185 lines
22 KiB
XML
185 lines
22 KiB
XML
<document ID-CLB-Dataset="7244" ID-DOI="http://dx.doi.org/10.3897/asp.79.e66021" ID-GBIF-Dataset="1f858329-d759-403a-829c-08f2ba4e1403" ID-Pensoft-Pub="1864-8312-79-57" ID-Pensoft-UUID="B1CDE34C960C59639EA47231EE68A71F" ID-ZooBank="A42504B7FFAF4856884A7B74F3E03AA3" ModsDocID="1864-8312-79-57" checkinTime="1618938684678" checkinUser="pensoft" docAuthor="Grebennikov, Vasily V." docDate="2021" docId="340A628155E5598B88B266968BEBDA27" docLanguage="en" docName="ArthropodSystPhyl 79: 57-74" docOrigin="Arthropod Systematics & Phylogeny 79" docPubDate="2021-04-20" docSource="http://dx.doi.org/10.3897/asp.79.e66021" docTitle="Typoderus iphitus Grebennikov 2021, sp. nov." docType="treatment" docUuid="83775EEA-6CA3-47FB-99BB-0B6EEC752B35" docUuidSource="ZooBank" docVersion="7" id="B1CDE34C960C59639EA47231EE68A71F" lastPageNumber="57" masterDocId="B1CDE34C960C59639EA47231EE68A71F" masterDocTitle="Sky islands of the Cameroon Volcanic Line support the westernmost clade of five new Typoderus weevils (Coleoptera: Curculionidae: Molytinae)" masterLastPageNumber="74" masterPageNumber="57" pageNumber="57" updateTime="1732750079891" updateUser="ExternalLinkService">
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<mods:titleInfo id="5E5326578ABDA5CCC15CBD2452D97F3C">
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<mods:title id="3973FCA0852D441727B2A84FA3D4F660">Sky islands of the Cameroon Volcanic Line support the westernmost clade of five new Typoderus weevils (Coleoptera: Curculionidae: Molytinae)</mods:title>
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<mods:name id="A39BF4F55B81C0360D54DE098D737321" type="personal">
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<mods:namePart id="9BF19E9D9DE66FA815A06EE270B571F9">Grebennikov, Vasily V.</mods:namePart>
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<mods:affiliation id="8E89EB9A3DC93687F91281D6EEAA6F62">Canadian Food Inspection Agency, 960 Carling Ave., Ottawa, ON, K 1 A 0 Y 9, Canada</mods:affiliation>
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<mods:nameIdentifier id="2111308B08AF72E17B6C4CA640D72C48" type="email">vasily.grebennikov@canada.ca</mods:nameIdentifier>
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<mods:typeOfResource id="1F644AA2F19A5256EFD1DE8440B8E99D">text</mods:typeOfResource>
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<mods:titleInfo id="1E5EEB42737730D803EC3428DB90453E">
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<mods:title id="B488840AA6C803AC3712915D3AD92804">Arthropod Systematics & Phylogeny</mods:title>
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</mods:titleInfo>
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<mods:part id="A94745D72FAA40A9377FA638D38569C8">
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<mods:date id="3C3957B0B7312664477E832D62F8F41C">2021</mods:date>
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<mods:detail id="624F7CCE2329F17F074A5C0C16E1E71F" type="pubDate">
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<mods:number id="0327E68B93E33A2FC659D81F69759A5B">2021-04-20</mods:number>
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<mods:detail id="F0F9A48E57EF840CDB7477611DB03285" type="volume">
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<mods:number id="25A911B812802AB7834489F8627DC65D">79</mods:number>
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<mods:extent id="EB95239E677CCF306508F2545DE75A29" unit="page">
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<mods:start id="FD23B1A61E854AFF45A2CDB0A3B1AE7B">57</mods:start>
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<mods:end id="523D9FF9D46EF77FA13D29C864FBFC4D">74</mods:end>
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<mods:url id="045B2E974C4D561B7B841947B7D38EF9">http://dx.doi.org/10.3897/asp.79.e66021</mods:url>
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<mods:classification id="938BF1C0F02D66BB5F8049BC873F8C03">journal article</mods:classification>
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<mods:identifier id="D9A7BA31226E0BE547538D59DA22454D" type="DOI">http://dx.doi.org/10.3897/asp.79.e66021</mods:identifier>
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<mods:identifier id="C2AAB6425A78971E01AD177A616B946D" type="Pensoft-Pub">1864-8312-79-57</mods:identifier>
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<mods:identifier id="47B7D62E94FFFDFF8AB9D1451E251F5B" type="ZooBank">A42504B7FFAF4856884A7B74F3E03AA3</mods:identifier>
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<mods:identifier id="9C787B5999D55964020A2EBA0B2151F4" type="Pensoft-UUID">B1CDE34C960C59639EA47231EE68A71F</mods:identifier>
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<treatment id="340A628155E5598B88B266968BEBDA27" ID-GBIF-Taxon="181520728" LSID="urn:lsid:zoobank.org:act:83775EEA-6CA3-47FB-99BB-0B6EEC752B35" httpUri="http://treatment.plazi.org/id/340A628155E5598B88B266968BEBDA27" lastPageNumber="57" pageId="0" pageNumber="57" scope_family="Curculionidae" scope_order="Coleoptera">
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<subSubSection id="FE9882B95A42D9737A6301E35EDEA4A2" pageId="0" pageNumber="57" type="nomenclature">
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<paragraph id="F95CA906D81531E89D38E8AB683FF8FF" pageId="0" pageNumber="57">
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4.2.5.
