0FE8EDDAC0EFFF35395DA65FE92FADF" box="[223,283,1346,1372]" httpUri="http://grbio.org/cool/sggj-9jsz" name="Fundacion Miguel Lillo" pageId="11" pageNumber="44">FML</collectionCode>
|
||
0 0 528, 1 ex.,
|
||
<quantity id="1117BBFDAC0EFFF3528EDA65FDB4FADF" box="[452,573,1346,1372]" metricMagnitude="-1" metricUnit="m" metricValue="1.115" pageId="11" pageNumber="44" unit="mm" value="111.5">111.5 mm</quantity>
|
||
, Salta, Luna Muerta, Hickman. MACN-ict 3656, 2 ex., 116.0–
|
||
<quantity id="1117BBFDAC0EFFF35654DA65FA11FADF" box="[1310,1432,1346,1372]" metricMagnitude="-1" metricUnit="m" metricValue="1.657" pageId="11" pageNumber="44" unit="mm" value="165.7">165.7 mm</quantity>
|
||
,
|
||
<collectingCountry id="AEF85688AC0EFFF353DDDA4FFE8BFA01" box="[151,258,1384,1410]" name="Taiwan" pageId="11" pageNumber="44">Formosa</collectingCountry>
|
||
, Riacho de Oro. MACN-ict 7275, 1 ex.,
|
||
<quantity id="1117BBFDAC0EFFF351A0DA4FFCEFFA01" box="[746,870,1384,1410]" metricMagnitude="-1" metricUnit="m" metricValue="1.516" pageId="11" pageNumber="44" unit="mm" value="151.6">151.6 mm</quantity>
|
||
, Chaco, Esteros del Palmar.
|
||
<collectionCode id="B0FE8EDDAC0EFFF357F1DA4FFB71FA01" box="[1211,1272,1384,1410]" httpUri="http://grbio.org/cool/sggj-9jsz" name="Fundacion Miguel Lillo" pageId="11" pageNumber="44">FML</collectionCode>
|
||
0 0 312, 1 ex., 138.0 mm, Corrientes,
|
||
<collectingCountry id="AEF85688AC0EFFF352F3DAAAFE6FFA24" box="[441,486,1421,1447]" name="Iceland" pageId="11" pageNumber="44">Isla</collectingCountry>
|
||
Apipé Grande, Ituzaingó. MACN-ict 4091, 1 ex.,
|
||
<quantity id="1117BBFDAC0EFFF35729DAAAFB5FFA24" box="[1123,1238,1421,1447]" metricMagnitude="-2" metricUnit="m" metricValue="9.84" pageId="11" pageNumber="44" unit="mm" value="98.4">98.4 mm</quantity>
|
||
, Entre Ríos, río
|
||
<collectingCountry id="AEF85688AC0EFFF353DDDA95FF76FA4F" box="[151,255,1458,1484]" name="Uruguay" pageId="11" pageNumber="44">Uruguay</collectingCountry>
|
||
, Concepción del
|
||
<collectingCountry id="AEF85688AC0EFFF3528CDA95FDA7FA4F" box="[454,558,1458,1484]" name="Uruguay" pageId="11" pageNumber="44">Uruguay</collectingCountry>
|
||
. MACN-ict 2314, 6 ex.,
|
||
<quantity id="1117BBFDAC0EFFF35007DA95FB81FA4F" box="[845,1032,1458,1484]" metricMagnitude="-2" metricUnit="m" metricValue="8.205" metricValueMax="10.42" metricValueMin="5.99" pageId="11" pageNumber="44" unit="mm" value="82.05" valueMax="104.2" valueMin="59.9">59.9–104.2 mm</quantity>
|
||
, Buenos Aires,
|
||
<collectingCountry id="AEF85688AC0EFFF357F5DA95FB60FA4F" box="[1215,1257,1458,1484]" name="Iceland" pageId="11" pageNumber="44">Isla</collectingCountry>
|
||
Martín García.
