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<bibRefCitation id="A11BFE8BFFA1FF993DBAFA850AEF8F59" author="Ziegler T &amp; Tran VA &amp; Babb RD &amp; Jones TR &amp; Moler PE &amp; Van Devender RW &amp; Nguyen TQ" box="[901,1152,1381,1406]" pageId="17" pageNumber="548" pagination="17 - 26" refId="ref17377" refString="Ziegler T, Tran VA, Babb RD, Jones TR, Moler PE, Van Devender RW, Nguyen TQ. 2019. A new species of reed snake, Calamaria Boie, 1827 from the Central Highlands of Vietnam (Squamata: Colubridae). Rev Suisse Zool. 126 (1): 17 - 26. doi: 10.5281 / zenodo. 2619512" type="journal article" year="2019">Ziegler et al. (2019: 23)</bibRefCitation>
</treatmentCitation>
, 
<treatmentCitation id="442BA56BFFA1FF993E9FFA680FD28F86" author="Poyarkov NA Jr &amp; Van Nguyen T &amp; Orlov NL &amp; Vogel G" box="[160,445,1416,1441]" page="341" pageId="17" pageNumber="548" year="2019">
<bibRefCitation id="A11BFE8BFFA1FF993E9FFA680FD28F86" author="Poyarkov NA Jr &amp; Van Nguyen T &amp; Orlov NL &amp; Vogel G" box="[160,445,1416,1441]" pageId="17" pageNumber="548" pagination="335 - 348" refId="ref16014" refString="Poyarkov NA Jr, Van Nguyen T, Orlov NL, Vogel G. 2019. A new species of the genus Calamaria Boie, 1827 from the highlands of the Langbian Plateau, Southern Vietnam (Squamata: Colubridae). Russ J Herpeto. 26 (6): 335 - 348. doi: 10.30906 / 1026 - 2296 - 2019 - 26 - 6 - 335 - 348" type="journal article" year="2019">
Poyarkov et al. (2019: 
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<emphasis id="F7FE5F68FFA1FF993FFBFA680C638F86" box="[452,524,1416,1441]" italics="true" pageId="17" pageNumber="548">in part</emphasis>
), 
<treatmentCitation id="442BA56BFFA1FF993C20FA680D5E8F86" author="Wallach V &amp; Williams KL &amp; Boundy J" box="[543,817,1416,1441]" page="142" pageId="17" pageNumber="548" year="2014">
<bibRefCitation id="A11BFE8BFFA1FF993C20FA680D5E8F86" author="Wallach V &amp; Williams KL &amp; Boundy J" box="[543,817,1416,1441]" pageId="17" pageNumber="548" refId="ref16724" refString="Wallach V, Williams KL, Boundy J. 2014. Snakes of the world: a catalogue of living and extinct species. Boca Raton: CRC Press; 1209 pp." type="book" year="2014">Wallach et al. (2014: 142</bibRefCitation>
</treatmentCitation>
, 
<emphasis id="F7FE5F68FFA1FF993D05FA680DED8F86" box="[826,898,1416,1441]" italics="true" pageId="17" pageNumber="548">in part</emphasis>
), 
<treatmentCitation id="442BA56BFFA1FF993DAAFA680AEC8F86" author="Weinell J &amp; Leviton AE &amp; Brown RM" box="[917,1155,1416,1441]" page="3" pageId="17" pageNumber="548" year="2021">
<bibRefCitation id="A11BFE8BFFA1FF993DAAFA680AEC8F86" author="Weinell J &amp; Leviton AE &amp; Brown RM" box="[917,1155,1416,1441]" pageId="17" pageNumber="548" pagination="1 - 14" refId="ref16878" refString="Weinell J, Leviton AE, Brown RM. 2021. A new species of reed snake, genus Calamaria (Colubridae: Calamariinae), from Mindoro Island, Philippines. Philipp J Syst Biol. 14 (2): 1 - 14. doi: 10.26757 / pjsb 2020 b 14006" type="journal article" year="2021">
Weinell et al. (2021: 
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</bibRefCitation>
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<emphasis id="F7FE5F68FFA1FF993E9FFA4A0E868FE4" box="[160,233,1450,1475]" italics="true" pageId="17" pageNumber="548">in part</emphasis>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="8D90D0F1FFA1FF9A3E9FFA190F538824" lastPageId="18" lastPageNumber="549" pageId="17" pageNumber="548" type="materials_examined">
<paragraph id="C535837AFFA1FF9A3E9FFA190C878B72" blockId="17.[160,1156,1529,1728]" lastBlockId="18.[160,1156,142,341]" lastPageId="18" lastPageNumber="549" pageId="17" pageNumber="548">
<materialsCitation id="75E28927FFA1FF993E9FFA190AEE8CE7" ID-GBIF-Occurrence="3412503307" collectingDate="1956-05-31" collectingDateMax="1975-06-10" collectingDateMin="1956-05-31" collectionCode="NZMC" collectorName="Alexei K. Zaguliayev" country="China" county="Jingdong Yi Autonomous County" elevation="0" location="in the vicinity of Tszindun, in the gorge of monsoon forest" municipality="Jingdong Yi Autonomous County" pageId="17" pageNumber="548" specimenCode="ZISP 17073" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan Province" typeStatus="holotype">
<emphasis id="F7FE5F68FFA1FF993E9FFA190F7D8C34" bold="true" box="[160,274,1529,1555]" italics="true" pageId="17" pageNumber="548">
<typeStatus id="1A313DD8FFA1FF993E9FFA190F628C34" box="[160,269,1529,1555]" pageId="17" pageNumber="548" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="A39B1BBFFFA1FF993F1EFA1A0F068C34" box="[289,361,1530,1555]" collectionName="NZMC" pageId="17" pageNumber="548">NZMC</collectionCode>
, original catalogue number 
<specimenCode id="952C2B01FFA1FF993C93FA1A0D468C34" box="[684,809,1530,1555]" collectionCode="ZISP" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="17" pageNumber="548">ZISP 17073</specimenCode>
(
<figureCitation id="5DB19FFFFFA1FF993D05FA1A0DFB8C34" box="[826,916,1530,1555]" captionStart="Figure 1" captionStartId="2.[160,225,916,939]" captionTargetBox="[163,1153,142,885]" captionTargetId="figure-192@2.[163,1153,142,885]" captionTargetPageId="2" captionText="Figure 1. (a) Portrait of Soviet herpetologist Sergey A. Chernov, who would describe the type specimen of C. yunnanensis; (b) the original illustration of the holotype of C. yunnanensis (ZISP 17073) from Jingdong, Yunnan Province, China, from Chernov (1962), used with permission; (c) the left lateral view of the head of a referred specimen of C. yunnanensis (ROM 41547) from Simao, Yunnan Province, China; and (d) a photograph of a live C. yunnanensis from Nanjian Yi Autonomous County, Yunnan Province, China exhibiting a different colour pattern on the head compared to the original description. Photo of (a) from the Zoological Institute of the Russian Academy of Sciences archive published in Ananjeva and Doronin (2020), used with permission by Dr. Natalia J. Ananjeva; photo of (c) taken by Justin L. Lee; photo of (d) taken by Chung-Wei Logan You, used with permission." figureDoi="http://doi.org/10.5281/zenodo.5476208" httpUri="https://zenodo.org/record/5476208/files/figure.png" pageId="17" pageNumber="548">Figure 1</figureCitation>
(b)), collected ‘ 
<location id="C055D5A1FFA1FF993A01FA1A0CB68C11" LSID="urn:lsid:plazi:treatment:4D23326CFFA1FF903FA2FBDD0A758B72:C055D5A1FFA1FF993A01FA1A0CB68C11" country="China" county="Jingdong Yi Autonomous County" municipality="Jingdong Yi Autonomous County" name="in the vicinity of Tszindun, in the gorge of monsoon forest" pageId="17" pageNumber="548" stateProvince="Yunnan Province">in the vicinity of Tszindun, in the gorge of monsoon forest</location>
<elevation id="4EA76449FFA1FF993CE1F9FD0D368C11" box="[734,857,1565,1590]" pageId="17" pageNumber="548">1100–1200</elevation>
meters’ [= in the vicinity of Jingdong, Jingdong Yi Autonomous County, southern 
<collectingRegion id="074E4D98FFA1FF993D2BF9DF0DB88C7F" box="[788,983,1599,1624]" country="China" name="Yunnan" pageId="17" pageNumber="548">Yunnan Province</collectingRegion>
, 
<collectingCountry id="BD9DC3EAFFA1FF993DDBF9DF0A488C7F" box="[996,1063,1599,1624]" name="China" pageId="17" pageNumber="548">China</collectingCountry>
] a 
<specimenCount id="D38C48F3FFA1FF993A73F9DF0AEC8C7F" box="[1100,1155,1599,1624]" pageId="17" pageNumber="548" type="male">male</specimenCount>
specimen (maturity unknown) collected on 
<date id="B134A5BAFFA1FF993C9FF9820D588C5C" box="[672,823,1634,1659]" pageId="17" pageNumber="548" value="1956-05-31">
<collectingDate id="A1705C52FFA1FF993C9FF9820D588C5C" box="[672,823,1634,1659]" pageId="17" pageNumber="548" value="1956-05-31">31 May 1956</collectingDate>
</date>
by Soviet Researcher 
<collectorName id="687FE6ACFFA1FF993A00F9820F568CB9" pageId="17" pageNumber="548">Alexei K. Zaguliayev</collectorName>
, as part of the joint Soviet–Chinese field expeditions to Yunnan Province, China. A translation of the original description can be found in the Supplementary data.
