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<document id="42644EE9139142EFF6EA8CE325376B92" ID-DOI="10.11646/zootaxa.4020.1.7" ID-GBIF-Dataset="82b2f539-5d25-42d6-95c5-e8b16a353e12" ID-ISSN="1175-5326" ID-Zenodo-Dep="237970" ID-ZooBank="BD8D1880-79A9-4672-99AF-5B3D1BEFAFEE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461269068152" checkinUser="plazi" docAuthor="Bickel, Daniel J." docDate="2015" docId="172D87FDFFB4FF87FF3DA69301D8F9B5" docLanguage="en" docName="zt04020p182.pdf" docOrigin="Zootaxa 4020 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Systenus zurqui Bickel, sp. nov." docType="treatment" docVersion="8" lastPageNumber="178" masterDocId="EB14FF85FFBDFF8EFFAAA70A0459FFCC" masterDocTitle="The Costa Rican Systenus Loew (Diptera: Dolichopodidae): rich local sympatry in an otherwise rare genus" masterLastPageNumber="182" masterPageNumber="169" pageNumber="178" updateTime="1698611248703" updateUser="plazi">
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<mods:title id="32F3833B1C391D154D6B5FF9572E7AE5">The Costa Rican Systenus Loew (Diptera: Dolichopodidae): rich local sympatry in an otherwise rare genus</mods:title>
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<mods:namePart id="9019B3099B202EB29EC23882E37FE798">Bickel, Daniel J.</mods:namePart>
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<mods:date id="F6466E0D02AEC6B06BD5993D963C9C17">2015</mods:date>
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<mods:number id="66B1B198EED75492434A37DD2158549C">4020</mods:number>
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<treatment id="172D87FDFFB4FF87FF3DA69301D8F9B5" ID-DOI="http://doi.org/10.5281/zenodo.6121813" ID-GBIF-Taxon="119638523" ID-Zenodo-Dep="6121813" LSID="urn:lsid:plazi:treatment:172D87FDFFB4FF87FF3DA69301D8F9B5" httpUri="http://treatment.plazi.org/id/172D87FDFFB4FF87FF3DA69301D8F9B5" lastPageNumber="178" pageId="9" pageNumber="178">
<subSubSection id="D79E6560FFB4FF87FF3DA693054EFE19" pageId="9" pageNumber="178" type="nomenclature">
<paragraph id="9F3B36EBFFB4FF87FF3DA6930648FE7F" blockId="9.[151,529,409,469]" box="[151,529,409,435]" pageId="9" pageNumber="178">
<heading id="C4738187FFB4FF87FF3DA6930648FE7F" bold="true" box="[151,529,409,435]" fontSize="11" level="1" pageId="9" pageNumber="178" reason="1">
<emphasis id="ADF0EAF9FFB4FF87FF3DA6930648FE7F" bold="true" box="[151,529,409,435]" pageId="9" pageNumber="178">
<taxonomicName id="58844D68FFB4FF87FF3DA69305F2FE7F" ID-CoL="9HVW9" authority="Bickel" authorityName="Bickel" box="[151,427,409,435]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="178" phylum="Arthropoda" rank="species" species="zurqui" status="sp. nov.">
<emphasis id="ADF0EAF9FFB4FF87FF3DA693050FFE7F" bold="true" box="[151,342,409,435]" italics="true" pageId="9" pageNumber="178">Systenus zurqui</emphasis>
Bickel
</taxonomicName>
<taxonomicNameLabel id="B6C35782FFB4FF87FE19A6930648FE7F" box="[435,529,409,435]" pageId="9" pageNumber="178" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="9F3B36EBFFB4FF87FF3DA6B6054EFE19" blockId="9.[151,529,409,469]" box="[151,279,444,469]" pageId="9" pageNumber="178">
(
<figureCitation id="07BF2A6EFFB4FF87FF35A6B604BCFE19" box="[159,229,444,469]" captionStart="FIGURE 3" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[166,1436,813,1836]" captionTargetId="figure@8.[151,1436,813,1836]" captionTargetPageId="8" captionText="FIGURE 3. Systenus divericatus sp. nov. (holotype): a. male habitus, left lateral; b. hypopygium, left lateral. S. zurqui sp. nov.: c. male habitus, left lateral; d. hypopygium, left lateral." httpUri="https://zenodo.org/record/237973/files/figure.png" pageId="9" pageNumber="178">Figs 3</figureCitation>
c, d)
</paragraph>
</subSubSection>
<subSubSection id="D79E6560FFB4FF87FF3DA50E0675FD45" pageId="9" pageNumber="178" type="materials_examined">
<paragraph id="9F3B36EBFFB4FF87FF3DA50E0675FD45" blockId="9.[151,1437,516,1657]" pageId="9" pageNumber="178">
<emphasis id="ADF0EAF9FFB4FF87FF3DA50E051DFDD1" bold="true" box="[151,324,516,541]" pageId="9" pageNumber="178">
<typeStatus id="403F8849FFB4FF87FF3DA50E048BFDD1" box="[151,210,516,541]" pageId="9" pageNumber="178">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="403F8849FFB4FF87FEF2A50E05AEFDD1" box="[344,503,516,541]" pageId="9" pageNumber="178" type="holotype">
<emphasis id="ADF0EAF9FFB4FF87FEF2A50E05AEFDD1" bold="true" box="[344,503,516,541]" pageId="9" pageNumber="178">HOLOTYPE</emphasis>
</typeStatus>
♂,
<collectingCountry id="E793767BFFB4FF87FD84A50F0685FDD1" box="[558,732,517,541]" name="Costa Rica" pageId="9" pageNumber="178">COSTA RICA</collectingCountry>
:
<emphasis id="ADF0EAF9FFB4FF87FD47A50E0702FDD1" bold="true" box="[749,859,516,541]" pageId="9" pageNumber="178">San José</emphasis>
:
<taxonomicName id="58844D68FFB4FF87FCDAA50F07E5FDD1" box="[880,956,517,541]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="181" phylum="Arthropoda" rank="species" species="zurqui">Zurquí</taxonomicName>
de Moravia.
