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<document id="942A0D1EF24B7FCFEE1AFD190B41B569" ID-CLB-Dataset="26919" ID-DOI="10.11646/zootaxa.4598.1.1" ID-GBIF-Dataset="475fd53e-c5c6-4a99-bb2e-cc2382f0171f" ID-ISSN="1175-5326" ID-Zenodo-Dep="2668063" ID-ZooBank="6BA5F086-1014-46B5-B6D5-77FE0CD07AEC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1557127411994" checkinUser="plazi" docAuthor="Seldon, David S. &amp; Buckley, Thomas R." docDate="2019" docId="03BBD0260247B74A90FBB3BBEEE3FB04" docLanguage="en" docName="zootaxa.4598.1.1.pdf.imf" docOrigin="Zootaxa 4598 (1)" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Mecodema longicolle * Broun 1923" docType="treatment" docVersion="11" lastPageNumber="69" masterDocId="FF82A85E0205B70E906CB267ED0BFFA4" masterDocTitle="The genus Mecodema Blanchard 1853 (Coleoptera: Carabidae: Broscini) from the North Island, New Zealand" masterLastPageNumber="148" masterPageNumber="1" pageNumber="67" updateTime="1698726038790" updateUser="ExternalLinkService">
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<mods:title id="341A428951AC2620B290FCE349D2EC29">The genus Mecodema Blanchard 1853 (Coleoptera: Carabidae: Broscini) from the North Island, New Zealand</mods:title>
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<mods:namePart id="60F194F8AA0966068FDF9E3E3F35A6D4">Seldon, David S.</mods:namePart>
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<mods:namePart id="5E7948FFDEB274C98D2656C8A0D19A22">Buckley, Thomas R.</mods:namePart>
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<treatment id="03BBD0260247B74A90FBB3BBEEE3FB04" ID-DOI="http://doi.org/10.5281/zenodo.5678394" ID-GBIF-Taxon="157062680" ID-Zenodo-Dep="5678394" LSID="urn:lsid:plazi:treatment:03BBD0260247B74A90FBB3BBEEE3FB04" httpUri="http://treatment.plazi.org/id/03BBD0260247B74A90FBB3BBEEE3FB04" lastPageId="68" lastPageNumber="69" pageId="66" pageNumber="67">
<subSubSection id="C30832BB0247B74C90FBB3BBEF4BFE52" box="[151,576,476,502]" pageId="66" pageNumber="67" type="nomenclature">
<paragraph id="8BAD61300247B74C90FBB3BBEF4BFE52" blockId="66.[151,576,476,535]" box="[151,576,476,502]" pageId="66" pageNumber="67">
<heading id="D0E5D65C0247B74C90FBB3BBEF4BFE52" bold="true" box="[151,576,476,502]" fontSize="11" level="1" pageId="66" pageNumber="67" reason="1">
<emphasis id="B966BD220247B74C90FBB3BBEF4BFE52" bold="true" box="[151,576,476,502]" pageId="66" pageNumber="67">
<taxonomicName id="4C121AB30247B74C90FBB3BBEF30FE52" ID-CoL="6QWMX" authority="Broun, 1923" authorityName="* Broun" authorityYear="1923" box="[151,571,476,502]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="longicolle">
<emphasis id="B966BD220247B74C90FBB3BBEC92FE52" bold="true" box="[151,409,476,502]" italics="true" pageId="66" pageNumber="67">Mecodema longicolle</emphasis>
<bibRefCitation id="EF831CC10247B74C91CCB3BBEF30FE52" author="Broun, T." box="[416,571,476,502]" pageId="66" pageNumber="67" pagination="667 - 708" refId="ref91555" refString="Broun, T. (1923) Descriptions of new genera and species of Coleoptera. [Part VIII]. Bulletin of the New Zealand Institute, 1, 667 - 708." type="journal article" year="1923">Broun, 1923</bibRefCitation>
</taxonomicName>
.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30832BB0247B74C90FBB399EC01FDB3" box="[151,266,510,535]" pageId="66" pageNumber="67" type="description">
<paragraph id="8BAD61300247B74C90FBB399EC01FDB3" blockId="66.[151,576,476,535]" box="[151,266,510,535]" pageId="66" pageNumber="67">
<figureCitation id="13297DB50247B74C90FBB399EC0DFDB3" box="[151,262,510,535]" captionStart="FIGURE 36" captionStartId="67.[151,250,1881,1903]" captionTargetBox="[298,1286,200,1844]" captionTargetId="figure@67.[290,1296,193,1860]" captionTargetPageId="67" captionText="FIGURE 36. Mecodema longicolle plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668135/files/figure.png" pageId="66" pageNumber="67">Figure 36</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30832BB0247B74C90FBB021E925FDF8" box="[151,1070,582,605]" pageId="66" pageNumber="67" type="reference_group">
<paragraph id="8BAD61300247B74C90FBB021E925FDF8" blockId="66.[151,1070,582,605]" box="[151,1070,582,605]" pageId="66" pageNumber="67">