</materialsCitation>
<materialsCitation id="75E28927FFA2FF9A3E9FFF6E0A3A8ACA" ID-GBIF-Occurrence="3412503314" collectingDate="1975-05-11" collectingDateMax="1975-06-10" collectingDateMin="1956-05-31" collectionCode="NZMC" collectorName="Topotypic &amp; collector unknown" country="China" county="Jingdong Yi Autonomous County" elevation="0" location="Jingdong Yi Autonomous County" municipality="Jingdong Yi Autonomous County" pageId="18" pageNumber="549" specimenCode="KIZ 054176, 75II0198" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan Province" typeStatus="holotype">
<emphasis id="F7FE5F68FFA2FF9A3E9FFF6E0FCD8A8F" bold="true" box="[160,418,142,168]" italics="true" pageId="18" pageNumber="549">
<collectorName id="687FE6ACFFA2FF9A3E9FFF6E0F7A8A8F" box="[160,277,142,168]" pageId="18" pageNumber="549">Topotypic</collectorName>
specimens.
</emphasis>
(
<figureCitation id="5DB19FFFFFA2FF9A3F85FF6F0C728A8F" box="[442,541,143,168]" captionStart="Figure 5" captionStartId="20.[160,225,1659,1682]" captionTargetBox="[163,1153,142,1627]" captionTargetId="figure-11@20.[163,1153,142,1627]" captionTargetPageId="20" captionText="Figure 5. Examples of Calamaria yunnanensis from the type locality: adult female (KIZ 054176), (a) dorsal and (b) ventral views; adult male (KIZ 056010), (c) dorsal and (d) ventral views; male (KIZ 056009), (e) dorsal view; and immature specimen, sex unknown (KIZ 056011), (f) dorsolateral view. Photographs taken by Shuo Liu, used with permission." figureDoi="http://doi.org/10.5281/zenodo.5476218" httpUri="https://zenodo.org/record/5476218/files/figure.png" pageId="18" pageNumber="549">Figure 5</figureCitation>
). 
<specimenCode id="952C2B01FFA2FF9A3C08FF6F0CD18A8F" box="[567,702,143,168]" collectionCode="KIZ" country="China" httpUri="http://grbio.org/cool/ncht-ribt" name="Kunming Institute of Zoology, Chinese Academy of Sciences" pageId="18" pageNumber="549">KIZ 054176</specimenCode>
(alternate catalogue: 
<specimenCode id="952C2B01FFA2FF9A3D96FF6F0A658A8F" box="[937,1034,143,168]" pageId="18" pageNumber="549">75II0198</specimenCode>
), an 
<specimenCount id="D38C48F3FFA2FF9A3A76FF6F0E838AEC" pageId="18" pageNumber="549" type="adult">adult female</specimenCount>
collected from 
<collectingMunicipality id="25511900FFA2FF9A3F91FF520D568AEC" box="[430,825,178,203]" pageId="18" pageNumber="549">Jingdong Yi Autonomous County</collectingMunicipality>
, southern 
<collectingRegion id="074E4D98FFA2FF9A3D86FF520AEF8AEC" box="[953,1152,178,203]" country="China" name="Yunnan" pageId="18" pageNumber="549">Yunnan Province</collectingRegion>
, 
<collectingCountry id="BD9DC3EAFFA2FF9A3E9FFF340E8F8ACA" box="[160,224,212,237]" name="China" pageId="18" pageNumber="549">China</collectingCountry>
(same locality as the type specimen) on 
<date id="B134A5BAFFA2FF9A3CF6FF340D0F8ACA" box="[713,864,212,237]" pageId="18" pageNumber="549" value="1975-05-11">
<collectingDate id="A1705C52FFA2FF9A3CF6FF340D0F8ACA" box="[713,864,212,237]" pageId="18" pageNumber="549" value="1975-05-11">11 May 1975</collectingDate>
</date>
(
<collectorName id="687FE6ACFFA2FF9A3D4EFF340A248ACA" box="[881,1099,212,237]" pageId="18" pageNumber="549">collector unknown</collectorName>
);
</materialsCitation>
<materialsCitation id="75E28927FFA2FF9A3A5FFF340D748B14" ID-GBIF-Occurrence="3412503318" collectingDate="1975-06-10" collectingDateMax="1975-06-10" collectingDateMin="1956-05-31" collectionCode="NZMC" collectorName="collector unknown" country="China" county="Jingdong Yi Autonomous County" elevation="0" location="Tszindun" municipality="Jingdong Yi Autonomous County" pageId="18" pageNumber="549" specimenCode="KIZ 056010-011, 75II0224-225" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<specimenCode id="952C2B01FFA2FF9A3A5FFF340F478B37" collectionCode="KIZ" country="China" httpUri="http://grbio.org/cool/ncht-ribt" name="Kunming Institute of Zoology, Chinese Academy of Sciences" pageId="18" pageNumber="549">KIZ 056010–011</specimenCode>
(alternate catalogues: 
<specimenCode id="952C2B01FFA2FF9A3C17FF170CAE8B37" box="[552,705,247,272]" pageId="18" pageNumber="549">75II0224–225</specimenCode>
, respectively), 
<specimenCount id="D38C48F3FFA2FF9A3D5BFF170DB88B37" box="[868,983,247,272]" pageId="18" pageNumber="549" type="male">two males</specimenCount>
from the same locality collected on 
<date id="B134A5BAFFA2FF9A3FB0FEFA0C468B14" box="[399,553,282,307]" pageId="18" pageNumber="549" value="1975-06-10">
<collectingDate id="A1705C52FFA2FF9A3FB0FEFA0C468B14" box="[399,553,282,307]" pageId="18" pageNumber="549" value="1975-06-10">10 June 1975</collectingDate>
</date>
(
<collectorName id="687FE6ACFFA2FF9A3C07FEFA0D7E8B14" box="[568,785,282,307]" pageId="18" pageNumber="549">collector unknown</collectorName>
);
</materialsCitation>
<materialsCitation id="75E28927FFA2FF9A3D1BFEFA0C878B72" ID-GBIF-Occurrence="3412503316" collectingDate="1956-05-31" collectingDateMax="1975-06-10" collectingDateMin="1956-05-31" collectionCode="NZMC" collectorName="Soviet Researcher Alexei K. Zaguliayev &amp; Soviet &amp; Chinese &amp; Supplementary &amp; Topotypic" country="China" county="Jingdong Yi Autonomous County" elevation="0" location="Tszindun" municipality="Jingdong Yi Autonomous County" pageId="18" pageNumber="549" specimenCode="KIZ 056009, KIZ 056009-010" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
and 
<specimenCode id="952C2B01FFA2FF9A3D67FEFA0DB58B14" box="[856,986,282,307]" collectionCode="KIZ" country="China" httpUri="http://grbio.org/cool/ncht-ribt" name="Kunming Institute of Zoology, Chinese Academy of Sciences" pageId="18" pageNumber="549">KIZ 056009</specimenCode>
, sex unknown, from the same locality and date as 
<specimenCode id="952C2B01FFA2FF9A3C11FEDC0C8B8B72" box="[558,740,316,341]" collectionCode="KIZ" country="China" httpUri="http://grbio.org/cool/ncht-ribt" name="Kunming Institute of Zoology, Chinese Academy of Sciences" pageId="18" pageNumber="549">KIZ 056009–010</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="C535837AFFA2FF9A3E9FFE610F538824" blockId="18.[160,1156,385,515]" pageId="18" pageNumber="549">
<emphasis id="F7FE5F68FFA2FF9A3E9FFE610F108BBC" bold="true" box="[160,383,385,411]" italics="true" pageId="18" pageNumber="549">Referred specimen.</emphasis>
<materialsCitation id="75E28927FFA2FF9A3FAFFE620F538824" ID-GBIF-Occurrence="3412503320" accessionNumber="ROM 41547" collectingDate="2002-06-14" collectionCode="ROM" collectorName="Robert W. Murphy &amp; Nikolai L. Orlov" country="China" elevation="0" isTailEnumeration="true" location="Simao" pageId="18" pageNumber="549" specimenCount="1" stateProvince="Yunnan Province">
ROM 41547 (
<figureCitation id="5DB19FFFFFA2FF9A3C1FFE620CE98BBC" box="[544,646,386,411]" captionStart="Figure 4" captionStartId="17.[160,225,945,968]" captionTargetBox="[163,1153,142,913]" captionTargetId="figure-284@17.[163,1153,142,914]" captionTargetPageId="17" captionText="Figure 4. Referred specimen of Calamaria yunnanensis (ROM 41547) with (a) dorsal and (b) ventral views of the body, along with (c) right lateral,(d) dorsal and (e) ventral views of the head.Scale bars: a, b = 10 mm; c–e = 5 mm. Photographs taken by Justin L. Lee." figureDoi="http://doi.org/10.5281/zenodo.5476216" httpUri="https://zenodo.org/record/5476216/files/figure.png" pageId="18" pageNumber="549">Figures 4</figureCitation>
and 
<figureCitation id="5DB19FFFFFA2FF9A3C81FE620CA38BBC" box="[702,716,386,411]" captionStart="Figure 6" captionStartId="21.[160,225,916,939]" captionTargetBox="[163,1153,142,885]" captionTargetId="figure-313@21.[163,1153,142,885]" captionTargetPageId="21" captionText="Figure 6. Lateral views of the tail tips of (a) Calamaria nebulosa sp. nov. holotype (FMNH 258666) and (b) Calamaria yunnanensis (ROM 41547). Scale bar: 10 mm. Photographs by Justin L. Lee." figureDoi="http://doi.org/10.5281/zenodo.5476220" httpUri="https://zenodo.org/record/5476220/files/figure.png" pageId="18" pageNumber="549">6</figureCitation>
), a male specimen (maturity unknown) found 
<quantity id="02722E9FFFA2FF9A3ED6FE440F5C8B9A" box="[233,307,420,445]" metricMagnitude="3" metricUnit="m" metricValue="8.8" pageId="18" pageNumber="549" unit="km" value="8.8">
<locationDeviation id="D4879A58FFA2FF9A3ED6FE440F5C8B9A" box="[233,307,420,445]" metricMagnitude="3" metricUnit="m" metricValue="8.8" pageId="18" pageNumber="549" unit="km" value="8.8">8.8 km</locationDeviation>
</quantity>
west of 
<location id="C055D5A1FFA2FF9A3FAFFE440FB78B9A" LSID="urn:lsid:plazi:treatment:4D23326CFFA1FF903FA2FBDD0A758B72:C055D5A1FFA2FF9A3FAFFE440FB78B9A" box="[400,472,420,445]" country="China" name="Simao" pageId="18" pageNumber="549" stateProvince="Yunnan Province">Simao</location>
, 
<collectingRegion id="074E4D98FFA2FF9A3FDEFE440CCF8B9A" box="[481,672,420,445]" country="China" name="Yunnan" pageId="18" pageNumber="549">Yunnan Province</collectingRegion>
