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<document id="13FDB7778F6D268B93B83DBC67115BA5" ID-CLB-Dataset="117405" ID-DOI="10.35929/RSZ.0014" ID-GBIF-Dataset="0d204a58-f06b-423f-b85c-06f6e4ad2b71" ID-ISSN="0035-418" ID-Zenodo-Dep="5743429" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638275271213" checkinUser="carolina" docAuthor="Sabatinelli, Guido" docDate="2020" docId="AA41AD4E7156FFEF7E1BFEF7FA85FED4" docLanguage="en" docName="RevueSuisseZool.127.1.157-181.pdf" docOrigin="Revue suisse de Zoologie 127 (1)" docSource="http://dx.doi.org/10.35929/rsz.0014" docStyle="DocumentStyle:EA5207814960044D272DF5BE47DFF256.3:RevSuisseZool.2020-.journal_article.1cover" docStyleId="EA5207814960044D272DF5BE47DFF256" docStyleName="RevSuisseZool.2020-.journal_article.1cover" docStyleVersion="3" docTitle="Cyphochilus obscurus Sharp 1876" docType="treatment" docVersion="10" lastPageNumber="172" masterDocId="5678D5367146FFFF7F1FFFF7FFF3FF8D" masterDocTitle="Taxonomic notes on the genus Cyphochilus Waterhouse, 1867 (Coleoptera, Scarabaeoidea, Melolonthinae) with description of 10 new species" masterLastPageNumber="181" masterPageNumber="157" pageNumber="172" updateTime="1699471448908" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="FFAAED12FC522D594AEB3ADE7D9B8ADB">Taxonomic notes on the genus Cyphochilus Waterhouse, 1867 (Coleoptera, Scarabaeoidea, Melolonthinae) with description of 10 new species</mods:title>
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<mods:namePart id="D9D4DD6B28C3314B598F313148343531">Sabatinelli, Guido</mods:namePart>
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<mods:identifier id="D1BB4F77C10D5BF7AC770CBC74B66AD5" type="DOI">10.35929/RSZ.0014</mods:identifier>
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<treatment id="AA41AD4E7156FFEF7E1BFEF7FA85FED4" ID-DOI="http://doi.org/10.5281/zenodo.5743496" ID-GBIF-Taxon="198967501" ID-Zenodo-Dep="5743496" LSID="urn:lsid:plazi:treatment:AA41AD4E7156FFEF7E1BFEF7FA85FED4" httpUri="http://treatment.plazi.org/id/AA41AD4E7156FFEF7E1BFEF7FA85FED4" lastPageNumber="172" pageId="16" pageNumber="172">
<subSubSection id="6AF24FD37156FFEF7E1BFEF7FD74FE95" box="[260,647,256,280]" pageId="16" pageNumber="172" type="nomenclature">
<paragraph id="22571C587156FFEF7E1BFEF7FD74FE95" blockId="16.[260,647,256,313]" box="[260,647,256,280]" pageId="16" pageNumber="172">
<heading id="791FAB347156FFEF7E1BFEF7FD74FE95" bold="true" box="[260,647,256,280]" centered="true" fontSize="10" level="2" pageId="16" pageNumber="172" reason="2">
<taxonomicName id="E5E867DB7156FFEF7E1BFEF7FD74FE95" authority="Sharp, 1876" authorityName="Sharp" authorityYear="1876" box="[260,647,256,280]" class="Insecta" family="Melolonthidae" genus="Cyphochilus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="172" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="109CC04A7156FFEF7E1BFEF7FD74FE95" bold="true" box="[260,647,256,280]" pageId="16" pageNumber="172">
<emphasis id="109CC04A7156FFEF7E1BFEF7FE0BFE95" bold="true" box="[260,504,256,280]" italics="true" pageId="16" pageNumber="172">Cyphochilus obscurus</emphasis>
<bibRefCitation id="467961A97156FFEF7EE0FEF6FD74FE95" author="Sharp D." box="[511,647,257,280]" pageId="16" pageNumber="172" pagination="65 - 90" refId="ref20538" refString="Sharp D. 1876. Description of some new species of Scarabaeidae from tropical Asia and Malaisia. Part III (Melolonthini). Coleopterologische Hefte 15: 65 - 90." type="journal article" year="1876">Sharp, 1876</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="6AF24FD37156FFEF7E74FED5FDD2FEB4" box="[363,545,290,313]" pageId="16" pageNumber="172" type="description">
<paragraph id="22571C587156FFEF7E74FED5FDD2FEB4" blockId="16.[260,647,256,313]" box="[363,545,290,313]" pageId="16" pageNumber="172">