|
||
<collectingCountry id="AEF85688AC0EFFF353DDDAFFFE82FA71" box="[151,267,1496,1522]" name="Uruguay" pageId="11" pageNumber="44">Uruguay</collectingCountry>
|
||
.
|
||
<collectionCode id="B0FE8EDDAC0EFFF35252DAFFFE1AFA71" box="[280,403,1496,1522]" httpUri="http://grbio.org/cool/nctz-br8x" name="Museo Nacional de Historia Natural, Mexico City" pageId="11" pageNumber="44">MNHNM</collectionCode>
|
||
2087, 1 ex.,
|
||
<quantity id="1117BBFDAC0EFFF35108DAFFFD30FA71" box="[578,697,1496,1522]" metricMagnitude="-2" metricUnit="m" metricValue="7.29" pageId="11" pageNumber="44" unit="mm" value="72.9">72.9 mm</quantity>
|
||
, Departamento Colonia, arroyo Limetas.
|
||
<taxonomicName id="11EF6D9BAC0EFFF35790DAFFFECFF995" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="44" phylum="Chordata" rank="species" species="mandelburgeri">
|
||
<emphasis id="E49BCA0AAC0EFFF35790DAFFFECFF995" italics="true" pageId="11" pageNumber="44">Crenicichla cf. mandelburgeri</emphasis>
|
||
</taxonomicName>
|
||
: MACN-ict 9439, 2 ex., 83.7–93.0 mm, Misiones, arroyo Guaruhape en ruta 220, río Paraná basin. MACN-ict 9440, 2 ex.,
|
||
<quantity id="1117BBFDAC0EFFF352B0D905FD2EF9BF" box="[506,679,1570,1596]" metricMagnitude="-2" metricUnit="m" metricValue="7.744999999999999" metricValueMax="8.23" metricValueMin="7.26" pageId="11" pageNumber="44" unit="mm" value="77.44999999999999" valueMax="82.3" valueMin="72.6">72.6–82.3 mm</quantity>
|
||
, Misiones, arroyo Cuñapirú, in route 223 near Ruiz de Montoya, río Paraná basin. MACN-ict 9441, 7 ex., 56.0–93.0 mm, Misiones, arroyo Cuñapirú (arroyo Tucangua), río Paraná basin. MACN-ict 9442, 2 ex.,
|
||
<quantity id="1117BBFDAC0EFFF35124D94AFCA5F904" box="[622,812,1645,1671]" metricMagnitude="-1" metricUnit="m" metricValue="1.551" metricValueMax="2.08" metricValueMin="1.022" pageId="11" pageNumber="44" unit="mm" value="155.1" valueMax="208.0" valueMin="102.2">102.2–208 mm</quantity>
|
||
, Misiones, arroyo Chapa, ruta 6, río Paraná basin.
|
||
<taxonomicName id="11EF6D9BAC0EFFF353DDD9B4FEF9F92F" box="[151,368,1683,1708]" class="Actinopterygii" family="Cichlidae" genus="Boggiana" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="44" phylum="Chordata" rank="species" species="ocellata">
|
||
<emphasis id="E49BCA0AAC0EFFF353DDD9B4FEF9F92F" box="[151,368,1683,1708]" italics="true" pageId="11" pageNumber="44">Boggiana ocellata</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<collectingCountry id="AEF85688AC0EFFF352CAD9B5FE66F92F" box="[384,495,1682,1708]" name="Paraguay" pageId="11" pageNumber="44">Paraguay</collectingCountry>
|
||
.