<taxonomicName id="4C121AB30247B74C90FBB021EF08FDF8" ID-CoL="738N6" authority="Broun, 1923" authorityName="Broun" authorityYear="1923" box="[151,515,582,604]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="subaeneum">
<emphasis id="B966BD220247B74C90FBB021EC75FDF8" box="[151,382,582,604]" italics="true" pageId="66" pageNumber="67">Mecodema subaeneum</emphasis>
<bibRefCitation id="EF831CC10247B74C91E8B020EF08FDF8" author="Broun, T." box="[388,515,582,604]" pageId="66" pageNumber="67" pagination="667 - 708" refId="ref91555" refString="Broun, T. (1923) Descriptions of new genera and species of Coleoptera. [Part VIII]. Bulletin of the New Zealand Institute, 1, 667 - 708." type="journal article" year="1923">Broun, 1923</bibRefCitation>
</taxonomicName>
(Martinborough, WA), synonymised by
<bibRefCitation id="EF831CC10247B74C93CCB021E921FDF8" author="Britton, E. B." box="[928,1066,582,604]" pageId="66" pageNumber="67" pagination="533 - 581" refId="ref90973" refString="Britton, E. B. (1949) The Carabidae (Coleoptera) of New Zealand part 3: A revision of the tribe Broscini. Transactions of the Royal Society of New Zealand, 77, 533 - 581." type="journal article" year="1949">Britton, 1949</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30832BB0247B74C90FBB0EEEF8FFD4E" pageId="66" pageNumber="67" type="diagnosis">
<paragraph id="8BAD61300247B74C90FBB0EEEF8FFD4E" blockId="66.[151,1437,649,2012]" pageId="66" pageNumber="67">
<emphasis id="B966BD220247B74C90FBB0EEEC18FD06" bold="true" box="[151,275,649,674]" pageId="66" pageNumber="67">Diagnosis:</emphasis>
Distinguished from other North Island
<taxonomicName id="4C121AB30247B74C92A7B0EEEE4DFD05" authorityName="Blanchard" authorityYear="1853" box="[715,838,649,673]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="B966BD220247B74C92A7B0EEEE4DFD05" box="[715,838,649,673]" italics="true" pageId="66" pageNumber="67">Mecodema</emphasis>
</taxonomicName>
species by having:
<emphasis id="B966BD220247B74C944CB0EEE925FD06" bold="true" box="[1056,1070,649,674]" pageId="66" pageNumber="67">1</emphasis>
, stipes with 3 basal setae seta;
<emphasis id="B966BD220247B74C95EBB0EEE89EFD06" bold="true" box="[1415,1429,649,674]" pageId="66" pageNumber="67">2</emphasis>
, prothoracic carina narrow the entire length, moderately crenulated with 79 setae each side;
<emphasis id="B966BD220247B74C94C9B0CAE9B8FD62" bold="true" box="[1189,1203,685,710]" pageId="66" pageNumber="67">3</emphasis>
, distinctive shape of apical portion of the penis lobe (
<figureCitation id="13297DB50247B74C926EB0B6EF58FD4E" box="[514,595,721,746]" captionStart="FIGURE 36" captionStartId="67.[151,250,1881,1903]" captionTargetBox="[298,1286,200,1844]" captionTargetId="figure@67.[290,1296,193,1860]" captionTargetPageId="67" captionText="FIGURE 36. Mecodema longicolle plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668135/files/figure.png" pageId="66" pageNumber="67">Fig. 36</figureCitation>
PL).
</paragraph>
</subSubSection>
<subSubSection id="C30832BB0247B74A90ABB092EE24FD0C" lastPageId="68" lastPageNumber="69" pageId="66" pageNumber="67" type="description">
<paragraph id="8BAD61300247B74C90ABB092EEA9FC96" blockId="66.[151,1437,649,2012]" pageId="66" pageNumber="67">
<emphasis id="B966BD220247B74C90ABB092EC50FCAA" bold="true" box="[199,347,757,782]" pageId="66" pageNumber="67">Description:</emphasis>
Length
<quantity id="4CEACCD50247B74C91D7B092EF47FCA9" box="[443,588,757,782]" metricMagnitude="-2" metricUnit="m" metricValue="1.775" metricValueMax="2.0" metricValueMin="1.55" pageId="66" pageNumber="67" unit="mm" value="17.75" valueMax="20.0" valueMin="15.5">15.520 mm</quantity>
, pronotal width
<quantity id="4CEACCD50247B74C9364B092EE75FCA9" box="[776,894,757,782]" metricMagnitude="-3" metricUnit="m" metricValue="4.45" metricValueMax="5.0" metricValueMin="3.9" pageId="66" pageNumber="67" unit="mm" value="4.45" valueMax="5.0" valueMin="3.9">3.95 mm</quantity>
, elytral width
<quantity id="4CEACCD50247B74C944FB091E992FCA9" box="[1059,1177,757,782]" metricMagnitude="-3" metricUnit="m" metricValue="5.35" metricValueMax="6.0" metricValueMin="4.7" pageId="66" pageNumber="67" unit="mm" value="5.35" valueMax="6.0" valueMin="4.7">4.76 mm</quantity>
. Colour of entire body dark reddish-brown to glossy black, coxae reddish-brown, legs black.