, 
<collectingCountry id="BD9DC3EAFFA2FF9A3C95FE440C858B9A" box="[682,746,420,445]" name="China" pageId="18" pageNumber="549">China</collectingCountry>
(
<geoCoordinate id="A0BEE5BDFFA2FF9A3CC9FE440DD88B9A" box="[758,951,420,445]" pageId="18" pageNumber="549">
22° 46 
<emphasis id="F7FE5F68FFA2FF9A3D02FE450D2C8B9B" bold="true" box="[829,835,421,444]" italics="true" pageId="18" pageNumber="549">ʹ</emphasis>
47.928 
<emphasis id="F7FE5F68FFA2FF9A3DACFE450DF68B9B" bold="true" box="[915,921,421,444]" italics="true" pageId="18" pageNumber="549">ʹ</emphasis>
’ N
</geoCoordinate>
, 
<geoCoordinate id="A0BEE5BDFFA2FF9A3DFEFE440EC08BC7" pageId="18" pageNumber="549">
100° 54 
<emphasis id="F7FE5F68FFA2FF9A3A2BFE450A758B9B" bold="true" box="[1044,1050,421,444]" italics="true" pageId="18" pageNumber="549">ʹ</emphasis>
24.0372 
<emphasis id="F7FE5F68FFA2FF9A3A47FE450A118B9B" bold="true" box="[1144,1150,421,444]" italics="true" pageId="18" pageNumber="549">ʹ</emphasis>
’ E
</geoCoordinate>
; 
<elevation id="4EA76449FFA2FF9A3E85FE270F6E8BC7" box="[186,257,455,480]" pageId="18" pageNumber="549">~1270</elevation>
metres above sea level) collected by 
<collectorName id="687FE6ACFFA2FF9A3C94FE270D168BC7" box="[683,889,455,480]" pageId="18" pageNumber="549">Robert W. Murphy</collectorName>
and 
<collectorName id="687FE6ACFFA2FF9A3D8CFE270A318BC7" box="[947,1118,455,480]" pageId="18" pageNumber="549">Nikolai L. Orlov</collectorName>
on 
<date id="B134A5BAFFA2FF9A3E9FFE0A0F578824" box="[160,312,490,515]" pageId="18" pageNumber="549" value="2002-06-14">
<collectingDate id="A1705C52FFA2FF9A3E9FFE0A0F578824" box="[160,312,490,515]" pageId="18" pageNumber="549" value="2002-06-14">14 June 2002</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="8D90D0F1FFA2FF9A3E9FFDCE0C788E77" pageId="18" pageNumber="549" type="diagnosis">
<paragraph id="C535837AFFA2FF9A3E9FFDCE0F7B886F" blockId="18.[160,1156,558,1104]" box="[160,276,558,584]" pageId="18" pageNumber="549">
<emphasis id="F7FE5F68FFA2FF9A3E9FFDCE0F7B886F" bold="true" box="[160,276,558,584]" italics="true" pageId="18" pageNumber="549">Diagnosis</emphasis>
</paragraph>
<paragraph id="C535837AFFA2FF9A3E9FFDB20C788E77" blockId="18.[160,1156,558,1104]" pageId="18" pageNumber="549">
A species of 
<taxonomicName id="028AF8F9FFA2FF9A3F04FDB20FC3884C" baseAuthorityName="Inger and Marx" baseAuthorityYear="1965" box="[315,428,594,619]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="549" phylum="Chordata" rank="genus">
<emphasis id="F7FE5F68FFA2FF9A3F04FDB20FC3884C" box="[315,428,594,619]" italics="true" pageId="18" pageNumber="549">Calamaria</emphasis>
</taxonomicName>
distinguished from all other members of the genus found in mainland Southeast Asia by having the following combination of morphological characters: (1) rostral scale wider than high; (2) portion of rostral scale visible from above less than half the length of the prefrontal suture; (3) six scales and shields surrounding the paraparietal scale; (4) absence of a preocular scale; (5) four supralabials, with the second and third touching the eye; (6) five or six infralabials, the first pair separating the mental scale from the anterior chin shields; (7) 167–184 ventrals in males, 
<quantity id="02722E9FFFA2FF9A3DA1FCC20D8A891C" box="[926,997,802,827]" metricMagnitude="0" metricUnit="m" metricValue="5.0546" pageId="18" pageNumber="549" unit="in" value="199.0">199 in</quantity>
female, three gular scales; (8) 15–20 subcaudals in males, 
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female, all paired; (9) dorsal colour pattern variable, usually bluish-grey or olive-brown with indistinct dark brown longitudinal stripes across each dorsal scale row forming a network-like pattern, sometimes restricted to the flanks; (10) ventral surface red or yellow usually with dark brown margins on the ventral scales and a black midventral stripe under the tail, completely immaculate in some specimens; and (11) a light yellow nuchal collar posterior to the head usually present, absent in some specimens. A summary of diagnostic characters and comparisons is given below and in 
<tableCitation id="8808B6C1FFA2FF9A3FA6FBD70C7F8E77" box="[409,528,1079,1104]" captionStart-0="Table 3" captionStart-1="Table 4" captionStartId-0="10.[133,189,442,465]" captionStartId-1="12.[133,189,438,461]" captionTargetBox-0="[144,1684,565,1386]" captionTargetBox-1="[143,1691,532,1389]" captionTargetPageId-0="10" captionTargetPageId-1="12" captionText-0="Table 3. Morphological comparisons between Calamaria nebulosa sp. nov. and specimens of Calamaria yunnanensis. Data with asterisks are adapted from Zhao et al. (1998) and Yang and Rao (2008). Abbreviations are noted in the materials and methods, except for DSR6R, which stands for the number of dorsal scales reducing to six rows above position of subcaudal anterior to tail tip; DSR4R, which stands for the number of dorsal scales reducing to four rows above position of subcaudal anterior to tail tip; and ‘SC’, which stands for subcaudal. Entries marked with ‘ – ’ are not available." captionText-1="Table 4. Morphological comparisons between Calamaria nebulosa sp. nov. (FMNH 258666), Calamaria yunnanensis and all species of Calamaria known from mainland Southeast Asia (see Materials and methods for list of literature sources). Symbols used in this table include: (M) = male; (F) = female; (m) = modified maxillary teeth." httpUri-0="http://table.plazi.org/id/91F5D3F2FFBAFF823EBAFE5A0AA8880E" httpUri-1="http://table.plazi.org/id/91F5D3F2FFBCFF843EBAFE560F70882F" pageId="18" pageNumber="549" tableUuid-0="91F5D3F2FFBAFF823EBAFE5A0AA8880E" tableUuid-1="91F5D3F2FFBCFF843EBAFE560F70882F">Tables 3–4</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8D90D0F1FFA2FF9E3E9FFB9C0DBA8FA0" lastPageId="22" lastPageNumber="553" pageId="18" pageNumber="549" type="description">
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<emphasis id="F7FE5F68FFA2FF9A3E9FFB9C0C3F8EB1" bold="true" box="[160,592,1148,1174]" italics="true" pageId="18" pageNumber="549">Description of the referred specimen</emphasis>
</paragraph>
<paragraph id="C535837AFFA2FF9A3E9FFB7F0A268F8C" blockId="18.[160,1156,1148,1728]" pageId="18" pageNumber="549">
Male (maturity unknown) in good condition, with one incision at the tail base and another along the posterior portion of body. SVL 
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, TailL 
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, TotalL 
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, HeadL 
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(
<quantity id="02722E9FFFA2FF9A3F36FB050F0C8ED9" box="[265,355,1253,1278]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="18" pageNumber="549" unit="mm" value="6.4">6.4 mm</quantity>
measured from the parietal tip), HeadW 4.0 mm, SnL 
<quantity id="02722E9FFFA2FF9A3DD7FB050A2C8ED9" box="[1000,1091,1253,1278]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="18" pageNumber="549" unit="mm" value="2.6">2.6 mm</quantity>
, SnW 2.0 mm, EyeD 1.0 mm, NarEye 
<quantity id="02722E9FFFA2FF9A3FC5FAE70C3D8F07" box="[506,594,1287,1312]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="18" pageNumber="549" unit="mm" value="1.5">1.5 mm</quantity>
, IOD 
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. Body height at midbody 
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, at tail base 
<quantity id="02722E9FFFA2FF9A3F3CFACA0F368F64" box="[259,345,1322,1347]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="18" pageNumber="549" unit="mm" value="3.3">3.3 mm</quantity>
; eye–mouth distance 
<quantity id="02722E9FFFA2FF9A3C73FACA0CCD8F64" box="[588,674,1322,1347]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="18" pageNumber="549" unit="mm" value="0.8">0.8 mm</quantity>
; rostral scale width 
<quantity id="02722E9FFFA2FF9A3D43FACA0DBD8F64" box="[892,978,1322,1347]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="18" pageNumber="549" unit="mm" value="0.9">0.9 mm</quantity>
, height 
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; prefrontal scale length 
<quantity id="02722E9FFFA2FF9A3F9EFAAD0F998F41" box="[417,502,1357,1382]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="18" pageNumber="549" unit="mm" value="1.8">1.8 mm</quantity>
, suture 1.0 mm; frontal scale length 
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, width 
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; parietal scale length 
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, suture 
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. TailL/TotalL 0.081; HeadL/W 1.6; HeadL/SVL 2.83; SnL/HeadL 0.41; EyeD/SnL 0.38; NarEye/SnL 0.58; SnW/IOD 0.66; IOD/HW 0.76.