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<taxonomicName id="AF9C2BF1360C17DB42CA5E4CAE6DD765" LSID="http://zoobank.org/83775EEA-6CA3-47FB-99BB-0B6EEC752B35" authority="Grebennikov, 2021" authorityName="Grebennikov" authorityYear="2021" class="Insecta" family="Curculionidae" genus="Typoderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typoderus iphitus" order="Coleoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="iphitus" status="sp. nov.">Typoderus iphitus</taxonomicName>
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<taxonomicNameLabel id="0FE4C25EC55575BB9E3CB8F35B2D3313" pageId="0" pageNumber="57">sp. nov.</taxonomicNameLabel>
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<figureCitation id="C7A78688C9EFEC29B22A0C5280C13B95" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of the genus Typoderus. Map designed using the online SimpleMappr tool (Shorthouse 2010)." figureDoi="10.3897/asp.79.e66021.figure1" httpUri="https://binary.pensoft.net/fig/533979" pageId="0" pageNumber="57">Figs 1</figureCitation>
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<figureCitation id="49618D2942A8F20E207EF8A3E25DFE90" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Neighbour Joining clustering of 20 DNA barcodes of Cameroonian Typoderus (analysis A 1). Terminal names consist of BOLD sample ID (its last four digits correspond to specimen number), litter sifting sample number (Table 1), length of the DNA barcode (number of ambiguous nucleotides is in square brackets), BIN three-letter four-digit number and GenBank accession number. Names next to clusters are those of the new species described in section 4.2. HT denote holotypes." figureDoi="10.3897/asp.79.e66021.figure2" httpUri="https://binary.pensoft.net/fig/533980" pageId="0" pageNumber="57">, 2</figureCitation>
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<figureCitation id="CFA507F458B6282A52CF5B1A866850FC" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Maximum Likelihood inference phylogram of Typoderus weevils rooted on Lupangus (analysis A 3, root is not shown). Cameroonian species form a clade. Digits at internodes are bootstrap values of 50 % and above. Non-Cameroonian (= Tanzanian) terminals are collapsed in species. Terminal names consist of specimen number (Table 3) and sifting sample number (Table 1). Habitus images are to scale; their numbers indicate exact Cameroonian specimens imaged. Names next to clades are those of the new species described in section 4.2." figureDoi="10.3897/asp.79.e66021.figure3" httpUri="https://binary.pensoft.net/fig/533981" pageId="0" pageNumber="57">, 3</figureCitation>
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<figureCitation id="523AD02DDB5DB7A6C1C44534B40578C8" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Maximum clade credibility Cameroonian Typoderus tree with median ages derived from the BEAST analysis A 4. The 95 % credibility interval for each node is given with a horizontal bar. Digits at nodes are million years before present. Terminal names consist of specimen number (Table 3), GenBank accession number and sifting sample number (Table 1). Names next to clades are those of the new species described in section 4.2." figureDoi="10.3897/asp.79.e66021.figure4" httpUri="https://binary.pensoft.net/fig/533982" pageId="0" pageNumber="57">, 4</figureCitation>
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<figureCitation id="F4CE6DBDF9548CBA5AF11A073FC88B29" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male holotype of T. iphitus sp. nov., habitus (A: dorsal, B: left lateral, C: ventral, D: left fronto-lateral), aedeagus (E: dorsal, F: ventral, G: right lateral) and H: sternite 9." figureDoi="10.3897/asp.79.e66021.figure8" httpUri="https://binary.pensoft.net/fig/533986" pageId="0" pageNumber="57">, 8A-H.</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="45C05CF6A5E486CBBC7729AC40BC82F8" pageId="0" pageNumber="57" type="description">
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<paragraph id="D73AF07046250C55EB543E65F599D0AF" pageId="0" pageNumber="57">Species diagnosis and description.</paragraph>
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<paragraph id="FC239ACCB3E529381C92A163E775219A" pageId="0" pageNumber="57">
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Holotype male (Figs