</paragraph>
<paragraph id="8BAD61300247B74C90ABB15AE9D5FB3D" blockId="66.[151,1437,649,2012]" pageId="66" pageNumber="67">
<emphasis id="B966BD220247B74C90ABB15AEC06FCF1" box="[199,269,829,853]" italics="true" pageId="66" pageNumber="67">Head:</emphasis>
Narrow and convex (
<figureCitation id="13297DB50247B74C927DB15AEF76FCF2" box="[529,637,829,854]" captionStart="FIGURE 14" captionStartId="20.[151,250,936,958]" captionTargetBox="[171,1412,204,911]" captionTargetId="figure@20.[160,1412,193,921]" captionTargetPageId="20" captionText="FIGURE 14. Diagrammatic cross sections that represent the width and convexity of head or deflection of pronotum (frontal view): (A) broad and flat; (B) broad and convex; (C) narrow and flat; (D) narrow and convex." httpUri="https://zenodo.org/record/2668091/files/figure.png" pageId="66" pageNumber="67">Fig. 14D</figureCitation>
). Vertex with very fine wrinkles (almost indistinct); vertexal groove slightly impressed, defined by punctures and short rugose wrinkles the entire length (
<figureCitation id="13297DB50247B74C9405B106E9B4FCDE" box="[1129,1215,865,890]" captionStart="FIGURE 36" captionStartId="67.[151,250,1881,1903]" captionTargetBox="[298,1286,200,1844]" captionTargetId="figure@67.[290,1296,193,1860]" captionTargetPageId="67" captionText="FIGURE 36. Mecodema longicolle plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668135/files/figure.png" pageId="66" pageNumber="67">Fig. 36</figureCitation>
); large supraorbital puncture bearing 34 setae; 34 shallow and short supraorbital grooves; frons (
<figureCitation id="13297DB50247B74C9448B1E2E962FC3A" box="[1060,1129,901,926]" captionStart="FIGURE 9" captionStartId="15.[151,250,1706,1728]" captionTargetBox="[157,1433,759,1683]" captionTargetId="figure@15.[151,1436,757,1683]" captionTargetPageId="15" captionText="FIGURE 9. Dorsal view of head with detail of the morphological structures used in the species descriptions. Ƒ = vertex; SPS = supraorbital puncture setae; SP = supraorbital puncture; F = frons area; FCS = frontoclypeal suture; C = clypeus; M = mandible; LS = labrum setae; L = labrum; ACM = anterior clypeal microsculpture; CSP = clypeal setose puncture; TP = tentorial pit; SG = supraorbital grooves; E = eye; ƑG = vertexal groove." httpUri="https://zenodo.org/record/2668081/files/figure.png" pageId="66" pageNumber="67">Fig. 9</figureCitation>
) with indistinct grooves, a large and shallow depression each side of midline anterad; frontoclypeal suture well-defined, tentorial pits small; clypeus convex medially, anterior with 3 narrow and indistinct grooves, 1 setose puncture on each side bearing 2 setae. Labrum rectangular, anterior edge slightly emarginated with 2 proximate central setae, 2 each side evenly spaced. Mentum lobes squared (
<figureCitation id="13297DB50247B74C9262B672EF71FB8A" box="[526,634,1045,1070]" captionStart="FIGURE 12" captionStartId="18.[151,250,1905,1927]" captionTargetBox="[311,1244,697,1878]" captionTargetId="figure@18.[310,1250,689,1889]" captionTargetPageId="18" captionText="FIGURE 12. Ventral view of the mentum with a number of different character states. Illustrations AD are the different forms of the indentation in the apex of the median process: (A) distinctly indentate (M. manaia); (B) moderately indentate (M. pluto); (C) slightly indentate (M. infimate); (D) indentation notched (M. parataiko). Illustrations EG are the three general forms (i.e., hashed lines) of the mentum lobes: (E) triangular; (F) rounded; (G) squared." httpUri="https://zenodo.org/record/2668087/files/figure.png" pageId="66" pageNumber="67">Fig. 12G</figureCitation>
), median process broad and very short, slightly angled upward (15°) apically, strongly indentate (
<figureCitation id="13297DB50247B74C91B0B65EEF4CFBF6" box="[476,583,1081,1106]" captionStart="FIGURE 12" captionStartId="18.[151,250,1905,1927]" captionTargetBox="[311,1244,697,1878]" captionTargetId="figure@18.[310,1250,689,1889]" captionTargetPageId="18" captionText="FIGURE 12. Ventral view of the mentum with a number of different character states. Illustrations AD are the different forms of the indentation in the apex of the median process: (A) distinctly indentate (M. manaia); (B) moderately indentate (M. pluto); (C) slightly indentate (M. infimate); (D) indentation notched (M. parataiko). Illustrations EG are the three general forms (i.e., hashed lines) of the mentum lobes: (E) triangular; (F) rounded; (G) squared." httpUri="https://zenodo.org/record/2668087/files/figure.png" pageId="66" pageNumber="67">Fig. 12A</figureCitation>
); mentum setae present. Submentum sclerite constriction narrow with 68 evenly spaced setae. Stipes with 3 basal setae. Gula pits (
<figureCitation id="13297DB50247B74C9371B63AEE6BFBD2" box="[797,864,1117,1142]" captionStart="FIGURE 8" captionStartId="13.[151,250,1097,1119]" captionTargetBox="[155,1432,309,1072]" captionTargetId="figure@13.[151,1436,303,1075]" captionTargetPageId="13" captionText="FIGURE 8. Detail of the ventral head with the taxonomic structures used in the species descriptions. SBMS = submentum setae; SBM = submentum sclerite; ML = mentum lobe; MP = mentum process (with indentation); MPW = mentum process width; SBS = stipes basal setae; MPL = mentum process length; MPS = mentum process setae; SBMC = submentum sclerite constriction; GP = gula pits; GS = gula suture." httpUri="https://zenodo.org/record/2668079/files/figure.png" pageId="66" pageNumber="67">Fig. 8</figureCitation>
) relatively large, gula suture defined entire length, gula flat and smooth. Gena with wrinkled lines forming an isodiametric pattern across entire area.
