152 lines
21 KiB
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152 lines
21 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.778.25775" ID-GBIF-Dataset="a051d02c-d2a6-48d8-9b52-243409f7735c" ID-PMC="PMC6082922" ID-Pensoft-Pub="1313-2970-778-1" ID-PubMed="30104914" ID-ZBK="C61BB5E5A1DB483E83403DB71A69191F" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-778-1" ModsDocOrigin="ZooKeys 778" ModsDocTitle="Revision of the World species of the genus Chromoteleia Ashmead (Hymenoptera, Platygastridae, Scelioninae)" checkinTime="1533208250173" checkinUser="pensoft" docAuthor="Chen, Hua-yan, Talamas, Elijah J., Valerio, Alejandro A., Masner, Lubomir & Johnson, Norman F." docDate="2018" docId="5FA364839D92AC01DD426FF5AB983D5B" docLanguage="en" docName="ZooKeys 778: 1-95" docOrigin="ZooKeys 778" docSource="http://dx.doi.org/10.3897/zookeys.778.25775" docTitle="Chromoteleia connectens Kieffer" docType="treatment" docVersion="4" lastPageNumber="5" masterDocId="FF9C4529B34F3746FFDD5736FB20853F" masterDocTitle="Revision of the World species of the genus Chromoteleia Ashmead (Hymenoptera, Platygastridae, Scelioninae)" masterLastPageNumber="95" masterPageNumber="1" pageNumber="5" updateTime="1668166106366" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the World species of the genus Chromoteleia Ashmead (Hymenoptera, Platygastridae, Scelioninae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Chen, Hua-yan</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Talamas, Elijah J.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Valerio, Alejandro A.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Masner, Lubomir</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Johnson, Norman F.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2018</mods:date>
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<mods:detail type="volume">
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<mods:number>778</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>95</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.778.25775</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.778.25775</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-778-1</mods:identifier>
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<mods:identifier type="ZBK">C61BB5E5A1DB483E83403DB71A69191F</mods:identifier>
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<mods:identifier type="ZooBank">C61BB5E5A1DB483E83403DB71A69191F</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="146770667" LSID="urn:lsid:plazi:treatment:5FA364839D92AC01DD426FF5AB983D5B" httpUri="http://treatment.plazi.org/id/5FA364839D92AC01DD426FF5AB983D5B" lastPageNumber="5" pageId="4" pageNumber="5">
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<subSubSection pageId="4" pageNumber="5" type="nomenclature">
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<paragraph pageId="4" pageNumber="5">
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<taxonomicName LSID="http://zoobank.org/7409F8CA-0594-4232-8481-FCC20CDC290E" authority="Kieffer" class="Insecta" family="Scelionidae" genus="Chromoteleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chromoteleia connectens" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="connectens">Chromoteleia connectens Kieffer</taxonomicName>
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Figures 14, 58-63, 64-69, 237
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="5" type="reference_group">
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<paragraph pageId="4" pageNumber="5">
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<taxonomicName class="Insecta" family="Scelionidae" genus="Chromoteleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chromoteleia (Oxyscelio) connectens" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="connectens" subGenus="Oxyscelio">Chromoteleia (Oxyscelio) connectens</taxonomicName>
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Kieffer, 1910a: 313 (original description, keyed).
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</paragraph>
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<paragraph pageId="4" pageNumber="5">
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<taxonomicName class="Insecta" family="Scelionidae" genus="Oxyscelio" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oxyscelio (Oxyscelio) connectens" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="connectens" subGenus="Oxyscelio">Oxyscelio (Oxyscelio) connectens</taxonomicName>
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(Kieffer): Kieffer, 1910b: 69 (generic transfer).
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</paragraph>
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<paragraph pageId="4" pageNumber="5">
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<taxonomicName class="Insecta" family="Scelionidae" genus="Oxyscelio" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oxyscelio connectens" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="connectens">Oxyscelio connectens</taxonomicName>
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(Kieffer): Kieffer, 1926: 361, 362 (description, keyed); Dodd, 1931: 77 (excluded from
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<taxonomicName class="Insecta" family="Scelionidae" genus="Oxyscelio" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oxyscelio" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">Oxyscelio</taxonomicName>
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, position uncertain).
