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<document id="988958B5305E4628EF69138991E88B97" ID-DOI="10.11646/zootaxa.4500.3.9" ID-ISSN="1175-5326" ID-Zenodo-Dep="5297742" ID-ZooBank="D4CE78C0-77A2-46B5-95BD-F0CB2E209676" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630086432396" checkinUser="felipe" docAuthor="Santis, Marcelo Domingos De" docDate="2018" docId="631787C0FFC4FFE166D5FD4F27E1601D" docLanguage="en" docName="zootaxa.4500.3.9.pdf" docOrigin="Zootaxa 4500 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cryptosophia Santis 2018, gen. nov." docType="treatment" docVersion="4" lastPageNumber="436" masterDocId="9F2EFFB8FFC6FFE26642FFD525416100" masterDocTitle="A new genus and species of Sophiini Townsend, 1936 (Diptera: Tachinidae) from Brazil, with an updated key to the genera of the tribe" masterLastPageNumber="442" masterPageNumber="433" pageNumber="435" updateTime="1699141723882" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="BF294ED301E213B7AC23126E87C3787A">A new genus and species of Sophiini Townsend, 1936 (Diptera: Tachinidae) from Brazil, with an updated key to the genera of the tribe</mods:title>
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<mods:name id="887EE407B33587FCEB8E9EE2AB7BA677" type="personal">
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<mods:namePart id="1AB0C30D9BBEE8DB5555A4867CE39D91">Santis, Marcelo Domingos De</mods:namePart>
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<mods:date id="4D09EBCD0B4DEC9C6EEB36AFCDA63546">2018</mods:date>
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<mods:number id="DE529E39453D6C887BD21EEDB67DFE53">2018-10-17</mods:number>
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<mods:identifier id="3F5DEFD8641CEC064D8646138D13E592" type="ISSN">1175-5326</mods:identifier>
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<treatment id="631787C0FFC4FFE166D5FD4F27E1601D" ID-DOI="http://doi.org/10.5281/zenodo.5304529" ID-Zenodo-Dep="5304529" LSID="urn:lsid:plazi:treatment:631787C0FFC4FFE166D5FD4F27E1601D" httpUri="http://treatment.plazi.org/id/631787C0FFC4FFE166D5FD4F27E1601D" lastPageId="3" lastPageNumber="436" pageId="2" pageNumber="435">
<subSubSection id="A3A4655DFFC4FFE066D5FD4F24EC63B4" box="[151,429,666,692]" pageId="2" pageNumber="435" type="nomenclature">
<paragraph id="EB0136D6FFC4FFE066D5FD4F24EC63B4" blockId="2.[151,429,666,692]" box="[151,429,666,692]" pageId="2" pageNumber="435">
<heading id="B04981BAFFC4FFE066D5FD4F24EC63B4" bold="true" box="[151,429,666,692]" fontSize="11" level="1" pageId="2" pageNumber="435" reason="1">
<emphasis id="D9CAEAC4FFC4FFE066D5FD4F24EC63B4" bold="true" box="[151,429,666,692]" pageId="2" pageNumber="435">
<taxonomicName id="2CBE4D55FFC4FFE066D5FD4F247963B4" ID-CoL="89PZQ" authority="Santis, 2018" authorityName="Santis" authorityYear="2018" box="[151,312,666,692]" class="Insecta" family="Tachinidae" genus="Cryptosophia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="435" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="D9CAEAC4FFC4FFE066D5FD4F247963B4" bold="true" box="[151,312,666,692]" italics="true" pageId="2" pageNumber="435">Cryptosophia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C2F957BFFFC4FFE0677DFD4F24EC63B4" box="[319,429,666,692]" pageId="2" pageNumber="435" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A3A4655DFFC4FFE066D5FD34245C6242" pageId="2" pageNumber="435" type="etymology">
<paragraph id="EB0136D6FFC4FFE066D5FD34245C6242" blockId="2.[151,1437,737,2024]" pageId="2" pageNumber="435">
<emphasis id="D9CAEAC4FFC4FFE066D5FD34245F63FA" bold="true" box="[151,286,737,762]" inLineHeading="true" pageId="2" pageNumber="435" reason="1">Etymology.</emphasis>
Feminine name derived from the combination of the Greek adjective “
<emphasis id="D9CAEAC4FFC4FFE0621CFD3421E463FA" box="[1118,1189,737,762]" italics="true" pageId="2" pageNumber="435">krypto</emphasis>
” (= hidden), and the generic name
<taxonomicName id="2CBE4D55FFC4FFE0677AFCD124C7621D" authorityName="Robineau-Desvoidy" authorityYear="1830" box="[312,390,772,797]" class="Insecta" family="Tachinidae" genus="Sophia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="435" phylum="Arthropoda" rank="genus">
<emphasis id="D9CAEAC4FFC4FFE0677AFCD124C7621D" box="[312,390,772,797]" italics="true" pageId="2" pageNumber="435">Sophia</emphasis>
</taxonomicName>
, in reference to the meron with pale yellow setae almost indistinguishable from the yellow integument.