</paragraph>
<paragraph id="8BAD61300247B74C90ABB6C2EF33FA1E" blockId="66.[151,1437,649,2012]" pageId="66" pageNumber="67">
<emphasis id="B966BD220247B74C90ABB6C2EC49FB19" box="[199,322,1189,1213]" italics="true" pageId="66" pageNumber="67">Prothorax:</emphasis>
Prothoracic carina narrow the entire length, moderately crenulated with 79 setae each side (
<figureCitation id="13297DB50247B74C9503B6C2EDBDFB46" captionStart="FIGURE 36" captionStartId="67.[151,250,1881,1903]" captionTargetBox="[298,1286,200,1844]" captionTargetId="figure@67.[290,1296,193,1860]" captionTargetPageId="67" captionText="FIGURE 36. Mecodema longicolle plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668135/files/figure.png" pageId="66" pageNumber="67">Fig. 36</figureCitation>
), extended to anterior angle; posterior lateral sinuation indistinctly carinate, angled inward; pronotum narrow and greatly deflected, overall shape ovate, disc with transverse wrinkles, anterior and posterior edges with short rugose lines perpendicular to edge (
<figureCitation id="13297DB50247B74C925FB776EF81FA8E" box="[563,650,1297,1322]" captionStart="FIGURE 36" captionStartId="67.[151,250,1881,1903]" captionTargetBox="[298,1286,200,1844]" captionTargetId="figure@67.[290,1296,193,1860]" captionTargetPageId="67" captionText="FIGURE 36. Mecodema longicolle plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668135/files/figure.png" pageId="66" pageNumber="67">Fig. 36</figureCitation>
); midline well-defined, anterior medial triangular-shaped impression and posterior medial diamond-shaped impression present; pronotal foveae shallow and narrow (may be indistinct); anterior and posterior edges inwardly curved (
<figureCitation id="13297DB50247B74C92CEB73EEFFCFAD6" box="[674,759,1369,1394]" captionStart="FIGURE 36" captionStartId="67.[151,250,1881,1903]" captionTargetBox="[298,1286,200,1844]" captionTargetId="figure@67.[290,1296,193,1860]" captionTargetPageId="67" captionText="FIGURE 36. Mecodema longicolle plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668135/files/figure.png" pageId="66" pageNumber="67">Fig. 36</figureCitation>
). Prosternum flat with fine transverse lines, a small circular depression each side; proepisternum with numerous obsolescent punctures. Procoxal setae absent; protibia distally expanded and shovel-like (
<figureCitation id="13297DB50247B74C91A8B7C6EF20FA1E" box="[452,555,1441,1466]" captionStart="FIGURE 10" captionStartId="16.[151,250,1442,1464]" captionTargetBox="[152,1428,191,1421]" captionTargetId="figure@16.[151,1435,183,1427]" captionTargetPageId="16" captionText="FIGURE 10. The protibia form is a character used mainly to determine between genera, except for one species in Mecodema where the difference between (A) greatly distally expanded (M. antarctica) and (B) expanded distally (all other Mecodema species) is used." httpUri="https://zenodo.org/record/2668083/files/figure.png" pageId="66" pageNumber="67">Fig. 10B</figureCitation>
).
</paragraph>
<paragraph id="8BAD61300247B74C90ABB7A2EE7CF97B" blockId="66.[151,1437,649,2012]" pageId="66" pageNumber="67">
<emphasis id="B966BD220247B74C90ABB7A2EC1DFA79" box="[199,278,1477,1501]" italics="true" pageId="66" pageNumber="67">Elytra:</emphasis>
Narrow and greatly deflected, apex with steep slope; humeral angle anteriorly convergent (
<figureCitation id="13297DB50247B74C955EB7A2E884FA7A" box="[1330,1423,1477,1502]" captionStart="FIGURE 6" captionStartId="10.[151,250,519,541]" captionTargetBox="[156,1432,193,498]" captionTargetId="figure@10.[151,1436,193,498]" captionTargetPageId="10" captionText="FIGURE 6. Detail of the three different forms of the elytral humeral angle: (A) = anteriorly convergent (M. parataiko), (B) = evenly convex (M. oconnori), (C) = subangulate (M. atuanui)." httpUri="https://zenodo.org/record/2668075/files/figure.png" pageId="66" pageNumber="67">Fig. 6A</figureCitation>
); basal margin moderately curved and bevelled to base, interval 1 extended to margin; lateral carina very narrow the entire length, extended to humeral angle; humerus with 12 proximate setose punctures (
<figureCitation id="13297DB50247B74C94C8B46AE9F7F982" box="[1188,1276,1549,1574]" captionStart="FIGURE 36" captionStartId="67.[151,250,1881,1903]" captionTargetBox="[298,1286,200,1844]" captionTargetId="figure@67.[290,1296,193,1860]" captionTargetPageId="67" captionText="FIGURE 36. Mecodema longicolle plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668135/files/figure.png" pageId="66" pageNumber="67">Fig. 36</figureCitation>
); suture welldefined; striae 14 defined by long, narrow asetose punctures, irregularly spaced, striae 58 well-defined by larger asetose punctures; intervals 14 indistinct but anterior area flattened, elytra greatly deflected, interval 5 weakly convex, intervals 68 indistinct, interval microsculpture present as very fine lines emanating from punctures; 7
<superScript id="7C67CC780247B74C95FCB41FE897F922" attach="none" box="[1424,1436,1656,1670]" fontSize="6" pageId="66" pageNumber="67">th</superScript>
strial setal (
<figureCitation id="13297DB50247B74C9171B4C4EC6BF91F" box="[285,352,1699,1724]" captionStart="FIGURE 1" captionStartId="5.[151,250,1912,1934]" captionTargetBox="[159,1427,1029,1884]" captionTargetId="figure@5.[151,1436,1023,1890]" captionTargetPageId="5" captionText="FIGURE 1. Dorsal view of a Mecodema specimen showing taxonomic structures used in the species descriptions. H =head; P = pronotum; E = elytra; S = striae; I1 = interval 1; A = apex; 7S = 7 th strial setae; HSP = humerus setose puncture; HA = humeral angle; BM = basal margin; P = protibia." httpUri="https://zenodo.org/record/2668065/files/figure.png" pageId="66" pageNumber="67">Fig. 1</figureCitation>
) pattern with 3 setose punctures in anterior ½ (1
<superScript id="7C67CC780247B74C93F4B4F8EEA9F909" attach="left" box="[920,930,1695,1709]" fontSize="6" pageId="66" pageNumber="67">st</superScript>
setae is on junction of basal margin and 7
<superScript id="7C67CC780247B74C95FCB4F8E897F909" attach="none" box="[1424,1436,1695,1709]" fontSize="6" pageId="66" pageNumber="67">th</superScript>
stria), 56 setose punctures in posterior ½, setose punctures large.