<geoCoordinate id="FAB0502CFFB4FF87FBF3A50E00EFFDD1" box="[1113,1206,516,541]" direction="north" orientation="latitude" pageId="9" pageNumber="178" precision="555" value="10.05">10.05°N</geoCoordinate>
<geoCoordinate id="FAB0502CFFB4FF87FB69A50F0171FDD1" box="[1219,1320,517,541]" direction="west" orientation="longitude" pageId="9" pageNumber="178" precision="555" value="-84.02">84.02°W</geoCoordinate>
,
<quantity id="587C9B0EFFB4FF87FA90A50E01CEFDD0" box="[1338,1431,516,541]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="9" pageNumber="178" unit="m" value="1600.0">1600 m</quantity>
, Malaise trap,
<date id="EB3A102BFFB4FF87FE9DA5220595FD8C" box="[311,460,552,576]" pageId="9" pageNumber="178" value="2014-08-10">10.VIII.2014</date>
, ZADBI-909 (
<collectionCode id="F995AE2EFFB4FF87FDDCA5220695FD8C" LSID="urn:lsid:biocol.org:col:34802" box="[630,716,552,576]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="9" pageNumber="178">LACM</collectionCode>
;
<collectionCode id="F995AE2EFFB4FF87FD71A5220775FD8C" LSID="urn:lsid:biocol.org:col:34802" box="[731,812,552,576]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="9" pageNumber="178">LACM</collectionCode>
Ent 320001). (
<collectionCode id="F995AE2EFFB4FF87FC74A5220068FD8C" LSID="urn:lsid:biocol.org:col:34802" box="[990,1073,552,576]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="9" pageNumber="178">LACM</collectionCode>
Ent 300001);
<typeStatus id="403F8849FFB4FF87FB77A522012CFD8D" box="[1245,1397,552,577]" pageId="9" pageNumber="178" type="paratype">
<emphasis id="ADF0EAF9FFB4FF87FB77A522012CFD8D" bold="true" box="[1245,1397,552,577]" pageId="9" pageNumber="178">PARATYPE</emphasis>
</typeStatus>
♂,
<emphasis id="ADF0EAF9FFB4FF87FF3DA5460546FDA9" bold="true" box="[151,287,588,613]" pageId="9" pageNumber="178">Puntarenas</emphasis>
: Estación La Casona, R.B. Monteverde, A.C. Arenal,
<quantity id="587C9B0EFFB4FF87FC2BA5460781FDA8" box="[897,984,588,613]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="9" pageNumber="178" unit="m" value="1520.0">1520 m</quantity>
,
<date id="EB3A102BFFB4FF87FC4EA5470015FDA9" box="[996,1100,589,613]" pageId="9" pageNumber="178" value="1993-12">XII.1993</date>
, 253250 449700, #2605, N.G. Obando, (
<collectionCode id="F995AE2EFFB4FF87FEAFA57A050EFD44" box="[261,343,624,648]" pageId="9" pageNumber="178">INBIO</collectionCode>
, CR1001 865191).
</paragraph>
</subSubSection>
<subSubSection id="D79E6560FFB4FF87FF6DA59E06A6FAB1" pageId="9" pageNumber="178" type="description">
<paragraph id="9F3B36EBFFB4FF87FF6DA59E05B8FB90" blockId="9.[151,1437,516,1657]" pageId="9" pageNumber="178">
<emphasis id="ADF0EAF9FFB4FF87FF6DA59E0508FD61" bold="true" box="[199,337,660,685]" pageId="9" pageNumber="178">Description</emphasis>
.