|
||
<collectionCode id="B0FE8EDDAC0EFFF352B1D9B5FDD9F92F" box="[507,592,1682,1708]" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="11" pageNumber="44">MSNG</collectionCode>
|
||
33700 (
|
||
<typeStatus id="0954A8BAAC0EFFF351F8D9B5FC96F92F" box="[690,799,1682,1708]" pageId="11" pageNumber="44" type="holotype">holotype</typeStatus>
|
||
),
|
||
<quantity id="1117BBFDAC0EFFF35079D9B5FC27F92F" box="[819,942,1682,1708]" metricMagnitude="-1" metricUnit="m" metricValue="2.575" pageId="11" pageNumber="44" unit="mm" value="257.5">257.5 mm</quantity>
|
||
, Puerto 14 de Mayo, Bahía Negra, Chaco Boreal.
|
||
<taxonomicName id="11EF6D9BAC0EFFF353B2D99FFDA0F952" box="[248,553,1720,1745]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="44" phylum="Chordata" rank="species" species="semifasciata">
|
||
<emphasis id="E49BCA0AAC0EFFF353B2D99FFDA0F952" box="[248,553,1720,1745]" italics="true" pageId="11" pageNumber="44">Crenicichla semifasciata</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<collectingCountry id="AEF85688AC0EFFF35170D99FFD31F951" box="[570,696,1720,1746]" name="Argentina" pageId="11" pageNumber="44">Argentina</collectingCountry>
|
||
. MACN-ict 3683, 1 ex.,
|
||
<quantity id="1117BBFDAC0EFFF350A0D99FFBD3F951" box="[1002,1114,1720,1746]" metricMagnitude="-2" metricUnit="m" metricValue="6.88" pageId="11" pageNumber="44" unit="mm" value="68.8">68.8 mm</quantity>
|
||
,
|
||
<collectingCountry id="AEF85688AC0EFFF3572DD99FFB5DF951" box="[1127,1236,1720,1746]" name="Taiwan" pageId="11" pageNumber="44">Formosa</collectingCountry>
|
||
, Riacho de Oro. MACN-ict 6239, 1 ex., 176,6 mm, Entre Ríos, arroyo Curupí.
|
||
<taxonomicName id="11EF6D9BAC0EFFF3503FD9FAFBCBF975" box="[885,1090,1757,1782]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="44" phylum="Chordata" rank="species" species="tesay">
|
||
<emphasis id="E49BCA0AAC0EFFF3503FD9FAFBCBF975" box="[885,1090,1757,1782]" italics="true" pageId="11" pageNumber="44">Crenicichla tesay</emphasis>
|
||
</taxonomicName>
|
||
: MACN-ict 9016 (
|
||
<typeStatus id="0954A8BAAC0EFFF35669D9FAFA06F974" box="[1315,1423,1757,1783]" pageId="11" pageNumber="44" type="holotype">holotype</typeStatus>
|
||
),
|
||
<quantity id="1117BBFDAC0EFFF353DDD825FE9CF89F" box="[151,277,1794,1820]" metricMagnitude="-1" metricUnit="m" metricValue="1.151" pageId="11" pageNumber="44" unit="mm" value="115.1">115.1 mm</quantity>
|
||
,
|
||
<collectingCountry id="AEF85688AC0EFFF35268D825FE29F89F" box="[290,416,1794,1820]" name="Argentina" pageId="11" pageNumber="44">Argentina</collectingCountry>
|
||
, Misiones, río Iguazú basin, arroyo Verde.