<figureCitation id="BAD300DD7156FFEF7E74FED5FE5DFEB4" box="[363,430,290,313]" captionStart="Figs 1-14" captionStartId="7.[151,192,957,978]" captionTargetBox="[157,1435,257,896]" captionTargetId="figure-87@7.[151,1436,257,914]" captionTargetPageId="7" captionText="Figs 1-14. Original labels of type material. (1) Cyphochilus pygidialis Nonfried, 1893. (2) C. pygidialis Brenske [sic]. (3) C. septentrionalis Waterhouse, 1867. (4) C. obscurus Sharp, 1876. (5) C. proximus Sharp, 1876. (6) C. flavomarginatus Frey, 1971. (7) C. cylindricus Brenske, 1903. (8) Melolontha candida (Olivier, 1789). (9) Cyphochilus pygidialis v. angeri Nonfried, 1893. (10) C. feae Brenske, 1903. (11) C. vestitus Sharp, 1876. (12) C. tricolor Waterhouse, 1867. (13) C. testaceipes Fairmaire, 1902. (14) C. peninsularis Arrow, 1938." figureDoi="http://doi.org/10.5281/zenodo.5743431" httpUri="https://zenodo.org/record/5743431/files/figure.png" pageId="16" pageNumber="172">Figs 4</figureCitation>
,
<figureCitation id="BAD300DD7156FFEF7EA5FED5FE08FEB4" box="[442,507,290,313]" captionStart="Figs 47-58" captionStartId="13.[151,192,1959,1980]" captionTargetBox="[164,1425,260,1900]" captionTargetId="figure-7@13.[151,1436,254,1945]" captionTargetPageId="13" captionText="Figs 47-58. Parameres in right, dorsal, and left view. (47-49) Cyphochilus carinchebanus Brenske, 1903. (50-52) C. obscurus Sharp, 1876, holotype. (53-55) C. proximus Sharp, 1876, holotype. (56-58) C. tricolor Waterhouse, 1867, lectotype. – Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743441" httpUri="https://zenodo.org/record/5743441/files/figure.png" pageId="16" pageNumber="172">50-52</figureCitation>
,
<figureCitation id="BAD300DD7156FFEF7D19FED5FDD2FEB4" box="[518,545,290,313]" captionStart="Figs 80-91" captionStartId="23.[151,192,1883,1904]" captionTargetBox="[206,1382,268,1813]" captionTargetId="figure-116@23.[201,1387,257,1827]" captionTargetPageId="23" captionText="Figs 80-91. Habitus. (80) Cyphochilus candidus (Olivier, 1789), lectotype ♂ MNHN. (81) C. candidus (Olivier, 1789), Nepal: Godavari, ♂ MHNG, yellow scales aberration. (82) C. carinchebanus Brenske, 1903, Myanmar: Carin Cheba, ♂ ZMHB. (83) C. feae Brenske, 1903, holotype ZMHB. (84) C. gandhii n. sp., holotype MHNG. (85) C. leducthoi n. sp., holotype MHNG.(86) C. septentrionalis Waterhouse, 1867, lectotype BMNH. (87) C. obscurus Sharp, 1876, holotype MNHN. (88) C. sansuukyii n. sp., paratype ♀ MHNG. (89) C. peninsularis Arrow, 1938, paralectotype ♂ MHNG. (90) C. rohingyae n. sp., holotype MHNG. (91) C. rohingyae n. sp., paratype ♂ MHNG, Malaysia: Perak, bicolor aberration. – Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5743449" httpUri="https://zenodo.org/record/5743449/files/figure.png" pageId="16" pageNumber="172">87</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="6AF24FD37156FFEF7F88FEA7FDE2FEE8" box="[151,529,336,357]" pageId="16" pageNumber="172" type="reference_group">
<paragraph id="22571C587156FFEF7F88FEA7FDE2FEE8" blockId="16.[151,529,336,357]" box="[151,529,336,357]" pageId="16" pageNumber="172">
<treatmentCitationGroup id="02F83B767156FFEF7F88FEA7FDE2FEE8" box="[151,529,336,357]" pageId="16" pageNumber="172">
<treatmentCitation id="A3493A497156FFEF7F88FEA7FDFDFEE8" author="Sharp D." box="[151,526,336,357]" page="83" pageId="16" pageNumber="172" year="1876">
<taxonomicName id="E5E867DB7156FFEF7F88FEA7FDFDFEE8" authority="Sharp, 1876: 83" authorityName="Sharp" authorityPageNumber="83" authorityYear="1876" box="[151,526,336,357]" class="Insecta" family="Melolonthidae" genus="Cyphochilus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="172" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="109CC04A7156FFEF7F88FEA7FE9FFEE8" box="[151,364,336,357]" italics="true" pageId="16" pageNumber="172">Cyphochilus obscurus</emphasis>