</paragraph>
</subSubSection>
<subSubSection id="A3A4655DFFC4FFE06685FC9926A96265" box="[199,1000,844,869]" pageId="2" pageNumber="435" type="type_taxon">
<paragraph id="EB0136D6FFC4FFE06685FC9926A96265" blockId="2.[151,1437,737,2024]" box="[199,1000,844,869]" pageId="2" pageNumber="435">
<emphasis id="D9CAEAC4FFC4FFE06685FC9924236265" bold="true" box="[199,354,844,869]" inLineHeading="true" pageId="2" pageNumber="435" reason="1">
<typeStatus id="34058874FFC4FFE06685FC9924436265" box="[199,258,844,869]" pageId="2" pageNumber="435">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="2CBE4D55FFC4FFE0672BFC9927346265" ID-CoL="8BM67" authority="Santis, 2018" authorityName="Santis" authorityYear="2018" box="[361,629,844,869]" class="Insecta" family="Tachinidae" genus="Cryptosophia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="435" phylum="Arthropoda" rank="species" species="aurulenta" status="sp. nov.">
<emphasis id="D9CAEAC4FFC4FFE0672BFC9927346265" box="[361,629,844,869]" italics="true" pageId="2" pageNumber="435">Cryptosophia aurulenta</emphasis>
</taxonomicName>
<emphasis id="D9CAEAC4FFC4FFE0643EFC9927976265" bold="true" box="[636,726,844,869]" pageId="2" pageNumber="435">
<taxonomicNameLabel id="C2F957BFFFC4FFE0643EFC9927976265" box="[636,726,844,869]" pageId="2" pageNumber="435" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, by present designation.
</paragraph>
</subSubSection>
<subSubSection id="A3A4655DFFC4FFE06685FCA426A8653E" pageId="2" pageNumber="435" type="diagnosis">
<paragraph id="EB0136D6FFC4FFE06685FCA426A8653E" blockId="2.[151,1437,737,2024]" pageId="2" pageNumber="435">
<emphasis id="D9CAEAC4FFC4FFE06685FCA42400628A" bold="true" box="[199,321,881,906]" inLineHeading="true" pageId="2" pageNumber="435" reason="1">Diagnosis.</emphasis>
<materialsCitation id="5BD63C8BFFC4FFE06709FCA426A5653E" collectionCode="V" pageId="2" pageNumber="435" specimenCount="1" typeStatus="paratype">
Head with occiput clearly swollen medially. Thorax with short setae; proepisternal seta lacking; anepisternum with pale yellow setae (but
<specimenCount id="FDB8FD5FFFC4FFE0642AFC4027B962AD" box="[616,760,917,941]" pageId="2" pageNumber="435" type="generic" typeStatus="paratype">one paratype</specimenCount>
with 23 black setae); meron with pale yellow setae, almost indistinguishable from the yellow integument; scutellum with one pair of weak, short and straight apical setae. Fore, mid and hind coxae with anterior surface bare. Abdomen with tergites III to
<collectionCode id="8DAFAE13FFC4FFE0627FFC08211062F4" box="[1085,1105,989,1012]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="435" type="Museum">V</collectionCode>
entirely pale yellow, without lateral marginal setae. First instar larva with segments 5 to 11 with small and subrectangular plates dorsally (without a pore), each with a spine-like projection on posterolateral surface
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A3A4655DFFC4FFE16685FB9C27E1601D" lastPageId="3" lastPageNumber="436" pageId="2" pageNumber="435" type="description">
<paragraph id="EB0136D6FFC4FFE06685FB9C24616712" blockId="2.[151,1437,737,2024]" pageId="2" pageNumber="435">
<emphasis id="D9CAEAC4FFC4FFE06685FB9C24186562" bold="true" box="[199,345,1097,1122]" inLineHeading="true" pageId="2" pageNumber="435" reason="1">Description.</emphasis>