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</paragraph>
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<paragraph pageId="4" pageNumber="5">
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<taxonomicName class="Insecta" family="Scelionidae" genus="Chromoteleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chromoteleia connectens" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="connectens">Chromoteleia connectens</taxonomicName>
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Kieffer: Masner, 1976: 25 (type information); Hoebeke, 1980: 26 (type information); Johnson, 1992: 364 (cataloged, type information).
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</paragraph>
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<paragraph pageId="4" pageNumber="5">
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<taxonomicName class="Insecta" family="Scelionidae" genus="Chromoteleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chromoteleia brevitarsis" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="brevitarsis">Chromoteleia brevitarsis</taxonomicName>
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Kieffer, 1910a: 313, 315 (original description, keyed); Masner, 1976: 25 (type information, junior synonym of
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<taxonomicName class="Insecta" family="Scelionidae" genus="Chromoteleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chromoteleia connectens" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="connectens">Chromoteleia connectens</taxonomicName>
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Kieffer); Johnson, 1992: 364 (type information).
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</paragraph>
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<paragraph pageId="4" pageNumber="5">
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<taxonomicName class="Insecta" family="Scelionidae" genus="Petalosema" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Petalosema brevitarsis" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="brevitarsis">Petalosema brevitarsis</taxonomicName>
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(Kieffer): Kieffer, 1926: 358, 360 (generic transfer, description, keyed).
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</paragraph>
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<paragraph pageId="4" pageNumber="5">http://zoobank.org/C7398CF1-267D-4D5D-B89C-55697354254C</paragraph>
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<paragraph pageId="4" pageNumber="5">http://bioguid.osu.edu/xbiod_concepts/8526</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="5" type="description">
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<paragraph pageId="4" pageNumber="5">Description.</paragraph>
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<paragraph pageId="4" pageNumber="5">
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Body length of female: 5.38-5.95 mm (n = 20). Body length of male: 5.10-5.86 mm (n = 20). Color of A1: yellow to orange. A6 in female: as wide as long. A5 in female: distinctly longer than wide. A6 in male: approximately 2.0
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<normalizedToken originalValue="×">x</normalizedToken>
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longer than wide. Number of basiconic sensilla on A7: 0. Number of basiconic sensilla on A12: 1. Sculpture of dorsal A1: striate. Color of head: black. Sculpture of frons directly above interantennal process: transversely striate to rugose. Central keel: complete, extending from interantennal process to median ocellus. Ventral margin of clypeus: straight. Granulate microsculpture of dorsal frons: present. Occipital carina: complete. Granulate microsculpture of vertex: absent; present. Sculpture of occiput: rugose. Sculpture of gena: dorsoventrally strigose.Color of mesosoma: variably orange to black. Sculpture of epicoxal lobe posterior of propleural epicoxal sulcus: smooth. Sculpture of lateral pronotal area above pronotal cervical sulcus: smooth throughout. Sculpture of netrion: rugose. Microsculpture of mesoscutum: granulate. Macrosculpture of mesoscutal midlobe: punctate rugose throughout. Macrosculpture of lateral lobe of mesoscutum: punctate rugose. Sculpture of notaulus: foveate. Notaular foveae: interconnected. Median mesoscutal carina: present along full length of mesoscutum. Mesoscutellum in lateral view: convex. Sculpture of mesoscutellum: smooth medially, densely punctate laterally; longitudinally carinate medially, densely punctate laterally. Shape of metascutellum: trapezoidal with broad posterior margin. Median metascutellar carina: present. Sculpture of metascutellum: rugose. Sculpture of lateral propodeal area: rugose; smooth. Mesopleural carina: present. Sculpture of mesepisternum below femoral depression: rugose anteriorly, sparsely punctate posteriorly. Sculpture of dorsal metapleural area: rugose. Setation of dorsal metapleural area: absent. Setation of area directly dorsal to the metapleural triangle: absent. Sculpture of ventral metapleural area: rugose anteriorly, smooth posteriorly. Color of legs: orange yellow throughout; orange to pale brown. Length of hind basitarsus: about as long as remaining segments combined. Sculpture of hind coxa: densely punctate.