</paragraph>
<paragraph id="8BAD61300247B74C90ABB48CEF4FF878" blockId="66.[151,1437,649,2012]" pageId="66" pageNumber="67">
<emphasis id="B966BD220247B74C90ABB48CEC88F8A7" box="[199,387,1771,1795]" italics="true" pageId="66" pageNumber="67">Ventral surface:</emphasis>
Mesepisternum (
<figureCitation id="13297DB50247B74C923BB48CEFA8F8A7" box="[599,675,1771,1796]" captionStart="FIGURE 2" captionStartId="6.[151,250,800,822]" captionTargetBox="[159,1432,197,773]" captionTargetId="figure@6.[151,1436,193,779]" captionTargetPageId="6" captionText="FIGURE 2. Ventral view of a Mecodema specimen showing specific morphological structures, excluding taxonomic structures indicated in detail figures, used in the species descriptions. ƑLF = ventrite lateral foveae; ƑSP = ventrite setose punctures; MTC = metacoxa; MTƑP = metaventrite process; MSC = mesocoxa; PC = procoxa; PS = prosternum; G = gena; PES = proepisternum; MSE = mesepisternum; MTE = metepisternum; Ƒ1Ƒ6 = ventrites 1-6 (ventrites 1-3 may be fused); M = midline (dashed line, not a taxonomic structure)." httpUri="https://zenodo.org/record/2668067/files/figure.png" pageId="66" pageNumber="67">Fig. 2</figureCitation>
) with numerous obsolescent punctures; metepisternum without microsculpture; setose punctures present on mesocoxa (2) and metacoxa (3). Abdominal ventrites 1 and 2 finely lineate with sparse obsolescent punctures; ventrites 35 lineate laterally, 1 setose puncture each side of midline; ventrite 6 (
<figureCitation id="13297DB50247B74C9175B531EC6AF8CB" box="[281,353,1878,1903]" captionStart="FIGURE 2" captionStartId="6.[151,250,800,822]" captionTargetBox="[159,1432,197,773]" captionTargetId="figure@6.[151,1436,193,779]" captionTargetPageId="6" captionText="FIGURE 2. Ventral view of a Mecodema specimen showing specific morphological structures, excluding taxonomic structures indicated in detail figures, used in the species descriptions. ƑLF = ventrite lateral foveae; ƑSP = ventrite setose punctures; MTC = metacoxa; MTƑP = metaventrite process; MSC = mesocoxa; PC = procoxa; PS = prosternum; G = gena; PES = proepisternum; MSE = mesepisternum; MTE = metepisternum; Ƒ1Ƒ6 = ventrites 1-6 (ventrites 1-3 may be fused); M = midline (dashed line, not a taxonomic structure)." httpUri="https://zenodo.org/record/2668067/files/figure.png" pageId="66" pageNumber="67">Fig. 2</figureCitation>
) setae present:
<specimenCount id="9D14AAB90247B74C9270B53FEF3AF8CB" box="[540,561,1880,1903]" pageId="66" pageNumber="67" type="male"></specimenCount>
with 2 closely-spaced, short setae each side, apical edge squared between punctures,
<specimenCount id="9D14AAB90247B74C917FB51BEC23F837" box="[275,296,1916,1939]" pageId="66" pageNumber="67" type="female"></specimenCount>
with 2 widely spaced, long setae on each side of midline, apical edge a obtusely rounded; ventrites foveate laterally (may be indistinct). Anterior metaventrite process a short triangle with a well-defined carina, apically broad and narrowed posterad.
</paragraph>
<caption id="DF6D31B80246B74D90FBB53EE808F808" httpUri="https://zenodo.org/record/2668135/files/figure.png" pageId="67" pageNumber="68" startId="67.[151,250,1881,1903]" targetBox="[298,1286,200,1844]" targetPageId="67">
<paragraph id="8BAD61300246B74D90FBB53EE808F808" blockId="67.[151,1436,1881,1964]" pageId="67" pageNumber="68">
<emphasis id="B966BD220246B74D90FBB53EEC2AF8CB" bold="true" box="[151,289,1881,1903]" pageId="67" pageNumber="68">FIGURE 36.</emphasis>
<taxonomicName id="4C121AB30246B74D914BB53EEF09F8CB" authorityName="* Broun" authorityYear="1923" box="[295,514,1881,1903]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="longicolle">
<emphasis id="B966BD220246B74D914BB53EEF09F8CB" box="[295,514,1881,1903]" italics="true" pageId="67" pageNumber="68">Mecodema longicolle</emphasis>
</taxonomicName>
plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe.