<bibRefCitation id="467961A97156FFEF7E6DFEA7FDFDFEE8" author="Sharp D." box="[370,526,336,357]" pageId="16" pageNumber="172" pagination="65 - 90" refId="ref20538" refString="Sharp D. 1876. Description of some new species of Scarabaeidae from tropical Asia and Malaisia. Part III (Melolonthini). Coleopterologische Hefte 15: 65 - 90." type="journal article" year="1876">Sharp, 1876: 83</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="6AF24FD37156FFEF7F88FE89FDD0FCA1" pageId="16" pageNumber="172" type="materials_examined">
<paragraph id="22571C587156FFEF7F88FE89FE00FD95" blockId="16.[151,756,382,536]" pageId="16" pageNumber="172">
<emphasis id="109CC04A7156FFEF7F88FE89FE3BFE18" bold="true" box="[151,456,382,405]" pageId="16" pageNumber="172">Type material examined:</emphasis>
<materialsCitation id="928016057156FFEF7EC8FE89FE00FD95" ID-GBIF-Occurrence="3886339303" collectionCode="MNHN" collectorName="D. Sharp &amp; Coll. R. Oberthur" country="Laos" location="LAOS" pageId="16" pageNumber="172" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="FD53A2FA7156FFEF7EC8FE89FDB3FE18" box="[471,576,382,405]" pageId="16" pageNumber="172" type="holotype">Holotype</typeStatus>
:
<collectionCode id="44F9849D7156FFEF7D4CFE89FD5AFE18" box="[595,681,382,405]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="172">MNHN</collectionCode>
:
<specimenCount id="34EED7D17156FFEF7DA3FE89FD1FFE18" box="[700,748,382,405]" pageId="16" pageNumber="172" type="male">1 ♂</specimenCount>
;
<collectingCountry id="5AFF5CC87156FFEF7F88FE68FF12FE3B" box="[151,225,415,438]" name="Laos" pageId="16" pageNumber="172">
<collectionCode id="44F9849D7156FFEF7F88FE68FF12FE3B" box="[151,225,415,438]" collectionName="LAOS" pageId="16" pageNumber="172">LAOS</collectionCode>
</collectingCountry>
: //
<taxonomicName id="E5E867DB7156FFEF7E1DFE68FE1DFE3B" authorityName="Sharp" authorityYear="1876" box="[258,494,415,438]" class="Insecta" family="Melolonthidae" genus="Cyphochilus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="172" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="109CC04A7156FFEF7E1DFE68FE1DFE3B" box="[258,494,415,438]" italics="true" pageId="16" pageNumber="172">Cyphochilus obscurus</emphasis>
</taxonomicName>
/
<collectingCountry id="5AFF5CC87156FFEF7D1AFE68FDCFFE3B" box="[517,572,415,438]" name="Laos" pageId="16" pageNumber="172">Laos</collectingCountry>
.
<typeStatus id="FD53A2FA7156FFEF7D59FE68FD88FE3B" box="[582,635,415,438]" pageId="16" pageNumber="172">Type</typeStatus>
/
<collectorName id="8F1D798E7156FFEF7D8EFE68FD34FE3B" box="[657,711,415,438]" name="D. Sharp" pageId="16" pageNumber="172">D. S.</collectorName>
(
<collectionCode id="44F9849D7156FFEF7DC8FE68FD1FFE3B" box="[727,748,415,438]" pageId="16" pageNumber="172">H</collectionCode>
) //
<collectingCountry id="5AFF5CC87156FFEF7FB1FE37FF11FE5A" box="[174,226,448,471]" name="Laos" pageId="16" pageNumber="172">Laos</collectingCountry>
(
<collectionCode id="44F9849D7156FFEF7FEDFE37FEF4FE5A" box="[242,263,448,471]" pageId="16" pageNumber="172">H</collectionCode>
) / Mouhot (
<collectionCode id="44F9849D7156FFEF7E93FE37FE6FFE5A" box="[396,412,448,471]" pageId="16" pageNumber="172">P</collectionCode>
) // Ex Musaeo /
<collectorName id="8F1D798E7156FFEF7D49FE37FD45FE5A" box="[598,694,448,471]" pageId="16" pageNumber="172">D. Sharp</collectorName>
1890 (
<collectionCode id="44F9849D7156FFEF7F80FE17FF43FE7A" box="[159,176,480,503]" pageId="16" pageNumber="172">P</collectionCode>
) // Muséum Paris / 1952 /
<collectorName id="8F1D798E7156FFEF7EF4FE17FD5CFE7A" box="[491,687,480,503]" pageId="16" pageNumber="172">Coll. R. Oberthur</collectorName>
(
<collectionCode id="44F9849D7156FFEF7DDEFE17FD21FE7A" box="[705,722,480,503]" pageId="16" pageNumber="172">P</collectionCode>
) //
<typeStatus id="FD53A2FA7156FFEF7F88FDF6FF2BFD95" box="[151,216,513,536]" pageId="16" pageNumber="172">Typus</typeStatus>
(
<collectionCode id="44F9849D7156FFEF7FF9FDF6FF09FD95" box="[230,250,513,536]" pageId="16" pageNumber="172">H</collectionCode>
on red) //. (