</paragraph>
<paragraph id="C535837AFFA2FF9B3EFFFA550D7088F5" blockId="18.[160,1156,1148,1728]" lastBlockId="19.[160,1156,142,861]" lastPageId="19" lastPageNumber="550" pageId="18" pageNumber="549">Body elongate and vermiform, round in cross section; tail thick, roughly equal in diameter to rest of body; posterior portion of tail slightly tapering, then abruptly tapering at the tip, which appears obtusely pointed in dorsal view (weakly acute), round in lateral view; head sub-oblong in dorsal view, indistinct from neck; snout blunt, weakly depressed posteriorly; nostrils small and elliptical; eyes small, round, diameter larger than eye– mouth distance; pupil indistinct, round; rostral scale triangular in frontal view, wider than high, portion visible from above roughly 4/5 the length of the prefrontal suture, suture bordering prefrontal and rostral scale deep-‘V’-shaped; prefrontal scales subhexagonal, shorter than frontal; frontal scale shield-shaped, hexagonal, 1.3× longer than wide, around 1.4× longer than parietal suture; posterior angle formed by the frontal/ parietal sutures producing the posterior vertex of the frontal right (~90°); parietals subhexagonal, longer than wide, longer than frontal; parietal suture 1.4× longer than prefrontal suture; anterior parietal angle formed by the sutures between the parietal/frontal and the suture between the supraocular/parietal broadly obtuse, pointing laterally (~127°); supraoculars subrectangular, width of frontal around 2.7× maximum width of supraoculars; paraparietal shaped like a temporal scale with 6/6 scales and shields in contact; postocular scale rectangular, not as high as eye diameter; preocular absent; nasal scale small, subrectangular, barely surrounding nostrils; supralabials 4/4, 1st and 2nd smallest, 1st in contact with nasal, 1st and 2nd in contact with prefrontal, 2nd and 3rd in contact with eye, 4th largest and rectangular; infralabials 5/5, 1st smallest and blocking mental from contacting upper genials, 1st–3rd in contact with upper genials; 3rd–4th in contact with lower genials, 5th largest; mental triangular, unremarkable; upper genial scales paired and rectangular, completely in contact; lower genials in contact anteriorly and separated at midsection by the first gular scale, posterior portion round-pointed; three gular scales before first ventral scale, all rhomboid.</paragraph>
<paragraph id="C535837AFFA3FF9B3EFFFD3C0DC6897A" blockId="19.[160,1156,142,861]" pageId="19" pageNumber="550">Dorsal scale rows smooth, in 13–13–13 rows across entire body; dorsal scales reducing to six rows above the 3rd subcaudal scale, reducing to four scales above the 16th (third-tolast) subcaudal scale; ventrals 167; subcaudals 19, all paired; cloacal plate single; total body scales 187; subcaudal ratio 0.102; maxillary teeth 8/8, modified.</paragraph>
<paragraph id="C535837AFFA3FF9B3E9FFC690D628984" blockId="19.[160,1156,905,1589]" box="[160,781,905,931]" pageId="19" pageNumber="550">
<emphasis id="F7FE5F68FFA3FF9B3E9FFC690D628984" bold="true" box="[160,781,905,931]" italics="true" pageId="19" pageNumber="550">Colouration of the referred specimen in preservative</emphasis>
</paragraph>
<paragraph id="C535837AFFA3FF9B3E9FFC4C0F5C8C12" blockId="19.[160,1156,905,1589]" pageId="19" pageNumber="550">After formalin fixation and preservation in 70% ethanol for 18 years, the dorsum is greybrown and highly iridescent; margins of all dorsal scales dark brown with faint light brown vermiculations on the medial portion of most dorsal scales; dark margins of dorsal scales forming a weak network-like pattern; along flanks, three dark brown longitudinal stripes, lower two stripes 0.5–1.0 dorsal scale rows wide, separated from each other by 0.5 dorsal scale rows; first two stripes on flanks broader (one dorsal scales wide) than highest stripe (0.5 dorsal scale rows wide) along the anterior portion of body; highest stripe two dorsal scales above lower stripes, reduced to one dorsal scale row at midbody; medial portion between stripes filled with light white vermiculations; all longitudinal stripes fade posteriorly, except for one pair that continues onto the posterior portion of tail; dorsal portion of head dark brown with occasional light brown hueing along cephalic scales; nuchal collar whitish-tan, around one dorsal scale in width at vertebrum then widening in lateral view, originating at the third and fourth row of dorsal scales; supralabials and nasal tannish, fourth supralabial mostly white; flanks along nape also white, connecting to nuchal collar; eye blue, pupil greyish-blue; underside of head cream with grey-brown mottling scattered along mental, infralabials, upper genials and nuchal region of first dorsal scales; venter white, margins dark brown; underside of tail with small dark spotting along margins and dark brown midventral stripe starting from tail tip, breaking up near the start of the cloacal plate.</paragraph>
<paragraph id="C535837AFFA3FF9B3E9FF9810D7A8C5C" blockId="19.[160,1155,1633,1728]" box="[160,789,1633,1659]" pageId="19" pageNumber="550">
<emphasis id="F7FE5F68FFA3FF9B3E9FF9810D7A8C5C" bold="true" box="[160,789,1633,1659]" italics="true" pageId="19" pageNumber="550">Description of the topotypic specimens and variation</emphasis>
</paragraph>
<paragraph id="C535837AFFA3FF9D3E9FF9650A298C93" blockId="19.[160,1155,1633,1728]" lastBlockId="21.[160,1156,1032,1716]" lastPageId="21" lastPageNumber="552" pageId="19" pageNumber="550">
<bibRefCitation id="A11BFE8BFFA3FF9B3E9FF9650F318CB9" author="Zhao EM &amp; Huang MH &amp; Zong Y &amp; Zheng J &amp; Huang ZJ &amp; Yang D &amp; Li DJ" box="[160,350,1669,1694]" pageId="19" pageNumber="550" refId="ref17192" refString="Zhao EM, Huang MH, Zong Y, Zheng J, Huang ZJ, Yang D, Li DJ, Eds. 1998. Fauna Sinica. Reptilia Vol. 3. Squamata Serpentes. Beijing: Science Press; p. i - xvii + 1 - 522, Pls. I - VIII, Col. Pls. I - IV. [in Chinese]." type="book" year="1998">Zhao et al. (1998)</bibRefCitation>
and 
<bibRefCitation id="A11BFE8BFFA3FF9B3FACF9650C188CB9" author="Yang D &amp; Rao DQ" box="[403,631,1669,1694]" pageId="19" pageNumber="550" refId="ref16995" refString="Yang D, Rao DQ. 2008. Amphibians and reptiles of Yunnan. Kunming: Yunnan Science and Technology Press; 409 pp. [in Chinese]." type="book" year="2008">Yang and Rao (2008)</bibRefCitation>
published descriptions of 
<taxonomicName id="028AF8F9FFA3FF9B3DA5F9650A548CB9" authorityName=": Chernov" authorityYear="1962" box="[922,1083,1669,1694]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="550" phylum="Chordata" rank="species" species="yunnanensis">
<emphasis id="F7FE5F68FFA3FF9B3DA5F9650A548CB9" box="[922,1083,1669,1694]" italics="true" pageId="19" pageNumber="550">C. yunnanensis</emphasis>
</taxonomicName>
based on the same three topotypic specimens (
<specimenCount id="D38C48F3FFA3FF9B3C58F9470CAF8CE7" box="[615,704,1703,1728]" pageId="19" pageNumber="550" type="male">2 males</specimenCount>
, 
<specimenCount id="D38C48F3FFA3FF9B3CF6F9470D438CE7" box="[713,812,1703,1728]" pageId="19" pageNumber="550" type="female">1 female</specimenCount>
). Although they originate from the 
<typeStatus id="1A313DD8FFA5FF9D3EF4FBE80E928E06" box="[203,253,1032,1057]" pageId="21" pageNumber="552">type</typeStatus>
locality, the written descriptions of these specimens by these authors have slight differences from 
<bibRefCitation id="A11BFE8BFFA5FF9D3F5EFBCB0C628E63" author="Chernov SA" box="[353,525,1067,1092]" pageId="21" pageNumber="552" pagination="382 - 384" refId="ref14538" refString="Chernov SA. 1962. On one yet unknown snake of the genus Calamaria from Yunnan (Scientific results of the Chinese-Soviet expeditions in 1955 - 1957 to South-Western China). Proc Zool Inst Russ Acad Sci USSR, Leningrad. 30: 382 - 384. [in Russian]." type="journal article" year="1962">Chernov (1962)</bibRefCitation>
’s original description and from ROM 41547. 