</paragraph>
</caption>
<paragraph id="8BAD61300241B74A90ABB2FFEDC4FD99" blockId="68.[151,1437,151,1184]" pageId="68" pageNumber="69">
<emphasis id="B966BD220241B74A90ABB2FFEC7FFF14" box="[199,372,152,176]" italics="true" pageId="68" pageNumber="69">Male genitalia:</emphasis>
Apical portion of penis lobe asymmetrically hooked (
<figureCitation id="13297DB50241B74A93BEB2F0E928FF14" box="[978,1059,151,176]" captionStart="FIGURE 36" captionStartId="67.[151,250,1881,1903]" captionTargetBox="[298,1286,200,1844]" captionTargetId="figure@67.[290,1296,193,1860]" captionTargetPageId="67" captionText="FIGURE 36. Mecodema longicolle plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668135/files/figure.png" pageId="68" pageNumber="69">Fig. 36</figureCitation>
PL) with a moderate deflection to right of vertical axis (VV) (
<figureCitation id="13297DB50241B74A91ABB2DBEF27FF71" box="[455,556,188,213]" captionStart="FIGURE 15" captionStartId="20.[151,250,1943,1965]" captionTargetBox="[162,1424,1039,1914]" captionTargetId="figure@20.[162,1424,1038,1921]" captionTargetPageId="20" captionText="FIGURE 15. The penis lobe in ventral view showing the relative deflection to right of vertical: (A) distinctly deflected (M. regulus); (B) moderately deflected (M. haunoho); (C) slightly deflected (M. oregoides). Scale bars = 1 mm. Structures of penis lobe indicated are: base (B); lateral edge (LE); apical portion (AP); ventral edge (ƑE)." httpUri="https://zenodo.org/record/2668093/files/figure.png" pageId="68" pageNumber="69">Fig. 15B</figureCitation>
); apex of penis lobe shaft broadened ventrally curved symmetrically upward to form the dorsal process, a narrowly rounded and recurved hook (
<figureCitation id="13297DB50241B74A9301B2B8EEB4FF5C" box="[877,959,223,248]" captionStart="FIGURE 36" captionStartId="67.[151,250,1881,1903]" captionTargetBox="[298,1286,200,1844]" captionTargetId="figure@67.[290,1296,193,1860]" captionTargetPageId="67" captionText="FIGURE 36. Mecodema longicolle plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668135/files/figure.png" pageId="68" pageNumber="69">Fig. 36</figureCitation>
PL), evenly curved to shaft dorsad; ventral edge of penis lobe shaft straight (VV); penis lobe shaft very narrow, overall length slightly curved ventrally (
<figureCitation id="13297DB50241B74A9503B363EDBFFEE4" captionStart="FIGURE 36" captionStartId="67.[151,250,1881,1903]" captionTargetBox="[298,1286,200,1844]" captionTargetId="figure@67.[290,1296,193,1860]" captionTargetPageId="67" captionText="FIGURE 36. Mecodema longicolle plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668135/files/figure.png" pageId="68" pageNumber="69">Fig. 36</figureCitation>
PL), almost straight (LV). Structures of the endophallus (
<figureCitation id="13297DB50241B74A9350B340EE88FEE4" box="[828,899,295,320]" captionStart="FIGURE 4" captionStartId="7.[151,250,973,995]" captionTargetBox="[354,1232,193,952]" captionTargetId="figure@7.[354,1232,193,952]" captionTargetPageId="7" captionText="FIGURE 4. An Environmental Scanning Electron Microscope (ESEM) photograph of the four main structures of the apical plate of the endophallus within the penis lobe. This is an example from Mecodema haunoho showing the central spicule (c.s), left (l.s.f) and the right setose flange (r.s.f), and microtrichial field (m.f). These structures are variable among species but within a species, they are very consistent, even over distant geographical ranges. The microtrichial field can be covered in flattened scales as in photo above, or with elongated setae-like scales." httpUri="https://zenodo.org/record/2668071/files/figure.png" pageId="68" pageNumber="69">Fig. 4</figureCitation>
): lateral form of the apex of the central spicule pointed; dorsal form of the apex of the central spicule long and narrow; setal (scales) coverage of the apical plate extensive (76100%); size of left setose flange small; size of right setose flange large. Left paramere basal lobe narrow and rectangular without dorsal hump, short curved slope to arm; arm short and narrow (
<figureCitation id="13297DB50241B74A9578B3F3E860FE09" box="[1300,1387,404,429]" captionStart="FIGURE 36" captionStartId="67.[151,250,1881,1903]" captionTargetBox="[298,1286,200,1844]" captionTargetId="figure@67.[290,1296,193,1860]" captionTargetPageId="67" captionText="FIGURE 36. Mecodema longicolle plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668135/files/figure.png" pageId="68" pageNumber="69">Fig. 36</figureCitation>
LP); terminal lobe indistinct from arm with apical tuft of short, very sparse setae, extended along arm only; ventral edge straight (
<figureCitation id="13297DB50241B74A9096B3BBEC40FE51" box="[250,331,476,501]" captionStart="FIGURE 36" captionStartId="67.[151,250,1881,1903]" captionTargetBox="[298,1286,200,1844]" captionTargetId="figure@67.[290,1296,193,1860]" captionTargetPageId="67" captionText="FIGURE 36. Mecodema longicolle plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668135/files/figure.png" pageId="68" pageNumber="69">Fig. 36</figureCitation>
LP). Right paramere a narrowly rectangular for ½ length then gradually narrowed to terminal lobe, double row of setae extended along apical ¾ of ventral edge (
<figureCitation id="13297DB50241B74A933AB398EEA2FDBC" box="[854,937,511,536]" captionStart="FIGURE 36" captionStartId="67.[151,250,1881,1903]" captionTargetBox="[298,1286,200,1844]" captionTargetId="figure@67.[290,1296,193,1860]" captionTargetPageId="67" captionText="FIGURE 36. Mecodema longicolle plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668135/files/figure.png" pageId="68" pageNumber="69">Fig. 36</figureCitation>
RP), apical ⅓ inwardly curved toward penis lobe.
</paragraph>
<paragraph id="8BAD61300241B74A90ABB02FEE24FD0C" blockId="68.[151,1437,151,1184]" pageId="68" pageNumber="69">
<emphasis id="B966BD220241B74A90ABB02FEC9BFDC4" box="[199,400,584,608]" italics="true" pageId="68" pageNumber="69">Female genitalia:</emphasis>
Basal gonocoxite 1 long and narrow the entire length, internal dorso-lateral carina without setae, ventral surface smooth. Gonocoxite 2 (
<figureCitation id="13297DB50241B74A92F9B00BEFD3FD21" box="[661,728,620,645]" captionStart="FIGURE 5" captionStartId="9.[151,250,1714,1736]" captionTargetBox="[167,1436,790,1674]" captionTargetId="figure@9.[151,1436,790,1693]" captionTargetPageId="9" captionText="FIGURE 5. The entire female reproductive tract (ventral view) including taxonomic structures (for all other abbreviated structure names see Seldon et al. 2012). However, for this revision only the following structures are important: HS = helminthoid sclerite; G1 = gonocoxite 1; G2 = gonocoxite 2; TDC = transverse dorsal carina; R = ramus." httpUri="https://zenodo.org/record/2668073/files/figure.png" pageId="68" pageNumber="69">Fig. 5</figureCitation>
) a rounded diamond-shaped, terminal dorsal carina with small undulations and reflexed. Ramus long and relatively broad.