<figureCitation id="BAD300DD7156FFEF7E67FDF6FE4EFD95" box="[376,445,513,536]" captionStart="Figs 1-14" captionStartId="7.[151,192,957,978]" captionTargetBox="[157,1435,257,896]" captionTargetId="figure-87@7.[151,1436,257,914]" captionTargetPageId="7" captionText="Figs 1-14. Original labels of type material. (1) Cyphochilus pygidialis Nonfried, 1893. (2) C. pygidialis Brenske [sic]. (3) C. septentrionalis Waterhouse, 1867. (4) C. obscurus Sharp, 1876. (5) C. proximus Sharp, 1876. (6) C. flavomarginatus Frey, 1971. (7) C. cylindricus Brenske, 1903. (8) Melolontha candida (Olivier, 1789). (9) Cyphochilus pygidialis v. angeri Nonfried, 1893. (10) C. feae Brenske, 1903. (11) C. vestitus Sharp, 1876. (12) C. tricolor Waterhouse, 1867. (13) C. testaceipes Fairmaire, 1902. (14) C. peninsularis Arrow, 1938." figureDoi="http://doi.org/10.5281/zenodo.5743431" httpUri="https://zenodo.org/record/5743431/files/figure.png" pageId="16" pageNumber="172">Figs 4</figureCitation>
,
<figureCitation id="BAD300DD7156FFEF7ED6FDF6FE14FD95" box="[457,487,513,536]" captionStart="Figs 80-91" captionStartId="23.[151,192,1883,1904]" captionTargetBox="[206,1382,268,1813]" captionTargetId="figure-116@23.[201,1387,257,1827]" captionTargetPageId="23" captionText="Figs 80-91. Habitus. (80) Cyphochilus candidus (Olivier, 1789), lectotype ♂ MNHN. (81) C. candidus (Olivier, 1789), Nepal: Godavari, ♂ MHNG, yellow scales aberration. (82) C. carinchebanus Brenske, 1903, Myanmar: Carin Cheba, ♂ ZMHB. (83) C. feae Brenske, 1903, holotype ZMHB. (84) C. gandhii n. sp., holotype MHNG. (85) C. leducthoi n. sp., holotype MHNG.(86) C. septentrionalis Waterhouse, 1867, lectotype BMNH. (87) C. obscurus Sharp, 1876, holotype MNHN. (88) C. sansuukyii n. sp., paratype ♀ MHNG. (89) C. peninsularis Arrow, 1938, paralectotype ♂ MHNG. (90) C. rohingyae n. sp., holotype MHNG. (91) C. rohingyae n. sp., paratype ♂ MHNG, Malaysia: Perak, bicolor aberration. – Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5743449" httpUri="https://zenodo.org/record/5743449/files/figure.png" pageId="16" pageNumber="172">87</figureCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="22571C587156FFEF7F88FDC6FD75FDC5" blockId="16.[151,756,561,812]" box="[151,646,561,584]" pageId="16" pageNumber="172">
<emphasis id="109CC04A7156FFEF7F88FDC6FE06FDC5" bold="true" box="[151,501,561,584]" pageId="16" pageNumber="172">Additional material examined:</emphasis>
<specimenCount id="34EED7D17156FFEF7EE4FDC6FD71FDC5" box="[507,642,561,584]" pageId="16" pageNumber="172" type="generic">8 specimens</specimenCount>
.
</paragraph>
<paragraph id="22571C587156FFEF7F88FDA6FDB8FD47" blockId="16.[151,756,561,812]" pageId="16" pageNumber="172">
<materialsCitation id="928016057156FFEF7F88FDA6FF0FFD04" ID-GBIF-Occurrence="3886339370" collectionCode="ISNB" country="Laos" location="Paklay" pageId="16" pageNumber="172" specimenCount="1" specimenCount-male="1">
<emphasis id="109CC04A7156FFEF7F88FDA6FF24FDE5" box="[151,215,593,616]" italics="true" pageId="16" pageNumber="172">
<collectingCountry id="5AFF5CC87156FFEF7F88FDA6FF24FDE5" box="[151,215,593,616]" name="Laos" pageId="16" pageNumber="172">
<collectionCode id="44F9849D7156FFEF7F88FDA6FF24FDE5" box="[151,215,593,616]" collectionName="LAOS" pageId="16" pageNumber="172">LAOS</collectionCode>
</collectingCountry>
</emphasis>
:
<collectionCode id="44F9849D7156FFEF7FF7FDA6FEDBFDE5" box="[232,296,593,616]" collectionName="Belgium, Brussels, Institut Royal des Sciences Naturelles de Belgique" pageId="16" pageNumber="172">ISNB</collectionCode>
;
<specimenCount id="34EED7D17156FFEF7E29FDA6FE91FDE5" box="[310,354,593,616]" pageId="16" pageNumber="172" type="male">1 ♂</specimenCount>
;
<location id="27374A837156FFEF7E6CFDA6FE4CFDE5" LSID="urn:lsid:plazi:treatment:AA41AD4E7156FFEF7E1BFEF7FA85FED4:27374A837156FFEF7E6CFDA6FE4CFDE5" box="[371,447,593,616]" country="Laos" name="Paklay" pageId="16" pageNumber="172">Paklay</location>
;
<date id="56563A987156FFEF7ED2FDA6FDAFFDE5" box="[461,604,593,616]" pageId="16" pageNumber="172" value="1917-08-15">15.VIII.1917</date>
; R.