<emphasis id="ADF0EAF9FFB4FF87FECAA59E05C5FD61" bold="true" box="[352,412,660,685]" pageId="9" pageNumber="178">Male</emphasis>
: body length:
<quantity id="587C9B0EFFB4FF87FDE9A59E06C4FD60" box="[579,669,660,685]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="9" pageNumber="178" unit="mm" value="2.6">2.6 mm</quantity>
; wing 2.5 x
<quantity id="587C9B0EFFB4FF87FC81A59F07DBFD60" box="[811,898,661,685]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="9" pageNumber="178" unit="mm" value="0.7">0.7 mm</quantity>
(
<figureCitation id="07BF2A6EFFB4FF87FC38A59E078FFD61" box="[914,982,660,685]" captionStart="FIGURE 3" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[166,1436,813,1836]" captionTargetId="figure@8.[151,1436,813,1836]" captionTargetPageId="8" captionText="FIGURE 3. Systenus divericatus sp. nov. (holotype): a. male habitus, left lateral; b. hypopygium, left lateral. S. zurqui sp. nov.: c. male habitus, left lateral; d. hypopygium, left lateral." httpUri="https://zenodo.org/record/237973/files/figure.png" pageId="9" pageNumber="178">Fig. 3</figureCitation>
c); similar to
<taxonomicName id="58844D68FFB4FF87FBC5A59F0084FD60" box="[1135,1245,661,684]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="178" phylum="Arthropoda" rank="species" species="tenorio">
<emphasis id="ADF0EAF9FFB4FF87FBC5A59F0084FD60" box="[1135,1245,661,684]" italics="true" pageId="9" pageNumber="178">S. tenorio</emphasis>
</taxonomicName>
except as noted:
<emphasis id="ADF0EAF9FFB4FF87FF3DA5B2048DFD1D" box="[151,212,696,721]" italics="true" pageId="9" pageNumber="178">Head</emphasis>
: proboscis yellowish, projecting anteriorly, keel-like; antenna dark brown with yellowish near base of postpedicel on median side; scape and pedicel short; postpedicel elongate, triangular and tapering gradually, covered with short pubescence, about 3 times as long as basal width, and with short apical arista.
<emphasis id="ADF0EAF9FFB4FF87FB4DA40A0161FCD5" box="[1255,1336,768,793]" italics="true" pageId="9" pageNumber="178">Thorax</emphasis>
: dorsum dark metallic green but covered with thick grey pruinosity; pleura green with dense grey pruinosity; some 12 pairs of short ac.
<emphasis id="ADF0EAF9FFB4FF87FEB1A4430509FCAC" box="[283,336,841,864]" italics="true" pageId="9" pageNumber="178">Legs</emphasis>
: CI yellow; CII and CIII mostly dark brown, but distal eighth pale yellow; all trochanters, and the remainder of legs I and II yellow; FIII yellow basally but distal quarter mostly dark brown; TIII yellow but infuscated distally; tarsus III mostly yellow but with brown infuscation; CI and CII with white anterior setae and shorter vestiture, CIII with strong white lateral seta at ½; major leg setae black; I: 3.7; 3.6; 2.0/ 0.9/ 0.7/ 0.4/ 0.3; TI with normal short black vestiture; II: 4.3; 4.8; 2.4/ 1.5/ 1.0/ 0.8/ 0.5; TII with strong ad seta at 1/5 and ½, and weaker pd seta at 1/6 and 2/3, and with apical ring of ad, pd, av and ventral seta; III: 4.7; 6.0; 1.1/ 2.1/ 1.2/ 0.8/ 0.4; TIII with normal black vestiture, with ad-pd setal pair at 1/5 and with 4 spaced black dorsal setae, and with apical ring of dorsal, ad and av seta.
</paragraph>
<paragraph id="9F3B36EBFFB4FF87FF6DA36306A6FAB1" blockId="9.[151,1437,516,1657]" pageId="9" pageNumber="178">
<emphasis id="ADF0EAF9FFB4FF87FF6DA36304A6FB4C" box="[199,255,1129,1152]" italics="true" pageId="9" pageNumber="178">Wing</emphasis>
: CuAx ratio: 0.7.