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<figureCitation id="59BC6261FD1FDE3144B337430466603E" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male holotype of T. iphitus sp. nov., habitus (A: dorsal, B: left lateral, C: ventral, D: left fronto-lateral), aedeagus (E: dorsal, F: ventral, G: right lateral) and H: sternite 9." figureDoi="10.3897/asp.79.e66021.figure8" httpUri="https://binary.pensoft.net/fig/533986" pageId="0" pageNumber="57">8A-H</figureCitation>
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), length between anterior edge of pronotum and elytral apex 4.6 mm, DNA barcode: MH917891. Diagnostic combination of morphological characters as in Table
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<tableCitation id="8DEABF1388189099640F76BB81CD7C23" captionStart="Table 4" captionStartId="T4" captionText="Table 4. Character matrix with discrete morphological characters for diagnostics of new Cameroonian Typoderus weevils (see also Discussion). - Characters: 1. - Body, colour of darkest specimens (Fig. 3, that is of a beetle at least a few days old, because specimens freshly emerged from the pupa are much paler): greyish: 0; brownish: 1; blackish: 2. 2. - Elytra at middle in cross-section: triangular (Figs 5 A-H, 6 A-H); elytral disk flat, ridge in interstriae 6 - 7 forming elytral lateral contour in dorsal view: 0; cylindrical (Figs 7 A-H, 8 A-H, 9 A-H); elytral disk evenly rounded laterally, ridge in interstriae 6 - 7 positioned inside lateral contour in dorsal view: 1. 3. - Hind tibiae, additional apical projection near inner apical spur (in at least some males): absent (Figs 7 B, 8 B, 9 B): 0; present (Figs 5 B, 6 B): 1. 4. - Pilosity on elytral declivity (excluding that on elytral protuberances), whether similar to that on the rest of elytra; similar: 0; dissimilar, notably denser and longer (Fig. 9 A, except abraded specimens): 1. 5. - Ratio of length to width of sclerotized part of aedeagus, dorsal view: one (aedeagus subquadrate as in Figs 5 E, 6 E, 9 E or notably unsclerotized, as in Fig. 8 E): 1; two (aedeagus elongate, Fig. 7 E): 2. 6. - Numerous (about 10) " teeth " inside internal sac of aedeagus: absent (Figs 5 E-F, 6 E-F, 8 E-F, 9 E-F): 0; present (Figs 7 E-F): 1. 7. - Tegminal apodeme: absent (Figs 6 F, 8 F) or indistinguishably short (Figs 5 F, 9 F): 0; present, consisting of two rods (Fig. 7 F): 1." httpUri="http://table.plazi.org/id/9CBF17CBDB7A16608D6711C763B03F8C" pageId="0" pageNumber="57" tableUuid="9CBF17CBDB7A16608D6711C763B03F8C">4</tableCitation>
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. Sister clade to
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<taxonomicName id="DD172B5DEAF0CD485C3FFDBBE8747536" lsidName="T. amphion" pageId="0" pageNumber="57" rank="species" species="amphion">
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<emphasis id="618328C86D7499BAF20725FAD3455B78" italics="true" pageId="0" pageNumber="57">T. amphion</emphasis>
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</taxonomicName>
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<emphasis id="4E23C8BF611833E80E5E48BAE5CA9759" bold="true" pageId="0" pageNumber="57">sp. nov.</emphasis>
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,
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<taxonomicName id="374828F1EC1742A229391B34830E67D0" lsidName="T. telamon" pageId="0" pageNumber="57" rank="species" species="telamon">
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<emphasis id="720842CB96AF5BC373A04BE303B7BA6E" italics="true" pageId="0" pageNumber="57">T. telamon</emphasis>
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</taxonomicName>
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<emphasis id="AC2D79C9B4587685C4371D0F002DA72C" bold="true" pageId="0" pageNumber="57">sp. nov.</emphasis>
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and
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<taxonomicName id="82637EE943CE712176A46CDE2DD1EAD5" lsidName="T. clytius" pageId="0" pageNumber="57" rank="species" species="clytius">
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<emphasis id="7F0336F4EFFC105BAEE28C4007256588" italics="true" pageId="0" pageNumber="57">T. clytius</emphasis>
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</taxonomicName>
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<emphasis id="95D6D01AEAED6F9E81A5BB467562F89C" bold="true" pageId="0" pageNumber="57">sp. nov.</emphasis>
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(Fig.