<bibRefCitation id="A11BFE8BFFA5FF9D3A34FBCB0E898E41" author="Zhao EM &amp; Huang MH &amp; Zong Y &amp; Zheng J &amp; Huang ZJ &amp; Yang D &amp; Li DJ" pageId="21" pageNumber="552" refId="ref17192" refString="Zhao EM, Huang MH, Zong Y, Zheng J, Huang ZJ, Yang D, Li DJ, Eds. 1998. Fauna Sinica. Reptilia Vol. 3. Squamata Serpentes. Beijing: Science Press; p. i - xvii + 1 - 522, Pls. I - VIII, Col. Pls. I - IV. [in Chinese]." type="book" year="1998">Zhao et al. (1998)</bibRefCitation>
indicated that the midventral stripe and nuchal collar in 
<taxonomicName id="028AF8F9FFA5FF9D3D40FBAD0A4B8E41" authorityName=": Chernov" authorityYear="1962" box="[895,1060,1101,1126]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="552" phylum="Chordata" rank="species" species="yunnanensis">
<emphasis id="F7FE5F68FFA5FF9D3D40FBAD0A4B8E41" box="[895,1060,1101,1126]" italics="true" pageId="21" pageNumber="552">C. yunnanensis</emphasis>
</taxonomicName>
may be present or absent, while 
<bibRefCitation id="A11BFE8BFFA5FF9D3F8CFB900CF48EAE" author="Yang D &amp; Rao DQ" box="[435,667,1136,1161]" pageId="21" pageNumber="552" refId="ref16995" refString="Yang D, Rao DQ. 2008. Amphibians and reptiles of Yunnan. Kunming: Yunnan Science and Technology Press; 409 pp. [in Chinese]." type="book" year="2008">Yang and Rao (2008)</bibRefCitation>
did not note the presence of either feature. Photographs of these specimens from the KIZ collection allow me to clarify this issue (
<figureCitation id="5DB19FFFFFA5FF9D3E97FB550F668EE9" box="[168,265,1205,1230]" captionStart="Figure 5" captionStartId="20.[160,225,1659,1682]" captionTargetBox="[163,1153,142,1627]" captionTargetId="figure-11@20.[163,1153,142,1627]" captionTargetPageId="20" captionText="Figure 5. Examples of Calamaria yunnanensis from the type locality: adult female (KIZ 054176), (a) dorsal and (b) ventral views; adult male (KIZ 056010), (c) dorsal and (d) ventral views; male (KIZ 056009), (e) dorsal view; and immature specimen, sex unknown (KIZ 056011), (f) dorsolateral view. Photographs taken by Shuo Liu, used with permission." figureDoi="http://doi.org/10.5281/zenodo.5476218" httpUri="https://zenodo.org/record/5476218/files/figure.png" pageId="21" pageNumber="552">Figure 5</figureCitation>
; Shuo Liu, pers. comm.) The official museum catalogue numbers for these specimens are different than those given in 
<bibRefCitation id="A11BFE8BFFA5FF9D3C89FB380DDF8ED6" author="Yang D &amp; Rao DQ" box="[694,944,1240,1265]" pageId="21" pageNumber="552" refId="ref16995" refString="Yang D, Rao DQ. 2008. Amphibians and reptiles of Yunnan. Kunming: Yunnan Science and Technology Press; 409 pp. [in Chinese]." type="book" year="2008">Yang and Rao (2008)</bibRefCitation>
, but they can be associated with their specimen numbers based on scale counts and body measurements. In the single female specimen (KIZ 054176), the distinct nuchal collar and midventral stripe on the tail is absent. The dorsum is brown, becoming darker posteriorly near the tail, with indistinct longitudinal stripes along the edges of the dorsal scales forming a ‘net-like’ pattern. The venter is immaculate yellow, with the light pigment extending onto the first outer row of dorsal scales. 
<bibRefCitation id="A11BFE8BFFA5FF9D3FDDFA480CDC8FE6" author="Zhao EM &amp; Huang MH &amp; Zong Y &amp; Zheng J &amp; Huang ZJ &amp; Yang D &amp; Li DJ" box="[482,691,1448,1473]" pageId="21" pageNumber="552" refId="ref17192" refString="Zhao EM, Huang MH, Zong Y, Zheng J, Huang ZJ, Yang D, Li DJ, Eds. 1998. Fauna Sinica. Reptilia Vol. 3. Squamata Serpentes. Beijing: Science Press; p. i - xvii + 1 - 522, Pls. I - VIII, Col. Pls. I - IV. [in Chinese]." type="book" year="1998">Zhao et al. (1998)</bibRefCitation>
and 
<bibRefCitation id="A11BFE8BFFA5FF9D3CCBFA480D858FE6" author="Yang D &amp; Rao DQ" box="[756,1002,1448,1473]" pageId="21" pageNumber="552" refId="ref16995" refString="Yang D, Rao DQ. 2008. Amphibians and reptiles of Yunnan. Kunming: Yunnan Science and Technology Press; 409 pp. [in Chinese]." type="book" year="2008">Yang and Rao (2008)</bibRefCitation>
also provide conflicting data on the number of ventrals and subcaudals for this specimen, with the former reporting 198 ventrals and 19 subcaudals and the latter reporting 201 ventrals with 21 subcaudals. A re-count of the scales for this specimen indicates that it has 199 ventrals and 19 subcaudals, agreeing more with 
<bibRefCitation id="A11BFE8BFFA5FF9D3CD4F9D30DAE8C6B" author="Zhao EM &amp; Huang MH &amp; Zong Y &amp; Zheng J &amp; Huang ZJ &amp; Yang D &amp; Li DJ" box="[747,961,1587,1612]" pageId="21" pageNumber="552" refId="ref17192" refString="Zhao EM, Huang MH, Zong Y, Zheng J, Huang ZJ, Yang D, Li DJ, Eds. 1998. Fauna Sinica. Reptilia Vol. 3. Squamata Serpentes. Beijing: Science Press; p. i - xvii + 1 - 522, Pls. I - VIII, Col. Pls. I - IV. [in Chinese]." type="book" year="1998">Zhao et al. (1998)</bibRefCitation>
. This number is significantly higher than for the male specimens of 
<taxonomicName id="028AF8F9FFA5FF9D3CD7F9B50DE48C49" authorityName=": Chernov" authorityYear="1962" box="[744,907,1621,1646]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="552" phylum="Chordata" rank="species" species="yunnanensis">
<emphasis id="F7FE5F68FFA5FF9D3CD7F9B50DE48C49" box="[744,907,1621,1646]" italics="true" pageId="21" pageNumber="552">C. yunnanensis</emphasis>
</taxonomicName>
(167–184), but sexual dimorphism in the number of scale counts is common in 
<taxonomicName id="028AF8F9FFA5FF9D3D1CF9980DFB8CB6" baseAuthorityName="Inger and Marx" baseAuthorityYear="1965" box="[803,916,1656,1681]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="552" phylum="Chordata" rank="genus">
<emphasis id="F7FE5F68FFA5FF9D3D1CF9980DFB8CB6" box="[803,916,1656,1681]" italics="true" pageId="21" pageNumber="552">Calamaria</emphasis>
</taxonomicName>
, and most females in the genus have a higher number of ventrals than the males (
<bibRefCitation id="A11BFE8BFFA5FF9D3D71F97B0A568C93" author="Inger RF &amp; Marx H" box="[846,1081,1691,1716]" pageId="21" pageNumber="552" pagination="1 - 304" refId="ref15062" refString="Inger RF, Marx H. 1965. The systematics and evolution of the oriental colubrid snakes of the genus Calamaria. Fieldiana Zool. 49: 1 - 304." type="journal article" year="1965">Inger and Marx 1965</bibRefCitation>
).
</paragraph>
<caption id="91F5D3F2FFA4FF9C3E9FF99B0CD18CCD" ID-DOI="http://doi.org/10.5281/zenodo.5476218" ID-Zenodo-Dep="5476218" httpUri="https://zenodo.org/record/5476218/files/figure.png" pageId="20" pageNumber="551" startId="20.[160,225,1659,1682]" targetBox="[163,1153,142,1627]" targetPageId="20">
<paragraph id="C535837AFFA4FF9C3E9FF99B0CD18CCD" blockId="20.[160,1157,1659,1770]" pageId="20" pageNumber="551">
<emphasis id="F7FE5F68FFA4FF9C3E9FF99B0E938CB5" bold="true" box="[160,252,1659,1682]" pageId="20" pageNumber="551">Figure 5.</emphasis>
Examples of 
<taxonomicName id="028AF8F9FFA4FF9C3FB9F99B0C088CB5" authorityName=": Chernov" authorityYear="1962" box="[390,615,1659,1682]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="551" phylum="Chordata" rank="species" species="yunnanensis">
<emphasis id="F7FE5F68FFA4FF9C3FB9F99B0C088CB5" box="[390,615,1659,1682]" italics="true" pageId="20" pageNumber="551">Calamaria yunnanensis</emphasis>
</taxonomicName>
from the type locality: adult female (KIZ 054176), (a) dorsal and (b) ventral views; adult male (KIZ 056010), (c) dorsal and (d) ventral views; male (KIZ 056009), (e) dorsal view; and immature specimen, sex unknown (KIZ 056011), (f) dorsolateral view. Photographs taken by Shuo Liu, used with permission.
</paragraph>
</caption>
<caption id="91F5D3F2FFA5FF9D3E9FFC740A7D89EE" ID-DOI="http://doi.org/10.5281/zenodo.5476220" ID-Zenodo-Dep="5476220" httpUri="https://zenodo.org/record/5476220/files/figure.png" pageId="21" pageNumber="552" startId="21.[160,225,916,939]" targetBox="[163,1153,142,885]" targetPageId="21">
<paragraph id="C535837AFFA5FF9D3E9FFC740A7D89EE" blockId="21.[160,1155,916,969]" pageId="21" pageNumber="552">
<emphasis id="F7FE5F68FFA5FF9D3E9FFC740E96898C" bold="true" box="[160,249,916,939]" pageId="21" pageNumber="552">Figure 6.</emphasis>
Lateral views of the tail tips of (a) 
<taxonomicName id="028AF8F9FFA5FF9D3C73FC740D65898C" authority="Lee, 2021" authorityName="Lee" authorityYear="2021" box="[588,778,916,939]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="552" phylum="Chordata" rank="species" species="nebulosa" status="sp. nov.">
<emphasis id="F7FE5F68FFA5FF9D3C73FC740D65898C" box="[588,778,916,939]" italics="true" pageId="21" pageNumber="552">Calamaria nebulosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ECCDE213FFA5FF9D3D2FFC740D32898C" box="[784,861,916,939]" pageId="21" pageNumber="552" rank="species">sp. nov.</taxonomicNameLabel>
holotype (FMNH 258666) and (b) 
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<emphasis id="F7FE5F68FFA5FF9D3EFDFC520FCD89EE" box="[194,418,946,969]" italics="true" pageId="21" pageNumber="552">Calamaria yunnanensis</emphasis>
</taxonomicName>
(ROM 41547). Scale bar: 10 mm. Photographs by Justin L. Lee.