<location id="27374A837156FFEF7D94FDA6FF3AFD04" LSID="urn:lsid:plazi:treatment:AA41AD4E7156FFEF7E1BFEF7FA85FED4:27374A837156FFEF7D94FDA6FF3AFD04" country="Laos" name="Vitalis de Salv" pageId="16" pageNumber="172">Vitalis de Salv</location>
[aza]
</materialsCitation>
–
<materialsCitation id="928016057156FFEF7E3BFD85FE6AFD24" ID-GBIF-Occurrence="3886339481" collectionCode="ISNB" country="Laos" location="Luang Prabang" pageId="16" pageNumber="172" specimenCount="1" specimenCount-male="1" stateProvince="Louangphabang">
<collectionCode id="44F9849D7156FFEF7E3BFD85FE97FD04" box="[292,356,626,649]" collectionName="Belgium, Brussels, Institut Royal des Sciences Naturelles de Belgique" pageId="16" pageNumber="172">ISNB</collectionCode>
;
<specimenCount id="34EED7D17156FFEF7E6EFD85FE6FFD04" box="[369,412,626,649]" pageId="16" pageNumber="172" type="male">1 ♂</specimenCount>
;
<collectingRegion id="E02CD2BA7156FFEF7EB3FD85FDA7FD04" box="[428,596,626,649]" country="Laos" name="Louangphabang" pageId="16" pageNumber="172">Luang Prabang</collectingRegion>
;
<date id="56563A987156FFEF7D7EFD85FD1CFD04" box="[609,751,626,649]" pageId="16" pageNumber="172" value="1917-08-27">27.VIII.1917</date>
; R.
<location id="27374A837156FFEF7FA5FD65FE97FD24" LSID="urn:lsid:plazi:treatment:AA41AD4E7156FFEF7E1BFEF7FA85FED4:27374A837156FFEF7FA5FD65FE97FD24" box="[186,356,658,681]" country="Laos" name="Vitalis de Salv" pageId="16" pageNumber="172">Vitalis de Salv</location>
[aza]
</materialsCitation>
. –
<materialsCitation id="928016057156FFEF7EDFFD65FDB4FD47" ID-GBIF-Occurrence="3886339465" collectionCode="ISNB" country="Laos" isTailEnumeration="true" location="Pak." municipality="Laos" pageId="16" pageNumber="172" specimenCount="2" specimenCount-male="2">
<collectionCode id="44F9849D7156FFEF7EDFFD65FDF3FD24" box="[448,512,658,681]" collectionName="Belgium, Brussels, Institut Royal des Sciences Naturelles de Belgique" pageId="16" pageNumber="172">ISNB</collectionCode>
;
<specimenCount id="34EED7D17156FFEF7D10FD65FDCFFD24" box="[527,572,658,681]" pageId="16" pageNumber="172" type="male">2 ♂</specimenCount>
;
<collectingMunicipality id="C23386227156FFEF7D50FD65FD75FD24" box="[591,646,658,681]" pageId="16" pageNumber="172">
<collectingCountry id="5AFF5CC87156FFEF7D50FD65FD75FD24" box="[591,646,658,681]" name="Laos" pageId="16" pageNumber="172">Laos</collectingCountry>
</collectingMunicipality>
,
<location id="27374A837156FFEF7D8AFD65FD37FD24" LSID="urn:lsid:plazi:treatment:AA41AD4E7156FFEF7E1BFEF7FA85FED4:27374A837156FFEF7D8AFD65FD37FD24" box="[661,708,658,681]" country="Laos" municipality="Laos" name="Pak." pageId="16" pageNumber="172">Pak.</location>
[?];
<date id="56563A987156FFEF7F88FD44FEE7FD47" box="[151,276,691,714]" pageId="16" pageNumber="172" value="1919-09-24">24.IX.1919</date>
; coll. R.
<location id="27374A837156FFEF7E6CFD44FDE0FD47" LSID="urn:lsid:plazi:treatment:AA41AD4E7156FFEF7E1BFEF7FA85FED4:27374A837156FFEF7E6CFD44FDE0FD47" box="[371,531,691,714]" country="Laos" name="Vitalis de Salv" pageId="16" pageNumber="172">Vitalis de Salv</location>
[aza]
</materialsCitation>
.
</paragraph>
<paragraph id="22571C587156FFEF7F88FD23FDD0FCA1" blockId="16.[151,756,561,812]" pageId="16" pageNumber="172">
<materialsCitation id="928016057156FFEF7F88FD23FD20FD66" ID-GBIF-Occurrence="3886339421" box="[151,723,724,747]" collectionCode="PCMN" country="Thailand" location="Chiang Mai" pageId="16" pageNumber="172" specimenCount="2" specimenCount-male="2" stateProvince="Chiang Mai">
<emphasis id="109CC04A7156FFEF7F88FD23FEEBFD66" box="[151,280,724,747]" italics="true" pageId="16" pageNumber="172">
<collectingCountry id="5AFF5CC87156FFEF7F88FD23FEEBFD66" box="[151,280,724,747]" name="Thailand" pageId="16" pageNumber="172">THAILAND</collectingCountry>
</emphasis>
:
<collectionCode id="44F9849D7156FFEF7E2FFD23FE73FD66" box="[304,384,724,747]" pageId="16" pageNumber="172">PCMN</collectionCode>
;
<specimenCount id="34EED7D17156FFEF7E8BFD23FE35FD66" box="[404,454,724,747]" pageId="16" pageNumber="172" type="male">2 ♂</specimenCount>
;
<collectingRegion id="E02CD2BA7156FFEF7EC1FD23FD98FD66" box="[478,619,724,747]" country="Thailand" name="Chiang Mai" pageId="16" pageNumber="172">Chiang Mai</collectingRegion>