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<figureCitation id="D15E0640616A8D18EB157B2A220A4F68" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Maximum Likelihood inference phylogram of Typoderus weevils rooted on Lupangus (analysis A 3, root is not shown). Cameroonian species form a clade. Digits at internodes are bootstrap values of 50 % and above. Non-Cameroonian (= Tanzanian) terminals are collapsed in species. Terminal names consist of specimen number (Table 3) and sifting sample number (Table 1). Habitus images are to scale; their numbers indicate exact Cameroonian specimens imaged. Names next to clades are those of the new species described in section 4.2." figureDoi="10.3897/asp.79.e66021.figure3" httpUri="https://binary.pensoft.net/fig/533981" pageId="0" pageNumber="57">3</figureCitation>
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).
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</paragraph>
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<caption id="7E0D3F4BFD6ADDC723C76ECEC170C102" doi="10.3897/asp.79.e66021.figure8" httpUri="https://binary.pensoft.net/fig/533986" pageId="0" pageNumber="57" start="Figure 8" startId="F8">
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<paragraph id="E4F243A452A308F43BB2AE4609EF6819" pageId="0" pageNumber="57">
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<emphasis id="136CC036905B77F307CEB00F704BBE55" bold="true" pageId="0" pageNumber="57">Figure 8.</emphasis>
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Male holotype of
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<taxonomicName id="947791EA73D76DCC7E7613480596C532" lsidName="T. iphitus" pageId="0" pageNumber="57" rank="species" species="iphitus">
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<emphasis id="86652D846D3C60648BD27517E0C9D3BA" italics="true" pageId="0" pageNumber="57">T. iphitus</emphasis>
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</taxonomicName>
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<emphasis id="9FC99FAD066EF3F9F77A9182CFB8EF88" bold="true" pageId="0" pageNumber="57">sp. nov.</emphasis>
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, habitus (
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<emphasis id="F09BC2EDB031636B5A482DA4DEFC8B8E" bold="true" pageId="0" pageNumber="57">A</emphasis>
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: dorsal,
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<emphasis id="06DD74A32F841E6414D6F39EBD599CC5" bold="true" pageId="0" pageNumber="57">B</emphasis>
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: left lateral,
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<emphasis id="B03448AE65A932FAB9DDFD2AEC783A27" bold="true" pageId="0" pageNumber="57">C</emphasis>
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: ventral,
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<emphasis id="338F62E7FCE13ACA889557CBDEC0D9C7" bold="true" pageId="0" pageNumber="57">D</emphasis>
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: left fronto-lateral), aedeagus (
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<emphasis id="253811760A3009A3D39A1F844FB7A5AF" bold="true" pageId="0" pageNumber="57">E</emphasis>
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: dorsal,
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<emphasis id="040AAB414C0EBCD833F9716C551D0230" bold="true" pageId="0" pageNumber="57">F</emphasis>
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: ventral,
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<emphasis id="B87EBC9114EA59FA23B39E480B374FB1" bold="true" pageId="0" pageNumber="57">G</emphasis>
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: right lateral) and
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<emphasis id="9DB60A15318E7417DE77277DD4057E16" bold="true" pageId="0" pageNumber="57">H</emphasis>
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: sternite 9.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="F3C2F8CF44C35ED4D2703B0A9E5DDC46" pageId="0" pageNumber="57" type="materials_examined">
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<paragraph id="799D77B522CD4EE67E54956E8F20AA75" pageId="0" pageNumber="57">Material examined.</paragraph>
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<paragraph id="74CD9DFBB0A067C95521EA87B40B5A2B" pageId="0" pageNumber="57">
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<materialsCitation id="51B392D834AD3865720E9EAD1979CEE8" ID-GBIF-Occurrence="3080451310" collectingDate="2015-12-29" collectorName="V. Grebennikov" country="CAMEROON" elevation="1977" latitude="4.8017" location="Mt. Kupe" longLatPrecision="7" longitude="9.7015" specimenCount="1" typeStatus="Holotype">