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</paragraph>
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<paragraph pageId="4" pageNumber="5">Length of postmarginal vein: distinctly longer than stigmal vein.</paragraph>
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<paragraph pageId="4" pageNumber="5">Color of metasoma in female: black; mostly black with T1-T3 orange to dark brown. Color of metasoma in male: black. Horn on T1 in female: present. Striae of posterior margin of T1 in female: sparse. Striae of T1 in male: sparse. Transverse sulcus on T2: present. Sculpture of T2: densely longitudinally striate, punctate rugulose in interstices. Sculpture of T6 in female: densely punctate and granulate. Length of T6 in female: approximately as long as wide. Shape of T6 in female in lateral view: flat. Apical spine on female T6: absent. Sculpture of T6 in male: densely punctate. Sculpture of T7 in male: granulate. Posterior margin of T7 in male: emarginate medially between rounded projections. Sculpture of medial S2: densely longitudinally striate with fine punctures in interstices.</paragraph>
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<caption pageId="4" pageNumber="5">
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<paragraph pageId="4" pageNumber="5">
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Figures 58-63.
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<taxonomicName class="Insecta" family="Scelionidae" genus="Chromoteleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chromoteleia connectens" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="connectens">Chromoteleia connectens</taxonomicName>
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Kieffer, female, holotype (CAS TYPE9618). 58 Lateral habitus 59 Head and mesosoma, lateral view 60 Dorsal habitus 61 Head and mesosoma, dorsal view 62 Head, anterior view 63 Metasoma, dorsal view.
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</paragraph>
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</caption>
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<caption pageId="4" pageNumber="5">
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<paragraph pageId="4" pageNumber="5">
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Figures 64-69.
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<taxonomicName class="Insecta" family="Scelionidae" genus="Chromoteleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chromoteleia brevitarsis" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="brevitarsis">Chromoteleia brevitarsis</taxonomicName>
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Kieffer, female, holotype (CAS TYPE9597). 64 Lateral habitus 65 Head and mesosoma, lateral view 66 Dorsal habitus 67 Head and mesosoma, dorsal view 68 Head, anterior view 69 Metasoma, dorsal view.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="5" type="diagnosis">
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<paragraph pageId="4" pageNumber="5">Diagnosis.</paragraph>
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<paragraph pageId="4" pageNumber="5">
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This species can be distinguished from other
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<taxonomicName class="Insecta" family="Scelionidae" genus="Chromoteleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chromoteleia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">Chromoteleia</taxonomicName>
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by the following combination of characters: female A7 without basiconic sensillum, occipital carina complete, median mesoscutal carina present along full length of mesoscutum, ventral metapleural area smooth posteriorly, postmarginal vein distinctly longer than stigma vein, male T7 with posterior margin deeply emarginated between rounded projections.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="5" type="distribution">
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<paragraph pageId="4" pageNumber="5">Link to distribution map.</paragraph>
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<paragraph pageId="4" pageNumber="5">[http://hol.osu.edu/map-large.html?id=4212]</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="5" type="materials_examined">
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<paragraph pageId="4" pageNumber="5">Material examined.</paragraph>
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<paragraph pageId="4" pageNumber="5">
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Holotype, female,
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<taxonomicName lsidName="C. brevitarsis" pageId="4" pageNumber="5" rank="species" species="brevitarsis">C. brevitarsis</taxonomicName>
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: BRAZIL: PA, no date, Baker, CAS TYPE9597 (deposited in CAS). Holotype, female,
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<taxonomicName lsidName="C. connectens" pageId="4" pageNumber="5" rank="species" species="connectens">C. connectens</taxonomicName>
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: BRAZIL: PA, no date, Baker, CAS TYPE9618 (deposited in CAS).