<emphasis id="ADF0EAF9FFB4FF87FE7FA3620666FB4D" box="[469,575,1128,1153]" italics="true" pageId="9" pageNumber="178">Abdomen</emphasis>
: tergites 16 metallic blue- green with only dusting of pruinosity posterior margin of tergite 1 with posterior row of long black setae, tergites otherwise covered with short black vestiture; segment 7 dark brown and forming unusually long peduncle, distinctly longer than length of epandrium, with sternite 7 glabrous and tergite 7 covered with black vestiture; sternite 8 forming cap-like cover over hypopygial foramen on left side of epandrium; hypopygium (
<figureCitation id="07BF2A6EFFB4FF87FD79A3F2074FFADD" box="[723,790,1272,1297]" captionStart="FIGURE 3" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[166,1436,813,1836]" captionTargetId="figure@8.[151,1436,813,1836]" captionTargetPageId="8" captionText="FIGURE 3. Systenus divericatus sp. nov. (holotype): a. male habitus, left lateral; b. hypopygium, left lateral. S. zurqui sp. nov.: c. male habitus, left lateral; d. hypopygium, left lateral." httpUri="https://zenodo.org/record/237973/files/figure.png" pageId="9" pageNumber="178">Fig. 3</figureCitation>
d) dark brown with yellow surstylus and cercus; ventral epandrium extended ventrally with 2 subapical setae; surstylus forming long curved and tapering arm, bearing curved ventral lobe with apical seta, and with curved dorsal digitiform projection; cercus elongate and clavate, with subapical field of strong pale setae.
<emphasis id="ADF0EAF9FFB4FF87FD83A26E06D9FAB1" bold="true" box="[553,640,1380,1405]" pageId="9" pageNumber="178">Female</emphasis>
: unknown.
</paragraph>
</subSubSection>
<subSubSection id="D79E6560FFB4FF87FF6DA2820661F998" pageId="9" pageNumber="178" type="discussion">
<paragraph id="9F3B36EBFFB4FF87FF6DA2820661F998" blockId="9.[151,1437,516,1657]" pageId="9" pageNumber="178">
<emphasis id="ADF0EAF9FFB4FF87FF6DA282056DFA6D" bold="true" box="[199,308,1416,1441]" pageId="9" pageNumber="178">Remarks</emphasis>
.
<taxonomicName id="58844D68FFB4FF87FEE9A28305ABFA6C" box="[323,498,1417,1440]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="178" phylum="Arthropoda" rank="species" species="zurqui">
<emphasis id="ADF0EAF9FFB4FF87FEE9A28305ABFA6C" box="[323,498,1417,1440]" italics="true" pageId="9" pageNumber="178">Systenus zurqui</emphasis>
</taxonomicName>
is known from the
<taxonomicName id="58844D68FFB4FF87FD7EA2820779FA6D" box="[724,800,1416,1441]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="181" phylum="Arthropoda" rank="species" species="zurqui">Zurqui</taxonomicName>
site (collected in August) and Monteverde (collected in December),
<collectingCountry id="E793767BFFB4FF87FE89A2A705FAFA09" box="[291,419,1452,1477]" name="Costa Rica" pageId="9" pageNumber="178">Costa Rica</collectingCountry>
. Both sites are above
<quantity id="587C9B0EFFB4FF87FD0BA2A706A1FA08" box="[673,760,1453,1477]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="178" unit="m" value="1200.0">1200 m</quantity>
in elevation. The Monteverde specimen is distinctly larger (wing length 3.0) than the
<taxonomicName id="58844D68FFB4FF87FE63A2DA064CFA25" box="[457,533,1488,1513]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="181" phylum="Arthropoda" rank="species" species="zurqui">Zurqui</taxonomicName>
specimen (
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), but they are otherwise similar. It has the diagnostic set of male characters of black tapering postpedicel, femora yellow except femur III dark brown in the distal quarter, tibia I without ivory colored vestiture, an unusually long hypopygial peduncle, longer than epandrium, and a subapical field of long pale setae on the cercus.
</paragraph>
</subSubSection>
<subSubSection id="D79E6560FFB4FF87FF6DA16A01D8F9B5" box="[199,1409,1632,1657]" pageId="9" pageNumber="178" type="etymology">
<paragraph id="9F3B36EBFFB4FF87FF6DA16A01D8F9B5" blockId="9.[151,1437,516,1657]" box="[199,1409,1632,1657]" pageId="9" pageNumber="178">
<emphasis id="ADF0EAF9FFB4FF87FF6DA16A051FF9B5" bold="true" box="[199,326,1632,1657]" pageId="9" pageNumber="178">Etymology</emphasis>
. The specific epithet “
<emphasis id="ADF0EAF9FFB4FF87FDEEA16B06C2F9B4" box="[580,667,1633,1656]" italics="true" pageId="9" pageNumber="178">
<taxonomicName id="58844D68FFB4FF87FDEEA16B06CBF9B4" box="[580,658,1633,1656]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="178" phylum="Arthropoda" rank="species" species="zurqui">zurqui</taxonomicName>
</emphasis>
is a place name and should be considered as a noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>