</paragraph>
</caption>
<paragraph id="C535837AFFA6FF9E3EFFFF6E0A0788F5" blockId="22.[160,1156,142,1415]" pageId="22" pageNumber="553">
In the other specimens (KIZ 056009–010), both males, the midventral stripe is present on the tail and a nuchal collar is present in one of the specimens (KIZ 056010). The other specimen (KIZ 056009) is partially destroyed with only the midbody and tail remaining, so it is uncertain whether or not it had a nuchal collar. Both specimens also have some dark brown vermiculate-shaped spots present on the underside of the tail in addition to the midventral stripe (KIZ 056009–010). It is unknown whether or not the 
<typeStatus id="1A313DD8FFA6FF9E3DE3FEDC0A628B72" box="[988,1037,316,341]" pageId="22" pageNumber="553">type</typeStatus>
specimen contained this characteristic, but a few small dark vermiculations are present on ROM 41547. Otherwise, the colouration of these 
<specimenCount id="D38C48F3FFA6FF9E3CBDFE610D578BBD" box="[642,824,385,410]" pageId="22" pageNumber="553" type="generic">three specimens</specimenCount>
agrees with 
<bibRefCitation id="A11BFE8BFFA6FF9E3DF5FE610A1C8BBD" author="Chernov SA" box="[970,1139,385,410]" pageId="22" pageNumber="553" pagination="382 - 384" refId="ref14538" refString="Chernov SA. 1962. On one yet unknown snake of the genus Calamaria from Yunnan (Scientific results of the Chinese-Soviet expeditions in 1955 - 1957 to South-Western China). Proc Zool Inst Russ Acad Sci USSR, Leningrad. 30: 382 - 384. [in Russian]." type="journal article" year="1962">Chernov (1962)</bibRefCitation>
’s original description of 
<taxonomicName id="028AF8F9FFA6FF9E3FA0FE440C2F8B9A" authorityName=": Chernov" authorityYear="1962" box="[415,576,420,445]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="553" phylum="Chordata" rank="species" species="yunnanensis">
<emphasis id="F7FE5F68FFA6FF9E3FA0FE440C2F8B9A" box="[415,576,420,445]" italics="true" pageId="22" pageNumber="553">C. yunnanensis</emphasis>
</taxonomicName>
and with ROM 41547, with the only differing characteristic being the presence of indistinct dark brown longitudinal stripes on the dorsum. 
<bibRefCitation id="A11BFE8BFFA6FF9E3E9FFE090F288825" author="Chernov SA" box="[160,327,489,514]" pageId="22" pageNumber="553" pagination="382 - 384" refId="ref14538" refString="Chernov SA. 1962. On one yet unknown snake of the genus Calamaria from Yunnan (Scientific results of the Chinese-Soviet expeditions in 1955 - 1957 to South-Western China). Proc Zool Inst Russ Acad Sci USSR, Leningrad. 30: 382 - 384. [in Russian]." type="journal article" year="1962">Chernov (1962)</bibRefCitation>
did not note this feature in his description but stated that the margins of the dorsal scales were often dark brown and ‘continuous’, suggesting that there were indistinct stripes as described in these other topotypic specimens. Indeed, the striped pattern is barely visible on some the other topotypic specimens in the KIZ collection due to their state of preservation, and this artefact could have caused Chernov to miss this character when he was writing his description. Another characteristic shared by these specimens is the presence of dark brown margins along the ventral scales, which is shared with ROM 41547.
</paragraph>
<paragraph id="C535837AFFA6FF9E3EFFFD3C0FC68E2D" blockId="22.[160,1156,142,1415]" pageId="22" pageNumber="553">
The scale counts for both specimens are identical to those given by 
<bibRefCitation id="A11BFE8BFFA6FF9E3DFFFD3C0AEB88D2" author="Zhao EM &amp; Huang MH &amp; Zong Y &amp; Zheng J &amp; Huang ZJ &amp; Yang D &amp; Li DJ" box="[960,1156,732,757]" pageId="22" pageNumber="553" refId="ref17192" refString="Zhao EM, Huang MH, Zong Y, Zheng J, Huang ZJ, Yang D, Li DJ, Eds. 1998. Fauna Sinica. Reptilia Vol. 3. Squamata Serpentes. Beijing: Science Press; p. i - xvii + 1 - 522, Pls. I - VIII, Col. Pls. I - IV. [in Chinese]." type="book" year="1998">Zhao et al. (1998)</bibRefCitation>
and 
<bibRefCitation id="A11BFE8BFFA6FF9E3EEEFD1E0FD58930" author="Yang D &amp; Rao DQ" box="[209,442,766,791]" pageId="22" pageNumber="553" refId="ref16995" refString="Yang D, Rao DQ. 2008. Amphibians and reptiles of Yunnan. Kunming: Yunnan Science and Technology Press; 409 pp. [in Chinese]." type="book" year="2008">Yang and Rao (2008)</bibRefCitation>
, except KIZ 056009 has 20 subcaudal scales instead of 22 scales as reported by these authors. A fourth specimen (KIZ 056011), unreported in either study, was also found in the collection. It is an immature individual and its sex cannot be accurately determined, but it clearly possesses the same colour pattern characteristics as KIZ 056009–010 and ROM 41547. Overall, the colour polymorphism observed in these topotypic specimens agrees with the description provided by 
<bibRefCitation id="A11BFE8BFFA6FF9E3D52FC4C0A5689E2" author="Zhao EM &amp; Huang MH &amp; Zong Y &amp; Zheng J &amp; Huang ZJ &amp; Yang D &amp; Li DJ" box="[877,1081,940,965]" pageId="22" pageNumber="553" refId="ref17192" refString="Zhao EM, Huang MH, Zong Y, Zheng J, Huang ZJ, Yang D, Li DJ, Eds. 1998. Fauna Sinica. Reptilia Vol. 3. Squamata Serpentes. Beijing: Science Press; p. i - xvii + 1 - 522, Pls. I - VIII, Col. Pls. I - IV. [in Chinese]." type="book" year="1998">Zhao et al. (1998)</bibRefCitation>
, while 
<bibRefCitation id="A11BFE8BFFA6FF9E3E9FFC2E0FE089C0" author="Yang D &amp; Rao DQ" box="[160,399,974,999]" pageId="22" pageNumber="553" refId="ref16995" refString="Yang D, Rao DQ. 2008. Amphibians and reptiles of Yunnan. Kunming: Yunnan Science and Technology Press; 409 pp. [in Chinese]." type="book" year="2008">Yang and Rao (2008)</bibRefCitation>
’s omission of these characteristics may have caused confusion in diagnosing the species.
</paragraph>
<paragraph id="C535837AFFA6FF9E3EFFFBF40DBA8FA0" blockId="22.[160,1156,142,1415]" pageId="22" pageNumber="553">
Photographs of a live 
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<emphasis id="F7FE5F68FFA6FF9E3F95FBF40C288E0A" box="[426,583,1044,1069]" italics="true" pageId="22" pageNumber="553">C. yunnanensis</emphasis>
</taxonomicName>
shown in 
<bibRefCitation id="A11BFE8BFFA6FF9E3C8BFBF40DC58E0A" author="Yang JH &amp; Zheng X" box="[692,938,1044,1069]" pageId="22" pageNumber="553" pagination="485 - 491" refId="ref17025" refString="Yang JH, Zheng X. 2018. A new species of the genus Calamaria (Squamata: Colubridae) from Yunnan Province, China. Copeia. 106 (3): 485 - 491. doi: 10.1643 / CH- 17 - 663" type="journal article" year="2018">Yang and Zheng (2018</bibRefCitation>
, fig. 4) and in 
<figureCitation id="5DB19FFFFFA6FF9E3A02FBF40EC18E68" captionStart="Figure 1" captionStartId="2.[160,225,916,939]" captionTargetBox="[163,1153,142,885]" captionTargetId="figure-192@2.[163,1153,142,885]" captionTargetPageId="2" captionText="Figure 1. (a) Portrait of Soviet herpetologist Sergey A. Chernov, who would describe the type specimen of C. yunnanensis; (b) the original illustration of the holotype of C. yunnanensis (ZISP 17073) from Jingdong, Yunnan Province, China, from Chernov (1962), used with permission; (c) the left lateral view of the head of a referred specimen of C. yunnanensis (ROM 41547) from Simao, Yunnan Province, China; and (d) a photograph of a live C. yunnanensis from Nanjian Yi Autonomous County, Yunnan Province, China exhibiting a different colour pattern on the head compared to the original description. Photo of (a) from the Zoological Institute of the Russian Academy of Sciences archive published in Ananjeva and Doronin (2020), used with permission by Dr. Natalia J. Ananjeva; photo of (c) taken by Justin L. Lee; photo of (d) taken by Chung-Wei Logan You, used with permission." figureDoi="http://doi.org/10.5281/zenodo.5476208" httpUri="https://zenodo.org/record/5476208/files/figure.png" pageId="22" pageNumber="553">Figure 1</figureCitation>
offer some insight into the live colouration of this species. The specimen agrees with the colour pattern observed in KIZ 054176. However, the dorsum is predominately olive-brown, becoming dark grey-brown posteriorly, with well-defined dark brown longitudinal stripes across the body forming a network-like pattern on the dorsum. The venter in this specimen is a bright coral red, extending onto the lower flanks of the first two dorsal scale rows and on the labial region of the head and nostrils. A streak of red pigment forms a small marking posterior to the parietals but is too indistinct to be considered a nuchal collar, as it is nowhere near as pronounced as the markings present in other specimens. Red colouration may have been present on the ventral surface of other topotypic 
<taxonomicName id="028AF8F9FFA6FF9E3D55FAAC0A678F42" authorityName=": Chernov" authorityYear="1962" box="[874,1032,1356,1381]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="553" phylum="Chordata" rank="species" species="yunnanensis">
<emphasis id="F7FE5F68FFA6FF9E3D55FAAC0A678F42" box="[874,1032,1356,1381]" italics="true" pageId="22" pageNumber="553">C. yunnanensis</emphasis>
</taxonomicName>
specimens and ROM 41547, but this pigment tends to fade quickly after preservation.