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</paragraph>
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<paragraph pageId="4" pageNumber="5">Other material: (524 females, 594 males) BELIZE: 1 female, OSUC586444 (CNCI). BOLIVIA: 5 females, 7 males, OSUC584139-584140, 584298-584300 (CNCI); DPI_FSCA 00010202-00010208 (CSCA). BRAZIL: 77 females, 73 males, OSUC202485, 202488, 202490, 202493, 202495-202497, 202499-202501, 202515-202517, 202519, 202523, 202532, 202535 (AEIC); OSUC232991 (BPBM); OSUC583467-583468, 583471, 583473-583475, 583484-583485, 583492, 583495-583496, 583498, 583510, 583516, 584146, 584152, 584161, 584164-584165, 584171-584175, 584177, 584330, 584334, 584336, 586124-586126, 586131-586132, 586134, 586352, 586356, 586358, 586365, 586368, 586394-586395, 586397-586398, 586404, 586406, 586418-586419, 586567-586568, 586796 (CNCI); OSUC199611, 199634 (FSCA); OSUC181648, 225221, 225223, 225240, 231875, 233121, 233201, 233295, 233345, 235221, 267216, 322173, 322599, 326391, 326465, 344484, 354669, 359051, 359057, 359059, 359065, 359067, 359076, 370978, 379216, 380242, 449172, 449186-449187 (MPEG); OSUC112474, 112594, 112933, 132283, 132337-132338 (MZSP); OSUC132223, 132546, 132576, 132614, 225220, 225239, 233200, 233320, 255000, 322166, 322175, 322598, 322639, 326569, 345343, 359056, 359063-359064, 359068, 370976-370977, 379214, 381994, 449168, 449178, 48530, 48535, 56248, 56250, 56257-56258, 56260, 56262 (OSUC); OSUC204987, 204989 (UCDC); OSUC225298-225300, 225302-225304, 225309, 225312, 225314-225315 (USNM). COLOMBIA: 49 females, 55 males, OSUC584137, 584301 (CNCI); OSUC143884, 143894-143895, 143897, 143899, 143901, 144022, 152042-152043, 152069, 152071, 162521, 162680-162681, 166453, 166456-166457, 166521, 166555, 170459, 176951, 182582-182583, 182713, 188629-188630, 188632-188633, 188691-188692, 188783, 189043-189044, 189046, 190347-190348, 190350, 190901, 191362, 193136, 193185, 193429, 193456-193458, 193460, 193661, 193664, 193680, 193819, 193960-193962, 228571, 228601, 228603-228605, 228607, 230379, 230434, 230436, 230439, 232724, 232726, 249925-249926, 253478, 269375, 270003, 270005-270007, 321676 (IAVH); OSUC143896, 143898, 152070, 152072, 162520, 162679, 166450, 166458, 166520, 166565, 166588, 170460, 176950, 182581, 189045, 190349, 191043, 191173, 193168, 193184, 193793, 228602, 228606, 232723, 249928, 269377, 270004, 273444 (OSUC). COSTA RICA: 145 females, 195 males, 1 unknown, OSUC202537, 202539-202540, 202549, 205189 (AEIC); OSUC232993 (BPBM); OSUC232189, 253965, 253967-253968, 253970-253971, 556950, 556971, 556973-556977, 556980, 556982-556985, 556993, 556995-556996, 556999, 557001, 557004, 557010, 557014, 557018, 557023, 557048, 557050-557051, 557053, 557058, 557084, 557087, 577888-577893, 577895-577896, 577899, 577908, 577927, 577929, 577938, 577940, 577943-577944, 577947, 577949-577951, 577954-577955, 577962, 577964, 577971, 577974-577978, 577980-577984, 577986-577987, 577989-577990, 577992-577999, 578001, 578007, 578009, 578012-578013, 578018, 578020-578022, 578025-578043, 578045, 578047-578052, 578066, 578073, 578076, 578080, 578085, 578092, 578097, 578099, 578104, 583413, 583415-583416, 583422-583423, 583461, 583479, 583482, 583508, 583511-583513, 583731, 583739, 583756, 583760, 583763-583764, 583768-583771, 583773, 583787, 583794, 583796, 583834, 583850, 583878, 583881, 583890-583892, 583896-583897, 583901, 583910, 583916, 583926-583928, 583931-583938, 583947-583948, 583978, 583980, 583982, 583988-583990, 583992, 583994, 583997-583998, 584003-584006, 584008-584009, 584012, 584014, 584016-584017, 584019, 584023, 584026-584028, 584031, 584226, 584228, 584231-584232, 584234, 584237, 584239, 584241, 584245-584246, 584248-584249, 584251, 584253-584259, 584271-584273, 584277-584280, 584284-584286, 584288-584293, 586121, 586123, 586130, 586150-586151, 586161, 586163-586164, 586166-586168, 586171, 