;
<date id="56563A987156FFEF7D60FD23FD20FD66" box="[639,723,724,747]" pageId="16" pageNumber="172" value="1989-10">X.1989</date>
</materialsCitation>
. –
<materialsCitation id="928016057156FFEF7F88FD03FE15FC86" ID-GBIF-Occurrence="3886339503" box="[151,486,756,779]" collectionCode="MHNG" country="Thailand" location="Nan" pageId="16" pageNumber="172" specimenCount="3" specimenCount-male="3" stateProvince="Nan">
<collectionCode id="44F9849D7156FFEF7F88FD03FF1FFC86" box="[151,236,756,779]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="16" pageNumber="172" type="Museum">MHNG</collectionCode>
;
<specimenCount id="34EED7D17156FFEF7FE4FD03FED4FC86" box="[251,295,756,779]" pageId="16" pageNumber="172" type="male">3 ♂</specimenCount>
;
<collectingRegion id="E02CD2BA7156FFEF7E26FD03FE9AFC86" box="[313,361,756,779]" country="Thailand" name="Nan" pageId="16" pageNumber="172">Nan</collectingRegion>
;
<date id="56563A987156FFEF7E67FD03FE15FC86" box="[376,486,756,779]" pageId="16" pageNumber="172" value="1993-08">VIII.1993</date>
</materialsCitation>
. –
<materialsCitation id="928016057156FFEF7D13FD03FDECFCA1" ID-GBIF-Occurrence="3886339447" collectionCode="MHNG" country="Thailand" isTailEnumeration="true" location="Wiang Papao" pageId="16" pageNumber="172" specimenCount="3" specimenCount-male="3" stateProvince="Chiang Mai">
<collectionCode id="44F9849D7156FFEF7D13FD03FD92FC86" box="[524,609,756,779]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="16" pageNumber="172" type="Museum">MHNG</collectionCode>
;
<specimenCount id="34EED7D17156FFEF7D6FFD03FD6FFC86" box="[624,668,756,779]" pageId="16" pageNumber="172" type="male">3 ♂</specimenCount>
;
<location id="27374A837156FFEF7DB1FD03FF28FCA1" LSID="urn:lsid:plazi:treatment:AA41AD4E7156FFEF7E1BFEF7FA85FED4:27374A837156FFEF7DB1FD03FF28FCA1" country="Thailand" name="Wiang Papao" pageId="16" pageNumber="172">Wiang Papao</location>
,
<quantity id="E510B1BD7156FFEF7FF9FCE2FEDAFCA1" box="[230,297,789,812]" metricMagnitude="4" metricUnit="m" metricValue="8.0" pageId="16" pageNumber="172" unit="km" value="80.0">80 km</quantity>
NE
<collectingRegion id="E02CD2BA7156FFEF7E45FCE2FE2FFCA1" box="[346,476,789,812]" country="Thailand" name="Chiang Mai" pageId="16" pageNumber="172">Chiang Mai</collectingRegion>
, 1992
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="6AF24FD37156FFEF7F88FCB2FE99F924" pageId="16" pageNumber="172" type="description">
<paragraph id="22571C587156FFEF7F88FCB2FD2AFA65" blockId="16.[151,757,837,1512]" pageId="16" pageNumber="172">
<emphasis id="109CC04A7156FFEF7F88FCB2FD76FCD1" bold="true" box="[151,645,837,860]" pageId="16" pageNumber="172">
Redescription of the
<typeStatus id="FD53A2FA7156FFEF7EDAFCB2FDDAFCD1" box="[453,553,837,860]" pageId="16" pageNumber="172" type="holotype">holotype</typeStatus>
male:
</emphasis>
<emphasis id="109CC04A7156FFEF7DBFFCB2FD3FFCD1" box="[672,716,837,860]" italics="true" pageId="16" pageNumber="172">Size</emphasis>
– BL:
<quantity id="E510B1BD7156FFEF7FD4FC92FEC3FCF1" box="[203,304,869,892]" metricMagnitude="-2" metricUnit="m" metricValue="1.64" pageId="16" pageNumber="172" unit="mm" value="16.4">16.4 mm</quantity>
, BW:
<quantity id="E510B1BD7156FFEF7E65FC92FE4DFCF1" box="[378,446,869,892]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="16" pageNumber="172" unit="mm" value="7.0">7 mm</quantity>
, BWX:
<quantity id="E510B1BD7156FFEF7D02FC92FD87FCF1" box="[541,628,869,892]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="16" pageNumber="172" unit="mm" value="8.7">8.7 mm</quantity>
, situated at midpoint of elytra.
<emphasis id="109CC04A7156FFEF7E64FC71FE49FC10" box="[379,442,902,925]" italics="true" pageId="16" pageNumber="172">Color</emphasis>
– Integument light brown; surface with white scales densely aggregated on the sides of pronotum.
<emphasis id="109CC04A7156FFEF7E71FC30FE5BFC53" box="[366,424,967,990]" italics="true" pageId="16" pageNumber="172">Head</emphasis>
– CW/L: 2.6; clypeus largely rounded; (F/O: 3.4); antennal club slightly longer than antennomeres 2-7 (A2-7/CL: 0.8).