Coloration. Occiput with golden pruinosity, but dorsal 1/4 light brown, with surrounding golden pruinose vitta along median occipital sclerite. Median occipital sclerite light brown. Occiput with short and pale setulae. Head with pale golden pruinosity, with anterior part of gena light brown. Face dark yellow. Thorax with prescutum mostly light brown, with golden pruinosity on postpronotal lobe, forming an anterodorsal spot adjacent to postpronotal lobe, and on median vittae and posterior horizontal vittae; postscutum mostly light brown, with golden pruinosity in the form of a triangle (reaching posterior margin), and with a paired lateral vitta slightly fading to light brown (ground color) anteriorly; postalar callus with golden pruinosity anteriorly (
<figureCitation id="73852A53FFC4FFE062F5FAF42053643A" box="[1207,1298,1313,1338]" captionStart="FIGURE 2" captionStartId="5.[151,250,1817,1839]" captionTargetBox="[151,1435,193,1792]" captionTargetId="figure-88@5.[150,1436,193,1796]" captionTargetPageId="5" captionText="FIGURE 2. Cryptosophia aurulenta gen. and sp nov., holotype ♀ (INPA). A. Lateral habitus. B. Dorsal habitus. C. Head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5297746" httpUri="https://zenodo.org/record/5297746/files/figure.png" pageId="2" pageNumber="435">Fig. 2B</figureCitation>
). Scutellum golden pruinose, with anterior horizontal vittae light brown. Subscutellum and postnotum light brown. Wing hyaline. Anepisternum with pale yellow setae (but
<specimenCount id="FDB8FD5FFFC4FFE064AFFABF26C16482" box="[749,896,1386,1410]" pageId="2" pageNumber="435" type="generic" typeStatus="paratype">one paratype</specimenCount>
with 23 black setae) setae. Meron with pale yellow setae, almost indistinguishable from the yellow integument. Thorax light brown with golden pruinosity. Anepimeron with two irregular light brown spots. Anterior (plus adjacent area) and posterior spiracles light brown. Abdomen mostly pale yellow, but syntergite I+II with a light brown median longitudinal stripe covering about anterior 1/4.
</paragraph>
<paragraph id="EB0136D6FFC4FFE06685F9C824D467C4" blockId="2.[151,1437,737,2024]" pageId="2" pageNumber="435">
Head (
<figureCitation id="73852A53FFC4FFE06751F9C8242F6736" box="[275,366,1565,1590]" captionStart="FIGURE 2" captionStartId="5.[151,250,1817,1839]" captionTargetBox="[151,1435,193,1792]" captionTargetId="figure-88@5.[150,1436,193,1796]" captionTargetPageId="5" captionText="FIGURE 2. Cryptosophia aurulenta gen. and sp nov., holotype ♀ (INPA). A. Lateral habitus. B. Dorsal habitus. C. Head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5297746" httpUri="https://zenodo.org/record/5297746/files/figure.png" pageId="2" pageNumber="435">Fig. 2C</figureCitation>
). Occiput clearly swollen medially. Fronto-orbital plate with 5 to 6 pairs of proclinate frontal setae, lowermost frontal seta not reaching antennal insertion. Fronto-orbital plate and parafacial bare. Frontal vitta somewhat widened anteriorly. Facial ridge with 1 setula in lower third. Lower facial margin not protruding, invisible in profile. Gena height about 5.5 x eye height. Vibrissa long, inserted at level of lower facial margin. Five to 7 subvibrissal setae.