|
||
<taxonomicName id="11EF6D9BAC0EFFF350FFD824FB01F89F" box="[949,1160,1795,1820]" class="Actinopterygii" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="44" phylum="Chordata" rank="species" species="yaha">
|
||
<emphasis id="E49BCA0AAC0EFFF350FFD824FB01F89F" box="[949,1160,1795,1820]" italics="true" pageId="11" pageNumber="44">Crenicichla yaha</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<collectingCountry id="AEF85688AC0EFFF357D0D825FA9EF89F" box="[1178,1303,1794,1820]" name="Argentina" pageId="11" pageNumber="44">Argentina</collectingCountry>
|
||
, Misiones. MACN-ict 8924 (
|
||
<typeStatus id="0954A8BAAC0EFFF3522DD80FFE5AF8C1" box="[359,467,1832,1858]" pageId="11" pageNumber="44" type="holotype">holotype</typeStatus>
|
||
),
|
||
<quantity id="1117BBFDAC0EFFF352A2D80FFDEBF8C1" box="[488,610,1832,1858]" metricMagnitude="-1" metricUnit="m" metricValue="1.037" pageId="11" pageNumber="44" unit="mm" value="103.7">103.7 mm</quantity>
|
||
, arroyo Urugua–í in
|
||
<collectingCountry id="AEF85688AC0EFFF3501FD80FFCF6F8C1" box="[853,895,1832,1858]" name="Iceland" pageId="11" pageNumber="44">Isla</collectingCountry>
|
||
Palacios. AI 199, 1 ex.,
|
||
<quantity id="1117BBFDAC0EFFF357D5D80FFA90F8C1" box="[1183,1305,1832,1858]" metricMagnitude="-1" metricUnit="m" metricValue="1.166" pageId="11" pageNumber="44" unit="mm" value="116.6">116.6 mm</quantity>
|
||
, río Iguazú basin, arroyo Benavente. MTD-F 30606 (
|
||
<typeStatus id="0954A8BAAC0EFFF351C9D86AFD66F8E4" box="[643,751,1869,1895]" pageId="11" pageNumber="44" type="paratype">paratype</typeStatus>
|
||
), 1 ex.,
|
||
<quantity id="1117BBFDAC0EFFF35004D86AFC40F8E4" box="[846,969,1869,1895]" metricMagnitude="-1" metricUnit="m" metricValue="1.059" pageId="11" pageNumber="44" unit="mm" value="105.9">105.9 mm</quantity>
|
||
, arroyo Urugua–í in ruta provincial 19, Parque Provincial
|
||
<collectingCountry id="AEF85688AC0EFFF35237D855FDBFF80F" box="[381,566,1906,1932]" name="Falkland Islands" pageId="11" pageNumber="44">Islas Malvinas</collectingCountry>
|
||
. AI 200 (
|
||
<typeStatus id="0954A8BAAC0EFFF351E1D855FC95F80F" box="[683,796,1906,1932]" pageId="11" pageNumber="44" type="paratype">paratype</typeStatus>
|
||
), 1 ex.,
|
||
<quantity id="1117BBFDAC0EFFF350CAD855FC74F80F" box="[896,1021,1906,1932]" metricMagnitude="-1" metricUnit="m" metricValue="1.358" pageId="11" pageNumber="44" unit="mm" value="135.8">135.8 mm</quantity>
|
||
SL, arroyo Uruzú (affluent of A. Urugua–í) in ruta provincial 19, Parque Provincial
|
||
<collectingCountry id="AEF85688AC0EFFF351BCD8BFFC2FF831" box="[758,934,1944,1970]" name="Falkland Islands" pageId="11" pageNumber="44">Islas Malvinas</collectingCountry>
|
||
. AI 202 (
|
||
<typeStatus id="0954A8BAAC0EFFF3575FD8BFFB05F831" box="[1045,1164,1944,1970]" pageId="11" pageNumber="44" type="paratype">paratypes</typeStatus>
|
||
), 4 ex., 1 (C&S)
|
||
<quantity id="1117BBFDAC0EFFF35613D8BFFE8BF854" metricMagnitude="-2" metricUnit="m" metricValue="4.295" metricValueMax="4.85" metricValueMin="3.7399999999999998" pageId="11" pageNumber="44" unit="mm" value="42.95" valueMax="48.5" valueMin="37.4">37.4– 48.5 mm</quantity>
|
||
, arroyo Urugua–í in
|
||
<collectingCountry id="AEF85688AC0EFFF352BCD89AFDA9F854" box="[502,544,1981,2007]" name="Iceland" pageId="11" pageNumber="44">Isla</collectingCountry>
|
||
Palacios.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |