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<document id="44ED89F6A13A8F04012F18D3EAF0B372" ID-DOI="10.5281/zenodo.183706" ID-GBIF-Dataset="e274d1d3-7bc2-46f8-a22e-e7d99d667298" ID-ISSN="1175­5326" ID-Zenodo-Dep="183706" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459929192289" checkinUser="plazi" docAuthor="Schwabe, Enrico &amp; Engl, Winfried" docDate="2008" docId="F976879AFFDFE353FF5EFAF0FE03EF8F" docLanguage="en" docName="zt01866p204.pdf" docOrigin="Zootaxa 1866" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Brookula bohni Schwabe &amp; Engl, 2008, sp. nov." docType="treatment" docVersion="11" lastPageNumber="198" masterDocId="054FFFE2FFD6E358FFC9FFA3FF89EC0A" masterDocTitle="Description of two new deep­water species of the genus Brookula Iredale, 1912 (Mollusca, Gastropoda, Trochoidea), with a revision of the genus for the Subantarctic and Arctic Sector of the Atlantic Ocean *" masterLastPageNumber="204" masterPageNumber="187" pageNumber="196" updateTime="1698229735954" updateUser="plazi">
<mods:mods id="DE90C696C7609EE3AFFA90ADDD4C4D06" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1D896951BF27633ED9504800392CA705">
<mods:title id="F5944DC1D6EDA062FBC1A14DEEE2FA36">Description of two new deep­water species of the genus Brookula Iredale, 1912 (Mollusca, Gastropoda, Trochoidea), with a revision of the genus for the Subantarctic and Arctic Sector of the Atlantic Ocean *</mods:title>
</mods:titleInfo>
<mods:name id="9024EE1FC3AB00782D0547BBA3DED5A6" type="personal">
<mods:role id="9940C82C6D9EE720506D8A52FA0479B8">
<mods:roleTerm id="CE3D9609E8DD2F5A18EB321236C959C0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="991995EC515D27BF352FA8E3CE21F861">Schwabe, Enrico</mods:namePart>
</mods:name>
<mods:name id="449DE3B404FDC389998953800C88D939" type="personal">
<mods:role id="FEA677ACA18DA0F1D89354E27AAC509A">
<mods:roleTerm id="6F1CB6243FF337E1E0D98DC89C8F904F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="23859B8FA0E13E0042F48F645D166EB2">Engl, Winfried</mods:namePart>
</mods:name>
<mods:typeOfResource id="7041796062BD0F9458E1644FCE630B98">text</mods:typeOfResource>
<mods:relatedItem id="B1CFE7D6E3A9B6C2447C8736E67DA4CB" type="host">
<mods:titleInfo id="09B2095B25D7FD2CC0631B028705A224">
<mods:title id="CBD47C4AE78FCDEE2BBE7C5A6E161F73">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D278601E239310760E9B90B9A58B00C9">
<mods:date id="D827A36D59CD0ED291E3CB92A53DEBDC">2008</mods:date>
<mods:detail id="C819683A53686C490B18BE21A3FA578B" type="volume">
<mods:number id="AF08AC6B90D0AB76C9652AEE824940BE">1866</mods:number>
</mods:detail>
<mods:extent id="97838C7E332F4536C6A3022A5145FA2F" unit="page">
<mods:start id="261266CFFE5A5C16BC686057B709FAD3">187</mods:start>
<mods:end id="3AD0DDF0035E2A1C6240A4F7598E84ED">204</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="374C39029F0B610E47A0FB2407F67E04">journal article</mods:classification>
<mods:identifier id="C8D7E905BB2ED79FD13D3A55918FD64A" type="DOI">10.5281/zenodo.183706</mods:identifier>
<mods:identifier id="40860A84AC67081C5ECBD9B319206C2C" type="GBIF-Dataset">e274d1d3-7bc2-46f8-a22e-e7d99d667298</mods:identifier>
<mods:identifier id="9398932599B917828359CCB73A3F6F12" type="ISSN">1175­5326</mods:identifier>
<mods:identifier id="18097AF3164D561487C5840485C6E28C" type="Zenodo-Dep">183706</mods:identifier>
</mods:mods>
<treatment id="F976879AFFDFE353FF5EFAF0FE03EF8F" ID-DOI="http://doi.org/10.5281/zenodo.5631753" ID-GBIF-Taxon="119374588" ID-Zenodo-Dep="5631753" LSID="urn:lsid:plazi:treatment:F976879AFFDFE353FF5EFAF0FE03EF8F" httpUri="http://treatment.plazi.org/id/F976879AFFDFE353FF5EFAF0FE03EF8F" lastPageId="11" lastPageNumber="198" pageId="9" pageNumber="196">
<subSubSection id="39C56507FFDFE351FF5EFAF0FE99E99F" pageId="9" pageNumber="196" type="nomenclature">
<paragraph id="7160368CFFDFE351FF5EFAF0FE32E966" blockId="9.[151,443,1362,1429]" box="[151,443,1362,1389]" pageId="9" pageNumber="196">
<heading id="2A2881E0FFDFE351FF5EFAF0FE32E966" bold="true" box="[151,443,1362,1389]" fontSize="11" level="1" pageId="9" pageNumber="196" reason="1">
<emphasis id="43ABEA9EFFDFE351FF5EFAF0FE32E966" bold="true" box="[151,443,1362,1389]" pageId="9" pageNumber="196">
<taxonomicName id="B6DF4D0FFFDFE351FF5EFAF0FEDFE967" ID-CoL="NDTW" box="[151,342,1363,1389]" class="Gastropoda" family="Liotiidae" genus="Brookula" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="9" pageNumber="196" phylum="Mollusca" rank="species" species="bohni" status="sp. nov.">
<emphasis id="43ABEA9EFFDFE351FF5EFAF0FEDFE967" bold="true" box="[151,342,1363,1389]" italics="true" pageId="9" pageNumber="196">Brookula bohni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="589857E5FFDFE351FE94FAF1FE32E966" box="[349,443,1362,1388]" pageId="9" pageNumber="196" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="7160368CFFDFE351FF5EFAD8FE99E99F" blockId="9.[151,443,1362,1429]" box="[151,272,1403,1429]" pageId="9" pageNumber="196">
(
<figureCitation id="E9E42A09FFDFE351FF56FAD8FE8EE99F" box="[159,263,1403,1429]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStartId-0="10.[151,244,1628,1649]" captionStartId-1="11.[151,244,1966,1987]" captionTargetBox-0="[151,1435,637,1600]" captionTargetBox-1="[151,1435,960,1939]" captionTargetId-0="figure@10.[151,1436,637,1601]" captionTargetId-1="figure@11.[151,1436,960,1939]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURE 7: A – D, Holotype of Brookula bohni sp. nov. (ZSM Mol 20070895) Antarctica, South Shetlands, St. PS 61 / 043 – 08 (60 ° 27.12 ’ S 56 ° 05.10 ’ W to 60 ° 27.18 ’ S 56 ° 04.80 ’ W), 3959 – 3961 m. A, Ventral view of the complete shell. B, View of the shell base showing the narrow opening of the umbilicus. C, Apical view to show the protoconch and the teleoconch whorls. D, Detail of fig. C to show the fine protoconch ornamentation. Scale bars: 100 µm." captionText-1="FIGURE 8: The geographic distribution of Brookula bohni sp. nov. with a 2500 m depth contour in light grey." httpUri-0="https://zenodo.org/record/183711/files/figure.png" httpUri-1="https://zenodo.org/record/183712/files/figure.png" pageId="9" pageNumber="196">Figs 7–8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="39C56507FFDFE351FF5EFA68FDC1EAF7" pageId="9" pageNumber="196" type="materials_examined">
<paragraph id="7160368CFFDFE351FF5EFA68FEB1E9EE" blockId="9.[151,1436,1483,2029]" box="[151,312,1483,1508]" pageId="9" pageNumber="196">
<emphasis id="43ABEA9EFFDFE351FF5EFA68FEB1E9EE" box="[151,312,1483,1508]" italics="true" pageId="9" pageNumber="196">
<typeStatus id="AE64882EFFDFE351FF5EFA68FF47E9EE" box="[151,206,1483,1508]" pageId="9" pageNumber="196">Type</typeStatus>
material
</emphasis>
</paragraph>
<paragraph id="7160368CFFDFE351FF0FFA51FAA2EA07" blockId="9.[151,1436,1483,2029]" box="[198,1323,1522,1549]" pageId="9" pageNumber="196">
<typeStatus id="AE64882EFFDFE351FF0FFA51FEB1EA06" box="[198,312,1522,1548]" pageId="9" pageNumber="196" type="holotype">
<emphasis id="43ABEA9EFFDFE351FF0FFA51FEB1EA06" bold="true" box="[198,312,1522,1548]" pageId="9" pageNumber="196">Holotype</emphasis>
</typeStatus>
(
<collectionCode id="17CEAE49FFDFE351FE8EFA50FE0CEA07" box="[327,389,1523,1549]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="9" pageNumber="196">ZSM</collectionCode>
Mol 20070895), leg. Katrin Linse,
<date id="0561104CFFDFE351FCEFFA50FC63EA07" box="[806,1002,1523,1549]" pageId="9" pageNumber="196" value="2002-02-04">February 4 2002</date>
, ANT XIX/3 (ANDEEP I).
</paragraph>
<paragraph id="7160368CFFDFE351FF0FF9B9FD3FEAA7" blockId="9.[151,1436,1483,2029]" pageId="9" pageNumber="196">
<typeStatus id="AE64882EFFDFE351FF0FF9B9FECAEA3E" box="[198,323,1562,1588]" pageId="9" pageNumber="196" type="paratype">
<emphasis id="43ABEA9EFFDFE351FF0FF9B9FECAEA3E" bold="true" box="[198,323,1562,1588]" pageId="9" pageNumber="196">Paratypes</emphasis>
</typeStatus>
, one from
<typeStatus id="AE64882EFFDFE351FE00F9B8FE72EA3F" box="[457,507,1563,1589]" pageId="9" pageNumber="196">type</typeStatus>
locality (
<collectionCode id="17CEAE49FFDFE351FDA4F9B8FD22EA3F" box="[621,683,1563,1589]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="9" pageNumber="196">ZSM</collectionCode>
Mol
<date id="0561104CFFDFE351FD26F9B8FCE0EA3F" box="[751,873,1563,1589]" pageId="9" pageNumber="196" value="1185-02-20">20021185</date>
). Two
<typeStatus id="AE64882EFFDFE351FC70F9B8FBA0EA3F" box="[953,1065,1563,1589]" pageId="9" pageNumber="196" type="paratype">paratypes</typeStatus>
(
<collectionCode id="17CEAE49FFDFE351FBF2F9B8FBF0EA3F" box="[1083,1145,1563,1589]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="9" pageNumber="196">ZSM</collectionCode>
Mol
<date id="0561104CFFDFE351FB75F9B8FABBEA3F" box="[1212,1330,1563,1589]" pageId="9" pageNumber="196" value="1177-02-20">20021177</date>
)
<taxonomicName id="B6DF4D0FFFDFE351FA8AF9B8FF4DEA57" class="Gastropoda" family="Liotiidae" genus="Brookula" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="9" pageNumber="201" phylum="Mollusca" rank="species" species="antarctica">
<collectingCountry id="09C8761CFFDFE351FA8AF9B8FF4DEA57" name="Antarctica" pageId="9" pageNumber="196">Antarctica</collectingCountry>
</taxonomicName>
, Drake Passage, St. PS 61
<emphasis id="43ABEA9EFFDFE351FE37F9E1FD8FEA56" bold="true" box="[510,518,1602,1628]" pageId="9" pageNumber="196">/</emphasis>
042–02 (
<geoCoordinate id="14EB504BFFDFE351FDB9F9E0FD7FEA57" box="[624,758,1603,1629]" direction="south" orientation="latitude" pageId="9" pageNumber="196" precision="9" value="-59.6715">59°40.29’S</geoCoordinate>
<geoCoordinate id="14EB504BFFDFE351FD36F9E0FC06EA57" box="[767,911,1603,1629]" direction="west" orientation="longitude" pageId="9" pageNumber="196" precision="9" value="-57.5905">57°35.43’W</geoCoordinate>
to
<geoCoordinate id="14EB504BFFDFE351FC71F9E0FBB4EA57" box="[952,1085,1603,1629]" direction="south" orientation="latitude" pageId="9" pageNumber="196" precision="9" value="-59.673668">59°40.42’S</geoCoordinate>
<geoCoordinate id="14EB504BFFDFE351FB8FF9E0FB52EA57" box="[1094,1243,1603,1629]" direction="west" orientation="longitude" pageId="9" pageNumber="196" precision="9" value="-57.587833">57°35.27’W</geoCoordinate>
),
<quantity id="B6279B69FFDFE351FB38F9E0FA11EA57" box="[1265,1432,1603,1629]" metricMagnitude="3" metricUnit="m" metricValue="3.6814999999999998" metricValueMax="3.683" metricValueMin="3.6799999999999997" pageId="9" pageNumber="196" unit="m" value="3681.5" valueMax="3683.0" valueMin="3680.0">3680–3683 m</quantity>
, Katrin Linse,
<date id="0561104CFFDFE351FEF1F9C8FE75EA8F" box="[312,508,1643,1669]" pageId="9" pageNumber="196" value="2002-01-27">January 27 2002</date>
, ANT XIX/3 (ANDEEP I). Besides the SEM­mounted
<typeStatus id="AE64882EFFDFE351FB4BF9C8FB62EA8F" box="[1154,1259,1643,1669]" pageId="9" pageNumber="196" type="holotype">holotype</typeStatus>
, all other specimens are fixed and preserved in 96% ethanol.
</paragraph>
<paragraph id="7160368CFFDFE351FF0FF918FDC1EAF7" blockId="9.[151,1436,1483,2029]" pageId="9" pageNumber="196">
<emphasis id="43ABEA9EFFDFE351FF0FF918FEE8EADE" box="[198,353,1723,1748]" italics="true" pageId="9" pageNumber="196">
<typeStatus id="AE64882EFFDFE351FF0FF918FF74EADE" box="[198,253,1723,1748]" pageId="9" pageNumber="196">Type</typeStatus>
locality
</emphasis>
:
<taxonomicName id="B6DF4D0FFFDFE351FEBFF918FE7AEADF" box="[374,499,1723,1749]" class="Gastropoda" family="Liotiidae" genus="Brookula" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="9" pageNumber="201" phylum="Mollusca" rank="species" species="antarctica">
<collectingCountry id="09C8761CFFDFE351FEBFF918FE7AEADF" box="[374,499,1723,1749]" name="Antarctica" pageId="9" pageNumber="196">Antarctica</collectingCountry>
</taxonomicName>
, South Shetlands, St. PS 61/043–08 (
<geoCoordinate id="14EB504BFFDFE351FC0CF918FBC2EADF" box="[965,1099,1723,1749]" direction="south" orientation="latitude" pageId="9" pageNumber="196" precision="9" value="-60.452">60°27.12’S</geoCoordinate>
<geoCoordinate id="14EB504BFFDFE351FB9EF918FB61EADF" box="[1111,1256,1723,1749]" direction="west" orientation="longitude" pageId="9" pageNumber="196" precision="9" value="-56.085">56°05.10’W</geoCoordinate>
to
<geoCoordinate id="14EB504BFFDFE351FADEF918FA15EADF" box="[1303,1436,1723,1749]" direction="south" orientation="latitude" pageId="9" pageNumber="196" precision="9" value="-60.453">60°27.18’S</geoCoordinate>
<geoCoordinate id="14EB504BFFDFE351FF5EF940FEA5EAF7" box="[151,300,1763,1789]" direction="west" orientation="longitude" pageId="9" pageNumber="196" precision="9" value="-56.08">56°04.80’W</geoCoordinate>
),
<quantity id="B6279B69FFDFE351FE89F940FE6BEAF7" box="[320,482,1763,1789]" metricMagnitude="3" metricUnit="m" metricValue="3.96" metricValueMax="3.961" metricValueMin="3.9589999999999996" pageId="9" pageNumber="196" unit="m" value="3960.0" valueMax="3961.0" valueMin="3959.0">3959–3961 m</quantity>
(
<figureCitation id="E9E42A09FFDFE351FE3BF940FDB1EAF7" box="[498,568,1763,1789]" captionStart="FIGURE 8" captionStartId="11.[151,244,1966,1987]" captionTargetBox="[151,1435,960,1939]" captionTargetId="figure@11.[151,1436,960,1939]" captionTargetPageId="11" captionText="FIGURE 8: The geographic distribution of Brookula bohni sp. nov. with a 2500 m depth contour in light grey." httpUri="https://zenodo.org/record/183712/files/figure.png" pageId="9" pageNumber="196">Fig. 8</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="39C56507FFDFE351FF0FF8A8FA9BEB47" pageId="9" pageNumber="196" type="etymology">
<paragraph id="7160368CFFDFE351FF0FF8A8FA9BEB47" blockId="9.[151,1436,1483,2029]" pageId="9" pageNumber="196">
<emphasis id="43ABEA9EFFDFE351FF0FF8A8FECAEB2E" box="[198,323,1803,1828]" italics="true" pageId="9" pageNumber="196">Etymology</emphasis>
: The species is named after our friend Jens Michael Bohn (ZSM), a holothurian specialist and participant of the relevant expeditions. We selected his surname because his nickname is “
<taxonomicName id="B6DF4D0FFFDFE351FB7AF890FA8BEB47" box="[1203,1282,1843,1869]" class="Gastropoda" family="Liotiidae" genus="Brookula" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="9" pageNumber="202" phylum="Mollusca" rank="species" species="bohni">Bohni</taxonomicName>
”.
</paragraph>
</subSubSection>
<subSubSection id="39C56507FFDFE353FF0FF8F8FDBFEE9F" lastPageId="11" lastPageNumber="198" pageId="9" pageNumber="196" type="description">
<paragraph id="7160368CFFDFE352FF0FF8F8FBA2EE10" blockId="9.[151,1436,1483,2029]" lastBlockId="10.[151,1438,152,538]" lastPageId="10" lastPageNumber="197" pageId="9" pageNumber="196">
<emphasis id="43ABEA9EFFDFE351FF0FF8F8FED8EB7E" box="[198,337,1883,1908]" italics="true" pageId="9" pageNumber="196">Description</emphasis>
: Shell minute, up to
<quantity id="B6279B69FFDFE351FD91F8F8FD4DEB7F" box="[600,708,1883,1909]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="9" pageNumber="196" unit="mm" value="1.3">1.30 mm</quantity>
long,
<quantity id="B6279B69FFDFE351FCDDF8F8FC08EB7F" box="[788,897,1883,1909]" metricMagnitude="-3" metricUnit="m" metricValue="1.28" pageId="9" pageNumber="196" unit="mm" value="1.28">1.28 mm</quantity>
wide (
<figureCitation id="E9E42A09FFDFE351FC1EF8F8FBB0EB7F" box="[983,1081,1883,1909]" captionStart="FIGURE 7" captionStartId="10.[151,244,1628,1649]" captionTargetBox="[151,1435,637,1600]" captionTargetId="figure@10.[151,1436,637,1601]" captionTargetPageId="10" captionText="FIGURE 7: A – D, Holotype of Brookula bohni sp. nov. (ZSM Mol 20070895) Antarctica, South Shetlands, St. PS 61 / 043 – 08 (60 ° 27.12 ’ S 56 ° 05.10 ’ W to 60 ° 27.18 ’ S 56 ° 04.80 ’ W), 3959 – 3961 m. A, Ventral view of the complete shell. B, View of the shell base showing the narrow opening of the umbilicus. C, Apical view to show the protoconch and the teleoconch whorls. D, Detail of fig. C to show the fine protoconch ornamentation. Scale bars: 100 µm." httpUri="https://zenodo.org/record/183711/files/figure.png" pageId="9" pageNumber="196">Fig. 7A</figureCitation>
), globosely turbinate, fragile, translucent white in color, with inflated whorls slightly depressed at the shoulder, slightly expanded laterally, low spired, spire angle ranging from 80° (smallest
<typeStatus id="AE64882EFFDFE351FD38F808FCD1EBCF" box="[753,856,1963,1989]" pageId="9" pageNumber="196" type="paratype">paratype</typeStatus>
) to 85° (
<typeStatus id="AE64882EFFDFE351FC09F808FBA4EBCF" box="[960,1069,1963,1989]" pageId="9" pageNumber="196" type="holotype">holotype</typeStatus>
). Protoconch (
<figureCitation id="E9E42A09FFDFE351FB11F808FABDEBCF" box="[1240,1332,1963,1989]" captionStart="FIGURE 7" captionStartId="10.[151,244,1628,1649]" captionTargetBox="[151,1435,637,1600]" captionTargetId="figure@10.[151,1436,637,1601]" captionTargetPageId="10" captionText="FIGURE 7: A – D, Holotype of Brookula bohni sp. nov. (ZSM Mol 20070895) Antarctica, South Shetlands, St. PS 61 / 043 – 08 (60 ° 27.12 ’ S 56 ° 05.10 ’ W to 60 ° 27.18 ’ S 56 ° 04.80 ’ W), 3959 – 3961 m. A, Ventral view of the complete shell. B, View of the shell base showing the narrow opening of the umbilicus. C, Apical view to show the protoconch and the teleoconch whorls. D, Detail of fig. C to show the fine protoconch ornamentation. Scale bars: 100 µm." httpUri="https://zenodo.org/record/183711/files/figure.png" pageId="9" pageNumber="196">Fig. 7D</figureCitation>
) of 1.25 whorls, 280 µm in diameter, 100 µm high, globose, sculptured with fine anastomosing threads, forming microscopic pits of irregular arrangment. Teleoconch (
<figureCitation id="E9E42A09FFDCE352FCD4FF3BFC2BECB8" box="[797,930,152,178]" captionStart="FIGURE 7" captionStartId="10.[151,244,1628,1649]" captionTargetBox="[151,1435,637,1600]" captionTargetId="figure@10.[151,1436,637,1601]" captionTargetPageId="10" captionText="FIGURE 7: A – D, Holotype of Brookula bohni sp. nov. (ZSM Mol 20070895) Antarctica, South Shetlands, St. PS 61 / 043 – 08 (60 ° 27.12 ’ S 56 ° 05.10 ’ W to 60 ° 27.18 ’ S 56 ° 04.80 ’ W), 3959 – 3961 m. A, Ventral view of the complete shell. B, View of the shell base showing the narrow opening of the umbilicus. C, Apical view to show the protoconch and the teleoconch whorls. D, Detail of fig. C to show the fine protoconch ornamentation. Scale bars: 100 µm." httpUri="https://zenodo.org/record/183711/files/figure.png" pageId="10" pageNumber="197">Figs 7A, C</figureCitation>
) of up to 2.75 whorls with convex profile. Suture incised. Last adult whorl globose, slightly depressed at shoulder. Aperture oval, peritreme holostomate, although very weak at periphery of preceeding whorl; with thick inner, thin outer lip. Shell base roundly inflated. Umbilicus (
<figureCitation id="E9E42A09FFDCE352FE59FEB3FE64ED20" box="[400,493,272,298]" captionStart="FIGURE 7" captionStartId="10.[151,244,1628,1649]" captionTargetBox="[151,1435,637,1600]" captionTargetId="figure@10.[151,1436,637,1601]" captionTargetPageId="10" captionText="FIGURE 7: A – D, Holotype of Brookula bohni sp. nov. (ZSM Mol 20070895) Antarctica, South Shetlands, St. PS 61 / 043 – 08 (60 ° 27.12 ’ S 56 ° 05.10 ’ W to 60 ° 27.18 ’ S 56 ° 04.80 ’ W), 3959 – 3961 m. A, Ventral view of the complete shell. B, View of the shell base showing the narrow opening of the umbilicus. C, Apical view to show the protoconch and the teleoconch whorls. D, Detail of fig. C to show the fine protoconch ornamentation. Scale bars: 100 µm." httpUri="https://zenodo.org/record/183711/files/figure.png" pageId="10" pageNumber="197">Fig. 7B</figureCitation>
) tightly open, circular, deep. Axial sculpture of narrow ribs, ranging from 29 (largest
<typeStatus id="AE64882EFFDCE352FF3EFE9BFED7ED58" box="[247,350,312,338]" pageId="10" pageNumber="197" type="paratype">paratype</typeStatus>
) to 31 on last whorl in
<typeStatus id="AE64882EFFDCE352FDB0FE9BFD6AED58" box="[633,739,312,338]" pageId="10" pageNumber="197" type="holotype">holotype</typeStatus>
, 25 on penultimate whorl. Ribs regularly spaced, rounded, low, interspaces about three times rib width. Spiral sculpture of numerous flat threads, seven on first teleoconch whorl, about 12 on last adult whorl, narrower than axial ribs, crowded in upper and lower half of whorl, centrally absent (in the two small
<typeStatus id="AE64882EFFDCE352FDE3FE13FD15EDC0" box="[554,668,432,458]" pageId="10" pageNumber="197" type="paratype">paratypes</typeStatus>
) or interrupted by distinctly separated slightly higher spirals, not overlaying the axial ribs distally. Nine flat spirals, two spiral cords (with inner being largest), which surround umbilicus, at shell base. Operculum circular, multispiral, with centric nucleus.
</paragraph>
<caption id="25A06604FFDCE352FF5EF9FFFD6CEAD6" httpUri="https://zenodo.org/record/183711/files/figure.png" pageId="10" pageNumber="197" targetBox="[151,1435,637,1600]" targetPageId="10">
<paragraph id="7160368CFFDCE352FF5EF9FFFD6CEAD6" blockId="10.[151,1436,1628,1756]" pageId="10" pageNumber="197">
<emphasis id="43ABEA9EFFDCE352FF5EF9FFFEC5EA7B" bold="true" box="[151,332,1628,1649]" pageId="10" pageNumber="197">FIGURE 7: A–D</emphasis>
, Holotype of
<taxonomicName id="B6DF4D0FFFDCE352FE16F9FFFDF3EA7B" box="[479,634,1628,1649]" class="Gastropoda" family="Liotiidae" genus="Brookula" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="10" pageNumber="197" phylum="Mollusca" rank="species" species="bohni" status="sp. nov.">
<emphasis id="43ABEA9EFFDCE352FE16F9FFFDF3EA7B" box="[479,634,1628,1649]" italics="true" pageId="10" pageNumber="197">Brookula bohni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="589857E5FFDCE352FD4DF9FFFD5AEA7B" box="[644,723,1628,1649]" pageId="10" pageNumber="197" rank="species">sp. nov.</taxonomicNameLabel>
(ZSM Mol 20070895)
<taxonomicName id="B6DF4D0FFFDCE352FC0FF9FFFBA5EA7B" box="[966,1068,1628,1649]" class="Gastropoda" family="Liotiidae" genus="Brookula" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="10" pageNumber="201" phylum="Mollusca" rank="species" species="antarctica">Antarctica</taxonomicName>
, South Shetlands, St. PS 61/043–08 (60°27.12’S 56°05.10’W to 60°27.18’S 56°04.80’W), 3959–3961 m.
<emphasis id="43ABEA9EFFDCE352FCFFF9DDFCCEEA99" bold="true" box="[822,839,1662,1683]" pageId="10" pageNumber="197">A</emphasis>
, Ventral view of the complete shell.
<emphasis id="43ABEA9EFFDCE352FB63F9DDFB33EA99" bold="true" box="[1194,1210,1662,1683]" pageId="10" pageNumber="197">B</emphasis>
, View of the shell base showing the narrow opening of the umbilicus.
<emphasis id="43ABEA9EFFDCE352FD9DF902FDECEABC" bold="true" box="[596,613,1697,1718]" pageId="10" pageNumber="197">C</emphasis>
, Apical view to show the protoconch and the teleoconch whorls.
<emphasis id="43ABEA9EFFDCE352FB1FF902FB6EEABC" bold="true" box="[1238,1255,1697,1718]" pageId="10" pageNumber="197">D</emphasis>
, Detail of fig. C to show the fine protoconch ornamentation. Scale bars: 100 µm.
</paragraph>
</caption>
<paragraph id="7160368CFFDCE352FF0FF89FFE5CEBC4" blockId="10.[151,1437,1852,1998]" pageId="10" pageNumber="197">
Comparisons:
<taxonomicName id="B6DF4D0FFFDCE352FEB8F89FFC6FEB5C" authority="Absalao, Miyaji &amp; Pimenta, 2001" authorityName="Absalao, Miyaji &amp; Pimenta" authorityYear="2001" box="[369,998,1852,1878]" class="Gastropoda" family="Liotiidae" genus="Brookula" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="10" pageNumber="197" phylum="Mollusca" rank="species" species="spinulata">
<emphasis id="43ABEA9EFFDCE352FEB8F89FFDD8EB5F" box="[369,593,1852,1877]" italics="true" pageId="10" pageNumber="197">Brookula spinulata</emphasis>
<bibRefCitation id="154E4B7DFFDCE352FD90F89FFC6FEB5C" author="Absalao" box="[601,998,1852,1878]" pageId="10" pageNumber="203" refString="Absalao, R. S., Miyaji, C. &amp; Pimenta, A. D. (2001) The genus Brookula Iredale, 1912 (Gastropoda, Trochidae) from Brazil: description of a new species, with notes on other South American species. Zoosystema, 23 (4), 675 - 687." type="journal article" year="2001">Absalão, Miyaji &amp; Pimenta, 2001</bibRefCitation>
</taxonomicName>
differs in 1) its coarse sculpture with spinulate projections at the axial ribs, 2) the presence of fine axial lines between the spiral threads, 3) the lower number (6 vs 12) of spirals on the last body whorl, despite the general larger size, 4) the lower number (20 vs 29–31) of axial ribs.
</paragraph>
<caption id="25A06604FFDDE353FF5EFF34FCB4ECA5" ID-Table-UUID="25A06604FFDDE353FF5EFF34FCB4ECA5" box="[151,829,151,175]" httpUri="http://table.plazi.org/id/25A06604FFDDE353FF5EFF34FCB4ECA5" pageId="11" pageNumber="198" targetBox="[156,1407,213,419]" targetIsTable="true" targetPageId="11">
<paragraph id="7160368CFFDDE353FF5EFF34FCB4ECA5" blockId="11.[151,829,151,175]" box="[151,829,151,175]" pageId="11" pageNumber="198">
<emphasis id="43ABEA9EFFDDE353FF5EFF34FE85ECA5" bold="true" box="[151,268,151,175]" pageId="11" pageNumber="198">TABLE 2:</emphasis>
Shell morphometric data of
<taxonomicName id="B6DF4D0FFFDDE353FDF7FF3BFD6DECA5" box="[574,740,152,175]" class="Gastropoda" family="Liotiidae" genus="Brookula" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="11" pageNumber="198" phylum="Mollusca" rank="species" species="bohni" status="sp. nov.">
<emphasis id="43ABEA9EFFDDE353FDF7FF3BFD6DECA5" box="[574,740,152,175]" italics="true" pageId="11" pageNumber="198">Brookula bohni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="589857E5FFDDE353FD23FF3BFCB4ECA5" box="[746,829,152,175]" pageId="11" pageNumber="198" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="7160368CFFDDE353FF55FF76FADFEDA9" pageId="11" pageNumber="198">
<table id="03DFC42CFFDD1CA7FF55FF76FAF6EDA9" box="[156,1407,213,419]" gridcols="6" gridrows="5" pageId="11" pageNumber="198">
<tr id="CFEF34CEFFDD1CA7FF55FF76FAF6ECE6" box="[156,1407,213,236]" gridrow="0" pageId="11" pageNumber="198">
<th id="8C3E5DB2FFDD1CA7FF55FF76FE8CECE6" box="[156,261,213,236]" gridcol="0" gridrow="0" pageId="11" pageNumber="198">Specimen</th>
<th id="8C3E5DB2FFDD1CA7FE47FF76FE34ECE6" box="[398,445,213,236]" gridcol="1" gridrow="0" pageId="11" pageNumber="198">SL</th>
<th id="8C3E5DB2FFDD1CA7FD8FFF76FDFFECE6" box="[582,630,213,236]" gridcol="2" gridrow="0" pageId="11" pageNumber="198">SW</th>
<th id="8C3E5DB2FFDD1CA7FD1AFF76FB95ECE6" box="[723,1052,213,236]" gridcol="3" gridrow="0" pageId="11" pageNumber="198">SL/SW ratio Spire angle (°)</th>
<th id="8C3E5DB2FFDD1CA7FBA7FF76FB14ECE6" box="[1134,1181,213,236]" gridcol="4" gridrow="0" pageId="11" pageNumber="198">AL</th>
<th id="8C3E5DB2FFDD1CA7FB34FF76FAF6ECE6" box="[1277,1407,213,236]" gridcol="5" gridrow="0" pageId="11" pageNumber="198">SL/AL ratio</th>
</tr>
<tr id="CFEF34CEFFDD1CA7FF55FEA0FAF6ED10" box="[156,1407,259,282]" gridrow="1" pageId="11" pageNumber="198">
<th id="8C3E5DB2FFDD1CA7FF55FEA0FE8CED10" box="[156,261,259,282]" gridcol="0" gridrow="1" pageId="11" pageNumber="198">holotype</th>
<td id="8C3E5DB2FFDD1CA7FE47FEA0FE34ED10" box="[398,445,259,282]" gridcol="1" gridrow="1" pageId="11" pageNumber="198">1.30</td>
<td id="8C3E5DB2FFDD1CA7FD8FFEA0FDFFED10" box="[582,630,259,282]" gridcol="2" gridrow="1" pageId="11" pageNumber="198">1.28</td>
<td id="8C3E5DB2FFDD1CA7FD1AFEA0FB95ED10" box="[723,1052,259,282]" gridcol="3" gridrow="1" pageId="11" pageNumber="198">1.02 85</td>
<td id="8C3E5DB2FFDD1CA7FBA7FEA0FB14ED10" box="[1134,1181,259,282]" gridcol="4" gridrow="1" pageId="11" pageNumber="198">0.73</td>
<td id="8C3E5DB2FFDD1CA7FB34FEA0FAF6ED10" box="[1277,1407,259,282]" gridcol="5" gridrow="1" pageId="11" pageNumber="198">1.78</td>
</tr>
<tr id="CFEF34CEFFDD1CA7FF55FE92FAF6ED42" box="[156,1407,305,328]" gridrow="2" pageId="11" pageNumber="198">
<th id="8C3E5DB2FFDD1CA7FF55FE92FE8CED42" box="[156,261,305,328]" gridcol="0" gridrow="2" pageId="11" pageNumber="198">paratype</th>
<td id="8C3E5DB2FFDD1CA7FE47FE92FE34ED42" box="[398,445,305,328]" gridcol="1" gridrow="2" pageId="11" pageNumber="198">1.30</td>
<td id="8C3E5DB2FFDD1CA7FD8FFE92FDFFED42" box="[582,630,305,328]" gridcol="2" gridrow="2" pageId="11" pageNumber="198">1.25</td>
<td id="8C3E5DB2FFDD1CA7FD1AFE92FB95ED42" box="[723,1052,305,328]" gridcol="3" gridrow="2" pageId="11" pageNumber="198">1.04 81</td>
<td id="8C3E5DB2FFDD1CA7FBA7FE92FB14ED42" box="[1134,1181,305,328]" gridcol="4" gridrow="2" pageId="11" pageNumber="198">0.74</td>
<td id="8C3E5DB2FFDD1CA7FB34FE92FAF6ED42" box="[1277,1407,305,328]" gridcol="5" gridrow="2" pageId="11" pageNumber="198">1.76</td>
</tr>
<tr id="CFEF34CEFFDD1CA7FF55FEFDFAF6ED7F" box="[156,1407,350,373]" gridrow="3" pageId="11" pageNumber="198">
<th id="8C3E5DB2FFDD1CA7FF55FEFDFE8CED7F" box="[156,261,350,373]" gridcol="0" gridrow="3" pageId="11" pageNumber="198">paratype</th>
<td id="8C3E5DB2FFDD1CA7FE47FEFDFE34ED7F" box="[398,445,350,373]" gridcol="1" gridrow="3" pageId="11" pageNumber="198">1.00</td>
<td id="8C3E5DB2FFDD1CA7FD8FFEFDFDFFED7F" box="[582,630,350,373]" gridcol="2" gridrow="3" pageId="11" pageNumber="198">0.90</td>
<td id="8C3E5DB2FFDD1CA7FD1AFEFDFB95ED7F" box="[723,1052,350,373]" gridcol="3" gridrow="3" pageId="11" pageNumber="198">1.11 84</td>
<td id="8C3E5DB2FFDD1CA7FBA7FEFDFB14ED7F" box="[1134,1181,350,373]" gridcol="4" gridrow="3" pageId="11" pageNumber="198">0.48</td>
<td id="8C3E5DB2FFDD1CA7FB34FEFDFAF6ED7F" box="[1277,1407,350,373]" gridcol="5" gridrow="3" pageId="11" pageNumber="198">2.08</td>
</tr>
<tr id="CFEF34CEFFDD1CA7FF55FE2FFAF6EDA9" box="[156,1407,396,419]" gridrow="4" pageId="11" pageNumber="198">
<th id="8C3E5DB2FFDD1CA7FF55FE2FFE8CEDA9" box="[156,261,396,419]" gridcol="0" gridrow="4" pageId="11" pageNumber="198">paratype</th>
<td id="8C3E5DB2FFDD1CA7FE47FE2FFE34EDA9" box="[398,445,396,419]" gridcol="1" gridrow="4" pageId="11" pageNumber="198">0.95</td>
<td id="8C3E5DB2FFDD1CA7FD8FFE2FFDFFEDA9" box="[582,630,396,419]" gridcol="2" gridrow="4" pageId="11" pageNumber="198">1.05</td>
<td id="8C3E5DB2FFDD1CA7FD1AFE2FFB95EDA9" box="[723,1052,396,419]" gridcol="3" gridrow="4" pageId="11" pageNumber="198">0.91 80</td>
<td id="8C3E5DB2FFDD1CA7FBA7FE2FFB14EDA9" box="[1134,1181,396,419]" gridcol="4" gridrow="4" pageId="11" pageNumber="198">0.63</td>
<td id="8C3E5DB2FFDD1CA7FB34FE2FFAF6EDA9" box="[1277,1407,396,419]" gridcol="5" gridrow="4" pageId="11" pageNumber="198">1.51</td>
</tr>
</table>
</paragraph>
<paragraph id="7160368CFFDDE353FF0FFDA0FDBFEE9F" blockId="11.[151,1437,515,901]" pageId="11" pageNumber="198">
<taxonomicName id="B6DF4D0FFFDDE353FF0FFDA0FD45EE17" authority="Melvill &amp; Standen, 1912" authorityName="Melvill &amp; Standen" authorityYear="1912" box="[198,716,515,541]" class="Gastropoda" family="Liotiidae" genus="Brookula" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="11" pageNumber="198" phylum="Mollusca" rank="species" species="calypso">
<emphasis id="43ABEA9EFFDDE353FF0FFDA0FE1AEE16" box="[198,403,515,540]" italics="true" pageId="11" pageNumber="198">Brookula calypso</emphasis>
(Melvill &amp; Standen, 1912)
</taxonomicName>
differs in 1) having cancellate sculpture, built up by regularly arranged axial ribs and spiral threads, 2) the presence of small axial threads between the axial ribs, 3) having a slightly larger protoconch (295 vs 280 µm) although it has only one whorl, 4) having a larger spire angle (90 vs 85°), 5) having a shallow suture.
</paragraph>
</subSubSection>
<subSubSection id="39C56507FFDDE353FF0FFD00FE03EF8F" pageId="11" pageNumber="198" type="reference_group">
<paragraph id="7160368CFFDDE353FF0FFD00FDFDEF3F" blockId="11.[151,1437,515,901]" pageId="11" pageNumber="198">
<taxonomicName id="B6DF4D0FFFDDE353FF0FFD00FDBDEEB7" ID-CoL="NDV6" authority="Clarke, 1961" authorityName="Clarke" authorityYear="1961" box="[198,564,675,701]" class="Gastropoda" family="Liotiidae" genus="Brookula" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="11" pageNumber="198" phylum="Mollusca" rank="species" species="powelli">
<emphasis id="43ABEA9EFFDDE353FF0FFD00FE1BEEB6" box="[198,402,675,700]" italics="true" pageId="11" pageNumber="198">Brookula powelli</emphasis>
<bibRefCitation id="154E4B7DFFDDE353FE52FD00FDBDEEB7" author="Clarke" box="[411,564,675,701]" pageId="11" pageNumber="203" refString="Clarke, A. H. (1961) Abyssal mollusks from the South Atlantic Ocean. Bulletin of the Museum of Comparative Zoology, 125, 345 - 387." type="journal article" year="1961">Clarke, 1961</bibRefCitation>
</taxonomicName>
differs in 1) shell proportions (0.81–0.93 vs 0.91–1.11, SL/SW ratio), 2) the lesser acute spire angle (min. 93.5 vs max. 85°), 3) the wider umbilicus (440 vs 135 µm), 4) the larger protoconch (one whorl of c. 350 µm vs 1.25 whorl of 280 µm), 5) the higher number of axial ribs at last whorl (37 vs 29–31), 6) the round vs oval aperture.
</paragraph>
<paragraph id="7160368CFFDDE353FF0FFCE0FE03EF8F" blockId="11.[151,1437,515,901]" pageId="11" pageNumber="198">
The oval aperture and the characteristic sculpture make the species easily separable from the remaining
<collectingCountry id="09C8761CFFDDE353FF5EFCC8FE8DEF8F" box="[151,260,875,901]" name="Antarctica" pageId="11" pageNumber="198">Antarctic</collectingCountry>
congeners.
</paragraph>
</subSubSection>
</treatment>
</document>
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