</paragraph>
</subSubSection>
<subSubSection id="C30832BB0241B74A90ABB0D3EC53FCB1" pageId="68" pageNumber="69" type="discussion">
<paragraph id="8BAD61300241B74A90ABB0D3EC53FCB1" blockId="68.[151,1437,151,1184]" pageId="68" pageNumber="69">
<emphasis id="B966BD220241B74A90ABB0D3EC59FD69" bold="true" box="[199,338,692,717]" pageId="68" pageNumber="69">Comments:</emphasis>
Externally
<taxonomicName id="4C121AB30241B74A91BFB0D2EF63FD68" authorityName="* Broun" authorityYear="1923" box="[467,616,692,716]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="longicolle">
<emphasis id="B966BD220241B74A91BFB0D2EF63FD68" box="[467,616,692,716]" italics="true" pageId="68" pageNumber="69">M. longicolle</emphasis>
</taxonomicName>
and
<taxonomicName id="4C121AB30241B74A92F1B0D2EE0DFD68" authorityName="* Britton" authorityYear="1949" box="[669,774,692,716]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="florae">
<emphasis id="B966BD220241B74A92F1B0D2EE0DFD68" box="[669,774,692,716]" italics="true" pageId="68" pageNumber="69">M. florae</emphasis>
</taxonomicName>
look very similar, and both species overlap geographically. However, the male internal aedeagal character are completely different, especially the apical portion (see
<figureCitation id="13297DB50241B74A9500B0B0EDB9FCB0" captionStart="FIGURE 27" captionStartId="46.[151,250,1952,1974]" captionTargetBox="[296,1289,262,1919]" captionTargetId="figure@46.[285,1301,257,1931]" captionTargetPageId="46" captionText="FIGURE 27. Mecodema florae plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668117/files/figure.png" pageId="68" pageNumber="69">Figs 27</figureCitation>
PL and 36PL).
</paragraph>
</subSubSection>
<subSubSection id="C30832BB0241B74A90ABB178E816FC9C" box="[199,1309,799,824]" pageId="68" pageNumber="69" type="distribution">
<paragraph id="8BAD61300241B74A90ABB178E816FC9C" blockId="68.[151,1437,151,1184]" box="[199,1309,799,824]" pageId="68" pageNumber="69">
<emphasis id="B966BD220241B74A90ABB178EC69FC9C" bold="true" box="[199,354,799,824]" pageId="68" pageNumber="69">Distribution:</emphasis>
<collectingCountry id="F30521A00241B74A9105B147EF09FC9C" box="[361,514,799,824]" name="New Zealand" pageId="68" pageNumber="69">New Zealand</collectingCountry>
, North Island, Mt Taranaki, Mt Hikurangi, Ruahine Range, Rangitikei.
</paragraph>
</subSubSection>
<subSubSection id="C30832BB0241B74A90ABB123EEE3FB04" pageId="68" pageNumber="69" type="materials_examined">
<paragraph id="8BAD61300241B74A90ABB123EF53FC01" blockId="68.[151,1437,151,1184]" pageId="68" pageNumber="69">
<materialsCitation id="3B7A6B6D0241B74A90ABB123EF53FC01" ID-GBIF-Occurrence="2237851758" collectingDate="1918-04-06" collectionCode="BMNH" collectorName="Type &amp; New Zeal. &amp; Broun Coll. Brit. Mus. &amp; Coll. Brit. Mus. &amp; Mt Egmont &amp; Hudson &amp; Exposed" pageId="68" pageNumber="69" specimenCount="1" typeStatus="holotype">
<emphasis id="B966BD220241B74A90ABB123EC36FCF9" bold="true" box="[199,317,836,861]" pageId="68" pageNumber="69">
<typeStatus id="54A9DF920241B74A90ABB123EC33FCF9" box="[199,312,836,861]" pageId="68" pageNumber="69" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectionCode id="ED03F9F50241B74A9129B122EC92FCF8" box="[325,409,837,860]" pageId="68" pageNumber="69">BMNH</collectionCode>
male labelled.
<collectorName id="26E704E60241B74A922BB122EF74FCF9" box="[583,639,837,861]" pageId="68" pageNumber="69">Type</collectorName>
[round label with red border] /
<collectorName id="26E704E60241B74A9384B122E954FCF9" box="[1000,1119,836,861]" pageId="68" pageNumber="69">New Zeal.</collectorName>
[red underline]
<collectorName id="26E704E60241B74A957AB122EC01FCDB" pageId="68" pageNumber="69">Broun Coll. Brit. Mus.</collectorName>
1922—482. / 4259 [hw] /
<collectorName id="26E704E60241B74A9254B10FEFB2FC24" box="[568,697,872,896]" pageId="68" pageNumber="69">Mt Egmont</collectorName>
<date id="FFAC47F00241B74A92ACB100EE2EFC24" box="[704,805,871,896]" pageId="68" pageNumber="69" value="1918-04-06">
<collectingDate id="EFE8BE180241B74A92ACB100EE2EFC24" box="[704,805,871,896]" pageId="68" pageNumber="69" value="1918-04-06">6.4.1918</collectingDate>
</date>
. [hw] /
<collectorName id="26E704E60241B74A9310B100EEDDFC24" box="[892,982,871,896]" pageId="68" pageNumber="69">Hudson</collectorName>
, 6 G7. n.sp [hw] /
<taxonomicName id="4C121AB30241B74A94C4B100E897FC24" authorityName="* Broun" authorityYear="1923" box="[1192,1436,871,896]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="longicolle">Mecodema longicolle</taxonomicName>
[hw].