586179-586181, 586183, 586194, 586196, 586198, 586200-586201, 586203, 586205-586206, 586209, 586212-586214, 586216, 586260-586261, 586268-586269, 586271-586274, 586293-586295, 586302, 586357, 586390, 586399, 586415, 586449, 586451, 586471, 586474-586476, 586485, 586493, 586499, 586501, 586510, 586513, 586516-586520, 586541-586543, 586559, 586619-586620, 586638-586640, 586645, 586648-586649, 586656, 586658-586659, 586679, 586681, 586764, 586778, 586781, 586783, 586785-586787, 586831, 586833, 586837, 586839, 586841 (CNCI); DPI_FSCA 00010229-00010230, SM0810076, SM0810559 (KUNH); OSUC205812 (UCDC). ECUADOR: 122 females, 166 males, OSUC149644, 202502 (AEIC); OSUC181428, 557030, 557034, 557037, 557042-557045, 557092-557094, 557096, 557098, 583476-583477, 583751, 584047-584059, 584061-584073, 584075-584082, 584084-584096, 584099-584101, 584105, 584110-584111, 584120-584121, 584123-584130, 584132, 584134-584136, 584399, 584401, 584403, 584405-584406, 584408, 584411-584413, 584415, 584427, 584430, 584432-584451, 584807-584816, 584818-584837, 584839-584856, 585089-585090, 585097, 586142-586144, 586178, 586311-586313, 586317-586319, 586335, 586359, 586413, 586546-586553, 586555-586556, 586558, 586625, 586633, 586668-586670, 586672-586676, 586693, 586696-586707, 586709-586713, 586715-586720, 586723, 586725-586726, 586728-586730, 586732-586746, 586873 (CNCI); DPI_FSCA 00010184-00010201, 00010209 (CSCA); OSUC199615-199629 (FSCA); OSUC221931, 221939 (TAMU); OSUC204993, 204995-204996, 204998 (UCDC). FRENCH GUIANA: 36 females, 28 males, OSUC555804, 555814, 555819, 586222, 586225, 586228-586230, 586232, 586234-586238, 586240-586242, 586244-586245, 586247-586255, 586257, 586423, 586427, 586434, 586436, 586439, 586443, 586446, 586453-586458, 586461-586464, 586609-586612, 586830, 586844-586845, 586850-586851, 586854-586855, 586859 (CNCI); SM0096678 (KUNH); OSUC267215, 47019-47021, 47023 (OSUC). GUATEMALA: 2 females, 1 male, OSUC203118 (AEIC); OSUC584925 (CNCI); OSUC204701 (UCDC). HONDURAS: 1 male, OSUC413764 (MZLU). PANAMA: 8 females, 8 males, OSUC556951, 584182, 584379, 586623v586624, 586626, 586629, 586824 (CNCI); DPI_FSCA 00010227 (KUNH); OSUC221921, 233027, 271017-271020, 320637 (TAMU). PERU: 24 females, 17 males, OSUC149639 (AEIC); OSUC578057, 578060, 578062, 584305, 584307-584308, 584314-584316, 586135-586137, 586185-586188, 586190-586192, 586421-586422, 586441-586442, 586805, 586808-586814, 586846, 586865 (CNCI); OSUC218803-218805 (INHS); DPI_FSCA 00010228 (KUNH); OSUC223890, 323994 (OSUC); OSUC225399 (USNM). VENEZUELA: 55 females, 43 males, OSUC149646 (AEIC); OSUC578015, 583517, 586145 (CNCI); OSUC230357-230359, 230361, 232281, 232827-232830, 251645-251647, 251649, 251651-251652, 251655, 251657-251661, 251663-251666, 251668-251670, 251672-251675, 251677-251681, 251683-251684, 251686-251688, 251690, 320743-320745, 320747-320753, 321367, 321369-321371, 321373-321374, 321376-321379, 321382, 323405, 323407-323412, 323414, 381986-381987, 381989, 381991-381992 (MIZA); OSUC230360, 232282, 251643, 251648, 251650, 251653, 251656, 251671, 251682, 251685, 251692, 321368, 321372, 321375, 321380-321381, 323406 (OSUC).</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="5" type="comments">
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<paragraph pageId="4" pageNumber="5">Comments.</paragraph>
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<paragraph pageId="4" pageNumber="5">This species exhibits variation in color and microsculpture. The development of microsculpture on vertex could be absent or present; mesoscutum smooth or carinate medially; mesoscutellum, mesopleuron, and metapleuron ranging from orange to dark brown or black.</paragraph>
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</subSubSection>
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</treatment>
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</document> |