</paragraph>
</subSubSection>
<subSubSection id="8D90D0F1FFA6FF9F3E9FFA540CE98C23" lastPageId="23" lastPageNumber="554" pageId="22" pageNumber="553" type="distribution">
<paragraph id="C535837AFFA6FF9E3E9FFA540C768FE9" blockId="22.[160,1156,1460,1728]" box="[160,537,1460,1486]" pageId="22" pageNumber="553">
<emphasis id="F7FE5F68FFA6FF9E3E9FFA540C768FE9" bold="true" box="[160,537,1460,1486]" italics="true" pageId="22" pageNumber="553">Distribution and natural history</emphasis>
</paragraph>
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<location id="C055D5A1FFA6FF9E3E9FFA370ED38FD7" LSID="urn:lsid:plazi:treatment:4D23326CFFA1FF903FA2FBDD0A758B72:C055D5A1FFA6FF9E3E9FFA370ED38FD7" box="[160,188,1495,1520]" name="So" pageId="22" pageNumber="553">So</location>
far, 
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<emphasis id="F7FE5F68FFA6FF9E3ECDFA370F6C8FD7" box="[242,259,1495,1520]" italics="true" pageId="22" pageNumber="553">C</emphasis>
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<emphasis id="F7FE5F68FFA6FF9E3F3CFA370F688FD7" box="[259,263,1495,1520]" italics="true" pageId="22" pageNumber="553">.</emphasis>
<materialsCitation id="75E28927FFA6FF9E3F2FFA370FA88C34" ID-GBIF-Occurrence="3412503309" country="China" location="Yunnan Province" pageId="22" pageNumber="553" specimenCount="1" stateProvince="Yunnan">
<emphasis id="F7FE5F68FFA6FF9E3F2FFA370FF88FD7" box="[272,407,1495,1520]" italics="true" pageId="22" pageNumber="553">yunnanensis</emphasis>
has been collected from three localities in south-central 
<collectingRegion id="074E4D98FFA6FF9E3A12FA370F6C8C34" country="China" name="Yunnan" pageId="22" pageNumber="553">Yunnan Province</collectingRegion>
, 
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(
<figureCitation id="5DB19FFFFFA6FF9E3F5EFA1A0FD08C34" box="[353,447,1530,1555]" captionStart="Figure 7" captionStartId="23.[160,225,1054,1077]" captionTargetBox="[163,1153,142,1022]" captionTargetId="figure-184@23.[163,1153,142,1022]" captionTargetPageId="23" captionText="Figure 7. Map showing the distribution of Calamaria nebulosa sp. nov. (red) and Calamaria yunnanensis (blue). Stars represent type localities. Map created using simplemappr.net." figureDoi="http://doi.org/10.5281/zenodo.5476224" httpUri="https://zenodo.org/record/5476224/files/figure.png" pageId="22" pageNumber="553">Figure 7</figureCitation>
)
</materialsCitation>
. 
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<typeStatus id="1A313DD8FFA6FF9E3C38FA1A0C568C34" box="[519,569,1530,1555]" pageId="22" pageNumber="553">type</typeStatus>
locality, 
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, is situated 1100–1200 metres above sea level, while the locality of 
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41547 was at 1270 metres above sea level
</materialsCitation>
. 
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<location id="C055D5A1FFA6FF9E3F66F9DF0FED8C7F" LSID="urn:lsid:plazi:treatment:4D23326CFFA1FF903FA2FBDD0A758B72:C055D5A1FFA6FF9E3F66F9DF0FED8C7F" box="[345,386,1599,1624]" country="China" name="The" pageId="22" pageNumber="553" stateProvince="Yunnan">The</location>
<typeStatus id="1A313DD8FFA6FF9E3FB5F9DF0FD38C7F" box="[394,444,1599,1624]" pageId="22" pageNumber="553">type</typeStatus>
specimen was found underneath a rock in a forested montane valley
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. 
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photographed individual in 
<bibRefCitation id="A11BFE8BFFA6FF9E3C6EF9820D3B8C5C" author="Yang JH &amp; Zheng X" box="[593,852,1634,1659]" pageId="22" pageNumber="553" pagination="485 - 491" refId="ref17025" refString="Yang JH, Zheng X. 2018. A new species of the genus Calamaria (Squamata: Colubridae) from Yunnan Province, China. Copeia. 106 (3): 485 - 491. doi: 10.1643 / CH- 17 - 663" type="journal article" year="2018">Yang and Zheng (2018)</bibRefCitation>
was located approximately 
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north-west of the 
<typeStatus id="1A313DD8FFA6FF9E3FFAF9650F988CB9" box="[453,503,1669,1694]" pageId="22" pageNumber="553">type</typeStatus>
locality in 
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, 
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, 
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, 
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(
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24°48 
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09.3” 
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</geoCoordinate>
, 
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100°32 
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58.5” 
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; ~1700 metres above sea level)
</materialsCitation>
. 
<materialsCitation id="75E28927FFA6FF9F3DC6F9470D5D8EEB" ID-GBIF-Occurrence="3412503302" collectingDate="2011-07-27" collectionCode="UTC" collectorName="Chung-Wei You" country="China" lastPageId="23" lastPageNumber="554" location="It" pageId="22" pageNumber="553" specimenCount="1" stateProvince="Yunnan">
<location id="C055D5A1FFA6FF9E3DC6F9470A678CE7" LSID="urn:lsid:plazi:treatment:4D23326CFFA1FF903FA2FBDD0A758B72:C055D5A1FFA6FF9E3DC6F9470A678CE7" box="[1017,1032,1703,1728]" country="China" name="It" pageId="22" pageNumber="553" stateProvince="Yunnan">It</location>
was found on 
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<collectingDate id="A1705C52FFA7FF9F3EFBFB700F208E8E" box="[196,335,1168,1193]" pageId="23" pageNumber="554" value="2011-07-27">27 July 2011</collectingDate>
</date>
at night between 17:00 and 18:00 
<collectionCode id="A39B1BBFFFA7FF9F3CECFB700D6B8E8E" box="[723,772,1168,1193]" country="USA" httpUri="http://grbio.org/cool/tba0-w5ye" name="University of Tennessee at Chattanooga" pageId="23" pageNumber="554" type="University or college">UTC</collectionCode>
, attempting to climb across a rock along the road’s shoulder (
<collectorName id="687FE6ACFFA7FF9F3FE4FB530CFC8EEB" box="[475,659,1203,1228]" pageId="23" pageNumber="554">Chung-Wei You</collectorName>
, pers. comm)
</materialsCitation>
. 
<materialsCitation id="75E28927FFA7FF9F3D7FFB530F6A8F13" ID-GBIF-Occurrence="3412503321" collectorName="Nanjing Yi &amp; Jingdong Autonomous Counties" country="China" location="The" pageId="23" pageNumber="554" specimenCount="1" stateProvince="Yunnan">
<location id="C055D5A1FFA7FF9F3D7FFB530D068EEB" LSID="urn:lsid:plazi:treatment:4D23326CFFA1FF903FA2FBDD0A758B72:C055D5A1FFA7FF9F3D7FFB530D068EEB" box="[832,873,1203,1228]" country="China" name="The" pageId="23" pageNumber="554" stateProvince="Yunnan">The</location>
surrounding habitats in these localities include a mix of tropical montane deciduous forests near Simao, and montane evergreen broadleaf forests near 
<collectorName id="687FE6ACFFA7FF9F3C8FFB180D438F36" box="[688,812,1272,1297]" pageId="23" pageNumber="554">Nanjing Yi</collectorName>
and 
<collectorName id="687FE6ACFFA7FF9F3D47FB180F6A8F13" pageId="23" pageNumber="554">Jingdong Autonomous Counties</collectorName>
</materialsCitation>
. 
<materialsCitation id="75E28927FFA7FF9F3F2CFAFB0F0D8F5E" ID-GBIF-Occurrence="3412503315" collectorName="Cwb &amp; Koppen Classification" country="China" location="The" municipality="Cwa" pageId="23" pageNumber="554" specimenCount="1" stateProvince="Yunnan">
<location id="C055D5A1FFA7FF9F3F2CFAFB0F538F13" LSID="urn:lsid:plazi:treatment:4D23326CFFA1FF903FA2FBDD0A758B72:C055D5A1FFA7FF9F3F2CFAFB0F538F13" box="[275,316,1307,1332]" country="China" municipality="Cwa" name="The" pageId="23" pageNumber="554" stateProvince="Yunnan">The</location>
climate in these regions is monsoon influenced and considered humid subtropical (
<collectingMunicipality id="25511900FFA7FF9F3F13FADD0F308F71" box="[300,351,1341,1366]" pageId="23" pageNumber="554">Cwa</collectingMunicipality>
) or subtropical highland (
<collectorName id="687FE6ACFFA7FF9F3CB8FADD0CD38F71" box="[647,700,1341,1366]" pageId="23" pageNumber="554">Cwb</collectorName>
) according to the 
<collectorName id="687FE6ACFFA7FF9F3DAEFADD0AEC8F71" box="[913,1155,1341,1366]" pageId="23" pageNumber="554">Köppen Classification</collectorName>
(
<bibRefCitation id="A11BFE8BFFA7FF9F3E99FA800F368F5E" author="Peel MC &amp; Finlayson BL &amp; McMahon TA" box="[166,345,1376,1401]" pageId="23" pageNumber="554" pagination="1633 - 1644" refId="ref15927" refString="Peel MC, Finlayson BL, McMahon TA. 2007. Updated world map of the Koppen-Geiger climate classification. Hydrol Earth Syst Sci. 11 (5): 1633 - 1644. doi: 10.5194 / hess- 11 - 1633 - 2007" type="journal article" year="2007">Peel et al. 2007</bibRefCitation>
)
</materialsCitation>
. 