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<typeStatus id="82C55013E4B38FD585FBE223207A12B5">Holotype</typeStatus>
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male (CNC): "
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<collectingCountry id="857FA6B803A219A85C9E8D725B207CEE" name="Cameroon">CAMEROON</collectingCountry>
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,
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<location id="4D18AF91B6C6FE2A8C86886E6C8DA884" LSID="urn:lsid:plazi:treatment:340A628155E5598B88B266968BEBDA27:4D18AF91B6C6FE2A8C86886E6C8DA884" country="CAMEROON" latitude="4.8017" longLatPrecision="7" longitude="9.7015" name="Mt. Kupe">Mt. Kupe</location>
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,
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<geoCoordinate id="410B82CC8A6E751741F78084CA989FD8" degrees="4.8017" direction="north" orientation="latitude" precision="5" value="4.8017">4.8017</geoCoordinate>
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,
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<geoCoordinate id="E33278BE69CC447463DEE0AB262E7849" degrees="9.7015" direction="east" orientation="longitude" precision="5" value="9.7015">9.7015</geoCoordinate>
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,
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<quantity id="35EFF6B5AE37DCDF3310BBBC0AA27579" metricMagnitude="3" metricUnit="m" metricValue="1.9769999999999999" unit="m" value="1977.0">
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<elevation id="6F8FA290CAFF8C6C37DCF5FEF52488B0" metricMagnitude="3" metricUnit="m" metricValue="1.9769999999999999" unit="m" value="1977.0">1977m</elevation>
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</quantity>
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,
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<collectingDate id="5FE1E983970059597F126F1FA20C5843" value="2015-12-29">29.xii.2015</collectingDate>
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, sift.CM15,
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<collectorName id="F0787456B0C16571C75D058D29C04EE7">V.Grebennikov</collectorName>
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",
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<normalizedToken id="7CD125E08AE0F140B48E009244275F90" originalValue="“CNCCOLVG00009722”">"CNCCOLVG00009722"</normalizedToken>
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</materialsCitation>
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.
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<materialsCitation id="3997BF93F6669723A6012AC797203C9F" ID-GBIF-Occurrence="3080451303" collectingDate="2015-12-29" collectorName="V. Grebennikov" country="CAMEROON" elevation="1525" latitude="4.8193" location="Mt. Kupe" longLatPrecision="7" longitude="9.7075" specimenCount="1" typeStatus="Paratypes">
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<typeStatus id="A479A0F5442D9F2C742AA72B5BF51A6D">Paratypes</typeStatus>
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(CNC): 4, as in Fig.
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<figureCitation id="2DC85D8189E68B797B8BB534D4BD08E7" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Maximum Likelihood inference phylogram of Typoderus weevils rooted on Lupangus (analysis A 3, root is not shown). Cameroonian species form a clade. Digits at internodes are bootstrap values of 50 % and above. Non-Cameroonian (= Tanzanian) terminals are collapsed in species. Terminal names consist of specimen number (Table 3) and sifting sample number (Table 1). Habitus images are to scale; their numbers indicate exact Cameroonian specimens imaged. Names next to clades are those of the new species described in section 4.2." figureDoi="10.3897/asp.79.e66021.figure3" httpUri="https://binary.pensoft.net/fig/533981" pageId="0" pageNumber="57">3</figureCitation>
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; specimens 9556, 9723 and 9724: same data as holotype; specimen 9531: "
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<collectingCountry id="91089A3E9AAF736B89DF7511A5F3345C" name="Cameroon">CAMEROON</collectingCountry>
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,
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<location id="6A962D642815F16FFF8ADD215A0AC3DA" LSID="urn:lsid:plazi:treatment:340A628155E5598B88B266968BEBDA27:6A962D642815F16FFF8ADD215A0AC3DA" country="CAMEROON" latitude="4.8193" longLatPrecision="7" longitude="9.7075" name="Mt. Kupe">Mt. Kupe</location>