<collectorName id="26E704E60241B74A90B6B1EBEC31FC01" box="[218,314,908,933]" pageId="68" pageNumber="69">Exposed</collectorName>
aedeagus apex damaged.
</materialsCitation>
</paragraph>
<paragraph id="8BAD61300241B74A90ABB1C8EC7FFBB4" blockId="68.[151,1437,151,1184]" pageId="68" pageNumber="69">
<materialsCitation id="3B7A6B6D0241B74A90ABB1C8EC64FBB4" ID-GBIF-Occurrence="2237851673" collectingDate="1919-01-20" collectionCode="BMNH" collectorName="Type &amp; New Zeal. &amp; Broun Coll. Brit. Mus. &amp; Coll. Brit. Mus. &amp; Martinborough" pageId="68" pageNumber="69" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<emphasis id="B966BD220241B74A90ABB1C8EC3FFC6C" bold="true" box="[199,308,943,968]" pageId="68" pageNumber="69">
<typeStatus id="54A9DF920241B74A90ABB1C8EC3FFC6C" box="[199,308,943,968]" pageId="68" pageNumber="69" type="paratype">Paratype</typeStatus>
</emphasis>
(
<taxonomicName id="4C121AB30241B74A9128B1D7EF4EFC6C" box="[324,581,944,968]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="subaeanum">
<emphasis id="B966BD220241B74A9128B1D7EF4EFC6C" box="[324,581,944,968]" italics="true" pageId="68" pageNumber="69">Mecodema subaeanum</emphasis>
</taxonomicName>
):
<collectionCode id="ED03F9F50241B74A9230B1D7EFBBFC63" box="[604,688,944,967]" pageId="68" pageNumber="69">BMNH</collectionCode>
male labelled.
<collectorName id="26E704E60241B74A9337B1D7EE99FC6C" box="[859,914,944,968]" pageId="68" pageNumber="69">Type</collectorName>
[round label with red outer circle] /
<collectorName id="26E704E60241B74A9544B1D7E896FC6C" box="[1320,1437,943,968]" pageId="68" pageNumber="69">New Zeal.</collectorName>
[red underlined]
<collectorName id="26E704E60241B74A9139B1B2EF5DFC48" box="[341,598,980,1005]" pageId="68" pageNumber="69">Broun Coll. Brit. Mus.</collectorName>
1922—482. / 4262
<specimenCount id="9D14AAB90241B74A9355B1B2EE45FC48" box="[825,846,981,1004]" pageId="68" pageNumber="69" type="male"></specimenCount>
[hw] /
<collectorName id="26E704E60241B74A93CFB1B3E947FC49" box="[931,1100,980,1005]" pageId="68" pageNumber="69">Martinborough</collectorName>
<date id="FFAC47F00241B74A9438B1B3E9C3FC48" box="[1108,1224,980,1004]" pageId="68" pageNumber="69" value="1919-01-20">
<collectingDate id="EFE8BE180241B74A9438B1B3E9C3FC48" box="[1108,1224,980,1004]" pageId="68" pageNumber="69" value="1919-01-20">20.1.1919</collectingDate>
</date>
. [hw] /
<taxonomicName id="4C121AB30241B74A9573B1B3EC1EFBB4" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="subaeanum">Mecodema subaeanum</taxonomicName>
<specimenCount id="9D14AAB90241B74A9170B19FEC3AFBAB" box="[284,305,1016,1039]" pageId="68" pageNumber="69" type="male"></specimenCount>
[hw]
</materialsCitation>
.
</paragraph>
<paragraph id="8BAD61300241B74A90ABB67BEEE3FB04" blockId="68.[151,1437,151,1184]" pageId="68" pageNumber="69">
<emphasis id="B966BD220241B74A90ABB67BECBEFB91" bold="true" box="[199,437,1052,1077]" pageId="68" pageNumber="69">Material examined:</emphasis>
<specimenCount id="9D14AAB90241B74A91D2B67BECE8FB90" box="[446,483,1052,1076]" pageId="68" pageNumber="69" type="male">1♂</specimenCount>
,
<collectingCountry id="F30521A00241B74A919EB67AEFCBFB90" box="[498,704,1052,1076]" name="New Zealand" pageId="68" pageNumber="69">NEW ZEALAND</collectingCountry>
511 [hw] G.V. Hudson /
<taxonomicName id="4C121AB30241B74A9389B67BE96AFB91" authorityName="Blanchard" authorityYear="1853" box="[997,1121,1052,1077]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="genus">Mecodema</taxonomicName>
sp [hw] Det. G.E. Bryant /
<taxonomicName id="4C121AB30241B74A90FBB658ECC3FBF3" authority="Br" authorityName="Br" box="[151,456,1087,1112]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="subaeanum">Mecodema subaeanum Br</taxonomicName>
[hw] E.B. Britton det. 1936 [hw] / Genitalia card mounted (BMNH);
<specimenCount id="9D14AAB90241B74A9549B627E843FBFC" box="[1317,1352,1088,1112]" pageId="68" pageNumber="69" type="female">1♀</specimenCount>
,
<collectingCountry id="F30521A00241B74A9530B627EC10FBD8" name="New Zealand" pageId="68" pageNumber="69">NEW ZEALAND</collectingCountry>
RI Ruahine SFP, Rangitane Rd end, Kawhatau Base,
<date id="FFAC47F00241B74A93E2B603E94EFBD8" box="[910,1093,1124,1149]" pageId="68" pageNumber="69" value="1994-02-25" valueMax="1994-03-01" valueMin="1994-02-25">25.II1.III.1994</date>
, Larivière. Larochelle /
<quantity id="4CEACCD50241B74A9530B603E897FBD8" box="[1372,1436,1124,1148]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="68" pageNumber="69" unit="m" value="800.0">800m</quantity>
Very wet, well drained horopito-Melicytus-tree fern for.; rich litter. Pittraps.
</paragraph>
</subSubSection>
</treatment>
</document>