<emphasis id="109CC04A7156FFEF7D0FFBFFFD89FB92" box="[528,634,1032,1055]" italics="true" pageId="16" pageNumber="172">Pronotum</emphasis>
– (PnW/L: 2); apical angles and lateral margins largely flattened, apical angles right, not protuberant; basal lateral margins straight, basal angles obtuse.
<emphasis id="109CC04A7156FFEF7D24FB9DFD8DFB0C" box="[571,638,1130,1153]" italics="true" pageId="16" pageNumber="172">Elytra</emphasis>
– Without any visible striae.
<emphasis id="109CC04A7156FFEF7E41FB7CFDFAFB2F" box="[350,521,1163,1186]" italics="true" pageId="16" pageNumber="172">Thoracic sterna</emphasis>
– Surface with short, white pubescence and scales; mesosternum feebly developed (MstL:
<quantity id="E510B1BD7156FFEF7E7DFB3BFE35FB6E" box="[354,454,1228,1251]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" pageId="16" pageNumber="172" unit="mm" value="0.47">0.47 mm</quantity>
) however, visible in lateral view between the mesocoxae.
<emphasis id="109CC04A7156FFEF7D1CFB1AFD9AFA89" box="[515,617,1261,1284]" italics="true" pageId="16" pageNumber="172">Abdomen</emphasis>
– Sternites only with scattered scales; pygidium convex with apical margin very narrowly reflected.
<emphasis id="109CC04A7156FFEF7D55FAD9FD8EFAC8" box="[586,637,1326,1349]" italics="true" pageId="16" pageNumber="172">Legs</emphasis>
– Protibia tridentate with the basal tooth vestigial; slender apical tooth as long as the middle tooth; claws of all tarsi bifid at apex.
<emphasis id="109CC04A7156FFEF7E1FFA67FE9AFA2A" box="[256,361,1424,1447]" italics="true" pageId="16" pageNumber="172">Aedeagus</emphasis>
– Parameres strongly asymmetric with 5 appendices 50-52; the right paramere in form of club with an apical spike directed externally (
<figureCitation id="BAD300DD7156FFEF7D62FA26FD3EFA65" box="[637,717,1489,1512]" captionStart="Figs 47-58" captionStartId="13.[151,192,1959,1980]" captionTargetBox="[164,1425,260,1900]" captionTargetId="figure-7@13.[151,1436,254,1945]" captionTargetPageId="13" captionText="Figs 47-58. Parameres in right, dorsal, and left view. (47-49) Cyphochilus carinchebanus Brenske, 1903. (50-52) C. obscurus Sharp, 1876, holotype. (53-55) C. proximus Sharp, 1876, holotype. (56-58) C. tricolor Waterhouse, 1867, lectotype. – Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743441" httpUri="https://zenodo.org/record/5743441/files/figure.png" pageId="16" pageNumber="172">Fig. 51</figureCitation>
).
</paragraph>
<paragraph id="22571C587156FFEF7F88F9F6FE67F9F4" blockId="16.[151,756,1537,1657]" pageId="16" pageNumber="172">
<emphasis id="109CC04A7156FFEF7F88F9F6FEE9F995" bold="true" box="[151,282,1537,1560]" pageId="16" pageNumber="172">Variability:</emphasis>
BL:
<quantity id="E510B1BD7156FFEF7E4EF9F6FE24F995" box="[337,471,1537,1560]" metricMagnitude="-2" metricUnit="m" metricValue="1.675" metricValueMax="1.75" metricValueMin="1.6" pageId="16" pageNumber="172" unit="mm" value="16.75" valueMax="17.5" valueMin="16.0">16-17.5 mm</quantity>
. Specimens from
<collectingCountry id="5AFF5CC87156FFEF7D8AF9F6FD07F995" box="[661,756,1537,1560]" name="Thailand" pageId="16" pageNumber="172">Thailand</collectingCountry>
have the vestiture with ochre scales instead of white, however, this is a variation observed also in other species of
<taxonomicName id="E5E867DB7156FFEF7E17F995FE7EF9F4" authorityName="Waterhouse" authorityYear="1867" box="[264,397,1634,1657]" class="Insecta" family="Melolonthidae" genus="Cyphochilus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="109CC04A7156FFEF7E17F995FE7EF9F4" box="[264,397,1634,1657]" italics="true" pageId="16" pageNumber="172">Cyphochilus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="22571C587156FFEF7F88F965FE99F924" blockId="16.[151,362,1682,1705]" box="[151,362,1682,1705]" pageId="16" pageNumber="172">
<emphasis id="109CC04A7156FFEF7F88F965FF00F924" bold="true" box="[151,243,1682,1705]" pageId="16" pageNumber="172">Female:</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="6AF24FD37156FFEF7F88F935FD55F954" box="[151,678,1730,1753]" pageId="16" pageNumber="172" type="materials_examined">
<paragraph id="22571C587156FFEF7F88F935FD55F954" blockId="16.[151,678,1730,1753]" box="[151,678,1730,1753]" pageId="16" pageNumber="172">
<emphasis id="109CC04A7156FFEF7F88F935FEC1F954" bold="true" box="[151,306,1730,1753]" pageId="16" pageNumber="172">
<typeStatus id="FD53A2FA7156FFEF7F88F935FF3CF954" box="[151,207,1730,1753]" pageId="16" pageNumber="172">Type</typeStatus>
locality:
</emphasis>