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,
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<geoCoordinate id="86F8F293BD964ED0AA9407536323C373" degrees="4.8193" direction="north" orientation="latitude" precision="5" value="4.8193">4.8193</geoCoordinate>
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,
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<geoCoordinate id="309B524E0F26BED2217569D768BC7BC8" degrees="9.7075" direction="east" orientation="longitude" precision="5" value="9.7075">9.7075</geoCoordinate>
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,
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<quantity id="CCF46309EE9373CB8522317B3889F3DB" metricMagnitude="3" metricUnit="m" metricValue="1.525" unit="m" value="1525.0">
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<elevation id="D62FE8559A27272067450AEBF0FFF826" metricMagnitude="3" metricUnit="m" metricValue="1.525" unit="m" value="1525.0">1525m</elevation>
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|
</quantity>
|
|
,
|
|
<collectingDate id="D96CAC95355ED048EC203F6CBAA27DA3" value="2015-12-29">29.xii.2015</collectingDate>
|
|
, sift.CM13,
|
|
<collectorName id="EAEE8FA6ED738E170956D9B5619F8058">V.Grebennikov</collectorName>
|
|
"
|
|
</materialsCitation>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="A7C81B2DCE56B897EEC71E35294525EA" pageId="0" pageNumber="57" type="distribution">
|
|
<paragraph id="D3D9176463ECFD9A8E7896CCA3C4D265" pageId="0" pageNumber="57">Distribution.</paragraph>
|
|
<paragraph id="3C893F16663B5EBB669371B89B1889B7" pageId="0" pageNumber="57">
|
|
Known only from Mt. Kupe in Cameroon, in sympatry with
|
|
<taxonomicName id="7C6691AB6BEB27ED1A1E9BAEDF281E10" lsidName="T. telamon" pageId="0" pageNumber="57" rank="species" species="telamon">
|
|
<emphasis id="51643BCF2976CB44B17D1BBA5B5C9E47" italics="true" pageId="0" pageNumber="57">T. telamon</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="4D8D7EF05D10E5567A08768430E59EC6" bold="true" pageId="0" pageNumber="57">sp. nov.</emphasis>
|
|
Elevation: 1,525-1,977 m.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="D0F67D7F1AACC6BC3B254CBCFE9F11BB" pageId="0" pageNumber="57" type="etymology">
|
|
<paragraph id="4BEA8D1DD31F47C9B82327DCAE93F4AA" pageId="0" pageNumber="57">Etymology.</paragraph>
|
|
<paragraph id="FBC3E7AE3CDBB992EF7E20AD37661485" pageId="0" pageNumber="57">Iphitus, from ancient Greek mythology, one of the Argonauts, brother of Clytius; noun in apposition.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="5A9EB9E35AF8BF29C7195FC78F415EE4" pageId="0" pageNumber="57" type="comments">
|
|
<paragraph id="CF2D317ABF69026FBF0A5CFACCFA7A3B" pageId="0" pageNumber="57">Comment.</paragraph>
|
|
<paragraph id="29C5B79531A7A15F1A5CF15145271819" pageId="0" pageNumber="57">
|
|
Being morphologically indistinguishable and sympatric, it is more parsimonious to treat all specimens assigned to this species as the same biological entity, despite a relatively deep intraspecific divergence (Figs
|
|
<figureCitation id="EEE6F2A1BC133006F8A8CD2D312910ED" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Neighbour Joining clustering of 20 DNA barcodes of Cameroonian Typoderus (analysis A 1). Terminal names consist of BOLD sample ID (its last four digits correspond to specimen number), litter sifting sample number (Table 1), length of the DNA barcode (number of ambiguous nucleotides is in square brackets), BIN three-letter four-digit number and GenBank accession number. Names next to clusters are those of the new species described in section 4.2. HT denote holotypes." figureDoi="10.3897/asp.79.e66021.figure2" httpUri="https://binary.pensoft.net/fig/533980" pageId="0" pageNumber="57">2</figureCitation>
|
|
-
|
|
<figureCitation id="6EDD599153C7A3D28EC3B684734F14F1" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Maximum clade credibility Cameroonian Typoderus tree with median ages derived from the BEAST analysis A 4. The 95 % credibility interval for each node is given with a horizontal bar. Digits at nodes are million years before present. Terminal names consist of specimen number (Table 3), GenBank accession number and sifting sample number (Table 1). Names next to clades are those of the new species described in section 4.2." figureDoi="10.3897/asp.79.e66021.figure4" httpUri="https://binary.pensoft.net/fig/533982" pageId="0" pageNumber="57">4</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
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</document> |