</paragraph>
<paragraph id="EB0136D6FFC4FFE06685F90427C366E8" blockId="2.[151,1437,737,2024]" pageId="2" pageNumber="435">
Thorax (
<figureCitation id="73852A53FFC4FFE0676AF90424C067EA" box="[296,385,1745,1770]" captionStart="FIGURE 2" captionStartId="5.[151,250,1817,1839]" captionTargetBox="[151,1435,193,1792]" captionTargetId="figure-88@5.[150,1436,193,1796]" captionTargetPageId="5" captionText="FIGURE 2. Cryptosophia aurulenta gen. and sp nov., holotype ♀ (INPA). A. Lateral habitus. B. Dorsal habitus. C. Head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5297746" httpUri="https://zenodo.org/record/5297746/files/figure.png" pageId="2" pageNumber="435">Fig. 2B</figureCitation>
). Acrostichal setae 0+0. Dorsocentral setae 34+4, first post-sutural seta weak. Proepimeron with 1 weak and short seta. Meron with 24 setae. Anepisternum with 36 weak setae. Scutellum with one pair of weak basal, one pair of long and divergent subapical, and one pair of weak, short and straight apical setae. Subscutellum not well developed (in contrast to the well-developed subscutellum usually present in
<taxonomicName id="2CBE4D55FFC4FFE06350F8E820D36656" box="[1298,1426,1853,1878]" class="Insecta" family="Tachinidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="435" phylum="Arthropoda" rank="family">Tachinidae</taxonomicName>
). Postmetacoxal area membranous.
<emphasis id="D9CAEAC4FFC4FFE06455F8B727106679" box="[535,593,1890,1913]" italics="true" pageId="2" pageNumber="435">Wing</emphasis>
. Costal vein with only one strong and long seta. M vein bent towards vein R
<subScript id="773A3493FFC4FFE066EBF841258066A2" attach="left" box="[169,193,1940,1954]" fontSize="6" pageId="2" pageNumber="435">4+5</subScript>
at an angle of less than 90º, and somewhat convex after bend.
<emphasis id="D9CAEAC4FFC4FFE0653FF85326F3669D" box="[893,946,1926,1949]" italics="true" pageId="2" pageNumber="435">Legs</emphasis>
. Fore tarsus enlarged, compressed laterally, and nearly as long as tibia. Fore, mid and hind coxae with anterior face bare, with setae only in distal margin; hind tibia with one seta on posterodorsal margin.
</paragraph>
<paragraph id="EB0136D6FFC5FFE16685FF4227DE61D5" blockId="3.[151,1437,151,285]" pageId="3" pageNumber="436">
Abdomen (
<figureCitation id="73852A53FFC5FFE16707FF42248061B0" box="[325,449,151,176]" captionStart="FIGURE 2" captionStartId="5.[151,250,1817,1839]" captionTargetBox="[151,1435,193,1792]" captionTargetId="figure-88@5.[150,1436,193,1796]" captionTargetPageId="5" captionText="FIGURE 2. Cryptosophia aurulenta gen. and sp nov., holotype ♀ (INPA). A. Lateral habitus. B. Dorsal habitus. C. Head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5297746" httpUri="https://zenodo.org/record/5297746/files/figure.png" pageId="3" pageNumber="436">Fig. 2AB</figureCitation>
). Syntergite I+II with mid-dorsal longitudinal depression extending to posterior margin. Sternites II to V absent (wholly membranous).
</paragraph>
<paragraph id="EB0136D6FFC5FFE16685FF3527E1601D" blockId="3.[151,1437,151,285]" pageId="3" pageNumber="436">
<emphasis id="D9CAEAC4FFC5FFE16685FF3524CE61F8" box="[199,399,223,248]" italics="true" pageId="3" pageNumber="436">First instar larva.</emphasis>
Segments 5 to 11 with small and subrectangular plates dorsally (without a pore), each with a spine-like projection on posterolateral surface.
</paragraph>
</subSubSection>
</treatment>
</document>