The
<typeStatus id="FD53A2FA7156FFEF7E77F935FE65F954" box="[360,406,1730,1753]" pageId="16" pageNumber="172">type</typeStatus>
locality is given as
<collectingCountry id="5AFF5CC87156FFEF7D74F935FD51F954" box="[619,674,1730,1753]" name="Laos" pageId="16" pageNumber="172">Laos</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="6AF24FD37156FFEF7F88F905FD34F8A4" pageId="16" pageNumber="172" type="distribution">
<paragraph id="22571C587156FFEF7F88F905FD34F8A4" blockId="16.[151,756,1778,1833]" pageId="16" pageNumber="172">
<emphasis id="109CC04A7156FFEF7F88F905FEDEF884" bold="true" box="[151,301,1778,1801]" pageId="16" pageNumber="172">Distribution:</emphasis>
The species is known from only a few specimens, all from North
<collectingCountry id="5AFF5CC87156FFEF7EABF8E5FE1BF8A4" box="[436,488,1810,1833]" name="Laos" pageId="16" pageNumber="172">Laos</collectingCountry>
and North
<collectingCountry id="5AFF5CC87156FFEF7D7FF8E5FD31F8A4" box="[608,706,1810,1833]" name="Thailand" pageId="16" pageNumber="172">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="6AF24FD37156FFEF7F88F8B5FA85FED4" pageId="16" pageNumber="172" type="discussion">
<paragraph id="22571C587156FFEF7F88F8B5FA85FED4" blockId="16.[151,757,1858,2077]" lastBlockId="16.[831,1437,258,345]" pageId="16" pageNumber="172">
<emphasis id="109CC04A7156FFEF7F88F8B5FEFAF8D4" bold="true" box="[151,265,1858,1881]" pageId="16" pageNumber="172">Remarks:</emphasis>
<bibRefCitation id="467961A97156FFEF7E04F8B5FE40F8D4" author="Sharp D." box="[283,435,1858,1881]" pageId="16" pageNumber="172" pagination="65 - 90" refId="ref20538" refString="Sharp D. 1876. Description of some new species of Scarabaeidae from tropical Asia and Malaisia. Part III (Melolonthini). Coleopterologische Hefte 15: 65 - 90." type="journal article" year="1876">Sharp (1876)</bibRefCitation>
thought that the
<typeStatus id="FD53A2FA7156FFEF7D8AF8B5FD07F8D4" box="[661,756,1858,1881]" pageId="16" pageNumber="172" type="holotype">holotype</typeStatus>
of
<taxonomicName id="E5E867DB7156FFEF7FA4F894FEB2F8F7" authorityName="Sharp" authorityYear="1876" box="[187,321,1891,1914]" class="Insecta" family="Melolonthidae" genus="Cyphochilus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="172" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="109CC04A7156FFEF7FA4F894FEB2F8F7" box="[187,321,1891,1914]" italics="true" pageId="16" pageNumber="172">C. obscurus</emphasis>
</taxonomicName>
was a female, however it is a male.
<taxonomicName id="E5E867DB7156FFEF7F88F874FE76F817" authorityName="Sharp" authorityYear="1876" box="[151,389,1923,1946]" class="Insecta" family="Melolonthidae" genus="Cyphochilus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="172" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="109CC04A7156FFEF7F88F874FE76F817" box="[151,389,1923,1946]" italics="true" pageId="16" pageNumber="172">Cyphochilus obscurus</emphasis>
</taxonomicName>
can be readily separated from all other congeners by the unique form of the parameres. Moreover, the bifid claws are an astonishing character since all other
<taxonomicName id="E5E867DB7156FFEF7E57F812FE3CF871" authorityName="Waterhouse" authorityYear="1867" box="[328,463,2021,2044]" class="Insecta" family="Melolonthidae" genus="Cyphochilus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="109CC04A7156FFEF7E57F812FE3CF871" box="[328,463,2021,2044]" italics="true" pageId="16" pageNumber="172">Cyphochilus</emphasis>
</taxonomicName>
have claws with a thorn in medial position, never bifid. Actually, this is a quite strong character utilized to separate group of genera in the
<taxonomicName id="E5E867DB7156FFEF7C76FED5FC00FEB4" baseAuthorityName="Medvedev" baseAuthorityYear="1951" box="[873,1011,290,313]" class="Insecta" family="Melolonthidae" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="172" phylum="Arthropoda" rank="tribe" tribe="Rhizotrogini">Rhizotrogini</taxonomicName>
, however, all other characters attribute this species to genus
<taxonomicName id="E5E867DB7156FFEF7B3EFEB5FB55FED4" authorityName="Waterhouse" authorityYear="1867" box="[1057,1190,322,345]" class="Insecta" family="Melolonthidae" genus="Cyphochilus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="109CC04A7156FFEF7B3EFEB5FB55FED4" box="[1057,1190,322,345]" italics="true" pageId="16" pageNumber="172">Cyphochilus</emphasis>
</taxonomicName>
without any doubt.
</paragraph>
</subSubSection>
</treatment>
</document>
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