<materialsCitation id="75E28927FFA7FF9F3F4FFA800DD08FBB" ID-GBIF-Occurrence="3412503311" country="China" location="Besides" pageId="23" pageNumber="554" specimenCount="1" stateProvince="Yunnan">
<location id="C055D5A1FFA7FF9F3F4FFA800FAB8F5E" LSID="urn:lsid:plazi:treatment:4D23326CFFA1FF903FA2FBDD0A758B72:C055D5A1FFA7FF9F3F4FFA800FAB8F5E" box="[368,452,1376,1401]" country="China" name="Besides" pageId="23" pageNumber="554" stateProvince="Yunnan">Besides</location>
this information, nothing else is known regarding its natural history, but given these observations, it is likely a nocturnal species
</materialsCitation>
. Similar to other species of 
<taxonomicName id="028AF8F9FFA7FF9F3F2BFA450FEA8F99" baseAuthorityName="Inger and Marx" baseAuthorityYear="1965" box="[276,389,1445,1470]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="554" phylum="Chordata" rank="genus">
<emphasis id="F7FE5F68FFA7FF9F3F2BFA450FEA8F99" box="[276,389,1445,1470]" italics="true" pageId="23" pageNumber="554">Calamaria</emphasis>
</taxonomicName>
, it is presumed to be a fossorial species of snake that spends most of its life underground or below the forest floor, feeding on small soft-bodied invertebrates such as earthworms (
<bibRefCitation id="A11BFE8BFFA7FF9F3FB1FA0B0C168C23" author="Inger RF &amp; Marx H" box="[398,633,1515,1540]" pageId="23" pageNumber="554" pagination="1 - 304" refId="ref15062" refString="Inger RF, Marx H. 1965. The systematics and evolution of the oriental colubrid snakes of the genus Calamaria. Fieldiana Zool. 49: 1 - 304." type="journal article" year="1965">Inger and Marx 1965</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="91F5D3F2FFA7FF9F3E9FFBFE0DAC8E75" ID-DOI="http://doi.org/10.5281/zenodo.5476224" ID-Zenodo-Dep="5476224" httpUri="https://zenodo.org/record/5476224/files/figure.png" pageId="23" pageNumber="554" startId="23.[160,225,1054,1077]" targetBox="[163,1153,142,1022]" targetPageId="23">
<paragraph id="C535837AFFA7FF9F3E9FFBFE0DAC8E75" blockId="23.[160,1155,1054,1106]" pageId="23" pageNumber="554">
<emphasis id="F7FE5F68FFA7FF9F3E9FFBFE0E938E12" bold="true" box="[160,252,1054,1077]" pageId="23" pageNumber="554">Figure 7.</emphasis>
Map showing the distribution of 
<taxonomicName id="028AF8F9FFA7FF9F3C6DFBFE0D7C8E12" authority="Lee, 2021" authorityName="Lee" authorityYear="2021" box="[594,787,1054,1077]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="554" phylum="Chordata" rank="species" species="nebulosa" status="sp. nov.">
<emphasis id="F7FE5F68FFA7FF9F3C6DFBFE0D7C8E12" box="[594,787,1054,1077]" italics="true" pageId="23" pageNumber="554">Calamaria nebulosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ECCDE213FFA7FF9F3D24FBFE0D058E12" box="[795,874,1054,1077]" pageId="23" pageNumber="554" rank="species">sp. nov.</taxonomicNameLabel>
(red) and 
<taxonomicName id="028AF8F9FFA7FF9F3DEAFBFE0EB58E75" authorityName=": Chernov" authorityYear="1962" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="554" phylum="Chordata" rank="species" species="yunnanensis">
<emphasis id="F7FE5F68FFA7FF9F3DEAFBFE0EB58E75" italics="true" pageId="23" pageNumber="554">Calamaria yunnanensis</emphasis>
</taxonomicName>
(blue). Stars represent type localities. Map created using simplemappr.net.
</paragraph>
</caption>
<subSubSection id="8D90D0F1FFA7FF903E9FF9DF0A668AED" lastPageId="24" lastPageNumber="555" pageId="23" pageNumber="554" type="conservation">
<paragraph id="C535837AFFA7FF9F3E9FF9DF0FE48C7E" blockId="23.[160,1155,1599,1694]" box="[160,395,1599,1625]" pageId="23" pageNumber="554">
<emphasis id="F7FE5F68FFA7FF9F3E9FF9DF0FE48C7E" bold="true" box="[160,395,1599,1625]" italics="true" pageId="23" pageNumber="554">Conservation status</emphasis>
</paragraph>
<paragraph id="C535837AFFA7FF903E9FF9830A668AED" blockId="23.[160,1155,1599,1694]" lastBlockId="24.[160,1156,142,202]" lastPageId="24" lastPageNumber="555" pageId="23" pageNumber="554">
<taxonomicName id="028AF8F9FFA7FF9F3E9FF9830FC88C5B" authorityName=": Chernov" authorityYear="1962" box="[160,423,1635,1660]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="554" phylum="Chordata" rank="species" species="yunnanensis">
<emphasis id="F7FE5F68FFA7FF9F3E9FF9830FC88C5B" box="[160,423,1635,1660]" italics="true" pageId="23" pageNumber="554">Calamaria yunnanensis</emphasis>
</taxonomicName>
<materialsCitation id="75E28927FFA7FF9F3F89F9830DE88CB9" ID-GBIF-Occurrence="3412503304" country="China" location="Yunnan Province" pageId="23" pageNumber="554" specimenCount="1" stateProvince="Yunnan">
has been recorded from two localities in southern 
<collectingRegion id="074E4D98FFA7FF9F3A12F9830F6E8CB9" country="China" name="Yunnan" pageId="23" pageNumber="554">Yunnan Province</collectingRegion>
<collectingCountry id="BD9DC3EAFFA7FF9F3F37F9650F278CB9" box="[264,328,1669,1694]" name="China" pageId="23" pageNumber="554">China</collectingCountry>
based on a total of six to seven known specimens
</materialsCitation>
. 
<materialsCitation id="75E28927FFA7FF903DADF9650C8B8A80" ID-GBIF-Occurrence="3412503312" collectorName="The" country="China" lastPageId="24" lastPageNumber="555" location="Yunnan Province" pageId="23" pageNumber="554" specimenCount="1" stateProvince="Yunnan">
<collectorName id="687FE6ACFFA7FF9F3DADF9650DD48CB9" box="[914,955,1669,1694]" pageId="23" pageNumber="554">The</collectorName>
full extent of this species distribution in 
<collectingRegion id="074E4D98FFA8FF903FA3FF6E0C358A80" box="[412,602,142,167]" country="China" name="Yunnan" pageId="24" pageNumber="555">Yunnan Province</collectingRegion>
is unknown
</materialsCitation>
. I recommend that 
<taxonomicName id="028AF8F9FFA8FF903D81FF6E0A0F8A80" authorityName=": Chernov" authorityYear="1962" box="[958,1120,142,167]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="555" phylum="Chordata" rank="species" species="yunnanensis">
<emphasis id="F7FE5F68FFA8FF903D81FF6E0A0F8A80" box="[958,1120,142,167]" italics="true" pageId="24" pageNumber="555">
C. 
<materialsCitation id="75E28927FFA8FF903DE6FF6E0A0F8A80" ID-GBIF-Occurrence="3412503317" box="[985,1120,142,167]" collectorName="Data Deficient'" country="China" location="Red List" pageId="24" pageNumber="555" specimenCount="1" stateProvince="Yunnan">yunnanensis</materialsCitation>
</emphasis>
</taxonomicName>
be listed as ‘ 
<collectorName id="687FE6ACFFA8FF903F3AFF510FC18AED" box="[261,430,177,202]" pageId="24" pageNumber="555">Data Deficient’</collectorName>
following the IUCN’s 
<location id="C055D5A1FFA8FF903C9CFF510C948AED" LSID="urn:lsid:plazi:treatment:4D23326CFFA1FF903FA2FBDD0A758B72:C055D5A1FFA8FF903C9CFF510C948AED" box="[675,763,177,202]" country="China" name="Red List" pageId="24" pageNumber="555" stateProvince="Yunnan">Red List</location>
categories (
<bibRefCitation id="A11BFE8BFFA8FF903DBCFF510D938AED" author="IUCN" box="[899,1020,177,202]" pageId="24" pageNumber="555" refId="ref15093" refString="IUCN. 2019. IUCN International Union for Conservation of Nature. [accessed 2020 Apr 3]. https: // www. iucnredlist. org /." type="journal volume" year="2019">IUCN 2019</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8D90D0F1FFA8FF903E9FFF160A758B72" pageId="24" pageNumber="555" type="etymology">
<paragraph id="C535837AFFA8FF903E9FFF160F728B37" blockId="24.[160,1155,246,341]" box="[160,285,246,272]" pageId="24" pageNumber="555">
<emphasis id="F7FE5F68FFA8FF903E9FFF160F728B37" bold="true" box="[160,285,246,272]" italics="true" pageId="24" pageNumber="555">Etymology</emphasis>
</paragraph>
<paragraph id="C535837AFFA8FF903E9FFEF90A758B72" blockId="24.[160,1155,246,341]" pageId="24" pageNumber="555">
<materialsCitation id="75E28927FFA8FF903E9FFEF90F3E8B72" ID-GBIF-Occurrence="3412503306" country="China" location="Yunnan Province" pageId="24" pageNumber="555" specimenCount="1" stateProvince="Yunnan">
The species epithet ‘ 
<taxonomicName id="028AF8F9FFA8FF903FB2FEF90C7B8B15" authorityName=": Chernov" authorityYear="1962" box="[397,532,281,306]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="555" phylum="Chordata" rank="species" species="yunnanensis">
<emphasis id="F7FE5F68FFA8FF903FB2FEF90C7B8B15" box="[397,532,281,306]" italics="true" pageId="24" pageNumber="555">yunnanensis</emphasis>
</taxonomicName>
’ is a reference to the 
<typeStatus id="1A313DD8FFA8FF903D29FEF90D278B15" box="[790,840,281,306]" pageId="24" pageNumber="555">type</typeStatus>
locality situated in 
<collectingRegion id="074E4D98FFA8FF903A12FEF90F6C8B72" country="China" name="Yunnan" pageId="24" pageNumber="555">Yunnan Province</collectingRegion>
, 
<collectingCountry id="BD9DC3EAFFA8FF903F31FEDC0F3E8B72" box="[270,337,316,341]" name="China" pageId="24" pageNumber="555">China</collectingCountry>
</materialsCitation>
. I recommend the English common name ‘ 
<collectingRegion id="074E4D98FFA8FF903D0DFEDC0DE48B72" box="[818,907,316,341]" country="China" name="Yunnan" pageId="24" pageNumber="555">Yunnan</collectingRegion>
reed snake’.
</paragraph>
</subSubSection>
</treatment>
</document>